773 lines
105 KiB
XML
773 lines
105 KiB
XML
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<mods:title id="8D04490603B8165B4DACA70608FA2C7E">Taxonomy of freshwater gobies of the genus Rhinogobius (Oxudercidae, Gobiiformes) from central Vietnam, with descriptions of two new species</mods:title>
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<mods:namePart id="92C3ABACB1328E688EE555AE2544F915">Maeda, Ken</mods:namePart>
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<mods:affiliation id="166467CD3EEFB189F9754B5DB04AB4C8">Marine Eco-Evo-Devo Unit, Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 -</mods:affiliation>
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<mods:namePart id="447879954A163C01B949E4CB8B43CD8C">Kobayashi, Hirozumi</mods:namePart>
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<mods:affiliation id="11950B959DE18BD44E067CFD4A7E90BC">Tropical Biosphere Research Center, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903 - 0213, Japan. & Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:namePart id="2941F450F4C3247DA0A510A1D302374C">Iida, Midori</mods:namePart>
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<mods:affiliation id="32EB7D8815A0DC0DC4C2A2B4BD555496">Marine Biological Station, Sado Island Center for Ecological Sustainability, Niigata University, 87 Tassha, Sado, Niigata 952 - 2135,</mods:affiliation>
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<mods:namePart id="4C519FAA10372A4C75413B69B7A0D582">Tran, Hau Duc</mods:namePart>
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<mods:affiliation id="D668459B48930DA66B932AFCE9EEFFAF">Institute of Natural Science, Hanoi National University of Education, 136 Xuan Thuy, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:date id="B69A23D37940CF64951A35583ECA99F6">2024</mods:date>
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<taxonomicName id="399B48AAFFF87D4CA0C8FDBFFCCCFDAA" ID-CoL="4S934" authority="Chen, Kottelat & Miller, 1999" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[151,812,565,592]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="523" phylum="Chordata" rank="species" species="taenigena">
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<emphasis id="CCEFEF3BFFF87D4CA0C8FDBFFE50FDAA" bold="true" box="[151,432,565,592]" italics="true" pageId="16" pageNumber="523">Rhinogobius taenigena</emphasis>
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<bibRefCitation id="9A0A4ED8FFF87D4CA1E7FDBFFCCCFDAA" author="Chen, I. - S. & Kottelat, M. & Miller, P. J." box="[440,812,565,591]" pageId="16" pageNumber="523" pagination="19 - 32" refId="ref23100" refString="Chen, I. - S., Kottelat, M. & Miller, P. J. (1999) Freshwater gobies of the genus Rhinogobius from the Mekong basin in Thailand and Laos, with descriptions of three new species. Zoological Studies, 38, 19 - 32." type="journal article" year="1999">Chen, Kottelat & Miller, 1999</bibRefCitation>
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(
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<figureCitation id="66A02FACFFF87D4CA0C0FDD3FF0FFD96" box="[159,239,601,627]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="16" pageNumber="523">
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Figs.
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,
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<figureCitation id="66A02FACFFF87D4CA0A4FDD3FECFFD96" box="[251,303,601,627]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="16" pageNumber="523">6e, f</figureCitation>
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,
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<figureCitation id="66A02FACFFF87D4CA164FDD3FE9FFD96" box="[315,383,601,627]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="16" pageNumber="523">10a–c</figureCitation>
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,
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<figureCitation id="66A02FACFFF87D4CA1D4FDD3FE48FD96" box="[395,424,601,627]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="16" pageNumber="523">11</figureCitation>
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,
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<figureCitation id="66A02FACFFF87D4CA1EBFDD3FE33FD96" box="[436,467,601,627]" captionStart="FIGURE 12" captionStartId="20.[151,250,1061,1085]" captionTargetBox="[151,1435,181,1037]" captionTargetId="figure-18@20.[151,1435,181,1037]" captionTargetPageId="20" captionText="FIGURE 12. In situ underwater photographs of Rhinogobius taenigena in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietnam. a, b, males, 20 June 2019; c, d, females, 20 June 2019 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330593" httpUri="https://zenodo.org/record/13330593/files/figure.png" pageId="16" pageNumber="523">12</figureCitation>
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;
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<tableCitation id="B3190692FFF87D4CA180FDD3FDDCFD96" box="[479,572,601,627]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="16" pageNumber="523">Tables 1</tableCitation>
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,
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<tableCitation id="B3190692FFF87D4CA215FDD3FDBAFD96" box="[586,602,601,627]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="16" pageNumber="523">2</tableCitation>
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,
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<tableCitation id="B3190692FFF87D4CA239FDD3FD73FD96" box="[614,659,601,627]" captionStart-0="TABLE 4" captionStart-1="TABLE 5" captionStart-2="TABLE 6" captionStart-3="TABLE 7" captionStartId-0="7.[151,237,151,175]" captionStartId-1="7.[151,234,868,892]" captionStartId-2="8.[151,236,151,175]" captionStartId-3="8.[151,237,868,892]" captionText-0="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-1="TABLE 5. Longitudinal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-2="TABLE 6. Transverse scale counts of various species of Rhinogobius examined in the present study. *Includes the holotype." captionText-3="TABLE 7. Predorsal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="16" pageNumber="523">4–7</tableCitation>
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,
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<tableCitation id="B3190692FFF87D4CA2C1FDD3FD4DFD96" box="[670,685,601,627]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="16" pageNumber="523">9</tableCitation>
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)
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<emphasis id="CCEFEF3BFFF87D4CA0C8FD2BFE7DFD5E" box="[151,413,673,699]" italics="true" pageId="16" pageNumber="523">Rhinogobius taenigena</emphasis>
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locality: Xe Pon between rapids upstream and downstream of Ban Fuang, Xe Bang Hiang basin, Mekong basin,
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(Xe Bang Hiang basin,
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<bibRefCitation id="9A0A4ED8FFF87D4CA39EFD63FB5DFCE1" author="Serov, D. V. & Nezdoliy, V. K. & Pavlov, D. S." box="[961,1213,745,772]" pageId="16" pageNumber="523" refId="ref24523" refString="Serov, D. V., Nezdoliy, V. K. & Pavlov, D. S. (2006) The freshwater fishes of central Vietnam. KMK Scientific Press Ltd., Moscow, 364 pp." type="book" year="2006">
|
||
Serov
|
||
<emphasis id="CCEFEF3BFFF87D4CA454FD60FBA4FCE6" box="[1035,1092,745,771]" italics="true" pageId="16" pageNumber="523">et al.</emphasis>
|
||
2006: 295
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(Gianh River basin to Phong Nha River,
|
||
<collectingRegion id="3C5FFDCBFFF87D4CA1DEFC87FD95FCC2" box="[385,629,781,807]" country="Vietnam" name="Quang Binh" pageId="16" pageNumber="523">Quang Binh Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="868C73B9FFF87D4CA2DFFC87FD04FCC2" box="[640,740,781,807]" name="Vietnam" pageId="16" pageNumber="523">Vietnam</collectingCountry>
|
||
);
|
||
<treatmentCitation id="7F3A1538FFF87D4CA2A9FC84FACFFCC2" author="Nagao Natural Environment Foundation" box="[758,1327,781,808]" page="441" pageId="16" pageNumber="523" year="2021">
|
||
<bibRefCitation id="9A0A4ED8FFF87D4CA2A9FC84FACFFCC2" author="Nagao Natural Environment Foundation" box="[758,1327,781,808]" pageId="16" pageNumber="523" refId="ref24278" refString="Nagao Natural Environment Foundation (2021) Fishes of the Indochinese Mekong. Nagao Natural Environment Foundation, Tokyo, xii + 546 pp." type="book" year="2021">Nagao Natural Environment Foundation 2021: 441</bibRefCitation>
|
||
</treatmentCitation>
|
||
(Mekong basin in
|
||
<collectingRegion id="3C5FFDCBFFF87D4CA0A9FCBBFE12FCAE" box="[246,498,817,843]" country="Laos" name="Savannakhet" pageId="16" pageNumber="523">Savannakhet Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="868C73B9FFF87D4CA1A1FCBBFDD9FCAE" box="[510,569,817,843]" name="Laos" pageId="16" pageNumber="523">Laos</collectingCountry>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B68160A2FFF87D4CA098FCDFFE14FAA6" pageId="16" pageNumber="523" type="materials_examined">
|
||
<paragraph id="FE243329FFF87D4CA098FCDFFC69FC76" blockId="16.[151,1437,673,2032]" pageId="16" pageNumber="523">
|
||
<emphasis id="CCEFEF3BFFF87D4CA098FCDFFE53FC8A" bold="true" box="[199,435,853,879]" pageId="16" pageNumber="523">Material examined.</emphasis>
|
||
<materialsCitation id="4EF33974FFF87D4CA1E3FCDFFC03FC8A" box="[444,995,853,879]" location="Mekong" pageId="16" pageNumber="523" specimenCount="13">
|
||
<specimenCount id="E89DF8A0FFF87D4CA1E3FCDFFD7AFC8A" box="[444,666,853,879]" count="13" pageId="16" pageNumber="523" type="generic">Thirteen specimens</specimenCount>
|
||
from the
|
||
<location id="FB4465F2FFF87D4CA350FCDFFC8CFC8A" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA350FCDFFC8CFC8A" box="[783,876,853,879]" name="Mekong" pageId="16" pageNumber="523">Mekong</location>
|
||
basin and
|
||
</materialsCitation>
|
||
<materialsCitation id="4EF33974FFF87D4CA3B2FCDFFC65FC76" country="Vietnam" county="Thua Thien Hue Province" location="Song Huong" municipality="Perfume River" pageId="16" pageNumber="523" specimenCount="5">
|
||
<specimenCount id="E89DF8A0FFF87D4CA3B2FCDFFB77FC8A" box="[1005,1175,853,879]" count="5" pageId="16" pageNumber="523" type="generic">five specimens</specimenCount>
|
||
from the
|
||
<location id="FB4465F2FFF87D4CA552FCDFFA7CFC95" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA552FCDFFA7CFC95" box="[1293,1436,853,880]" country="Vietnam" county="Thua Thien Hue Province" municipality="Perfume River" name="Song Huong" pageId="16" pageNumber="523">Song Huong</location>
|
||
(
|
||
<collectingMunicipality id="1E40A953FFF87D4CA0C1FCF3FEA6FC76" box="[158,326,889,915]" pageId="16" pageNumber="523">Perfume River</collectingMunicipality>
|
||
) basin, both in
|
||
<collectingCounty id="17454BA5FFF87D4CA1ACFCF3FCF7FC76" box="[499,791,889,916]" pageId="16" pageNumber="523">Thua Thien Hue Province</collectingCounty>
|
||
,
|
||
<collectingCountry id="868C73B9FFF87D4CA37DFCF3FC65FC76" box="[802,901,889,915]" name="Vietnam" pageId="16" pageNumber="523">Vietnam</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFF87D4CA098FC17FBA0FB56" blockId="16.[151,1437,673,2032]" pageId="16" pageNumber="523">
|
||
<emphasis id="CCEFEF3BFFF87D4CA098FC17FE91FC52" bold="true" box="[199,369,925,951]" pageId="16" pageNumber="523">Mekong basin</emphasis>
|
||
.
|
||
<materialsCitation id="4EF33974FFF87D4CA120FC14FBF1FC3E" collectingDate="2019-06-20" collectionCode="HNUE-F" latitude="16.356556" location="Xe Kong in Hong Trung" longLatPrecision="1" longitude="107.15439" municipality="Luoi District" pageId="16" pageNumber="523" specimenCode="HNUE-F 355-356" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCode id="AE3D9B52FFF87D4CA120FC14FDB1FC5D" box="[383,593,925,952]" collectionCode="HNUE-F" pageId="16" pageNumber="523">HNUE-F 355–356</specimenCode>
|
||
,
|
||
<specimenCount id="E89DF8A0FFF87D4CA202FC17FD48FC52" box="[605,680,925,951]" count="1" pageId="16" pageNumber="523" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA2E7FC17FCFEFC5D" box="[696,798,925,952]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="16" pageNumber="523" unit="mm" value="19.3">19.3 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFF87D4CA3D9FC17FC07FC52" box="[902,999,925,951]" count="1" pageId="16" pageNumber="523" type="female">1 female</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA3A8FC17FBBDFC5D" box="[1015,1117,925,952]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="16" pageNumber="523" unit="mm" value="29.5">29.5 mm</quantity>
|
||
SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA569FC14FEA2FC39" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA569FC14FEA2FC39" latitude="16.356556" longLatPrecision="1" longitude="107.15439" municipality="Luoi District" name="Xe Kong in Hong Trung" pageId="16" pageNumber="523">Xe Kong in Hong Trung</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA136FC4BFE1AFC3E" box="[361,506,961,987]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA254FC4BFD4BFC3E" box="[523,683,961,987]" degrees="16" direction="north" minutes="21" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="23.6" value="16.356556">16°21’23.6”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA2ECFC4BFCBFFC39" box="[691,863,961,988]" degrees="107" direction="east" minutes="09" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="15.8" value="107.15439">107°09’15.8”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA32AFC4BFBF1FC3E" box="[885,1041,961,988]" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA32AFC4BFBF1FC3E" box="[885,1041,961,988]" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFF87D4CA441FC4BFE11FBC6" collectingDate="2019-06-20" collectionCode="NSMT-P" latitude="16.356556" location="Xe Kong in Hong Trung" longLatPrecision="1" longitude="107.15439" municipality="Luoi District" pageId="16" pageNumber="523" specimenCode="NSMT-P 147891-147892" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCode id="AE3D9B52FFF87D4CA441FC4BFAA2FC3E" box="[1054,1346,961,987]" collectionCode="NSMT-P" pageId="16" pageNumber="523">NSMT-P 147891–147892</specimenCode>
|
||
,
|
||
<specimenCount id="E89DF8A0FFF87D4CA510FC4BFA7CFC3E" box="[1359,1436,961,987]" count="1" pageId="16" pageNumber="523" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA0C0FC6FFEE5FBE5" box="[159,261,997,1024]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="16" pageNumber="523" unit="mm" value="28.1">28.1 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFF87D4CA131FC6FFE30FC1A" box="[366,464,997,1023]" count="1" pageId="16" pageNumber="523" type="female">1 female</specimenCount>
|
||
(30.0 mm SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA37EFC6CFBD8FBE5" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA37EFC6CFBD8FBE5" box="[801,1080,997,1024]" latitude="16.356556" longLatPrecision="1" longitude="107.15439" municipality="Luoi District" name="Xe Kong in Hong Trung" pageId="16" pageNumber="523">Xe Kong in Hong Trung</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA402FC6FFB0DFC1A" box="[1117,1261,997,1023]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA4A2FC6FFA7DFC1A" box="[1277,1437,997,1023]" degrees="16" direction="north" minutes="21" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="23.6" value="16.356556">16°21’23.6”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA0C8FB83FEA3FBC1" box="[151,323,1033,1060]" degrees="107" direction="east" minutes="09" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="15.8" value="107.15439">107°09’15.8”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA107FB83FE11FBC6" box="[344,497,1033,1060]" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA107FB83FE11FBC6" box="[344,497,1033,1060]" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFF87D4CA1A2FB83FAC4FBA2" collectingDate="2019-06-20" collectionCode="NSMT-P" latitude="16.356556" location="Xe Kong in Hong Trung" longLatPrecision="1" longitude="107.15439" municipality="Luoi District" pageId="16" pageNumber="523" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
|
||
OISTICH 291, 292, 294, 299,
|
||
<specimenCount id="E89DF8A0FFF87D4CA30BFB83FC7FFBC6" box="[852,927,1033,1059]" count="1" pageId="16" pageNumber="523" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA3F0FB83FBF5FBC1" box="[943,1045,1033,1060]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="16" pageNumber="523" unit="mm" value="23.6">23.6 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFF87D4CA423FB83FB09FBC6" box="[1148,1257,1033,1059]" count="3" pageId="16" pageNumber="523" type="female">3 females</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA4A7FB83FA7DFBC1" box="[1272,1437,1033,1060]" metricMagnitude="-2" metricUnit="m" metricValue="3.04" metricValueMax="3.17" metricValueMin="2.91" pageId="16" pageNumber="523" unit="mm" value="30.4" valueMax="31.7" valueMin="29.1">29.1–31.7 mm</quantity>
|
||
SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA13DFBA4FD8FFBAD" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA13DFBA4FD8FFBAD" box="[354,623,1069,1096]" latitude="16.356556" longLatPrecision="1" longitude="107.15439" municipality="Luoi District" name="Xe Kong in Hong Trung" pageId="16" pageNumber="523">Xe Kong in Hong Trung</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA2CFFBA7FCFEFBA2" box="[656,798,1069,1095]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA374FBA7FC2BFBA2" box="[811,971,1069,1095]" degrees="16" direction="north" minutes="21" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="23.6" value="16.356556">16°21’23.6”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA38FFBA7FB9CFBAD" box="[976,1148,1069,1096]" degrees="107" direction="east" minutes="09" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="15.8" value="107.15439">107°09’15.8”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA4D0FBA7FAC4FBA2" box="[1167,1316,1069,1096]" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA4D0FBA7FAC4FBA2" box="[1167,1316,1069,1096]" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFF87D4CA571FBA7FEEEFB6A" collectingDate="2019-06-20" collectionCode="NSMT-P" latitude="16.29597" location="Xe Kong in Hong Kim" longLatPrecision="1" longitude="107.20939" municipality="Luoi District" pageId="16" pageNumber="523" specimenCount="1">
|
||
OISTICH 301, male (
|
||
<quantity id="39639ECCFFF87D4CA14AFBDBFE9AFB89" box="[277,378,1105,1132]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="16" pageNumber="523" unit="mm" value="29.7">29.7 mm</quantity>
|
||
SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA20CFBD8FCB7FB8E" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA20CFBD8FCB7FB8E" box="[595,855,1105,1132]" latitude="16.29597" longLatPrecision="1" longitude="107.20939" municipality="Luoi District" name="Xe Kong in Hong Kim" pageId="16" pageNumber="523">Xe Kong in Hong Kim</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA325FBDBFBEAFB8E" box="[890,1034,1105,1131]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA445FBDBFB59FB89" box="[1050,1209,1105,1132]" degrees="16" direction="north" minutes="17" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="45.5" value="16.29597">16°17’45.5”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA49FFBDBFA8CFB8E" box="[1216,1388,1105,1131]" degrees="107" direction="east" minutes="12" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="33.8" value="107.20939">107°12’33.8”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA5DFFBDBFEEEFB6A" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA5DFFBDBFEEEFB6A" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFF87D4CA144FBFFFBDCFB56" collectingDate="2019-06-20" collectionCode="NSMT-P" latitude="16.304722" location="Xe Kong in Hong Kim" longLatPrecision="1" longitude="107.212776" municipality="Luoi District" pageId="16" pageNumber="523" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
|
||
OISTICH 302–305,
|
||
<specimenCount id="E89DF8A0FFF87D4CA25CFBFFFDBAFB6A" box="[515,602,1141,1167]" count="2" pageId="16" pageNumber="523" type="male">2 males</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA235FBFFFCF0FB75" box="[618,784,1141,1168]" metricMagnitude="-2" metricUnit="m" metricValue="3.285" metricValueMax="3.4899999999999998" metricValueMin="3.08" pageId="16" pageNumber="523" unit="mm" value="32.85" valueMax="34.9" valueMin="30.8">30.8–34.9 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFF87D4CA324FBFFFC08FB6A" box="[891,1000,1141,1167]" count="2" pageId="16" pageNumber="523" type="female">2 females</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA3A7FBFFFB7EFB75" box="[1016,1182,1141,1168]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" metricValueMax="3.41" metricValueMin="3.05" pageId="16" pageNumber="523" unit="mm" value="32.3" valueMax="34.1" valueMin="30.5">30.5–34.1 mm</quantity>
|
||
SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA523FBFCFE93FB56" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA523FBFCFE93FB56" latitude="16.304722" longLatPrecision="1" longitude="107.212776" municipality="Luoi District" name="Xe Kong in Hong Kim" pageId="16" pageNumber="523">Xe Kong in Hong Kim</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA1C9FB13FDC9FB56" box="[406,553,1177,1203]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
, (
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA262FB13FD3CFB56" box="[573,732,1177,1203]" degrees="16" direction="north" minutes="18" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="17.0" value="16.304722">16°18’17.0”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA2BCFB13FC6FFB56" box="[739,911,1177,1203]" degrees="107" direction="east" minutes="12" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="46.0" value="107.212776">107°12’46.0”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA3FCFB13FBDCFB56" box="[931,1084,1177,1204]" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA3FCFB13FBDCFB56" box="[931,1084,1177,1204]" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFF87D4CA098FB37FE14FAA6" blockId="16.[151,1437,673,2032]" pageId="16" pageNumber="523">
|
||
<emphasis id="CCEFEF3BFFF87D4CA098FB37FE41FB32" bold="true" box="[199,417,1213,1239]" pageId="16" pageNumber="523">Song Huong basin</emphasis>
|
||
.
|
||
<materialsCitation id="4EF33974FFF87D4CA1F0FB34FBF7FB1E" collectingDate="2019-06-20" collectionCode="HNUE-F" collectorName="Hong Ha, A" latitude="16.3" location="Song Bo in Hong Ha" longLatPrecision="1" longitude="107.324745" municipality="Luoi District" pageId="16" pageNumber="523" specimenCode="HNUE-F 357-358" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCode id="AE3D9B52FFF87D4CA1F0FB34FD62FB3D" box="[431,642,1213,1240]" collectionCode="HNUE-F" pageId="16" pageNumber="523">HNUE-F 357–358</specimenCode>
|
||
,
|
||
<specimenCount id="E89DF8A0FFF87D4CA2D2FB37FD38FB32" box="[653,728,1213,1239]" count="1" pageId="16" pageNumber="523" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA2B7FB37FCADFB3D" box="[744,845,1213,1240]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="16" pageNumber="523" unit="mm" value="29.4">29.4 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFF87D4CA3EAFB37FBF6FB32" box="[949,1046,1213,1239]" count="1" pageId="16" pageNumber="523" type="female">1 female</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA47AFB37FB6AFB3D" box="[1061,1162,1213,1240]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="16" pageNumber="523" unit="mm" value="31.1">31.1 mm</quantity>
|
||
SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA53CFB37FEA8FB19" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA53CFB37FEA8FB19" latitude="16.3" longLatPrecision="1" longitude="107.324745" municipality="Luoi District" name="Song Bo in Hong Ha" pageId="16" pageNumber="523">
|
||
Song Bo in
|
||
<collectorName id="536E56FFFFF87D4CA080FB68FEA8FB19" box="[223,328,1250,1276]" pageId="16" pageNumber="523">Hong Ha</collectorName>
|
||
</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA130FB6BFDE1FB1E" box="[367,513,1249,1275]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA24DFB6BFD51FB1E" box="[530,689,1249,1275]" degrees="16" direction="north" minutes="18" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="00.0" value="16.3">16°18’00.0”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA2E6FB6BFC86FB1E" box="[697,870,1249,1275]" degrees="107" direction="east" minutes="19" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="29.1" value="107.324745">107°19’29.1”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA324FB6BFBF7FB1E" box="[891,1047,1249,1276]" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA324FB6BFBF7FB1E" box="[891,1047,1249,1276]" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFF87D4CA47BFB6BFE11FAA6" collectingDate="2019-06-20" collectionCode="HNUE-F" collectorName="Hong Ha, A" latitude="16.3" location="Song Bo in Hong Ha" longLatPrecision="1" longitude="107.324745" municipality="Luoi District" pageId="16" pageNumber="523" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
|
||
OISTICH 310–312,
|
||
<specimenCount id="E89DF8A0FFF87D4CA553FB6BFAB9FB1E" box="[1292,1369,1249,1275]" count="1" pageId="16" pageNumber="523" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA535FB6BFF23FAC5" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="16" pageNumber="523" unit="mm" value="28.8">28.8 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFF87D4CA16EFA8FFE7FFAFA" box="[305,415,1285,1311]" count="2" pageId="16" pageNumber="523" type="female">2 females</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFF87D4CA1EFFA8FFDB7FAC5" box="[432,599,1285,1312]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.34" metricValueMin="2.96" pageId="16" pageNumber="523" unit="mm" value="31.5" valueMax="33.4" valueMin="29.6">29.6–33.4 mm</quantity>
|
||
SL), a tributary of
|
||
<location id="FB4465F2FFF87D4CA365FA8FFBD1FAC5" LSID="urn:lsid:plazi:treatment:7632823FFFF87D49A0C8FDBFFEA9FDD8:FB4465F2FFF87D4CA365FA8FFBD1FAC5" box="[826,1073,1285,1312]" latitude="16.3" longLatPrecision="1" longitude="107.324745" municipality="Luoi District" name="Song Bo in Hong Ha" pageId="16" pageNumber="523">
|
||
Song Bo in
|
||
<collectorName id="536E56FFFFF87D4CA399FA8CFBD1FAC5" box="[966,1073,1286,1312]" pageId="16" pageNumber="523">Hong Ha</collectorName>
|
||
</location>
|
||
, A
|
||
<collectingMunicipality id="1E40A953FFF87D4CA406FA8FFB0BFAFA" box="[1113,1259,1285,1311]" pageId="16" pageNumber="523">Luoi District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA4A2FA8FFA7CFAFA" box="[1277,1436,1285,1311]" degrees="16" direction="north" minutes="18" orientation="latitude" pageId="16" pageNumber="523" precision="1" seconds="00.0" value="16.3">16°18’00.0”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFF87D4CA0C8FAA3FEA4FAA6" box="[151,324,1321,1347]" degrees="107" direction="east" minutes="19" orientation="longitude" pageId="16" pageNumber="523" precision="1" seconds="29.1" value="107.324745">107°19’29.1”E</geoCoordinate>
|
||
),
|
||
<date id="8A2515E9FFF87D4CA107FAA3FE11FAA6" box="[344,497,1321,1348]" pageId="16" pageNumber="523" value="2019-06-20">
|
||
<collectingDate id="9A61EC01FFF87D4CA107FAA3FE11FAA6" box="[344,497,1321,1348]" pageId="16" pageNumber="523" value="2019-06-20">20 June 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B68160A2FFF87D48A098FAC7FD67F90F" lastPageId="20" lastPageNumber="527" pageId="16" pageNumber="523" type="description">
|
||
<paragraph id="FE243329FFF87D4CA098FAC7FDC3F80A" blockId="16.[151,1437,673,2032]" pageId="16" pageNumber="523">
|
||
<emphasis id="CCEFEF3BFFF87D4CA098FAC7FEB9FA82" bold="true" box="[199,345,1357,1383]" pageId="16" pageNumber="523">Description.</emphasis>
|
||
Body nearly cylindrical anteriorly and somewhat compressed posteriorly. Anterior tip of head pointed in female, but rounded in male. Head larger in male than female (head length 30.9–33.5 vs. 28.1–30.5% of SL;
|
||
<figureCitation id="66A02FACFFF87D4CA0B7FA1FFECDFA4A" box="[232,301,1429,1455]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="16" pageNumber="523">Fig. 2</figureCitation>
|
||
,
|
||
<tableCitation id="B3190692FFF87D4CA164FA1FFE71FA4A" box="[315,401,1429,1455]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="16" pageNumber="523">Table 9</tableCitation>
|
||
). Eyes located dorsolaterally. Mouth terminal and oblique with thick upper and lower lips. When mouth closed, anterior tips of upper and lower jaws almost even, but lower jaw sometimes slightly protruding beyond upper jaw. Upper jaw extending posteriorly to about anterior margin of pupil. Mouth larger in male than female (upper-jaw length 10.2–14.6 vs. 8.5–9.6% of SL;
|
||
<figureCitation id="66A02FACFFF87D4CA37FF98BFC84F9FE" box="[800,868,1537,1563]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="16" pageNumber="523">Fig. 2</figureCitation>
|
||
,
|
||
<tableCitation id="B3190692FFF87D4CA32EF98BFC27F9FE" box="[881,967,1537,1563]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="16" pageNumber="523">Table 9</tableCitation>
|
||
). Upper and lower jaws usually with 2–4 rows of conical teeth; those in outermost row enlarged and those in inner rows irregularly aligned. Mental flap on chin usually rectangular or trapezoidal (posterior end forming long side), often with a small central projection. Anterior nostril short and tubular, posterior nostril a pore. Cephalic sensory pore system with B´, C, D(S), E, F, H´, K´, and L´ in oculoscapular canal and M´, N, and O´ in preopercular canal of both left and right sides in 11 of
|
||
<specimenCount id="E89DF8A0FFF87D4CA5DFF91BFEECF92A" count="18" pageId="16" pageNumber="523" type="generic">18 specimens</specimenCount>
|
||
examined (
|
||
<figureCitation id="66A02FACFFF87D4CA1D1F93FFDE9F92A" box="[398,521,1717,1743]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="16" pageNumber="523">Fig. 10a–c</figureCitation>
|
||
), but
|
||
<specimenCount id="E89DF8A0FFF87D4CA219F93CFD3FF92A" box="[582,735,1717,1744]" count="1" pageId="16" pageNumber="523" type="generic">one specimen</specimenCount>
|
||
having an additional pore behind pore F (not G, but near F) on left side,
|
||
<specimenCount id="E89DF8A0FFF87D4CA15FF950FE46F916" box="[256,422,1753,1780]" count="2" pageId="16" pageNumber="523" type="generic">two specimens</specimenCount>
|
||
having pore G on one side and pore H´ longitudinal groove-like on another side of one of
|
||
<specimenCount id="E89DF8A0FFF87D4CA0C8F974FEA1F8F2" box="[151,321,1789,1816]" count="2" pageId="16" pageNumber="523" type="generic">two specimens</specimenCount>
|
||
,
|
||
<specimenCount id="E89DF8A0FFF87D4CA113F974FE05F8F2" box="[332,485,1789,1816]" count="1" pageId="16" pageNumber="523" type="generic">one specimen</specimenCount>
|
||
having an additional pore between pores E and F in right side,
|
||
<specimenCount id="E89DF8A0FFF87D4CA4F0F974FAA9F8F2" box="[1199,1353,1789,1816]" count="1" pageId="16" pageNumber="523" type="generic">one specimen</specimenCount>
|
||
having a single heart-shaped pore C,
|
||
<specimenCount id="E89DF8A0FFF87D4CA1B7F8A8FD6EF8DE" box="[488,654,1825,1852]" count="2" pageId="16" pageNumber="523" type="generic">two specimens</specimenCount>
|
||
having two pores aligned longitudinally on one side of C (total three pores at C) and oculoscapular canal of one side without pore M´ and with vertical groove-like N´ instead in one of
|
||
<specimenCount id="E89DF8A0FFF87D4CA0C8F8E0FEA1F866" box="[151,321,1897,1924]" count="2" pageId="16" pageNumber="523" type="generic">two specimens</specimenCount>
|
||
, and oculoscapular canal of one side lacking N in another one. Arrangement of cutaneous sensory papillae of head shown in
|
||
<figureCitation id="66A02FACFFF87D4CA196F807FDA6F842" box="[457,582,1933,1959]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="16" pageNumber="523">Fig. 10a–c</figureCitation>
|
||
. Cheek having no significant transverse papilla rows. Vertebrae 10+16=26 (n=1), 10+17=27 (n=11), 10+18=28 (n=1), or 11+17=28 (n=1;
|
||
<tableCitation id="B3190692FFF87D4CA300F83BFC52F82E" box="[863,946,1969,1995]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="16" pageNumber="523">Table 1</tableCitation>
|
||
), P-V 3/II II I I 0/9 (n=12), P-V 3/II II I I I 0/10 (n=1), or 3/II III 0 I 0/9 (n=1).
|
||
</paragraph>
|
||
<caption id="AAE463A1FFF97D4DA0C8FF1DFDB0FF12" pageId="17" pageNumber="524" startId="17.[151,239,151,175]" targetType="table">
|
||
<paragraph id="FE243329FFF97D4DA0C8FF1DFDB0FF12" blockId="17.[151,1436,151,247]" pageId="17" pageNumber="524">
|
||
<emphasis id="CCEFEF3BFFF97D4DA0C8FF1DFEEDFF4A" bold="true" box="[151,269,151,175]" pageId="17" pageNumber="524">TABLE 9.</emphasis>
|
||
Morphometric data of
|
||
<taxonomicName id="399B48AAFFF97D4DA24FFF1DFCEAFF4A" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[528,778,151,175]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="524" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFF97D4DA24FFF1DFCEAFF4A" box="[528,778,151,175]" italics="true" pageId="17" pageNumber="524">Rhinogobius taenigena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="CCEFEF3BFFF97D4DA345FF12FC70FF4A" box="[794,912,151,175]" italics="true" pageId="17" pageNumber="524">
|
||
<taxonomicName id="399B48AAFFF97D4DA345FF12FC6CFF4A" box="[794,908,151,175]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="524" phylum="Chordata" rank="species" species="leavelli">R. leavelli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="399B48AAFFF97D4DA395FF12FBD0FF4A" box="[970,1072,151,175]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="524" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="CCEFEF3BFFF97D4DA395FF12FBD0FF4A" box="[970,1072,151,175]" italics="true" pageId="17" pageNumber="524">R. similis</emphasis>
|
||
</taxonomicName>
|
||
from central Vietnam, expressed as percentages of standard length. D
|
||
<subScript id="621F316CFFF97D4DA27DFF41FDCAFF3C" attach="right" box="[546,554,203,217]" fontSize="6" pageId="17" pageNumber="524">1</subScript>
|
||
, first dorsal fin; D
|
||
<subScript id="621F316CFFF97D4DA2AAFF41FD1DFF3C" attach="left" box="[757,765,203,217]" fontSize="6" pageId="17" pageNumber="524">2</subScript>
|
||
, second dorsal fin; A, anal fin; C, caudal fin; P
|
||
<subScript id="621F316CFFF97D4DA55DFF41FAEAFF3C" attach="left" box="[1282,1290,203,217]" fontSize="6" pageId="17" pageNumber="524">2</subScript>
|
||
, pelvic fin. * Includes five specimens of unknown sex.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FE243329FFF97D4DA1AEFE82FA95FB70" pageId="17" pageNumber="524">
|
||
<table id="8C9BC189FFF982A3A0FCFE8DFA96FB70" box="[163,1398,263,1173]" gridcols="7" gridrows="23" pageId="17" pageNumber="524">
|
||
<tr id="40AB316BFFF982A3A0FCFE8DFA96FEFA" box="[163,1398,263,287]" gridrow="0" pageId="17" pageNumber="524" rowspan-0="1" rowspan-4="1" rowspan-6="1">
|
||
<th id="037A5817FFF982A3A1AEFE8DFD0BFEFA" box="[497,747,263,287]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="17" pageNumber="524">
|
||
<taxonomicName id="399B48AAFFF97D4DA1AEFE82FD91FEFA" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[497,625,264,287]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="524" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFF97D4DA1AEFE82FD91FEFA" box="[497,625,264,287]" italics="true" pageId="17" pageNumber="524">R. taenigena</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="037A5817FFF982A3A369FE8DFC7DFEFA" box="[822,925,263,287]" gridcol="3" gridrow="0" pageId="17" pageNumber="524">
|
||
<taxonomicName id="399B48AAFFF97D4DA369FE82FC7DFEFA" box="[822,925,263,287]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="524" phylum="Chordata" rank="species" species="leavelli">
|
||
<emphasis id="CCEFEF3BFFF97D4DA369FE82FC7DFEFA" box="[822,925,263,287]" italics="true" pageId="17" pageNumber="524">R. leavelli</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="037A5817FFF982A3A423FE8DFB01FEFA" box="[1148,1249,263,287]" gridcol="5" gridrow="0" pageId="17" pageNumber="524">
|
||
<taxonomicName id="399B48AAFFF97D4DA423FE82FB3AFEFA" box="[1148,1242,263,287]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="524" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="CCEFEF3BFFF97D4DA423FE82FB3AFEFA" box="[1148,1242,263,287]" italics="true" pageId="17" pageNumber="524">R. similis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFEBAFA96FEAD" box="[163,1398,304,328]" gridrow="1" pageId="17" pageNumber="524" rowspan-0="1">
|
||
<td id="037A5817FFF982A3A1AEFEBAFDB6FEAD" box="[497,598,304,328]" gridcol="1" gridrow="1" pageId="17" pageNumber="524">Male</td>
|
||
<td id="037A5817FFF982A3A2DAFEBAFD0BFEAD" box="[645,747,304,328]" gridcol="2" gridrow="1" pageId="17" pageNumber="524">Female</td>
|
||
<td id="037A5817FFF982A3A369FEBAFC7DFEAD" box="[822,925,304,328]" gridcol="3" gridrow="1" pageId="17" pageNumber="524">Male</td>
|
||
<td id="037A5817FFF982A3A394FEBAFBD0FEAD" box="[971,1072,304,328]" gridcol="4" gridrow="1" pageId="17" pageNumber="524">Female</td>
|
||
<td id="037A5817FFF982A3A423FEBAFB01FEAD" box="[1148,1249,304,328]" gridcol="5" gridrow="1" pageId="17" pageNumber="524">Male</td>
|
||
<td id="037A5817FFF982A3A54FFEBAFA96FEAD" box="[1296,1398,304,328]" gridcol="6" gridrow="1" pageId="17" pageNumber="524">Female*</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFED2FA96FE95" box="[163,1398,344,368]" gridrow="2" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFED2FE39FE95" box="[163,473,344,368]" gridcol="0" gridrow="2" pageId="17" pageNumber="524">Number of specimens</th>
|
||
<td id="037A5817FFF982A3A1AEFED2FDB6FE95" box="[497,598,344,368]" gridcol="1" gridrow="2" pageId="17" pageNumber="524">8</td>
|
||
<td id="037A5817FFF982A3A2DAFED2FD0BFE95" box="[645,747,344,368]" gridcol="2" gridrow="2" pageId="17" pageNumber="524">10</td>
|
||
<td id="037A5817FFF982A3A369FED2FC7DFE95" box="[822,925,344,368]" gridcol="3" gridrow="2" pageId="17" pageNumber="524">12</td>
|
||
<td id="037A5817FFF982A3A394FED2FBD0FE95" box="[971,1072,344,368]" gridcol="4" gridrow="2" pageId="17" pageNumber="524">10</td>
|
||
<td id="037A5817FFF982A3A423FED2FB01FE95" box="[1148,1249,344,368]" gridcol="5" gridrow="2" pageId="17" pageNumber="524">9</td>
|
||
<td id="037A5817FFF982A3A54FFED2FA96FE95" box="[1296,1398,344,368]" gridcol="6" gridrow="2" pageId="17" pageNumber="524">15</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFE0AFA96FE7D" box="[163,1398,384,408]" gridrow="3" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFE0AFE39FE7D" box="[163,473,384,408]" gridcol="0" gridrow="3" pageId="17" pageNumber="524">Standard length (mm)</th>
|
||
<td id="037A5817FFF982A3A1AEFE0AFDB6FE7D" box="[497,598,384,408]" gridcol="1" gridrow="3" pageId="17" pageNumber="524">23.6–34.9</td>
|
||
<td id="037A5817FFF982A3A2DAFE0AFD0BFE7D" box="[645,747,384,408]" gridcol="2" gridrow="3" pageId="17" pageNumber="524">29.1–34.1</td>
|
||
<td id="037A5817FFF982A3A369FE0AFC7DFE7D" box="[822,925,384,408]" gridcol="3" gridrow="3" pageId="17" pageNumber="524">27.3–43.8</td>
|
||
<td id="037A5817FFF982A3A394FE0AFBD0FE7D" box="[971,1072,384,408]" gridcol="4" gridrow="3" pageId="17" pageNumber="524">27.0–41.1</td>
|
||
<td id="037A5817FFF982A3A423FE0AFB01FE7D" box="[1148,1249,384,408]" gridcol="5" gridrow="3" pageId="17" pageNumber="524">42.5–67.1</td>
|
||
<td id="037A5817FFF982A3A54FFE0AFA96FE7D" box="[1296,1398,384,408]" gridcol="6" gridrow="3" pageId="17" pageNumber="524">30.5–62.4</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFE22FA96FE25" box="[163,1398,424,448]" gridrow="4" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFE22FE39FE25" box="[163,473,424,448]" gridcol="0" gridrow="4" pageId="17" pageNumber="524">Head length</th>
|
||
<td id="037A5817FFF982A3A1AEFE22FDB6FE25" box="[497,598,424,448]" gridcol="1" gridrow="4" pageId="17" pageNumber="524">30.9–33.5</td>
|
||
<td id="037A5817FFF982A3A2DAFE22FD0BFE25" box="[645,747,424,448]" gridcol="2" gridrow="4" pageId="17" pageNumber="524">28.1–30.5</td>
|
||
<td id="037A5817FFF982A3A369FE22FC7DFE25" box="[822,925,424,448]" gridcol="3" gridrow="4" pageId="17" pageNumber="524">30.7–34.1</td>
|
||
<td id="037A5817FFF982A3A394FE22FBD0FE25" box="[971,1072,424,448]" gridcol="4" gridrow="4" pageId="17" pageNumber="524">28.7–31.1</td>
|
||
<td id="037A5817FFF982A3A423FE22FB01FE25" box="[1148,1249,424,448]" gridcol="5" gridrow="4" pageId="17" pageNumber="524">30.5–34.0</td>
|
||
<td id="037A5817FFF982A3A54FFE22FA96FE25" box="[1296,1398,424,448]" gridcol="6" gridrow="4" pageId="17" pageNumber="524">28.9–34.0</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFE5BFA96FE0C" box="[163,1398,465,489]" gridrow="5" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFE5BFE39FE0C" box="[163,473,465,489]" gridcol="0" gridrow="5" pageId="17" pageNumber="524">Snout length</th>
|
||
<td id="037A5817FFF982A3A1AEFE5BFDB6FE0C" box="[497,598,465,489]" gridcol="1" gridrow="5" pageId="17" pageNumber="524">7.8–10.3</td>
|
||
<td id="037A5817FFF982A3A2DAFE5BFD0BFE0C" box="[645,747,465,489]" gridcol="2" gridrow="5" pageId="17" pageNumber="524">7.1–8.2</td>
|
||
<td id="037A5817FFF982A3A369FE5BFC7DFE0C" box="[822,925,465,489]" gridcol="3" gridrow="5" pageId="17" pageNumber="524">9.7–12.1</td>
|
||
<td id="037A5817FFF982A3A394FE5BFBD0FE0C" box="[971,1072,465,489]" gridcol="4" gridrow="5" pageId="17" pageNumber="524">8.2–9.8</td>
|
||
<td id="037A5817FFF982A3A423FE5BFB01FE0C" box="[1148,1249,465,489]" gridcol="5" gridrow="5" pageId="17" pageNumber="524">11.1–13.9</td>
|
||
<td id="037A5817FFF982A3A54FFE5BFA96FE0C" box="[1296,1398,465,489]" gridcol="6" gridrow="5" pageId="17" pageNumber="524">9.0–12.3</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFE73FA96FDF4" box="[163,1398,505,529]" gridrow="6" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFE73FE39FDF4" box="[163,473,505,529]" gridcol="0" gridrow="6" pageId="17" pageNumber="524">Eye diameter</th>
|
||
<td id="037A5817FFF982A3A1AEFE73FDB6FDF4" box="[497,598,505,529]" gridcol="1" gridrow="6" pageId="17" pageNumber="524">6.0–7.1</td>
|
||
<td id="037A5817FFF982A3A2DAFE73FD0BFDF4" box="[645,747,505,529]" gridcol="2" gridrow="6" pageId="17" pageNumber="524">5.9–6.8</td>
|
||
<td id="037A5817FFF982A3A369FE73FC7DFDF4" box="[822,925,505,529]" gridcol="3" gridrow="6" pageId="17" pageNumber="524">6.6–7.7</td>
|
||
<td id="037A5817FFF982A3A394FE73FBD0FDF4" box="[971,1072,505,529]" gridcol="4" gridrow="6" pageId="17" pageNumber="524">6.6–8.1</td>
|
||
<td id="037A5817FFF982A3A423FE73FB01FDF4" box="[1148,1249,505,529]" gridcol="5" gridrow="6" pageId="17" pageNumber="524">6.2–7.3</td>
|
||
<td id="037A5817FFF982A3A54FFE73FA96FDF4" box="[1296,1398,505,529]" gridcol="6" gridrow="6" pageId="17" pageNumber="524">6.9–8.5</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFDABFA96FDDC" box="[163,1398,545,569]" gridrow="7" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFDABFE39FDDC" box="[163,473,545,569]" gridcol="0" gridrow="7" pageId="17" pageNumber="524">Postorbital length of head</th>
|
||
<td id="037A5817FFF982A3A1AEFDABFDB6FDDC" box="[497,598,545,569]" gridcol="1" gridrow="7" pageId="17" pageNumber="524">16.1–18.1</td>
|
||
<td id="037A5817FFF982A3A2DAFDABFD0BFDDC" box="[645,747,545,569]" gridcol="2" gridrow="7" pageId="17" pageNumber="524">14.7–16.4</td>
|
||
<td id="037A5817FFF982A3A369FDABFC7DFDDC" box="[822,925,545,569]" gridcol="3" gridrow="7" pageId="17" pageNumber="524">14.1–16.4</td>
|
||
<td id="037A5817FFF982A3A394FDABFBD0FDDC" box="[971,1072,545,569]" gridcol="4" gridrow="7" pageId="17" pageNumber="524">13.4–14.8</td>
|
||
<td id="037A5817FFF982A3A423FDABFB01FDDC" box="[1148,1249,545,569]" gridcol="5" gridrow="7" pageId="17" pageNumber="524">13.3–16.0</td>
|
||
<td id="037A5817FFF982A3A54FFDABFA96FDDC" box="[1296,1398,545,569]" gridcol="6" gridrow="7" pageId="17" pageNumber="524">12.8–15.3</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFDC3FA96FD84" box="[163,1398,585,609]" gridrow="8" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFDC3FE39FD84" box="[163,473,585,609]" gridcol="0" gridrow="8" pageId="17" pageNumber="524">Upper-jaw length</th>
|
||
<td id="037A5817FFF982A3A1AEFDC3FDB6FD84" box="[497,598,585,609]" gridcol="1" gridrow="8" pageId="17" pageNumber="524">10.2–14.6</td>
|
||
<td id="037A5817FFF982A3A2DAFDC3FD0BFD84" box="[645,747,585,609]" gridcol="2" gridrow="8" pageId="17" pageNumber="524">8.5–9.6</td>
|
||
<td id="037A5817FFF982A3A369FDC3FC7DFD84" box="[822,925,585,609]" gridcol="3" gridrow="8" pageId="17" pageNumber="524">11.0–15.8</td>
|
||
<td id="037A5817FFF982A3A394FDC3FBD0FD84" box="[971,1072,585,609]" gridcol="4" gridrow="8" pageId="17" pageNumber="524">8.8–11.2</td>
|
||
<td id="037A5817FFF982A3A423FDC3FB01FD84" box="[1148,1249,585,609]" gridcol="5" gridrow="8" pageId="17" pageNumber="524">12.6–17.6</td>
|
||
<td id="037A5817FFF982A3A54FFDC3FA96FD84" box="[1296,1398,585,609]" gridcol="6" gridrow="8" pageId="17" pageNumber="524">9.5–12.2</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFDFBFA96FD6A" box="[163,1398,625,655]" gridrow="9" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFDFBFE39FD6A" box="[163,473,625,655]" gridcol="0" gridrow="9" pageId="17" pageNumber="524">Body depth at P2 origin</th>
|
||
<td id="037A5817FFF982A3A1AEFDFBFDB6FD6A" box="[497,598,625,655]" gridcol="1" gridrow="9" pageId="17" pageNumber="524">14.2–16.5</td>
|
||
<td id="037A5817FFF982A3A2DAFDFBFD0BFD6A" box="[645,747,625,655]" gridcol="2" gridrow="9" pageId="17" pageNumber="524">14.2–17.2</td>
|
||
<td id="037A5817FFF982A3A369FDFBFC7DFD6A" box="[822,925,625,655]" gridcol="3" gridrow="9" pageId="17" pageNumber="524">14.8–17.3</td>
|
||
<td id="037A5817FFF982A3A394FDFBFBD0FD6A" box="[971,1072,625,655]" gridcol="4" gridrow="9" pageId="17" pageNumber="524">15.5–18.5</td>
|
||
<td id="037A5817FFF982A3A423FDFBFB01FD6A" box="[1148,1249,625,655]" gridcol="5" gridrow="9" pageId="17" pageNumber="524">16.2–19.3</td>
|
||
<td id="037A5817FFF982A3A54FFDFBFA96FD6A" box="[1296,1398,625,655]" gridcol="6" gridrow="9" pageId="17" pageNumber="524">16.4–19.6</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFD10FA96FD57" box="[163,1398,666,690]" gridrow="10" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFD10FE39FD57" box="[163,473,666,690]" gridcol="0" gridrow="10" pageId="17" pageNumber="524">Body depth at A origin</th>
|
||
<td id="037A5817FFF982A3A1AEFD10FDB6FD57" box="[497,598,666,690]" gridcol="1" gridrow="10" pageId="17" pageNumber="524">13.6–17.4</td>
|
||
<td id="037A5817FFF982A3A2DAFD10FD0BFD57" box="[645,747,666,690]" gridcol="2" gridrow="10" pageId="17" pageNumber="524">14.5–17.7</td>
|
||
<td id="037A5817FFF982A3A369FD10FC7DFD57" box="[822,925,666,690]" gridcol="3" gridrow="10" pageId="17" pageNumber="524">14.4–17.2</td>
|
||
<td id="037A5817FFF982A3A394FD10FBD0FD57" box="[971,1072,666,690]" gridcol="4" gridrow="10" pageId="17" pageNumber="524">15.1–17.8</td>
|
||
<td id="037A5817FFF982A3A423FD10FB01FD57" box="[1148,1249,666,690]" gridcol="5" gridrow="10" pageId="17" pageNumber="524">14.6–17.9</td>
|
||
<td id="037A5817FFF982A3A54FFD10FA96FD57" box="[1296,1398,666,690]" gridcol="6" gridrow="10" pageId="17" pageNumber="524">16.5–18.1</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFD48FA96FD3F" box="[163,1398,706,730]" gridrow="11" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFD48FE39FD3F" box="[163,473,706,730]" gridcol="0" gridrow="11" pageId="17" pageNumber="524">Depth at caudal peduncle</th>
|
||
<td id="037A5817FFF982A3A1AEFD48FDB6FD3F" box="[497,598,706,730]" gridcol="1" gridrow="11" pageId="17" pageNumber="524">10.9–12.5</td>
|
||
<td id="037A5817FFF982A3A2DAFD48FD0BFD3F" box="[645,747,706,730]" gridcol="2" gridrow="11" pageId="17" pageNumber="524">11.1–12.9</td>
|
||
<td id="037A5817FFF982A3A369FD48FC7DFD3F" box="[822,925,706,730]" gridcol="3" gridrow="11" pageId="17" pageNumber="524">10.4–12.3</td>
|
||
<td id="037A5817FFF982A3A394FD48FBD0FD3F" box="[971,1072,706,730]" gridcol="4" gridrow="11" pageId="17" pageNumber="524">10.1–12.0</td>
|
||
<td id="037A5817FFF982A3A423FD48FB01FD3F" box="[1148,1249,706,730]" gridcol="5" gridrow="11" pageId="17" pageNumber="524">10.1–12.1</td>
|
||
<td id="037A5817FFF982A3A54FFD48FA96FD3F" box="[1296,1398,706,730]" gridcol="6" gridrow="11" pageId="17" pageNumber="524">9.5–11.5</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFD60FA96FCE7" box="[163,1398,746,770]" gridrow="12" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFD60FE39FCE7" box="[163,473,746,770]" gridcol="0" gridrow="12" pageId="17" pageNumber="524">Length of caudal peduncle</th>
|
||
<td id="037A5817FFF982A3A1AEFD60FDB6FCE7" box="[497,598,746,770]" gridcol="1" gridrow="12" pageId="17" pageNumber="524">25.3–28.4</td>
|
||
<td id="037A5817FFF982A3A2DAFD60FD0BFCE7" box="[645,747,746,770]" gridcol="2" gridrow="12" pageId="17" pageNumber="524">25.7–29.0</td>
|
||
<td id="037A5817FFF982A3A369FD60FC7DFCE7" box="[822,925,746,770]" gridcol="3" gridrow="12" pageId="17" pageNumber="524">25.7–29.1</td>
|
||
<td id="037A5817FFF982A3A394FD60FBD0FCE7" box="[971,1072,746,770]" gridcol="4" gridrow="12" pageId="17" pageNumber="524">25.6–28.8</td>
|
||
<td id="037A5817FFF982A3A423FD60FB01FCE7" box="[1148,1249,746,770]" gridcol="5" gridrow="12" pageId="17" pageNumber="524">26.5–28.6</td>
|
||
<td id="037A5817FFF982A3A54FFD60FA96FCE7" box="[1296,1398,746,770]" gridcol="6" gridrow="12" pageId="17" pageNumber="524">26.1–30.2</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFC98FA96FCCE" box="[163,1398,786,811]" gridrow="13" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFC98FE39FCCE" box="[163,473,786,811]" gridcol="0" gridrow="13" pageId="17" pageNumber="524">Predorsal length</th>
|
||
<td id="037A5817FFF982A3A1AEFC98FDB6FCCE" box="[497,598,786,811]" gridcol="1" gridrow="13" pageId="17" pageNumber="524">38.1–40.9</td>
|
||
<td id="037A5817FFF982A3A2DAFC98FD0BFCCE" box="[645,747,786,811]" gridcol="2" gridrow="13" pageId="17" pageNumber="524">35.0–38.0</td>
|
||
<td id="037A5817FFF982A3A369FC98FC7DFCCE" box="[822,925,786,811]" gridcol="3" gridrow="13" pageId="17" pageNumber="524">37.4–39.8</td>
|
||
<td id="037A5817FFF982A3A394FC98FBD0FCCE" box="[971,1072,786,811]" gridcol="4" gridrow="13" pageId="17" pageNumber="524">36.7–39.4</td>
|
||
<td id="037A5817FFF982A3A423FC98FB01FCCE" box="[1148,1249,786,811]" gridcol="5" gridrow="13" pageId="17" pageNumber="524">36.8–40.4</td>
|
||
<td id="037A5817FFF982A3A54FFC98FA96FCCE" box="[1296,1398,786,811]" gridcol="6" gridrow="13" pageId="17" pageNumber="524">36.1–40.2</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFCB1FA96FCB6" box="[163,1398,827,851]" gridrow="14" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFCB1FE39FCB6" box="[163,473,827,851]" gridcol="0" gridrow="14" pageId="17" pageNumber="524">Preanal length</th>
|
||
<td id="037A5817FFF982A3A1AEFCB1FDB6FCB6" box="[497,598,827,851]" gridcol="1" gridrow="14" pageId="17" pageNumber="524">55.8–59.0</td>
|
||
<td id="037A5817FFF982A3A2DAFCB1FD0BFCB6" box="[645,747,827,851]" gridcol="2" gridrow="14" pageId="17" pageNumber="524">56.0–59.7</td>
|
||
<td id="037A5817FFF982A3A369FCB1FC7DFCB6" box="[822,925,827,851]" gridcol="3" gridrow="14" pageId="17" pageNumber="524">54.6–57.5</td>
|
||
<td id="037A5817FFF982A3A394FCB1FBD0FCB6" box="[971,1072,827,851]" gridcol="4" gridrow="14" pageId="17" pageNumber="524">55.0–60.8</td>
|
||
<td id="037A5817FFF982A3A423FCB1FB01FCB6" box="[1148,1249,827,851]" gridcol="5" gridrow="14" pageId="17" pageNumber="524">53.7–56.3</td>
|
||
<td id="037A5817FFF982A3A54FFCB1FA96FCB6" box="[1296,1398,827,851]" gridcol="6" gridrow="14" pageId="17" pageNumber="524">49.2–58.0</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFCE9FA96FC64" box="[163,1398,867,897]" gridrow="15" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFCE9FE39FC64" box="[163,473,867,897]" gridcol="0" gridrow="15" pageId="17" pageNumber="524">Length of D 1 base</th>
|
||
<td id="037A5817FFF982A3A1AEFCE9FDB6FC64" box="[497,598,867,897]" gridcol="1" gridrow="15" pageId="17" pageNumber="524">15.7–18.9</td>
|
||
<td id="037A5817FFF982A3A2DAFCE9FD0BFC64" box="[645,747,867,897]" gridcol="2" gridrow="15" pageId="17" pageNumber="524">16.4–19.5</td>
|
||
<td id="037A5817FFF982A3A369FCE9FC7DFC64" box="[822,925,867,897]" gridcol="3" gridrow="15" pageId="17" pageNumber="524">14.4–19.4</td>
|
||
<td id="037A5817FFF982A3A394FCE9FBD0FC64" box="[971,1072,867,897]" gridcol="4" gridrow="15" pageId="17" pageNumber="524">16.7–19.3</td>
|
||
<td id="037A5817FFF982A3A423FCE9FB01FC64" box="[1148,1249,867,897]" gridcol="5" gridrow="15" pageId="17" pageNumber="524">17.5–19.4</td>
|
||
<td id="037A5817FFF982A3A54FFCE9FA96FC64" box="[1296,1398,867,897]" gridcol="6" gridrow="15" pageId="17" pageNumber="524">15.7–19.8</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFC01FA96FC4C" box="[163,1398,907,937]" gridrow="16" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFC01FE39FC4C" box="[163,473,907,937]" gridcol="0" gridrow="16" pageId="17" pageNumber="524">D1 length</th>
|
||
<td id="037A5817FFF982A3A1AEFC01FDB6FC4C" box="[497,598,907,937]" gridcol="1" gridrow="16" pageId="17" pageNumber="524">20.1–24.1</td>
|
||
<td id="037A5817FFF982A3A2DAFC01FD0BFC4C" box="[645,747,907,937]" gridcol="2" gridrow="16" pageId="17" pageNumber="524">18.0–21.1</td>
|
||
<td id="037A5817FFF982A3A369FC01FC7DFC4C" box="[822,925,907,937]" gridcol="3" gridrow="16" pageId="17" pageNumber="524">18.3–29.9</td>
|
||
<td id="037A5817FFF982A3A394FC01FBD0FC4C" box="[971,1072,907,937]" gridcol="4" gridrow="16" pageId="17" pageNumber="524">17.6–21.2</td>
|
||
<td id="037A5817FFF982A3A423FC01FB01FC4C" box="[1148,1249,907,937]" gridcol="5" gridrow="16" pageId="17" pageNumber="524">20.1–23.8</td>
|
||
<td id="037A5817FFF982A3A54FFC01FA96FC4C" box="[1296,1398,907,937]" gridcol="6" gridrow="16" pageId="17" pageNumber="524">17.7–22.3</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFC39FA96FC34" box="[163,1398,947,977]" gridrow="17" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFC39FE39FC34" box="[163,473,947,977]" gridcol="0" gridrow="17" pageId="17" pageNumber="524">Interval between D1 and D2 bases</th>
|
||
<td id="037A5817FFF982A3A1AEFC39FDB6FC34" box="[497,598,947,977]" gridcol="1" gridrow="17" pageId="17" pageNumber="524">1.7–4.4</td>
|
||
<td id="037A5817FFF982A3A2DAFC39FD0BFC34" box="[645,747,947,977]" gridcol="2" gridrow="17" pageId="17" pageNumber="524">2.3–4.8</td>
|
||
<td id="037A5817FFF982A3A369FC39FC7DFC34" box="[822,925,947,977]" gridcol="3" gridrow="17" pageId="17" pageNumber="524">1.1–7.2</td>
|
||
<td id="037A5817FFF982A3A394FC39FBD0FC34" box="[971,1072,947,977]" gridcol="4" gridrow="17" pageId="17" pageNumber="524">3.2–5.4</td>
|
||
<td id="037A5817FFF982A3A423FC39FB01FC34" box="[1148,1249,947,977]" gridcol="5" gridrow="17" pageId="17" pageNumber="524">1.0–3.0</td>
|
||
<td id="037A5817FFF982A3A54FFC39FA96FC34" box="[1296,1398,947,977]" gridcol="6" gridrow="17" pageId="17" pageNumber="524">1.6–4.3</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFC56FA96FC1C" box="[163,1398,988,1017]" gridrow="18" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFC56FE39FC1C" box="[163,473,988,1017]" gridcol="0" gridrow="18" pageId="17" pageNumber="524">Length of D2 base</th>
|
||
<td id="037A5817FFF982A3A1AEFC56FDB6FC1C" box="[497,598,988,1017]" gridcol="1" gridrow="18" pageId="17" pageNumber="524">15.7–19.1</td>
|
||
<td id="037A5817FFF982A3A2DAFC56FD0BFC1C" box="[645,747,988,1017]" gridcol="2" gridrow="18" pageId="17" pageNumber="524">14.3–18.2</td>
|
||
<td id="037A5817FFF982A3A369FC56FC7DFC1C" box="[822,925,988,1017]" gridcol="3" gridrow="18" pageId="17" pageNumber="524">15.6–17.1</td>
|
||
<td id="037A5817FFF982A3A394FC56FBD0FC1C" box="[971,1072,988,1017]" gridcol="4" gridrow="18" pageId="17" pageNumber="524">14.9–16.6</td>
|
||
<td id="037A5817FFF982A3A423FC56FB01FC1C" box="[1148,1249,988,1017]" gridcol="5" gridrow="18" pageId="17" pageNumber="524">17.2–18.7</td>
|
||
<td id="037A5817FFF982A3A54FFC56FA96FC1C" box="[1296,1398,988,1017]" gridcol="6" gridrow="18" pageId="17" pageNumber="524">16.0–18.8</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFB8EFA96FBC7" box="[163,1398,1028,1058]" gridrow="19" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFB8EFE39FBC7" box="[163,473,1028,1058]" gridcol="0" gridrow="19" pageId="17" pageNumber="524">D 2 length</th>
|
||
<td id="037A5817FFF982A3A1AEFB8EFDB6FBC7" box="[497,598,1028,1058]" gridcol="1" gridrow="19" pageId="17" pageNumber="524">28.8–36.1</td>
|
||
<td id="037A5817FFF982A3A2DAFB8EFD0BFBC7" box="[645,747,1028,1058]" gridcol="2" gridrow="19" pageId="17" pageNumber="524">25.7–30.8</td>
|
||
<td id="037A5817FFF982A3A369FB8EFC7DFBC7" box="[822,925,1028,1058]" gridcol="3" gridrow="19" pageId="17" pageNumber="524">26.4–36.5</td>
|
||
<td id="037A5817FFF982A3A394FB8EFBD0FBC7" box="[971,1072,1028,1058]" gridcol="4" gridrow="19" pageId="17" pageNumber="524">24.1–28.2</td>
|
||
<td id="037A5817FFF982A3A423FB8EFB01FBC7" box="[1148,1249,1028,1058]" gridcol="5" gridrow="19" pageId="17" pageNumber="524">31.3–43.7</td>
|
||
<td id="037A5817FFF982A3A54FFB8EFA96FBC7" box="[1296,1398,1028,1058]" gridcol="6" gridrow="19" pageId="17" pageNumber="524">23.6–30.0</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFBA6FA96FBA1" box="[163,1398,1068,1092]" gridrow="20" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFBA6FE39FBA1" box="[163,473,1068,1092]" gridcol="0" gridrow="20" pageId="17" pageNumber="524">Length of A base</th>
|
||
<td id="037A5817FFF982A3A1AEFBA6FDB6FBA1" box="[497,598,1068,1092]" gridcol="1" gridrow="20" pageId="17" pageNumber="524">13.3–15.7</td>
|
||
<td id="037A5817FFF982A3A2DAFBA6FD0BFBA1" box="[645,747,1068,1092]" gridcol="2" gridrow="20" pageId="17" pageNumber="524">11.0–15.5</td>
|
||
<td id="037A5817FFF982A3A369FBA6FC7DFBA1" box="[822,925,1068,1092]" gridcol="3" gridrow="20" pageId="17" pageNumber="524">13.2–15.8</td>
|
||
<td id="037A5817FFF982A3A394FBA6FBD0FBA1" box="[971,1072,1068,1092]" gridcol="4" gridrow="20" pageId="17" pageNumber="524">12.0–14.2</td>
|
||
<td id="037A5817FFF982A3A423FBA6FB01FBA1" box="[1148,1249,1068,1092]" gridcol="5" gridrow="20" pageId="17" pageNumber="524">14.7–16.5</td>
|
||
<td id="037A5817FFF982A3A54FFBA6FA96FBA1" box="[1296,1398,1068,1092]" gridcol="6" gridrow="20" pageId="17" pageNumber="524">13.7–15.9</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFBDEFA96FB88" box="[163,1398,1108,1133]" gridrow="21" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFBDEFE39FB88" box="[163,473,1108,1133]" gridcol="0" gridrow="21" pageId="17" pageNumber="524">A length</th>
|
||
<td id="037A5817FFF982A3A1AEFBDEFDB6FB88" box="[497,598,1108,1133]" gridcol="1" gridrow="21" pageId="17" pageNumber="524">25.0–33.0</td>
|
||
<td id="037A5817FFF982A3A2DAFBDEFD0BFB88" box="[645,747,1108,1133]" gridcol="2" gridrow="21" pageId="17" pageNumber="524">22.0–28.4</td>
|
||
<td id="037A5817FFF982A3A369FBDEFC7DFB88" box="[822,925,1108,1133]" gridcol="3" gridrow="21" pageId="17" pageNumber="524">24.0–30.6</td>
|
||
<td id="037A5817FFF982A3A394FBDEFBD0FB88" box="[971,1072,1108,1133]" gridcol="4" gridrow="21" pageId="17" pageNumber="524">22.2–25.1</td>
|
||
<td id="037A5817FFF982A3A423FBDEFB01FB88" box="[1148,1249,1108,1133]" gridcol="5" gridrow="21" pageId="17" pageNumber="524">26.4–38.5</td>
|
||
<td id="037A5817FFF982A3A54FFBDEFA96FB88" box="[1296,1398,1108,1133]" gridcol="6" gridrow="21" pageId="17" pageNumber="524">22.6–28.2</td>
|
||
</tr>
|
||
<tr id="40AB316BFFF982A3A0FCFBF6FA96FB70" box="[163,1398,1148,1173]" gridrow="22" pageId="17" pageNumber="524">
|
||
<th id="037A5817FFF982A3A0FCFBF6FE39FB70" box="[163,473,1148,1173]" gridcol="0" gridrow="22" pageId="17" pageNumber="524">C length</th>
|
||
<td id="037A5817FFF982A3A1AEFBF6FDB6FB70" box="[497,598,1148,1173]" gridcol="1" gridrow="22" pageId="17" pageNumber="524">24.3–28.7</td>
|
||
<td id="037A5817FFF982A3A2DAFBF6FD0BFB70" box="[645,747,1148,1173]" gridcol="2" gridrow="22" pageId="17" pageNumber="524">23.3–25.9</td>
|
||
<td id="037A5817FFF982A3A369FBF6FC7DFB70" box="[822,925,1148,1173]" gridcol="3" gridrow="22" pageId="17" pageNumber="524">21.6–28.7</td>
|
||
<td id="037A5817FFF982A3A394FBF6FBD0FB70" box="[971,1072,1148,1173]" gridcol="4" gridrow="22" pageId="17" pageNumber="524">22.4–24.4</td>
|
||
<td id="037A5817FFF982A3A423FBF6FB01FB70" box="[1148,1249,1148,1173]" gridcol="5" gridrow="22" pageId="17" pageNumber="524">24.9–30.2</td>
|
||
<td id="037A5817FFF982A3A54FFBF6FA96FB70" box="[1296,1398,1148,1173]" gridcol="6" gridrow="22" pageId="17" pageNumber="524">23.0–28.1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FE243329FFF97D4DA098FB4EFE2FF98F" blockId="17.[151,1437,1220,1931]" pageId="17" pageNumber="524">
|
||
First dorsal fin usually with six spines, but
|
||
<specimenCount id="E89DF8A0FFF97D4DA2EEFB4FFCACFB3B" box="[689,844,1220,1247]" count="1" pageId="17" pageNumber="524" type="generic">one specimen</specimenCount>
|
||
with seven spines. Second dorsal fin with one spine and seven (n=4), eight (n=11), or nine (n=3) soft rays (
|
||
<tableCitation id="B3190692FFF97D4DA348FB62FC92FAE7" box="[791,882,1256,1282]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="17" pageNumber="524">Table 2</tableCitation>
|
||
). First and second dorsal-fin bases separated by a small interval (1.7–4.8% of SL;
|
||
<tableCitation id="B3190692FFF97D4DA278FA86FD9FFAC3" box="[551,639,1292,1318]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="17" pageNumber="524">Table 9</tableCitation>
|
||
). In females, first dorsal fin rounded, almost semi-circular; posterior tip of fin (usually tips of sixth spines) not reaching second dorsal-fin origin. In males, first dorsal fin rounded or parallelogram; posterior tip of fin (usually fourth or fifth spine) extending to, or exceeding base of second dorsal-fin spine, but never exceeding base of second soft ray of second dorsal fin. Anal fin with one spine and six (n=2), seven (n=10), or eight (n=6) soft rays (
|
||
<tableCitation id="B3190692FFF97D4DA20FFA16FD48FA53" box="[592,680,1436,1462]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="17" pageNumber="524">Table 2</tableCitation>
|
||
). Caudal fin with 17 segmented rays including 12 (n=3), 13 (n=2), 14 (n=12), or 15 (n=1) branched rays; posterior margin rounded. Males having larger second dorsal, anal, and caudal fins, than females (second dorsal-fin length 28.8–36.1 vs. 25.7–30.8% of SL, anal-fin length 25.0–33.0 vs. 22.0–28.4% of SL, and caudal-fin length 24.3–28.7 vs. 23.3–25.9% of SL;
|
||
<figureCitation id="66A02FACFFF97D4DA381F982FBC1F9C7" box="[990,1057,1544,1570]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="17" pageNumber="524">Fig. 2</figureCitation>
|
||
,
|
||
<tableCitation id="B3190692FFF97D4DA473F982FB61F9C7" box="[1068,1153,1544,1570]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="17" pageNumber="524">Table 9</tableCitation>
|
||
). Pectoral fin with 15–17 rays (mode 16 rays) (
|
||
<tableCitation id="B3190692FFF97D4DA1D5F9A6FE03F9A3" box="[394,483,1580,1606]" captionStart="TABLE 4" captionStartId="7.[151,237,151,175]" captionText="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="17" pageNumber="524">Table 4</tableCitation>
|
||
). Pelvic fin with one spine and five soft rays; joined together to form a cuplike disc with fleshy bilobed frenum.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFF97D4DA098F9FEFDC3F86E" blockId="17.[151,1437,1220,1931]" pageId="17" pageNumber="524">
|
||
Ctenoid scales covering lateral, dorsal, and ventral surfaces of body posterior to flank except belly. Nape and posterior part of occipital region covered by cycloid scales; scaled area extending anteriorly to area above opercle both in males and females. Other regions on head naked. Belly and flank covered with cycloid scales. First and second dorsal, and anal-fin bases covered with ctenoid scales, and mixed with a few small cycloid scales. Proximal part of caudal fin covered with cycloid scales. Pectoral-fin base and breast (pre-pelvic area) probably naked (at least invisible without staining with alizarin red; see
|
||
<bibRefCitation id="9A0A4ED8FFF97D4DA2B5F8A2FC5DF8A7" author="Suzuki, T. & Shibukawa, K. & Senou, H. & Chen, I. - S." box="[746,957,1832,1858]" pageId="17" pageNumber="524" pagination="227 - 238" refId="ref24564" refString="Suzuki, T., Shibukawa, K., Senou, H. & Chen, I. - S. (2016) Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, 63, 227 - 238. https: // doi. org / 10.1007 / s 10228 - 015 - 0494 - 3" type="journal article" year="2016">
|
||
Suzuki
|
||
<emphasis id="CCEFEF3BFFF97D4DA31FF8A3FC98F8A7" box="[832,888,1832,1858]" italics="true" pageId="17" pageNumber="524">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). Longitudinal scales 30–33 (usually 31 or 32), transverse scales 8–10, transverse scales in caudal peduncle usually seven but six in a single specimen, and predorsal scales 5–12 (
|
||
<tableCitation id="B3190692FFF97D4DA1C8F8FAFDF6F86E" box="[407,534,1904,1931]" captionStart-0="TABLE 5" captionStart-1="TABLE 6" captionStart-2="TABLE 7" captionStartId-0="7.[151,234,868,892]" captionStartId-1="8.[151,236,151,175]" captionStartId-2="8.[151,237,868,892]" captionText-0="TABLE 5. Longitudinal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-1="TABLE 6. Transverse scale counts of various species of Rhinogobius examined in the present study. *Includes the holotype." captionText-2="TABLE 7. Predorsal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="17" pageNumber="524">Tables 5–7</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AAE463A1FFFA7D4EA0C8FAC2FEBDFA4D" ID-DOI="http://doi.org/10.5281/zenodo.13330589" ID-Zenodo-Dep="13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="18" pageNumber="525" startId="18.[151,250,1352,1376]" targetBox="[160,1427,188,1320]" targetPageId="18" targetType="figure">
|
||
<paragraph id="FE243329FFFA7D4EA0C8FAC2FEBDFA4D" blockId="18.[151,1436,1352,1448]" pageId="18" pageNumber="525">
|
||
<emphasis id="CCEFEF3BFFFA7D4EA0C8FAC2FEC0FA85" bold="true" box="[151,288,1352,1376]" pageId="18" pageNumber="525">FIGURE 10.</emphasis>
|
||
Arrangement of cephalic sensory pores and cutaneous sensory papillae in
|
||
<taxonomicName id="399B48AAFFFA7D4EA44FFAC2FB1AFA85" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[1040,1274,1352,1376]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="525" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFFA7D4EA44FFAC2FB1AFA85" box="[1040,1274,1352,1376]" italics="true" pageId="18" pageNumber="525">Rhinogobius taenigena</emphasis>
|
||
</taxonomicName>
|
||
(a–c, OISTICH 303) and
|
||
<taxonomicName id="399B48AAFFFA7D4EA0A9FAE6FE27FA61" box="[246,455,1388,1412]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="525" phylum="Chordata" rank="species" species="leavelli">
|
||
<emphasis id="CCEFEF3BFFFA7D4EA0A9FAE6FE27FA61" box="[246,455,1388,1412]" italics="true" pageId="18" pageNumber="525">Rhinogobius leavelli</emphasis>
|
||
</taxonomicName>
|
||
(d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FE243329FFFA7D4EA098FA5DFDDFF8F4" blockId="18.[151,1437,1495,2025]" pageId="18" pageNumber="525">
|
||
Color in life and fresh (
|
||
<figureCitation id="66A02FACFFFA7D4EA190FA5DFDC9FA14" box="[463,553,1495,1521]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="18" pageNumber="525">Figs. 11</figureCitation>
|
||
and
|
||
<figureCitation id="66A02FACFFFA7D4EA23FFA5DFD60FA14" box="[608,640,1495,1521]" captionStart="FIGURE 12" captionStartId="20.[151,250,1061,1085]" captionTargetBox="[151,1435,181,1037]" captionTargetId="figure-18@20.[151,1435,181,1037]" captionTargetPageId="20" captionText="FIGURE 12. In situ underwater photographs of Rhinogobius taenigena in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietnam. a, b, males, 20 June 2019; c, d, females, 20 June 2019 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330593" httpUri="https://zenodo.org/record/13330593/files/figure.png" pageId="18" pageNumber="525">12</figureCitation>
|
||
): in males, background of head and body grey or yellowish-brown, but whitish on ventral side. Reddish-brown stripe connecting posterior margin of center of upper lip and anterior margin of left and right eyes; thinner and unclear stripe continuing posteriorly from posterior margin of eye, but ending immediately and not reaching posterior margin of preopercle. No or faint reddish-brown marking at center of upper lip. Dorsal side of head often darker than lateral side. Lower half of cheek and opercular region often darker than upper half. Opercular membrane, interopercle, and lower part of subopercle light bluish-grey with many small red or orange spots. Six to eight, dark-brown transverse bands regularly aligned along lateral side of body from flank to caudal-fin base, and intervals light blue, but this pattern sometimes irregular and unclear. Many small, orange spots sometimes scattered on sides of body.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFFA7D48A098F891FD55FAF3" blockId="18.[151,1437,1495,2025]" lastBlockId="20.[151,1437,1168,2022]" lastPageId="20" lastPageNumber="527" pageId="18" pageNumber="525">
|
||
First dorsal-fin with series of black or reddish-brown markings on middle parts of membranes between first to fifth spines; anterior two markings (between first and second, and second and third spines) usually black and clear, but posterior ones reddish-brown and often unclear. Distal part of membrane light blue or white between first and second spines, and orange or red between second and fifth spines. Distal parts of spines black or grey. Reddish-brown spots on proximal part of first to fourth spines; spines and membranes light bluish-grey, both above and below reddish-brown spots. Membranes posterior to fifth spine light bluish-grey with some reddish-brown spots. Second dorsal-fin reddish-brown with four to seven light bluish-grey stripes across fin, and with a broad, orange or pale-yellow band at distal margin. A black, vertical rectangular or elliptical spot at middle proximal part of caudal fin, with a yellowish-brown or grey blotch above black spot, but this blotch often unclear. Other part of caudal fin yellowish or greyish translucent with five or six reddish-brown vertical bands with a pale-yellow margin (
|
||
<figureCitation id="66A02FACFFFB7D4FA56EFE89FA6FFEF8" box="[1329,1423,259,285]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="19" pageNumber="526">Fig. 11b</figureCitation>
|
||
), or dusky grey or reddish-brown with a pale-yellow margin without distinct vertical bands (
|
||
<figureCitation id="66A02FACFFFB7D4FA42EFEADFB2DFEA4" box="[1137,1229,295,321]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="19" pageNumber="526">Fig. 11a</figureCitation>
|
||
). Anal fin orange, but dark brown distally with broad, bluish-white margin. Pectoral fin grey or greyish translucent, its proximal part pale yellowish-brown or pale grey with a black spot at middle of base; an additional black spot often appearing at lower part of pectoral-fin base, which is sometimes combined with middle one forming a large black blotch. Pelvic disc dark grey with a white or light blue margin.
|
||
</paragraph>
|
||
<caption id="AAE463A1FFFB7D4FA0C8F83AFBA0F809" ID-DOI="http://doi.org/10.5281/zenodo.13330591" ID-Zenodo-Dep="13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="19" pageNumber="526" startId="19.[151,250,1968,1992]" targetBox="[151,1435,337,1952]" targetPageId="19" targetType="figure">
|
||
<paragraph id="FE243329FFFB7D4FA0C8F83AFBA0F809" blockId="19.[151,1436,1968,2028]" pageId="19" pageNumber="526">
|
||
<emphasis id="CCEFEF3BFFFB7D4FA0C8F83AFEFFF82D" bold="true" box="[151,287,1968,1992]" pageId="19" pageNumber="526">FIGURE 11.</emphasis>
|
||
<taxonomicName id="399B48AAFFFB7D4FA17AF83AFDF0F82D" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[293,528,1968,1992]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="526" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFFB7D4FA17AF83AFDF0F82D" box="[293,528,1968,1992]" italics="true" pageId="19" pageNumber="526">Rhinogobius taenigena</emphasis>
|
||
</taxonomicName>
|
||
immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AAE463A1FFFC7D48A0C8FBAFFAEFFB87" ID-DOI="http://doi.org/10.5281/zenodo.13330593" ID-Zenodo-Dep="13330593" httpUri="https://zenodo.org/record/13330593/files/figure.png" pageId="20" pageNumber="527" startId="20.[151,250,1061,1085]" targetBox="[151,1435,181,1037]" targetPageId="20" targetType="figure">
|
||
<paragraph id="FE243329FFFC7D48A0C8FBAFFAEFFB87" blockId="20.[151,1437,1061,1122]" pageId="20" pageNumber="527">
|
||
<emphasis id="CCEFEF3BFFFC7D48A0C8FBAFFEC1FBD8" bold="true" box="[151,289,1061,1085]" pageId="20" pageNumber="527">FIGURE 12.</emphasis>
|
||
<emphasis id="CCEFEF3BFFFC7D48A176FBACFE8BFBD8" box="[297,363,1062,1085]" italics="true" pageId="20" pageNumber="527">In situ</emphasis>
|
||
underwater photographs of
|
||
<taxonomicName id="399B48AAFFFC7D48A2D0FBACFC9BFBD8" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[655,891,1062,1086]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="527" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFFC7D48A2D0FBACFC9BFBD8" box="[655,891,1062,1086]" italics="true" pageId="20" pageNumber="527">Rhinogobius taenigena</emphasis>
|
||
</taxonomicName>
|
||
in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietnam. a, b, males, 20 June 2019; c, d, females, 20 June 2019 (photo by K. Maeda).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FE243329FFFC7D48A098FAAAFDB8F9D3" blockId="20.[151,1437,1168,2022]" pageId="20" pageNumber="527">In females, markings and background color of head and body similar to those of males, but opercular membrane, interopercle, and lower part of subopercle yellowish-brown or dark grey without red spots. First dorsal fin having two to four, indefinite brown stripes; intervals translucent or white; distal part of fin yellow or bluish-white. Second dorsal fin with four to five, brown stripes, intervals white or translucent; with a yellow or bluish-white margin. Caudal fin translucent or white with five or six brown bands with a thin, pale-yellow margin; color and markings of proximal part of fin similar to those in males. Anal fin translucent, but reddish-brown or dark brown distally with a bluish-white margin. Pectoral fin translucent with a pattern on its base similar to that in males. Pelvic disc translucent or grey with a white margin.</paragraph>
|
||
<paragraph id="FE243329FFFC7D48A098F9CAFD67F90F" blockId="20.[151,1437,1168,2022]" pageId="20" pageNumber="527">
|
||
Color in preservative (
|
||
<figureCitation id="66A02FACFFFC7D48A196F9CAFDD8F9BF" box="[457,568,1600,1626]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="20" pageNumber="527">Fig. 6e, f</figureCitation>
|
||
): background of head and body cream and those of fins translucent or light brown. Reddish-brown markings observed in life become dark brown, and dusky-brown markings become black. Light bluish-grey color on opercular membrane, interopercle, and lower part of subopercle of male become dark brown, and red and orange spots become light brown (dark and pale pattern opposite from that in life). Red, orange, white, and bluish colors of fins disappeared.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B68160A2FFFC7D49A098F97EFEA9FDD8" lastPageId="21" lastPageNumber="528" pageId="20" pageNumber="527" type="discussion">
|
||
<paragraph id="FE243329FFFC7D49A098F97EFAD1FF30" blockId="20.[151,1437,1168,2022]" lastBlockId="21.[151,1437,151,573]" lastPageId="21" lastPageNumber="528" pageId="20" pageNumber="527">
|
||
<emphasis id="CCEFEF3BFFFC7D48A098F97EFEDBF8EB" bold="true" box="[199,315,1780,1806]" pageId="20" pageNumber="527">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="21208D8BFFFC7D48A125F97EFE4AF8EB" box="[378,426,1780,1806]" pageId="20" pageNumber="527">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="399B48AAFFFC7D48A26BF97FFD24F8EB" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[564,708,1781,1806]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="527" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFFC7D48A26BF97FFD24F8EB" box="[564,708,1781,1806]" italics="true" pageId="20" pageNumber="527">R. taenigena</emphasis>
|
||
</taxonomicName>
|
||
is Xe Pon (Se Pon), a tributary of the Banghiang River in the Mekong basin (
|
||
<bibRefCitation id="9A0A4ED8FFFC7D48A11CF892FE1EF8D7" author="Chen, I. - S. & Kottelat, M. & Miller, P. J." box="[323,510,1816,1842]" pageId="20" pageNumber="527" pagination="19 - 32" refId="ref23100" refString="Chen, I. - S., Kottelat, M. & Miller, P. J. (1999) Freshwater gobies of the genus Rhinogobius from the Mekong basin in Thailand and Laos, with descriptions of three new species. Zoological Studies, 38, 19 - 32." type="journal article" year="1999">
|
||
Chen
|
||
<emphasis id="CCEFEF3BFFFC7D48A1D9F893FE5DF8D7" box="[390,445,1816,1842]" italics="true" pageId="20" pageNumber="527">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
). Though located in
|
||
<collectingCountry id="868C73B9FFFC7D48A280F892FCF8F8D7" box="[735,792,1816,1842]" name="Laos" pageId="20" pageNumber="527">Laos</collectingCountry>
|
||
, it is close to the border with
|
||
<collectingCountry id="868C73B9FFFC7D48A405F892FB59F8D7" box="[1114,1209,1816,1842]" name="Vietnam" pageId="20" pageNumber="527">Vietnam</collectingCountry>
|
||
(a little downstream from the Xe Pon reaches shown in
|
||
<figureCitation id="66A02FACFFFC7D48A274F8B6FD8DF8B3" box="[555,621,1852,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." figureDoi="http://doi.org/10.5281/zenodo.13330563" httpUri="https://zenodo.org/record/13330563/files/figure.png" pageId="20" pageNumber="527">Fig. 1</figureCitation>
|
||
). This species was also reported from
|
||
<collectingRegion id="3C5FFDCBFFFC7D48A475F8B6FAC0F8B3" box="[1066,1312,1852,1878]" country="Vietnam" name="Quang Binh" pageId="20" pageNumber="527">Quang Binh Province</collectingRegion>
|
||
(
|
||
<figureCitation id="66A02FACFFFC7D48A56FF8B6FA92F8B3" box="[1328,1394,1852,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." figureDoi="http://doi.org/10.5281/zenodo.13330563" httpUri="https://zenodo.org/record/13330563/files/figure.png" pageId="20" pageNumber="527">Fig. 1</figureCitation>
|
||
) in
|
||
<collectingCountry id="868C73B9FFFC7D48A0C8F8EAFF16F89F" box="[151,246,1888,1914]" name="Vietnam" pageId="20" pageNumber="527">Vietnam</collectingCountry>
|
||
from the Gianh River basin to Phong Nha River (
|
||
<bibRefCitation id="9A0A4ED8FFFC7D48A36AF8EAFBE0F89F" author="Serov, D. V. & Nezdoliy, V. K. & Pavlov, D. S." box="[821,1024,1888,1914]" pageId="20" pageNumber="527" refId="ref24523" refString="Serov, D. V., Nezdoliy, V. K. & Pavlov, D. S. (2006) The freshwater fishes of central Vietnam. KMK Scientific Press Ltd., Moscow, 364 pp." type="book" year="2006">
|
||
Serov
|
||
<emphasis id="CCEFEF3BFFFC7D48A3DEF8EBFC5BF89F" box="[897,955,1888,1914]" italics="true" pageId="20" pageNumber="527">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
). We found this species in two river systems in Thua Thien Hue Province of
|
||
<collectingCountry id="868C73B9FFFC7D48A23CF80EFD22F87B" box="[611,706,1924,1950]" name="Vietnam" pageId="20" pageNumber="527">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="66A02FACFFFC7D48A28CF80EFCF5F87B" box="[723,789,1924,1950]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." figureDoi="http://doi.org/10.5281/zenodo.13330563" httpUri="https://zenodo.org/record/13330563/files/figure.png" pageId="20" pageNumber="527">Fig. 1</figureCitation>
|
||
). One of them, Xe Kong (Se Kong), is a tributary of the Mekong, originating in Thua Thien Hue Province and flowing through southern
|
||
<collectingCountry id="868C73B9FFFC7D48A450F822FBA5F827" box="[1039,1093,1960,1986]" name="Laos" pageId="20" pageNumber="527">Laos</collectingCountry>
|
||
and eastern
|
||
<collectingCountry id="868C73B9FFFC7D48A490F822FAA6F827" box="[1231,1350,1960,1986]" name="Cambodia" pageId="20" pageNumber="527">Cambodia</collectingCountry>
|
||
. It joins the Mekong near
|
||
<collectingRegion id="3C5FFDCBFFFC7D48A106F846FE05F803" box="[345,485,1996,2022]" country="Cambodia" name="Stung Treng" pageId="20" pageNumber="527">Stung Treng</collectingRegion>
|
||
,
|
||
<collectingCountry id="868C73B9FFFC7D48A1B0F846FD86F803" box="[495,614,1996,2022]" name="Cambodia" pageId="20" pageNumber="527">Cambodia</collectingCountry>
|
||
.Another site is in the upper reaches of the Song Bo (Bo River), a tributary of the Song Huong (Perfume River), which flows directly to the South
|
||
<collectingCountry id="868C73B9FFFD7D49A3FCFF1DFC06FF54" box="[931,998,151,177]" name="China" pageId="21" pageNumber="528">China</collectingCountry>
|
||
Sea near the city of Hue. These suggest that this species could be widely distributed in river systems near the Vietnam-Laos border in this region.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFFD7D49A098FF55FEA9FDD8" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="528">
|
||
<bibRefCitation id="9A0A4ED8FFFD7D49A098FF55FE71FF1C" author="Chen, I. - S. & Kottelat, M. & Miller, P. J." box="[199,401,223,249]" pageId="21" pageNumber="528" pagination="19 - 32" refId="ref23100" refString="Chen, I. - S., Kottelat, M. & Miller, P. J. (1999) Freshwater gobies of the genus Rhinogobius from the Mekong basin in Thailand and Laos, with descriptions of three new species. Zoological Studies, 38, 19 - 32." type="journal article" year="1999">
|
||
Chen
|
||
<emphasis id="CCEFEF3BFFFD7D49A156FF55FEA0FF1C" box="[265,320,223,249]" italics="true" pageId="21" pageNumber="528">et al.</emphasis>
|
||
(1999)
|
||
</bibRefCitation>
|
||
described “four small blackish-brown spots on ventral edge of cheek” (on the ventral margin of the preopercle according to their fig. 16) of a male as a diagnostic character of
|
||
<taxonomicName id="399B48AAFFFD7D49A411FE89FB3EFEF9" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[1102,1246,259,284]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="528" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFFD7D49A411FE89FB3EFEF9" box="[1102,1246,259,284]" italics="true" pageId="21" pageNumber="528">R. taenigena</emphasis>
|
||
</taxonomicName>
|
||
. They described the coloration based on “observations from color slides taken of freshly preserved specimens.”
|
||
<bibRefCitation id="9A0A4ED8FFFD7D49A4ABFEADFE1EFE80" author="Nagao Natural Environment Foundation" pageId="21" pageNumber="528" refId="ref24278" refString="Nagao Natural Environment Foundation (2021) Fishes of the Indochinese Mekong. Nagao Natural Environment Foundation, Tokyo, xii + 546 pp." type="book" year="2021">Nagao Natural Environment Foundation (2021)</bibRefCitation>
|
||
noted that “the spots are not always evident” in specimens freshly collected near the
|
||
<typeStatus id="21208D8BFFFD7D49A0C8FEE5FF27FE6C" box="[151,199,367,393]" pageId="21" pageNumber="528">type</typeStatus>
|
||
locality, and “if present, the number varies from one to four.” Photographs by the
|
||
<bibRefCitation id="9A0A4ED8FFFD7D49A400FEE5FE46FE48" author="Nagao Natural Environment Foundation" pageId="21" pageNumber="528" refId="ref24278" refString="Nagao Natural Environment Foundation (2021) Fishes of the Indochinese Mekong. Nagao Natural Environment Foundation, Tokyo, xii + 546 pp." type="book" year="2021">Nagao Natural Environment Foundation (2021: 441)</bibRefCitation>
|
||
show orange spots similar to spots on the opercular membrane, interopercle, and lower part of subopercle. Our material from
|
||
<collectingCountry id="868C73B9FFFD7D49A21EFE3DFD40FE34" box="[577,672,439,465]" name="Vietnam" pageId="21" pageNumber="528">Vietnam</collectingCountry>
|
||
has neither blackish-brown spots nor spots of any other color on the preopercle, either in fresh or after preservation. This is the only point on which our observations do not agree with previous descriptions. No significant morphological differences were also found between the Xe Kong and Song Bo populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |