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<document id="85BFBB05BBCC7A935B03FDD18C458ADF" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331808FF94473431D3AFA1EBDF" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba rinha Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="228BE132-A946-439C-986D-0871BF187E7E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="79" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="77" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="195A414359A448C8CCFD2F5AD2A9A3E6">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="F32BC09FA95C3C8BDA6B808F7AB960F4">Griswold, Charles</mods:namePart>
<mods:affiliation id="7857AC1E855404662C8B8D26EDE8A67F">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="C4CB8E4A98DE9D0D39ACC5AEC0A64564">Ubick, Darrell</mods:namePart>
<mods:affiliation id="278E267A82C3372B87301FDDDD996BE6">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="D82B069D2DF285AD6A60E13608B8819F">Ledford, Joel</mods:namePart>
<mods:affiliation id="CB20D117A56702EAD2651889940C348D">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="3FF09E2421F35EECED8FAABE26346FD3">Polotow, Daniele</mods:namePart>
<mods:affiliation id="48CE48A628F0389A35FBDAA92B212FDF">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:date id="CFFA68F50A4082B0CF87DCF3DA96AA32">2022</mods:date>
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<taxonomicName id="B48399A61808FF96473431D3AA3EE9BF" ID-CoL="C8KJZ" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,265,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha" status="sp. nov.">
<emphasis id="41F73E371808FF96473431D3AA3EE9BF" bold="true" box="[123,265,220,243]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1808FF96465831D3AAACE9BF" box="[279,411,220,243]" pageId="76" pageNumber="77" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="3B99B1AE1808FF96473431F3A819E85D" box="[123,814,252,273]" pageId="76" pageNumber="77" type="description">
<paragraph id="733CE2251808FF96473431F3A819E85D" blockId="76.[123,814,220,273]" box="[123,814,252,273]" pageId="76" pageNumber="77">
<figureCitation id="EBB8FEA01808FF96473431F3AA3CE85D" box="[123,267,252,273]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: AE = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="76" pageNumber="77">Figures 6 D, E</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96465A31F3AA0EE85D" box="[277,313,252,273]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route dAnosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="76" pageNumber="77">8 A</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96460B31F3AA42E85D" box="[324,373,252,273]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne dAmbre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route dAnosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="76" pageNumber="77">11 D</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9646CF31F3AA9EE85D" box="[384,425,252,273]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="76" pageNumber="77">29 I</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9646FC31F3A937E85D" box="[435,512,252,273]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">59 AC</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96454531F3A96DE85D" box="[522,602,252,273]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">71 G, H</figureCitation>
,
<figureCitation id="EBB8FEA01808FF96452A31F3A986E85D" box="[613,689,252,273]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="76" pageNumber="77">81 B, E</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9645F431F3A833E85D" box="[699,772,252,273]" captionStart="MAP" captionStartId="177.[155,174,1068,1087]" captionTargetBox="[129,1176,216,1047]" captionTargetId="figure-17@177.[648,1176,216,1059]" captionTargetPageId="177" captionText="MAP 3. Distributions of Uduba rinha and Uduba irwini. MAP 4. Distributions of Uduba madagascariensis, Uduba barbarae, Uduba pseudoevanescens and Uduba orona." figureDoi="http://doi.org/10.5281/zenodo.13158775" httpUri="https://zenodo.org/record/13158775/files/figure.png" pageId="76" pageNumber="77">Maps 3</figureCitation>
, 18.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E4302EAA76E8D4" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="733CE2251808FF9647E4302EAA76E8D4" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E4302EAA67E874" bold="true" box="[171,336,289,312]" pageId="76" pageNumber="77">Type material.</emphasis>
<materialsCitation id="C3EBE8781808FF964618302EAA76E8D4" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CAS" country="Madagascar" elevation="140" location="Parc National de Namoroka" municipality="Fisher-Griswold Arthropod Team" pageId="76" pageNumber="77" specimenCode="CASENT9018660, CASENT9082218" specimenCount="2" stateProvince="Mahajanga" typeStatus="holotype">
<typeStatus id="AC385C871808FF964618302EAA8DE874" box="[343,442,289,312]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
male (
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) and
<typeStatus id="AC385C871808FF964444302EA850E874" box="[779,871,289,312]" pageId="76" pageNumber="77" type="paratype">paratype</typeStatus>
female (
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)
<collectingMethod id="AAC29A321808FF964734304EAAAFE814" box="[123,408,321,344]" pageId="76" pageNumber="77">collected from pitfall traps</collectingMethod>
at
<quantity id="B47B4FC01808FF9646F5304EAAC0E814" box="[442,503,321,344]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="76" pageNumber="77" unit="m" value="140.0">
<elevation id="F8AE05161808FF9646F5304EAAC0E814" box="[442,503,321,344]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="76" pageNumber="77" unit="m" value="140.0">140m</elevation>
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elevation in dry tropical forest at
<location id="765CB4FE1808FF96442B304EAFA6E814" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE1808FF96442B304EAFA6E814" box="[868,1169,321,344]" country="Madagascar" municipality="Fisher-Griswold Arthropod Team" name="Parc National de Namoroka" pageId="76" pageNumber="77" stateProvince="Mahajanga">Parc National de Namoroka</location>
,
<collectingRegion id="B1472CC71808FF964734306EAA52E834" box="[123,357,353,376]" country="Madagascar" name="Mahajanga" pageId="76" pageNumber="77">Mahajanga Province</collectingRegion>
,
<collectingCountry id="0B94A2B51808FF964635306EAACEE834" box="[378,505,353,376]" name="Madagascar" pageId="76" pageNumber="77">Madagascar</collectingCountry>
,
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<collectingDate id="17793D0D1808FF96455F306EA833E834" box="[528,772,353,376]" pageId="76" pageNumber="77" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">48 November 2002</collectingDate>
</date>
,
<collectingMunicipality id="9358785F1808FF964455306EAFA5E834" box="[794,1170,353,376]" pageId="76" pageNumber="77">Fisher-Griswold Arthropod Team</collectingMunicipality>
, deposited in
<collectionCode id="15927AE01808FF964649308EAA0AE8D4" box="[262,317,385,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="76" pageNumber="77" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E430AEA816EBF5" pageId="76" pageNumber="77" type="discussion">
<paragraph id="733CE2251808FF9647E430AEA816EBF5" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E430AEAA24E8F4" bold="true" box="[171,275,417,440]" pageId="76" pageNumber="77">Remarks</emphasis>
. Males of the similar species
<taxonomicName id="B48399A61808FF96451530AEA9DEE8F4" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[602,745,417,440]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96451530AEA9DEE8F4" bold="true" box="[602,745,417,440]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61808FF96446F30AEA8B3E8F4" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[800,900,417,440]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96446F30AEA8B3E8F4" bold="true" box="[800,900,417,440]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
may be distinguished by the TA2 form. Association of the males and females of
<taxonomicName id="B48399A61808FF96459830CDA801E895" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[727,822,450,473]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96459830CDA801E895" bold="true" box="[727,822,450,473]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
is unproblematic in that several were collected together at Parc National de Namoroka. Two similar females, one from Manongarivo (CASENT9006108) and a second from Tsingy de Bemeraha (CASENT9009383), are more difficult to associate with species. Females of
<taxonomicName id="B48399A61808FF96452C332DA9C6EB75" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[611,753,546,569]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96452C332DA9C6EB75" bold="true" box="[611,753,546,569]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
have the epigynal plate posterior of the ML smooth or grooved and the CO hidden beneath the ML hood. The female from Manongarivo (CASENT9006108) that has the CO hidden beneath the ML hood is assigned to species
<taxonomicName id="B48399A61808FF96437E336DAFB8EB35" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1073,1167,610,633]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96437E336DAFB8EB35" bold="true" box="[1073,1167,610,633]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
. The female from Tsingy de Bemeraha (CASENT9009383) that has exposed copulatory openings differs from
<taxonomicName id="B48399A61808FF96464C33ADAA55EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[259,354,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96464C33ADAA55EBF5" bold="true" box="[259,354,674,697]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
and is assigned to
<taxonomicName id="B48399A61808FF96457C33ADA9A1EBF5" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[563,662,674,697]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96457C33ADA9A1EBF5" bold="true" box="[563,662,674,697]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E433CCAA20EA76" pageId="76" pageNumber="77" type="etymology">
<paragraph id="733CE2251808FF9647E433CCAA20EA76" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E433CCAA1DEB96" bold="true" box="[171,298,707,730]" pageId="76" pageNumber="77">Etymology.</emphasis>
The species epithet is a patronym in honor of Mr. HarinHala RinHa, entomologist and explorer-naturalist who conducted most of the fieldwork for the Schlinger
<emphasis id="41F73E371808FF9644B333ECAF1BEBB6" box="[1020,1068,739,762]" italics="true" pageId="76" pageNumber="77">et al</emphasis>
., Malaise Trap Survey of
<collectingCountry id="0B94A2B51808FF96466E320CAA95EA56" box="[289,418,771,794]" name="Madagascar" pageId="76" pageNumber="77">Madagascar</collectingCountry>
arthropods. His efforts have revealed numerous new species of insects and arachnids.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9647E4324CAF4FED77" pageId="76" pageNumber="77" type="diagnosis">
<paragraph id="733CE2251808FF9647E4324CAF4FED77" blockId="76.[122,1174,289,1725]" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E4324CAA8FEA16" bold="true" box="[171,440,835,858]" pageId="76" pageNumber="77">
Diagnosis.
<taxonomicName id="B48399A61808FF964667324CAA8FEA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,440,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF964667324CAA8FEA16" bold="true" box="[296,440,835,858]" italics="true" pageId="76" pageNumber="77">Uduba rinha</emphasis>
</taxonomicName>
</emphasis>
are members of Group IV, the
<taxonomicName id="B48399A61808FF96445D324CA8AEEA16" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[786,921,835,858]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">Uduba rinha</taxonomicName>
group (Map 18). They have a divided cribellum. Males can be distinguished from those of other
<taxonomicName id="B48399A61808FF9644DA326CA8E8EA36" box="[917,991,867,890]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371808FF9644DA326CA8E8EA36" bold="true" box="[917,991,867,890]" italics="true" pageId="76" pageNumber="77">Uduba</emphasis>
</taxonomicName>
except
<taxonomicName id="B48399A61808FF96437C326CAFA2EA36" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1075,1173,867,890]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96437C326CAFA2EA36" bold="true" box="[1075,1173,867,890]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
by the simple, trapezoidal MA (
<figureCitation id="EBB8FEA01808FF96469B328CA953EAD6" box="[468,612,899,922]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">Figs. 59 AC</figureCitation>
<emphasis id="41F73E371808FF96452B328CA95AEAD6" bold="true" box="[612,621,899,922]" pageId="76" pageNumber="77">)</emphasis>
;
<taxonomicName id="B48399A61808FF964532328CA9EBEAD6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[637,732,899,922]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF964532328CA9EBEAD6" bold="true" box="[637,732,899,922]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="B48399A61808FF964340328CAF45EAD6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[1039,1138,899,922]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF964340328CAF45EAD6" bold="true" box="[1039,1138,899,922]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
by having the TA2 a flexible hook or curlicue (
<figureCitation id="EBB8FEA01808FF96451C32ACA985EAF6" box="[595,690,931,954]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="76" pageNumber="77">Fig. 29 I</figureCitation>
) (
<taxonomicName id="B48399A61808FF96458532ACA81BEAF6" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[714,812,931,954]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF96458532ACA81BEAF6" bold="true" box="[714,812,931,954]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
has the TA2 an erect, rectangular spike,
<figureCitation id="EBB8FEA01808FF96478E32CCAA1DEA96" box="[193,298,963,986]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="76" pageNumber="77">Fig. 46 B</figureCitation>
). The females can be distinguished from those of other cribellate
<taxonomicName id="B48399A61808FF9644BD32CCAF0BEA96" box="[1010,1084,963,986]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371808FF9644BD32CCAF0BEA96" bold="true" box="[1010,1084,963,986]" italics="true" pageId="76" pageNumber="77">Uduba</emphasis>
</taxonomicName>
by having the epigynum lateral lobes fused medially (
<figureCitation id="EBB8FEA01808FF96452732EBA9EDEAB7" box="[616,730,996,1019]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">Figs. 71 G</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9645AC32EBA820EAB7" box="[739,791,996,1019]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="76" pageNumber="77">81 B</figureCitation>
). The females of
<taxonomicName id="B48399A61808FF96448332EBAF1EEAB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[972,1065,996,1019]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371808FF96448332EBAF1EEAB7" bold="true" box="[972,1065,996,1019]" italics="true" pageId="76" pageNumber="77">U. rinha</emphasis>
</taxonomicName>
have copulatory openings that are hidden beneath the sides of the ML (
<figureCitation id="EBB8FEA01808FF964416350BA8E0ED57" box="[857,983,1028,1051]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">Figs. 71 G</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9644A8350BAF16ED57" box="[999,1057,1028,1051]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="76" pageNumber="77">81 B</figureCitation>
); the lone female of
<taxonomicName id="B48399A61808FF9647A4352BAA79ED77" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[235,334,1060,1083]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371808FF9647A4352BAA79ED77" bold="true" box="[235,334,1060,1083]" italics="true" pageId="76" pageNumber="77">U. irwini</emphasis>
</taxonomicName>
has the copulatory openings exposed just posteriad of the ML (
<figureCitation id="EBB8FEA01808FF96434E352BAF5BED77" box="[1025,1132,1060,1083]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="76" pageNumber="77">Fig. 71 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1808FF9447E4354BAFA1EB1F" lastPageId="78" lastPageNumber="79" pageId="76" pageNumber="77" type="description">
<paragraph id="733CE2251808FF9747E4354BA96BEA94" blockId="76.[122,1174,289,1725]" lastBlockId="77.[123,1174,220,1725]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<emphasis id="41F73E371808FF9647E4354BAA40ED17" bold="true" box="[171,375,1092,1115]" pageId="76" pageNumber="77">Description. Male</emphasis>
(
<typeStatus id="AC385C871808FF9646C8354BAAC6ED17" box="[391,497,1092,1115]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
): Total length 10.00. Markings as in
<figureCitation id="EBB8FEA01808FF964431354BA8D4ED17" box="[894,995,1092,1115]" captionStart="FIGURE" captionStartId="112.[155,167,1162,1181]" captionTargetBox="[120,1176,216,1143]" captionTargetId="figure-11@112.[120,1176,216,1143]" captionTargetPageId="112" captionText="FIGURE 8. Habitus photos of Uduba spp., males, dorsal views. A. U. rinha, MRAC174.445 from 25 km W Morarano Chrome, Toamasina Province. B. U. pseudoevanescens, Paratype (CASENT9006006) collected at 57 km Route dAnosibe de Moramanga. C. U. evanescens (Dahl, 1901), (CASENT900687) from Ranomafana. D. U. fisheri, (CASENT9002693) from montane rainforest near Anjozorobe. E. U. rajery, Holotype (CASENT9026393) from littoral rainforest in Forêt Ambohidena, Ile Sainte Marie. F. U. valiha, Holotype (CASENT9026394) from Forêt de Vevembe. G. U. rakotozafy, Holotype (CASENT9006024) from forest at Parc National Périnet. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158576" httpUri="https://zenodo.org/record/13158576/files/figure.png" pageId="76" pageNumber="77">Figs. 8 A</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9644A0354BAF11ED17" box="[1007,1062,1092,1115]" captionStart="FIGURE" captionStartId="115.[155,167,1199,1218]" captionTargetBox="[120,1176,216,1180]" captionTargetId="figure-16@115.[120,1176,216,1180]" captionTargetPageId="115" captionText="FIGURE 11. Habitus drawings, Uduba ♂♂, dorsal views. A. U. salegy, Holotype (CASENT9006044) from Parc National Montagne dAmbre. B. U. halabe, Paratype, (CASENT9006017) from Parc National Ranomafana. C. U. pseudoevanescens, Paratype (CASENT9006006) from Route dAnosibe de Moramanga. D. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. E. U. rakotozafy, Holotype (CASENT9006024) from Parc National Périnet. F. U. ibonia, Holotype (CASENT9006038) from 7 km W Ranomafana. G. U. schlingeri (CASENT9006061) from 7 km W Ranomafana. Illustrations by Jenny Speckels. All to same scale, scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158584" httpUri="https://zenodo.org/record/13158584/files/figure.png" pageId="76" pageNumber="77">11 D</figureCitation>
. Carapace 5.50 long, 3.60 wide, 1.70 high; clypeus 0.20 high. Eye diameters: AME 0.16, ALE 0.20, PME 0.22, PLE 0.24. Chelicerae 2.10 long; sternum 2.30 long, 1.80 wide; labium 1.00 long; palpal coxae 1.50 long. Divided cribellum. Spination. (MRAC 174.445): palpusfemur d1-1-1; leg Ifemur d0- 0-2, p0-1-2, r1-1-1-1, patella p1, r1, tibia d1-0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p1-1- 2, v2-2-2, r1-1-1; leg IIfemur d1-1-1, p0-1-0-1, r0-1-0-1, patella p1, tibia d0-1-0, p0-1-1-0, v1-2- 2, r0-1-0-0, metatarsus p1-1-2, v2-2-2, r1-1-1; leg IIIfemur d1-1-1, p1-1-1-1, r1-1-1-1, patella r1, tibia d0-1-0, p0-1-1-0, v1-0-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2-2-2, r1-1-2; leg IVfemur d1-1-1, p0-1-0-1, r0-0-0-1, tibia d0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v1- 1-1-2-1, r1-2-1. Scopulae: cymbium, apicodorsal; tarsi IIV, ventral. Leg measurements (
<typeStatus id="AC385C871808FF96431A346AAB99ECD0" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
): I: 4.50 + 2.00 + 4.30 + 4.30 + 2.60 = 17.70; II: 3.80 + 1.80 + 3.30 + 3.30 + 2.20 = 14.40; III: 3.20 + 1.40 + 2.00 + 2.50 + 1.60 = 10.70; IV: 3.20 + 1.80 + 3.30 + 2.60 + 1.60 = 12.50; palpus: 2.00 + 0.80 + 0.80 + NA + 2.30 = 5.90. Leg formula 1243.
<emphasis id="41F73E371808FF96446934CAA8AEEC90" bold="true" box="[806,921,1477,1500]" pageId="76" pageNumber="77">Male palp</emphasis>
(MRAC 174.445): palpal tibia 0.36 cymbial length, RTA long, curved to ventral, RTA length 0.59 times tibia length, length 2.00 times width, with sharply-pointed apex (
<figureCitation id="EBB8FEA01808FF9645E5370AA826EF50" box="[682,785,1541,1564]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">Fig. 59 C</figureCitation>
), VTA short, conical, blunt (
<figureCitation id="EBB8FEA01808FF964371370AAB87EF71" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">Figs. 59 AC</figureCitation>
), length 1.44 times width, length 0.48 times tibia width; tibia lacking stout spines; tegulum convex, without ridge, tegulum length 1.08 times width; TA1 extends distad of tegulum apex, TA1 extends past TA3 apex by 1.21 times tegulum length; TA3 a transverse hook, TA3 extends apicad of TA2 by 1.175 tegulum length, TA2 small, curlicue-shaped, soft and flexible, extends partially in front of TA3 (
<figureCitation id="EBB8FEA01808FF96465C37A9AAB6EFF1" box="[275,385,1702,1725]" captionStart="FIGURE" captionStartId="129.[155,167,1079,1098]" captionTargetBox="[120,1176,216,1061]" captionTargetId="figure-16@129.[120,1176,216,1061]" captionTargetPageId="129" captionText="FIGURE 29. Uduba spp., color-coded comparisons of male left palpal tarsi, ventral views. A. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. B. U. rakotofrah, paratype (CASENT9062794), from Anosyenne Mts. C. U. madagascariensis (Vinson, 1863) (CASENT9006026) from Ambohimanga. D. U. halabe, holotype (CASENT9006018) from Parc National Ranomafana. E. U. schlingeri, (CASENT9006062) from Parc National Ranomafana. F. U. ibonia, (CASENT9006036) from Parc National Ranomafana. G. U. rakotozafy, holotype (CASENT9006024) from Parc National Périnet. H. U. fisheri, (CASENT9007941) from Anjozorobe. I. U. rinha, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province. Conductor (C), yellow; Median apophysis (MA), blue; TA1, purple; TA2, green; TA3, red. Illustrations by Charles Griswold and Jenny Speckels, not to scale." figureDoi="http://doi.org/10.5281/zenodo.13158631" httpUri="https://zenodo.org/record/13158631/files/figure.png" pageId="76" pageNumber="77">Figs. 29 I</figureCitation>
,
<figureCitation id="EBB8FEA01808FF9646C137A9AAF1EFF1" box="[398,454,1702,1725]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="76" pageNumber="77">59 A</figureCitation>
); MA a broad, flat trapezoid, origin at 0.40 times tegulum length, extends nearly to tegulum apex, tegulum apex at 0.09 tegulum length from MA apex, length (including apical processes) 1.05 times width, MA large, length 0.38 tegulum length, width 0.39 tegulum width (
<figureCitation id="EBB8FEA01809FF9747B73013AABDE87F" box="[248,394,284,307]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="77" pageNumber="78">Figs. 59 A, B</figureCitation>
); conductor fan entire (
<figureCitation id="EBB8FEA01809FF9745C73013A9C6E87F" box="[648,753,284,307]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="77" pageNumber="78">Fig. 59 C</figureCitation>
).
<emphasis id="41F73E371809FF97444A3013A86FE87F" bold="true" box="[773,856,284,307]" pageId="77" pageNumber="78">Female</emphasis>
(
<typeStatus id="AC385C871809FF9744283013A8FDE87F" box="[871,970,284,307]" pageId="77" pageNumber="78" type="paratype">paratype</typeStatus>
): Total length 8.60. Markings as in
<figureCitation id="EBB8FEA01809FF9746533033AAACE81F" box="[284,411,316,339]" captionStart="FIGURE" captionStartId="110.[155,167,951,970]" captionTargetBox="[120,1176,216,932]" captionTargetId="figure-11@110.[120,1176,216,932]" captionTargetPageId="110" captionText="FIGURE 6. Habitus photos of Uduba spp. A. U. valiha, female Paratype (CASENT9006071) from Massif Andringitra, dorsal. B. Uduba andriamihajai, new species, male (CAS9006077) R. S. Ivohibe, dorsal. C. U. madagascariensis (Vinson, 1863), CASENT9006029 male from forest near Ambohimanga, village (CASENT9006029), dorsal. D, E. U. rinha, female (CASENT9006108) from R.S. Manongarivo, dorsal and ventral. F, G. Uduba lamba, Betampona male, (CASENT9042542) dorsal and ventral. Scale bars: AE = 5mm, F, G = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13158570" httpUri="https://zenodo.org/record/13158570/files/figure.png" pageId="77" pageNumber="78">Figs. 6 D, E</figureCitation>
. Carapace 4.80 long, 3.40 wide, 1.90 high; clypeus 0.30 high. Eye diameter: AME 0.14, ALE 0.18, PME 0.20. Chelicera 2.20 long; sternum 2.10 long, 0.80 wide; labium 1.20 long; palpal coxae 1.60 long. Divided cribellum. Spination: palpusfemur d0-1-2, patella p1- 0, tibia p2-0, tarsus p2-1; leg Ifemur d1-0-0, p0-0-2, tibia v2-2-2-2, metatarsus v2-2-3; leg II femur d1-0-0, p0-0-1, tibia p0-1-0, v1-1-2, metatarsus p0-1-0, v2-2-3; leg IIIfemur d1-0-0-1, p1- 0-1, r0-1-0, tibia d1-1-0, p0-1-1-0, v1-1-2, r0-1-1-0, metatarsus d1-0-0-0, p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IVfemur d1-0-0-0, tibia v1-2-2, r1-0-1, metatarsus p0-0-0-2, v2-1-2-2, r0-0-1- 2. Scopulae: ventral on tarsi IIV and apices of metatarsi I and II. Leg measurements: I: 3.50 + 1.80 + 2.80 + 2.40 + 1.90 = 12.40; II: 3.00 + 1.60 + 2.30 + 2.10 + 1.70 = 10.70; III: 2.50 + 1.20 + 1.30 + 1.70 + 1.30 = 8.00; IV: 3.50 + 1.60 + 2.70 + 2.80 + 1.50 = 12.10; palpus: 1.80 + 0.80 + 0.90 + NA + 1.50 = 5.00. Leg formula 1423.
<emphasis id="41F73E371809FF9745313371A80CEBD9" bold="true" box="[638,827,638,661]" pageId="77" pageNumber="78">Female genitalia</emphasis>
(
<typeStatus id="AC385C871809FF9744053371A89AEBD9" box="[842,941,638,661]" pageId="77" pageNumber="78" type="paratype">paratype</typeStatus>
): epigynal plate without depressed atrium, plate width 1.32 times length (
<figureCitation id="EBB8FEA01809FF9745E03390A82EEBFA" box="[687,793,671,694]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Fig. 71 G</figureCitation>
); median lobe and lateral lobes differentiated only in anterior half of plate, distance from posterior margin of ML to epigastric furrow 0.83 times epigynum length; median lobe (
<figureCitation id="EBB8FEA01809FF97450033D0A98DEBBA" box="[591,698,735,758]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="77" pageNumber="78">Fig. 81 B</figureCitation>
), a flat plate with lateral margins extending around to posterior, nearly meeting in middle, broad, ML width between bases of LL 0.59 times epigynum width, ML width 4.20 times LL width at sides of ML; median lobe width 0.38 times epigynum width, ML width 0.45 times ML length, with copulatory openings hidden beneath lateral margins of ML (
<figureCitation id="EBB8FEA01809FF974609326FAA85EA3B" box="[326,434,864,887]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Fig. 71 G</figureCitation>
), distance between copulatory openings 0.56 times epigynum width, CO origin at 0.70 of epigynum length. Vulva (
<figureCitation id="EBB8FEA01809FF974538328FA9D5EADB" box="[631,738,896,919]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Fig. 71 H</figureCitation>
) with spermathecal ducts forming three longitudinal loops, vulva length 0.61 times width, fertilization ducts widely separated, vulva width 2.68 times distance between FD (
<figureCitation id="EBB8FEA01809FF9746A932CEA978EA94" box="[486,591,961,984]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="77" pageNumber="78">Fig. 81 E</figureCitation>
).
</paragraph>
<paragraph id="733CE2251809FF9747E432EEA993EF50" blockId="77.[123,1174,220,1725]" pageId="77" pageNumber="78">
<emphasis id="41F73E371809FF9747E432EEAA6BEAB4" bold="true" box="[171,348,993,1016]" pageId="77" pageNumber="78">Variation. Male</emphasis>
(N= 5): Total length 8.1410.00; carapace length / width = 1.401.54, carapace height / width = 0.420.56, PER / carapace width = 0.360.81, PER / OAL = 2.482.92, PER / AER = 1.311.42, OAL / OQL = 1.071.15, OQP / OQA = 1.101.43, clypeus height / AME = 1.181.56, cheliceral length / clypeus height = 8.0011.60, sternum length / width = 1.281.41, palpal coxa length / width = 2.413.35, femur I length / carapace width = 1.221.26, metatarsus I length / carapace width = 1.171.35, femur IV length / carapace width = 0.891.45, cymbium length / carapace width = 0.560.70, cymbium length / palpal patella length = 2.362.91, cymbium length / palpal tibia length = 2.502.91, cymbium length / palpal femur length = 0.711.15, palpal tibia length / palpal patella length = 0.931.09.
<emphasis id="41F73E371809FF9745ED35ECA9C2EDB6" bold="true" box="[674,757,1251,1274]" pageId="77" pageNumber="78">Female</emphasis>
(N= 3): Total length 8.6010.60; carapace length / width = 1.411.56, carapace height / width = 0.540.62, PER / carapace width = 0.39 0.44, PER / OAL = 3.133.19, PER / AER = 1.321.39, OAL / OQL = 1.051.29, OQP / OQA = 1.121.23, clypeus height / AME = 1.752.50, cheliceral length / clypeus height = 6.507.71, sternum length / width = 1.171.35, palpal coxa length / width = 2.292.38, femur I length / carapace width = 0.971.05, metatarsus I length / carapace width = 0.710.85, femur IV length / carapace width = 0.951.05, cymbium length / carapace width = 0.410.44, palpal tarsus length / palpal patella length = 1.671.88, palpal tarsus length / palpal tibia length = 1.501.67, palpal tarsus length / palpal femur length = 0.811.00, palpal tibia length / palpal patella length = 1.101.13. Female genitalia vary as in
<figureCitation id="EBB8FEA01809FF9746E8370AA909EF50" box="[423,574,1541,1564]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="77" pageNumber="78">Figs. 71 G, H</figureCitation>
,
<figureCitation id="EBB8FEA01809FF974505370AA993EF50" box="[586,676,1541,1564]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="77" pageNumber="78">81 B, E.</figureCitation>
</paragraph>
<paragraph id="733CE2251809FF9447E4372AA9FCE89F" blockId="77.[123,1174,220,1725]" lastBlockId="78.[123,1174,220,659]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
<materialsCitation id="C3EBE8781809FF9747E4372AA96AEF10" collectingDate="1988-10" collectionCode="USNM" collectorName="W. Steiner" country="Madagascar" location="Antananarivo Province" pageId="77" pageNumber="78" specimenCode="CASENT9006040" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
M
<emphasis id="41F73E371809FF97478D372AAABFEF70" bold="true" box="[194,392,1573,1596]" pageId="77" pageNumber="78">aterial examined.</emphasis>
<collectingCountry id="0B94A2B51809FF9746C1372AA967EF70" box="[398,592,1573,1596]" name="Madagascar" pageId="77" pageNumber="78">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371809FF974515372AA865EF70" box="[602,850,1573,1596]" italics="true" pageId="77" pageNumber="78">
<collectingRegion id="B1472CC71809FF974515372AA865EF70" box="[602,850,1573,1596]" country="Madagascar" name="Antananarivo" pageId="77" pageNumber="78">Antananarivo Province</collectingRegion>
</emphasis>
:
<collectingRegion id="B1472CC71809FF974411372AA8C5EF70" box="[862,1010,1573,1596]" country="Madagascar" name="Antananarivo" pageId="77" pageNumber="78">Antananarivo</collectingRegion>
,
<date id="073DC4E51809FF9744B3372AAFA6EF70" box="[1020,1169,1573,1596]" pageId="77" pageNumber="78" value="1988-10">
<collectingDate id="17793D0D1809FF9744B3372AAFA6EF70" box="[1020,1169,1573,1596]" pageId="77" pageNumber="78" value="1988-10">October 1988</collectingDate>
</date>
,
<collectorName id="DE7687F31809FF974734374AABDCEF10" box="[123,235,1605,1628]" pageId="77" pageNumber="78">W. Steiner</collectorName>
(
<specimenCode id="23254A5E1809FF9747B4374AAAF2EF10" box="[251,453,1605,1628]" collectionCode="CASENT" pageId="77" pageNumber="78">CASENT9006040</specimenCode>
,
<specimenCount id="658529AC1809FF97469D374AAAC4EF10" box="[466,499,1605,1628]" count="1" pageId="77" pageNumber="78" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01809FF97454B374AA961EF10" box="[516,598,1605,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="77" pageNumber="78" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781809FF974525374AA83BEFD1" collectingDate="1998-11-08" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="400" latitude="-13.961666" location="Manongarivo" longLatPrecision="21" longitude="48.433334" pageId="77" pageNumber="78" specimenCode="CASENT9006108" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
<emphasis id="41F73E371809FF974525374AA862EF10" box="[618,853,1605,1628]" italics="true" pageId="77" pageNumber="78">
<collectingRegion id="B1472CC71809FF974525374AA862EF10" box="[618,853,1605,1628]" country="Madagascar" name="Antsiranana" pageId="77" pageNumber="78">Antsiranana Province</collectingRegion>
</emphasis>
: R.S.
<location id="765CB4FE1809FF9744D1374AAF07EF10" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE1809FF9744D1374AAF07EF10" box="[926,1072,1605,1628]" country="Madagascar" latitude="-13.961666" longLatPrecision="21" longitude="48.433334" name="Manongarivo" pageId="77" pageNumber="78" stateProvince="Antsiranana">Manongarivo</location>
,
<locationDeviation id="628EFB071809FF974371374AAA52EF30" location="Antanambao" pageId="77" pageNumber="78">
<quantity id="B47B4FC01809FF974371374AAFA2EF10" box="[1086,1173,1605,1628]" metricMagnitude="4" metricUnit="m" metricValue="1.08" pageId="77" pageNumber="78" unit="km" value="10.8">10.8 km</quantity>
229°SW Antanambao
</locationDeviation>
,
<geoCoordinate id="16B784E21809FF97463D376AAADBEF31" box="[370,492,1637,1661]" degrees="13" direction="south" minutes="57" orientation="latitude" pageId="77" pageNumber="78" precision="15" seconds="42" value="-13.961666">13°57ʹ42ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21809FF9746B53769A9B2EF31" box="[506,645,1638,1661]" degrees="048" direction="east" minutes="26" orientation="longitude" pageId="77" pageNumber="78" precision="15" seconds="00" value="48.433334">048°26ʹ00ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01809FF9745853769A83EEF31" box="[714,777,1638,1661]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="77" pageNumber="78" unit="m" value="400.0">
<elevation id="F8AE05161809FF9745853769A83EEF31" box="[714,777,1638,1661]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="77" pageNumber="78" unit="m" value="400.0">400m</elevation>
</quantity>
, sifted litter, rainforest,
<date id="073DC4E51809FF9743403769AB84EFD1" pageId="77" pageNumber="78" value="1998-11-08">
<collectingDate id="17793D0D1809FF9743403769AB84EFD1" pageId="77" pageNumber="78" value="1998-11-08">8 November 1998</collectingDate>
</date>
,
<collectorName id="DE7687F31809FF9747F23789AA0AEFD1" box="[189,317,1670,1693]" pageId="77" pageNumber="78">B. L. Fisher</collectorName>
[BLF1996] (
<specimenCode id="23254A5E1809FF9746863789A9A5EFD1" box="[457,658,1670,1693]" collectionCode="CASENT" pageId="77" pageNumber="78">CASENT9006108</specimenCode>
,
<specimenCount id="658529AC1809FF9745D23789A989EFD1" box="[669,702,1670,1693]" count="1" pageId="77" pageNumber="78" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01809FF9745833789A833EFD1" box="[716,772,1670,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="77" pageNumber="78" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781809FF9444593789A82DE81F" collectingDate="2002-11-04" collectingDateMax="2002-11-08" collectingDateMin="2002-11-04" collectionCode="CAS" collectorName="Fisher &amp; Griswold &amp; Male &amp; Female" country="Madagascar" elevation="140" lastPageId="78" lastPageNumber="79" latitude="-16.466667" location="Parc National de Namoroka" longLatPrecision="21" longitude="45.35" pageId="77" pageNumber="78" specimenCode="CASENT9018660, CASENT9082218, CASENT9065644, CASENT9065645, CASENT9065646, CASENT9065647, CASENT9065648" specimenCount="13" specimenCount-female="2" specimenCount-male="11" stateProvince="Mahajanga" typeStatus="holotype">
<emphasis id="41F73E371809FF9744593789A8C2EFD1" box="[790,1013,1670,1693]" italics="true" pageId="77" pageNumber="78">
<collectingRegion id="B1472CC71809FF9744593789A8C2EFD1" box="[790,1013,1670,1693]" country="Madagascar" name="Mahajanga" pageId="77" pageNumber="78">Mahajanga Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1809FF97434B3789AA39EFF1" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE1809FF97434B3789AA39EFF1" country="Madagascar" latitude="-16.466667" longLatPrecision="21" longitude="45.35" name="Parc National de Namoroka" pageId="77" pageNumber="78" stateProvince="Mahajanga">Parc National de Namoroka</location>
,
<locationDeviation id="628EFB071809FF97465637A9A956EFF1" box="[281,609,1702,1725]" location="Vilanandro" pageId="77" pageNumber="78">
<quantity id="B47B4FC01809FF97465637A9AA53EFF1" box="[281,356,1702,1725]" metricMagnitude="3" metricUnit="m" metricValue="9.8" pageId="77" pageNumber="78" unit="km" value="9.8">9.8 km</quantity>
300°WNW Vilanandro
</locationDeviation>
,
<geoCoordinate id="16B784E21809FF97452437A9A9D2EFF1" box="[619,741,1702,1725]" degrees="16" direction="south" minutes="28" orientation="latitude" pageId="77" pageNumber="78" precision="15" seconds="00" value="-16.466667">16°28ʹ00ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21809FF9745BD37A9A84AEFF1" box="[754,893,1702,1725]" degrees="045" direction="east" minutes="21" orientation="longitude" pageId="77" pageNumber="78" precision="15" seconds="00" value="45.35">045°21ʹ00ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01809FF9744F037A9A8C9EFF1" box="[959,1022,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="77" pageNumber="78" unit="m" value="140.0">
<elevation id="F8AE05161809FF9744F037A9A8C9EFF1" box="[959,1022,1702,1725]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="77" pageNumber="78" unit="m" value="140.0">140m</elevation>
</quantity>
,
<collectingMethod id="AAC29A321809FF97434637A9AF4EEFF1" box="[1033,1145,1702,1725]" pageId="77" pageNumber="78">pitfall trap</collectingMethod>
in tropical dry forest,
<date id="073DC4E5180AFF94461931D3A909E9BF" box="[342,574,220,243]" pageId="78" pageNumber="79" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">
<collectingDate id="17793D0D180AFF94461931D3A909E9BF" box="[342,574,220,243]" pageId="78" pageNumber="79" value="2002-11-04" valueMax="2002-11-08" valueMin="2002-11-04">48 November 2002</collectingDate>
</date>
,
<collectorName id="DE7687F3180AFF94450131D3A9B9E9BF" box="[590,654,220,243]" pageId="78" pageNumber="79">Fisher</collectorName>
,
<collectorName id="DE7687F3180AFF9445EE31D3A833E9BF" box="[673,772,220,243]" pageId="78" pageNumber="79">Griswold</collectorName>
<emphasis id="41F73E37180AFF94445F31D3A87DE9BF" box="[784,842,220,243]" italics="true" pageId="78" pageNumber="79">et al.</emphasis>
[BLF6444], (
<typeStatus id="AC385C87180AFF9444A831D3AF7BE9BF" box="[999,1100,220,243]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
<collectorName id="DE7687F3180AFF94431731D3AFA6E9BF" box="[1112,1169,220,243]" pageId="78" pageNumber="79">Male</collectorName>
,
<specimenCode id="23254A5E180AFF94473431F3AA7AE85F" box="[123,333,252,275]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9018660</specimenCode>
,
<specimenCount id="658529AC180AFF94462031F3AAA6E85F" box="[367,401,252,275]" count="1" pageId="78" pageNumber="79" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9446FC31F3AADCE85F" box="[435,491,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<typeStatus id="AC385C87180AFF94455231F3A9B3E85F" box="[541,644,252,275]" pageId="78" pageNumber="79" type="paratype">Paratype</typeStatus>
<collectorName id="DE7687F3180AFF9445D031F3A9C4E85F" box="[671,755,252,275]" pageId="78" pageNumber="79">Female</collectorName>
,
<specimenCode id="23254A5E180AFF94445A31F3A8D0E85F" box="[789,999,252,275]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9082218</specimenCode>
,
<specimenCount id="658529AC180AFF94434631F3AF1CE85F" box="[1033,1067,252,275]" count="1" pageId="78" pageNumber="79" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0180AFF94430131F3AFBEE85F" box="[1102,1161,252,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9447CD3013AA7BE87F" box="[130,332,284,307]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065644</specimenCode>
,
<specimenCount id="658529AC180AFF9446193013AA40E87F" box="[342,375,284,307]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<specimenCount id="658529AC180AFF9446323013AAA8E87F" box="[381,415,284,307]" count="1" pageId="78" pageNumber="79" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE0180AFF9446E33013AAD4E87F" box="[428,483,284,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9446B23013A9F1E87F" box="[509,710,284,307]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065645</specimenCode>
,
<specimenCount id="658529AC180AFF94459E3013A9C5E87F" box="[721,754,284,307]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9445B03013A801E87F" box="[767,822,284,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF94441F3013AF2DE87F" box="[848,1050,284,307]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065646</specimenCode>
,
<specimenCount id="658529AC180AFF94436C3013AF72E87F" box="[1059,1093,284,307]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF94431D3013AFBEE87F" box="[1106,1161,284,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9447CD3033AA7CE81F" box="[130,331,316,339]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065647</specimenCode>
,
<specimenCount id="658529AC180AFF9446193033AA40E81F" box="[342,375,316,339]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9446CA3033AA8BE81F" box="[389,444,316,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
), (
<specimenCode id="23254A5E180AFF9446993033A997E81F" box="[470,672,316,339]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9065648</specimenCode>
,
<specimenCount id="658529AC180AFF9445E43033A9FBE81F" box="[683,716,316,339]" count="2" pageId="78" pageNumber="79" type="male">2♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9445953033A825E81F" box="[730,786,316,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE878180AFF94446D3033A82EE8DF" collectingDate="2007-10-09" collectingDateMax="2007-10-16" collectingDateMin="2007-10-09" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="20" location="Analamanitra Forest" pageId="78" pageNumber="79" specimenCode="CASENT9042527" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="holotype">
<location id="765CB4FE180AFF94446D3033A8C8E81F" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE180AFF94446D3033A8C8E81F" box="[802,1023,316,339]" country="Madagascar" name="Analamanitra Forest" pageId="78" pageNumber="79" stateProvince="Mahajanga">Analamanitra Forest</location>
,
<locationDeviation id="628EFB07180AFF9443453033ABE6E83F" location="Misinjo" pageId="78" pageNumber="79">
<quantity id="B47B4FC0180AFF9443453033AF79E81F" box="[1034,1102,316,339]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="78" pageNumber="79" unit="km" value="14.0">14 km</quantity>
NE of Misinjo
</locationDeviation>
, 16°8ʹS, 45°42ʹE, elev.
<quantity id="B47B4FC0180AFF9446813053A937E83F" box="[462,512,348,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="78" pageNumber="79" unit="m" value="20.0">
<elevation id="F8AE0516180AFF9446813053A937E83F" box="[462,512,348,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="78" pageNumber="79" unit="m" value="20.0">20m</elevation>
</quantity>
, dense dry forest, malaise,
<date id="073DC4E5180AFF9444673053A8CCE83F" box="[808,1019,348,371]" pageId="78" pageNumber="79" value="2007-10-09" valueMax="2007-10-16" valueMin="2007-10-09">
<collectingDate id="17793D0D180AFF9444673053A8CCE83F" box="[808,1019,348,371]" pageId="78" pageNumber="79" value="2007-10-09" valueMax="2007-10-16" valueMin="2007-10-09">916 October 2007</collectingDate>
</date>
,
<collectorName id="DE7687F3180AFF9443483053AF50E83F" box="[1031,1127,348,371]" pageId="78" pageNumber="79">M. Irwin</collectorName>
and
<collectorName id="DE7687F3180AFF9447343073AA2CE8DF" box="[123,283,380,403]" pageId="78" pageNumber="79">R. HarinHala</collectorName>
[MG-38-04] (
<specimenCode id="23254A5E180AFF94468A3073A9B9E8DF" box="[453,654,380,403]" collectionCode="CASENT" pageId="78" pageNumber="79">CASENT9042527</specimenCode>
,
<specimenCount id="658529AC180AFF9445ED3073A9F4E8DF" box="[674,707,380,403]" count="1" pageId="78" pageNumber="79" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180AFF9445963073A826E8DF" box="[729,785,380,403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE878180AFF9444633073A9F0E89F" collectingDate="1991-10-01" collectingDateMax="1991-10-25" collectingDateMin="1991-10-01" collectionCode="MRAC" collectorName="A. Pauly" country="Madagascar" location="25 km W Morarano Chrome" pageId="78" pageNumber="79" specimenCode="MRAC174.445" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="41F73E37180AFF9444633073AF3BE8DF" box="[812,1036,380,403]" italics="true" pageId="78" pageNumber="79">
<collectingRegion id="B1472CC7180AFF9444633073AF3BE8DF" box="[812,1036,380,403]" country="Madagascar" name="Toamasina" pageId="78" pageNumber="79">Toamasina Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE180AFF94436C3073AA7AE8FF" LSID="urn:lsid:plazi:treatment:FB2A53331808FF94473431D3AFA1EBDF:765CB4FE180AFF94436C3073AA7AE8FF" country="Madagascar" name="25 km W Morarano Chrome" pageId="78" pageNumber="79" stateProvince="Toamasina">
<quantity id="B47B4FC0180AFF94436C3073AF59E8DF" box="[1059,1134,380,403]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="78" pageNumber="79" unit="km" value="25.0">25 km</quantity>
W Morarano Chrome
</location>
, (coordinates approximate) 17º45ʹS, 48º10ʹE, forest, yellow plate,
<date id="073DC4E5180AFF94432F3093AA27E89F" pageId="78" pageNumber="79" value="1991-10-01" valueMax="1991-10-25" valueMin="1991-10-01">
<collectingDate id="17793D0D180AFF94432F3093AA27E89F" pageId="78" pageNumber="79" value="1991-10-01" valueMax="1991-10-25" valueMin="1991-10-01">125 October 1991</collectingDate>
</date>
,
<collectorName id="DE7687F3180AFF94465630B3AA4FE89F" box="[281,376,444,467]" pageId="78" pageNumber="79">A. Pauly</collectorName>
(
<specimenCode id="23254A5E180AFF9446C830B3A907E89F" box="[391,560,444,467]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="78" pageNumber="79" type="Museum">MRAC174.445</specimenCode>
,
<specimenCount id="658529AC180AFF94457330B3A969E89F" box="[572,606,444,467]" count="1" pageId="78" pageNumber="79" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE0180AFF94452330B3A988E89F" box="[620,703,444,467]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="78" pageNumber="79" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE225180AFF9447E430D3AFA1EB1F" blockId="78.[123,1174,220,659]" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E430D3AA56E8BF" bold="true" box="[171,353,476,499]" pageId="78" pageNumber="79">Natural history.</emphasis>
Collections are from dry forest and there is even
<specimenCount id="658529AC180AFF9444C430D3AF17E8BF" box="[907,1056,476,499]" count="1" pageId="78" pageNumber="79" type="generic">one specimen</specimenCount>
labeled as coming from
<collectingRegion id="B1472CC7180AFF94464330F3AA97EB5F" box="[268,416,508,531]" country="Madagascar" name="Antananarivo" pageId="78" pageNumber="79">Antananarivo</collectingRegion>
, although the habitat for this specimen, urban or not, is not specified. Specimens were taken on the forest floor in sifted litter and yellow pan traps and above the floor in a malaise trap.
<taxonomicName id="B48399A6180AFF94467A3333AAF5EB1F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[309,450,572,595]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E37180AFF94467A3333AAF5EB1F" bold="true" box="[309,450,572,595]" italics="true" pageId="78" pageNumber="79">Uduba rinha</emphasis>
</taxonomicName>
are cribellate but we have no records of how they use cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE180AFF9447E43353AFA1EBDF" pageId="78" pageNumber="79" type="distribution">
<paragraph id="733CE225180AFF9447E43353AFA1EBDF" blockId="78.[123,1174,220,659]" pageId="78" pageNumber="79">
<emphasis id="41F73E37180AFF9447E43353AA0AEB3F" bold="true" box="[171,317,604,627]" pageId="78" pageNumber="79">Distribution.</emphasis>
The species
<taxonomicName id="B48399A6180AFF9446843353A96DEB3F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[459,602,604,627]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E37180AFF9446843353A96DEB3F" bold="true" box="[459,602,604,627]" italics="true" pageId="78" pageNumber="79">Uduba rinha</emphasis>
</taxonomicName>
is recorded on the west side of the great escarpment across the northern half of
<collectingCountry id="0B94A2B5180AFF9446D83373A92FEBDF" box="[407,536,636,659]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
on the high plateau and in western dry forests (
<figureCitation id="EBB8FEA0180AFF9443413373AF57EBDF" box="[1038,1120,636,659]" captionStart="MAP" captionStartId="177.[155,174,1068,1087]" captionTargetBox="[129,1176,216,1047]" captionTargetId="figure-17@177.[648,1176,216,1059]" captionTargetPageId="177" captionText="MAP 3. Distributions of Uduba rinha and Uduba irwini. MAP 4. Distributions of Uduba madagascariensis, Uduba barbarae, Uduba pseudoevanescens and Uduba orona." figureDoi="http://doi.org/10.5281/zenodo.13158775" httpUri="https://zenodo.org/record/13158775/files/figure.png" pageId="78" pageNumber="79">Maps 3</figureCitation>
, 18).
</paragraph>
</subSubSection>
</treatment>
</document>