160 lines
13 KiB
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160 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza & Marusik, Yuri M." docDate="2021" docId="02249EDF73C059D1843C3DE96E998ED5" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Zodarion Walckenaer 1826" docType="treatment" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia & Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1035</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>145</mods:start>
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<mods:end>193</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
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<mods:identifier type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
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<mods:identifier type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181753573" LSID="urn:lsid:plazi:treatment:02249EDF73C059D1843C3DE96E998ED5" httpUri="http://treatment.plazi.org/id/02249EDF73C059D1843C3DE96E998ED5" lastPageNumber="145" pageId="0" pageNumber="145">
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<subSubSection pageId="0" pageNumber="145" type="nomenclature">
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<paragraph pageId="0" pageNumber="145">
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Genus
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<taxonomicName LSID="02249EDF-73C0-59D1-843C-3DE96E998ED5" authority="Walckenaer, 1826" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodarion Walckenaer, 1826</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="type species">
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<paragraph pageId="0" pageNumber="145">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName class="Arachnida" family="Zodariidae" genus="Enyo" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Enyo nitida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitida">
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<emphasis italics="true" pageId="0" pageNumber="145">Enyo nitida</emphasis>
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</taxonomicName>
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Audouin, 1826 from Egypt.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="comments">
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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With 177 named species, this is the largest genus within
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<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
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(
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<bibRefCitation author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
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). Based on the copulatory organs of the species currently considered in this genus,
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<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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does not appear to be monophyletic. There are only two species that are morphologically similar to the generotype:
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<taxonomicName lsidName="Z. luctuosum" pageId="0" pageNumber="145" rank="species" species="luctuosum">
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<emphasis italics="true" pageId="0" pageNumber="145">Z. luctuosum</emphasis>
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</taxonomicName>
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(O. Pickard-Cambridge, 1872) and
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<taxonomicName lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
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<emphasis italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
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</taxonomicName>
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(O. Pickard-Cambridge, 1872), from the eastern Mediterranean (east of Tunisia to Iran).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="materials_examined">
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<paragraph pageId="0" pageNumber="145">Comparative material.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
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<emphasis italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
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</taxonomicName>
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(O. Pickard-Cambridge, 1876),
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<materialsCitation ID-GBIF-Occurrence="3093634307" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Israel" location="Israel" specimenCount="2" specimenCount-male="2">
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<specimenCount type="male">1♂</specimenCount>
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from
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<collectingCountry name="Israel">Israel</collectingCountry>
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(Figs
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19D-F</figureCitation>
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,
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25D-F</figureCitation>
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) and
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<taxonomicName baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
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<emphasis italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
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</taxonomicName>
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(Audouin, 1826):
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<specimenCount type="male">1♂</specimenCount>
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from
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<collectingCountry name="Israel">Israel</collectingCountry>
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(Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28A-C</figureCitation>
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)
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="comments">
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<paragraph pageId="0" pageNumber="145">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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Until recently,
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<taxonomicName lsidName="Z. expers" pageId="0" pageNumber="145" rank="species" species="expers">
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<emphasis italics="true" pageId="0" pageNumber="145">Z. expers</emphasis>
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</taxonomicName>
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was placed in
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<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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, 1991, but now it is placed in
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<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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. It differs from the generotype of
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<taxonomicName authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
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</taxonomicName>
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, but the copulatory organs are also very different from those of
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<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
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<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
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</taxonomicName>
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, and most likely it represents a separate genus. Although it is absent in Iran or Turkmenistan, we have provided figures of this species to illustrate the differences with the generotype of
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<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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. Furthermore, although
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<taxonomicName lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
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<emphasis italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
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</taxonomicName>
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has not been recorded in the current study area, because it is the type species of the genus, we have provided illustartions for this species as well, to demonstrate its differences with other
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<taxonomicName authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
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genera in Iran and Turkmenistan, as well as with other species currently placed in
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<taxonomicName authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |