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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.410.7156" ID-GBIF-Dataset="2949043f-90ae-45b1-934d-7bb69b00ecd4" ID-PMC="PMC4042711" ID-Pensoft-Pub="1313-2970-410-121" ID-PubMed="24899853" ID-ZBK="148429B0C4774B3CB24ADDC55BD2769D" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-410-121" ModsDocOrigin="ZooKeys 410" ModsDocTitle="Four new Mouse Spider species (Araneae, Mygalomorphae, Actinopodidae, Missulena) from Western Australia" checkinTime="1451245866751" checkinUser="pensoft" docAuthor="Miglio, Laura Tavares, Harms, Danilo, Framenau, Volker Wilhelm &amp; Harvey, Mark Stephen" docDate="2014" docId="0283886FB81A37A21D92F4EE38F96EC0" docLanguage="en" docName="ZooKeys 410: 121-148" docOrigin="ZooKeys 410" docSource="http://dx.doi.org/10.3897/zookeys.410.7156" docTitle="Missulena melissae Miglio, Harms, Framenau &amp; Harvey, 2014, sp. n." docType="treatment" docUuid="ABC49948-F3B9-4F6C-9C86-67B18B20605A" docUuidSource="ZooBank" docVersion="4" lastPageNumber="127" masterDocId="FFABFFC79500FFFEFFA55244B422FFBF" masterDocTitle="Four new Mouse Spider species (Araneae, Mygalomorphae, Actinopodidae, Missulena) from Western Australia" masterLastPageNumber="148" masterPageNumber="121" pageNumber="124" updateTime="1668158506528" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Four new Mouse Spider species (Araneae, Mygalomorphae, Actinopodidae, Missulena) from Western Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miglio, Laura Tavares</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harms, Danilo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Framenau, Volker Wilhelm</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark Stephen</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>410</mods:number>
</mods:detail>
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<mods:start>121</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.410.7156</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.410.7156</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-410-121</mods:identifier>
<mods:identifier type="ZBK">148429B0C4774B3CB24ADDC55BD2769D</mods:identifier>
<mods:identifier type="ZooBank">148429B0C4774B3CB24ADDC55BD2769D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152053094" LSID="urn:lsid:zoobank.org:act:ABC49948-F3B9-4F6C-9C86-67B18B20605A" httpUri="http://treatment.plazi.org/id/0283886FB81A37A21D92F4EE38F96EC0" lastPageId="6" lastPageNumber="127" pageId="3" pageNumber="124">
<subSubSection pageId="3" pageNumber="124" type="nomenclature">
<paragraph pageId="3" pageNumber="124">
<taxonomicName LSID="http://zoobank.org/ABC49948-F3B9-4F6C-9C86-67B18B20605A" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena melissae" order="Araneae" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="melissae">Missulena melissae</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="124">sp. n.</taxonomicNameLabel>
Figs 1A, 2
<normalizedToken originalValue="AJ">A-J</normalizedToken>
, 3
<normalizedToken originalValue="AJ">A-J</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124" type="type material">
<paragraph pageId="3" pageNumber="124">Type material.</paragraph>
<paragraph lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124">
AUSTRALIA:
<pageBreakToken pageId="4" pageNumber="125" start="start">Western</pageBreakToken>
Australia: holotype male, Millstream-Chichester National Park, 6 km N. of Millstream Homestead, site PW11,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.540222">21°32'24.8&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="117.05701">117°03'25.2&quot;E</geoCoordinate>
, 15 July 200311 October 2004, wet pitfall trap, Department of CALM staff (WAM T97323). Paratypes: 2 males, same data as holotype, except Corunna Downs, 52.5 km N. of Nullagine, Pilbara Biological Survey site NW11,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.407694">21°24'27.7&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="120.071304">120°04'16.7&quot;E</geoCoordinate>
, 3 August 200320 October 2004, Department of CALM staff (WAM T120931).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="125" type="etymology">
<paragraph pageId="4" pageNumber="125">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="125">
The specific epithet is a patronym in honour of Melissa Thomas, the third
<normalizedToken originalValue="authors">author's</normalizedToken>
partner, for her continuing support of late-night arachnological endeavours.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="125" type="diagnosis">
<paragraph pageId="4" pageNumber="125">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="125">
Males of
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena melissae" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="melissae">Missulena melissae</taxonomicName>
sp. n. differ from the two other species with a brown body colour, strongly-sclerotised rastellum with thick spines, smooth and glabrous chelicerae with prominent horizontal ridges and short claws (i.e.
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena faulderi" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="faulderi">Missulena faulderi</taxonomicName>
and
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena rutraspina" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="rutraspina">Missulena rutraspina</taxonomicName>
) by details of the bulb and somatic morphology: embolus short and with a distal tooth (exceeding length of the bulb and without distal tooth in
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena faulderi" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="faulderi">Missulena faulderi</taxonomicName>
), carapace length ca. 4 cm (3.5 cm in
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena rutraspina" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="rutraspina">Missulena rutraspina</taxonomicName>
), patella I-III with rasps (patella III only in
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena faulderi" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="faulderi">Missulena faulderi</taxonomicName>
), rastellum on a low mound (mound distinct in
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena faulderi" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="faulderi">Missulena faulderi</taxonomicName>
), inner row of cheliceral teeth divided (first six teeth fused in
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena faulderi" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="faulderi">Missulena faulderi</taxonomicName>
), and pedipalp patella and tibia swollen (much more slender in
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena faulderi" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="faulderi">Missulena faulderi</taxonomicName>
). Males of
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena rutraspina" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="rutraspina">Missulena rutraspina</taxonomicName>
differ in having a simple embolus tip without processes, a straight embolus, rasps on patella III only, rastellum on a distinct mound, inner row of cheliceral teeth with six spaced teeth, and pedipalp tibia and patella more slender. The female of
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena melissae" order="Araneae" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="melissae">Missulena melissae</taxonomicName>
sp. n. is unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="127" pageId="5" pageNumber="126" type="description">
<paragraph pageId="5" pageNumber="126">
<pageBreakToken pageId="5" pageNumber="126" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="5" pageNumber="126">Adult male, based on holotype (WAM T97323). Medium-sized mygalomorph spider (total length 8.00).</paragraph>
<paragraph pageId="5" pageNumber="126">
Colour: carapace (Fig. 1A) dark reddish-brown, margins dark brown; eye region (Fig. 2B) dark brown, anterior median eyes on black tubercle; chelicerae (Fig. 3I) dark reddish-brown, fangs reddish-brown; abdomen (Fig. 1A) pale brown with a grey metallic spot in dorsal region; sternum (Fig. 3H) yellowish-brown, margins contoured dark yellowish-brown, sigillae yellowish-brown; labium (Fig. 3G) and maxillae (Fig. 3C) dark reddish-brown; legs (Figs 1A, 3
<normalizedToken originalValue="AB">A-B</normalizedToken>
) yellowish brown, tarsi and metatarsi ventrally yellow; spinnerets (Fig. 3J) pale gray, spigots white.
</paragraph>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 1. Dorsal habitus of species described in this paper: A
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena melissae" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="melissae">Missulena melissae</taxonomicName>
sp. n., holotype male from Millstream-Chichester National Park (WAM T97323) B
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena pinguipes" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="pinguipes">Missulena pinguipes</taxonomicName>
sp. n., holotype male from Digger Rocks (WAM T92331) C
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena leniae" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="leniae">Missulena leniae</taxonomicName>
sp. n., holotype male from Nanga Station, Shark Bay (WAM T96784) D
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena mainae" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="mainae">Missulena mainae</taxonomicName>
sp. n., holotype male from Quobba Station, Cape Cuvier (WAM T96782).
</paragraph>
</caption>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 2.
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena melissae" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="melissae">Missulena melissae</taxonomicName>
sp. n., holotype male (WAM T97323): A carapace, lateral view B eye group, dorsal view C carapace, dorsal view D rastellum, dorsal view E same, ventral view F pedipalp, proventral view G bulb and embolus, retrolateral view H embolus, ventral view I bulb and embolus, prolateral view J same, ventral view. Arrows: (EL) embolar lamella, (DET) distal embolar tooth, and (BEI) basal embolar intumescence.
</paragraph>
</caption>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 3.
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena melissae" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="melissae">Missulena melissae</taxonomicName>
sp. n., holotype male (WAM T97323): A patella II, dorsal view B patella III, dorsal view C coxae and labium of pedipalp, ventral view D coxae of pedipalp, ventral view E cheliceral groove, retroventral view F abdomen, dorsal view G labium, ventral view H sternum and sigillae, ventral view I chelicerae, dorsal view J spinnerets, ventral view.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="126">
Carapace: 3.81 long, 4.0 wide; clypeus 0.13; fovea 2.18; caput and eye region (Fig. 2
<normalizedToken originalValue="AC">A-C</normalizedToken>
) laterally elevated, strongly arched; fovea (Fig. 2A, C) very deep, wide and strongly procurved, medially extending as triangular depression, pars cephalica smooth around the eyes and between the eyes and fovea, pars thoracica rugose with bands of fine, random fissures centered around fovea (Fig. 2C).
</paragraph>
<paragraph pageId="5" pageNumber="126">Eyes: OQ 3.61 times wider than long, occupying 1.12 of cephalic width; OAW 3.54; OAL 2.68; IPF 0.40; width of anterior eye group 2.95, with of posterior group 2.36, OQ length 0.81; PME 0.19; PLE 0.22; ALE 0.26; AME 0.22, AME on tubercle, 0.27 long, 0.62 wide; AME inter-distance 0.65; AME to ALE 1; AME to PME 0.40; PLE to ALE 0.55; PLE to PME 0.41; PME inter-distance 1.41; PME to ALE 0.55; eye region (Fig. 2B) with reduced setation although some setae present anterior to AME and between posterior eyes and fovea.</paragraph>
<paragraph pageId="5" pageNumber="126">Chelicerae: 2.95 long, 1.54 wide; distally broad, diagonal, slightly conical; edges smoothly rounded; with weak transverse ridges which distally extend over entire length (Fig. 3I), without setae in area of transverse ridges but with ca. 60 short setae along inner margin of chelicera; rastellum developed, slightly pronounced, consisting of a sclerotised process with 3 strong conical spines and 1214 disordered setae (Fig. 3I), 1213 long setae extend forward from anterior margin of each chelicera and cover base of fang, setae largest on latero-ventral side; inner margin of cheliceral furrow with 3 rows of teeth (Fig. 3E); prolateral (inner) row with ca. 12 teeth, 3 proximal teeth fused together and the rest spaced; intermediate row with 9 proximal, spaced teeth; retrolateral (outer) row with 3 proximal, spaced teeth.</paragraph>
<paragraph pageId="5" pageNumber="126">Maxillae: 1.86 long; 1.31 wide, longer than wide (Fig. 3C, D), with ca. 57 pointed cuspules along entire anterior margin, distally pointed and extended into a prominent heel.</paragraph>
<paragraph pageId="5" pageNumber="126">Labium: 1.04 long, 0.77 wide; conical, ca. 20 pointed cuspules anteriorly (Fig. 3G); labiosternal suture developed as a shallow groove; a pair of sigilla near labiosternal suture (Fig. 3H), developed as irregular, poorly-defined patches.</paragraph>
<paragraph pageId="5" pageNumber="126">Sternum: 2.27 long, 2.09 wide; oval and rebordered (Fig. 3H), with prominent setae, arranged irregularly but denser lateral to labium; 4 pairs of sigillae, anterior and second pair (anterior-posterior) smallest and poorly defined, third pair bigger than 2 anterior pairs and poorly defined, and posterior pair bigger than all others, roughly oval and well defined, 3 posterior sigillae slightly depressed.</paragraph>
<paragraph pageId="5" pageNumber="126">Abdomen: 4 long, 3.54 wide; roughly oval (Fig. 3F); 4 spinnerets (Fig. 3J), PLS 0.72 long, 0.45 wide, apical segment domed; PMS 0.40 long, 0.13 wide.</paragraph>
<paragraph pageId="6" pageNumber="127">
<pageBreakToken pageId="6" pageNumber="127" start="start">Pedipalp</pageBreakToken>
: length of trochanter 0.85, femur 2.11, patella 0.33, tibia 0.66, tarsus 0.18; entire palp is aspinose, femur more than 3 times longer than tibia, tarsus terminally blunt (Fig. 2F); bulb pyriform and rather stout than globular (Fig. 2G, IJ), 2 strongly sclerotised sections connected by a velar median structure (
<normalizedToken originalValue="“haematodocha”">&quot;haematodocha&quot;</normalizedToken>
, Fig. 2G, I); bulb strongly twisted proventrally (Fig. 2I); embolus short, with an intumescence in proximal region (BEI), a strong curvature in the duct in prolateral view, tapering and slightly twisted medially (Fig. 2I, J); embolus tip triangular, with a lamella (EL) well-developed and a prominent tooth (DET) in all views (Fig. 2
<normalizedToken originalValue="GJ">G-J</normalizedToken>
).
</paragraph>
<paragraph pageId="6" pageNumber="127">Legs: with few brown setae, ventral setae of tibiae and metatarsi generally much longer and thicker than dorsal setae and bent towards the exterior; dorsal, lateral and ventral setae of tibiae and metatarsi longer than the diameter of respective segment; preening comb distal in tarsi, very small and plain; metatarsi and tarsi I and II ascopulate, metatarsi and tarsi III and IV densely scopulate but in metatarsi, the length of scopula reaches only 80% of the segment length. Metatarsi I and II with ca. 23, 33 fine ventral setae distally, respectively. Leg measurements: Leg I: femur 2.37, patella 2.00, tibia 2.00, metatarsus 2.00, tarsus 1.00, total 9.37. Leg II: 2.25, 1.87, 1.62, 2.12, 1.00, 8.87. Leg III: 2.5, 1.62, 1.5, 2.37, 1.00, 9.00. Leg IV: 3.25, 1.62, 2.00, 2.25, 1.37, 10.5. Formula 4123.</paragraph>
<paragraph pageId="6" pageNumber="127">
Trichobothria: arranged in discontinuous rows; tibiae
<normalizedToken originalValue="III">I-II</normalizedToken>
with 2 rows of 2 in retrodorsal and prodorsal position, respectively; tibiae III with 2 rows of 2 in retrolatero-dorsal and proximo-prodorsal position, respectively; tibiae IV with 2 rows, the first row with 2 in retrolatero-dorsal and the second row with 3 in proximolateral position; metatarsi with 5 in proximo-dorsal position, tarsi I+II with 4 and 6 medio-dorsally, respectively, III+IV with 5 and 6 medio-dorsally, respectively, all trichobothria in medio-dorsal position.
</paragraph>
<paragraph pageId="6" pageNumber="127">Leg spination: pedipalp aspinose; leg I: tibia rv000, v337, pv120, d000; metatarsus rv211, v235, pv000, d000; tarsus rv143, v273, pv222, d000; leg II: tibia rv000, v000, pv010, d000; metatarsus rv000, v000, pv000, d000; tarsus rv354, v122, pv132, d000; leg III: tibia rv000, v032, pv222, d213; metatarsus rv223, v000, pv334, d842; tarsus rv354, v001, pv134, d022; leg IV: tibia rv020, v244, pv112, d300; metatarsus rv132, v000, pv134, d001; tarsus rv4913, v001, pv146, d002; patellae I and II with ca. 53 and 35 rasps, in 8 and 6 oblique rows prolatero-dorsally, respectively; patella III with ca. 59 rasps widespread in dorsal view (Fig. 4B); patella IV with 19 rasps, in 8 and 6 oblique rows prolatero-dorsally, median rows shorter than lateral rows and with less spines, distal spines forming an interrupted crown of spines in the border of the article (Fig. 4B).</paragraph>
<paragraph pageId="6" pageNumber="127">Tarsal claws: leg I: 54/1; leg II: 66/1; leg III: 54/1; leg IV: 33/1; claws slightly shorter than spines of tarsi.</paragraph>
<paragraph pageId="6" pageNumber="127">Variation in paratypes (N=2): total length 6.727.36; carapace 3.45 long, 3.904.36 wide; number of labial cuspules 1416, maxillary cuspules 4867; rastellum with 14 thick and conical spines.</paragraph>
<caption pageId="6" pageNumber="127">
<paragraph pageId="6" pageNumber="127">
Figure 4. Distribution records of
<taxonomicName class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Missulena</taxonomicName>
spp. in Western Australia.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="6" pageNumber="127" type="distribution">
<paragraph pageId="6" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="127">This species is known from the type locality Millstream-Chichester National Park and Corunna Downs in the Pilbara biogeographic region of Western Australia (Fig. 4).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="127" type="habitat">
<paragraph pageId="6" pageNumber="127">Habitat.</paragraph>
<paragraph pageId="6" pageNumber="127">
All specimens were collected in pitfall traps. The collecting sites were dominated by
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
spp., with one site having a eucalypt over-storey (
<bibRefCitation author="McKenzie, NL" journalOrPublisher="Records of the Western Australian Museum, Supplement" pageId="17" pageNumber="138" pagination="3 - 89" title="Introduction to the Pilbara Biodiversity Survey, 2002 2007." volume="78" year="2009">McKenzie et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>