treatments-xml/data/02/1D/87/021D87FC8868FF99FF540DC6B736F8E4.xml

<document id="95D669FF37550C3C1CE7F9DD5A11E30A" ID-CLB-Dataset="30193" ID-DOI="10.11646/zootaxa.4413.3.7" ID-GBIF-Dataset="333e0858-f187-417b-8f38-23f615f9210c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1227963" ID-ZooBank="7B5A5CDE-E4D3-402D-99C6-BDF5585EA69D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524562695397" checkinUser="plazi" docAuthor="Ebert, David A." docDate="2018" docId="021D87FC8868FF99FF540DC6B736F8E4" docLanguage="en" docName="zootaxa.4413.3.7.pdf" docOrigin="Zootaxa 4413 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parmaturus nigripalatum Ebert 2018, new species" docType="treatment" docVersion="7" lastPageNumber="539" masterDocId="FE24FF84886AFF91FFC30D51B63FFF94" masterDocTitle="Parmaturus nigripalatum n. sp., a new species of deep-sea catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from Indonesia" masterLastPageNumber="540" masterPageNumber="531" pageNumber="533" updateTime="1698540879463" updateUser="ExternalLinkService">
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<mods:title id="E3491C0658CEA90D4A29A708895DA7E2">Parmaturus nigripalatum n. sp., a new species of deep-sea catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from Indonesia</mods:title>
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<mods:namePart id="BABC7EC944DA0A4460F2EAFA6A573929">Ebert, David A.</mods:namePart>
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<taxonomicName id="4DB44D698868FF93FF540DC6B7F2FF25" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[151,461,151,177]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="533" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88868FF93FF540DC6B7F2FF25" bold="true" box="[151,461,151,177]" italics="true" pageId="2" pageNumber="533">Parmaturus nigripalatum</emphasis>
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, 
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<paragraph id="8A0B36EA8868FF93FF540DEBB7AFFF63" blockId="2.[151,860,151,247]" pageId="2" pageNumber="533">
Indonesian Filetail Catshark (English), Hiu minyak (
<collectingCountry id="F2A3767A8868FF93FD200DEBB56CFF47" box="[739,851,186,211]" name="Indonesia" pageId="2" pageNumber="533">Indonesia</collectingCountry>
) (
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; 
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)
</paragraph>
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<paragraph id="8A0B36EA8868FF93FF540C78B479FEF2" blockId="2.[151,1436,294,1006]" pageId="2" pageNumber="533">
<materialsCitation id="3ADC3CB78868FF93FF540C78B479FEF2" ID-GBIF-Occurrence="1838430544" collectingDate="2016-04-16" collectionCode="NCIP" country="Indonesia" elevation="180" latitude="-9.04" location="Sumbawa" longLatPrecision="780" longitude="117.92" pageId="2" pageNumber="533" specimenCode="NCIP 6567, 548" specimenCount="1" typeStatus="holotype">
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<typeStatus id="550F88488868FF93FF540C78B739FED5" box="[151,262,297,321]" pageId="2" pageNumber="533" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DA129E918868FF93FED10C7BB7ABFED6" box="[274,404,297,322]" collectionCode="NCIP" country="Indonesia" httpUri="http://grbio.org/cool/h4qb-ekf6" name="Pusat Penelitian dan Pengembangan Oseanologi" pageId="2" pageNumber="533">
NCIP 
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, 
<specimenCode id="DA129E918868FF93FE5C0C78B7F6FED6" box="[415,457,297,322]" collectionCode="NCIP" country="Indonesia" httpUri="http://grbio.org/cool/h4qb-ekf6" name="Pusat Penelitian dan Pengembangan Oseanologi" pageId="2" pageNumber="533">548</specimenCode>
mm TL, adult male, south of 
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, 
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, 
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, 
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, deep water long line, 
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<elevation id="0199D1D98868FF93FF0C0C1FB771FEF1" box="[207,334,334,358]" metricMagnitude="2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="2" pageNumber="533" unit="m" value="180.0" valueMax="190.0" valueMin="170.0">170–190 m</elevation>
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depth, 
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<date id="FE0A102A8868FF93FE620C1CB47DFEF2" box="[417,578,333,358]" pageId="2" pageNumber="533" value="2016-04-16">16 April 2016</date>
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.
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</paragraph>
</subSubSection>
<subSubSection id="C2AE65618868FF93FF040C20B746FD5A" pageId="2" pageNumber="533" type="diagnosis">
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<emphasis id="B8C0EAF88868FF93FF040C20B77EFE1E" bold="true" box="[199,321,369,394]" pageId="2" pageNumber="533">Diagnosis.</emphasis>
A moderate-sized species of 
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<emphasis id="B8C0EAF88868FF93FD600C23B517FE1D" box="[675,808,370,393]" italics="true" pageId="2" pageNumber="533">Parmaturus</emphasis>
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with combination of characteristics as follows: body slender, soft-velvety skin, dermal denticles on side and upper body tricuspidate; uniformly plain medium brown body coloration; head slightly depress; snout relatively short; mouth curved, upper and lower labial furrows conspicuous with the uppers and lowers of equal lengths; the roof of mouth blackish with darker pores; teeth mainly tricuspidate with greatly elongate median cusps, about 50 rows in both jaws; origin of the first dorsal fin posterior to the insertion of the pelvic fins, pre-first dorsal fin slightly posterior of body mid-length; origin of second dorsal fin posterior to mid-length of anal fin base; pelvic fins origin and vent slightly posterior to midlength; second dorsal fin slightly larger than first; caudal peduncle relatively short, pronounced caudal crests on upper and lower anterior caudal fin margins with enlarged denticles; vertebral counts monospondylous centra 42, precaudal centra 90.
</paragraph>
</subSubSection>
<subSubSection id="C2AE65618868FF97FF040F88B5CAFA9E" lastPageId="6" lastPageNumber="537" pageId="2" pageNumber="533" type="description">
<paragraph id="8A0B36EA8868FF93FF040F88B753FC82" blockId="2.[151,1436,294,1006]" pageId="2" pageNumber="533">
<emphasis id="B8C0EAF88868FF93FF040F88B766FD66" bold="true" box="[199,345,729,754]" pageId="2" pageNumber="533">Description.</emphasis>
Proportional measurements expressed as a percentage of total length (TL) are provided for the 
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(
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).
</paragraph>
<paragraph id="8A0B36EA8868FF93FF040E70B323FC7A" blockId="2.[151,1436,294,1006]" pageId="2" pageNumber="533">
Body not tadpole-shaped, flabby, soft, slightly depressed anteriorly, tapering posteriorly from about pelvic fin bases towards caudal fin origin; abdomen longer than head length; pre-pelvic length 49.3% TL and pre-vent length 51.6% TL; pectoral-pelvic space 27.2% TL, 0.8 of head length; pelvic-anal space 1.5 times anal-fin base length (
<figureCitation id="128F2A6F8868FF93FF5C0EDCB6DAFC32" box="[159,229,909,934]" captionStart="FIGURE 2" captionStartId="2.[151,250,1665,1687]" captionTargetBox="[165,1426,1068,1630]" captionTargetId="figure@2.[151,1436,1058,1643]" captionTargetPageId="2" captionText="FIGURE 2. Parmaturus nigripalatum n. sp. holotype adult male 548 mm TL (NCIP 6567) in lateral and ventral views." httpUri="https://zenodo.org/record/1227967/files/figure.png" pageId="2" pageNumber="533">Fig. 2</figureCitation>
). Head relatively short and broad, length 20.7% TL, width 10.8% TL; slightly depressed, narrowly rounded in lateral view, height 7.2% TL. Snout short, pre-narial length 4.7% TL, 1.1 times eye length; rounded-parabolic in dorsal view, tip moderately rounded, conical in lateral view; preoral length 5% TL, 
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mouth width.
</paragraph>
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<emphasis id="B8C0EAF88868FF93FF540BD0B72DF903" bold="true" box="[151,274,1665,1687]" pageId="2" pageNumber="533">FIGURE 2.</emphasis>
<taxonomicName id="4DB44D698868FF93FEDA0BD3B423F902" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[281,540,1665,1687]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="533" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88868FF93FEDA0BD3B423F902" box="[281,540,1665,1687]" italics="true" pageId="2" pageNumber="533">Parmaturus nigripalatum</emphasis>
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<emphasis id="B8C0EAF88868FF93FDE00BD0B46AF903" bold="true" box="[547,597,1665,1687]" pageId="2" pageNumber="533">n. sp</emphasis>
.
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holotype adult male 548 mm TL (NCIP 6567) in lateral and ventral views.
</paragraph>
</caption>
<paragraph id="8A0B36EA8868FF93FF040B95B52BF8F8" blockId="2.[151,1437,1731,2008]" pageId="2" pageNumber="533">
Eyes relatively large, length 4.2% TL, 
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head length, dorsolateral on head, slit-like, not elevated, slightly above lateral midline; interorbital width 0.8 times pre-narial snout length, 
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head length; subocular ridges welldeveloped. Spiracle small, sub-oval, length 14.5 eye length, and set behind and below posterior eye notch. Nostrils well separated, tube-like, internarial length 2.6% TL; anterior nasal flaps triangular, posterior tip expanded, forming a conspicuous lobe, above level with the mouth.
</paragraph>
<paragraph id="8A0B36EA8868FF92FF040A26B7CEFE89" blockId="2.[151,1437,1731,2008]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533">
Mouth large, semi-angular, width 9.0% TL, 2.1 times its length, with symphysial notch on both jaws; lower symphysis not reaching upper symphysis (
<figureCitation id="128F2A6F8868FF93FDB00ACAB48DF820" box="[627,690,1947,1972]" captionStart="FIGURE 3" captionStartId="3.[151,250,1312,1334]" captionTargetBox="[357,1247,355,1275]" captionTargetId="figure@3.[340,1247,337,1291]" captionTargetPageId="3" captionText="FIGURE 3. Ventral view of head of Parmaturus nigripalatum n. sp. holotype adult male 548 mm TL (NCIP 6567)." httpUri="https://zenodo.org/record/1227969/files/figure.png" pageId="2" pageNumber="533">Fig 3</figureCitation>
). Labial furrows well developed, upper labial furrow almost equal to lower furrow, upper and lower labial furrows length 1.9% TL. Teeth of upper and lower jaws exposed when mouth closed; tricuspidate with enlarged, erect, acutely pointed central cusp flanked by a single smaller cusplet on each side; teeth on upper jaw smaller relative to lower jaw, but otherwise similar shaped; tooth count on upper jaw 52, lower jaw 48; longest teeth adjacent to symphysis, 2–3 rows of shorter teeth at symphysis (
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); lateral teeth relatively shorter than anterior.
</paragraph>
<caption id="DECB66628869FF92FF540871B310FAA2" box="[151,1327,1312,1334]" httpUri="https://zenodo.org/record/1227969/files/figure.png" pageId="3" pageNumber="534" startId="3.[151,250,1312,1334]" targetBox="[357,1247,355,1275]" targetPageId="3">
<paragraph id="8A0B36EA8869FF92FF540871B310FAA2" blockId="3.[151,1327,1312,1334]" box="[151,1327,1312,1334]" pageId="3" pageNumber="534">
<emphasis id="B8C0EAF88869FF92FF540871B72DFAA2" bold="true" box="[151,274,1312,1334]" pageId="3" pageNumber="534">FIGURE 3.</emphasis>
Ventral view of head of 
<taxonomicName id="4DB44D698869FF92FDCE0870B52FFAA1" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[525,784,1312,1334]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="3" pageNumber="534" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88869FF92FDCE0870B52FFAA1" box="[525,784,1312,1334]" italics="true" pageId="3" pageNumber="534">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F357838869FF92FCD50871B570FAA2" box="[790,847,1312,1334]" pageId="3" pageNumber="534" rank="species">
<emphasis id="B8C0EAF88869FF92FCD50871B576FAA2" bold="true" box="[790,841,1312,1334]" pageId="3" pageNumber="534">n. sp</emphasis>
.
</taxonomicNameLabel>
holotype adult male 548 mm TL (NCIP 6567).
</paragraph>
</caption>
<paragraph id="8A0B36EA8869FF92FF040832B5B6F92B" blockId="3.[151,1437,1379,2015]" pageId="3" pageNumber="534">
Dermal denticles on side small, tricuspidate, erect, closely spaced but not overlapping, crown-like with a long pointed median cusp, ending in a longitudinal ridge; denticles on head, body and caudal in similar shape; denticles on the lateral trunk with shorter median cusps. Upper and lower caudal crests well developed, dorsal crests extending from mid-dorsal caudal margin to the second dorsal fin insertion, length of dorsal crests 10.8% TL; ventral crests extending well along caudal peduncle from the origin of lower caudal to almost reaching anal fin insertion or just below mid second dorsal base, length of lower crests 9.3% TL. Denticles of caudal crests have similar shape to body denticles but greatly enlarged, consisting of two rows of enlarged denticles directed posterolaterally and one slightly smaller denticles in the middle row (
<figureCitation id="128F2A6F8869FF92FC620B0FB5DFF9E3" box="[929,992,1630,1655]" captionStart="FIGURE 5" captionStartId="7.[151,250,716,738]" captionTargetBox="[151,1436,193,695]" captionTargetId="figure@7.[151,1436,193,695]" captionTargetPageId="7" captionText="FIGURE 5. Parmaturus nigripalatum n. sp., close-up of denticles on flank (left) and caudal crests of dorsal caudal margin (right)." httpUri="https://zenodo.org/record/1227973/files/figure.png" pageId="3" pageNumber="534">Fig 5</figureCitation>
); denticles on anterior crests increasing in size posteriorly until the middle of crests and then slightly smaller posteriorly onto mid-dorsal caudal margin; denticles on ventral crests slightly smaller to those on dorsal crests.
</paragraph>
<paragraph id="8A0B36EA8869FF92FF040B9AB6CDF84B" blockId="3.[151,1437,1379,2015]" pageId="3" pageNumber="534">
Pectoral fins relatively small, lobate, rounded at apex; anterior margin more convex than posterior margin, anterior margin length 9.6% TL, inner margin rounded. Pelvic fins relatively large, semi-oval, apex broadly rounded, rear corner narrowly rounded, length 9.2% TL. Adult claspers relatively short, outer length 5.4% TL, clasper base width 29.8% of clasper outer length, tapering from base to tip, clasper tip angular and pointed. Denticles covering entire ventral surface and almost absent from dorsal surface. Clasper groove beginning from near pelvic insertion, obliquely running from midline to the lateral edge. Lateral fold and cover rhipidion are not well developed or nearly absent. Rhipidion present along the clasper gland with posterior end beside exorhipidion (
<figureCitation id="128F2A6F8869FF92FF5C0A97B6DBF84B" box="[159,228,1990,2015]" captionStart="FIGURE6" captionStartId="7.[151,250,1934,1956]" captionTargetBox="[340,1180,827,1899]" captionTargetId="figure@7.[340,1247,827,1899]" captionTargetPageId="7" captionText="FIGURE6. Dorsalview of claspers of preserved specimenof Parmaturus nigripalatum n. sp. holotypeadult male 548 mm TL (NCIP 6567). Abbreviations: AP: apopyle; CG: clasper groove; HP: hypopyle; CRH: cover rhipidion; RH: rhipidion; ERH: exorhipidion." httpUri="https://zenodo.org/record/1227975/files/figure.png" pageId="3" pageNumber="534">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DECB6662886EFF95FF540DC9B71CFF5B" pageId="4" pageNumber="535">
<paragraph id="8A0B36EA886EFF95FF540DC9B71CFF5B" blockId="4.[151,1435,151,207]" pageId="4" pageNumber="535">
<emphasis id="B8C0EAF8886EFF95FF540DC9B734FF3B" bold="true" box="[151,267,152,175]" pageId="4" pageNumber="535">TABLE 1.</emphasis>
Morphological measurement of 
<taxonomicName id="4DB44D69886EFF95FDB30DC9B5BDFF3B" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[624,898,151,175]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="535" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886EFF95FDB30DC9B5BDFF3B" box="[624,898,151,175]" italics="true" pageId="4" pageNumber="535">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886EFF95FC480DC9B5F6FF3B" box="[907,969,152,175]" pageId="4" pageNumber="535" rank="species">
<emphasis id="B8C0EAF8886EFF95FC480DC9B5F6FF3B" bold="true" box="[907,969,152,175]" pageId="4" pageNumber="535">n. sp.</emphasis>
</taxonomicNameLabel>
, presented in in the percentage of the total length (mm).
</paragraph>
</caption>
<paragraph id="8A0B36EA886EFF95FF5C0DBBB4C8F856" pageId="4" pageNumber="535">
<table id="F8B4C44A886E006EFF5C0DBBB231F856" box="[159,1038,234,1986]" colsContinueIn="5.[159,1038,202,1568]" gridcols="2" gridrows="42" pageId="4" pageNumber="535">
<tr id="348434A8886E006EFF5C0DBBB231FE94" box="[159,1038,234,256]" gridrow="0" pageId="4" pageNumber="535">
<th id="77555DD4886E006EFF5C0DBBB7CBFE94" box="[159,500,234,256]" gridcol="0" gridrow="0" pageId="4" pageNumber="535">Species</th>
<th id="77555DD4886E006EFD080DBBB231FE94" box="[715,1038,234,256]" gridcol="1" gridrow="0" pageId="4" pageNumber="535">
<taxonomicName id="4DB44D69886EFF95FD0F0DBAB5F0FE94" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[716,975,235,256]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="535" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886EFF95FD0F0DBAB5F0FE94" box="[716,975,235,256]" italics="true" pageId="4" pageNumber="535">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886EFF95FC160DBBB231FE94" box="[981,1038,234,256]" pageId="4" pageNumber="535" rank="species">
<emphasis id="B8C0EAF8886EFF95FC160DBBB237FE94" bold="true" box="[981,1032,234,256]" pageId="4" pageNumber="535">n. sp</emphasis>
.
</taxonomicNameLabel>
</th>
</tr>
<tr id="348434A8886E006EFF5C0C45B231FEBE" box="[159,1038,276,298]" gridrow="1" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0C45B7CBFEBE" box="[159,500,276,298]" gridcol="0" gridrow="1" pageId="4" pageNumber="535">Catalogue number</td>
<td id="77555DD4886E006EFD080C45B231FEBE" box="[715,1038,276,298]" gridcol="1" gridrow="1" pageId="4" pageNumber="535">NCIP 6567</td>
</tr>
<tr id="348434A8886E006EFF5C0C13B231FECC" box="[159,1038,322,344]" gridrow="2" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0C13B7CBFECC" box="[159,500,322,344]" gridcol="0" gridrow="2" pageId="4" pageNumber="535">Type</td>
<td id="77555DD4886E006EFD080C13B231FECC" box="[715,1038,322,344]" gridcol="1" gridrow="2" pageId="4" pageNumber="535">holotype</td>
</tr>
<tr id="348434A8886E006EFF5C0C21B231FE12" box="[159,1038,368,390]" gridrow="3" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0C21B7CBFE12" box="[159,500,368,390]" gridcol="0" gridrow="3" pageId="4" pageNumber="535">Sex and maturity</td>
<td id="77555DD4886E006EFD080C21B231FE12" box="[715,1038,368,390]" gridcol="1" gridrow="3" pageId="4" pageNumber="535">adult male</td>
</tr>
<tr id="348434A8886E006EFF5C0CCBB231FE24" box="[159,1038,410,432]" gridrow="4" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0CCBB7CBFE24" box="[159,500,410,432]" gridcol="0" gridrow="4" pageId="4" pageNumber="535">Total length (mm)</td>
<td id="77555DD4886E006EFD080CCBB231FE24" box="[715,1038,410,432]" gridcol="1" gridrow="4" pageId="4" pageNumber="535">548</td>
</tr>
<tr id="348434A8886E006EFF5C0C95B231FE4E" box="[159,1038,452,474]" gridrow="5" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0C95B7CBFE4E" box="[159,500,452,474]" gridcol="0" gridrow="5" pageId="4" pageNumber="535">Pre-first dorsal length</td>
<td id="77555DD4886E006EFD080C95B231FE4E" box="[715,1038,452,474]" gridcol="1" gridrow="5" pageId="4" pageNumber="535">53.83</td>
</tr>
<tr id="348434A8886E006EFF5C0CBFB231FD90" box="[159,1038,494,516]" gridrow="6" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0CBFB7CBFD90" box="[159,500,494,516]" gridcol="0" gridrow="6" pageId="4" pageNumber="535">Pre-second dorsal length</td>
<td id="77555DD4886E006EFD080CBFB231FD90" box="[715,1038,494,516]" gridcol="1" gridrow="6" pageId="4" pageNumber="535">69.71</td>
</tr>
<tr id="348434A8886E006EFF5C0F49B231FDBA" box="[159,1038,536,558]" gridrow="7" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0F49B7CBFDBA" box="[159,500,536,558]" gridcol="0" gridrow="7" pageId="4" pageNumber="535">Precaudal length</td>
<td id="77555DD4886E006EFD080F49B231FDBA" box="[715,1038,536,558]" gridcol="1" gridrow="7" pageId="4" pageNumber="535">75.55</td>
</tr>
<tr id="348434A8886E006EFF5C0F13B231FDCC" box="[159,1038,578,600]" gridrow="8" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0F13B7CBFDCC" box="[159,500,578,600]" gridcol="0" gridrow="8" pageId="4" pageNumber="535">Prepectoral length</td>
<td id="77555DD4886E006EFD080F13B231FDCC" box="[715,1038,578,600]" gridcol="1" gridrow="8" pageId="4" pageNumber="535">19.42</td>
</tr>
<tr id="348434A8886E006EFF5C0F3DB231FD16" box="[159,1038,620,642]" gridrow="9" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0F3DB7CBFD16" box="[159,500,620,642]" gridcol="0" gridrow="9" pageId="4" pageNumber="535">Prepelvic length</td>
<td id="77555DD4886E006EFD080F3DB231FD16" box="[715,1038,620,642]" gridcol="1" gridrow="9" pageId="4" pageNumber="535">49.27</td>
</tr>
<tr id="348434A8886E006EFF5C0FC7B231FD38" box="[159,1038,662,684]" gridrow="10" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0FC7B7CBFD38" box="[159,500,662,684]" gridcol="0" gridrow="10" pageId="4" pageNumber="535">Preanal length</td>
<td id="77555DD4886E006EFD080FC7B231FD38" box="[715,1038,662,684]" gridcol="1" gridrow="10" pageId="4" pageNumber="535">62.23</td>
</tr>
<tr id="348434A8886E006EFF5C0F91B231FD42" box="[159,1038,704,726]" gridrow="11" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0F91B7CBFD42" box="[159,500,704,726]" gridcol="0" gridrow="11" pageId="4" pageNumber="535">Snout-vent length</td>
<td id="77555DD4886E006EFD080F91B231FD42" box="[715,1038,704,726]" gridcol="1" gridrow="11" pageId="4" pageNumber="535">51.64</td>
</tr>
<tr id="348434A8886E006EFF5C0FBBB231FC94" box="[159,1038,746,768]" gridrow="12" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0FBBB7CBFC94" box="[159,500,746,768]" gridcol="0" gridrow="12" pageId="4" pageNumber="535">Preorbital length (direct)</td>
<td id="77555DD4886E006EFD080FBBB231FC94" box="[715,1038,746,768]" gridcol="1" gridrow="12" pageId="4" pageNumber="535">6.82</td>
</tr>
<tr id="348434A8886E006EFF5C0E45B231FCBE" box="[159,1038,788,810]" gridrow="13" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0E45B7CBFCBE" box="[159,500,788,810]" gridcol="0" gridrow="13" pageId="4" pageNumber="535">Eye length</td>
<td id="77555DD4886E006EFD080E45B231FCBE" box="[715,1038,788,810]" gridcol="1" gridrow="13" pageId="4" pageNumber="535">4.19</td>
</tr>
<tr id="348434A8886E006EFF5C0E6FB231FCC0" box="[159,1038,830,852]" gridrow="14" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0E6FB7CBFCC0" box="[159,500,830,852]" gridcol="0" gridrow="14" pageId="4" pageNumber="535">Eye height</td>
<td id="77555DD4886E006EFD080E6FB231FCC0" box="[715,1038,830,852]" gridcol="1" gridrow="14" pageId="4" pageNumber="535">2.15</td>
</tr>
<tr id="348434A8886E006EFF5C0E39B231FCEA" box="[159,1038,872,894]" gridrow="15" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0E39B7CBFCEA" box="[159,500,872,894]" gridcol="0" gridrow="15" pageId="4" pageNumber="535">Interorbital space</td>
<td id="77555DD4886E006EFD080E39B231FCEA" box="[715,1038,872,894]" gridcol="1" gridrow="15" pageId="4" pageNumber="535">6.11</td>
</tr>
<tr id="348434A8886E006EFF5C0EC3B231FC3C" box="[159,1038,914,936]" gridrow="16" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0EC3B7CBFC3C" box="[159,500,914,936]" gridcol="0" gridrow="16" pageId="4" pageNumber="535">Prespiracular length</td>
<td id="77555DD4886E006EFD080EC3B231FC3C" box="[715,1038,914,936]" gridcol="1" gridrow="16" pageId="4" pageNumber="535">11.32</td>
</tr>
<tr id="348434A8886E006EFF5C0EEDB231FC46" box="[159,1038,956,978]" gridrow="17" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0EEDB7CBFC46" box="[159,500,956,978]" gridcol="0" gridrow="17" pageId="4" pageNumber="535">Spiracle length</td>
<td id="77555DD4886E006EFD080EEDB231FC46" box="[715,1038,956,978]" gridcol="1" gridrow="17" pageId="4" pageNumber="535">0.29</td>
</tr>
<tr id="348434A8886E006EFF5C0EB7B231FC68" box="[159,1038,998,1020]" gridrow="18" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0EB7B7CBFC68" box="[159,500,998,1020]" gridcol="0" gridrow="18" pageId="4" pageNumber="535">Prenarial length</td>
<td id="77555DD4886E006EFD080EB7B231FC68" box="[715,1038,998,1020]" gridcol="1" gridrow="18" pageId="4" pageNumber="535">4.66</td>
</tr>
<tr id="348434A8886E006EFF5C0941B231FBB2" box="[159,1038,1040,1062]" gridrow="19" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0941B7CBFBB2" box="[159,500,1040,1062]" gridcol="0" gridrow="19" pageId="4" pageNumber="535">Nostril width</td>
<td id="77555DD4886E006EFD080941B231FBB2" box="[715,1038,1040,1062]" gridcol="1" gridrow="19" pageId="4" pageNumber="535">2.59</td>
</tr>
<tr id="348434A8886E006EFF5C096BB231FBC4" box="[159,1038,1082,1104]" gridrow="20" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C096BB7CBFBC4" box="[159,500,1082,1104]" gridcol="0" gridrow="20" pageId="4" pageNumber="535">Internarial space</td>
<td id="77555DD4886E006EFD08096BB231FBC4" box="[715,1038,1082,1104]" gridcol="1" gridrow="20" pageId="4" pageNumber="535">2.55</td>
</tr>
<tr id="348434A8886E006EFF5C0935B231FBEE" box="[159,1038,1124,1146]" gridrow="21" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0935B7CBFBEE" box="[159,500,1124,1146]" gridcol="0" gridrow="21" pageId="4" pageNumber="535">Anterior nasal flap length</td>
<td id="77555DD4886E006EFD080935B231FBEE" box="[715,1038,1124,1146]" gridcol="1" gridrow="21" pageId="4" pageNumber="535">1.39</td>
</tr>
<tr id="348434A8886E006EFF5C09DFB231FB30" box="[159,1038,1166,1188]" gridrow="22" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C09DFB7CBFB30" box="[159,500,1166,1188]" gridcol="0" gridrow="22" pageId="4" pageNumber="535">Preoral length</td>
<td id="77555DD4886E006EFD0809DFB231FB30" box="[715,1038,1166,1188]" gridcol="1" gridrow="22" pageId="4" pageNumber="535">5.03</td>
</tr>
<tr id="348434A8886E006EFF5C09E9B231FB5A" box="[159,1038,1208,1230]" gridrow="23" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C09E9B7CBFB5A" box="[159,500,1208,1230]" gridcol="0" gridrow="23" pageId="4" pageNumber="535">Mouth width</td>
<td id="77555DD4886E006EFD0809E9B231FB5A" box="[715,1038,1208,1230]" gridcol="1" gridrow="23" pageId="4" pageNumber="535">8.97</td>
</tr>
<tr id="348434A8886E006EFF5C09B3B231FB6C" box="[159,1038,1250,1272]" gridrow="24" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C09B3B7CBFB6C" box="[159,500,1250,1272]" gridcol="0" gridrow="24" pageId="4" pageNumber="535">Mouth length</td>
<td id="77555DD4886E006EFD0809B3B231FB6C" box="[715,1038,1250,1272]" gridcol="1" gridrow="24" pageId="4" pageNumber="535">4.21</td>
</tr>
<tr id="348434A8886E006EFF5C085DB231FAB6" box="[159,1038,1292,1314]" gridrow="25" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C085DB7CBFAB6" box="[159,500,1292,1314]" gridcol="0" gridrow="25" pageId="4" pageNumber="535">Upper labial furrow length</td>
<td id="77555DD4886E006EFD08085DB231FAB6" box="[715,1038,1292,1314]" gridcol="1" gridrow="25" pageId="4" pageNumber="535">1.90</td>
</tr>
<tr id="348434A8886E006EFF5C0867B231FAD8" box="[159,1038,1334,1356]" gridrow="26" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0867B7CBFAD8" box="[159,500,1334,1356]" gridcol="0" gridrow="26" pageId="4" pageNumber="535">Lower labial furrow length</td>
<td id="77555DD4886E006EFD080867B231FAD8" box="[715,1038,1334,1356]" gridcol="1" gridrow="26" pageId="4" pageNumber="535">1.94</td>
</tr>
<tr id="348434A8886E006EFF5C0831B231FAE2" box="[159,1038,1376,1398]" gridrow="27" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0831B7CBFAE2" box="[159,500,1376,1398]" gridcol="0" gridrow="27" pageId="4" pageNumber="535">Prebranchial length</td>
<td id="77555DD4886E006EFD080831B231FAE2" box="[715,1038,1376,1398]" gridcol="1" gridrow="27" pageId="4" pageNumber="535">15.50</td>
</tr>
<tr id="348434A8886E006EFF5C08DBB231FA34" box="[159,1038,1418,1440]" gridrow="28" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C08DBB7CBFA34" box="[159,500,1418,1440]" gridcol="0" gridrow="28" pageId="4" pageNumber="535">Head length</td>
<td id="77555DD4886E006EFD0808DBB231FA34" box="[715,1038,1418,1440]" gridcol="1" gridrow="28" pageId="4" pageNumber="535">20.69</td>
</tr>
<tr id="348434A8886E006EFF5C08E5B231FA5E" box="[159,1038,1460,1482]" gridrow="29" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C08E5B7CBFA5E" box="[159,500,1460,1482]" gridcol="0" gridrow="29" pageId="4" pageNumber="535">Head height</td>
<td id="77555DD4886E006EFD0808E5B231FA5E" box="[715,1038,1460,1482]" gridcol="1" gridrow="29" pageId="4" pageNumber="535">7.21</td>
</tr>
<tr id="348434A8886E006EFF5C088FB231FA60" box="[159,1038,1502,1524]" gridrow="30" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C088FB7CBFA60" box="[159,500,1502,1524]" gridcol="0" gridrow="30" pageId="4" pageNumber="535">Head width</td>
<td id="77555DD4886E006EFD08088FB231FA60" box="[715,1038,1502,1524]" gridcol="1" gridrow="30" pageId="4" pageNumber="535">10.76</td>
</tr>
<tr id="348434A8886E006EFF5C0B59B231F98A" box="[159,1038,1544,1566]" gridrow="31" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0B59B7CBF98A" box="[159,500,1544,1566]" gridcol="0" gridrow="31" pageId="4" pageNumber="535">First gill slit height</td>
<td id="77555DD4886E006EFD080B59B231F98A" box="[715,1038,1544,1566]" gridcol="1" gridrow="31" pageId="4" pageNumber="535">2.22</td>
</tr>
<tr id="348434A8886E006EFF5C0B63B231F9DC" box="[159,1038,1586,1608]" gridrow="32" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0B63B7CBF9DC" box="[159,500,1586,1608]" gridcol="0" gridrow="32" pageId="4" pageNumber="535">Fifth gill slit height</td>
<td id="77555DD4886E006EFD080B63B231F9DC" box="[715,1038,1586,1608]" gridcol="1" gridrow="32" pageId="4" pageNumber="535">1.32</td>
</tr>
<tr id="348434A8886E006EFF5C0B0DB231F9E6" box="[159,1038,1628,1650]" gridrow="33" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0B0DB7CBF9E6" box="[159,500,1628,1650]" gridcol="0" gridrow="33" pageId="4" pageNumber="535">First dorsal-fin anterior margin</td>
<td id="77555DD4886E006EFD080B0DB231F9E6" box="[715,1038,1628,1650]" gridcol="1" gridrow="33" pageId="4" pageNumber="535">8.64</td>
</tr>
<tr id="348434A8886E006EFF5C0BD7B231F908" box="[159,1038,1670,1692]" gridrow="34" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0BD7B7CBF908" box="[159,500,1670,1692]" gridcol="0" gridrow="34" pageId="4" pageNumber="535">First dorsal-fin base</td>
<td id="77555DD4886E006EFD080BD7B231F908" box="[715,1038,1670,1692]" gridcol="1" gridrow="34" pageId="4" pageNumber="535">6.47</td>
</tr>
<tr id="348434A8886E006EFF5C0BE1B231F952" box="[159,1038,1712,1734]" gridrow="35" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0BE1B7CBF952" box="[159,500,1712,1734]" gridcol="0" gridrow="35" pageId="4" pageNumber="535">First dorsal-fin height</td>
<td id="77555DD4886E006EFD080BE1B231F952" box="[715,1038,1712,1734]" gridcol="1" gridrow="35" pageId="4" pageNumber="535">3.80</td>
</tr>
<tr id="348434A8886E006EFF5C0B8BB231F964" box="[159,1038,1754,1776]" gridrow="36" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0B8BB7CBF964" box="[159,500,1754,1776]" gridcol="0" gridrow="36" pageId="4" pageNumber="535">First dorsal-fin inner margin</td>
<td id="77555DD4886E006EFD080B8BB231F964" box="[715,1038,1754,1776]" gridcol="1" gridrow="36" pageId="4" pageNumber="535">2.05</td>
</tr>
<tr id="348434A8886E006EFF5C0A55B231F88E" box="[159,1038,1796,1818]" gridrow="37" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0A55B7CBF88E" box="[159,500,1796,1818]" gridcol="0" gridrow="37" pageId="4" pageNumber="535">First dorsal-fin posterior margin</td>
<td id="77555DD4886E006EFD080A55B231F88E" box="[715,1038,1796,1818]" gridcol="1" gridrow="37" pageId="4" pageNumber="535">3.28</td>
</tr>
<tr id="348434A8886E006EFF5C0A7FB231F8D0" box="[159,1038,1838,1860]" gridrow="38" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0A7FB7CBF8D0" box="[159,500,1838,1860]" gridcol="0" gridrow="38" pageId="4" pageNumber="535">Interdorsal space</td>
<td id="77555DD4886E006EFD080A7FB231F8D0" box="[715,1038,1838,1860]" gridcol="1" gridrow="38" pageId="4" pageNumber="535">9.46</td>
</tr>
<tr id="348434A8886E006EFF5C0A09B231F8FA" box="[159,1038,1880,1902]" gridrow="39" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0A09B7CBF8FA" box="[159,500,1880,1902]" gridcol="0" gridrow="39" pageId="4" pageNumber="535">Second dorsal-fin anterior margin</td>
<td id="77555DD4886E006EFD080A09B231F8FA" box="[715,1038,1880,1902]" gridcol="1" gridrow="39" pageId="4" pageNumber="535">10.75</td>
</tr>
<tr id="348434A8886E006EFF5C0AD3B231F80C" box="[159,1038,1922,1944]" gridrow="40" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0AD3B7CBF80C" box="[159,500,1922,1944]" gridcol="0" gridrow="40" pageId="4" pageNumber="535">Second dorsa-fin base</td>
<td id="77555DD4886E006EFD080AD3B231F80C" box="[715,1038,1922,1944]" gridcol="1" gridrow="40" pageId="4" pageNumber="535">8.62</td>
</tr>
<tr id="348434A8886E006EFF5C0AFDB231F856" box="[159,1038,1964,1986]" gridrow="41" pageId="4" pageNumber="535">
<td id="77555DD4886E006EFF5C0AFDB7CBF856" box="[159,500,1964,1986]" gridcol="0" gridrow="41" pageId="4" pageNumber="535">Second dorsal-fin height</td>
<td id="77555DD4886E006EFD080AFDB231F856" box="[715,1038,1964,1986]" gridcol="1" gridrow="41" pageId="4" pageNumber="535">4.85</td>
</tr>
</table>
</paragraph>
<paragraph id="8A0B36EA886EFF94FB9D0A86B79AF874" blockId="4.[1118,1428,2007,2028]" lastBlockId="5.[151,1437,1631,2017]" lastPageId="5" lastPageNumber="536" pageId="4" pageNumber="535">
<emphasis id="B8C0EAF8886EFF95FB9D0A86B3ABF878" box="[1118,1428,2007,2028]" italics="true" pageId="4" pageNumber="535">......continued on the next page</emphasis>
Dorsal fins subequal in size, first distinctly smaller than second dorsal fin. First dorsal fin moderately high, originating slightly behind pelvic fin insertion, anterior margin slightly longer than first dorsal fin base, narrowly rounded at apex, posterior margin slightly convex, length less than half of anterior margin length; free rear tip narrowly rounded, inner margin straight, inner margin length 31.7% base length. Second dorsal fin high, originating over posterior half of anal fin base, anterior margin slightly convex, narrowly rounded apex, posterior margin almost upright; rear corner of second dorsal almost forming a right angle, free rear tip rounded, inner margin shorter than first dorsal inner margin, only 20.2% of its base length. Anal fin subtriangular, broadly rounded apex, larger than second dorsal fin; anal fin base 10.7% TL, slightly greater to interdorsal space by 1.1 times. Caudal fin relatively short, dorsal caudal margin length 20.5% TL, with ventral caudal lobe moderately developed distally; terminal lobe well developed but relatively small, fan-like with convex posterior margin, terminal caudal margin length 3.5% TL.
</paragraph>
<caption id="DECB6662886FFF94FF5C0DF1B7B2FF22" box="[159,397,160,182]" pageId="5" pageNumber="536">
<paragraph id="8A0B36EA886FFF94FF5C0DF1B7B2FF22" blockId="5.[159,397,160,182]" box="[159,397,160,182]" pageId="5" pageNumber="536">
<emphasis id="B8C0EAF8886FFF94FF5C0DF1B733FF22" bold="true" box="[159,268,160,182]" pageId="5" pageNumber="536">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8A0B36EA886FFF94FF5C0D9BB4C8F9B4" pageId="5" pageNumber="536">
<table id="F8B4C44A886F006EFF5C0D9BB231F9B4" box="[159,1038,202,1568]" colsContinueFrom="4.[159,1038,234,1986]" gridcols="2" gridrows="33" pageId="5" pageNumber="536">
<tr id="348434A8886F006EFF5C0D9BB231FF74" box="[159,1038,202,224]" gridrow="0" pageId="5" pageNumber="536">
<th id="77555DD4886F006EFF5C0D9BB43FFF74" box="[159,512,202,224]" gridcol="0" gridrow="0" pageId="5" pageNumber="536">Species</th>
<th id="77555DD4886F006EFD080D9BB231FF74" box="[715,1038,202,224]" gridcol="1" gridrow="0" pageId="5" pageNumber="536">
<taxonomicName id="4DB44D69886FFF94FD0F0D9AB5F0FF4B" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[716,975,202,224]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886FFF94FD0F0D9AB5F0FF4B" box="[716,975,202,224]" italics="true" pageId="5" pageNumber="536">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886FFF94FC160D9BB231FF74" box="[981,1038,202,224]" pageId="5" pageNumber="536" rank="species">
<emphasis id="B8C0EAF8886FFF94FC160D9BB237FF74" bold="true" box="[981,1032,202,224]" pageId="5" pageNumber="536">n. sp</emphasis>
.
</taxonomicNameLabel>
</th>
</tr>
<tr id="348434A8886F006EFF5C0DA5B231FE9E" box="[159,1038,244,266]" gridrow="1" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0DA5B43FFE9E" box="[159,512,244,266]" gridcol="0" gridrow="1" pageId="5" pageNumber="536">Second dorsal-fin inner margin</td>
<td id="77555DD4886F006EFD080DA5B231FE9E" box="[715,1038,244,266]" gridcol="1" gridrow="1" pageId="5" pageNumber="536">1.74</td>
</tr>
<tr id="348434A8886F006EFF5C0C4CB231FEA7" box="[159,1038,285,307]" gridrow="2" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0C4CB43FFEA7" box="[159,512,285,307]" gridcol="0" gridrow="2" pageId="5" pageNumber="536">Second dorsal-fin posterior margin</td>
<td id="77555DD4886F006EFD080C4CB231FEA7" box="[715,1038,285,307]" gridcol="1" gridrow="2" pageId="5" pageNumber="536">4.06</td>
</tr>
<tr id="348434A8886F006EFF5C0C19B231FECA" box="[159,1038,328,350]" gridrow="3" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0C19B43FFECA" box="[159,512,328,350]" gridcol="0" gridrow="3" pageId="5" pageNumber="536">Pectoral-fin anterior margin</td>
<td id="77555DD4886F006EFD080C19B231FECA" box="[715,1038,328,350]" gridcol="1" gridrow="3" pageId="5" pageNumber="536">9.56</td>
</tr>
<tr id="348434A8886F006EFF5C0C23B231FE1C" box="[159,1038,370,392]" gridrow="4" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0C23B43FFE1C" box="[159,512,370,392]" gridcol="0" gridrow="4" pageId="5" pageNumber="536">Pectoral-fin base</td>
<td id="77555DD4886F006EFD080C23B231FE1C" box="[715,1038,370,392]" gridcol="1" gridrow="4" pageId="5" pageNumber="536">5.32</td>
</tr>
<tr id="348434A8886F006EFF5C0CCDB231FE26" box="[159,1038,412,434]" gridrow="5" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0CCDB43FFE26" box="[159,512,412,434]" gridcol="0" gridrow="5" pageId="5" pageNumber="536">Pectoral-fin inner margin</td>
<td id="77555DD4886F006EFD080CCDB231FE26" box="[715,1038,412,434]" gridcol="1" gridrow="5" pageId="5" pageNumber="536">3.99</td>
</tr>
<tr id="348434A8886F006EFF5C0C97B231FE48" box="[159,1038,454,476]" gridrow="6" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0C97B43FFE48" box="[159,512,454,476]" gridcol="0" gridrow="6" pageId="5" pageNumber="536">Pectoral-fin posterior margin</td>
<td id="77555DD4886F006EFD080C97B231FE48" box="[715,1038,454,476]" gridcol="1" gridrow="6" pageId="5" pageNumber="536">6.41</td>
</tr>
<tr id="348434A8886F006EFF5C0CA1B231FD92" box="[159,1038,496,518]" gridrow="7" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0CA1B43FFD92" box="[159,512,496,518]" gridcol="0" gridrow="7" pageId="5" pageNumber="536">Pectoral-pelvic space</td>
<td id="77555DD4886F006EFD080CA1B231FD92" box="[715,1038,496,518]" gridcol="1" gridrow="7" pageId="5" pageNumber="536">27.22</td>
</tr>
<tr id="348434A8886F006EFF5C0F4BB231FDA4" box="[159,1038,538,560]" gridrow="8" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0F4BB43FFDA4" box="[159,512,538,560]" gridcol="0" gridrow="8" pageId="5" pageNumber="536">Pelvic-fin anterior margin</td>
<td id="77555DD4886F006EFD080F4BB231FDA4" box="[715,1038,538,560]" gridcol="1" gridrow="8" pageId="5" pageNumber="536">6.77</td>
</tr>
<tr id="348434A8886F006EFF5C0F15B231FDCE" box="[159,1038,580,602]" gridrow="9" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0F15B43FFDCE" box="[159,512,580,602]" gridcol="0" gridrow="9" pageId="5" pageNumber="536">Pelvic-fin base</td>
<td id="77555DD4886F006EFD080F15B231FDCE" box="[715,1038,580,602]" gridcol="1" gridrow="9" pageId="5" pageNumber="536">5.53</td>
</tr>
<tr id="348434A8886F006EFF5C0F3FB231FD10" box="[159,1038,622,644]" gridrow="10" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0F3FB43FFD10" box="[159,512,622,644]" gridcol="0" gridrow="10" pageId="5" pageNumber="536">Pelvic-fin length</td>
<td id="77555DD4886F006EFD080F3FB231FD10" box="[715,1038,622,644]" gridcol="1" gridrow="10" pageId="5" pageNumber="536">9.16</td>
</tr>
<tr id="348434A8886F006EFF5C0FC9B231FD3A" box="[159,1038,664,686]" gridrow="11" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0FC9B43FFD3A" box="[159,512,664,686]" gridcol="0" gridrow="11" pageId="5" pageNumber="536">Pelvic-fin inner margin length</td>
<td id="77555DD4886F006EFD080FC9B231FD3A" box="[715,1038,664,686]" gridcol="1" gridrow="11" pageId="5" pageNumber="536">3.64</td>
</tr>
<tr id="348434A8886F006EFF5C0F93B231FD4C" box="[159,1038,706,728]" gridrow="12" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0F93B43FFD4C" box="[159,512,706,728]" gridcol="0" gridrow="12" pageId="5" pageNumber="536">Pelvic-fin posterior margin length</td>
<td id="77555DD4886F006EFD080F93B231FD4C" box="[715,1038,706,728]" gridcol="1" gridrow="12" pageId="5" pageNumber="536">4.59</td>
</tr>
<tr id="348434A8886F006EFF5C0FBDB231FC96" box="[159,1038,748,770]" gridrow="13" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0FBDB43FFC96" box="[159,512,748,770]" gridcol="0" gridrow="13" pageId="5" pageNumber="536">Pelvic-anal space</td>
<td id="77555DD4886F006EFD080FBDB231FC96" box="[715,1038,748,770]" gridcol="1" gridrow="13" pageId="5" pageNumber="536">6.96</td>
</tr>
<tr id="348434A8886F006EFF5C0E47B231FCB8" box="[159,1038,790,812]" gridrow="14" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0E47B43FFCB8" box="[159,512,790,812]" gridcol="0" gridrow="14" pageId="5" pageNumber="536">Anal-fin anterior margin</td>
<td id="77555DD4886F006EFD080E47B231FCB8" box="[715,1038,790,812]" gridcol="1" gridrow="14" pageId="5" pageNumber="536">9.32</td>
</tr>
<tr id="348434A8886F006EFF5C0E11B231FCC2" box="[159,1038,832,854]" gridrow="15" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0E11B43FFCC2" box="[159,512,832,854]" gridcol="0" gridrow="15" pageId="5" pageNumber="536">Anal-fin base</td>
<td id="77555DD4886F006EFD080E11B231FCC2" box="[715,1038,832,854]" gridcol="1" gridrow="15" pageId="5" pageNumber="536">10.74</td>
</tr>
<tr id="348434A8886F006EFF5C0E3BB231FC14" box="[159,1038,874,896]" gridrow="16" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0E3BB43FFC14" box="[159,512,874,896]" gridcol="0" gridrow="16" pageId="5" pageNumber="536">Anal-fin height</td>
<td id="77555DD4886F006EFD080E3BB231FC14" box="[715,1038,874,896]" gridcol="1" gridrow="16" pageId="5" pageNumber="536">3.88</td>
</tr>
<tr id="348434A8886F006EFF5C0EC5B231FC3E" box="[159,1038,916,938]" gridrow="17" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0EC5B43FFC3E" box="[159,512,916,938]" gridcol="0" gridrow="17" pageId="5" pageNumber="536">Anal-fin length</td>
<td id="77555DD4886F006EFD080EC5B231FC3E" box="[715,1038,916,938]" gridcol="1" gridrow="17" pageId="5" pageNumber="536">11.54</td>
</tr>
<tr id="348434A8886F006EFF5C0EEFB231FC40" box="[159,1038,958,980]" gridrow="18" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0EEFB43FFC40" box="[159,512,958,980]" gridcol="0" gridrow="18" pageId="5" pageNumber="536">Anal-fin inner margin</td>
<td id="77555DD4886F006EFD080EEFB231FC40" box="[715,1038,958,980]" gridcol="1" gridrow="18" pageId="5" pageNumber="536">0.84</td>
</tr>
<tr id="348434A8886F006EFF5C0EB9B231FC6A" box="[159,1038,1000,1022]" gridrow="19" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0EB9B43FFC6A" box="[159,512,1000,1022]" gridcol="0" gridrow="19" pageId="5" pageNumber="536">Anal-fin posterior margin</td>
<td id="77555DD4886F006EFD080EB9B231FC6A" box="[715,1038,1000,1022]" gridcol="1" gridrow="19" pageId="5" pageNumber="536">5.39</td>
</tr>
<tr id="348434A8886F006EFF5C0943B231FBBC" box="[159,1038,1042,1064]" gridrow="20" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0943B43FFBBC" box="[159,512,1042,1064]" gridcol="0" gridrow="20" pageId="5" pageNumber="536">Dorsal-caudal space</td>
<td id="77555DD4886F006EFD080943B231FBBC" box="[715,1038,1042,1064]" gridcol="1" gridrow="20" pageId="5" pageNumber="536">1.99</td>
</tr>
<tr id="348434A8886F006EFF5C096DB231FBC6" box="[159,1038,1084,1106]" gridrow="21" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C096DB43FFBC6" box="[159,512,1084,1106]" gridcol="0" gridrow="21" pageId="5" pageNumber="536">Anal-caudal space</td>
<td id="77555DD4886F006EFD08096DB231FBC6" box="[715,1038,1084,1106]" gridcol="1" gridrow="21" pageId="5" pageNumber="536">4.52</td>
</tr>
<tr id="348434A8886F006EFF5C0937B231FBE8" box="[159,1038,1126,1148]" gridrow="22" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0937B43FFBE8" box="[159,512,1126,1148]" gridcol="0" gridrow="22" pageId="5" pageNumber="536">Caudal peduncle height</td>
<td id="77555DD4886F006EFD080937B231FBE8" box="[715,1038,1126,1148]" gridcol="1" gridrow="22" pageId="5" pageNumber="536">2.18</td>
</tr>
<tr id="348434A8886F006EFF5C09C1B231FB32" box="[159,1038,1168,1190]" gridrow="23" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C09C1B43FFB32" box="[159,512,1168,1190]" gridcol="0" gridrow="23" pageId="5" pageNumber="536">Caudal peduncle width</td>
<td id="77555DD4886F006EFD0809C1B231FB32" box="[715,1038,1168,1190]" gridcol="1" gridrow="23" pageId="5" pageNumber="536">1.46</td>
</tr>
<tr id="348434A8886F006EFF5C09EBB231FB44" box="[159,1038,1210,1232]" gridrow="24" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C09EBB43FFB44" box="[159,512,1210,1232]" gridcol="0" gridrow="24" pageId="5" pageNumber="536">Dorsal caudal-fin margin</td>
<td id="77555DD4886F006EFD0809EBB231FB44" box="[715,1038,1210,1232]" gridcol="1" gridrow="24" pageId="5" pageNumber="536">20.49</td>
</tr>
<tr id="348434A8886F006EFF5C09B5B231FB6E" box="[159,1038,1252,1274]" gridrow="25" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C09B5B43FFB6E" box="[159,512,1252,1274]" gridcol="0" gridrow="25" pageId="5" pageNumber="536">Preventral caudal-fin margin</td>
<td id="77555DD4886F006EFD0809B5B231FB6E" box="[715,1038,1252,1274]" gridcol="1" gridrow="25" pageId="5" pageNumber="536">12.36</td>
</tr>
<tr id="348434A8886F006EFF5C085FB231FAB0" box="[159,1038,1294,1316]" gridrow="26" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C085FB43FFAB0" box="[159,512,1294,1316]" gridcol="0" gridrow="26" pageId="5" pageNumber="536">Subterminal caudal-fin margin</td>
<td id="77555DD4886F006EFD08085FB231FAB0" box="[715,1038,1294,1316]" gridcol="1" gridrow="26" pageId="5" pageNumber="536">3.58</td>
</tr>
<tr id="348434A8886F006EFF5C0869B231FADA" box="[159,1038,1336,1358]" gridrow="27" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0869B43FFADA" box="[159,512,1336,1358]" gridcol="0" gridrow="27" pageId="5" pageNumber="536">Terminal caudal-fin margin</td>
<td id="77555DD4886F006EFD080869B231FADA" box="[715,1038,1336,1358]" gridcol="1" gridrow="27" pageId="5" pageNumber="536">3.53</td>
</tr>
<tr id="348434A8886F006EFF5C0833B231FAEC" box="[159,1038,1378,1400]" gridrow="28" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0833B43FFAEC" box="[159,512,1378,1400]" gridcol="0" gridrow="28" pageId="5" pageNumber="536">Clasper outer length</td>
<td id="77555DD4886F006EFD080833B231FAEC" box="[715,1038,1378,1400]" gridcol="1" gridrow="28" pageId="5" pageNumber="536">5.43</td>
</tr>
<tr id="348434A8886F006EFF5C08DDB231FA36" box="[159,1038,1420,1442]" gridrow="29" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C08DDB43FFA36" box="[159,512,1420,1442]" gridcol="0" gridrow="29" pageId="5" pageNumber="536">Clasper inner length</td>
<td id="77555DD4886F006EFD0808DDB231FA36" box="[715,1038,1420,1442]" gridcol="1" gridrow="29" pageId="5" pageNumber="536">8.78</td>
</tr>
<tr id="348434A8886F006EFF5C08E7B231FA58" box="[159,1038,1462,1484]" gridrow="30" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C08E7B43FFA58" box="[159,512,1462,1484]" gridcol="0" gridrow="30" pageId="5" pageNumber="536">Clasper base width</td>
<td id="77555DD4886F006EFD0808E7B231FA58" box="[715,1038,1462,1484]" gridcol="1" gridrow="30" pageId="5" pageNumber="536">1.62</td>
</tr>
<tr id="348434A8886F006EFF5C08B1B231FA62" box="[159,1038,1504,1526]" gridrow="31" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C08B1B43FFA62" box="[159,512,1504,1526]" gridcol="0" gridrow="31" pageId="5" pageNumber="536">Upper caudal crests length</td>
<td id="77555DD4886F006EFD0808B1B231FA62" box="[715,1038,1504,1526]" gridcol="1" gridrow="31" pageId="5" pageNumber="536">10.77</td>
</tr>
<tr id="348434A8886F006EFF5C0B5BB231F9B4" box="[159,1038,1546,1568]" gridrow="32" pageId="5" pageNumber="536">
<td id="77555DD4886F006EFF5C0B5BB43FF9B4" box="[159,512,1546,1568]" gridcol="0" gridrow="32" pageId="5" pageNumber="536">Lower caudal crests length</td>
<td id="77555DD4886F006EFD080B5BB231F9B4" box="[715,1038,1546,1568]" gridcol="1" gridrow="32" pageId="5" pageNumber="536">9.31</td>
</tr>
</table>
</paragraph>
<caption id="DECB6662886CFF97FF540EE2B333FC5D" box="[151,1292,947,969]" httpUri="https://zenodo.org/record/1227971/files/figure.png" pageId="6" pageNumber="537" startId="6.[151,250,947,969]" targetBox="[210,1380,161,925]" targetPageId="6">
<paragraph id="8A0B36EA886CFF97FF540EE2B333FC5D" blockId="6.[151,1292,947,969]" box="[151,1292,947,969]" pageId="6" pageNumber="537">
<emphasis id="B8C0EAF8886CFF97FF540EE2B72DFC5D" bold="true" box="[151,274,947,969]" pageId="6" pageNumber="537">FIGURE 4.</emphasis>
Symphysial teeth of 
<taxonomicName id="4DB44D69886CFF97FE290EE5B4D2FC5C" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[490,749,947,969]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886CFF97FE290EE5B4D2FC5C" box="[490,749,947,969]" italics="true" pageId="6" pageNumber="537">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886CFF97FD370EE2B513FC5D" box="[756,812,947,969]" pageId="6" pageNumber="537" rank="species">
<emphasis id="B8C0EAF8886CFF97FD370EE2B519FC5D" bold="true" box="[756,806,947,969]" pageId="6" pageNumber="537">n. sp</emphasis>
.
</taxonomicNameLabel>
holotype adult male 548 mm TL (NCIP 6567).
</paragraph>
</caption>
<paragraph id="8A0B36EA886CFF97FF040EA4B4F3FBC2" blockId="6.[151,1437,1013,2013]" pageId="6" pageNumber="537">Total vertebral counts 130, monospondylous precaudal (MP) vertebrae 42, precaudal Monospondylous vertebrae 90, diplospondylous centra (DP) 48. Transition between the monospondylous and diplospondylous centra behind pelvic fin girdle and over clasper channels.</paragraph>
<paragraph id="8A0B36EA886CFF97FF040930B2F1FB72" blockId="6.[151,1437,1013,2013]" pageId="6" pageNumber="537">
<emphasis id="B8C0EAF8886CFF97FF040930B770FBEE" bold="true" box="[199,335,1121,1146]" pageId="6" pageNumber="537">Coloration.</emphasis>
Specimen prior to preservation is plain medium brownish, dorsal side of pectoral fins and posterior of caudal slightly darker, ventral side of pectoral fins with conspicuous dark margin; edges of dorsal, pelvic and anal fins slightly darker, abdomen paler. Roof of mouth is blackish with darker pores. Claspers and pelvic fins are whitish at insertions. After preservation, body and fins are uniformly dark brown.
</paragraph>
<paragraph id="8A0B36EA886CFF97FF0409A0B5CAFA9E" blockId="6.[151,1437,1013,2013]" box="[199,1013,1265,1291]" pageId="6" pageNumber="537">
<emphasis id="B8C0EAF8886CFF97FF0409A0B6C1FA9E" bold="true" box="[199,254,1265,1290]" pageId="6" pageNumber="537">Size.</emphasis>
The only known specimen (
<typeStatus id="550F8848886CFF97FDFF09A3B49DFA9F" box="[572,674,1266,1291]" pageId="6" pageNumber="537" type="holotype">holotype</typeStatus>
) is an adult male 
<quantity id="4D4C9B0F886CFF97FCAB09A3B5FAFA9E" box="[872,965,1266,1291]" metricMagnitude="-1" metricUnit="m" metricValue="5.4799999999999995" pageId="6" pageNumber="537" unit="mm" value="548.0">548 mm</quantity>
TL.
</paragraph>
</subSubSection>
<subSubSection id="C2AE6561886CFF97FF040849B480FAC0" pageId="6" pageNumber="537" type="distribution">
<paragraph id="8A0B36EA886CFF97FF040849B480FAC0" blockId="6.[151,1437,1013,2013]" pageId="6" pageNumber="537">
<emphasis id="B8C0EAF8886CFF97FF040849B75FFAA5" bold="true" box="[199,352,1304,1329]" pageId="6" pageNumber="537">Distribution.</emphasis>
Known only from south of Sumbawa Island, 
<collectingCountry id="F2A3767A886CFF97FCBD0849B5D4FAA5" box="[894,1003,1304,1329]" name="Indonesia" pageId="6" pageNumber="537">Indonesia</collectingCountry>
(
<geoCoordinate id="EF80502D886CFF97FC3C0848B26EFAA5" box="[1023,1105,1301,1329]" direction="south" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-9.033334">9o 02’S</geoCoordinate>
, 
<geoCoordinate id="EF80502D886CFF97FBA20848B2EDFAA5" box="[1121,1234,1301,1329]" direction="east" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="117.916664">117o 55’E</geoCoordinate>
). Possibly occurs more widespread in eastern Indian Ocean waters.
</paragraph>
</subSubSection>
<subSubSection id="C2AE6561886CFF99FF040830B7E9FE39" lastPageId="8" lastPageNumber="539" pageId="6" pageNumber="537" type="etymology">
<paragraph id="8A0B36EA886CFF97FF040830B2B1FA55" blockId="6.[151,1437,1013,2013]" pageId="6" pageNumber="537">
<emphasis id="B8C0EAF8886CFF97FF040830B771FAED" bold="true" box="[199,334,1377,1401]" pageId="6" pageNumber="537">Etymology.</emphasis>
The specific name is a combination of the Latin 
<emphasis id="B8C0EAF8886CFF97FCB20830B598FAEC" box="[881,935,1377,1400]" italics="true" pageId="6" pageNumber="537">nigri</emphasis>
(black or dark) and 
<emphasis id="B8C0EAF8886CFF97FB480830B2D8FAED" box="[1163,1255,1377,1401]" italics="true" pageId="6" pageNumber="537">palatum</emphasis>
(roof of mouth) in allusion to the roof of its mouth being a very distinctive blackish colored. The proposed English common name is Indonesian Filetail Catshark and the proposed Indonesian common name is Hiu minyak.
</paragraph>
<paragraph id="8A0B36EA886CFF97FF04089DB4EAF9E1" blockId="6.[151,1437,1013,2013]" pageId="6" pageNumber="537">
<emphasis id="B8C0EAF8886CFF97FF04089DB75EFA71" bold="true" box="[199,353,1484,1509]" pageId="6" pageNumber="537">Comparison.</emphasis>
<taxonomicName id="4DB44D69886CFF97FEAA089CB4B8FA71" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[361,647,1485,1509]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886CFF97FEAA089CB4B8FA71" box="[361,647,1485,1509]" italics="true" pageId="6" pageNumber="537">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886CFF97FD4C089CB526FA71" box="[655,793,1484,1509]" pageId="6" pageNumber="537" rank="species">new species</taxonomicNameLabel>
, can be separated from all other 
<taxonomicName id="4DB44D69886CFF97FB44089CB332FA70" authorityName="Garman" authorityYear="1906" box="[1159,1293,1485,1508]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B8C0EAF8886CFF97FB44089CB332FA70" box="[1159,1293,1485,1508]" italics="true" pageId="6" pageNumber="537">Parmaturus</emphasis>
</taxonomicName>
species by a combination of characteristics including a prominent dorsal and ventral caudal crest, well-developed labial furrows with the uppers and lowers of equal lengths, mouth roof blackish with dark pores, position of the first dorsal fin origin, and tooth counts. The new species has distinctly lower total upper and lower tooth counts (52/48) than all other 
<taxonomicName id="4DB44D69886CFF97FF140B0CB762F9E0" authorityName="Garman" authorityYear="1906" box="[215,349,1629,1652]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B8C0EAF8886CFF97FF140B0CB762F9E0" box="[215,349,1629,1652]" italics="true" pageId="6" pageNumber="537">Parmaturus</emphasis>
</taxonomicName>
species (&gt;60 uppers and lowers).
</paragraph>
<paragraph id="8A0B36EA886CFF97FF040BD1B4BEF849" blockId="6.[151,1437,1013,2013]" pageId="6" pageNumber="537">
The closest geographic congener, and only other 
<taxonomicName id="4DB44D69886CFF97FD2A0BD0B550F90C" authorityName="Garman" authorityYear="1906" box="[745,879,1665,1688]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B8C0EAF8886CFF97FD2A0BD0B550F90C" box="[745,879,1665,1688]" italics="true" pageId="6" pageNumber="537">Parmaturus</emphasis>
</taxonomicName>
species known to occur in Indonesian waters is 
<taxonomicName id="4DB44D69886CFF97FA440BD0B6D5F929" authorityName="Seret and Last" authorityYear="2007" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanatus">
<emphasis id="B8C0EAF8886CFF97FA440BD0B6D5F929" italics="true" pageId="6" pageNumber="537">P. lanatus</emphasis>
</taxonomicName>
, which was described from a single juvenile specimen (
<quantity id="4D4C9B0F886CFF97FCAA0BF5B5F6F928" box="[873,969,1700,1725]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="537" unit="mm" value="360.0">360 mm</quantity>
TL) collected from the continental slope off Tanimbar Island in the Arafura Sea (Séret &amp; Last, 2007). However, although the new species is an adult male (
<quantity id="4D4C9B0F886CFF97FF5C0BBDB6C1F890" box="[159,254,1772,1797]" metricMagnitude="-1" metricUnit="m" metricValue="5.4799999999999995" pageId="6" pageNumber="537" unit="mm" value="548.0">548 mm</quantity>
TL) it can be easily separated from 
<taxonomicName id="4DB44D69886CFF97FD540BBCB539F891" authorityName="Seret and Last" authorityYear="2007" box="[663,774,1773,1797]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="lanatus">
<emphasis id="B8C0EAF8886CFF97FD540BBCB539F891" box="[663,774,1773,1797]" italics="true" pageId="6" pageNumber="537">P. lanatus</emphasis>
</taxonomicName>
by several characteristics including tooth counts (52/48 vs 94/92); presence of a crest of enlarged dermal denticles along the upper and ventral caudal margins; presence of prominent upper and lower labial furrow vs rudimentary or absent labial furrows; a first dorsal fin originating more than one-half it body length (53.8% TL) vs originating less than body length (44.4% TL); roof of mouth blackish with darker pores vs grayish; pectoral-pelvic space is considerably longer (27% vs. 17% TL); interorbital width in head length is greater (3.4 vs. 2.6 times); and abdomen length longer than head length (1.3) vs (0.8); and a slightly higher precaudal vertebral count (90 vs 85).
</paragraph>
<caption id="DECB6662886DFF96FF540F9DB6E0FC95" httpUri="https://zenodo.org/record/1227973/files/figure.png" pageId="7" pageNumber="538" startId="7.[151,250,716,738]" targetBox="[151,1436,193,695]" targetPageId="7">
<paragraph id="8A0B36EA886DFF96FF540F9DB6E0FC95" blockId="7.[151,1435,716,769]" pageId="7" pageNumber="538">
<emphasis id="B8C0EAF8886DFF96FF540F9DB72AFD76" bold="true" box="[151,277,716,738]" pageId="7" pageNumber="538">FIGURE 5.</emphasis>
<taxonomicName id="4DB44D69886DFF96FEDE0F9CB41CFD75" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[285,547,716,738]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886DFF96FEDE0F9CB41CFD75" box="[285,547,716,738]" italics="true" pageId="7" pageNumber="538">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886DFF96FDE80F9DB459FD76" box="[555,614,716,738]" pageId="7" pageNumber="538" rank="species">
<emphasis id="B8C0EAF8886DFF96FDE80F9DB45FFD76" bold="true" box="[555,608,716,738]" pageId="7" pageNumber="538">n. sp</emphasis>
.
</taxonomicNameLabel>
, close-up of denticles on flank (left) and caudal crests of dorsal caudal margin (right).
</paragraph>
</caption>
<caption id="DECB6662886DFF96FF540ADFB720F876" httpUri="https://zenodo.org/record/1227975/files/figure.png" pageId="7" pageNumber="538" startId="7.[151,250,1934,1956]" targetBox="[340,1180,827,1899]" targetPageId="7">
<paragraph id="8A0B36EA886DFF96FF540ADFB720F876" blockId="7.[151,1437,1934,2018]" pageId="7" pageNumber="538">
<emphasis id="B8C0EAF8886DFF96FF540ADFB72DF830" bold="true" box="[151,274,1934,1956]" pageId="7" pageNumber="538">FIGURE 6.</emphasis>
Dorsal view of claspers of preserved specimen of 
<taxonomicName id="4DB44D69886DFF96FCCD0ADEB22EF837" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[782,1041,1934,1956]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF8886DFF96FCCD0ADEB22EF837" box="[782,1041,1934,1956]" italics="true" pageId="7" pageNumber="538">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F35783886DFF96FBD40ADFB26FF830" box="[1047,1104,1934,1956]" pageId="7" pageNumber="538" rank="species">
<emphasis id="B8C0EAF8886DFF96FBD40ADFB275F830" bold="true" box="[1047,1098,1934,1956]" pageId="7" pageNumber="538">n. sp</emphasis>
.
</taxonomicNameLabel>
holotype adult male 548 mm TL (NCIP 6567). Abbreviations: AP: apopyle; CG: clasper groove; HP: hypopyle; CRH: cover rhipidion; RH: rhipidion; ERH: exorhipidion.
</paragraph>
</caption>
<paragraph id="8A0B36EA8862FF99FF040DC8B7E9FE39" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
<bibRef id="5314A21C8862FF99FF040DC8B7E9FE39" pageId="8" pageNumber="539" refId="ref2630" type="journal article">
<title id="F8B0DDDC8862FF99FF040DC8B29FFF24" box="[199,1184,151,176]" jopScore="0.008184114" pageId="8" pageNumber="539">
<emphasis id="B8C0EAF88862FF99FF040DC8B785FF24" box="[199,442,152,176]" italics="true" pageId="8" pageNumber="539">Parmaturus albipenis</emphasis>
was described from a single adult male (
<quantity id="4D4C9B0F8862FF99FCBC0DC6B5E2FF3B" box="[895,989,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="4.15" pageId="8" pageNumber="539" unit="mm" value="415.0">415 mm</quantity>
TL) collected off
</title>
<collectingCountry id="F2A3767A8862FF99FB650DC9B36DFF24" box="[1190,1362,151,176]" name="New Caledonia" pageId="8" pageNumber="539">
<author id="5209DE0F8862FF99FB650DC9B36DFF24" box="[1190,1362,151,176]" fns="N" npo="FL" pageId="8" pageNumber="539">New Caledonia</author>
</collectingCountry>
<journalOrPublisher id="50753C268862FF99FA9A0DC9B350FF63" jopScore="0.19919395" pageId="8" pageNumber="539">
(Séret &amp; Last, 2007), but it can be easily separated from 
<emphasis id="B8C0EAF88862FF99FD0A0DECB549FF40" box="[713,886,188,212]" italics="true" pageId="8" pageNumber="539">P. nigripalatum</emphasis>
<emphasis id="B8C0EAF88862FF99FCBD0DEDB589FF40" bold="true" box="[894,950,188,212]" pageId="8" pageNumber="539">n. sp</emphasis>
. by the absence of an enlarged caudal crest of dermal denticles along the upper dorsal and ventral margins vs presence; a lower precaudal vertebral count
</journalOrPublisher>
(
<part id="FE109C778862FF99FABC0DB1B3A3FF6C" box="[1407,1436,224,248]" pageId="8" pageNumber="539" type="volume">79</part>
vs 90), a much higher upper tooth count (&gt;130 vs 
<pagination id="B37DC81E8862FF99FD020C55B4DEFE89" box="[705,737,260,285]" pageId="8" pageNumber="539" type="pagination">52</pagination>
); a first dorsal fin originating less than one-half its body length (46.2% TL) vs more than half its body length (53.8% TL); dorsal fins similar in size vs second dorsal being noticeably larger; anterior margins of both dorsal fins similar in length vs second dorsal fin anterior margin longer than first; lower labial furrows longer than uppers vs labial furrows of similar length; and a whitish colored mouth roof vs blackish mouth roof.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="C2AE65618862FF99FF040CE8B736F8E4" pageId="8" pageNumber="539" type="description">
<paragraph id="8A0B36EA8862FF99FF040CE8B248FD33" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
<bibRef id="5314A21C8862FF99FF040CE8B248FD33" pageId="8" pageNumber="539" refId="ref2799" type="journal article">
<title id="F8B0DDDC8862FF99FF040CE8B7C0FE44" box="[199,511,440,464]" jopScore="0.0138716325" pageId="8" pageNumber="539">
<emphasis id="B8C0EAF88862FF99FF040CE8B7D9FE44" box="[199,486,440,464]" italics="true" pageId="8" pageNumber="539">Parmaturus nigripalatum</emphasis>
<emphasis id="B8C0EAF88862FF99FE2D0CE9B7C0FE44" bold="true" box="[494,511,440,464]" pageId="8" pageNumber="539">n</emphasis>
</title>
. sp. differs from 
<emphasis id="B8C0EAF88862FF99FD000CE8B5B6FE44" box="[707,905,440,464]" italics="true" pageId="8" pageNumber="539">P. albimarginatus</emphasis>
, which was described from a single adult male (
<quantity id="4D4C9B0F8862FF99FF5C0C8DB6C2FE60" box="[159,253,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="5.77" pageId="8" pageNumber="539" unit="mm" value="577.0">577 mm</quantity>
TL) collected off 
<collectingCountry id="F2A3767A8862FF99FE040C8CB44BFE61" box="[455,628,476,501]" name="New Caledonia" pageId="8" pageNumber="539">
<author id="5209DE0F8862FF99FE040C8CB44BFE61" box="[455,628,476,501]" fns="N" npo="FL" pageId="8" pageNumber="539">New Caledonia</author>
</collectingCountry>
<journalOrPublisher id="50753C268862FF99FDB80C8DB4E5FD8C" jopScore="0.0951783" pageId="8" pageNumber="539" type="_" typeScore="4">(Séret &amp; Last, 2007) by a total upper and lower tooth count that is much lower (52/48 vs 92/92); a slightly lower precaudal</journalOrPublisher>
(
<part id="FE109C778862FF99FD280F51B537FD8C" box="[747,776,512,536]" pageId="8" pageNumber="539" type="volume">90</part>
vs 
<pagination id="B37DC81E8862FF99FCF70CAEB56EFD8C" box="[820,849,511,536]" pageId="8" pageNumber="539" type="pagination">95</pagination>
) and total vertebral (130 vs 136) counts; prepelvic length longer (49.3% vs. 46.9% TL); preanal length longer (62.2% vs. 59.1% TL); longer snout-vent length (51.6% vs. 49.7% TL); greater eye height (2.1% vs. 0.5% TL); a longer prenarial (4.7% vs. 3.7% TL); a narrower head width (10.8% vs. 12.9% TL); labial furrows about equal vs lowers longer than uppers; and darker colored posterior margins of dorsal, caudal and anal fins, compared to white margins in 
<emphasis id="B8C0EAF88862FF99FC690FC0B24FFD3C" box="[938,1136,656,680]" italics="true" pageId="8" pageNumber="539">P. albimarginatus</emphasis>
.
</bibRef>
</paragraph>
<paragraph id="8A0B36EA8862FF99FF040FE4B478FBCC" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
<bibRef id="5314A21C8862FF99FF040FE4B478FBCC" pageId="8" pageNumber="539" refId="ref2987" type="journal article">
<title id="F8B0DDDC8862FF99FF040FE4B437FD58" box="[199,520,692,716]" jopScore="0.014969108" pageId="8" pageNumber="539">
<emphasis id="B8C0EAF88862FF99FF040FE4B7D4FD58" box="[199,491,692,716]" italics="true" pageId="8" pageNumber="539">Parmaturus nigripalatum</emphasis>
<emphasis id="B8C0EAF88862FF99FE340FE5B437FD58" bold="true" box="[503,520,692,716]" pageId="8" pageNumber="539">n</emphasis>
</title>
. sp. differs from 
<emphasis id="B8C0EAF88862FF99FD1E0FE4B504FD58" box="[733,827,692,716]" italics="true" pageId="8" pageNumber="539">P. bigus</emphasis>
, described from a single adult female collected off northeastern 
<collectingCountry id="F2A3767A8862FF99FEEA0F86B7ACFD64" box="[297,403,727,752]" name="Australia" pageId="8" pageNumber="539">Australia</collectingCountry>
, south of 
<author id="5209DE0F8862FF99FDC10F89B4F3FD64" box="[514,716,727,752]" fns="N" npo="FL" pageId="8" pageNumber="539">Saumarez Plateau</author>
<journalOrPublisher id="50753C268862FF99FD100F89B334FD7B" box="[723,1291,727,752]" jopScore="0.40256032" pageId="8" pageNumber="539" type="_" typeScore="4">(Séret &amp; Last, 2007) by a lower upper tooth count</journalOrPublisher>
(
<part id="FE109C778862FF99FAD80F86B307FD64" box="[1307,1336,727,752]" pageId="8" pageNumber="539" type="volume">52</part>
vs 
<pagination id="B37DC81E8862FF99FAA20F89B3B0FD64" box="[1377,1423,728,752]" pageId="8" pageNumber="539" type="pagination">120</pagination>
); lower precaudal (90 vs 102) and total (130 vs 144) vertebral counts; shorter precaudal length (75.6% vs. 84% TL); greater first and second fin dorsal bases (6.5% vs. 4.9% TL and 8.6% vs. 7.3% TL); longer first and second dorsal fin anterior margins (8.6% vs. 5.2% TL and 10.8% vs. 9.3% TL, respectively); shorter pelvic-anal space (7.0% vs. 8.1% TL); longer anal fin length (11.5% vs. 9.7% TL); narrower dorsal-caudal space (2.0% vs. 5.7% TL); longer preventral caudal-fin margin (12.4% vs. 7.3% TL); smaller spiracle (0.3% vs. 0.7% TL); wider mouth (9.0% vs. 6.7% TL); near equal length for upper and lower labial furrows compared to lower furrow being considerably longer that upper furrow; and mouth roof color that is blackish compared to the greyish color found in 
<emphasis id="B8C0EAF88862FF99FAC80E84B355FC78" box="[1291,1386,980,1004]" italics="true" pageId="8" pageNumber="539">P. bigus.</emphasis>
The overall body coloration (after preservation) between the new species and 
<emphasis id="B8C0EAF88862FF99FC170EA8B211FB84" box="[980,1070,1016,1040]" italics="true" pageId="8" pageNumber="539">P. bigus</emphasis>
is also informative, with former being uniform dark brown with slightly darker dorsal, pelvic and anal fins compared to the uniform plain yellowish brown, with slightly paler fin margins.
</bibRef>
</paragraph>
<paragraph id="8A0B36EA8862FF99FF040934B459FAA4" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
<taxonomicName id="4DB44D698862FF99FF040934B7D8FBE8" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[199,487,1124,1148]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88862FF99FF040934B7D8FBE8" box="[199,487,1124,1148]" italics="true" pageId="8" pageNumber="539">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F357838862FF99FE330935B40EFBE8" box="[496,561,1124,1148]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B8C0EAF88862FF99FE330935B415FBE8" bold="true" box="[496,554,1124,1148]" pageId="8" pageNumber="539">n. sp</emphasis>
.
</taxonomicNameLabel>
differs from 
<taxonomicName id="4DB44D698862FF99FD080934B559FBE8" authorityName="Hardy" authorityYear="1985" box="[715,870,1124,1148]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="macmillani">
<emphasis id="B8C0EAF88862FF99FD080934B559FBE8" box="[715,870,1124,1148]" italics="true" pageId="8" pageNumber="539">P. macmillani</emphasis>
</taxonomicName>
in the position of the first dorsal origin (54% vs. 42–48% TL); longer snout length (5.0% vs. 3.2–3.7% TL); smaller mouth width (9.0% vs. 10.3–10.8% TL); a shorter interdorsal space (9.5% vs. 10.7–11.2% TL); a shorter pectoral-pelvic space (27.2% vs. 29.4–29.5% TL); a longer pelvic-anal space (7.0% vs. 5.2–5.8% TL); slightly higher monospondylous (42 vs 38) and total (130 vs 124) vertebral counts; and almost equal length of upper and lower labial furrows compared to lower labial furrow much longer than upper in 
<emphasis id="B8C0EAF88862FF99FE060848B459FAA4" box="[453,614,1304,1328]" italics="true" pageId="8" pageNumber="539">
<taxonomicName id="4DB44D698862FF99FE060848B45CFAA4" authorityName="Hardy" authorityYear="1985" box="[453,611,1304,1328]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="macmillani">P. macmillani</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8A0B36EA8862FF99FF0D086CB365FA54" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
<taxonomicName id="4DB44D698862FF99FF0D086CB7D2FAC0" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[206,493,1340,1364]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88862FF99FF0D086CB7D2FAC0" box="[206,493,1340,1364]" italics="true" pageId="8" pageNumber="539">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F357838862FF99FE36086DB40BFAC0" box="[501,564,1340,1364]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B8C0EAF88862FF99FE36086DB412FAC0" bold="true" box="[501,557,1340,1364]" pageId="8" pageNumber="539">n. sp</emphasis>
.
</taxonomicNameLabel>
differs from 
<taxonomicName id="4DB44D698862FF99FD0A086CB59FFAC0" baseAuthorityName="Chan" baseAuthorityYear="1966" box="[713,928,1340,1364]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="melanobranchus">
<emphasis id="B8C0EAF88862FF99FD0A086CB59FFAC0" box="[713,928,1340,1364]" italics="true" pageId="8" pageNumber="539">P. melanobranchus</emphasis>
</taxonomicName>
by a first dorsal fin being about equal in size to second dorsal fin vs a noticeably smaller first dorsal fin relative to second; a first dorsal fin originating more than one-half its body length (53.8% TL) vs less than half its body length (45.9% TL); near equal labial furrow lengths vs much longer lower labial furrow; and lower precaudal (90 vs 102) and total (130 vs 140) vertebral counts.
</paragraph>
<paragraph id="8A0B36EA8862FF99FF04089CB73BF90C" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
<taxonomicName id="4DB44D698862FF99FF04089CB7D9FA70" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[199,486,1484,1508]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88862FF99FF04089CB7D9FA70" box="[199,486,1484,1508]" italics="true" pageId="8" pageNumber="539">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F357838862FF99FE2D089DB411FA70" box="[494,558,1484,1508]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B8C0EAF88862FF99FE2D089DB418FA70" bold="true" box="[494,551,1484,1508]" pageId="8" pageNumber="539">n. sp</emphasis>
.
</taxonomicNameLabel>
differs from 
<taxonomicName id="4DB44D698862FF99FD06089CB511FA70" authorityName="Garman" authorityYear="1906" box="[709,814,1484,1508]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="pilosus">
<emphasis id="B8C0EAF88862FF99FD06089CB511FA70" box="[709,814,1484,1508]" italics="true" pageId="8" pageNumber="539">P. pilosus</emphasis>
</taxonomicName>
by a longer pectoral-pelvic fin space (27.2% vs 21%); second dorsal fin origin above anal fin base vs second dorsal fin origin opposite anal fin insertion; roof of mouth blackish with darker pores vs light without distinctly darker pores; uniform medium brown, slightly lighter on abdomen and with darker fin edges vs uniform reddish above, becoming lighter distally below first dorsal and extending to caudal region, white below along abdomen and laterally along flanks, and without darker markings or fin edges.
</paragraph>
<paragraph id="8A0B36EA8862FF99FF040BF5B736F8E4" blockId="8.[151,1437,151,1904]" pageId="8" pageNumber="539">
Geographically, 
<taxonomicName id="4DB44D698862FF99FEBC0BF4B400F928" authorityName="Springer" authorityYear="1979" box="[383,575,1700,1724]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="campechiensis">
<emphasis id="B8C0EAF88862FF99FEBC0BF4B400F928" box="[383,575,1700,1724]" italics="true" pageId="8" pageNumber="539">P. campechiensis</emphasis>
</taxonomicName>
is known only from the western Gulf of 
<collectingRegion id="4870F8088862FF99FBCB0BF5B25EF929" box="[1032,1121,1700,1725]" country="Mexico" name="Mexico" pageId="8" pageNumber="539">
<collectingCountry id="F2A3767A8862FF99FBCB0BF5B25EF929" box="[1032,1121,1700,1725]" name="Mexico" pageId="8" pageNumber="539">Mexico</collectingCountry>
</collectingRegion>
, Bay of 
<collectingRegion id="4870F8088862FF99FB7D0BF5B308F929" box="[1214,1335,1700,1725]" country="Mexico" name="Campeche" pageId="8" pageNumber="539">Campeche</collectingRegion>
, while 
<taxonomicName id="4DB44D698862FF99FA440BF4B6C6F94B" baseAuthorityName="Gilbert" baseAuthorityYear="1892" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="xaniurus">
<emphasis id="B8C0EAF88862FF99FA440BF4B6C6F94B" italics="true" pageId="8" pageNumber="539">P. xaniurus</emphasis>
</taxonomicName>
is only known from the eastern North Pacific from 
<collectingRegion id="4870F8088862FF99FC870B96B587F974" box="[836,952,1735,1760]" country="United States of America" name="California" pageId="8" pageNumber="539">California</collectingRegion>
, 
<collectingCountry id="F2A3767A8862FF99FC070B99B5C3F974" box="[964,1020,1736,1760]" name="United States of America" pageId="8" pageNumber="539">USA</collectingCountry>
to 
<collectingRegion id="4870F8088862FF99FBE10B96B2EDF974" box="[1058,1234,1735,1760]" country="Mexico" name="Baja California" pageId="8" pageNumber="539">
Baja 
<collectingRegion id="4870F8088862FF99FB9E0B96B2EDF974" box="[1117,1234,1735,1760]" country="United States of America" name="California" pageId="8" pageNumber="539">California</collectingRegion>
</collectingRegion>
, 
<collectingRegion id="4870F8088862FF99FB1E0B96B30DF974" box="[1245,1330,1735,1760]" country="Mexico" name="Mexico" pageId="8" pageNumber="539">
<collectingCountry id="F2A3767A8862FF99FB1E0B96B30DF974" box="[1245,1330,1735,1760]" name="Mexico" pageId="8" pageNumber="539">Mexico</collectingCountry>
</collectingRegion>
(Ebert 
<emphasis id="B8C0EAF88862FF99FA4B0B98B692F890" italics="true" pageId="8" pageNumber="539">et al</emphasis>
., 2013). In addition to biogeography, 
<taxonomicName id="4DB44D698862FF99FD910BBCB54FF890" authority="Ebert, 2018" authorityName="Ebert" authorityYear="2018" box="[594,880,1772,1796]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="nigripalatum" status="sp. nov.">
<emphasis id="B8C0EAF88862FF99FD910BBCB54FF890" box="[594,880,1772,1796]" italics="true" pageId="8" pageNumber="539">Parmaturus nigripalatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3F357838862FF99FCB40BBDB589F890" box="[887,950,1772,1796]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B8C0EAF88862FF99FCB40BBDB590F890" bold="true" box="[887,943,1772,1796]" pageId="8" pageNumber="539">n. sp</emphasis>
.
</taxonomicNameLabel>
can be separated from these two species by a upper and lower total tooth counts (52/48 vs&gt;60/60); higher total vertebral counts (130 vs 109–121); and upper and lower labial furrows near equal in length vs rudimentary (
<taxonomicName id="4DB44D698862FF99FC930A64B230F8D8" authorityName="Springer" authorityYear="1979" box="[848,1039,1844,1868]" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="campechiensis">
<emphasis id="B8C0EAF88862FF99FC930A64B230F8D8" box="[848,1039,1844,1868]" italics="true" pageId="8" pageNumber="539">P. campechiensis</emphasis>
</taxonomicName>
) or shorter upper labial furrows (
<taxonomicName id="4DB44D698862FF99FA440A64B6C6F8FB" baseAuthorityName="Gilbert" baseAuthorityYear="1892" class="Chondrichthyes" family="Scyliorhinidae" genus="Parmaturus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="xaniurus">
<emphasis id="B8C0EAF88862FF99FA440A64B6C6F8FB" italics="true" pageId="8" pageNumber="539">P. xaniurus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>