172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document id="2D9F8826E03AD709E1340F446D662B83" ID-CLB-Dataset="5336" ID-DOI="10.11646/zootaxa.4999.4.2" ID-GBIF-Dataset="6e1e8064-37e6-486f-b4fa-de0170edfb5d" ID-ISSN="1175-5326" ID-Zenodo-Dep="5119186" ID-ZooBank="4DD55A91-8739-4AC5-9208-36B7DF7AF22F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626871680260" checkinUser="plazi" docAuthor="Archibald, S. Bruce, Aase, Arvid & Nel, André" docDate="2021" docId="021387DAFF85FFD4FF7E0F17473A64A0" docLanguage="en" docName="zootaxa.4999.4.2.pdf" docOrigin="Zootaxa 4999 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eoteredon lacoi Archibald & Aase & Nel 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="326" masterDocId="FE2AFFA2FF84FFD0FFE90A6A454D6561" masterDocTitle="The second North American fossil horntail wood-wasp (Hymenoptera: Siricidae) from the early Eocene Green River Formation" masterLastPageNumber="334" masterPageNumber="325" pageNumber="326" updateTime="1699036931053" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="4365570AF43B0D406E502A23FBDD3AE5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1C75E0CECEAE7CC1F63DE3C9EAB17C5C">
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<mods:title id="EBAE2ED5268924F134769E08651AE759">The second North American fossil horntail wood-wasp (Hymenoptera: Siricidae) from the early Eocene Green River Formation</mods:title>
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</mods:titleInfo>
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<mods:name id="F87E39DF9F1746ED26193575F2FCD147" type="personal">
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<mods:roleTerm id="877BC7042EBFADB282FB29144176A526">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1122F891C0A10ECE3386F8F5572B755">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="40030D54F1CF343D1974B9032D798FB6">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada; and Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America; and Royal British Columbia Museum, 65 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada.</mods:affiliation>
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</mods:name>
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<mods:name id="8864E10D30A4599571EC9FD87126D017" type="personal">
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<mods:roleTerm id="2E1093B95D785249CE5591948D4FEA3D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1930DCD500E59AB321C00974DFA6D52">Aase, Arvid</mods:namePart>
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<mods:affiliation id="22D7C55226D8C63CC8582E1BB0DBB9E8">Fossil Butte National Monument, 864 Chicken Creek Road, Kemmerer, WY 83101, Wyoming, USA.</mods:affiliation>
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<mods:name id="C56D9FC43BB82E43995B361C3444814C" type="personal">
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<mods:roleTerm id="CED84B1A5946EDB3985EB1AD67EC0EA6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BAE6D66E23D8B4B6F8ABD9F92FCE9429">Nel, André</mods:namePart>
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<mods:affiliation id="5F3D164B979EDB724DCFC2BCC2239318">Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier 75005 Paris, France.</mods:affiliation>
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<mods:typeOfResource id="4D6723A02722F7C9EDB7003ECFD9B7D8">text</mods:typeOfResource>
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<mods:title id="F74B8DD24F8A8DCFC03B2CB36B7BDAA8">Zootaxa</mods:title>
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<mods:part id="B6753F26022DE7AC6843AA5D5AED094C">
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<mods:date id="6B285400D884756D88A7374C3E3C9670">2021</mods:date>
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<mods:number id="24D7A1B961E126AB3AFF3AA8DAD303A0">2021-07-13</mods:number>
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<mods:number id="5C6A4857C88B72F723CFDAACD9C70A26">4999</mods:number>
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<mods:number id="164ADE87C1EE99A2E809A22F40CC7716">4</mods:number>
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<mods:start id="EB618B02B145DC7001A4C183EE11BBF5">325</mods:start>
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<mods:classification id="74FB3AF3A9BAA270E734CEEDA99B5C8D">journal article</mods:classification>
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<mods:identifier id="57F714743106AD1C8EE3E56B689EF378" type="CLB-Dataset">5336</mods:identifier>
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<mods:identifier id="B15974FD7A5C421D8374C9F889B8B43F" type="DOI">10.11646/zootaxa.4999.4.2</mods:identifier>
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<mods:identifier id="CC26BE51B98F2DB8BD880A146CEFEA44" type="GBIF-Dataset">6e1e8064-37e6-486f-b4fa-de0170edfb5d</mods:identifier>
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<mods:identifier id="F5FA39E1E0E7CBBCE1BDE2B39783FBE2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BEE043E6082E15BB418F502834FE48B4" type="Zenodo-Dep">5119186</mods:identifier>
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<treatment id="021387DAFF85FFD4FF7E0F17473A64A0" ID-DOI="http://doi.org/10.5281/zenodo.5119192" ID-GBIF-Taxon="183718548" ID-Zenodo-Dep="5119192" LSID="urn:lsid:plazi:treatment:021387DAFF85FFD4FF7E0F17473A64A0" httpUri="http://treatment.plazi.org/id/021387DAFF85FFD4FF7E0F17473A64A0" lastPageId="4" lastPageNumber="326" pageId="1" pageNumber="326">
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<subSubSection id="C2A06547FF85FFD1FF7E0F1744F160F9" box="[151,444,1405,1432]" pageId="1" pageNumber="326" type="nomenclature">
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<paragraph id="8A0536CCFF85FFD1FF7E0F1744F160F9" blockId="1.[151,444,1405,1468]" box="[151,444,1405,1432]" pageId="1" pageNumber="326">
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<heading id="D14D81A0FF85FFD1FF7E0F1744F160F9" bold="true" box="[151,444,1405,1432]" fontSize="11" level="1" pageId="1" pageNumber="326" reason="1">
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||
<emphasis id="B8CEEADEFF85FFD1FF7E0F1744F160F9" bold="true" box="[151,444,1405,1432]" pageId="1" pageNumber="326">
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||
<taxonomicName id="4DBA4D4FFF85FFD1FF7E0F17441A60F9" authority="Archibald & Aase & Nel, 2021" authorityName="Archibald & Aase & Nel" authorityYear="2021" box="[151,343,1405,1432]" class="Insecta" family="Siricidae" genus="Eoteredon" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="326" phylum="Arthropoda" rank="species" species="lacoi" status="sp. nov.">
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<emphasis id="B8CEEADEFF85FFD1FF7E0F17441A60F9" bold="true" box="[151,343,1405,1432]" italics="true" pageId="1" pageNumber="326">Eoteredon lacoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A3FD57A5FF85FFD1FEB70F1444F160F9" box="[350,444,1406,1432]" pageId="1" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C2A06547FF85FFD1FF7E0FC8444660DD" box="[151,267,1442,1468]" pageId="1" pageNumber="326" type="description">
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||
<paragraph id="8A0536CCFF85FFD1FF7E0FC8444660DD" blockId="1.[151,444,1405,1468]" box="[151,267,1442,1468]" pageId="1" pageNumber="326">
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||
(
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<figureCitation id="12812A49FF85FFD1FF760FC845AA60DD" box="[159,231,1442,1468]" captionStart="FIGURE 3" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[168,1420,474,1733]" captionTargetId="figure-190@4.[168,1420,474,1733]" captionTargetPageId="4" captionText="FIGURE 3. Eoteredon lacoi gen. et sp. nov., holotype in ventral aspect, photographed dry. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5119196" httpUri="https://zenodo.org/record/5119196/files/figure.png" pageId="1" pageNumber="326">Figs 3</figureCitation>
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,
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<figureCitation id="12812A49FF85FFD1FF1D0FC8444E60DD" box="[244,259,1442,1468]" captionStart="FIGURE 4" captionStartId="6.[151,250,1727,1751]" captionTargetBox="[158,1429,194,1693]" captionTargetId="figure-28@6.[151,1436,181,1703]" captionTargetPageId="6" captionText="FIGURE 4. Drawings of wings of A, B, Teredon cubensis and C–F, Eoteredon lacoi gen. et sp. nov., holotype. A, T. cubensis male forewing, somewhat crushed in the basal posterior region. B, T. cubensis female forewing. C, right forewing of E. lacoi. D, left forewing of E. lacoi. E, E. lacoi female left hind wing. F, E. lacoi female right hind wing. Veins labelled in Roman font, cell names in italics. A, B re-drawn from Schiff et al. (2012, figs. C1.36, C34.5, no scale bars on originals). Scale bar = 5 mm (C–F)." figureDoi="http://doi.org/10.5281/zenodo.5119198" httpUri="https://zenodo.org/record/5119198/files/figure.png" pageId="1" pageNumber="326">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C2A06547FF85FFD1FF7E0F8346266349" pageId="1" pageNumber="326" type="materials_examined">
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<paragraph id="8A0536CCFF85FFD1FF7E0F8346266349" blockId="1.[151,1437,1513,1937]" pageId="1" pageNumber="326">
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<emphasis id="B8CEEADEFF85FFD1FF7E0F83444B6362" bold="true" box="[151,262,1513,1539]" pageId="1" pageNumber="326">Material.</emphasis>
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<materialsCitation id="3AD23C91FF85FFD1FEE40F8046266349" ID-GBIF-Occurrence="3326171301" collectionCode="FOBU" county="Housed" location="Kemmerer" municipality="Fossil Butte National Monument" pageId="1" pageNumber="326" specimenCode="FOBU13582" specimenCount="1" stateProvince="Wyoming" typeStatus="holotype">
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||
<typeStatus id="5501886EFF85FFD1FEE40F8044386365" box="[269,373,1514,1540]" pageId="1" pageNumber="326" type="holotype">Holotype</typeStatus>
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||
<specimenCode id="DA1C9EB7FF85FFD1FE950F8047406364" box="[380,525,1514,1541]" collectionCode="FOBU" pageId="1" pageNumber="326">FOBU13582</specimenCode>
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(part and counterpart of a nearly complete female) (
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<figureCitation id="12812A49FF85FFD1FBBC0F8041D46365" box="[1109,1177,1514,1540]" captionStart="FIGURE 3" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[168,1420,474,1733]" captionTargetId="figure-190@4.[168,1420,474,1733]" captionTargetPageId="4" captionText="FIGURE 3. Eoteredon lacoi gen. et sp. nov., holotype in ventral aspect, photographed dry. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5119196" httpUri="https://zenodo.org/record/5119196/files/figure.png" pageId="1" pageNumber="326">Fig. 3</figureCitation>
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).
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<collectingCounty id="63644E40FF85FFD1FB470F8040496365" box="[1198,1284,1514,1540]" pageId="1" pageNumber="326">Housed</collectingCounty>
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||
in the collections of
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<collectingMunicipality id="6A61ACB6FF85FFD1FF1A0C64472B6349" box="[243,614,1550,1576]" pageId="1" pageNumber="326">Fossil Butte National Monument</collectingMunicipality>
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||
,
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<location id="8F656017FF85FFD1FD980C6447A66349" LSID="urn:lsid:plazi:treatment:021387DAFF85FFD4FF7E0F17473A64A0:8F656017FF85FFD1FD980C6447A66349" box="[625,747,1550,1576]" county="Housed" municipality="Fossil Butte National Monument" name="Kemmerer" pageId="1" pageNumber="326" stateProvince="Wyoming">Kemmerer</location>
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,
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<collectingRegion id="487EF82EFF85FFD1FD1C0C64462A6349" box="[757,871,1550,1576]" country="United States of America" name="Wyoming" pageId="1" pageNumber="326">Wyoming</collectingRegion>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C2A06547FF85FFD1FF2E0C5846426311" pageId="1" pageNumber="326" type="etymology">
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<paragraph id="8A0536CCFF85FFD1FF2E0C5846426311" blockId="1.[151,1437,1513,1937]" pageId="1" pageNumber="326">
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<emphasis id="B8CEEADEFF85FFD1FF2E0C584403632D" bold="true" box="[199,334,1586,1612]" pageId="1" pageNumber="326">Etymology.</emphasis>
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The specific epithet is formed from the surname of Greg Laco, who donated the
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<typeStatus id="5501886EFF85FFD1FB3D0C58407B632D" box="[1236,1334,1586,1612]" pageId="1" pageNumber="326" type="holotype">holotype</typeStatus>
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to Fossil Butte National Monument, recognizing his contribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="C2A06547FF85FFD1FF2E0C13474A63F5" box="[199,519,1657,1684]" pageId="1" pageNumber="326" type="diagnosis">
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<paragraph id="8A0536CCFF85FFD1FF2E0C13474A63F5" blockId="1.[151,1437,1513,1937]" box="[199,519,1657,1684]" pageId="1" pageNumber="326">
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<emphasis id="B8CEEADEFF85FFD1FF2E0C13447763F2" bold="true" box="[199,314,1657,1683]" pageId="1" pageNumber="326">Diagnosis</emphasis>
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. As for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C2A06547FF85FFD1FF2E0CF440106261" pageId="1" pageNumber="326" type="materials_examined">
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<paragraph id="8A0536CCFF85FFD1FF2E0CF440106261" blockId="1.[151,1437,1513,1937]" pageId="1" pageNumber="326">
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<materialsCitation id="3AD23C91FF85FFD1FF2E0CF440176261" ID-GBIF-Occurrence="3326171302" collectorName="Ma. The Smith Hollow" country="United States of America" location="Green River Formation in Fossil Basin" municipality="Kemmerer" pageId="1" pageNumber="326" specimenCount="3" stateProvince="Wyoming" typeStatus="holotype">
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<emphasis id="B8CEEADEFF85FFD1FF2E0CF444B063D6" bold="true" box="[199,509,1693,1720]" pageId="1" pageNumber="326">
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||
<typeStatus id="5501886EFF85FFD1FF2E0CF4444F63D9" box="[199,258,1694,1720]" pageId="1" pageNumber="326">Type</typeStatus>
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locality and horizon
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</emphasis>
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. The
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<typeStatus id="5501886EFF85FFD1FDA90CF4473D63D9" box="[576,624,1694,1720]" pageId="1" pageNumber="326">type</typeStatus>
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and only specimen was found in the 18-inch Layer of the Fossil Butte Member of the
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<location id="8F656017FF85FFD1FEA30CA8464463BD" LSID="urn:lsid:plazi:treatment:021387DAFF85FFD4FF7E0F17473A64A0:8F656017FF85FFD1FEA30CA8464463BD" box="[330,777,1730,1756]" country="United States of America" municipality="Kemmerer" name="Green River Formation in Fossil Basin" pageId="1" pageNumber="326" stateProvince="Wyoming">Green River Formation in Fossil Basin</location>
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, approximately six meters below the K-spar Tuff dated at 51.98 ± 0.35
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<collectorName id="274F531AFF85FFD1FEC50C8C47626261" box="[300,559,1766,1792]" pageId="1" pageNumber="326">Ma. The Smith Hollow</collectorName>
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<typeStatus id="5501886EFF85FFD1FDDF0C8C472B6261" box="[566,614,1766,1792]" pageId="1" pageNumber="326">type</typeStatus>
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||
locality is
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||
<quantity id="4D429B29FF85FFD1FD0B0C8C460C6261" box="[738,833,1766,1792]" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="1" pageNumber="326" unit="mi" value="11.0">11 miles</quantity>
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west of downtown
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<collectingMunicipality id="6A61ACB6FF85FFD1FBF40C8C41DA6261" box="[1053,1175,1766,1792]" pageId="1" pageNumber="326">Kemmerer</collectingMunicipality>
|
||
,
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||
<collectingRegion id="487EF82EFF85FFD1FB480C8C405F6261" box="[1185,1298,1766,1792]" country="United States of America" name="Wyoming" pageId="1" pageNumber="326">Wyoming</collectingRegion>
|
||
,
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<collectingCountry id="F2AD765CFF85FFD1FAF70C8C40176261" box="[1310,1370,1766,1792]" name="United States of America" pageId="1" pageNumber="326">USA</collectingCountry>
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C2A06547FF85FFD4FF2E0D63473A64A0" lastPageId="4" lastPageNumber="329" pageId="1" pageNumber="326" type="description">
|
||
<paragraph id="8A0536CCFF85FFD4FF2E0D63473A64A0" blockId="1.[151,1437,1513,1937]" lastBlockId="4.[151,1437,135,450]" lastPageId="4" lastPageNumber="329" pageId="1" pageNumber="326">
|
||
<emphasis id="B8CEEADEFF85FFD1FF2E0D63441C6242" bold="true" box="[199,337,1801,1827]" pageId="1" pageNumber="326">Description</emphasis>
|
||
. Female, preserved in ventral aspect, except head in frontal aspect. Head poorly preserved,
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||
<quantity id="4D429B29FF85FFD1FA900D60458E6229" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="326" unit="mm" value="4.5">4.5 mm</quantity>
|
||
long,
|
||
<quantity id="4D429B29FF85FFD1FEE50D44442E6229" box="[268,355,1838,1865]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="326" unit="mm" value="3.5">3.5 mm</quantity>
|
||
wide, with compound eyes, mandibles poorly visible; ocelli, antenna, antennal sockets not discernible; thorax
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||
<quantity id="4D429B29FF85FFD1FEA60D3844E8620D" box="[335,421,1874,1900]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="326" unit="mm" value="8.7">8.7 mm</quantity>
|
||
long,
|
||
<quantity id="4D429B29FF85FFD1FE050D38470F620D" box="[492,578,1874,1901]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="326" unit="mm" value="5.5">5.5 mm</quantity>
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||
wide; only parts of metathoracic legs preserved, poorly: femur
|
||
<quantity id="4D429B29FF85FFD1FAE40D384029620D" box="[1293,1380,1874,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="326" unit="mm" value="2.2">2.2 mm</quantity>
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||
long and 1.0 mm wide, 3.5 as broad as metatibia; metatibia ca.
|
||
<quantity id="4D429B29FF85FFD1FCD80D1C46C462F1" box="[817,905,1910,1936]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="326" unit="mm" value="3.8">3.8 mm</quantity>
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||
long as preserved,
|
||
<quantity id="4D429B29FF85FFD1FB8E0D1C41F362F1" box="[1127,1214,1910,1936]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="326" unit="mm" value="0.3">0.3 mm</quantity>
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||
wide; fragments of metabasitarsomere possibly present. Forewing
|
||
<quantity id="4D429B29FF80FFD4FD4B0AED464865C3" box="[674,773,135,162]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="329" unit="mm" value="17.5">17.5 mm</quantity>
|
||
long, 5.0 mm wide, with apex poorly preserved; 2r–m present, joins cell 2M; cell 1Rs2 clearly longer than wide; cell 1R1 rather broad, only 2.8 times as long as broad; cell 2R1 about 0.7 times as long as cell 3R1; 2r-rs joins stigma in its distal half; stigma gradually attenuated evenly distal to junction with 2r-rs; Cu1 absent; 1cu–a joining Cu midway between
|
||
<quantity id="4D429B29FF80FFD4FC440A9E4683646F" box="[941,974,244,270]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="329" unit="m" value="1.0">1m</quantity>
|
||
–cu, M; Sc faint, present in basal third of wing; 2A extending along posterior edge of wing about 0.5 times cell 1A length; 3A possibly distorted (see text). Hind wing ca.
|
||
<quantity id="4D429B29FF80FFD4FED00B5644D66437" box="[313,411,316,342]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="329" unit="mm" value="11.3">11.3 mm</quantity>
|
||
long; anal cell presumed open (see below); hamuli not discernible; 1r–m clearly shorter than M; 1r–m at basal third of cell 1M; M markedly curved; abdomen
|
||
<quantity id="4D429B29FF80FFD4FC9D0B354695641B" box="[884,984,351,378]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="4" pageNumber="329" unit="mm" value="16.3">16.3 mm</quantity>
|
||
long, 7.0 mm wide, tergites not discernible (preserved in ventral aspect); distinct but incomplete ovipositor, preserved portion
|
||
<quantity id="4D429B29FF80FFD4FB8E0BEE41F064FF" box="[1127,1213,388,414]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="329" unit="mm" value="8.8">8.8 mm</quantity>
|
||
long, extending
|
||
<quantity id="4D429B29FF80FFD4FA900BEE458E64A3" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="329" unit="mm" value="2.2">2.2 mm</quantity>
|
||
from abdomen; annuli not discernible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |