treatments-xml/data/02/9F/BA/029FBA58E0EF5698BCD7A759AB9D7294.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="029FBA58E0EF5698BCD7A759AB9D7294" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila antidomedon Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="F3B6C9CA-F389-46AE-8C49-CA7223A0BE72" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
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<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:F3B6C9CA-F389-46AE-8C49-CA7223A0BE72" httpUri="http://treatment.plazi.org/id/029FBA58E0EF5698BCD7A759AB9D7294" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/F3B6C9CA-F389-46AE-8C49-CA7223A0BE72" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila antidomedon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="antidomedon" status="sp. nov.">
Angustopila antidomedon 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Angustopila antidomedon Pall-Gergely &amp; Hunyadi, sp. nov. A, C-F, I, J holotype (HNHM 105293) B, H paratype (specimen 2). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture on the protoconch (F), aperture (G, H); ventral (I) and frontal (J) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure61" httpUri="https://binary.pensoft.net/fig/808870" pageId="0" pageNumber="1">Fig. 61</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.94 mm, D: 0.83 mm); Thanh 
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province, 
<normalizedToken originalValue="Như">Như</normalizedToken>
Thanh District, Hải 
<normalizedToken originalValue="Vân">Van</normalizedToken>
, Hang 
<normalizedToken originalValue="Lò">Lo</normalizedToken>
Cao 
<normalizedToken originalValue="Kháng">Khang</normalizedToken>
Chiến, vicinity of the cave (locality code: 2020/41); 
<geoCoordinate degrees="19" direction="north" minutes="37.08" orientation="latitude" precision="9" value="19.618">19°37.08'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="34.63" orientation="longitude" precision="9" value="105.57716">105°34.63'E</geoCoordinate>
; 20 m a.s.l.; 14 Feb. 2020; A. Hunyadi leg.; HNHM 105294.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 1 imaged + 8 other shells; same data as for holotype; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized, conical 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a narrow umbilicus, a parietal tooth and a blunt but strong palatal tooth situated at the lower part of the palatal side.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell of normal size for the genus, higher than wide; off-white, conical; body whorl widest from standard apertural view; protoconch consists of 1.25 whorls, with weak spiral striation preceding the first teleoconch whorl; teleoconch with rather strong, irregularly spaced radial growth lines and equally strong, equidistantly and sparsely-arranged spiral striae (ca. 12-19 on body whorl from standard apertural view); whorls 4.5 or slightly less, rounded or very slightly depressed-rounded in form; aperture oblique to shell axis from lateral view; umbilicus narrow; aperture subovoid or reniform with very wide, not distinctly separated sinulus; peristome expanded, not reflected; parietal callus slightly protruding, somewhat detached from penultimate whorl; parietal tooth straight, strong, normally high and long, situated inside shell at some distance from peristome edge; lower palatal tooth, blunt but strong; approximately half of the available shells has a tiny subcolumellar tooth.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.86-0.94; D = 0.82-0.88, H/D*100 = 103.6-113.3 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4), RUD = 24.2-25.0 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila pustulata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pustulata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila pustulata</emphasis>
</taxonomicName>
sp. nov. is generally larger (wider), although the size ranges slightly overlap (D of 
<taxonomicName lsidName="A. antidomedon" pageId="0" pageNumber="1" rank="species" species="antidomedon">
<emphasis italics="true" pageId="0" pageNumber="1">A. antidomedon</emphasis>
</taxonomicName>
sp. nov.: 0.82-0.88 mm, D of 
<taxonomicName lsidName="A. pustulata" pageId="0" pageNumber="1" rank="species" species="pustulata">
<emphasis italics="true" pageId="0" pageNumber="1">A. pustulata</emphasis>
</taxonomicName>
sp. nov.: 0.87-0.93 mm), it has a lower spire (Fig. 
<figureCitation captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Angustopila pustulata Pall-Gergely &amp; Hunyadi, sp. nov. A paratype B-I holotype (HNHM 105312). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture of the protoconch showing protoconch-teleoconch boundary (F); aperture (G); ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure77" httpUri="https://binary.pensoft.net/fig/808886" pageId="0" pageNumber="1">77A</figureCitation>
is a rare example, nearly all shells have a low spire, similar to the holotype), and its palatal tooth is much lower (weaker). Moreover, some shells of 
<taxonomicName lsidName="A. antidomedon" pageId="0" pageNumber="1" rank="species" species="antidomedon">
<emphasis italics="true" pageId="0" pageNumber="1">A. antidomedon</emphasis>
</taxonomicName>
sp. nov. possess a small columellar tooth. 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila occidentalis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="occidentalis">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila occidentalis</emphasis>
</taxonomicName>
sp. nov. is larger, has a lower spire and a comparatively larger aperture with usually weaker palatal tooth. 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila uvula" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="uvula">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila uvula</emphasis>
</taxonomicName>
sp. nov. has a concave-conical shell and a stronger parietal lamella. See also under 
<taxonomicName lsidName="A. bathyodon" pageId="0" pageNumber="1" rank="species" species="bathyodon">
<emphasis italics="true" pageId="0" pageNumber="1">A. bathyodon</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="A. gracilis" pageId="0" pageNumber="1" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">A. gracilis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure61" httpUri="https://binary.pensoft.net/fig/808870" pageId="0" pageNumber="1" start="Figure 61" startId="F61">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 61.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila antidomedon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="antidomedon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila antidomedon</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A, C-F, I, J</emphasis>
holotype (HNHM 105293) 
<emphasis bold="true" pageId="0" pageNumber="1">B, H</emphasis>
paratype (specimen 2). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), aperture (
<emphasis bold="true" pageId="0" pageNumber="1">G, H</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
From the Greek 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ἀντιδομή">ἀντιδομή</normalizedToken>
</emphasis>
(= opposed) and 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὀδούς">ὀδούς</normalizedToken>
</emphasis>
(= tooth) referring to the position of the palatal tooth in relation to the parietal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from the type locality only (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure27" httpUri="https://binary.pensoft.net/fig/808836" pageId="0" pageNumber="1">27</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>