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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.53.5079" ID-PMC="PMC4547025" ID-Pensoft-Pub="1314-2003-53-83" ID-Pensoft-UUID="FF98DD26FF8AD752B817FF81FFE4AC22" ID-PubMed="26312042" ID-Zenodo-Dep="576292" ModsDocID="1314-2003-53-83" checkinTime="1451251407221" checkinUser="pensoft" docAuthor="Rodda, Michele" docDate="2015" docId="027C170A926EE486C2F4EA9FF7DFFD3D" docLanguage="en" docName="PhytoKeys 53: 83-93" docOrigin="PhytoKeys 53" docPubDate="2015-07-21" docSource="http://dx.doi.org/10.3897/phytokeys.53.5079" docTitle="Hoya ruthiae Rodda 2015, sp. nov." docType="treatment" docVersion="6" id="FF98DD26FF8AD752B817FF81FFE4AC22" lastPageNumber="84" masterDocId="FF98DD26FF8AD752B817FF81FFE4AC22" masterDocTitle="Two new species of Hoya R. Br. (Apocynaceae, Asclepiadoideae) from Borneo" masterLastPageNumber="93" masterPageNumber="83" pageNumber="84" updateTime="1668141425532" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of Hoya R. Br. (Apocynaceae, Asclepiadoideae) from Borneo</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rodda, Michele</mods:namePart>
<mods:affiliation>The Herbarium, Singapore Botanic Gardens, 1 Cluny Road, 259569 Singapore</mods:affiliation>
<mods:nameIdentifier type="email">rodda.michele@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-07-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>83</mods:start>
<mods:end>93</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.53.5079</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.53.5079</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-53-83</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF98DD26FF8AD752B817FF81FFE4AC22</mods:identifier>
<mods:identifier type="Zenodo-Dep">576292</mods:identifier>
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<treatment ID-GBIF-Taxon="152008562" LSID="urn:lsid:plazi:treatment:027C170A926EE486C2F4EA9FF7DFFD3D" httpUri="http://treatment.plazi.org/id/027C170A926EE486C2F4EA9FF7DFFD3D" lastPageNumber="84" pageId="1" pageNumber="84">
<subSubSection pageId="1" pageNumber="84" type="nomenclature">
<paragraph pageId="1" pageNumber="84">
<taxonomicName LSID="027C170A-926E-E486-C2F4-EA9FF7DFFD3D" authority="Rodda" authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae" status="sp. nov.">Hoya ruthiae Rodda</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="84">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya ruthiae photographed from Rodda M. MR 606 (SING) prior to pressing A Flower, lateral view with two corolla lobes removed B Corolla, underneath C Corolla and corona, top view D Corona, underneath E Pedicel, calyx and ovaries F Pollinarium with twin pollinia. (Photographs by M. Rodda)" figureDoi="10.3897/phytokeys.53.5079.figure1" httpUri="https://binary.pensoft.net/fig/47618" pageId="1" pageNumber="84">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya ruthiae photographed from Rodda M. MR 606 (SING) prior to pressing A Inflorescence D, E, F, G Two leaves (D, F adaxial surface E, G abaxial surface). Hoya uncinata photographed from Rodda M MR 607 (SING) prior to pressing B Flower, lateral view C Flower, top view. (Photographs by M. Rodda)" figureDoi="10.3897/phytokeys.53.5079.figure2" httpUri="https://binary.pensoft.net/fig/47619" pageId="1" pageNumber="84">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="diagnostic characters">
<paragraph pageId="1" pageNumber="84">Diagnostic characters.</paragraph>
<paragraph pageId="1" pageNumber="84">
Similar to
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
Teijsm. and Binn as both species have clear exudate, deeply lobed rotate corolla and corpusculum of the pollinarium almost as large as the pollinium. The flattened corolla is smaller (1.5-1.7 cm in diameter) with ovate lobes in
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
, while the corolla of
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
is 1.8-2.2 cm in diameter with narrowly lanceolate lobes.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="type">
<paragraph pageId="1" pageNumber="84">Type.</paragraph>
<paragraph pageId="1" pageNumber="84">
Malaysia, Sabah, Lahad Datu, Bukit Baturong, on limestone, 7 July 2000,
<emphasis italics="true" pageId="1" pageNumber="84">Kiew R RK5029</emphasis>
(SING, holotype; barcode SING0077484).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="description">
<paragraph pageId="1" pageNumber="84">Description.</paragraph>
<paragraph pageId="1" pageNumber="84">
Delicate lithophytic climber with clear exudate in all vegetative parts; all vegetative parts glabrous. Leafy
<emphasis italics="true" pageId="1" pageNumber="84">stems</emphasis>
cylindrical, slender, up to 4 mm in diameter, dark brown or grey, with membranaceous peeling bark; internodes 5-15 cm long.
<emphasis italics="true" pageId="1" pageNumber="84">Petioles</emphasis>
terete, fleshy, 5-15
<normalizedToken originalValue="×">x</normalizedToken>
1.5-3 mm in diameter,
<emphasis italics="true" pageId="1" pageNumber="84">lamina</emphasis>
lanceolate, fleshy, (5-)7-15
<normalizedToken originalValue="×">x</normalizedToken>
1.5-3 cm, apex acute-acuminate, base cuneate, light green above turning red in bright light with numerous grey spots, lighter green underneath; penninerved, secondary veins obscure.
<emphasis italics="true" pageId="1" pageNumber="84">Inflorescences</emphasis>
pseudo-umbelliform or globular, 3-4 cm in diameter, 4-20 flowered;
<emphasis italics="true" pageId="1" pageNumber="84">peduncles</emphasis>
persistent, extra-axillary, terete, 1-3 cm
<normalizedToken originalValue="×">x</normalizedToken>
1.5-2.5 mm in diameter, glabrous;
<emphasis italics="true" pageId="1" pageNumber="84">pedicels</emphasis>
10-15 mm
<normalizedToken originalValue="×">x</normalizedToken>
0.5-0.8 mm in diameter, glabrous.
<emphasis italics="true" pageId="1" pageNumber="84">Buds</emphasis>
conical with a 5-ridged base, ca. 10
<normalizedToken originalValue="×">x</normalizedToken>
6 mm.
<emphasis italics="true" pageId="1" pageNumber="84">Calyx</emphasis>
lobes triangular, white-pink 1.3-1.5
<normalizedToken originalValue="×">x</normalizedToken>
0.7-1 mm, apex rounded, glabrous; basal
<emphasis italics="true" pageId="1" pageNumber="84">colleters</emphasis>
1 in each sepal sinus, ovoid.
<emphasis italics="true" pageId="1" pageNumber="84">Corolla</emphasis>
rotate, deeply lobed, 1-1.5 cm in diameter, 1.8-2.2 cm when flattened, white tinged pink, thinly and minutely pubescent inside, outside glabrous, tube 1.5-2 mm long;
<emphasis italics="true" pageId="1" pageNumber="84">corolla lobes</emphasis>
narrowly lanceolate with a triangular acuminate apex, 9-10
<normalizedToken originalValue="×">x</normalizedToken>
3-4 mm, laterally revolute, lobe tips recurved.
<emphasis italics="true" pageId="1" pageNumber="84">Gynostegium</emphasis>
stalked, corona column conical 1-1.2
<normalizedToken originalValue="×">x</normalizedToken>
ca. 2 mm diam, glabrous;
<emphasis italics="true" pageId="1" pageNumber="84">corona</emphasis>
staminal, 2.5-3 mm high, 6-7 mm in diameter, fleshy, yellow with a purple centre;
<emphasis italics="true" pageId="1" pageNumber="84">corona lobes</emphasis>
laterally compressed, ovate above, with revolute margins beneath, 2.8-3.2 mm
<normalizedToken originalValue="×">x</normalizedToken>
1-1.2 mm, inner process erecto-patent, linear with an acute tip, as high as the anthers, outer process round.
<emphasis italics="true" pageId="1" pageNumber="84">Anthers</emphasis>
ovate, 650-750
<normalizedToken originalValue="×">x</normalizedToken>
300-400
<normalizedToken originalValue="µm">µm</normalizedToken>
, with apical round membranaceous appendage as high as the style-head apex.
<emphasis italics="true" pageId="1" pageNumber="84">Pollinia</emphasis>
oblong, with obliquely truncate apex and round base and evident pellucid margin, 550-630
<normalizedToken originalValue="×">x</normalizedToken>
150-200
<normalizedToken originalValue="μm">μm</normalizedToken>
;
<emphasis italics="true" pageId="1" pageNumber="84">caudicles</emphasis>
attached at the base of the retinaculum, elongate, ca. 150
<normalizedToken originalValue="µm">µm</normalizedToken>
long,
<emphasis italics="true" pageId="1" pageNumber="84">corpusculum</emphasis>
600-650
<normalizedToken originalValue="×">x</normalizedToken>
250-300
<normalizedToken originalValue="μm">μm</normalizedToken>
; style-head 5-angled in cross section, with 5 spreading lobes alternating with the stamens;
<emphasis italics="true" pageId="1" pageNumber="84">style-head</emphasis>
apex columnar, 1-1.2 mm long, ca. 0.5 mm broad at the base, apex conical;
<emphasis italics="true" pageId="1" pageNumber="84">ovary</emphasis>
linear, ca. 2 mm long, each carpel ca. 0.5 mm wide at the base.
<emphasis italics="true" pageId="1" pageNumber="84">Fruits</emphasis>
and
<emphasis italics="true" pageId="1" pageNumber="84">seeds</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="etymology">
<paragraph pageId="1" pageNumber="84">Etymology.</paragraph>
<paragraph pageId="1" pageNumber="84">
This species is named after Ruth Kiew (1946-), tropical botanist based at the Forest Research Institute Malaysia and author of numerous publications on
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia" order="Lamiales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="1" pageNumber="84">Begonia</emphasis>
</taxonomicName>
L. and
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Lamiales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="family">Gesneriaceae</taxonomicName>
, among others.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="distribution and ecology">
<paragraph pageId="1" pageNumber="84">Distribution and ecology.</paragraph>
<paragraph pageId="1" pageNumber="84">
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
is only known from Bukit Baturong, Sabah, Malaysia, where it was found growing on limestone.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="conservation status">
<paragraph pageId="1" pageNumber="84">Conservation status.</paragraph>
<paragraph pageId="1" pageNumber="84">
The distribution area, population size and possible threats to the habitat of
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
are not known, as it is only known from the type specimen and unlocalised cultivated material. It is therefore considered Data Deficient (DD) (
<bibRefCitation author="IUCN Standards and Petitions Subcommittee" journalOrPublisher="Standards and Petitions Subcommittee" pageId="5" pageNumber="88" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B3" refString="IUCN Standards and Petitions Subcommittee, 2014. Guidelines for using the IUCN Red List categories and criteria. Version 11. Standards and Petitions Subcommittee, http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for using the IUCN Red List categories and criteria. Version 11" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2014">IUCN 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="notes">
<paragraph pageId="1" pageNumber="84">Notes.</paragraph>
<paragraph pageId="1" pageNumber="84">
The most striking feature of
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
is its lack of coloured latex. Other species lacking coloured latex are the type species of the genus
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya carnosa" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="carnosa">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya carnosa</emphasis>
</taxonomicName>
R.Br. and several morphologically similar taxa (
<bibRefCitation author="Rodda, M" journalOrPublisher="Webbia" pageId="6" pageNumber="89" pagination="23 - 27" publicationUrl="10.1080/00837792.2012.10670904" refId="B12" refString="Rodda, M, Simonsson Juhonewee, N, 2012. Hoya vangviengiensis (Apocynaceae, Asclepiadoideae), a new species from limestone formations of Vang Vieng, Lao PDR. Webbia 67: 23 - 27, DOI: 10.1080/00837792.2012.10670904" title="Hoya vangviengiensis (Apocynaceae, Asclepiadoideae), a new species from limestone formations of Vang Vieng, Lao PDR." url="10.1080/00837792.2012.10670904" volume="67" year="2012">Rodda and Simonsson Juhonewe 2012</bibRefCitation>
). All these have rotate corollas with thickly pubescent lobes within and rhomboid corona lobes with narrow inner and outer corona lobe processes. In Borneo only
<taxonomicName authorityName="T. Green" authorityYear="2004" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya monetteae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="monetteae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya monetteae</emphasis>
</taxonomicName>
T. Green belongs to this group.
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
, as mentioned above, is morphologically very similar to
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
, another non-laticiferous species known to occur in Java and Sumatra. Both species have deeply lobed rotate corollas, laterally compressed corona lobes and corpusculum of the pollinarium almost as large as the pollinium.
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
has larger flowers (1.8-2.2 cm in diameter when flattened vs. 1.5-1.7 cm) and the corolla lobes are narrowly lanceolate (vs. ovate in
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
). The corona lobes of both species are ovate but in
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
the outer process is round while in
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
it terminates in an incurved membranaceous apiculate appendage. Flowers of
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
are illustrated in Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya ruthiae photographed from Rodda M. MR 606 (SING) prior to pressing A Inflorescence D, E, F, G Two leaves (D, F adaxial surface E, G abaxial surface). Hoya uncinata photographed from Rodda M MR 607 (SING) prior to pressing B Flower, lateral view C Flower, top view. (Photographs by M. Rodda)" figureDoi="10.3897/phytokeys.53.5079.figure2" httpUri="https://binary.pensoft.net/fig/47619" pageId="1" pageNumber="84">2</figureCitation>
.
</paragraph>
<caption ID-Zenodo-Dep="964876" doi="10.3897/phytokeys.53.5079.figure1" httpUri="https://binary.pensoft.net/fig/47618" pageId="1" pageNumber="84" start="Figure 1" startId="F1">
<paragraph pageId="1" pageNumber="84">
<emphasis bold="true" pageId="1" pageNumber="84">Figure 1.</emphasis>
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
photographed from
<emphasis italics="true" pageId="1" pageNumber="84">Rodda M. MR606</emphasis>
(SING) prior to pressing
<emphasis bold="true" pageId="1" pageNumber="84">A</emphasis>
Flower, lateral view with two corolla lobes removed
<emphasis bold="true" pageId="1" pageNumber="84">B</emphasis>
Corolla, underneath
<emphasis bold="true" pageId="1" pageNumber="84">C</emphasis>
Corolla and corona, top view
<emphasis bold="true" pageId="1" pageNumber="84">D</emphasis>
Corona, underneath
<emphasis bold="true" pageId="1" pageNumber="84">E</emphasis>
Pedicel, calyx and ovaries
<emphasis bold="true" pageId="1" pageNumber="84">F</emphasis>
Pollinarium with twin pollinia. (Photographs by M. Rodda)
</paragraph>
</caption>
<caption ID-Zenodo-Dep="964878" doi="10.3897/phytokeys.53.5079.figure2" httpUri="https://binary.pensoft.net/fig/47619" pageId="1" pageNumber="84" start="Figure 2" startId="F2">
<paragraph pageId="1" pageNumber="84">
<emphasis bold="true" pageId="1" pageNumber="84">Figure 2.</emphasis>
<taxonomicName authorityName="Rodda" authorityYear="2015" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya ruthiae" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya ruthiae</emphasis>
</taxonomicName>
photographed from
<emphasis italics="true" pageId="1" pageNumber="84">Rodda M. MR606</emphasis>
(SING) prior to pressing
<emphasis bold="true" pageId="1" pageNumber="84">A</emphasis>
Inflorescence
<emphasis bold="true" pageId="1" pageNumber="84">D, E, F, G</emphasis>
Two leaves (
<emphasis bold="true" pageId="1" pageNumber="84">D, F</emphasis>
adaxial surface
<emphasis bold="true" pageId="1" pageNumber="84">E, G</emphasis>
abaxial surface).
<taxonomicName authorityName="Teijsm &amp; Binn" authorityYear="1863" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya uncinata" order="Gentianales" pageId="1" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis italics="true" pageId="1" pageNumber="84">Hoya uncinata</emphasis>
</taxonomicName>
photographed from
<emphasis italics="true" pageId="1" pageNumber="84">Rodda M MR607</emphasis>
(SING) prior to pressing
<emphasis bold="true" pageId="1" pageNumber="84">B</emphasis>
Flower, lateral view
<emphasis bold="true" pageId="1" pageNumber="84">C</emphasis>
Flower, top view. (Photographs by M. Rodda)
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="1" pageNumber="84" type="additional specimens examined">
<paragraph pageId="1" pageNumber="84">Additional specimens examined.</paragraph>
<paragraph pageId="1" pageNumber="84">
Unlocalised (nursery origin), Cultivated in Thailand, Ratchaburi Prov., Ratchaburi, 23 March 2014,
<emphasis italics="true" pageId="1" pageNumber="84">Rodda M MR606</emphasis>
(SING).
</paragraph>
</subSubSection>
</treatment>
</document>