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<document id="B8547DCCF116374F28FDF21E1F93EB4D" ID-DOI="10.11646/zootaxa.5496.1.4" ID-ISSN="1175-5326" ID-ZooBank="24953EBD-C29F-4C0B-9CA5-C39F94938047" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723805608457" checkinUser="plazi" docAuthor="Tishechkin, Dmitri Yu." docDate="2024" docId="039D0A61FF932016C894FE23AE98F89C" docLanguage="en" docName="zootaxa.5496.1.4.pdf" docOrigin="Zootaxa 5496 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kybos virgator" docType="treatment" docVersion="1" lastPageNumber="97" masterDocId="FFA47219FF9B201EC803FFF9AB04FFED" masterDocTitle="Leafhoppers of the genus Kybos Fieber, 1866 (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) in Kyrgyzstan" masterLastPageNumber="100" masterPageNumber="89" pageNumber="97" updateTime="1723806196009" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="F32041A4307EBD10C481FFBC2ABD85C5">Leafhoppers of the genus Kybos Fieber, 1866 (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) in Kyrgyzstan</mods:title>
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<mods:namePart id="27429ABCCA5493C30A97EBD7B91C84E6">Tishechkin, Dmitri Yu.</mods:namePart>
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<mods:date id="EE00B93B6D56ACA0A8D5CDF97516E68B">2024</mods:date>
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<mods:number id="B1F4D80B792F03EF6B889E2218B75C10">5496</mods:number>
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<mods:number id="3E3A00479702CB6391B714313196C282">1</mods:number>
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<mods:identifier id="CAB9FB3A4130D59FD2F9B17B67DF50CC" type="DOI">10.11646/zootaxa.5496.1.4</mods:identifier>
<mods:identifier id="F0A92D00C4B60771CF9A5012A7496307" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039D0A61FF932016C894FE23AE98F89C" LSID="urn:lsid:plazi:treatment:039D0A61FF932016C894FE23AE98F89C" httpUri="http://treatment.plazi.org/id/039D0A61FF932016C894FE23AE98F89C" lastPageNumber="97" pageId="8" pageNumber="97">
<subSubSection id="C32EE8FCFF932016C894FE23A906FE19" box="[151,514,474,501]" pageId="8" pageNumber="97" type="nomenclature">
<paragraph id="8B8BBB77FF932016C894FE23A906FE19" blockId="8.[151,514,474,537]" box="[151,514,474,501]" pageId="8" pageNumber="97">
<heading id="D0C30C1BFF932016C894FE23A906FE19" bold="true" box="[151,514,474,501]" fontSize="11" level="1" pageId="8" pageNumber="97" reason="1">
<taxonomicName id="4C34C0F4FF932016C894FE23A906FE19" ID-CoL="3RGWV" authority="(Ribaut, 1933)" baseAuthorityName="Ribaut" baseAuthorityYear="1933" box="[151,514,474,501]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF932016C894FE23A906FE19" bold="true" box="[151,514,474,501]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C894FE23AA4CFE18" bold="true" box="[151,328,474,501]" italics="true" pageId="8" pageNumber="97">Kybos virgator</emphasis>
(Ribaut, 1933)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C894FE06AAB8FDF4" box="[151,444,510,537]" pageId="8" pageNumber="97" type="description">
<paragraph id="8B8BBB77FF932016C894FE06AAB8FDF4" blockId="8.[151,514,474,537]" box="[151,444,510,537]" pageId="8" pageNumber="97">
<figureCitation id="130FA7F2FF932016C894FE06ABE8FDF4" box="[151,236,511,537]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="8" pageNumber="97">Figs 12</figureCitation>
,
<figureCitation id="130FA7F2FF932016C8FBFE06AA12FDF4" box="[248,278,511,537]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="8" pageNumber="97">37</figureCitation>
, 48–49, 57, 63
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C894FDBDA964FD74" pageId="8" pageNumber="97" type="reference_group">
<paragraph id="8B8BBB77FF932016C894FDBDA912FDB6" blockId="8.[151,656,579,665]" box="[151,534,579,603]" pageId="8" pageNumber="97">
<taxonomicName id="4C34C0F4FF932016C894FDBDA912FDB6" ID-CoL="39LDZ" authority="Ribaut, 1933: 152" authorityName="Ribaut" authorityPageNumber="152" authorityYear="1933" box="[151,534,579,603]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF932016C894FDBDAA5DFDB6" box="[151,345,580,603]" italics="true" pageId="8" pageNumber="97">Empoasca virgator</emphasis>
Ribaut, 1933: 152
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF932016C894FD9AA994FD97" blockId="8.[151,656,579,665]" box="[151,656,610,634]" pageId="8" pageNumber="97">
<taxonomicName id="4C34C0F4FF932016C894FD9AA994FD97" ID-CoL="5NQHQ" authority="Wagner, 1935: 28" authorityName="Wagner" authorityPageNumber="28" authorityYear="1935" box="[151,656,610,634]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="variety" species="virgator" variety="saageri">
<emphasis id="B9406765FF932016C894FD9AAA5DFD94" box="[151,345,610,634]" italics="true" pageId="8" pageNumber="97">Empoasca virgator</emphasis>
var.
<emphasis id="B9406765FF932016C988FD9BAAD3FD94" box="[395,471,610,633]" italics="true" pageId="8" pageNumber="97">saageri</emphasis>
Wagner, 1935: 28
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF932016C894FD78A964FD74" blockId="8.[151,656,579,665]" box="[151,608,640,665]" pageId="8" pageNumber="97">
<taxonomicName id="4C34C0F4FF932016C894FD78A964FD74" ID-CoL="39LDZ" authority="Dlabola, 1951: 3" authorityName="Dlabola" authorityPageNumber="3" authorityYear="1951" box="[151,608,640,665]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="virgator" subGenus="Kybos">
<emphasis id="B9406765FF932016C894FD78AA05FD75" box="[151,257,641,664]" italics="true" pageId="8" pageNumber="97">Empoasca</emphasis>
(
<emphasis id="B9406765FF932016C90CFD79AA4AFD75" box="[271,334,640,664]" italics="true" pageId="8" pageNumber="97">Kybos</emphasis>
)
<emphasis id="B9406765FF932016C95DFD78AAB4FD75" box="[350,432,641,664]" italics="true" pageId="8" pageNumber="97">virgator</emphasis>
Dlabola, 1951: 3
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C894FD3FAF6DFC54" pageId="8" pageNumber="97" type="description">
<paragraph id="8B8BBB77FF932016C894FD3FAE15FCE8" blockId="8.[151,1437,710,1205]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C894FD3FAA2DFD0D" bold="true" box="[151,297,710,736]" pageId="8" pageNumber="97">Description.</emphasis>
Brownish green with lighter and darker spots in fore part of body. Forewings smoky hyaline in apical parts, veins of hindwings and dorsal side of abdomen dark, visible through folded forewings (
<figureCitation id="130FA7F2FF932016CCB3FD12AE07FCE8" box="[1200,1283,747,773]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="8" pageNumber="97">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8B8BBB77FF932016C8C4FCF6AF6DFC54" blockId="8.[151,1437,710,1205]" pageId="8" pageNumber="97">
Dorsal apodemes almost undeveloped, look like narrow transverse sclerotized band at base of abdomen. Ventral apodemes about twice as long as wide at base, parallel-sided, touching with inner margins or even converging and somewhat overlapping (
<figureCitation id="130FA7F2FF932016C9A0FCAEAAF2FC9C" box="[419,502,855,881]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="8" pageNumber="97">Fig. 37</figureCitation>
). Aedeagus with well-developed lateral processes (Figs 48–49); occasionally, one or both processes serrated or bear small appendages (visible in lateral view on Fig. 48). Style shape typical of genus. Pygofer appendages narrow (Fig. 57). Anal collar appendages narrow, falcate (Fig. 63).
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C8C4FC3AA966FBEC" pageId="8" pageNumber="97" type="biology_ecology">
<paragraph id="8B8BBB77FF932016C8C4FC3AA966FBEC" blockId="8.[151,1437,710,1205]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C8C4FC3AAA15FC30" bold="true" box="[199,273,963,989]" pageId="8" pageNumber="97">Hosts.</emphasis>
Mainly
<taxonomicName id="4C34C0F4FF932016C972FC3AAADBFC30" box="[369,479,963,989]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9406765FF932016C972FC3AAAADFC30" box="[369,425,963,989]" italics="true" pageId="8" pageNumber="97">Salix</emphasis>
spp.
</taxonomicName>
We once found a population on cultivated
<taxonomicName id="4C34C0F4FF932016CBC7FC3AAF61FC31" box="[964,1125,963,989]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="B9406765FF932016CBC7FC3AAF61FC31" box="[964,1125,963,989]" italics="true" pageId="8" pageNumber="97">Populus nigra</emphasis>
</taxonomicName>
; Dworakowska (1976) also reports
<taxonomicName id="4C34C0F4FF932016C8EFFC1EAA6CFBED" authorityName="Dlabola" authorityYear="1951" box="[236,360,999,1024]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF932016C8EFFC1EAA6CFBED" box="[236,360,999,1024]" italics="true" pageId="8" pageNumber="97">K. virgator</emphasis>
</taxonomicName>
from
<taxonomicName id="4C34C0F4FF932016C9AFFC1EA95FFBEC" box="[428,603,999,1025]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="glutinosa">
<emphasis id="B9406765FF932016C9AFFC1EA95FFBEC" box="[428,603,999,1025]" italics="true" pageId="8" pageNumber="97">Alnus glutinosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C8C4FBF3A807FB80" pageId="8" pageNumber="97" type="distribution">
<paragraph id="8B8BBB77FF932016C8C4FBF3A807FB80" blockId="8.[151,1437,710,1205]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C8C4FBF3AA65FBC9" bold="true" box="[199,353,1034,1060]" pageId="8" pageNumber="97">Distribution.</emphasis>
Europe, European
<collectingCountry id="F323FBE7FF932016CA39FBF2A98CFBC8" box="[570,648,1035,1061]" name="Russia" pageId="8" pageNumber="97">Russia</collectingCountry>
, eastern
<collectingCountry id="F323FBE7FF932016CAE9FBF2A874FBC8" box="[746,880,1035,1061]" name="Kazakhstan" pageId="8" pageNumber="97">Kazakhstan</collectingCountry>
,
<collectingCountry id="F323FBE7FF932016CB78FBF2A8ECFBC8" box="[891,1000,1035,1061]" name="Mongolia" pageId="8" pageNumber="97">Mongolia</collectingCountry>
(Dworakowska, 1976;
<bibRefCitation id="EFA5C686FF932016CCF3FBF2AE95FBC8" author="Mitjaev, I. D." box="[1264,1425,1035,1061]" pageId="8" pageNumber="97" pagination="1 - 170" refId="ref7271" refString="Mitjaev, I. D. (2002) Fauna, Ecology, and Zoogeography of leafhoppers (Homoptera, Cicadinea) of Kazakhstan. Tethys Entomological Research, 5, 1 - 170. [in Russian]" type="journal article" year="2002">Mitjaev, 2002</bibRefCitation>
). In
<collectingCountry id="F323FBE7FF932016C8B7FBD6AA33FBA4" box="[180,311,1071,1097]" name="Kyrgyzstan" pageId="8" pageNumber="97">Kyrgyzstan</collectingCountry>
,
<taxonomicName id="4C34C0F4FF932016C942FBD6AAB8FBA5" authorityName="Dlabola" authorityYear="1951" box="[321,444,1071,1096]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF932016C942FBD6AAB8FBA5" box="[321,444,1071,1096]" italics="true" pageId="8" pageNumber="97">K. virgator</emphasis>
</taxonomicName>
was found only in four localities in the northern part (
<bibRefCitation id="EFA5C686FF932016CC0FFBD6AFD8FBA4" author="Chelpakova, Zh. M." box="[1036,1244,1071,1097]" pageId="8" pageNumber="97" refId="ref6895" refString="Chelpakova, Zh. M. (1994) Auchenorrhyncha of South-Eastern Kyrgyzstan. &quot; Ilim &quot; Publ., Bishkek, 138 pp. [in Russian]" type="book" year="1994">Chelpakova, 1994</bibRefCitation>
), which probably indicates that it is an introduced species in this country.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C8C4FB8EAA58FB58" pageId="8" pageNumber="97" type="discussion">
<paragraph id="8B8BBB77FF932016C8C4FB8EAA58FB58" blockId="8.[151,1437,710,1205]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C8C4FB8EAA3FFB7C" bold="true" box="[199,315,1143,1169]" pageId="8" pageNumber="97">Remarks.</emphasis>
We did not collect this species in
<collectingCountry id="F323FBE7FF932016CACAFB8EA849FB7C" box="[713,845,1143,1169]" name="Kyrgyzstan" pageId="8" pageNumber="97">Kyrgyzstan</collectingCountry>
. The above description is based on specimens from European
<collectingCountry id="F323FBE7FF932016C90AFB62AA53FB58" box="[265,343,1179,1205]" name="Russia" pageId="8" pageNumber="97">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C894FAFEAE98F89C" pageId="8" pageNumber="97" type="key">
<paragraph id="8B8BBB77FF932016C894FAFEA9E9FACC" blockId="8.[151,749,1286,1313]" box="[151,749,1286,1313]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C894FAFEAAE1FACC" bold="true" box="[151,485,1286,1313]" pageId="8" pageNumber="97">Key to species of the genus</emphasis>
<taxonomicName id="4C34C0F4FF932016C9EFFAFEA931FACC" ID-CoL="8HL9H" authorityName="Fieber" authorityYear="1866" box="[492,565,1287,1313]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B9406765FF932016C9EFFAFEA931FACC" box="[492,565,1287,1313]" italics="true" pageId="8" pageNumber="97">Kybos</emphasis>
</taxonomicName>
<emphasis id="B9406765FF932016CA3FFAFFA9E9FACC" bold="true" box="[572,749,1286,1313]" pageId="8" pageNumber="97">
of
<collectingCountry id="F323FBE7FF932016CA5FFAFEA9E9FACC" box="[604,749,1287,1313]" name="Kyrgyzstan" pageId="8" pageNumber="97">Kyrgyzstan</collectingCountry>
</emphasis>
</paragraph>
<key id="FFA5EC46FF932016C894FAB5AE98F89C" pageId="8" pageNumber="97">
<keyStep id="30C0A3D2FF932016C894FAB5AE98FA76" pageId="8" pageNumber="97">
<paragraph id="8B8BBB77FF932016C894FAB5AE99FA8C" blockId="8.[151,1437,1356,1906]" box="[151,1437,1356,1378]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894FAB5AE99FA8C" box="[151,1437,1356,1378]" pageId="8" pageNumber="97">
1. Dorsal abdominal apodemes as long as their width at base or longer (
<figureCitation id="130FA7F2FF932016CB7BFAB5AF2AFA8C" box="[888,1070,1356,1378]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="8" pageNumber="97">Figs 13–14, 17–24</figureCitation>
)............................2.
</keyLead>
</paragraph>
<paragraph id="8B8BBB77FF932016C894FA90AE98FA76" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894FA90AE98FA76" pageId="8" pageNumber="97">
- Dorsal abdominal apodemes are much shorter than their width at base, sometimes are almost undeveloped (
<figureCitation id="130FA7F2FF932016CCFEFA90ABFFFA76" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="8" pageNumber="97">Figs 33–34, 36– 37</figureCitation>
).................................................................................................3.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C0A3D2FF932016C894FA5DAE98F9A1" pageId="8" pageNumber="97">
<paragraph id="8B8BBB77FF932016C894FA5DAE98FA19" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894FA5DAE98FA19" pageId="8" pageNumber="97">
2. Dorsal abdominal apodemes 1.5–2 times longer than wide at base, with strongly divergent inner margins separated by wide V-shaped notch (
<figureCitation id="130FA7F2FF932016C987FA38AAF0FA3B" box="[388,500,1473,1495]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="8" pageNumber="97">Figs 13–14</figureCitation>
). Green with lighter and darker spots in fore part of body, darkest males brownish (
<figureCitation id="130FA7F2FF932016CD11FA38AE6CFA3B" box="[1298,1384,1473,1495]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="8" pageNumber="97">Figs 1–4</figureCitation>
)............................................................................................
<taxonomicName id="4C34C0F4FF932016CD1EFA26AE92FA19" ID-CoL="3RGV6" authorityName="Mitjaev" authorityYear="1971" box="[1309,1430,1503,1524]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF932016CD1EFA26AE92FA19" box="[1309,1430,1503,1524]" italics="true" pageId="8" pageNumber="97">K. montanus</emphasis>
</taxonomicName>
.
</keyLead>
</paragraph>
<paragraph id="8B8BBB77FF932016C894FA05AE98F9A1" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894FA05AE98F9A1" pageId="8" pageNumber="97">
- Dorsal abdominal apodemes of approximately equal length and width at base, with parallel or slightly divergent inner margins separated by rather narrow notch (
<figureCitation id="130FA7F2FF932016CA28F9E0A998F9C3" box="[555,668,1561,1583]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="8" pageNumber="97">Figs 17–24</figureCitation>
). Bright green with lighter spots in fore part of body, old dry specimens slightly yellowish (
<figureCitation id="130FA7F2FF932016C94FF9CFAAA5F9A1" box="[332,417,1590,1612]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="8" pageNumber="97">Figs 5–8</figureCitation>
)..........................................................................
<taxonomicName id="4C34C0F4FF932016CD2BF9CEAE93F9A6" ID-CoL="9G637" authorityName="Zachvatkin" authorityYear="1953" box="[1320,1431,1590,1612]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="oshanini">
<emphasis id="B9406765FF932016CD2BF9CEAE93F9A6" box="[1320,1431,1590,1612]" italics="true" pageId="8" pageNumber="97">K. oshanini</emphasis>
</taxonomicName>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C0A3D2FF932016C894F9ADAE99F92C" pageId="8" pageNumber="97">
<paragraph id="8B8BBB77FF932016C894F9ADAE88F96B" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894F9ADAE88F96B" pageId="8" pageNumber="97">
3. Anal collar appendages wide, with sharply narrowed, extended apices (Fig. 61). Aedeagus with narrowly rounded apex in lateral view and with collar-like expansion at base, distinctly visible in back view (Figs 44–45). On birch. …
<taxonomicName id="4C34C0F4FF932016CCF8F98BAE82F96B" ID-CoL="3RGT3" authorityName="Dlabola" authorityYear="1963" box="[1275,1414,1649,1671]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="auricillatus">
<emphasis id="B9406765FF932016CCF8F98BAE82F96B" box="[1275,1414,1649,1671]" italics="true" pageId="8" pageNumber="97">K. auricillatus</emphasis>
</taxonomicName>
.
</keyLead>
</paragraph>
<paragraph id="8B8BBB77FF932016C894F976AE99F92C" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894F976AE99F92C" pageId="8" pageNumber="97">- Anal collar appendages narrow, falcate (Figs 60, 62–63).Aedeagus with wider apex in lateral view, without collar-like expansion at base (Figs 42–43, 46–49). On poplars or willows..........................................................4.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C0A3D2FF932016C894F930AE98F911" pageId="8" pageNumber="97">
<paragraph id="8B8BBB77FF932016C894F930AE98F933" blockId="8.[151,1437,1356,1906]" box="[151,1436,1737,1759]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894F930AE98F933" box="[151,1436,1737,1759]" pageId="8" pageNumber="97">
4. Pygofer appendages with distinct expansions in apical halves (Fig. 56). On poplars..........................
<taxonomicName id="4C34C0F4FF932016CD3EF933AE92F933" ID-CoL="3RGVN" baseAuthorityName="Edwards" baseAuthorityYear="1908" box="[1341,1430,1737,1759]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="populi">
<emphasis id="B9406765FF932016CD3EF933AE92F933" box="[1341,1430,1737,1759]" italics="true" pageId="8" pageNumber="97">K. populi</emphasis>
</taxonomicName>
.
</keyLead>
</paragraph>
<paragraph id="8B8BBB77FF932016C894F91EAE98F911" blockId="8.[151,1437,1356,1906]" box="[151,1436,1766,1788]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894F91EAE98F911" box="[151,1436,1766,1788]" pageId="8" pageNumber="97">- Pygofer appendages narrow, of uniform width throughout apical halves (Fig. 54, 57)................................5.</keyLead>
</paragraph>
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<paragraph id="8B8BBB77FF932016C894F8FAAE98F8DB" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894F8FAAE98F8DB" pageId="8" pageNumber="97">
5. Aedeagus without lateral processes (Figs 42–43). Live specimens white or greenish white, dry ones yellowish white, sometimes with slight greenish tinge (
<figureCitation id="130FA7F2FF932016C9E1F8D8A919F8DB" box="[482,541,1825,1847]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="8" pageNumber="97">Fig. 9</figureCitation>
). On silver poplar.................................................
<taxonomicName id="4C34C0F4FF932016CD1AF8DBAE92F8DB" ID-CoL="9G7GP" box="[1305,1430,1825,1847]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF932016CD1AF8DBAE92F8DB" box="[1305,1430,1825,1847]" italics="true" pageId="8" pageNumber="97">K. niveicolor</emphasis>
</taxonomicName>
.
</keyLead>
</paragraph>
<paragraph id="8B8BBB77FF932016C894F8C6AE98F89C" blockId="8.[151,1437,1356,1906]" pageId="8" pageNumber="97">
<keyLead id="30C51842FF932016C894F8C6AE98F89C" pageId="8" pageNumber="97">
- Aedeagus with lateral processes (Figs 48–49). Brownish green with lighter and darker spots in fore part of body (
<figureCitation id="130FA7F2FF932016CD25F8C7AE6AF8BE" box="[1318,1390,1854,1876]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="8" pageNumber="97">Fig. 12</figureCitation>
). On willows.....................................................................................
<taxonomicName id="4C34C0F4FF932016CD31F8A5AE98F89C" ID-CoL="39LDZ" authorityName="Dlabola" authorityYear="1951" box="[1330,1436,1884,1905]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF932016CD31F8A5AE98F89C" box="[1330,1436,1884,1905]" italics="true" pageId="8" pageNumber="97">K. virgator</emphasis>
</taxonomicName>
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<taxonomicName id="4C34C0F4FF9A201FC894FD93A916FD69" ID-CoL="3RGV6" authority="Mitjaev, 1971" authorityName="Mitjaev" authorityYear="1971" box="[151,530,618,645]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="montanus">
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<emphasis id="B9406765FF9A201FC894FD93AA65FD68" bold="true" box="[151,353,618,645]" italics="true" pageId="1" pageNumber="90">Kybos montanus</emphasis>
Mitjaev, 1971
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<subSubSection id="C32EE8FCFF9A201FC894FD76A933FBA2" pageId="1" pageNumber="90" type="description">
<paragraph id="8B8BBB77FF9A201FC894FD76A976FD44" blockId="1.[151,626,618,681]" box="[151,626,654,681]" pageId="1" pageNumber="90">
<figureCitation id="130FA7F2FF9A201FC894FD76ABFEFD44" box="[151,250,655,681]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="1" pageNumber="90">Figs 1–4</figureCitation>
,
<figureCitation id="130FA7F2FF9A201FC905FD76AA4AFD44" box="[262,334,655,681]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="1" pageNumber="90">13–16</figureCitation>
, 38–39, 50, 52, 58, 64–69
</paragraph>
<paragraph id="8B8BBB77FF9A201FC894FD2DAA66FCE7" blockId="1.[151,1437,723,778]" pageId="1" pageNumber="90">
<taxonomicName id="4C34C0F4FF9A201FC894FD2DA9C3FD06" authority="Dworakowska, 1976: 440" authorityName="Dworakowska" authorityPageNumber="440" authorityYear="1976" box="[151,711,723,747]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="mitjaevi" subGenus="Kybos">
<emphasis id="B9406765FF9A201FC894FD2DAAB7FD06" box="[151,435,723,747]" italics="true" pageId="1" pageNumber="90">Empoasca (Kybos) mitjaevi</emphasis>
Dworakowska, 1976: 440
</taxonomicName>
, n. nov. for
<taxonomicName id="4C34C0F4FF9A201FCB4DFD2AAF91FD06" authority="Mitjaev, 1971" authorityName="Mitjaev" authorityYear="1971" box="[846,1173,723,747]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9A201FCB4DFD2AA8FEFD06" box="[846,1018,723,747]" italics="true" pageId="1" pageNumber="90">Kybos montanus</emphasis>
Mitjaev, 1971
</taxonomicName>
, nec
<taxonomicName id="4C34C0F4FF9A201FCCD3FD2DAA66FCE7" authority="Caldwell, 1952" authorityName="Caldwell" authorityYear="1952" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="B9406765FF9A201FCCD3FD2DAE99FD06" box="[1232,1437,724,747]" italics="true" pageId="1" pageNumber="90">Empoasca montana</emphasis>
Caldwell, 1952
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF9A201FC894FCC1ABD5FC77" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
<emphasis id="B9406765FF9A201FC894FCC1AA2DFCBF" bold="true" box="[151,297,824,850]" pageId="1" pageNumber="90">Description.</emphasis>
Green with lighter and darker spots in fore part of body, darkest males brownish. Forewings smoky hyaline in apical parts, veins of hindwings and dorsal side of abdomen dark, visible through folded forewings (
<figureCitation id="130FA7F2FF9A201FCD69FCA5ABC1FC77" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="1" pageNumber="90">Figs 1–4</figureCitation>
).
</paragraph>
<paragraph id="8B8BBB77FF9A201FC8C4FC5DA933FBA2" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
Dorsal apodemes are 1.5–2 times longer than wide at base, separated by wide V-shaped notch, usually slightly tapering towards ends (
<figureCitation id="130FA7F2FF9A201FC99BFC31A91EFC0F" box="[408,538,968,994]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="1" pageNumber="90">Figs 13–14</figureCitation>
). Ventral apodemes about twice as long as wide at base, parallel-sided, with inner margins parallel or slightly diverging (
<figureCitation id="130FA7F2FF9A201FCA52FC15A9D2FBEA" box="[593,726,1004,1031]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="1" pageNumber="90">Figs 15–16</figureCitation>
). Aedeagus without lateral processes (Figs 38–39). Style with numerous long setae and serrated falcate apical part (Fig. 50). Pygofer appendages narrow (Fig. 52). Anal collar appendages narrow, falcate (Fig. 58).
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9A201FC8C4FBA1ABEAFA13" pageId="1" pageNumber="90" type="biology_ecology">
<paragraph id="8B8BBB77FF9A201FC8C4FBA1AF01FB9F" blockId="1.[151,1437,824,1823]" box="[199,1029,1112,1139]" pageId="1" pageNumber="90">
<emphasis id="B9406765FF9A201FC8C4FBA1AA15FB9F" bold="true" box="[199,273,1112,1138]" pageId="1" pageNumber="90">Hosts.</emphasis>
<taxonomicName id="4C34C0F4FF9A201FC91BFBA1AA54FB9F" box="[280,336,1112,1138]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="90" phylum="Tracheophyta" rank="genus">
<emphasis id="B9406765FF9A201FC91BFBA1AA54FB9F" box="[280,336,1112,1138]" italics="true" pageId="1" pageNumber="90">Salix</emphasis>
</taxonomicName>
sect.
<taxonomicName id="4C34C0F4FF9A201FC992FBA1AACAFB9F" authorityName="Linnaeus" authorityYear="1758" box="[401,462,1112,1138]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="90" phylum="Mollusca" rank="genus">
<emphasis id="B9406765FF9A201FC992FBA1AACAFB9F" box="[401,462,1112,1138]" italics="true" pageId="1" pageNumber="90">Helix</emphasis>
</taxonomicName>
, possibly, also willow species from other sections.
</paragraph>
<paragraph id="8B8BBB77FF9A201FC8C4FB85AF49FB7B" blockId="1.[151,1437,824,1823]" box="[199,1101,1148,1175]" pageId="1" pageNumber="90">
<emphasis id="B9406765FF9A201FC8C4FB85AA76FB7B" bold="true" box="[199,370,1148,1174]" pageId="1" pageNumber="90">Calling signal.</emphasis>
Signals of males from the following localities were investigated.
</paragraph>
<paragraph id="8B8BBB77FF9A201FC8C4FB59A993FB33" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
1.
<collectingCountry id="F323FBE7FF9A201FC8E5FB58AA6DFB56" box="[230,361,1185,1211]" name="Kyrgyzstan" pageId="1" pageNumber="90">Kyrgyzstan</collectingCountry>
, Alai Range, ca
<quantity id="4CCC1692FF9A201FCA28FB59A971FB57" box="[555,629,1184,1211]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="90" unit="km" value="25.0">25 km</quantity>
southeast of Kyzyl-Kiya, Abshirsay Gorge,
<taxonomicName id="4C34C0F4FF9A201FCC75FB59AFAAFB57" box="[1142,1198,1184,1210]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="90" phylum="Tracheophyta" rank="genus">
<emphasis id="B9406765FF9A201FCC75FB59AFAAFB57" box="[1142,1198,1184,1210]" italics="true" pageId="1" pageNumber="90">Salix</emphasis>
</taxonomicName>
sect.
<taxonomicName id="4C34C0F4FF9A201FCCF6FB59AE36FB57" authorityName="Linnaeus" authorityYear="1758" box="[1269,1330,1184,1210]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="90" phylum="Mollusca" rank="genus">
<emphasis id="B9406765FF9A201FCCF6FB59AE36FB57" box="[1269,1330,1184,1210]" italics="true" pageId="1" pageNumber="90">Helix</emphasis>
</taxonomicName>
,
<date id="FF8A9DB7FF9A201FCD41FB59ABD6FB33" pageId="1" pageNumber="90" value="2014-07-15">15. VII. 2014</date>
, signals of
<specimenCount id="9D3270FEFF9A201FC953FB3DAAC3FB33" box="[336,455,1220,1246]" count="4" pageId="1" pageNumber="90" type="male">four males</specimenCount>
recorded at 28
<superScript id="7C41163FFF9A201FCA76FB3DA990FB33" attach="right" box="[629,660,1220,1246]" fontSize="6" pageId="1" pageNumber="90">oC</superScript>
.
</paragraph>
<paragraph id="8B8BBB77FF9A201FC8C4FB11A902FACB" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
2.
<collectingCountry id="F323FBE7FF9A201FC8E0FB10AA62FAEE" box="[227,358,1257,1283]" name="Kyrgyzstan" pageId="1" pageNumber="90">Kyrgyzstan</collectingCountry>
, Suusamyr River ca
<quantity id="4CCC1692FF9A201FCA48FB11A987FAEF" box="[587,643,1256,1282]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="90" unit="km" value="4.0">4 km</quantity>
southeast of Suusamyr Village,
<taxonomicName id="4C34C0F4FF9A201FCBE9FB11AF26FAEF" box="[1002,1058,1256,1282]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="90" phylum="Tracheophyta" rank="genus">
<emphasis id="B9406765FF9A201FCBE9FB11AF26FAEF" box="[1002,1058,1256,1282]" italics="true" pageId="1" pageNumber="90">Salix</emphasis>
</taxonomicName>
sect.
<taxonomicName id="4C34C0F4FF9A201FCC61FB11AF9BFAEF" authorityName="Linnaeus" authorityYear="1758" box="[1122,1183,1256,1282]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="90" phylum="Mollusca" rank="genus">
<emphasis id="B9406765FF9A201FCC61FB11AF9BFAEF" box="[1122,1183,1256,1282]" italics="true" pageId="1" pageNumber="90">Helix</emphasis>
</taxonomicName>
,
<date id="FF8A9DB7FF9A201FCCAFFB11AE40FAEF" box="[1196,1348,1256,1283]" pageId="1" pageNumber="90" value="2023-07-12">12. VII. 2023</date>
, signals of
<specimenCount id="9D3270FEFF9A201FC8B5FAF5AA31FACB" box="[182,309,1292,1318]" count="3" pageId="1" pageNumber="90" type="male">three males</specimenCount>
recorded at 25
<superScript id="7C41163FFF9A201FC9E7FAF5A906FACB" attach="right" box="[484,514,1292,1318]" fontSize="6" pageId="1" pageNumber="90">oC</superScript>
.
</paragraph>
<paragraph id="8B8BBB77FF9A201FC8C4FAC9AFA5FA83" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
3.
<collectingCountry id="F323FBE7FF9A201FC8E5FAC8AA6DFAA6" box="[230,361,1329,1355]" name="Kyrgyzstan" pageId="1" pageNumber="90">Kyrgyzstan</collectingCountry>
, Western Karakol River (the Kokomeren River basin)
<quantity id="4CCC1692FF9A201FCBDDFAC9AF1EFAA7" box="[990,1050,1328,1354]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="90" unit="km" value="8.0">8 km</quantity>
from the mouth (ca
<quantity id="4CCC1692FF9A201FCD08FAC9AE43FAA7" box="[1291,1351,1328,1354]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="90" unit="km" value="8.0">8 km</quantity>
east of Suusamyr Village),
<taxonomicName id="4C34C0F4FF9A201FC975FAADAAAAFA83" box="[374,430,1364,1390]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="90" phylum="Tracheophyta" rank="genus">
<emphasis id="B9406765FF9A201FC975FAADAAAAFA83" box="[374,430,1364,1390]" italics="true" pageId="1" pageNumber="90">Salix</emphasis>
</taxonomicName>
sect.
<taxonomicName id="4C34C0F4FF9A201FC9EDFAADA92FFA83" authorityName="Linnaeus" authorityYear="1758" box="[494,555,1364,1390]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="90" phylum="Mollusca" rank="genus">
<emphasis id="B9406765FF9A201FC9EDFAADA92FFA83" box="[494,555,1364,1390]" italics="true" pageId="1" pageNumber="90">Helix</emphasis>
</taxonomicName>
,
<date id="FF8A9DB7FF9A201FCA3AFAADA9D6FA83" box="[569,722,1364,1391]" pageId="1" pageNumber="90" value="2023-07-12">12. VII. 2023</date>
, signals of
<specimenCount id="9D3270FEFF9A201FCB53FAADA8D4FA83" box="[848,976,1364,1390]" count="3" pageId="1" pageNumber="90" type="male">three males</specimenCount>
recorded at 24
<superScript id="7C41163FFF9A201FCC7CFAADAF99FA83" attach="right" box="[1151,1181,1364,1390]" fontSize="6" pageId="1" pageNumber="90">oC</superScript>
.
</paragraph>
<paragraph id="8B8BBB77FF9A201FC8C4FA81ABEAFA13" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
Calling signal is a phrase consisting of 2–6 syllables, following each other with a period of 350–900 ms (
<figureCitation id="130FA7F2FF9A201FC89CFA65AA22FA5A" box="[159,294,1436,1463]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="1" pageNumber="90">Figs 64–65</figureCitation>
). Each syllable consists of several partially merged pulses; in some signals pulses in syllables are indistinguishable (
<figureCitation id="130FA7F2FF9A201FC964FA39AAE8FA37" box="[359,492,1472,1498]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="1" pageNumber="90">Figs 66–69</figureCitation>
). Occasionally, the first syllable is preceded by several low-amplitude pulses (
<figureCitation id="130FA7F2FF9A201FCD69FA39ABE5FA13" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="1" pageNumber="90">Figs 64, 66</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9A201FC8C4F9F1AF7BF962" pageId="1" pageNumber="90" type="distribution">
<paragraph id="8B8BBB77FF9A201FC8C4F9F1AF7BF962" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
<emphasis id="B9406765FF9A201FC8C4F9F1AA65F9CF" bold="true" box="[199,353,1544,1570]" pageId="1" pageNumber="90">Distribution.</emphasis>
<bibRefCitation id="EFA5C686FF9A201FC968F9F1A913F9CF" author="Mitjaev, I. D." box="[363,535,1544,1570]" pageId="1" pageNumber="90" pagination="1 - 170" refId="ref7271" refString="Mitjaev, I. D. (2002) Fauna, Ecology, and Zoogeography of leafhoppers (Homoptera, Cicadinea) of Kazakhstan. Tethys Entomological Research, 5, 1 - 170. [in Russian]" type="journal article" year="2002">Mitjaev (2002)</bibRefCitation>
records this species from the mountains of southern and eastern
<collectingCountry id="F323FBE7FF9A201FCD10F9F1AE9DF9CF" box="[1299,1433,1544,1570]" name="Kazakhstan" pageId="1" pageNumber="90">Kazakhstan</collectingCountry>
,
<collectingCountry id="F323FBE7FF9A201FC894F9D4AA1EF9AA" box="[151,282,1581,1607]" name="Kyrgyzstan" pageId="1" pageNumber="90">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="F323FBE7FF9A201FC924F9D5AAACF9AB" box="[295,424,1580,1606]" name="Uzbekistan" pageId="1" pageNumber="90">Uzbekistan</collectingCountry>
, and
<collectingCountry id="F323FBE7FF9A201FC9E7F9D5A951F9AB" box="[484,597,1580,1606]" name="Tajikistan" pageId="1" pageNumber="90">Tajikistan</collectingCountry>
, although no specific locations in
<collectingCountry id="F323FBE7FF9A201FCBDAF9D4AF5EF9AA" box="[985,1114,1581,1607]" name="Kyrgyzstan" pageId="1" pageNumber="90">Kyrgyzstan</collectingCountry>
either in his works or in the articles of other authors were given. According to our observations, common on willows in North and Inner Tien Shan and in Alai Range; in West Tien Shan occurs only in the forest and subalpine zones.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9A201FC8C4F961A808F8F3" pageId="1" pageNumber="90" type="discussion">
<paragraph id="8B8BBB77FF9A201FC8C4F961A808F8F3" blockId="1.[151,1437,824,1823]" pageId="1" pageNumber="90">
<emphasis id="B9406765FF9A201FC8C4F961AA34F95F" bold="true" box="[199,304,1688,1714]" pageId="1" pageNumber="90">Remark.</emphasis>
Dworakowska (1976) treated
<taxonomicName id="4C34C0F4FF9A201FCA86F961A9CFF95F" authorityName="Fieber" authorityYear="1866" box="[645,715,1688,1714]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9406765FF9A201FCA86F961A9CFF95F" box="[645,715,1688,1714]" italics="true" pageId="1" pageNumber="90">Kybos</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="4C34C0F4FF9A201FCB92F960AF02F95F" authorityName="Walsh" authorityYear="1862" box="[913,1030,1689,1714]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9406765FF9A201FCB92F960AF02F95F" box="[913,1030,1689,1714]" italics="true" pageId="1" pageNumber="90">Empoasca</emphasis>
</taxonomicName>
and replaced the name
<taxonomicName id="4C34C0F4FF9A201FCD0CF960AE98F95F" authorityName="Mitjaev" authorityYear="1971" box="[1295,1436,1689,1714]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9A201FCD0CF960AE98F95F" box="[1295,1436,1689,1714]" italics="true" pageId="1" pageNumber="90">K. montanus</emphasis>
</taxonomicName>
by
<taxonomicName id="4C34C0F4FF9A201FC8BFF944AA6DF93B" authorityName="Dworakowska" authorityYear="1976" box="[188,361,1724,1750]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="mitjaevi" subGenus="Kybos">
<emphasis id="B9406765FF9A201FC8BFF944AA6DF93B" box="[188,361,1724,1750]" italics="true" pageId="1" pageNumber="90">E. (K.) mitjaevi</emphasis>
</taxonomicName>
as a junior secondary homonym. Here we treat
<taxonomicName id="4C34C0F4FF9A201FCB8CF945A8D1F93B" authorityName="Fieber" authorityYear="1866" box="[911,981,1724,1750]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9406765FF9A201FCB8CF945A8D1F93B" box="[911,981,1724,1750]" italics="true" pageId="1" pageNumber="90">Kybos</emphasis>
</taxonomicName>
as a separate genus; for this reason, the name
<taxonomicName id="4C34C0F4FF9A201FC8DFF918A914F917" authority="Mitjaev, 1971" authorityName="Mitjaev" authorityYear="1971" box="[220,528,1760,1786]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9A201FC8DFF918AA6FF917" box="[220,363,1761,1786]" italics="true" pageId="1" pageNumber="90">K. montanus</emphasis>
Mitjaev, 1971
</taxonomicName>
was reinstated. However,
<bibRefCitation id="EFA5C686FF9A201FCB3FF919A8E4F917" author="Mitjaev, I. D." box="[828,992,1760,1786]" pageId="1" pageNumber="90" pagination="1 - 170" refId="ref7271" refString="Mitjaev, I. D. (2002) Fauna, Ecology, and Zoogeography of leafhoppers (Homoptera, Cicadinea) of Kazakhstan. Tethys Entomological Research, 5, 1 - 170. [in Russian]" type="journal article" year="2002">Mitjaev (2002</bibRefCitation>
,
<bibRefCitation id="EFA5C686FF9A201FCBEFF919AF23F916" author="Mitjaev, I. D." box="[1004,1063,1760,1787]" pageId="1" pageNumber="90" pagination="43 - 81" refId="ref7310" refString="Mitjaev, I. D. (2015) Leafhoppers (Homoptera, Cicadinea) of Kazakhstan, annotated check-list of species. Selevinia, 23, 43 - 81. [in Russian with English summary]" type="journal article" year="2015">2015</bibRefCitation>
, etc.), treating
<taxonomicName id="4C34C0F4FF9A201FCCD2F919AE13F917" authorityName="Fieber" authorityYear="1866" box="[1233,1303,1760,1786]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9406765FF9A201FCCD2F919AE13F917" box="[1233,1303,1760,1786]" italics="true" pageId="1" pageNumber="90">Kybos</emphasis>
</taxonomicName>
as a genus, still used the name
<taxonomicName id="4C34C0F4FF9A201FC96CF8FCAAEEF8F3" authorityName="Dworakowska" authorityYear="1976" box="[367,490,1797,1822]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="mitjaevi">
<emphasis id="B9406765FF9A201FC96CF8FCAAEEF8F3" box="[367,490,1797,1822]" italics="true" pageId="1" pageNumber="90">K. mitjaevi</emphasis>
</taxonomicName>
in all his faunistic works.
</paragraph>
</subSubSection>
</treatment>
</document>

223
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@ -0,0 +1,223 @@
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<subSubSection id="C32EE8FCFF9D2018C894F9FEA90DF9CF" box="[151,521,1543,1570]" pageId="6" pageNumber="95" type="nomenclature">
<paragraph id="8B8BBB77FF9D2018C894F9FEA90DF9CF" blockId="6.[151,528,1543,1607]" box="[151,521,1543,1570]" pageId="6" pageNumber="95">
<heading id="D0C30C1BFF9D2018C894F9FEA90DF9CF" bold="true" box="[151,521,1543,1570]" fontSize="11" level="1" pageId="6" pageNumber="95" reason="1">
<taxonomicName id="4C34C0F4FF9D2018C894F9FEA90DF9CF" ID-CoL="3RGVN" authority="(Edwards, 1908)" baseAuthorityName="Edwards" baseAuthorityYear="1908" box="[151,521,1543,1570]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="populi">
<emphasis id="B9406765FF9D2018C894F9FEA90DF9CF" bold="true" box="[151,521,1543,1570]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C894F9FEAA31F9CF" bold="true" box="[151,309,1543,1570]" italics="true" pageId="6" pageNumber="95">Kybos populi</emphasis>
(Edwards, 1908)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C894F9D5A914F9AB" box="[151,528,1580,1607]" pageId="6" pageNumber="95" type="description">
<paragraph id="8B8BBB77FF9D2018C894F9D5A914F9AB" blockId="6.[151,528,1543,1607]" box="[151,528,1580,1607]" pageId="6" pageNumber="95">
<figureCitation id="130FA7F2FF9D2018C894F9D5ABEFF9AB" box="[151,235,1580,1606]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="6" pageNumber="95">Figs 11</figureCitation>
,
<figureCitation id="130FA7F2FF9D2018C8F4F9D5AA11F9AB" box="[247,277,1580,1606]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="6" pageNumber="95">36</figureCitation>
, 46–47, 56, 62, 77–78
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C894F988A912F909" pageId="6" pageNumber="95" type="reference_group">
<paragraph id="8B8BBB77FF9D2018C894F988A908F964" blockId="6.[151,683,1648,1764]" box="[151,524,1648,1673]" pageId="6" pageNumber="95">
<taxonomicName id="4C34C0F4FF9D2018C894F988A908F964" ID-CoL="8J3RM" authority="Edwards, 1908: 81" authorityName="Edwards" authorityPageNumber="81" authorityYear="1908" box="[151,524,1648,1673]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="populi">
<emphasis id="B9406765FF9D2018C894F988AA4CF965" box="[151,328,1648,1672]" italics="true" pageId="6" pageNumber="95">Empoasca populi</emphasis>
Edwards, 1908: 81
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF9D2018C894F969A9AFF94A" blockId="6.[151,683,1648,1764]" box="[151,683,1679,1703]" pageId="6" pageNumber="95">
<taxonomicName id="4C34C0F4FF9D2018C894F969A9AFF94A" ID-CoL="8J3RM" authority="Kloet &amp; Hincks, 1945: 57" authorityName="Kloet &amp; Hincks" authorityPageNumber="57" authorityYear="1945" box="[151,683,1679,1703]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="populi" subGenus="Kybos">
<emphasis id="B9406765FF9D2018C894F969AA05F94A" box="[151,257,1680,1703]" italics="true" pageId="6" pageNumber="95">Empoasca</emphasis>
(
<emphasis id="B9406765FF9D2018C90CF976AA4AF94A" box="[271,334,1679,1703]" italics="true" pageId="6" pageNumber="95">Kybos</emphasis>
)
<emphasis id="B9406765FF9D2018C95DF976AA9BF94A" box="[350,415,1679,1703]" italics="true" pageId="6" pageNumber="95">populi</emphasis>
Kloet &amp; Hincks, 1945: 57
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF9D2018C894F957A96CF92B" blockId="6.[151,683,1648,1764]" box="[151,616,1710,1734]" pageId="6" pageNumber="95">
<taxonomicName id="4C34C0F4FF9D2018C894F957A96CF92B" ID-CoL="9G7KR" authority="Zachvatkin, 1953: 208" authorityName="Zachvatkin" authorityPageNumber="208" authorityYear="1953" box="[151,616,1710,1734]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="populi" subSpecies="tremulae">
<emphasis id="B9406765FF9D2018C894F957AA79F92B" box="[151,381,1710,1734]" italics="true" pageId="6" pageNumber="95">Kybos populi tremulae</emphasis>
Zachvatkin, 1953: 208
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF9D2018C894F935A912F909" blockId="6.[151,683,1648,1764]" box="[151,534,1740,1764]" pageId="6" pageNumber="95">
<taxonomicName id="4C34C0F4FF9D2018C894F935A912F909" ID-CoL="3RGWY" authority="Mitjaev, 1968: 635" authorityName="Mitjaev" authorityPageNumber="635" authorityYear="1968" box="[151,534,1740,1764]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="zaisanensis">
<emphasis id="B9406765FF9D2018C894F935AA54F909" box="[151,336,1740,1764]" italics="true" pageId="6" pageNumber="95">Kybos zaisanensis</emphasis>
Mitjaev, 1968: 635
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2019C894F8EBAA4FF965" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95" type="description">
<paragraph id="8B8BBB77FF9D2018C894F8EBAF00F8BC" blockId="6.[151,1437,1810,2017]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C894F8EBAA2DF8C1" bold="true" box="[151,297,1810,1836]" pageId="6" pageNumber="95">Description.</emphasis>
Brownish green with lighter and darker spots in fore part of body. Forewings smoky hyaline in apical parts, dorsal side of abdomen dark, visible through folded forewings (
<figureCitation id="130FA7F2FF9D2018CBA7F8CEA8F2F8BC" box="[932,1014,1847,1873]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="6" pageNumber="95">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8B8BBB77FF9D2018C8C4F8A2A87CF80C" blockId="6.[151,1437,1810,2017]" pageId="6" pageNumber="95">
Dorsal apodemes almost undeveloped, look like narrow transverse sclerotized band at base of abdomen. Ventral apodemes about twice as long as wide at base, separated by narrow V-shaped notch (
<figureCitation id="130FA7F2FF9D2018CC66F886AFB8F874" box="[1125,1212,1919,1945]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="6" pageNumber="95">Fig. 36</figureCitation>
). Aedeagus without lateral processes (Figs 46–47). Style shape typical of genus. Pygofer appendages with distinct expansions in apical halves (Fig. 56). Anal collar appendages narrow, falcate (Fig. 62).
</paragraph>
<paragraph id="8B8BBB77FF9C2019C89BF9FDAA4FF965" blockId="7.[151,1436,1540,1672]" pageId="7" pageNumber="96">
<emphasis id="B9406765FF9C2019C89BF9FDAA58F9F1" bold="true" box="[152,348,1540,1564]" pageId="7" pageNumber="96">FIGURES 38–63.</emphasis>
<taxonomicName id="4C34C0F4FF9C2019C960F9FDAAD2F9F1" box="[355,470,1540,1564]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9406765FF9C2019C960F9FDAAA6F9F1" box="[355,418,1540,1564]" italics="true" pageId="7" pageNumber="96">Kybos</emphasis>
spp.
</taxonomicName>
, male genitalia. 38–39, 50, 52, 58—
<taxonomicName id="4C34C0F4FF9C2019CB4EF9FCA8CBF9F1" authorityName="Mitjaev" authorityYear="1971" box="[845,975,1541,1564]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9C2019CB4EF9FCA8CBF9F1" box="[845,975,1541,1564]" italics="true" pageId="7" pageNumber="96">K. montanus</emphasis>
</taxonomicName>
; 40–41, 53, 59—
<taxonomicName id="4C34C0F4FF9C2019CC87F9FCAFFDF9F1" authorityName="Zachvatkin" authorityYear="1953" box="[1156,1273,1540,1564]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="oshanini">
<emphasis id="B9406765FF9C2019CC87F9FCAFFDF9F1" box="[1156,1273,1540,1564]" italics="true" pageId="7" pageNumber="96">K. oshanini</emphasis>
</taxonomicName>
; 42–43, 51, 54, 60—
<taxonomicName id="4C34C0F4FF9C2019C8C9F9D0AA4BF9AD" box="[202,335,1576,1600]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9C2019C8C9F9D0AA4BF9AD" box="[202,335,1576,1600]" italics="true" pageId="7" pageNumber="96">K. niveicolor</emphasis>
</taxonomicName>
; 44–45, 55, 61—
<taxonomicName id="4C34C0F4FF9C2019CA07F9D0A99CF9AD" authorityName="Dlabola" authorityYear="1963" box="[516,664,1576,1600]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="auricillatus">
<emphasis id="B9406765FF9C2019CA07F9D0A99CF9AD" box="[516,664,1576,1600]" italics="true" pageId="7" pageNumber="96">K. auricillatus</emphasis>
</taxonomicName>
; 46–47, 56, 62—
<taxonomicName id="4C34C0F4FF9C2019CB4EF9D0A8A9F9AD" baseAuthorityName="Edwards" baseAuthorityYear="1908" box="[845,941,1576,1600]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="populi">
<emphasis id="B9406765FF9C2019CB4EF9D0A8A9F9AD" box="[845,941,1576,1600]" italics="true" pageId="7" pageNumber="96">K. populi</emphasis>
</taxonomicName>
; 48–49, 57, 63—
<taxonomicName id="4C34C0F4FF9C2019CC61F9D0AFD6F9AD" baseAuthorityName="Ribaut" baseAuthorityYear="1933" box="[1122,1234,1577,1600]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF9C2019CC61F9D0AFD6F9AD" box="[1122,1234,1577,1600]" italics="true" pageId="7" pageNumber="96">K. virgator</emphasis>
</taxonomicName>
. 38, 40, 42, 44, 46, 48—aedeagus, lateral view, 39, 41, 43, 45, 47, 49—same, dorsal view, 50–51—style, 52–57—pygofer appendage, 58–63—anal collar appendage.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9C2019C8C4F923A82AF820" pageId="7" pageNumber="96" type="biology_ecology">
<paragraph id="8B8BBB77FF9C2019C8C4F923AE9CF919" blockId="7.[151,1437,1754,1997]" box="[199,1432,1754,1781]" pageId="7" pageNumber="96">
<emphasis id="B9406765FF9C2019C8C4F923AA15F919" bold="true" box="[199,273,1754,1780]" pageId="7" pageNumber="96">Hosts.</emphasis>
On many poplar species including
<emphasis id="B9406765FF9C2019CA9DF923AF1CF919" box="[670,1048,1754,1780]" italics="true" pageId="7" pageNumber="96">
<taxonomicName id="4C34C0F4FF9C2019CA9DF923A85DF919" box="[670,857,1754,1780]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="species" species="tremula">Populus tremula</taxonomicName>
,
<taxonomicName id="4C34C0F4FF9C2019CB66F922A8B0F919" baseAuthorityName="Skvortsov" baseAuthorityYear="1972" box="[869,948,1754,1780]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="species" species="alba">P. alba</taxonomicName>
,
<taxonomicName id="4C34C0F4FF9C2019CBC3F922AF1CF919" box="[960,1048,1755,1780]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="species" species="nigra">P. nigra</taxonomicName>
</emphasis>
, and related species and cultivars.
</paragraph>
<paragraph id="8B8BBB77FF9C2019C8C4F907A8D3F8D1" blockId="7.[151,1437,1754,1997]" pageId="7" pageNumber="96">
<emphasis id="B9406765FF9C2019C8C4F907AA77F8F5" bold="true" box="[199,371,1790,1816]" pageId="7" pageNumber="96">Calling signal.</emphasis>
<materialsCitation id="3B5CB12AFF9C2019C978F907A8D7F8D1" collectingDate="2021-07-23" country="Russia" location="Voskresensk Region" municipality="Beloozerskiy Town" pageId="7" pageNumber="96" specimenCount="2" specimenCount-male="2" stateProvince="Moscow">
Signals of
<specimenCount id="9D3270FEFF9C2019C9F7F906A963F8F5" box="[500,615,1790,1817]" count="2" pageId="7" pageNumber="96" type="male">two males</specimenCount>
from
<collectingCountry id="F323FBE7FF9C2019CAAEF907A9FCF8F5" box="[685,760,1790,1816]" name="Russia" pageId="7" pageNumber="96">Russia</collectingCountry>
(
<collectingMunicipality id="6BEF210DFF9C2019CB04F907A8E2F8F4" box="[775,998,1790,1817]" pageId="7" pageNumber="96">Beloozerskiy Town</collectingMunicipality>
,
<location id="8EEBEDACFF9C2019CBF2F907AFDCF8F5" LSID="urn:lsid:plazi:treatment:039D0A61FF9D2016C894F9FEABEDFE64:8EEBEDACFF9C2019CBF2F907AFDCF8F5" box="[1009,1240,1790,1816]" country="Russia" municipality="Beloozerskiy Town" name="Voskresensk Region" pageId="7" pageNumber="96" stateProvince="Moscow">Voskresensk Region</location>
,
<collectingRegion id="49F07595FF9C2019CCE7F906AE9CF8F5" box="[1252,1432,1790,1817]" country="Russia" name="Moscow" pageId="7" pageNumber="96">Moscow Oblast</collectingRegion>
, on cultivated
<taxonomicName id="4C34C0F4FF9C2019C933F8DAAA79F8D1" baseAuthorityName="Skvortsov" baseAuthorityYear="1972" box="[304,381,1826,1852]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="B9406765FF9C2019C933F8DAAA79F8D1" box="[304,381,1826,1852]" italics="true" pageId="7" pageNumber="96">P. alba</emphasis>
</taxonomicName>
,
<date id="FF8A9DB7FF9C2019C988F8DBA920F8D1" box="[395,548,1826,1853]" pageId="7" pageNumber="96" value="2021-07-23">
<collectingDate id="EFCE645FFF9C2019C988F8DBA920F8D1" box="[395,548,1826,1853]" pageId="7" pageNumber="96" value="2021-07-23">23. VII. 2021</collectingDate>
</date>
, recording at 25
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) were investigated
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BBB77FF9C2019C8C4F8BFA82AF820" blockId="7.[151,1437,1754,1997]" pageId="7" pageNumber="96">
The temporal pattern of the calling signal is generally the same as in
<taxonomicName id="4C34C0F4FF9C2019CBD9F8BEAF6AF88D" box="[986,1134,1862,1888]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9C2019CBD9F8BEAF6AF88D" box="[986,1134,1862,1888]" italics="true" pageId="7" pageNumber="96">K. niveicolor</emphasis>
</taxonomicName>
. It is a succession of 5–10 pulses, following each other with a period of 300–480 ms; usually, the pulse repetition period increases towards the end of a signal (
<figureCitation id="130FA7F2FF9C2019C944F877AA9EF845" box="[327,410,1934,1960]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="7" pageNumber="96">Fig. 77</figureCitation>
). Pulse duration averages 65–95 ms (
<figureCitation id="130FA7F2FF9C2019CB35F877A88DF845" box="[822,905,1934,1960]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="7" pageNumber="96">Fig. 78</figureCitation>
). Differs from
<taxonomicName id="4C34C0F4FF9C2019CC2EF876AFBBF845" box="[1069,1215,1934,1960]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9C2019CC2EF876AFBBF845" box="[1069,1215,1934,1960]" italics="true" pageId="7" pageNumber="96">K. niveicolor</emphasis>
</taxonomicName>
by distinctly longer pulses with an abrupt amplitude outbreak in the initial part.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C8C4FF6FAA1BFEF0" pageId="8" pageNumber="97" type="distribution">
<paragraph id="8B8BBB77FF932016C8C4FF6FAA1BFEF0" blockId="8.[151,1436,150,393]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C8C4FF6FAA65FF5D" bold="true" box="[199,353,150,176]" pageId="8" pageNumber="97">Distribution.</emphasis>
Transpalearctic within the boreal zone. In
<collectingCountry id="F323FBE7FF932016CB52FF6EA8D0FF5C" box="[849,980,151,177]" name="Kyrgyzstan" pageId="8" pageNumber="97">Kyrgyzstan</collectingCountry>
, was found only in
<collectingRegion id="49F07595FF932016CCBEFF6EAE1DFF5C" box="[1213,1305,151,177]" country="Kyrgyzstan" name="Bishkek Shaary" pageId="8" pageNumber="97">Bishkek</collectingRegion>
and in one village on the northern coast of the Issyk-Kul Lake (
<bibRefCitation id="EFA5C686FF932016CAF9FF42A8CAFF38" author="Chelpakova, Zh. M." box="[762,974,187,213]" pageId="8" pageNumber="97" refId="ref6895" refString="Chelpakova, Zh. M. (1994) Auchenorrhyncha of South-Eastern Kyrgyzstan. &quot; Ilim &quot; Publ., Bishkek, 138 pp. [in Russian]" type="book" year="1994">Chelpakova, 1994</bibRefCitation>
), despite the fact that its host plants are among the most common trees in all regions of this country. This probably indicates that
<taxonomicName id="4C34C0F4FF932016CC7DFF26AFE3FF14" baseAuthorityName="Edwards" baseAuthorityYear="1908" box="[1150,1255,223,249]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="populi">
<emphasis id="B9406765FF932016CC7DFF26AFE3FF14" box="[1150,1255,223,249]" italics="true" pageId="8" pageNumber="97">K. populi</emphasis>
</taxonomicName>
is introduced in
<collectingCountry id="F323FBE7FF932016C894FEFAAA1FFEF0" box="[151,283,259,285]" name="Kyrgyzstan" pageId="8" pageNumber="97">Kyrgyzstan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF932016C8C4FEDEABEDFE64" pageId="8" pageNumber="97" type="discussion">
<paragraph id="8B8BBB77FF932016C8C4FEDEABEDFE64" blockId="8.[151,1436,150,393]" pageId="8" pageNumber="97">
<emphasis id="B9406765FF932016C8C4FEDEAA3FFEAC" bold="true" box="[199,315,295,321]" pageId="8" pageNumber="97">Remarks.</emphasis>
Despite collecting leafhoppers on different species of poplars throughout almost all regions of
<collectingCountry id="F323FBE7FF932016C894FEB2AA1CFE88" box="[151,280,331,357]" name="Kyrgyzstan" pageId="8" pageNumber="97">Kyrgyzstan</collectingCountry>
for many years, we never found this species. The above description is based on specimens from European
<collectingCountry id="F323FBE7FF932016C894FE96ABE1FE64" box="[151,229,367,393]" name="Russia" pageId="8" pageNumber="97">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

219
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<paragraph id="8B8BBB77FF9D2018C894FEDFA921FEAD" blockId="6.[151,549,294,357]" box="[151,549,294,321]" pageId="6" pageNumber="95">
<heading id="D0C30C1BFF9D2018C894FEDFA921FEAD" bold="true" box="[151,549,294,321]" fontSize="11" level="1" pageId="6" pageNumber="95" reason="1">
<taxonomicName id="4C34C0F4FF9D2018C894FEDFA921FEAD" ID-CoL="3RGT3" authority="Dlabola, 1963" authorityName="Dlabola" authorityYear="1963" box="[151,549,294,321]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillatus">
<emphasis id="B9406765FF9D2018C894FEDFA921FEAD" bold="true" box="[151,549,294,321]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C894FEDFAA77FEAC" bold="true" box="[151,371,294,321]" italics="true" pageId="6" pageNumber="95">Kybos auricillatus</emphasis>
<bibRefCitation id="EFA5C686FF9D2018C979FEDEA921FEAD" author="Dlabola, J." box="[378,549,294,321]" pageId="6" pageNumber="95" pagination="381 - 390" refId="ref6924" refString="Dlabola, J. (1963) Weitere neue Arten der Familie Cicadellidae aus Zentralasien und zoogeographische Bemerkungen zu einzelnen palaarktischen Zikadenarten (Homoptera). Acta Entomologica Musei Nationalis Pragae, 35, 381 - 390." type="journal article" year="1963">Dlabola, 1963</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C894FEB2AAE2FE88" box="[151,486,330,357]" pageId="6" pageNumber="95" type="description">
<paragraph id="8B8BBB77FF9D2018C894FEB2AAE2FE88" blockId="6.[151,549,294,357]" box="[151,486,330,357]" pageId="6" pageNumber="95">
<figureCitation id="130FA7F2FF9D2018C894FEB2ABE8FE88" box="[151,236,331,357]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="6" pageNumber="95">Figs 10</figureCitation>
,
<figureCitation id="130FA7F2FF9D2018C8FBFEB3AA45FE88" box="[248,321,330,357]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="6" pageNumber="95">34–35</figureCitation>
, 44–45, 55, 61
</paragraph>
</subSubSection>
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<paragraph id="8B8BBB77FF9D2018C894FE76A915FE4A" blockId="6.[151,724,399,454]" box="[151,529,399,423]" pageId="6" pageNumber="95">
<treatmentCitationGroup id="AB249C59FF9D2018C894FE76A915FE4A" box="[151,529,399,423]" pageId="6" pageNumber="95">
<treatmentCitation id="0A959D66FF9D2018C894FE76A915FE4A" author="Dlabola, J." box="[151,529,399,423]" page="382" pageId="6" pageNumber="95" year="1963">
<taxonomicName id="4C34C0F4FF9D2018C894FE76A915FE4A" ID-CoL="3RGT4" authority="Dlabola, 1963: 382" authorityName="Dlabola" authorityPageNumber="382" authorityYear="1963" box="[151,529,399,423]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillata">
<emphasis id="B9406765FF9D2018C894FE76AA4CFE4A" box="[151,328,399,423]" italics="true" pageId="6" pageNumber="95">Kybos auricillata</emphasis>
<bibRefCitation id="EFA5C686FF9D2018C94DFE76A915FE4A" author="Dlabola, J." box="[334,529,399,423]" pageId="6" pageNumber="95" pagination="381 - 390" refId="ref6924" refString="Dlabola, J. (1963) Weitere neue Arten der Familie Cicadellidae aus Zentralasien und zoogeographische Bemerkungen zu einzelnen palaarktischen Zikadenarten (Homoptera). Acta Entomologica Musei Nationalis Pragae, 35, 381 - 390." type="journal article" year="1963">Dlabola, 1963: 382</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8BBB77FF9D2018C894FE56A9D0FE2B" blockId="6.[151,724,399,454]" box="[151,724,430,454]" pageId="6" pageNumber="95">
<taxonomicName id="4C34C0F4FF9D2018C894FE56A9D0FE2B" ID-CoL="8J3PW" authority="Dworakowska, 1976: 427" authorityName="Dworakowska" authorityPageNumber="427" authorityYear="1976" box="[151,724,430,454]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillata" subGenus="Kybos">
<emphasis id="B9406765FF9D2018C894FE56AA05FE2B" box="[151,257,431,454]" italics="true" pageId="6" pageNumber="95">Empoasca</emphasis>
(
<emphasis id="B9406765FF9D2018C90CFE57AA4AFE2B" box="[271,334,430,454]" italics="true" pageId="6" pageNumber="95">Kybos</emphasis>
)
<emphasis id="B9406765FF9D2018C95DFE57AACDFE2B" box="[350,457,430,454]" italics="true" pageId="6" pageNumber="95">auricillata</emphasis>
Dworakowska, 1976: 427
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C894FE0DA98EFCE7" pageId="6" pageNumber="95" type="description">
<paragraph id="8B8BBB77FF9D2018C894FE0DA854FDDF" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C894FE0DAA2DFDE3" bold="true" box="[151,297,500,526]" pageId="6" pageNumber="95">Description.</emphasis>
Green with lighter and darker spots in fore part of body. Forewings smoky hyaline in apical parts, veins of hindwings and dorsal side of abdomen dark (
<figureCitation id="130FA7F2FF9D2018CAECFDE1A847FDDF" box="[751,835,536,562]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="6" pageNumber="95">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8B8BBB77FF9D2018C8C4FDC5A98EFCE7" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
Dorsal apodemes very short, look like wide transverse sclerotized band at base of abdomen (
<figureCitation id="130FA7F2FF9D2018CCE1FDC5AE3CFDBB" box="[1250,1336,572,598]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="6" pageNumber="95">Fig. 34</figureCitation>
). Ventral apodemes about 1.6 times as long as wide at base, parallel-sided, touching with inner margins (
<figureCitation id="130FA7F2FF9D2018CCC4FD99AE1FFD96" box="[1223,1307,608,635]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="6" pageNumber="95">Fig. 35</figureCitation>
). Aedeagus without lateral processes, relatively narrow, with narrowly rounded apex in lateral view and with collar-like expansion at base, distinctly visible in back view (Figs 44–45). Style with numerous long setae and serrated falcate apical part, as in other members of genus. Pygofer appendages narrow (Fig. 55). Anal collar appendages wide, with sharply narrowed, extended apices (Fig. 61).
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C8C4FCEDA9C6FBEB" pageId="6" pageNumber="95" type="biology_ecology">
<paragraph id="8B8BBB77FF9D2018C8C4FCEDA9C9FCC3" blockId="6.[151,1437,500,1463]" box="[199,717,788,814]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C8C4FCEDAA02FCC3" bold="true" box="[199,262,788,814]" pageId="6" pageNumber="95">
<collectingRegion id="49F07595FF9D2018C8C4FCEDAA05FCC3" box="[199,257,788,814]" country="Afghanistan" name="Khost" pageId="6" pageNumber="95">Host</collectingRegion>
.
</emphasis>
<taxonomicName id="4C34C0F4FF9D2018C90EFCEDAAE8FCC3" box="[269,492,788,814]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="species" species="tianschanica">
<emphasis id="B9406765FF9D2018C90EFCEDAAE8FCC3" box="[269,492,788,814]" italics="true" pageId="6" pageNumber="95">Betula tianschanica</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C34C0F4FF9D2018CA10FCECA9B9FCC3" box="[531,701,788,814]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="species" species="turkestanica">
<emphasis id="B9406765FF9D2018CA10FCECA9B9FCC3" box="[531,701,788,814]" italics="true" pageId="6" pageNumber="95">B. turkestanica</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8BBB77FF9D2018C8C4FCC1AF42FC9B" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C8C4FCC1AA76FCBF" bold="true" box="[199,370,824,850]" pageId="6" pageNumber="95">Calling signal.</emphasis>
<materialsCitation id="3B5CB12AFF9D2018C97BFCC1AF46FC9B" collectingDate="2008-07-26" county="West Tien Shan" elevation="1880" location="Signals" municipality="Chatkal Range" pageId="6" pageNumber="95" specimenCount="1" specimenCount-male="1">
<location id="8EEBEDACFF9D2018C97BFCC1AACEFCBF" LSID="urn:lsid:plazi:treatment:039D0A61FF9D2018C894FEDFA81DFA5B:8EEBEDACFF9D2018C97BFCC1AACEFCBF" box="[376,458,824,850]" county="West Tien Shan" municipality="Chatkal Range" name="Signals" pageId="6" pageNumber="95">Signals</location>
of
<specimenCount id="9D3270FEFF9D2018C9EDFCC0A956FCBF" box="[494,594,824,851]" count="1" pageId="6" pageNumber="95" type="male">one male</specimenCount>
from
<location id="8EEBEDACFF9D2018CA96FCC0A818FCBE" LSID="urn:lsid:plazi:treatment:039D0A61FF9D2018C894FEDFA81DFA5B:8EEBEDACFF9D2018CA96FCC0A818FCBE" box="[661,796,825,851]" county="West Tien Shan" municipality="Chatkal Range" name="Kyrgyzsyan" pageId="6" pageNumber="95">Kyrgyzsyan</location>
(the shore of the
<location id="8EEBEDACFF9D2018CBDFFCC1AFADFCBF" LSID="urn:lsid:plazi:treatment:039D0A61FF9D2018C894FEDFA81DFA5B:8EEBEDACFF9D2018CBDFFCC1AFADFCBF" box="[988,1193,824,850]" county="West Tien Shan" municipality="Chatkal Range" name="Sary-Chelek Lake" pageId="6" pageNumber="95">Sary-Chelek Lake</location>
,
<collectingMunicipality id="6BEF210DFF9D2018CCB7FCC1AE5FFCBE" box="[1204,1371,824,851]" pageId="6" pageNumber="95">Chatkal Range</collectingMunicipality>
,
<collectingCounty id="62EAC3FBFF9D2018CD66FCC0AA08FC9B" pageId="6" pageNumber="95">West Tien Shan</collectingCounty>
,
<elevation id="00195C44FF9D2018C914FCA5AAB1FC9B" box="[279,437,860,887]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="6" pageNumber="95" unit="m" value="1880.0">
<quantity id="4CCC1692FF9D2018C914FCA5AA68FC9A" box="[279,364,860,887]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="6" pageNumber="95" unit="m" value="1880.0">1880 m</quantity>
a. s. l.
</elevation>
,
<collectedFrom id="3828F88AFF9D2018C9C1FCA4A9C0FC9B" box="[450,708,860,887]" pageId="6" pageNumber="95">
on
<taxonomicName id="4C34C0F4FF9D2018C9E6FCA5A9C0FC9B" box="[485,708,860,886]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="species" species="tianschanica">
<emphasis id="B9406765FF9D2018C9E6FCA5A9C0FC9B" box="[485,708,860,886]" italics="true" pageId="6" pageNumber="95">Betula tianschanica</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="FF8A9DB7FF9D2018CAD0FCA5A86FFC9B" box="[723,875,860,887]" pageId="6" pageNumber="95" value="2008-07-26">
<collectingDate id="EFCE645FFF9D2018CAD0FCA5A86FFC9B" box="[723,875,860,887]" pageId="6" pageNumber="95" value="2008-07-26">26. VII. 2008</collectingDate>
</date>
) were investigated
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BBB77FF9D2018C8C4FC79A9C6FBEB" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">Calling signal distinctly differs from signals of three previous species and is a phrase lasting for about 4 s and consisting of two parts with different waveshape. The first part is a succession of partially merging pulses, in the second part the pulses are completely indistinguishable. Unfortunately, due to the low quality of the recordings, we do not have oscillograms suitable for publication.</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C8C4FBE9A900FB7B" pageId="6" pageNumber="95" type="distribution">
<paragraph id="8B8BBB77FF9D2018C8C4FBE9A900FB7B" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C8C4FBE9AA65FBC7" bold="true" box="[199,353,1040,1066]" pageId="6" pageNumber="95">Distribution.</emphasis>
Apparently, West Tien Shan within
<collectingCountry id="F323FBE7FF9D2018CAF5FBE8A873FBC6" box="[758,887,1041,1067]" name="Kyrgyzstan" pageId="6" pageNumber="95">Kyrgyzstan</collectingCountry>
and
<collectingCountry id="F323FBE7FF9D2018CBADFBE9AF34FBC7" box="[942,1072,1040,1066]" name="Uzbekistan" pageId="6" pageNumber="95">Uzbekistan</collectingCountry>
. Was described based on a small series of specimens from the environs of Arslanbob (= Arstanbap), Fergana Range, West Tien Shan,
<quantity id="4CCC1692FF9D2018CD24FBCDAE7BFBA2" box="[1319,1407,1076,1103]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="95" unit="m" value="2000.0">2000 m</quantity>
a. s. l. (
<bibRefCitation id="EFA5C686FF9D2018C8D1FBA1AA73FB9F" author="Dlabola, J." box="[210,375,1112,1138]" pageId="6" pageNumber="95" pagination="381 - 390" refId="ref6924" refString="Dlabola, J. (1963) Weitere neue Arten der Familie Cicadellidae aus Zentralasien und zoogeographische Bemerkungen zu einzelnen palaarktischen Zikadenarten (Homoptera). Acta Entomologica Musei Nationalis Pragae, 35, 381 - 390." type="journal article" year="1963">Dlabola, 1963</bibRefCitation>
;
<bibRefCitation id="EFA5C686FF9D2018C986FBA1A94EFB9F" author="Dubovskiy, G. K." box="[389,586,1112,1138]" pageId="6" pageNumber="95" refId="ref7176" refString="Dubovskiy, G. K. (1966) Cicadinea (Auchenorrhyncha) of Fergana Valley. &quot; Fan &quot; Publ., Tashkent, 256 pp. [in Russian]" type="book" year="1966">Dubovskiy, 1966</bibRefCitation>
). Later, we found it at the same altitudes on Chatkal Range about
<quantity id="4CCC1692FF9D2018CD46FBA1AE98FB9F" box="[1349,1436,1112,1138]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="95" unit="km" value="100.0">100 km</quantity>
northwest from the
<typeStatus id="548F05D5FF9D2018C970FB84AAA7FB7A" box="[371,419,1149,1175]" pageId="6" pageNumber="95">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9D2018C8C4FB59A81DFA5B" pageId="6" pageNumber="95" type="discussion">
<paragraph id="8B8BBB77FF9D2018C8C4FB59AAFAFB32" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C8C4FB59AA3FFB57" bold="true" box="[199,315,1184,1210]" pageId="6" pageNumber="95">Remarks.</emphasis>
Despite collecting leafhoppers on birches in different regions of
<collectingCountry id="F323FBE7FF9D2018CC31FB58AFB7FB56" box="[1074,1203,1185,1211]" name="Kyrgyzstan" pageId="6" pageNumber="95">Kyrgyzstan</collectingCountry>
for many years, we found
<taxonomicName id="4C34C0F4FF9D2018C8E3FB3CAA85FB33" authorityName="Dlabola" authorityYear="1963" box="[224,385,1220,1246]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillatus">
<emphasis id="B9406765FF9D2018C8E3FB3CAA85FB33" box="[224,385,1220,1246]" italics="true" pageId="6" pageNumber="95">K. auricillatus</emphasis>
</taxonomicName>
only once.
</paragraph>
<paragraph id="8B8BBB77FF9D2018C8C4FB11A8A5FAA7" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
Collar-like expansion at the base of the aedeagus stem gives the appearance of a tooth-like appendage in lateral view (Fig. 44). Apparently, for this reason, in the original description (
<bibRefCitation id="EFA5C686FF9D2018CBC9FAF5AFA8FACB" author="Dlabola, J." box="[970,1196,1292,1318]" pageId="6" pageNumber="95" pagination="381 - 390" refId="ref6924" refString="Dlabola, J. (1963) Weitere neue Arten der Familie Cicadellidae aus Zentralasien und zoogeographische Bemerkungen zu einzelnen palaarktischen Zikadenarten (Homoptera). Acta Entomologica Musei Nationalis Pragae, 35, 381 - 390." type="journal article" year="1963">Dlabola, 1963: 383</bibRefCitation>
, fig. 5) two partially separated upward-pointing processes at the aedeagus base are shown.
</paragraph>
<paragraph id="8B8BBB77FF9D2018C8C4FAACA81DFA5B" blockId="6.[151,1437,500,1463]" pageId="6" pageNumber="95">
In
<bibRefCitation id="EFA5C686FF9D2018C8E5FAADAA97FA83" author="Dlabola, J." box="[230,403,1364,1390]" pageId="6" pageNumber="95" pagination="381 - 390" refId="ref6924" refString="Dlabola, J. (1963) Weitere neue Arten der Familie Cicadellidae aus Zentralasien und zoogeographische Bemerkungen zu einzelnen palaarktischen Zikadenarten (Homoptera). Acta Entomologica Musei Nationalis Pragae, 35, 381 - 390." type="journal article" year="1963">Dlabola (1963)</bibRefCitation>
,
<bibRefCitation id="EFA5C686FF9D2018C99CFAADA96BFA83" author="Dubovskiy, G. K." box="[415,623,1364,1390]" pageId="6" pageNumber="95" refId="ref7176" refString="Dubovskiy, G. K. (1966) Cicadinea (Auchenorrhyncha) of Fergana Valley. &quot; Fan &quot; Publ., Tashkent, 256 pp. [in Russian]" type="book" year="1966">Dubovskiy (1966)</bibRefCitation>
, and Dworakowska (1976) this species is mentioned as
<taxonomicName id="4C34C0F4FF9D2018CCE5FAACAE79FA83" authorityName="Dworakowska" authorityYear="1976" box="[1254,1405,1364,1390]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillata">
<emphasis id="B9406765FF9D2018CCE5FAACAE79FA83" box="[1254,1405,1364,1390]" italics="true" pageId="6" pageNumber="95">K. auricillata</emphasis>
</taxonomicName>
or
<taxonomicName id="4C34C0F4FF9D2018C894FA80AA5AFA7F" authorityName="Dworakowska" authorityYear="1976" box="[151,350,1400,1426]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillata" subGenus="Kybos">
<emphasis id="B9406765FF9D2018C894FA80ABB4FA7F" box="[151,176,1401,1426]" italics="true" pageId="6" pageNumber="95">E.</emphasis>
(
<emphasis id="B9406765FF9D2018C8BCFA80ABDDFA7F" box="[191,217,1401,1426]" italics="true" pageId="6" pageNumber="95">K.</emphasis>
)
<emphasis id="B9406765FF9D2018C8EBFA81AA5AFA7F" box="[232,350,1400,1426]" italics="true" pageId="6" pageNumber="95">auricillata</emphasis>
</taxonomicName>
. However, the grammatical gender of the generic name is masculine, so the species name must be changed to
<taxonomicName id="4C34C0F4FF9D2018C91BFA65AAA2FA5B" authorityName="Dlabola" authorityYear="1963" box="[280,422,1436,1462]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="auricillatus">
“
<emphasis id="B9406765FF9D2018C926FA65AAA2FA5B" box="[293,422,1436,1462]" italics="true" pageId="6" pageNumber="95">auricillatus</emphasis>
</taxonomicName>
” (
<bibRefCitation id="EFA5C686FF9D2018C9C2FA65A9A9FA5B" author="Dmitriev, D. A. &amp; Anufriev, G. A. &amp; Bartlett, C. R. &amp; Blanco-Rodriguez, E. &amp; Borodin, O. I. &amp; Cao, Y. - H. &amp; Deitz, L. L. &amp; Dietrich, C. H. &amp; Dmitrieva, M. O. &amp; El-Sonbati, S. A. &amp; Evangelista de Souza, O. &amp; Gjonov, I. V. &amp; Goncalves, A. C. &amp; Hendrix, S. &amp; McKamey, S. &amp; Kohler, M. &amp; Kunz, G. &amp; Malenovsky, I. &amp; Morris, B. O. &amp; Novoselova, M. &amp; Pinedo-Escatel, J. A. &amp; Rakitov, R. A. &amp; Rothschild, M. J. &amp; Sanborn, A. F. &amp; Takiya, D. M. &amp; Wallace, M. S. &amp; Zahniser, J. N." box="[449,685,1436,1463]" pageId="6" pageNumber="95" refId="ref6962" refString="Dmitriev, D. A., Anufriev, G. A., Bartlett, C. R., Blanco-Rodriguez, E., Borodin, O. I., Cao, Y. - H., Deitz, L. L., Dietrich, C. H., Dmitrieva, M. O., El-Sonbati, S. A., Evangelista de Souza, O., Gjonov, I. V., Goncalves, A. C., Hendrix, S., McKamey, S., Kohler, M., Kunz, G., Malenovsky, I., Morris, B. O., Novoselova, M., Pinedo-Escatel, J. A., Rakitov, R. A., Rothschild, M. J., Sanborn, A. F., Takiya, D. M., Wallace, M. S. &amp; Zahniser, J. N. (2022 onward). Kybos Fieber, 1866. World Auchenorrhyncha Database. TaxonPages. Available from: https: // hoppers. speciesfile. org / otus / 43322 / overview (accessed 9 March 2024)" type="journal volume" year="2022">
Dmitriev
<emphasis id="B9406765FF9D2018CA2CFA64A963FA5B" box="[559,615,1436,1462]" italics="true" pageId="6" pageNumber="95">et al.</emphasis>
, 2022
</bibRefCitation>
onward).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="A81DB077AF02DD1C41404CF8C0B6B7FC">Leafhoppers of the genus Kybos Fieber, 1866 (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) in Kyrgyzstan</mods:title>
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<subSubSection id="C32EE8FCFF9F201AC894FAEFA945FADD" box="[151,577,1302,1329]" pageId="4" pageNumber="93" type="nomenclature">
<paragraph id="8B8BBB77FF9F201AC894FAEFA945FADD" blockId="4.[151,577,1302,1365]" box="[151,577,1302,1329]" pageId="4" pageNumber="93">
<heading id="D0C30C1BFF9F201AC894FAEFA945FADD" bold="true" box="[151,577,1302,1329]" fontSize="11" level="1" pageId="4" pageNumber="93" reason="1">
<taxonomicName id="4C34C0F4FF9F201AC894FAEFA945FADD" ID-CoL="9G635" authority="Zachvatkin, 1953" authorityName="Zachvatkin" authorityYear="1953" box="[151,577,1302,1329]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9F201AC894FAEFA945FADD" bold="true" box="[151,577,1302,1329]" pageId="4" pageNumber="93">
<emphasis id="B9406765FF9F201AC894FAEFAA65FADC" bold="true" box="[151,353,1302,1329]" italics="true" pageId="4" pageNumber="93">Kybos niveicolor</emphasis>
Zachvatkin, 1953
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9F201AC894FAC2A929FAB8" box="[151,557,1338,1365]" pageId="4" pageNumber="93" type="description">
<paragraph id="8B8BBB77FF9F201AC894FAC2A929FAB8" blockId="4.[151,577,1302,1365]" box="[151,557,1338,1365]" pageId="4" pageNumber="93">
<figureCitation id="130FA7F2FF9F201AC894FAC2ABDAFAB8" box="[151,222,1339,1365]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="4" pageNumber="93">Figs 9</figureCitation>
,
<figureCitation id="130FA7F2FF9F201AC8E9FAC2AA0CFAB8" box="[234,264,1339,1365]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="4" pageNumber="93">33</figureCitation>
, 42–43, 51, 54, 60, 74–76
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9F201AC894FA86A839FA5B" pageId="4" pageNumber="93" type="reference_group">
<paragraph id="8B8BBB77FF9F201AC894FA86A92BFA7A" blockId="4.[151,829,1407,1462]" box="[151,559,1407,1431]" pageId="4" pageNumber="93">
<taxonomicName id="4C34C0F4FF9F201AC894FA86A92BFA7A" ID-CoL="9G635" authority="Zachvatkin, 1953: 239" authorityName="Zachvatkin" authorityPageNumber="239" authorityYear="1953" box="[151,559,1407,1431]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9F201AC894FA86AA40FA7A" box="[151,324,1407,1431]" italics="true" pageId="4" pageNumber="93">Kybos niveicolor</emphasis>
Zachvatkin, 1953: 239
</taxonomicName>
</paragraph>
<paragraph id="8B8BBB77FF9F201AC894FA66A839FA5B" blockId="4.[151,829,1407,1462]" box="[151,829,1438,1462]" pageId="4" pageNumber="93">
<taxonomicName id="4C34C0F4FF9F201AC894FA66A839FA5B" ID-CoL="9G7GP" authority="Dworakowska, 1976: 440" authorityName="Dworakowska" authorityPageNumber="440" authorityYear="1976" box="[151,829,1438,1462]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="niveicolor" subGenus="Kybos" subSpecies="niveicolor">
<emphasis id="B9406765FF9F201AC894FA66AA05FA5B" box="[151,257,1439,1462]" italics="true" pageId="4" pageNumber="93">Empoasca</emphasis>
(
<emphasis id="B9406765FF9F201AC90CFA67AA4AFA5B" box="[271,334,1438,1462]" italics="true" pageId="4" pageNumber="93">Kybos</emphasis>
)
<emphasis id="B9406765FF9F201AC95DFA67A936FA5B" box="[350,562,1438,1462]" italics="true" pageId="4" pageNumber="93">niveicolor niveicolor</emphasis>
Dworakowska, 1976: 440
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9F201AC894FA1DAF0EF95F" pageId="4" pageNumber="93" type="description">
<paragraph id="8B8BBB77FF9F201AC894FA1DAA35F9CF" blockId="4.[151,1437,1508,2039]" pageId="4" pageNumber="93">
<emphasis id="B9406765FF9F201AC894FA1DAA2DFA13" bold="true" box="[151,297,1508,1534]" pageId="4" pageNumber="93">Description.</emphasis>
Live specimens white or greenish white, dry ones yellowish white, sometimes with slight greenish tinge (
<figureCitation id="130FA7F2FF9F201AC8DDF9F1AA27F9CF" box="[222,291,1544,1570]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." pageId="4" pageNumber="93">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B8BBB77FF9F201AC8C4F9D5AF0EF95F" blockId="4.[151,1437,1508,2039]" pageId="4" pageNumber="93">
Dorsal apodemes short, ventral apodemes about 1.6–1.7 times as long as wide at base, narrow, parallel-sided, separated by narrow notch (
<figureCitation id="130FA7F2FF9F201AC9D1F9A9A922F987" box="[466,550,1616,1642]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." pageId="4" pageNumber="93">Fig. 33</figureCitation>
). Genitalia almost same as in
<taxonomicName id="4C34C0F4FF9F201ACB78F9A8AF0DF987" authorityName="Mitjaev" authorityYear="1971" box="[891,1033,1617,1642]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9F201ACB78F9A8AF0DF987" box="[891,1033,1617,1642]" italics="true" pageId="4" pageNumber="93">K. montanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34C0F4FF9F201ACC42F9A8AFC7F987" authorityName="Zachvatkin" authorityYear="1953" box="[1089,1219,1616,1642]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
<emphasis id="B9406765FF9F201ACC42F9A8AFC7F987" box="[1089,1219,1616,1642]" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
</taxonomicName>
. Aedeagus without lateral processes (Figs 42–43). Style with numerous long setae and serrated falcate apical part (Fig. 51). Pygofer appendages narrow (Fig. 54). Anal collar appendages narrow, falcate (Fig. 60).
</paragraph>
</subSubSection>
<subSubSection id="C32EE8FCFF9F201AC8C4F945A8BBF81A" pageId="4" pageNumber="93" type="biology_ecology">
<paragraph id="8B8BBB77FF9F201AC8C4F945A894F917" blockId="4.[151,1437,1508,2039]" pageId="4" pageNumber="93">
<emphasis id="B9406765FF9F201AC8C4F945AA15F93B" bold="true" box="[199,273,1724,1750]" pageId="4" pageNumber="93">Hosts.</emphasis>
<taxonomicName id="4C34C0F4FF9F201AC918F945AAB6F93B" baseAuthorityName="Skvortsov" baseAuthorityYear="1972" box="[283,434,1724,1750]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="B9406765FF9F201AC918F945AAB6F93B" box="[283,434,1724,1750]" italics="true" pageId="4" pageNumber="93">Populus alba</emphasis>
</taxonomicName>
.
<taxonomicName id="4C34C0F4FF9F201AC9C0F944A91AF93B" box="[451,542,1725,1750]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="nivea">
<emphasis id="B9406765FF9F201AC9C0F944A91AF93B" box="[451,542,1725,1750]" italics="true" pageId="4" pageNumber="93">P. nivea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34C0F4FF9F201ACA59F944A9DEF93B" box="[602,730,1724,1750]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="bolleana">
<emphasis id="B9406765FF9F201ACA59F944A9DEF93B" box="[602,730,1724,1750]" italics="true" pageId="4" pageNumber="93">P. bolleana</emphasis>
</taxonomicName>
, listed as food plants in the original description (
<bibRefCitation id="EFA5C686FF9F201ACD16F945ABE5F916" author="Zachvatkin, A. A." pageId="4" pageNumber="93" pagination="237 - 245" refId="ref7692" refString="Zachvatkin, A. A. (1953 b) On the fauna of Eupterygidae (Homoptera, Cicadina) of Central Asia. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 237 - 245. [in Russian]" type="book chapter" year="1953">Zachvatkin, 1953b</bibRefCitation>
), are now considered synonyms of
<taxonomicName id="4C34C0F4FF9F201ACA6FF918A888F917" authority="(Skvortsov, 1972)" baseAuthorityName="Skvortsov" baseAuthorityYear="1972" box="[620,908,1760,1786]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="B9406765FF9F201ACA6FF918A9BEF917" box="[620,698,1760,1786]" italics="true" pageId="4" pageNumber="93">P. alba</emphasis>
(
<bibRefCitation id="EFA5C686FF9F201ACACBF919A887F917" author="Skvortsov, A. K." box="[712,899,1760,1786]" pageId="4" pageNumber="93" pagination="6 - 25" refId="ref7467" refString="Skvortsov, A. K. (1972) Family 29. Salicaceae. Keys to Plants of Central Asia. Revised Synopsis of Flora of Central Asia, 3, 6 - 25. [&quot; Fan &quot; Publ., Tashkent; in Russian]" type="journal article" year="1972">Skvortsov, 1972</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8BBB77FF9F201AC8C4F8FDA826F8AF" blockId="4.[151,1437,1508,2039]" pageId="4" pageNumber="93">
<emphasis id="B9406765FF9F201AC8C4F8FDAA70F8F3" bold="true" box="[199,372,1796,1822]" pageId="4" pageNumber="93">Calling signal.</emphasis>
<materialsCitation id="3B5CB12AFF9F201AC97FF8FDA81AF8AF" collectingDate="2011-07-16" country="Kyrgyzstan" county="West Tien Shan" location="Arkyt Village" municipality="Chatkal Range" pageId="4" pageNumber="93" specimenCount="5" specimenCount-male="5">
Signals of
<specimenCount id="9D3270FEFF9F201AC9F5F8FDA96FF8F3" box="[502,619,1796,1822]" count="5" pageId="4" pageNumber="93" type="male">five males</specimenCount>
from
<collectingCountry id="F323FBE7FF9F201ACAB1F8FCA837F8F2" box="[690,819,1797,1823]" name="Kyrgyzstan" pageId="4" pageNumber="93">Kyrgyzstan</collectingCountry>
(
<location id="8EEBEDACFF9F201ACB41F8FDA8E5F8F3" LSID="urn:lsid:plazi:treatment:039D0A61FF9F2018C894FAEFA9CFFF38:8EEBEDACFF9F201ACB41F8FDA8E5F8F3" box="[834,993,1796,1822]" country="Kyrgyzstan" county="West Tien Shan" municipality="Chatkal Range" name="Arkyt Village" pageId="4" pageNumber="93">Arkyt Village</location>
,
<collectingMunicipality id="6BEF210DFF9F201ACBEEF8FDAF92F8F2" box="[1005,1174,1796,1823]" pageId="4" pageNumber="93">Chatkal Range</collectingMunicipality>
,
<collectingCounty id="62EAC3FBFF9F201ACCA0F8FCAE53F8F3" box="[1187,1367,1796,1823]" pageId="4" pageNumber="93">West Tien Shan</collectingCounty>
,
<collectedFrom id="3828F88AFF9F201ACD60F8FCABCDF8AF" pageId="4" pageNumber="93">
on
<taxonomicName id="4C34C0F4FF9F201ACD8BF8FCABCDF8AF" baseAuthorityName="Skvortsov" baseAuthorityYear="1972" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="B9406765FF9F201ACD8BF8FCABCDF8AF" italics="true" pageId="4" pageNumber="93">P. alba</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="FF8A9DB7FF9F201AC8D4F8D1AA6BF8AF" box="[215,367,1832,1859]" pageId="4" pageNumber="93" value="2011-07-16">
<collectingDate id="EFCE645FFF9F201AC8D4F8D1AA6BF8AF" box="[215,367,1832,1859]" pageId="4" pageNumber="93" value="2011-07-16">16. VII. 2011</collectingDate>
</date>
, recording at 22
<superScript id="7C41163FFF9F201ACA28F8D1A943F8AF" attach="right" box="[555,583,1832,1858]" fontSize="6" pageId="4" pageNumber="93">oC</superScript>
) were investigated
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BBB77FF9F201AC8C4F8B5A8BBF81A" blockId="4.[151,1437,1508,2039]" pageId="4" pageNumber="93">
Calling signal is generally the same as in
<taxonomicName id="4C34C0F4FF9F201ACA8CF8B4A80AF88B" authorityName="Zachvatkin" authorityYear="1953" box="[655,782,1868,1894]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
<emphasis id="B9406765FF9F201ACA8CF8B4A80AF88B" box="[655,782,1868,1894]" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34C0F4FF9F201ACB40F8B4A8CBF88B" authorityName="Mitjaev" authorityYear="1971" box="[835,975,1869,1894]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9F201ACB40F8B4A8CBF88B" box="[835,975,1869,1894]" italics="true" pageId="4" pageNumber="93">K. montanus</emphasis>
</taxonomicName>
. It consists of 3–6 high-amplitude pulses, following each other with a period of 320–520 ms; sometimes, the pulse repetition period increases towards the end of a signal (
<figureCitation id="130FA7F2FF9F201AC91DF86DAAA7F842" box="[286,419,1940,1967]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="4" pageNumber="93">Figs 74–75</figureCitation>
). Pulse duration averages 30–45 ms (
<figureCitation id="130FA7F2FF9F201ACB49F86DA89BF843" box="[842,927,1940,1966]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." pageId="4" pageNumber="93">Fig. 76</figureCitation>
). Unlike two above-mentioned species, in
<taxonomicName id="4C34C0F4FF9F201ACD80F86CAA0DF83F" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9F201ACD80F86CAA0DF83F" italics="true" pageId="4" pageNumber="93">K. niveicolor</emphasis>
</taxonomicName>
, additional low-amplitude pulses are absent between the main high-amplitude ones. As a result, the signal is not a phrase, i.e. a sequence of syllables, but a simple pulse sequence.
</paragraph>
</subSubSection>
<caption id="DF4BEBFFFF9E201BC89BF8F5A81FF835" pageId="5" pageNumber="94" startId="5.[152,270,1804,1828]" targetBox="[216,1373,187,1774]" targetPageId="5" targetType="figure">
<paragraph id="8B8BBB77FF9E201BC89BF8F5A81FF835" blockId="5.[151,1436,1804,2009]" pageId="5" pageNumber="94">
<emphasis id="B9406765FF9E201BC89BF8F5AA66F8C9" bold="true" box="[152,354,1804,1828]" pageId="5" pageNumber="94">FIGURES 13–37.</emphasis>
<taxonomicName id="4C34C0F4FF9E201BC96CF8F5AAECF8C9" box="[367,488,1804,1828]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9406765FF9E201BC96CF8F5AAAAF8C9" box="[367,430,1804,1828]" italics="true" pageId="5" pageNumber="94">Kybos</emphasis>
spp.
</taxonomicName>
, abdominal apodemes. 13–16—
<taxonomicName id="4C34C0F4FF9E201BCB40F8F4A8CFF8C9" authorityName="Mitjaev" authorityYear="1971" box="[835,971,1805,1828]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9406765FF9E201BCB40F8F4A8CFF8C9" box="[835,971,1805,1828]" italics="true" pageId="5" pageNumber="94">K. montanus</emphasis>
</taxonomicName>
(13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—
<taxonomicName id="4C34C0F4FF9E201BC971F8C8AAEAF8A5" authorityName="Zachvatkin" authorityYear="1953" box="[370,494,1840,1864]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="oshanini">
<emphasis id="B9406765FF9E201BC971F8C8AAEAF8A5" box="[370,494,1840,1864]" italics="true" pageId="5" pageNumber="94">K. oshanini</emphasis>
</taxonomicName>
(17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of
<taxonomicName id="4C34C0F4FF9E201BCC58F880AFCBF87D" authorityName="Zachvatkin" authorityYear="1953" box="[1115,1231,1912,1936]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="oshanini">
<emphasis id="B9406765FF9E201BCC58F880AFCBF87D" box="[1115,1231,1912,1936]" italics="true" pageId="5" pageNumber="94">K. oshanini</emphasis>
</taxonomicName>
in Moscow Oblast); 33—
<taxonomicName id="4C34C0F4FF9E201BC8C9F864AA4BF859" box="[202,335,1948,1972]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="niveicolor">
<emphasis id="B9406765FF9E201BC8C9F864AA4BF859" box="[202,335,1948,1972]" italics="true" pageId="5" pageNumber="94">K. niveicolor</emphasis>
</taxonomicName>
, dorsal and ventral apodemes; 34–35—
<taxonomicName id="4C34C0F4FF9E201BCAE7F864A872F859" authorityName="Dlabola" authorityYear="1963" box="[740,886,1948,1972]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="auricillatus">
<emphasis id="B9406765FF9E201BCAE7F864A872F859" box="[740,886,1948,1972]" italics="true" pageId="5" pageNumber="94">K. auricillatus</emphasis>
</taxonomicName>
(34—dorsal apodemes, 35—ventral apodemes); 36—
<taxonomicName id="4C34C0F4FF9E201BC894F838ABF2F835" baseAuthorityName="Edwards" baseAuthorityYear="1908" box="[151,246,1984,2008]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="populi">
<emphasis id="B9406765FF9E201BC894F838ABF2F835" box="[151,246,1984,2008]" italics="true" pageId="5" pageNumber="94">K. populi</emphasis>
</taxonomicName>
, dorsal and ventral apodemes; 37—
<taxonomicName id="4C34C0F4FF9E201BCA60F838A9D7F835" baseAuthorityName="Ribaut" baseAuthorityYear="1933" box="[611,723,1985,2008]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="virgator">
<emphasis id="B9406765FF9E201BCA60F838A9D7F835" box="[611,723,1985,2008]" italics="true" pageId="5" pageNumber="94">K. virgator</emphasis>
</taxonomicName>
, same.
</paragraph>
</caption>
<subSubSection id="C32EE8FCFF9D2018C8C4FF6FA9CFFF38" pageId="6" pageNumber="95" type="distribution">
<paragraph id="8B8BBB77FF9D2018C8C4FF6FA9CFFF38" blockId="6.[151,1436,150,213]" pageId="6" pageNumber="95">
<emphasis id="B9406765FF9D2018C8C4FF6FAA65FF5D" bold="true" box="[199,353,150,176]" pageId="6" pageNumber="95">Distribution.</emphasis>
Widespread in southern and southeastern
<collectingCountry id="F323FBE7FF9D2018CB46FF6EA8CFFF5C" box="[837,971,151,177]" name="Kazakhstan" pageId="6" pageNumber="95">Kazakhstan</collectingCountry>
,
<collectingCountry id="F323FBE7FF9D2018CBD4FF6EAF5EFF5C" box="[983,1114,151,177]" name="Kyrgyzstan" pageId="6" pageNumber="95">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="F323FBE7FF9D2018CC6BFF6EAFEDFF5C" box="[1128,1257,151,177]" name="Uzbekistan" pageId="6" pageNumber="95">Uzbekistan</collectingCountry>
, and
<collectingCountry id="F323FBE7FF9D2018CD25FF6EAE9CFF5C" box="[1318,1432,151,177]" name="Tajikistan" pageId="6" pageNumber="95">Tajikistan</collectingCountry>
; also, was found in Xinjian,
<collectingCountry id="F323FBE7FF9D2018C9CFFF42A90BFF38" box="[460,527,187,213]" name="China" pageId="6" pageNumber="95">China</collectingCountry>
(
<bibRefCitation id="EFA5C686FF9D2018CA1EFF42A9BBFF38" author="Mitjaev, I. D." box="[541,703,187,213]" pageId="6" pageNumber="95" pagination="1 - 170" refId="ref7271" refString="Mitjaev, I. D. (2002) Fauna, Ecology, and Zoogeography of leafhoppers (Homoptera, Cicadinea) of Kazakhstan. Tethys Entomological Research, 5, 1 - 170. [in Russian]" type="journal article" year="2002">Mitjaev, 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

565
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<paragraph id="8BB09A41D76C911D168CF9A0FC90F9DB" blockId="0.[151,800,1635,1697]" box="[151,800,1635,1662]" pageId="0" pageNumber="140">
<heading id="D0F82D2DD76C911D168CF9A0FC90F9DB" bold="true" box="[151,800,1635,1662]" fontSize="11" level="1" pageId="0" pageNumber="140" reason="1">
<emphasis id="B97B4653D76C911D168CF9A0FC90F9DB" bold="true" box="[151,800,1635,1662]" pageId="0" pageNumber="140">
<taxonomicName id="4C0FE1C2D76C911D168CF9A0FE10F9D8" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[151,416,1635,1662]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="140" phylum="Arthropoda" rank="species" species="medeirosi" status="sp. nov.">
<emphasis id="B97B4653D76C911D168CF9A0FE10F9D8" bold="true" box="[151,416,1635,1662]" italics="true" pageId="0" pageNumber="140">Dentigaster medeirosi</emphasis>
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<taxonomicNameLabel id="A248FB28D76C911D17BCF9A0FDB5F9DB" box="[423,517,1635,1661]" pageId="0" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
Dias &amp; Penteado-Dias
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<paragraph id="8BB09A41D76C911D168CF944FEF3F907" blockId="0.[151,800,1635,1697]" box="[151,323,1671,1697]" pageId="0" pageNumber="140">
(
<figureCitation id="133486C4D76C911D1684F944FEA1F907" box="[159,273,1671,1697]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–10" captionStartId-0="1.[152,270,1878,1902]" captionStartId-1="2.[152,270,920,944]" captionTargetBox-0="[151,1436,309,1855]" captionTargetBox-1="[151,1435,181,896]" captionTargetId-0="figure-97@1.[151,1436,309,1855]" captionTargetId-1="figure-17@2.[151,1435,181,896]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." captionText-1="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.13330840" figureDoi-1="http://doi.org/10.5281/zenodo.13330844" httpUri-0="https://zenodo.org/record/13330840/files/figure.png" httpUri-1="https://zenodo.org/record/13330844/files/figure.png" pageId="0" pageNumber="140">Figs 1–10</figureCitation>
,
<figureCitation id="133486C4D76C911D1705F944FE8BF907" box="[286,315,1671,1697]" captionStart="FIGURE 18" captionStartId="5.[152,255,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-25@5.[151,1436,181,1089]" captionTargetPageId="5" captionText="FIGURE 18. Map of distribution of species: z, Dentigaster medeirosi sp. nov. ▲, Dentigaster fernandesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13330848" httpUri="https://zenodo.org/record/13330848/files/figure.png" pageId="0" pageNumber="140">18</figureCitation>
)
</paragraph>
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<typeStatus id="54B424E3D76C911D168CF90CFEB4F94F" box="[151,260,1743,1769]" pageId="0" pageNumber="140" type="holotype">Holotype</typeStatus>
: body length,
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</paragraph>
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Color. Scapus and pedicellus orange; first flagellomere and basal half of second, light yellow; remaining flagellomeres dark brown. Head black, with a triangular anterior orange patch below the base of the antennae and extended downwards. Propleuron light orange. Pronotum black, with anterior and lower parts light yellow. Scutellum black from the anterior area to the scutellar sulcus and orange on the disc and remaining areas. Other parts of the mesosoma and legs orange; coxa and trochanter of fore and mid legs orange yellow. Tegula dark brown and humeral plate light yellow. Fore wing (
<figureCitation id="133486C4D76C911D146DF864FD09F867" box="[630,697,1959,1985]" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="0" pageNumber="140">Fig. 7</figureCitation>
) orange in the basal third, brown transverse band crossing the first discal and first subdiscal cells, transverse white band at the parastigma and basal part of pterostigma; wide brown transverse band with a white bulla in r-m; whitish subapical area and border apical light brown. Dark brown veins in the brown bands and orange veins in the other parts of the fore wing. Hind wing hyaline, with orange veins, in basal half and light brown, with brown veins, in the apical half. Metasoma: first tergite orange; second tergite black, with a large anterior light yellow band; third tergite black (
<figureCitation id="133486C4D76D911C14F4FEC0FCE5FEBB" box="[751,853,259,285]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="1" pageNumber="141">Figs 1, 2</figureCitation>
).
</paragraph>
<caption id="DF70CAC9D76D911C1683F895FCF1F835" ID-DOI="http://doi.org/10.5281/zenodo.13330840" ID-Zenodo-Dep="13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="1" pageNumber="141" startId="1.[152,270,1878,1902]" targetBox="[151,1436,309,1855]" targetPageId="1" targetType="figure">
<paragraph id="8BB09A41D76D911C1683F895FCF1F835" blockId="1.[151,1436,1878,1939]" pageId="1" pageNumber="141">
<emphasis id="B97B4653D76D911C1683F895FEF1F8C8" bold="true" box="[152,321,1878,1902]" pageId="1" pageNumber="141">FIGURES 1–6.</emphasis>
<taxonomicName id="4C0FE1C2D76D911C1753F894FD99F8C9" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[328,553,1879,1903]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="141" phylum="Arthropoda" rank="species" species="medeirosi" status="sp. nov.">
<emphasis id="B97B4653D76D911C1753F894FD99F8C9" box="[328,553,1879,1903]" italics="true" pageId="1" pageNumber="141">Dentigaster medeirosi</emphasis>
</taxonomicName>
<emphasis id="B97B4653D76D911C142BF894FD31F8C9" bold="true" box="[560,641,1879,1903]" pageId="1" pageNumber="141">
<taxonomicNameLabel id="A248FB28D76D911C142BF894FD31F8C9" box="[560,641,1879,1903]" pageId="1" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal.
</paragraph>
</caption>
<caption id="DF70CAC9D76E911F1683FC5BFD98FC72" ID-DOI="http://doi.org/10.5281/zenodo.13330844" ID-Zenodo-Dep="13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="2" pageNumber="142" startId="2.[152,270,920,944]" targetBox="[151,1435,181,896]" targetPageId="2" targetType="figure">
<paragraph id="8BB09A41D76E911F1683FC5BFD98FC72" blockId="2.[151,1436,920,980]" pageId="2" pageNumber="142">
<emphasis id="B97B4653D76E911F1683FC5BFEE1FC16" bold="true" box="[152,337,920,944]" pageId="2" pageNumber="142">FIGURES 7–10.</emphasis>
<taxonomicName id="4C0FE1C2D76E911F1741FC5AFD8DFC16" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[346,573,920,944]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="medeirosi" status="sp. nov.">
<emphasis id="B97B4653D76E911F1741FC5AFD8DFC16" box="[346,573,920,944]" italics="true" pageId="2" pageNumber="142">Dentigaster medeirosi</emphasis>
</taxonomicName>
<emphasis id="B97B4653D76E911F145CFC5BFD2AFC16" bold="true" box="[583,666,920,944]" pageId="2" pageNumber="142">
<taxonomicNameLabel id="A248FB28D76E911F145CFC5BFD2AFC16" box="[583,666,920,944]" pageId="2" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal.
</paragraph>
</caption>
<paragraph id="8BB09A41D76E911F16DCFBE5FD1BFA9A" blockId="2.[151,1437,1062,2025]" pageId="2" pageNumber="142">Head: Antenna with 46 flagellomeres. Scapus 2.43x as long as maximally wide; 1.09x longer than first flagellomere. First flagellomere 4.27x as long as wide, 1.34x as long as the second one. Penultimate flagellomere 1.5x as long as wide, 0.06x as long as first flagellomere, 0.5x as long as apical flagellomere. Head 1.17x as wide as its median length. Eye with with the same length of temple (in dorsal view) and 2.44x as high as wide. Head coarsely punctate. Clypeus moderately swollen in lateral view, with two blunt teeth in lower margin; its surface moderately punctate. Malar space 0.81x basal width of mandible, 0.32x eye height. Face moderately convex, with slightly protruding median carina. Temple not swollen in dorsal view. Frons flat, not concave. POL 1.25x ocellar diameter, 0.33x OOL. Occipital carina present, complete.</paragraph>
<paragraph id="8BB09A41D76E911F16DCFA84FAE9F903" blockId="2.[151,1437,1062,2025]" pageId="2" pageNumber="142">
Mesosoma: Mesosoma 1.95x as long as wide in lateral view, 1.92x as long as wide in dorsal view. Pronotum coarsely punctate. Mesoscutum anteriorly prominent and rounded, coarsely punctate. Notauli slightly apparent. Scutellar sulcus with ten transversal complete carinae. Mesopleuron, metapleuron and propodeum coarsely punctate; propodeum with small protrusion on each side and medium longitudinal carina, slightly protruding. Fore wing (
<figureCitation id="133486C4D76E911F1374FA71FF16FA56" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="2" pageNumber="142">Fig. 7</figureCitation>
) with stigma 5.0x as long as wide. Marginal cell distally closed. Vein r 0.34x as long as vein 3-SR, 0.10x as long as vein SR 1, 0.19x as long as vein 2-SR. Vein 1-SR+M present. First discal cell 3.25x as long as wide. Vein r-m present, with a bulla. Vein m-cu postfurcal. Vein cu-a present, postfurcal. Vein Cu1b present. First subdiscal cell distally closed. Vein 2-1A long. Hind wing: Vein 1-SC+R present. Subbasal cell medium sized. Vein M+CU 0.78x as long as vein 1-M. Legs: Hind coxa large. Hind femur 3.13x as long as wide. Hind tibia 7.22x as long as wide, 2.03x as long as the hind basitarsomere. Hind basitarsomere 8.0x as long as wide, 1.01x as long as tarsomeres 2–5.
</paragraph>
<paragraph id="8BB09A41D76E911F16DCF96CFF05F806" blockId="2.[151,1437,1062,2025]" pageId="2" pageNumber="142">
Metasoma: Carapace with two transversal areolate sutures (
<figureCitation id="133486C4D76E911F1575F96DFC04F96E" box="[878,948,1710,1736]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="2" pageNumber="142">Fig. 2</figureCitation>
); the one between first and second tergites wider, mainly in the middle area of the tergite. First and second tergites rugose-areolate, with irregular longitudinal striae, poorly defined. Third tergite finely rugose-areolate (
<figureCitation id="133486C4D76E911F1521F935FC3FF8B6" box="[826,911,1782,1808]" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="2" pageNumber="142">Fig. 10</figureCitation>
). First tergite with anterior prominent median knob and 1.35x as long as apical width, in dorsal view. Median length of second tergite 1.10x its basal width, 0.81x as long as first tergite. Carapace 3.32x as long as wide, in dorsal view. Third tergite with apex not denticulated in dorsal view. Postero-ventral rim, on third tergite in lateral view, angular, with posterior half directed upward (
<figureCitation id="133486C4D76E911F1374F8A1FF17F806" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="2" pageNumber="142">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BB09A41D76E911F16DCF869FECEF862" blockId="2.[151,1437,1062,2025]" box="[199,382,1962,1988]" pageId="2" pageNumber="142">Male: Unknown</paragraph>
<paragraph id="8BB09A41D76E911E16DCF80DFF05FEC3" blockId="2.[151,1437,1062,2025]" lastBlockId="3.[151,1437,151,1077]" lastPageId="3" lastPageNumber="143" pageId="2" pageNumber="142">
Variations: Head of
<taxonomicName id="4C0FE1C2D76E911F17B1F80CFD8AF84E" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[426,570,1998,2024]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="medeirosi">
<emphasis id="B97B4653D76E911F17B1F80CFD8AF84E" box="[426,570,1998,2024]" italics="true" pageId="2" pageNumber="142">D. medeirosi</emphasis>
</taxonomicName>
has anterior orange patch near the base of the antennae (
<figureCitation id="133486C4D76E911F12A5F80DFAB1F84F" box="[1214,1281,1998,2025]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="2" pageNumber="142">Fig. 5</figureCitation>
) or expanded downwards in some specimens (
<figureCitation id="133486C4D76F911E1411FF54FDE1FF17" box="[522,593,151,177]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="3" pageNumber="143">Fig. 6</figureCitation>
). Pronotum is black in the superior area, adjacent to the mesoscutum and mesopleuron; light yellow in the remaining parts (
<figureCitation id="133486C4D76F911E14D3FF78FCBAFF73" box="[712,778,187,213]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="3" pageNumber="143">Fig. 1</figureCitation>
). About a third of the specimens examined have pronotum almost completely black, with median dorsal patch light yellow, which can be reduced or laterally widened (
<figureCitation id="133486C4D76F911E1354FF1CFF18FEBB" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="3" pageNumber="143">Figs 3, 4</figureCitation>
). Small brown or dark brown blackish patch on the upper part of mesopleuron (
<figureCitation id="133486C4D76F911E123BFEC0FBD3FEBB" box="[1056,1123,259,285]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="3" pageNumber="143">Fig. 3</figureCitation>
) occurs in some specimens, generally poor defined. A dark brown patch on each side of the first tergite was observed in only
<specimenCount id="9D0951C8D76F911E12D3FEE4FAD1FEE7" box="[1224,1377,295,321]" count="1" pageId="3" pageNumber="143" type="generic">one specimen</specimenCount>
(
<figureCitation id="133486C4D76F911E1374FEE4FF17FEC3" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="3" pageNumber="143">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BB09A41D76F911E16DCFEACFA3AFE2F" blockId="3.[151,1437,151,1077]" box="[199,1418,367,393]" pageId="3" pageNumber="143">
Etymology: In honour to our friend José Honorato Gago da Câmara do Botelho de Medeiros (
<emphasis id="B97B4653D76F911E12F8FEACFACAFE2E" box="[1251,1402,367,392]" italics="true" pageId="3" pageNumber="143">In memoriam</emphasis>
).
</paragraph>
<paragraph id="8BB09A41D76F911E16DCFE50FE66FE0B" blockId="3.[151,1437,151,1077]" box="[199,470,403,429]" pageId="3" pageNumber="143">Habitat: Atlantic Forest.</paragraph>
<paragraph id="8BB09A41D76F911E16DCFE74FE12FE77" blockId="3.[151,1437,151,1077]" box="[199,418,439,465]" pageId="3" pageNumber="143">Material examined:</paragraph>
<paragraph id="8BB09A41D76F911E16DCFE18FC0BFB93" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="143">
<materialsCitation id="3B67901CD76F911E16DCFE18FBC0FDBF" collectingDate="2015-08" collectionCode="DCBU" collectorName="Parque Nacional da Serra dos Orgaos &amp; R. F. Monteiro" country="Brazil" county="Teresopolis" elevation="563" latitude="-22.476667" location="Malaise" longLatPrecision="21" longitude="-42.991943" municipality="Parque Nacional da Serra" pageId="3" pageNumber="143" specimenCode="DCBU 473.862" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<typeStatus id="54B424E3D76F911E16DCFE18FE82FE53" box="[199,306,475,501]" pageId="3" pageNumber="143" type="holotype">Holotype</typeStatus>
, female.
<collectingCountry id="F318DAD1D76F911E1781FE18FDB3FE53" box="[410,515,475,501]" name="Brazil" pageId="3" pageNumber="143">BRAZIL</collectingCountry>
.
<emphasis id="B97B4653D76F911E1414FE18FD04FE52" box="[527,692,475,501]" italics="true" pageId="3" pageNumber="143">
<collectingRegion id="49CB54A3D76F911E1414FE18FD04FE52" box="[527,692,475,501]" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="143">Rio de Janeiro</collectingRegion>
</emphasis>
:
<collectingCounty id="62D1E2CDD76F911E14DFFE18FCF8FE53" box="[708,840,475,501]" pageId="3" pageNumber="143">Teresópolis</collectingCounty>
,
<collectorName id="26FAFF97D76F911E154FFE18FAB0FE51" box="[852,1280,475,503]" pageId="3" pageNumber="143">
<collectingMunicipality id="6BD4003BD76F911E154FFE18FBC6FE53" box="[852,1142,475,501]" pageId="3" pageNumber="143">Parque Nacional da Serra</collectingMunicipality>
dos Órgãos
</collectorName>
,
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22
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28’36”S
</geoCoordinate>
–
<geoCoordinate id="EE3BFC86D76F911E168CFE3CFE93FDBF" box="[151,291,510,537]" degrees="42" direction="west" minutes="59" orientation="longitude" pageId="3" pageNumber="143" precision="15" seconds="31" value="-42.991943">
42
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59’31”W
</geoCoordinate>
,
<location id="8ED0CC9AD76F911E172BFE3CFE39FDBF" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E172BFE3CFE39FDBF" box="[304,393,511,537]" country="Brazil" county="Teresopolis" latitude="-22.476667" longLatPrecision="21" longitude="-42.991943" municipality="Parque Nacional da Serra" name="Malaise" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Malaise</location>
4a,
<quantity id="4CF737A4D76F911E17A0FE3DFDB6FDBF" box="[443,518,510,537]" metricMagnitude="2" metricUnit="m" metricValue="5.63" pageId="3" pageNumber="143" unit="m" value="563.0">
<elevation id="00227D72D76F911E17A0FE3DFDB6FDBF" box="[443,518,510,537]" metricMagnitude="2" metricUnit="m" metricValue="5.63" pageId="3" pageNumber="143" unit="m" value="563.0">563 m</elevation>
</quantity>
,
<date id="FFB1BC81D76F911E1409FE3DFD34FDBF" box="[530,644,510,537]" pageId="3" pageNumber="143" value="2015-08">
<collectingDate id="EFF54569D76F911E1409FE3DFD34FDBF" box="[530,644,510,537]" pageId="3" pageNumber="143" value="2015-08">VIII.2015</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E148AFE3CFC9FFDBF" box="[657,815,511,537]" pageId="3" pageNumber="143">R.F. Monteiro</collectorName>
e eq. col. (
<specimenCode id="DBA9323AD76F911E15ABFE3CFBD8FDBF" box="[944,1128,511,537]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 473.862</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E1260FE3CFAE2FD9B" collectingDate="2015-08" collectionCode="DCBU" collectorName="Malaise &amp; R. F. Monteiro" country="Brazil" elevation="332" latitude="-22.494444" location="Same" longLatPrecision="21" longitude="-42.99778" pageId="3" pageNumber="143" specimenCode="DCBU 473.849" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<typeStatus id="54B424E3D76F911E1260FE3CFB5BFDBF" box="[1147,1259,511,537]" pageId="3" pageNumber="143" type="paratype">Paratypes</typeStatus>
, females.
<location id="8ED0CC9AD76F911E1345FE3CFA2CFDBF" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E1345FE3CFA2CFDBF" box="[1374,1436,511,537]" country="Brazil" latitude="-22.494444" longLatPrecision="21" longitude="-42.99778" name="Same" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Same</location>
data,
<geoCoordinate id="EE3BFC86D76F911E16CDFDE0FEE3FD9B" box="[214,339,547,573]" degrees="22" direction="south" minutes="29" orientation="latitude" pageId="3" pageNumber="143" precision="15" seconds="40" value="-22.494444">
22
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29’40”S
</geoCoordinate>
–
<geoCoordinate id="EE3BFC86D76F911E1778FDE0FE42FD9B" box="[355,498,546,573]" degrees="42" direction="west" minutes="59" orientation="longitude" pageId="3" pageNumber="143" precision="15" seconds="52" value="-42.99778">
42
<superScript id="7C7A3709D76F911E179AFDE0FE39FD97" attach="none" box="[385,393,547,561]" fontSize="6" pageId="3" pageNumber="143">o</superScript>
59’52”W
</geoCoordinate>
,
<collectorName id="26FAFF97D76F911E17E4FDE0FDE8FD9B" box="[511,600,547,573]" pageId="3" pageNumber="143">Malaise</collectorName>
3b,
<quantity id="4CF737A4D76F911E1496FDE0FD68FD9B" box="[653,728,547,573]" metricMagnitude="2" metricUnit="m" metricValue="3.32" pageId="3" pageNumber="143" unit="m" value="332.0">
<elevation id="00227D72D76F911E1496FDE0FD68FD9B" box="[653,728,547,573]" metricMagnitude="2" metricUnit="m" metricValue="3.32" pageId="3" pageNumber="143" unit="m" value="332.0">332 m</elevation>
</quantity>
,
<date id="FFB1BC81D76F911E14FDFDE1FCE8FD9B" box="[742,856,546,573]" pageId="3" pageNumber="143" value="2015-08">
<collectingDate id="EFF54569D76F911E14FDFDE1FCE8FD9B" box="[742,856,546,573]" pageId="3" pageNumber="143" value="2015-08">VIII.2015</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E157EFDE0FBBDFD9B" box="[869,1037,547,573]" pageId="3" pageNumber="143">R. F. Monteiro</collectorName>
e eq. col. (
<specimenCode id="DBA9323AD76F911E128AFDE0FAF9FD9B" box="[1169,1353,547,573]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 473.849</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E1345FDE0FAE1FDC7" collectingDate="2015-09" collectionCode="DCBU" collectorName="Malaise &amp; R. F. Monteiro" country="Brazil" elevation="544" latitude="-22.476389" location="Same" longLatPrecision="21" longitude="-42.99139" pageId="3" pageNumber="143" specimenCode="DCBU 495.369" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<location id="8ED0CC9AD76F911E1345FDE0FA2CFD9B" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E1345FDE0FA2CFD9B" box="[1374,1436,547,573]" country="Brazil" latitude="-22.476389" longLatPrecision="21" longitude="-42.99139" name="Same" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Same</location>
data,
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22
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28’35”S
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<geoCoordinate id="EE3BFC86D76F911E1775FD85FE4EFDC7" box="[366,510,582,609]" degrees="42" direction="west" minutes="59" orientation="longitude" pageId="3" pageNumber="143" precision="15" seconds="29" value="-42.99139">
42
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59’29”W
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<collectorName id="26FAFF97D76F911E1417FD84FDD5FDC7" box="[524,613,583,609]" pageId="3" pageNumber="143">Malaise</collectorName>
4b,
<quantity id="4CF737A4D76F911E1487FD85FD59FDC7" box="[668,745,582,609]" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" pageId="3" pageNumber="143" unit="m" value="544.0">
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,
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<collectingDate id="EFF54569D76F911E14E3FD85FCE7FDC7" box="[760,855,582,609]" pageId="3" pageNumber="143" value="2015-09">IX.2015</collectingDate>
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<collectorName id="26FAFF97D76F911E157DFD84FBB6FDC7" box="[870,1030,583,609]" pageId="3" pageNumber="143">R.F. Monteiro</collectorName>
e eq. col. (
<specimenCode id="DBA9323AD76F911E1295FD84FAF8FDC7" box="[1166,1352,582,609]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 495.369</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E1345FD84FCDDFD23" collectingDate="2009-11-03" collectionCode="DCBU" collectorName="R. F. Monteiro" country="Brazil" location="Malaise" municipality="Same" pageId="3" pageNumber="143" specimenCode="DCBU 495.362" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingMunicipality id="6BD4003BD76F911E1345FD84FA2CFDC7" box="[1374,1436,583,609]" pageId="3" pageNumber="143">Same</collectingMunicipality>
data,
<location id="8ED0CC9AD76F911E16CFFDA8FE9DFD23" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E16CFFDA8FE9DFD23" box="[212,301,619,645]" country="Brazil" municipality="Same" name="Malaise" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Malaise</location>
1,
<date id="FFB1BC81D76F911E174AFDA8FE75FD23" box="[337,453,619,645]" pageId="3" pageNumber="143" value="2009-11-03">
<collectingDate id="EFF54569D76F911E174AFDA8FE75FD23" box="[337,453,619,645]" pageId="3" pageNumber="143" value="2009-11-03">3.XI.2009</collectingDate>
</date>
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<collectorName id="26FAFF97D76F911E17CBFDA8FDDEFD23" box="[464,622,619,645]" pageId="3" pageNumber="143">R.F. Monteiro</collectorName>
col. (
<specimenCode id="DBA9323AD76F911E14B6FDA8FCD4FD23" box="[685,868,618,645]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 495.362</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E156DFDA8FAA8FD0F" collectingDate="2017-10" collectionCode="DCBU" collectorName="L. Campello" country="Brazil" county="Angra" elevation="5441" location="Parque Estadual da Ilha Grande" municipality="Reis" pageId="3" pageNumber="143" specimenCode="DCBU 495.363, DCBU 495.367" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingCounty id="62D1E2CDD76F911E156DFDA8FC0CFD23" box="[886,956,619,645]" pageId="3" pageNumber="143">Angra</collectingCounty>
dos
<collectingMunicipality id="6BD4003BD76F911E15EAFDA8FB95FD23" box="[1009,1061,619,645]" pageId="3" pageNumber="143">Reis</collectingMunicipality>
,
<location id="8ED0CC9AD76F911E122AFDA8FA28FD23" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E122AFDA8FA28FD23" box="[1073,1432,619,645]" country="Brazil" county="Angra" municipality="Reis" name="Parque Estadual da Ilha Grande" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Parque Estadual da Ilha Grande</location>
,
<location id="8ED0CC9AD76F911E168CFD4CFEECFD0F" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E168CFD4CFEECFD0F" box="[151,348,655,681]" country="Brazil" county="Angra" municipality="Reis" name="Pico do Papagaio" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Pico do Papagaio</location>
,
<location id="8ED0CC9AD76F911E177CFD4CFE70FD0F" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E177CFD4CFE70FD0F" box="[359,448,655,681]" country="Brazil" county="Angra" municipality="Reis" name="Malaise" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Malaise</location>
<quantity id="4CF737A4D76F911E17DDFD4DFD9BFD0F" box="[454,555,654,681]" metricMagnitude="3" metricUnit="m" metricValue="5.441" pageId="3" pageNumber="143" unit="m" value="5441.0">
<elevation id="00227D72D76F911E17DDFD4DFD9BFD0F" box="[454,555,654,681]" metricMagnitude="3" metricUnit="m" metricValue="5.441" pageId="3" pageNumber="143" unit="m" value="5441.0">5, 441 m</elevation>
</quantity>
,
<date id="FFB1BC81D76F911E142DFD4CFD3BFD0F" box="[566,651,655,681]" pageId="3" pageNumber="143" value="2017-10">
<collectingDate id="EFF54569D76F911E142DFD4CFD3BFD0F" box="[566,651,655,681]" pageId="3" pageNumber="143" value="2017-10">X.2017</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E148CFD4CFC93FD0F" box="[663,803,655,681]" pageId="3" pageNumber="143">L. Campello</collectorName>
e eq. col. (
<specimenCode id="DBA9323AD76F911E1587FD4CFBE0FD0F" box="[924,1104,654,681]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 495.363</specimenCode>
,
<specimenCode id="DBA9323AD76F911E1240FD4CFAA0FD0F" box="[1115,1296,654,681]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 495.367</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E1339FD4CFC37FD6B" collectingDate="2017-10" collectionCode="DCBU" collectorName="L. Campello" country="Brazil" elevation="3336" location="Malaise" municipality="Same" pageId="3" pageNumber="143" specimenCode="DCBU 495.365" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingMunicipality id="6BD4003BD76F911E1339FD4CFAD0FD0F" box="[1314,1376,655,681]" pageId="3" pageNumber="143">Same</collectingMunicipality>
data,
<location id="8ED0CC9AD76F911E168CFD70FF40FD6B" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E168CFD70FF40FD6B" box="[151,240,691,717]" country="Brazil" municipality="Same" name="Malaise" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Malaise</location>
<quantity id="4CF737A4D76F911E16EDFD70FEEAFD6B" box="[246,346,691,717]" metricMagnitude="3" metricUnit="m" metricValue="3.336" pageId="3" pageNumber="143" unit="m" value="3336.0">
<elevation id="00227D72D76F911E16EDFD70FEEAFD6B" box="[246,346,691,717]" metricMagnitude="3" metricUnit="m" metricValue="3.336" pageId="3" pageNumber="143" unit="m" value="3336.0">3, 336 m</elevation>
</quantity>
,
<date id="FFB1BC81D76F911E177EFD70FE0AFD6B" box="[357,442,691,717]" pageId="3" pageNumber="143" value="2017-10">
<collectingDate id="EFF54569D76F911E177EFD70FE0AFD6B" box="[357,442,691,717]" pageId="3" pageNumber="143" value="2017-10">X.2017</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E17DDFD70FDE1FD6B" box="[454,593,691,717]" pageId="3" pageNumber="143">L. Campello</collectorName>
e eq. col. (
<specimenCode id="DBA9323AD76F911E14D2FD70FCCFFD6B" box="[713,895,690,717]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 495.365</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E158AFD70FD0DFD57" collectingDate="2017-10" collectionCode="DCBU" collectorName="L. Campello" country="Brazil" county="Same" elevation="1160" location="Malaise" municipality="Pico do Soldado" pageId="3" pageNumber="143" specimenCode="DCBU 495.368" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingCounty id="62D1E2CDD76F911E158AFD70FC7FFD6B" box="[913,975,691,717]" pageId="3" pageNumber="143">Same</collectingCounty>
data,
<collectingMunicipality id="6BD4003BD76F911E120AFD70FB7BFD6B" box="[1041,1227,691,717]" pageId="3" pageNumber="143">Pico do Soldado</collectingMunicipality>
,
<location id="8ED0CC9AD76F911E12CEFD70FA9EFD6B" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E12CEFD70FA9EFD6B" box="[1237,1326,691,717]" country="Brazil" county="Same" municipality="Pico do Soldado" name="Malaise" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Malaise</location>
<quantity id="4CF737A4D76F911E132FFD70FA27FD6B" box="[1332,1431,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="3" pageNumber="143" unit="m" value="1160.0">
<elevation id="00227D72D76F911E132FFD70FA27FD6B" box="[1332,1431,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="3" pageNumber="143" unit="m" value="1160.0">1, 160 m</elevation>
</quantity>
,
<date id="FFB1BC81D76F911E168CFD14FF5DFD57" box="[151,237,727,753]" pageId="3" pageNumber="143" value="2017-10">
<collectingDate id="EFF54569D76F911E168CFD14FF5DFD57" box="[151,237,727,753]" pageId="3" pageNumber="143" value="2017-10">X.2017</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E16E3FD14FE34FD57" box="[248,388,727,753]" pageId="3" pageNumber="143">L. Campello</collectorName>
e eq. col. (
<specimenCode id="DBA9323AD76F911E17E5FD14FD05FD57" box="[510,693,726,753]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 495.368</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E14DCFD14FC20FCB3" collectingDate="2014-12-05" collectionCode="DCBU" collectorName="Malaise &amp; R. F. Monteiro" country="Brazil" elevation="1" latitude="-22.433722" location="Parque Nacional de Itatiaia" longLatPrecision="1" longitude="-44.613747" municipality="Itatiaia" pageId="3" pageNumber="143" specimenCode="DCBU 113.181" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingMunicipality id="6BD4003BD76F911E14DCFD14FCA8FD57" box="[711,792,727,753]" pageId="3" pageNumber="143">Itatiaia</collectingMunicipality>
,
<location id="8ED0CC9AD76F911E1539FD14FBE5FD57" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E1539FD14FBE5FD57" box="[802,1109,727,753]" country="Brazil" latitude="-22.433722" longLatPrecision="1" longitude="-44.613747" municipality="Itatiaia" name="Parque Nacional de Itatiaia" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Parque Nacional de Itatiaia</location>
,
<geoCoordinate id="EE3BFC86D76F911E127BFD14FB53FD57" box="[1120,1251,727,753]" degrees="22" direction="south" minutes="26" orientation="latitude" pageId="3" pageNumber="143" precision="1" seconds="1.4" value="-22.433722">
22
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26’1.4”S
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–
<geoCoordinate id="EE3BFC86D76F911E12EFFD14FA28FD57" box="[1268,1432,726,753]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="3" pageNumber="143" precision="1" seconds="49.5" value="-44.613747">
44
<superScript id="7C7A3709D76F911E1309FD14FAAAFD43" attach="none" box="[1298,1306,727,741]" fontSize="6" pageId="3" pageNumber="143">o</superScript>
36’49.5”W
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,
<collectorName id="26FAFF97D76F911E168CFD38FF40FCB3" box="[151,240,763,789]" pageId="3" pageNumber="143">Malaise</collectorName>
1,
<quantity id="4CF737A4D76F911E1714FD38FEDCFCB3" box="[271,364,763,789]" metricMagnitude="0" metricUnit="m" metricValue="1.07" pageId="3" pageNumber="143" unit="m" value="1.07">
<elevation id="00227D72D76F911E1714FD38FEDCFCB3" box="[271,364,763,789]" metricMagnitude="0" metricUnit="m" metricValue="1.07" pageId="3" pageNumber="143" unit="m" value="1.07">1.070 m</elevation>
</quantity>
,
<date id="FFB1BC81D76F911E176DFD39FE43FCB3" box="[374,499,762,789]" pageId="3" pageNumber="143" value="2014-12-05">
<collectingDate id="EFF54569D76F911E176DFD39FE43FCB3" box="[374,499,762,789]" pageId="3" pageNumber="143" value="2014-12-05">5.XII.2014</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E17E6FD38FD28FCB3" box="[509,664,763,789]" pageId="3" pageNumber="143">R.F. Monteiro</collectorName>
col. (
<specimenCode id="DBA9323AD76F911E14CFFD38FC38FCB3" box="[724,904,763,789]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 113.181</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E1582FD38FD50FC9F" collectingDate="2013-11-21" collectionCode="DCBU" collectorName="R. F. Monteiro" country="Brazil" latitude="-22.433722" location="Malaise" longLatPrecision="14" longitude="-44.61361" municipality="Same" pageId="3" pageNumber="143" specimenCode="DCBU 473.847" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingMunicipality id="6BD4003BD76F911E1582FD38FC67FCB3" box="[921,983,763,789]" pageId="3" pageNumber="143">Same</collectingMunicipality>
data,
<geoCoordinate id="EE3BFC86D76F911E120CFD38FB2AFCB3" box="[1047,1178,763,789]" degrees="22" direction="south" minutes="26" orientation="latitude" pageId="3" pageNumber="143" precision="1" seconds="1.4" value="-22.433722">
22
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26’1.4”S
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–
<geoCoordinate id="EE3BFC86D76F911E12B0FD38FA8AFCB3" box="[1195,1338,763,789]" degrees="44" direction="west" minutes="36" orientation="longitude" pageId="3" pageNumber="143" precision="15" seconds="49" value="-44.61361">
44
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36’49”W
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,
<location id="8ED0CC9AD76F911E135FFD38FA2DFCB3" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E135FFD38FA2DFCB3" box="[1348,1437,763,789]" country="Brazil" latitude="-22.433722" longLatPrecision="14" longitude="-44.61361" municipality="Same" name="Malaise" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Malaise</location>
1,
<date id="FFB1BC81D76F911E16AFFCDCFE86FC9F" box="[180,310,799,825]" pageId="3" pageNumber="143" value="2013-11-21">
<collectingDate id="EFF54569D76F911E16AFFCDCFE86FC9F" box="[180,310,799,825]" pageId="3" pageNumber="143" value="2013-11-21">21.XI.2013</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E1759FCDCFE50FC9F" box="[322,480,799,825]" pageId="3" pageNumber="143">R.F. Monteiro</collectorName>
col. (
<specimenCode id="DBA9323AD76F911E143AFCDCFD68FC9F" box="[545,728,799,825]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 473.847</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E14F0FCDCFD43FCFB" collectingDate="2019-08-14" collectingDateMax="2019-08-17" collectingDateMin="2019-08-14" collectionCode="DCBU" collectorName="A. S. Soares" country="Brazil" county="Paraty" location="Moericke" municipality="Estacao Ecologica de Tamoios" pageId="3" pageNumber="143" specimenCode="DCBU 496.364" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingCounty id="62D1E2CDD76F911E14F0FCDCFC84FC9F" box="[747,820,799,825]" pageId="3" pageNumber="143">Paraty</collectingCounty>
,
<collectingMunicipality id="6BD4003BD76F911E155BFCDCFB2BFC9F" box="[832,1179,799,825]" pageId="3" pageNumber="143">Estação Ecológica de Tamoios</collectingMunicipality>
, continente,
<location id="8ED0CC9AD76F911E1331FCDCFA28FC9F" LSID="urn:lsid:plazi:treatment:03A62B57D76C911E168CF9A0FC0BFB93:8ED0CC9AD76F911E1331FCDCFA28FC9F" box="[1322,1432,799,825]" country="Brazil" county="Paraty" municipality="Estacao Ecologica de Tamoios" name="Moericke" pageId="3" pageNumber="143" stateProvince="Rio de Janeiro">Moericke</location>
,
<date id="FFB1BC81D76F911E168CFC80FEE6FCFB" box="[151,342,835,861]" pageId="3" pageNumber="143" value="2019-08-14" valueMax="2019-08-17" valueMin="2019-08-14">
<collectingDate id="EFF54569D76F911E168CFC80FEE6FCFB" box="[151,342,835,861]" pageId="3" pageNumber="143" value="2019-08-14" valueMax="2019-08-17" valueMin="2019-08-14">14–17.VIII.2019</collectingDate>
</date>
,
<collectorName id="26FAFF97D76F911E1778FC80FE5BFCFB" box="[355,491,835,861]" pageId="3" pageNumber="143">A.S. Soares</collectorName>
col. (
<specimenCode id="DBA9323AD76F911E142AFC80FD5BFCFB" box="[561,747,835,861]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="3" pageNumber="143">DCBU 496.364</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B67901CD76F911E151AFC80FE21FC27" collectingDate="2018-06-20" collectionCode="DCBU" collectorName="A. S. Soares" country="Brazil" county="Same" location="Malaise" municipality="Ilha Sandri" pageId="3" pageNumber="143" specimenCode="DCBU 473.846" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingCounty id="62D1E2CDD76F911E151AFC80FC8FFCFB" box="[769,831,835,861]" pageId="3" pageNumber="143">Same</collectingCounty>
data,
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287
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<treatment id="03A62B57D76F911B168CFB44FB1EFC6F" ID-DOI="http://doi.org/10.5281/zenodo.13330782" ID-Zenodo-Dep="13330782" LSID="urn:lsid:plazi:treatment:03A62B57D76F911B168CFB44FB1EFC6F" httpUri="http://treatment.plazi.org/id/03A62B57D76F911B168CFB44FB1EFC6F" lastPageId="6" lastPageNumber="146" pageId="3" pageNumber="143">
<subSubSection id="C315C9CAD76F911E168CFB44FC9BFB06" box="[151,811,1158,1185]" pageId="3" pageNumber="143" type="nomenclature">
<paragraph id="8BB09A41D76F911E168CFB44FC9BFB06" blockId="3.[151,811,1158,1221]" box="[151,811,1158,1185]" pageId="3" pageNumber="143">
<heading id="D0F82D2DD76F911E168CFB44FC9BFB06" bold="true" box="[151,811,1158,1185]" fontSize="11" level="1" pageId="3" pageNumber="143" reason="1">
<emphasis id="B97B4653D76F911E168CFB44FC9BFB06" bold="true" box="[151,811,1158,1185]" pageId="3" pageNumber="143">
<taxonomicName id="4C0FE1C2D76F911E168CFB44FE1CFB07" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[151,428,1158,1185]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="fernandesi" status="sp. nov.">
<emphasis id="B97B4653D76F911E168CFB44FE1CFB07" bold="true" box="[151,428,1158,1185]" italics="true" pageId="3" pageNumber="143">Dentigaster fernandesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248FB28D76F911E17A8FB44FDA0FB07" box="[435,528,1159,1185]" pageId="3" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
Dias &amp; Penteado-Dias
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315C9CAD76F911B168CFB68FB1EFC6F" lastPageId="6" lastPageNumber="146" pageId="3" pageNumber="143" type="description">
<paragraph id="8BB09A41D76F911E168CFB68FE96FB63" blockId="3.[151,811,1158,1221]" box="[151,294,1195,1221]" pageId="3" pageNumber="143">
(
<figureCitation id="133486C4D76F911E1684FB68FEAEFB63" box="[159,286,1195,1221]" captionStart-0="FIGURES 11–17" captionStart-1="FIGURE 18" captionStartId-0="4.[152,270,1776,1800]" captionStartId-1="5.[152,255,1113,1137]" captionTargetBox-0="[151,1435,181,1753]" captionTargetBox-1="[151,1436,181,1089]" captionTargetId-0="figure-17@4.[151,1435,181,1753]" captionTargetId-1="figure-25@5.[151,1436,181,1089]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." captionText-1="FIGURE 18. Map of distribution of species: z, Dentigaster medeirosi sp. nov. ▲, Dentigaster fernandesi sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.13330846" figureDoi-1="http://doi.org/10.5281/zenodo.13330848" httpUri-0="https://zenodo.org/record/13330846/files/figure.png" httpUri-1="https://zenodo.org/record/13330848/files/figure.png" pageId="3" pageNumber="143">Figs 11–18</figureCitation>
)
</paragraph>
<paragraph id="8BB09A41D76F911E168CFB30FE4EFAAB" blockId="3.[151,1437,1267,2013]" box="[151,510,1267,1293]" pageId="3" pageNumber="143">
<typeStatus id="54B424E3D76F911E168CFB30FEB4FAAB" box="[151,260,1267,1293]" pageId="3" pageNumber="143" type="holotype">Holotype</typeStatus>
: body length, 7.0 mm.
</paragraph>
<paragraph id="8BB09A41D76F911E16DCFAD4FA26F91B" blockId="3.[151,1437,1267,2013]" pageId="3" pageNumber="143">
Color: Scapus and pedicellus orange; first and second flagellomeres light yellow (second brown at the apex); remaining flagellomeres brown. Head orange yellow, with a dark brown patch on the face and other on the front to the vertex (
<figureCitation id="133486C4D76F911E170EFA9CFE27FADF" box="[277,407,1375,1401]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="3" pageNumber="143">Figs 12, 13</figureCitation>
). Propleuron blackish dark brown. Pronotum yellow, with brown patch at the lower margin. Scutellum, in dorsal view, blackish dark brown; anterior area at the scutellar sulcus brown, on each side with small yellow patch not very distinct; lateral area to the disc light yellow. Mesoscutum blackish dark brown, with two orange longitudinal bands. Mesopleuron and metapleuron blackish dark brown, both with an anterior orange patch. Propodeum with same color, with posterior orange patch in each side. Fore leg mostly yellow, femur brownish. Mid leg partially yellow, femur and tibia brownish. Hind leg mostly brown; coxa with dark brown and yellow parts. Tegula light brown and humeral plate yellowish light brown. Fore wing (
<figureCitation id="133486C4D76F911E15A5F9F4FBBFF9F7" box="[958,1039,1591,1617]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="3" pageNumber="143">Fig. 14</figureCitation>
) grayish brown, partially hyaline in the basal cell; transverse white band at level of basal part of the pterostigma, white bulla in vein r-m and white patch in the third submarginal cell, slightly prolonged below 3M vein. Hind wing light brown. Fore and hind wings with dark brown veins. Metasoma black, with yellow lateral patches on the three tergites, reduced in the third (
<figureCitation id="133486C4D76F911E132CF960FA38F91B" box="[1335,1416,1699,1725]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="3" pageNumber="143">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BB09A41D76F911E16DCF904FDADF87B" blockId="3.[151,1437,1267,2013]" pageId="3" pageNumber="143">Head: Antenna with 52 flagellomeres. Scapus 2.70x as long as maximally wide. First flagellomere 1.06x longer than scapus, 7.13x as long as wide, 1.27x as long as the second one. Penultimate flagellomere 1.33x as long as wide, 0.07x as long as first flagellomere, 0.57x as long as apical flagellomere. Head 1.67x as wide as its median length. Eye 1.87x the length of the temple (in dorsal view), 1.97x as high as wide. Head moderately coarse punctate. Clypeus moderately swollen in lateral view, with two small blunt teeth in lower margin; surface moderately punctate. Malar space 1.14x basal width of mandible, 0.51x eye height. Face moderately convex, with slightly protruding median carina. Temple not swollen in dorsal view. Frons flat, not concave. POL 1.33x ocellar diameter, 0.41x OOL. Occipital carina present, complete.</paragraph>
<caption id="DF70CAC9D76891191683F933FEF2F8F7" ID-DOI="http://doi.org/10.5281/zenodo.13330846" ID-Zenodo-Dep="13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="4" pageNumber="144" startId="4.[152,270,1776,1800]" targetBox="[151,1435,181,1753]" targetPageId="4" targetType="figure">
<paragraph id="8BB09A41D76891191683F933FEF2F8F7" blockId="4.[151,1437,1776,1873]" pageId="4" pageNumber="144">
<emphasis id="B97B4653D76891191683F933FEECF8AF" bold="true" box="[152,348,1776,1801]" pageId="4" pageNumber="144">FIGURES 11–17.</emphasis>
<taxonomicName id="4C0FE1C2D7689119177EF932FDE1F8AF" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[357,593,1777,1801]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="fernandesi" status="sp. nov.">
<emphasis id="B97B4653D7689119177EF932FDE1F8AF" box="[357,593,1777,1801]" italics="true" pageId="4" pageNumber="144">Dentigaster fernandesi</emphasis>
</taxonomicName>
<emphasis id="B97B4653D76891191441F932FD1EF8AF" bold="true" box="[602,686,1777,1801]" pageId="4" pageNumber="144">
<taxonomicNameLabel id="A248FB28D76891191441F932FD1EF8AF" box="[602,686,1777,1801]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal.
</paragraph>
</caption>
<caption id="DF70CAC9D76991181683FB9AFA9DFBD7" ID-DOI="http://doi.org/10.5281/zenodo.13330848" ID-Zenodo-Dep="13330848" box="[152,1325,1113,1137]" httpUri="https://zenodo.org/record/13330848/files/figure.png" pageId="5" pageNumber="145" startId="5.[152,255,1113,1137]" targetBox="[151,1436,181,1089]" targetPageId="5" targetType="figure">
<paragraph id="8BB09A41D76991181683FB9AFA9DFBD7" blockId="5.[152,1325,1113,1137]" box="[152,1325,1113,1137]" pageId="5" pageNumber="145">
<emphasis id="B97B4653D76991181683FB9AFE95FBD7" bold="true" box="[152,293,1113,1137]" pageId="5" pageNumber="145">FIGURE 18.</emphasis>
Map of distribution of species:
<emphasis id="B97B4653D76991181472FB9FFDCCFBD7" bold="true" box="[617,636,1116,1137]" italics="true" pageId="5" pageNumber="145">z</emphasis>
,
<taxonomicName id="4C0FE1C2D76991181493FB99FCD9FBD7" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[648,873,1113,1137]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="medeirosi" status="sp. nov.">
<emphasis id="B97B4653D76991181493FB99FCD9FBD7" box="[648,873,1113,1137]" italics="true" pageId="5" pageNumber="145">Dentigaster medeirosi</emphasis>
</taxonomicName>
<emphasis id="B97B4653D7699118156BFB9AFC70FBD7" bold="true" box="[880,960,1113,1137]" pageId="5" pageNumber="145">
<taxonomicNameLabel id="A248FB28D7699118156BFB9AFC70FBD7" box="[880,960,1113,1137]" pageId="5" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
▲,
<taxonomicName id="4C0FE1C2D769911815F6FB99FB66FBD7" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[1005,1238,1113,1137]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="fernandesi" status="sp. nov.">
<emphasis id="B97B4653D769911815F6FB99FB66FBD7" box="[1005,1238,1113,1137]" italics="true" pageId="5" pageNumber="145">Dentigaster fernandesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248FB28D769911812C7FB9AFA9DFBD7" box="[1244,1325,1113,1137]" pageId="5" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB09A41D769911816DCFB07FE10F9E0" blockId="5.[151,1437,1219,2038]" pageId="5" pageNumber="145">Mesosoma: Mesosoma 1.74x as long as wide in lateral view, 1.56x as long as wide in dorsal view. Pronotum finely punctate. Mesoscutum moderately prominent, rounded anteriorly, coarsely to moderately punctate. Notauli slightly evident. Scutellar sulcus with nine complete transversal carinae. Mesopleuron, metapleuron and propodeum moderately coarse punctate; a small protrusion, longitudinally elongated on each side of propodeum; mediolongitudinal carina absent. Fore wing: stigma 6.17x as long as wide. Marginal cell distally closed. Vein r 0.64x as long as vein 3-SR, 0.21x as long as vein SR1, 0.39x as long as vein 2-SR. Vein 1-SR + M present. First discal cell 2.55x as long as wide. Vein r-m present, with one bulla. Vein m-cu and vein cu-a present, postfurcal. Vein CU1b present. First subdiscal cell distally closed. Vein 2-1A long. Hind wing: Vein 1-SC+R present. Subbasal cell medium sized. Vein M + CU 0.88x as long as vein 1-M. Legs: Hind coxa large. Hind femur 4.03x as long as wide. Hind tibia 7.4x as long as wide, 2.39x as long as the hind basitarsomere. Hind basitarsomere 7.15x as long as wide, 0.97x as long as tarsomeres 2–5.</paragraph>
<paragraph id="8BB09A41D769911816DCF993FAF6F95F" blockId="5.[151,1437,1219,2038]" pageId="5" pageNumber="145">Metasoma: Carapace with two transverse areolate sutures; between the first and second tergites wider, mainly in the middle area. First and second tergites rugose- areolate. Third tergite finely rugose-areolate. First tergite, in dorsal view 1.11x as long as apical width with anterior prominent median knob. Median length of second tergite 0.75x its basal width, 0.86x as long as first tergite. Carapace 2.25x as long as wide, in dorsal view. Third tergite with apex not denticulated in dorsal view. Postero-ventral rim on third tergite, in lateral view, moderately curved upward.</paragraph>
<paragraph id="8BB09A41D769911816DCF8C0FE36F8BB" blockId="5.[151,1437,1219,2038]" box="[199,390,1795,1821]" pageId="5" pageNumber="145">Male: Unknown.</paragraph>
<paragraph id="8BB09A41D769911B16DCF8E4FF73FF73" blockId="5.[151,1437,1219,2038]" lastBlockId="6.[151,1437,151,969]" lastPageId="6" lastPageNumber="146" pageId="5" pageNumber="145">
Variations: Scapus and pedicellus orange or orange brownish. Pronotum all yellow or with small to large brown patch at the lower margin. Propleuron, mesopleuron and metapleuron blackish dark brown with yellow patches variously small to large. Two orange longitudinal bands of the mesoscutum variously wide to narrow (
<figureCitation id="133486C4D76991181322F8ACFA3FF82F" box="[1337,1423,1903,1929]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="5" pageNumber="145">Fig. 12</figureCitation>
). Propodeum and metapleuron can be completely blackish dark brown or propodeum with yellow lateral patches visible or almost imperceptibles. Fore wing with white patch at the third submarginal cell (
<figureCitation id="133486C4D769911812A1F874FAA0F877" box="[1210,1296,1975,2001]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="5" pageNumber="145">Fig. 14</figureCitation>
), slightly to markedly prolonged below 3M vein. Carapace lateral yellow patches with variable sizes, that can be large, including those in the third tergite (
<figureCitation id="133486C4D76A911B17ADFF54FDD1FF17" box="[438,609,151,177]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="6" pageNumber="146">Figs 11, 16, 17</figureCitation>
) or represented only at the lower margin, not visible in dorsal view (
<figureCitation id="133486C4D76A911B1374FF54FF06FF73" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="6" pageNumber="146">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BB09A41D76A911B16DCFF1CFC83FF5F" blockId="6.[151,1437,151,969]" box="[199,819,223,249]" pageId="6" pageNumber="146">Etymology: In honour to our friend Ulysses Fernandes.</paragraph>
<paragraph id="8BB09A41D76A911B16DCFEC0FE3AFEBB" blockId="6.[151,1437,151,969]" box="[199,394,259,285]" pageId="6" pageNumber="146">Habitat: Cerrado.</paragraph>
<paragraph id="8BB09A41D76A911B16DCFEE4FE12FEE7" blockId="6.[151,1437,151,969]" box="[199,418,295,321]" pageId="6" pageNumber="146">Material examined:</paragraph>
<paragraph id="8BB09A41D76A911B16DCFE88FB51FE77" blockId="6.[151,1437,151,969]" pageId="6" pageNumber="146">
<materialsCitation id="3B67901CD76A911B16DCFE88FA29FE2F" collectingDate="2018-08-28" collectionCode="DCBU" collectorName="N. W. Perioto &amp; R. I. R Lara" country="Brazil" county="Chapada Gaucha" elevation="2764" latitude="-15.174723" location="Cerrado" longLatPrecision="21" longitude="-45.72139" municipality="Parque Nacional Grande Sertao Veredas" pageId="6" pageNumber="146" specimenCode="DCBU 473.863" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="54B424E3D76A911B16DCFE88FE82FEC3" box="[199,306,331,357]" pageId="6" pageNumber="146" type="holotype">Holotype</typeStatus>
, female.
<collectingCountry id="F318DAD1D76A911B178FFE88FE4DFEC3" box="[404,509,331,357]" name="Brazil" pageId="6" pageNumber="146">BRAZIL</collectingCountry>
.
<emphasis id="B97B4653D76A911B141DFE88FD2DFEC3" box="[518,669,331,357]" italics="true" pageId="6" pageNumber="146">
<collectingRegion id="49CB54A3D76A911B141DFE88FD2DFEC3" box="[518,669,331,357]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="146">Minas Gerais</collectingRegion>
</emphasis>
:
<collectingCounty id="62D1E2CDD76A911B14B0FE88FCDAFEC3" box="[683,874,331,357]" pageId="6" pageNumber="146">Chapada Gaúcha</collectingCounty>
,
<collectingMunicipality id="6BD4003BD76A911B156FFE88FA82FEC3" box="[884,1330,331,357]" pageId="6" pageNumber="146">Parque Nacional Grande Sertão Veredas</collectingMunicipality>
,
<location id="8ED0CC9AD76A911B1320FE88FA28FEC3" LSID="urn:lsid:plazi:treatment:03A62B57D76F911B168CFB44FB1EFC6F:8ED0CC9AD76A911B1320FE88FA28FEC3" box="[1339,1432,331,357]" country="Brazil" county="Chapada Gaucha" latitude="-15.174723" longLatPrecision="21" longitude="-45.72139" municipality="Parque Nacional Grande Sertao Veredas" name="Cerrado" pageId="6" pageNumber="146" stateProvince="Minas Gerais">Cerrado</location>
,
<geoCoordinate id="EE3BFC86D76A911B168CFEADFE91FE2F" box="[151,289,366,393]" degrees="15" direction="south" minutes="10" orientation="latitude" pageId="6" pageNumber="146" precision="15" seconds="29" value="-15.174723">
15
<superScript id="7C7A3709D76A911B16A8FEACFF0BFEDB" attach="none" box="[179,187,367,381]" fontSize="6" pageId="6" pageNumber="146">o</superScript>
10’29” S
</geoCoordinate>
–
<geoCoordinate id="EE3BFC86D76A911B172AFEADFE78FE2F" box="[305,456,366,393]" degrees="45" direction="west" minutes="43" orientation="longitude" pageId="6" pageNumber="146" precision="15" seconds="17" value="-45.72139">
45
<superScript id="7C7A3709D76A911B1756FEACFEE5FEDB" attach="none" box="[333,341,367,381]" fontSize="6" pageId="6" pageNumber="146">o</superScript>
43’17” W
</geoCoordinate>
,
<location id="8ED0CC9AD76A911B17CDFEACFD9FFE2F" LSID="urn:lsid:plazi:treatment:03A62B57D76F911B168CFB44FB1EFC6F:8ED0CC9AD76A911B17CDFEACFD9FFE2F" box="[470,559,367,393]" country="Brazil" county="Chapada Gaucha" latitude="-15.174723" longLatPrecision="21" longitude="-45.72139" municipality="Parque Nacional Grande Sertao Veredas" name="Malaise" pageId="6" pageNumber="146" stateProvince="Minas Gerais">Malaise</location>
<quantity id="4CF737A4D76A911B1422FEACFD15FE2F" box="[569,677,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.764" pageId="6" pageNumber="146" unit="m" value="2764.0">
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,
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<collectingDate id="EFF54569D76A911B14AFFEACFCF9FE2F" box="[692,841,367,393]" pageId="6" pageNumber="146" value="2018-08-28">28.VIII.2018</collectingDate>
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,
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&amp;
<collectorName id="26FAFF97D76A911B120CFEACFB21FE2F" box="[1047,1169,367,393]" pageId="6" pageNumber="146">R.I.R Lara</collectorName>
col. (
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)
</materialsCitation>
.
<materialsCitation id="3B67901CD76A911B168CFE50FB6DFE77" collectingDate="2018-09-25" collectionCode="DCBU" country="Brazil" elevation="3764" latitude="-15.175" longLatPrecision="21" longitude="-45.72111" pageId="6" pageNumber="146" specimenCode="DCBU 445.618, DCBU 445.620, DCBU 445.621, DCBU 445.625, DCBU 473.857" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<typeStatus id="54B424E3D76A911B168CFE50FEB7FE0B" box="[151,263,403,429]" pageId="6" pageNumber="146" type="paratype">Paratypes</typeStatus>
, females. Same data (
<specimenCode id="DBA9323AD76A911B17E2FE50FD1EFE0B" box="[505,686,402,429]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="6" pageNumber="146">DCBU 445.618</specimenCode>
,
<specimenCode id="DBA9323AD76A911B14A3FE50FCDDFE0B" box="[696,877,402,429]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="6" pageNumber="146">DCBU 445.620</specimenCode>
). Same data,
<date id="FFB1BC81D76A911B121AFE51FB30FE0B" box="[1025,1152,402,429]" pageId="6" pageNumber="146" value="2018-09-25">
<collectingDate id="EFF54569D76A911B121AFE51FB30FE0B" box="[1025,1152,402,429]" pageId="6" pageNumber="146" value="2018-09-25">25.IX.2018</collectingDate>
</date>
(
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,
<specimenCode id="DBA9323AD76A911B1355FE50FF47FE77" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="6" pageNumber="146">DCBU 445.625</specimenCode>
). Same data,
<geoCoordinate id="EE3BFC86D76A911B1790FE75FDA2FE77" box="[395,530,438,465]" degrees="15" direction="south" minutes="10" orientation="latitude" pageId="6" pageNumber="146" precision="15" seconds="30" value="-15.175">
15
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10’30” S
</geoCoordinate>
–
<geoCoordinate id="EE3BFC86D76A911B143AFE75FD05FE77" box="[545,693,438,465]" degrees="45" direction="west" minutes="43" orientation="longitude" pageId="6" pageNumber="146" precision="15" seconds="16" value="-45.72111">
45
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43’16” W
</geoCoordinate>
, Malaise
<quantity id="4CF737A4D76A911B153BFE74FC35FE77" box="[800,901,439,465]" metricMagnitude="3" metricUnit="m" metricValue="3.764" pageId="6" pageNumber="146" unit="m" value="3764.0">
<elevation id="00227D72D76A911B153BFE74FC35FE77" box="[800,901,439,465]" metricMagnitude="3" metricUnit="m" metricValue="3.764" pageId="6" pageNumber="146" unit="m" value="3764.0">3, 764 m</elevation>
</quantity>
,
<date id="FFB1BC81D76A911B158AFE75FBA0FE77" box="[913,1040,438,465]" pageId="6" pageNumber="146" value="2018-09-25">
<collectingDate id="EFF54569D76A911B158AFE75FBA0FE77" box="[913,1040,438,465]" pageId="6" pageNumber="146" value="2018-09-25">25.IX.2018</collectingDate>
</date>
(
<specimenCode id="DBA9323AD76A911B1204FE74FB65FE77" box="[1055,1237,438,465]" collectionCode="DCBU" country="Brazil" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="6" pageNumber="146">DCBU 473.857</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB09A41D76A911B16DCFE18FB1EFC6F" blockId="6.[151,1437,151,969]" pageId="6" pageNumber="146">
Comments: The two new species of
<taxonomicName id="4C0FE1C2D76A911B1449FE18FD66FE52" authorityName="Zettel" authorityYear="1990" box="[594,726,475,500]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B97B4653D76A911B1449FE18FD66FE52" box="[594,726,475,500]" italics="true" pageId="6" pageNumber="146">Dentigaster</emphasis>
</taxonomicName>
now described have the first tergite protruding medio-anteriorly, similar to
<taxonomicName id="4C0FE1C2D76A911B1713FE3CFCBAFDBF" authority="Dias &amp; Penteado-Dias, 2013" authorityName="Dias &amp; Penteado-Dias" authorityYear="2013" box="[264,778,511,537]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="B97B4653D76A911B1713FE3CFE73FDBF" box="[264,451,511,537]" italics="true" pageId="6" pageNumber="146">Dentigaster lutzi</emphasis>
<bibRefCitation id="EF9EE7B0D76A911B17D1FE3CFCBAFDBF" author="Dias, M. M. &amp; Penteado-Dias, A. M." box="[458,778,511,537]" pageId="6" pageNumber="146" pagination="409 - 412" refId="ref3681" refString="Dias, M. M. &amp; Penteado-Dias, A. M. (2013) New species of the genus Dentigaster Zettel (Hymenoptera, Braconidae, Cheloninae) from Brazil. Brazilian Journal of Biology, 73 (2), 409 - 412. https: // doi. org / 10.1590 / S 1519 - 69842013000200023" type="journal article" year="2013">Dias &amp; Penteado-Dias, 2013</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0FE1C2D76A911B155BFE3CFB1BFDBF" authority="Braet, 2014" authorityName="Braet" authorityYear="2014" box="[832,1195,511,537]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="ucayali">
<emphasis id="B97B4653D76A911B155BFE3CFBADFDBF" box="[832,1053,511,537]" italics="true" pageId="6" pageNumber="146">Dentigaster ucayali</emphasis>
Braet, 2014
</taxonomicName>
.
<taxonomicName id="4C0FE1C2D76A911B12AEFE3CFAB7FDBF" authorityName="Dias &amp; Penteado-Dias" authorityYear="2013" box="[1205,1287,511,537]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="B97B4653D76A911B12AEFE3CFAB7FDBF" box="[1205,1287,511,537]" italics="true" pageId="6" pageNumber="146">D. lutzi</emphasis>
</taxonomicName>
has a bicolor pattern, with head and anterior part of mesosoma orange and other parts of mesosoma and metasoma principally dark brown to blackish. Second tergite with yellowish patch basally, that can be reduced.
<taxonomicName id="4C0FE1C2D76A911B1294FD84FB4CFDC7" authorityName="Braet" authorityYear="2014" box="[1167,1276,583,609]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="ucayali">
<emphasis id="B97B4653D76A911B1294FD84FB4CFDC7" box="[1167,1276,583,609]" italics="true" pageId="6" pageNumber="146">D.ucayali</emphasis>
</taxonomicName>
is black, with parts white as fore tibia, fore and hind tarsi and flagellomeres 1–3. Has also parts light brown to yellow, as face around eyes, medio-apical patch on first tergite and large patch on second tergite.
<taxonomicName id="4C0FE1C2D76A911B125CFD4CFB69FD0F" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[1095,1241,655,681]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="medeirosi">
<emphasis id="B97B4653D76A911B125CFD4CFB69FD0F" box="[1095,1241,655,681]" italics="true" pageId="6" pageNumber="146">D. medeirosi</emphasis>
</taxonomicName>
has head mainly black. Pronotum black and yellow or all black, in lateral view; remaining parts of mesosoma mainly orange. First tergite orange, second light yellow and black, and third black. Fore wings orange, brown and white (
<figureCitation id="133486C4D76A911B1311FD14FAC1FD57" box="[1290,1393,727,753]" captionStart="FIGURES 1–6" captionStartId="1.[152,270,1878,1902]" captionTargetBox="[151,1436,309,1855]" captionTargetId="figure-97@1.[151,1436,309,1855]" captionTargetPageId="1" captionText="FIGURES 1–6. Dentigaster medeirosi sp. nov. Holotype, female: 1, habitus, lateral; 2, habitus, dorsal. Paratypes, females: 3, head and mesosoma, lateral; 4, the same, dorsal; 5, 6, head, frontal." figureDoi="http://doi.org/10.5281/zenodo.13330840" httpUri="https://zenodo.org/record/13330840/files/figure.png" pageId="6" pageNumber="146">Figs 1, 2</figureCitation>
,
<figureCitation id="133486C4D76A911B1365FD14FA3EFD57" box="[1406,1422,727,753]" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="6" pageNumber="146">7</figureCitation>
).
<taxonomicName id="4C0FE1C2D76A911B168CFD38FE81FCB3" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[151,305,763,789]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="fernandesi">
<emphasis id="B97B4653D76A911B168CFD38FE81FCB3" box="[151,305,763,789]" italics="true" pageId="6" pageNumber="146">D. fernandesi</emphasis>
</taxonomicName>
has head orange yellow, with dark brown patch on the face and on the vertex (
<figureCitation id="133486C4D76A911B12A3FD38FABDFCB3" box="[1208,1293,763,789]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="6" pageNumber="146">Fig. 13</figureCitation>
). Mesosoma blackish brown, with orange patches variously small to large. Two orange longitudinal bands on mesoscutum (
<figureCitation id="133486C4D76A911B1374FCDCFF06FCFB" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="6" pageNumber="146">Fig. 12</figureCitation>
). Metasoma with lateral yellow patches of variable sizes (
<figureCitation id="133486C4D76A911B1543FC80FBBAFCFB" box="[856,1034,834,861]" captionStart="FIGURES 11–17" captionStartId="4.[152,270,1776,1800]" captionTargetBox="[151,1435,181,1753]" captionTargetId="figure-17@4.[151,1435,181,1753]" captionTargetPageId="4" captionText="FIGURES 11–17. Dentigaster fernandesi sp. nov. Holotype, female, 11, habitus lateral. Paratypes, females: 12, head and mesosoma, dorsal; 13, head, frontal; 14, fore and hind wings; 15, metasoma, dorsal; 16, third tergite of metasoma, lateral; 17, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330846" httpUri="https://zenodo.org/record/13330846/files/figure.png" pageId="6" pageNumber="146">Figs 11, 15–17</figureCitation>
).
<taxonomicName id="4C0FE1C2D76A911B123BFC80FB02FCFB" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[1056,1202,835,861]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="medeirosi">
<emphasis id="B97B4653D76A911B123BFC80FB02FCFB" box="[1056,1202,835,861]" italics="true" pageId="6" pageNumber="146">D. medeirosi</emphasis>
</taxonomicName>
has color similar to
<taxonomicName id="4C0FE1C2D76A911B168CFCA4FEE8FC27" authorityName="Zettel" authorityYear="1992" box="[151,344,871,897]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="erythrothorax">
<emphasis id="B97B4653D76A911B168CFCA4FEE8FC27" box="[151,344,871,897]" italics="true" pageId="6" pageNumber="146">D. erythrothorax</emphasis>
</taxonomicName>
, but has metasoma 1.42x as long as mesosoma and 3.32x as long as wide in dorsal view.
<taxonomicName id="4C0FE1C2D76A911B139AFCA4FE82FC03" authorityName="Zettel" authorityYear="1992" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="erythrothorax">
<emphasis id="B97B4653D76A911B139AFCA4FE82FC03" italics="true" pageId="6" pageNumber="146">D. erythrothorax</emphasis>
</taxonomicName>
has metasoma 1.04x as long as mesosoma and 2.29x as long as wide in dorsal view.
<taxonomicName id="4C0FE1C2D76A911B12FAFC48FAC0FC03" authorityName="Dias &amp; Penteado-Dias" authorityYear="2024" box="[1249,1392,907,933]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="medeirosi">
<emphasis id="B97B4653D76A911B12FAFC48FAC0FC03" box="[1249,1392,907,933]" italics="true" pageId="6" pageNumber="146">D. medeirosi</emphasis>
</taxonomicName>
has an angular postero-ventral rim on third tergite (
<figureCitation id="133486C4D76A911B14BCFC6CFD5CFC6F" box="[679,748,943,969]" captionStart="FIGURES 7–10" captionStartId="2.[152,270,920,944]" captionTargetBox="[151,1435,181,896]" captionTargetId="figure-17@2.[151,1435,181,896]" captionTargetPageId="2" captionText="FIGURES 7–10. Dentigaster medeirosi sp. nov. Paratypes, females: 7, fore wing; 8, metasoma, lateral; 9, third tergite of metasoma, lateral; 10, the same, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13330844" httpUri="https://zenodo.org/record/13330844/files/figure.png" pageId="6" pageNumber="146">Fig. 9</figureCitation>
), character missing in
<emphasis id="B97B4653D76A911B15F2FC6CFB1EFC6F" box="[1001,1198,943,969]" italics="true" pageId="6" pageNumber="146">
<taxonomicName id="4C0FE1C2D76A911B15F2FC6CFB1AFC6F" authorityName="Zettel" authorityYear="1992" box="[1001,1194,943,969]" class="Insecta" family="Braconidae" genus="Dentigaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="erythrothorax">D. erythrothorax</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

295
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<mods:classification id="06C805088C4982E3BDDC74D0D2E53AE6">journal article</mods:classification>
<mods:identifier id="840734D607AB546A986ADFE9A3D22592" type="CLB-Dataset">301527</mods:identifier>
<mods:identifier id="400A9626881FCBE20B64936D066C624F" type="DOI">10.11646/zootaxa.5496.1.11</mods:identifier>
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<treatment id="03AD7372FFFA3C20FF1CFA36FD12FB3E" ID-DOI="http://doi.org/10.5281/zenodo.13330793" ID-Zenodo-Dep="13330793" LSID="urn:lsid:plazi:treatment:03AD7372FFFA3C20FF1CFA36FD12FB3E" httpUri="http://treatment.plazi.org/id/03AD7372FFFA3C20FF1CFA36FD12FB3E" lastPageId="1" lastPageNumber="148" pageId="0" pageNumber="147">
<subSubSection id="C31E91EFFFFA3C21FF1CFA36FE76F9EB" box="[151,461,1531,1558]" pageId="0" pageNumber="147" type="nomenclature">
<paragraph id="8BBBC264FFFA3C21FF1CFA36FE76F9EB" blockId="0.[151,1437,1531,2025]" box="[151,461,1531,1558]" pageId="0" pageNumber="147">
<heading id="D0F37508FFFA3C21FF1CFA36FE76F9EB" bold="true" box="[151,461,1531,1558]" fontSize="11" level="1" pageId="0" pageNumber="147" reason="1">
<emphasis id="B9701E76FFFA3C21FF1CFA36FE76F9EB" bold="true" box="[151,461,1531,1558]" pageId="0" pageNumber="147">
<taxonomicName id="4C04B9E7FFFA3C21FF1CFA36FED2F9E8" authorityName="Raines &amp; Vannozzi" authorityYear="2024" box="[151,361,1531,1558]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="0" pageNumber="147" phylum="Mollusca" rank="species" species="wakense" status="sp. nov.">
<emphasis id="B9701E76FFFA3C21FF1CFA36FED2F9E8" bold="true" box="[151,361,1531,1558]" italics="true" pageId="0" pageNumber="147">Caecum wakense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A243A30DFFFA3C21FEFBFA36FE76F9EB" box="[368,461,1531,1557]" pageId="0" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31E91EFFFFA3C21FF1CF9ECFEC7F9C7" box="[151,380,1569,1593]" pageId="0" pageNumber="147" type="description">
<paragraph id="8BBBC264FFFA3C21FF1CF9ECFEC7F9C7" blockId="0.[151,1437,1531,2025]" box="[151,380,1569,1593]" pageId="0" pageNumber="147">
<figureCitation id="133FDEE1FFFA3C21FF1CF9ECFE8AF9C7" box="[151,305,1569,1593]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2041]" captionTargetBox="[271,1315,1239,1992]" captionTargetId="figure-545@1.[271,1315,1239,1992]" captionTargetPageId="1" captionText="FIGURE 1. Caecum wakense sp. nov.: A–B. Holotype, LACM 2905, C–D. Paratypes, LACM 2906." figureDoi="http://doi.org/10.5281/zenodo.13330859" httpUri="https://zenodo.org/record/13330859/files/figure.png" pageId="0" pageNumber="147">Figures 1A–D</figureCitation>
,
<figureCitation id="133FDEE1FFFA3C21FEB7F9ECFEC7F9C7" box="[316,380,1569,1593]" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="0" pageNumber="147">2A–C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31E91EFFFFA3C21FF1CF988FC5AF97F" pageId="0" pageNumber="147" type="materials_examined">
<paragraph id="8BBBC264FFFA3C21FF1CF988FC7CF9A3" blockId="0.[151,1437,1531,2025]" box="[151,967,1605,1630]" pageId="0" pageNumber="147">
<emphasis id="B9701E76FFFA3C21FF1CF988FE86F9A3" bold="true" box="[151,317,1605,1629]" inLineHeading="true" pageId="0" pageNumber="147" reason="1">Type material.</emphasis>
<materialsCitation id="3B6CC839FFFA3C21FEC8F988FDFAF9A3" box="[323,577,1605,1629]" collectionCode="LACM" pageId="0" pageNumber="147" specimenCode="LACM 2905" specimenCount="1" typeStatus="holotype">
<typeStatus id="54BF7CC6FFFA3C21FEC8F988FE12F9A3" box="[323,425,1605,1629]" pageId="0" pageNumber="147" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DBA26A1FFFFA3C21FE3FF988FDFAF9A3" box="[436,577,1605,1629]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="0" pageNumber="147" type="Museum">LACM 2905</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B6CC839FFFA3C21FDC0F98BFC78F9A3" box="[587,963,1605,1630]" collectionCode="LACM" pageId="0" pageNumber="147" specimenCode="LACM 2906, 2" specimenCount="1" typeStatus="paratype">
<typeStatus id="54BF7CC6FFFA3C21FDC0F98BFD13F9A0" box="[587,680,1606,1630]" pageId="0" pageNumber="147" type="paratype">Paratype</typeStatus>
lot,
<specimenCode id="DBA26A1FFFFA3C21FD53F988FCDFF9A3" box="[728,868,1605,1629]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="0" pageNumber="147" type="Museum">LACM 2906</specimenCode>
,
<specimenCode id="DBA26A1FFFFA3C21FCE4F988FCC7F9A3" box="[879,892,1605,1629]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="0" pageNumber="147" type="Museum">2</specimenCode>
shells
</materialsCitation>
.
</paragraph>
<paragraph id="8BBBC264FFFA3C21FF4CF9A4FC5AF97F" blockId="0.[151,1437,1531,2025]" box="[199,993,1641,1665]" pageId="0" pageNumber="147">
<materialsCitation id="3B6CC839FFFA3C21FF4CF9A4FC66F97F" box="[199,989,1641,1665]" country="Wake Island" location="Along" municipality="Atoll" pageId="0" pageNumber="147" specimenCount="1" typeStatus="holotype">
<emphasis id="B9701E76FFFA3C21FF4CF9A4FEE5F97F" bold="true" box="[199,350,1641,1665]" inLineHeading="true" pageId="0" pageNumber="147" reason="1">
<typeStatus id="54BF7CC6FFFA3C21FF4CF9A4FF44F97F" box="[199,255,1641,1665]" pageId="0" pageNumber="147">Type</typeStatus>
locality.
</emphasis>
<location id="8EDB94BFFFFA3C21FEE8F9A4FE1DF97F" LSID="urn:lsid:plazi:treatment:03AD7372FFFA3C20FF1CFA36FD12FB3E:8EDB94BFFFFA3C21FEE8F9A4FE1DF97F" box="[355,422,1641,1665]" country="Wake Island" municipality="Atoll" name="Along" pageId="0" pageNumber="147">Along</location>
the western coastline of
<collectingCountry id="F31382F4FFFA3C21FD24F9A4FD50F97F" box="[687,747,1641,1665]" name="Wake Island" pageId="0" pageNumber="147">Wake</collectingCountry>
<collectingMunicipality id="6BDF581EFFFA3C21FD7BF9A4FC92F97F" box="[752,809,1641,1665]" pageId="0" pageNumber="147">Atoll</collectingMunicipality>
, sublittoral zone
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E91EFFFFA3C21FF4CF940FC21F95B" box="[199,922,1677,1702]" pageId="0" pageNumber="147" type="etymology">
<paragraph id="8BBBC264FFFA3C21FF4CF940FC21F95B" blockId="0.[151,1437,1531,2025]" box="[199,922,1677,1702]" pageId="0" pageNumber="147">
<emphasis id="B9701E76FFFA3C21FF4CF940FEF3F95B" bold="true" box="[199,328,1677,1701]" inLineHeading="true" pageId="0" pageNumber="147" reason="1">Etymology.</emphasis>
This species is named for the
<typeStatus id="54BF7CC6FFFA3C21FD00F943FD02F958" box="[651,697,1678,1702]" pageId="0" pageNumber="147">type</typeStatus>
locality,
<collectingCountry id="F31382F4FFFA3C21FC90F940FCECF95B" box="[795,855,1677,1701]" name="Wake Island" pageId="0" pageNumber="147">Wake</collectingCountry>
Atoll.
</paragraph>
</subSubSection>
<subSubSection id="C31E91EFFFFA3C21FF4CF97CFC24F937" box="[199,927,1713,1737]" pageId="0" pageNumber="147" type="distribution">
<paragraph id="8BBBC264FFFA3C21FF4CF97CFC24F937" blockId="0.[151,1437,1531,2025]" box="[199,927,1713,1737]" pageId="0" pageNumber="147">
<emphasis id="B9701E76FFFA3C21FF4CF97CFEE1F937" bold="true" box="[199,346,1713,1737]" inLineHeading="true" pageId="0" pageNumber="147" reason="1">Distribution.</emphasis>
Currently known only from
<collectingCountry id="F31382F4FFFA3C21FD07F97CFD73F937" box="[652,712,1713,1737]" name="Wake Island" pageId="0" pageNumber="147">Wake</collectingCountry>
Atoll at
<quantity id="4CFC6F81FFFA3C21FCAEF97CFC20F937" box="[805,923,1713,1737]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="0" pageNumber="147" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
20 to
<specimenCount id="9D0209EDFFFA3C21FCEAF97CFC20F937" box="[865,923,1713,1737]" count="30" pageId="0" pageNumber="147" type="generic">30 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E91EFFFFA3C20FF4CF918FA20FF2D" lastPageId="1" lastPageNumber="148" pageId="0" pageNumber="147" type="description">
<paragraph id="8BBBC264FFFA3C20FF4CF918FA20FF2D" blockId="0.[151,1437,1531,2025]" lastBlockId="1.[151,1437,151,1216]" lastPageId="1" lastPageNumber="148" pageId="0" pageNumber="147">
<emphasis id="B9701E76FFFA3C21FF4CF918FEE9F913" bold="true" box="[199,338,1749,1773]" inLineHeading="true" pageId="0" pageNumber="147" reason="1">Description.</emphasis>
Protoconch paucispiral (about 1.2 whorls), planispiral, smooth, transition to teleoconch I marked by pronounced sinuous line (
<figureCitation id="133FDEE1FFFA3C21FE51F934FD8DF8EF" box="[474,566,1785,1809]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2041]" captionTargetBox="[271,1315,1239,1992]" captionTargetId="figure-545@1.[271,1315,1239,1992]" captionTargetPageId="1" captionText="FIGURE 1. Caecum wakense sp. nov.: A–B. Holotype, LACM 2905, C–D. Paratypes, LACM 2906." figureDoi="http://doi.org/10.5281/zenodo.13330859" httpUri="https://zenodo.org/record/13330859/files/figure.png" pageId="0" pageNumber="147">Figs 1A</figureCitation>
,
<figureCitation id="133FDEE1FFFA3C21FDCDF934FDD1F8EF" box="[582,618,1785,1809]" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="0" pageNumber="147">2A</figureCitation>
). Teleoconchs I and II short, surface smooth except for faint longitudinal microsculpture (
<figureCitation id="133FDEE1FFFA3C21FECFF8D0FE2BF8CB" box="[324,400,1821,1845]" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="0" pageNumber="147">Fig 2B</figureCitation>
), transition to teleoconch III marked by very slight axial line (
<figureCitation id="133FDEE1FFFA3C21FB91F8D0FBD4F8CB" box="[1050,1135,1821,1845]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2041]" captionTargetBox="[271,1315,1239,1992]" captionTargetId="figure-545@1.[271,1315,1239,1992]" captionTargetPageId="1" captionText="FIGURE 1. Caecum wakense sp. nov.: A–B. Holotype, LACM 2905, C–D. Paratypes, LACM 2906." figureDoi="http://doi.org/10.5281/zenodo.13330859" httpUri="https://zenodo.org/record/13330859/files/figure.png" pageId="0" pageNumber="147">Figs 1B</figureCitation>
,
<figureCitation id="133FDEE1FFFA3C21FBF1F8D0FB25F8CB" box="[1146,1182,1821,1845]" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="0" pageNumber="147">2A</figureCitation>
). Teleoconch III surface smooth except for fine longitudinal microsculpture and occasional axial growth striae, transition to teleoconch IV marked by incised axial line (
<figureCitation id="133FDEE1FFFA3C21FE0AF8A8FDB8F883" box="[385,515,1893,1917]" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="0" pageNumber="147">Figs. 2A, C</figureCitation>
). Teleoconch IV small for genus [Tol
<quantity id="4CFC6F81FFFA3C21FC2AF8A8FB9CF883" box="[929,1063,1893,1917]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="0" pageNumber="147" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
], tubular, thin, narrow, moderately and regularly arched [Larc:
<quantity id="4CFC6F81FFFA3C21FE48F844FDDAF85F" box="[451,609,1929,1953]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="0" pageNumber="147" unit="mm" value="0.5" valueMax="0.52" valueMin="0.48">0.48–0.52 mm</quantity>
; Arc:
<quantity id="4CFC6F81FFFA3C21FD29F844FCFAF85F" box="[674,833,1929,1953]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="0" pageNumber="147" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
], subcylindrical, semi-translucent (
<figureCitation id="133FDEE1FFFA3C21FB33F844FA83F85F" box="[1208,1336,1929,1953]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2041]" captionTargetBox="[271,1315,1239,1992]" captionTargetId="figure-545@1.[271,1315,1239,1992]" captionTargetPageId="1" captionText="FIGURE 1. Caecum wakense sp. nov.: A–B. Holotype, LACM 2905, C–D. Paratypes, LACM 2906." figureDoi="http://doi.org/10.5281/zenodo.13330859" httpUri="https://zenodo.org/record/13330859/files/figure.png" pageId="0" pageNumber="147">Figs. 1A–B</figureCitation>
). Surface with axial growth striae, and occasionally fine longitudinal microsculpture is visible, especially on ventral side (
<figureCitation id="133FDEE1FFFA3C21FAFAF860FF01F817" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="0" pageNumber="147">Fig. 2C</figureCitation>
). Posterior [Dpe:
<quantity id="4CFC6F81FFFA3C21FEF2F81CFDADF817" box="[377,534,2001,2025]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="0" pageNumber="147" unit="mm" value="0.235" valueMax="0.24" valueMin="0.23">0.23–0.24 mm</quantity>
] with squared shoulder (
<figureCitation id="133FDEE1FFFA3C21FCABF81CFC1BF817" box="[800,928,2001,2025]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2041]" captionTargetBox="[271,1315,1239,1992]" captionTargetId="figure-545@1.[271,1315,1239,1992]" captionTargetPageId="1" captionText="FIGURE 1. Caecum wakense sp. nov.: A–B. Holotype, LACM 2905, C–D. Paratypes, LACM 2906." figureDoi="http://doi.org/10.5281/zenodo.13330859" httpUri="https://zenodo.org/record/13330859/files/figure.png" pageId="0" pageNumber="147">Figs. 1A–B</figureCitation>
). Septum low, slightly recessed, without mucro (
<figureCitation id="133FDEE1FFFB3C20FF2BFF5AFEA4FF4E" box="[160,287,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,2017,2041]" captionTargetBox="[271,1315,1239,1992]" captionTargetId="figure-545@1.[271,1315,1239,1992]" captionTargetPageId="1" captionText="FIGURE 1. Caecum wakense sp. nov.: A–B. Holotype, LACM 2905, C–D. Paratypes, LACM 2906." figureDoi="http://doi.org/10.5281/zenodo.13330859" httpUri="https://zenodo.org/record/13330859/files/figure.png" pageId="1" pageNumber="148">Figs.1 A–B</figureCitation>
). Aperture [Da:
<quantity id="4CFC6F81FFFB3C20FE46FF5AFDD7FF51" box="[461,620,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="1" pageNumber="148" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.33–0.35 mm</quantity>
] slightly inflated, with two fine annuli. No varix. Lip smooth. Periostracum not observed. Operculum and soft animal unknown. A conceptual reconstruction of the growth stages is shown in
<figureCitation id="133FDEE1FFFB3C20FADCFF76FA2CFF2D" box="[1367,1431,187,211]" captionStart="FIGURE 2" captionStartId="2.[151,250,900,924]" captionTargetBox="[151,1436,181,876]" captionTargetId="figure-23@2.[151,1436,181,876]" captionTargetPageId="2" captionText="FIGURE 2. Conceptual reconstruction of growth stages of Caecum wakense sp. nov.: A. Protoconch to teleoconch IV. B. Surface of teleoconchs I and II. C. Surface of teleoconchs III and IV. White dots indicate transition points." figureDoi="http://doi.org/10.5281/zenodo.13330863" httpUri="https://zenodo.org/record/13330863/files/figure.png" pageId="1" pageNumber="148">Fig. 2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E91EFFFFB3C20FF4CFF12FD12FB3E" pageId="1" pageNumber="148" type="discussion">
<paragraph id="8BBBC264FFFB3C20FF4CFF12FCE9FC3A" blockId="1.[151,1437,151,1216]" pageId="1" pageNumber="148">
<emphasis id="B9701E76FFFB3C20FF4CFF12FE8DFF09" bold="true" box="[199,310,223,247]" inLineHeading="true" pageId="1" pageNumber="148" reason="1">Remarks.</emphasis>
<taxonomicName id="4C04B9E7FFFB3C20FEB6FF2DFD08FF09" authority="de Folin, 1880" authorityName="de Folin" authorityYear="1880" box="[317,691,223,247]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="succineum">
<emphasis id="B9701E76FFFB3C20FEB6FF2DFDB6FF09" box="[317,525,224,247]" italics="true" pageId="1" pageNumber="148">Caecum succineum</emphasis>
de
<bibRefCitation id="EF95BF95FFFB3C20FDBEFF12FD08FF09" author="Folin, L. de" box="[565,691,223,247]" pageId="1" pageNumber="148" pagination="806 - 812" refId="ref1588" refString="Folin, L. de (1880) On the Mollusca of H. M. S. ' Challenger' Expedition. The Caecidae, comprising the genera Parastrophia, Watsonia, and Caecum. Proceedings of the Zoological Society of London, 1879, 806 - 812. https: // doi. org / 10.1111 / j. 1096 - 3642.1879. tb 02720. x" type="book chapter" year="1880">Folin, 1880</bibRefCitation>
</taxonomicName>
, which occurs throughout the entire Indo-West Pacific, is similar due to its small size, smooth tube and low dome-shaped septum. However, it is characterized by the presence of several clear rings close to the aperture and a microsculpture of fine collabral rings due to the periostracum, while no longitudinal striation is observed (
<bibRefCitation id="EF95BF95FFFB3C20FEF2FE86FD85FE9D" author="Pizzini, M. &amp; Raines, B. &amp; Vannozzi, A." box="[377,574,331,355]" pageId="1" pageNumber="148" pagination="1 - 78" refId="ref1878" refString="Pizzini, M., Raines, B. &amp; Vannozzi, A. (2013) The family Caecidae in the South-West Pacific (Gastropoda: Rissooidea). Bollettino Malacologico, 49 (10), 1 - 78." type="journal article" year="2013">
Pizzini
<emphasis id="B9701E76FFFB3C20FE40FE81FE44FE9D" box="[459,511,331,355]" italics="true" pageId="1" pageNumber="148">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EF95BF95FFFB3C20FDCCFE86FD51FE9D" author="Vannozzi, A." box="[583,746,331,355]" pageId="1" pageNumber="148" pagination="421 - 434" refId="ref2116" refString="Vannozzi, A. (2023) The Caecidae from BIOMAGLO Expedition in the Mayotte-Glorieuses area (Mollusca, Gastropoda). Zoosystema, 45 (12), 421 - 434. https: // doi. org / 10.5252 / zoosystema 2023 v 45 a 12" type="journal article" year="2023">Vannozzi 2023</bibRefCitation>
).
<taxonomicName id="4C04B9E7FFFB3C20FD71FE86FBE1FE9D" authority="(Habe, 1978)" baseAuthorityName="Habe" baseAuthorityYear="1978" box="[762,1114,331,355]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="japonicum">
<emphasis id="B9701E76FFFB3C20FD71FE86FC73FE9D" box="[762,968,331,355]" italics="true" pageId="1" pageNumber="148">Caecum japonicum</emphasis>
(
<bibRefCitation id="EF95BF95FFFB3C20FC5EFE86FBE9FE9D" author="Habe, T." box="[981,1106,331,355]" pageId="1" pageNumber="148" pagination="1 - 5" refId="ref1750" refString="Habe, T. (1978) Four new species of Japanese Caecidae. Venus, 37 (1), 1 - 5. https: // doi. org / 10.18941 / venusjjm. 37.1 _ 1" type="journal article" year="1978">Habe, 1978</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F31382F4FFFB3C20FB12FE86FB6DFE9C" box="[1177,1238,331,354]" name="Japan" pageId="1" pageNumber="148">Japan</collectingCountry>
and the Indo-West Pacific is easily distinguished due to the color pattern, which is unusual for the family, including annular or zigzag stripes, although colorless specimens do occur (
<bibRefCitation id="EF95BF95FFFB3C20FDC0FE5EFCA1FE55" author="Pizzini, M. &amp; Raines, B. &amp; Vannozzi, A." box="[587,794,403,427]" pageId="1" pageNumber="148" pagination="1 - 78" refId="ref1878" refString="Pizzini, M., Raines, B. &amp; Vannozzi, A. (2013) The family Caecidae in the South-West Pacific (Gastropoda: Rissooidea). Bollettino Malacologico, 49 (10), 1 - 78." type="journal article" year="2013">
Pizzini
<emphasis id="B9701E76FFFB3C20FD14FE5EFD6CFE55" box="[671,727,403,427]" italics="true" pageId="1" pageNumber="148">et al.</emphasis>
2013
</bibRefCitation>
). However, it can be also distinguished due to the tapering apex, the inclination of both aperture and cutting plane, as well as the swelling in the last part of the tube preceding the aperture.
<taxonomicName id="4C04B9E7FFFB3C20FF76FE16FC86FE0C" authority="Pizzini, Raines &amp; Vannozzi, 2013" authorityName="Pizzini, Raines &amp; Vannozzi" authorityYear="2013" box="[253,829,474,498]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="dakuwaqa">
<emphasis id="B9701E76FFFB3C20FF76FE16FE71FE0C" box="[253,458,474,498]" italics="true" pageId="1" pageNumber="148">Caecum dakuwaqa</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FE5AFE17FC86FE0C" author="Pizzini, M. &amp; Raines, B. &amp; Vannozzi, A." box="[465,829,474,498]" pageId="1" pageNumber="148" pagination="1 - 78" refId="ref1878" refString="Pizzini, M., Raines, B. &amp; Vannozzi, A. (2013) The family Caecidae in the South-West Pacific (Gastropoda: Rissooidea). Bollettino Malacologico, 49 (10), 1 - 78." type="journal article" year="2013">Pizzini, Raines &amp; Vannozzi, 2013</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F31382F4FFFB3C20FC0BFE17FC13FE0C" box="[896,936,474,498]" name="Fiji" pageId="1" pageNumber="148">Fiji</collectingCountry>
,
<collectingCountry id="F31382F4FFFB3C20FC38FE16FBB1FE0C" box="[947,1034,475,498]" name="Vanuatu" pageId="1" pageNumber="148">Vanuatu</collectingCountry>
and northeastern
<collectingCountry id="F31382F4FFFB3C20FB47FE16FA27FE0C" box="[1228,1436,474,498]" name="Papua New Guinea" pageId="1" pageNumber="148">Papua New Guinea</collectingCountry>
has a bell-shaped aperture and a clearly twisted tube.
<taxonomicName id="4C04B9E7FFFB3C20FD69FE32FAE4FDE8" authority="Pizzini, Raines &amp; Vannozzi, 2013" authorityName="Pizzini, Raines &amp; Vannozzi" authorityYear="2013" box="[738,1375,510,534]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="subcylindratum">
<emphasis id="B9701E76FFFB3C20FD69FE32FC53FDE8" box="[738,1000,510,534]" italics="true" pageId="1" pageNumber="148">Caecum subcylindratum</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FC7AFE33FAE4FDE8" author="Pizzini, M. &amp; Raines, B. &amp; Vannozzi, A." box="[1009,1375,510,534]" pageId="1" pageNumber="148" pagination="1 - 78" refId="ref1878" refString="Pizzini, M., Raines, B. &amp; Vannozzi, A. (2013) The family Caecidae in the South-West Pacific (Gastropoda: Rissooidea). Bollettino Malacologico, 49 (10), 1 - 78." type="journal article" year="2013">Pizzini, Raines &amp; Vannozzi, 2013</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F31382F4FFFB3C20FF1CFDEEFF55FDC4" box="[151,238,547,570]" name="Vanuatu" pageId="1" pageNumber="148">Vanuatu</collectingCountry>
has a clearly subcylindrical teleoconch and a thick shell.
<taxonomicName id="4C04B9E7FFFB3C20FCDDFDEFFADAFDC4" authority="Pizzini, Raines &amp; Vannozzi, 2013" authorityName="Pizzini, Raines &amp; Vannozzi" authorityYear="2013" box="[854,1377,546,570]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="lapita">
<emphasis id="B9701E76FFFB3C20FCDDFDEFFC49FDC4" box="[854,1010,546,570]" italics="true" pageId="1" pageNumber="148">Caecum lapita</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FC72FDEFFADAFDC4" author="Pizzini, M. &amp; Raines, B. &amp; Vannozzi, A." box="[1017,1377,546,570]" pageId="1" pageNumber="148" pagination="1 - 78" refId="ref1878" refString="Pizzini, M., Raines, B. &amp; Vannozzi, A. (2013) The family Caecidae in the South-West Pacific (Gastropoda: Rissooidea). Bollettino Malacologico, 49 (10), 1 - 78." type="journal article" year="2013">Pizzini, Raines &amp; Vannozzi, 2013</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F31382F4FFFB3C20FF1CFD8BFF04FDA0" box="[151,191,582,606]" name="Fiji" pageId="1" pageNumber="148">Fiji</collectingCountry>
, northeastern
<collectingCountry id="F31382F4FFFB3C20FEDAFD8BFDA6FDA0" box="[337,541,582,606]" name="Papua New Guinea" pageId="1" pageNumber="148">Papua New Guinea</collectingCountry>
and north Sulawesi has a similar smooth tube and recessed septum but the aperture is bell-shaped ending with a few strong annuli.
<taxonomicName id="4C04B9E7FFFB3C20FDFEFDA7FB3EFD7C" authority="Pizzini, Raines &amp; Vannozzi, 2013" authorityName="Pizzini, Raines &amp; Vannozzi" authorityYear="2013" box="[629,1157,618,642]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="bathus">
<emphasis id="B9701E76FFFB3C20FDFEFDA7FCA1FD7C" box="[629,794,618,642]" italics="true" pageId="1" pageNumber="148">Caecum bathus</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FCABFDA7FB3EFD7C" author="Pizzini, M. &amp; Raines, B. &amp; Vannozzi, A." box="[800,1157,618,642]" pageId="1" pageNumber="148" pagination="1 - 78" refId="ref1878" refString="Pizzini, M., Raines, B. &amp; Vannozzi, A. (2013) The family Caecidae in the South-West Pacific (Gastropoda: Rissooidea). Bollettino Malacologico, 49 (10), 1 - 78." type="journal article" year="2013">Pizzini, Raines &amp; Vannozzi, 2013</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F31382F4FFFB3C20FB4DFDA7FAD6FD7C" box="[1222,1389,618,642]" name="New Caledonia" pageId="1" pageNumber="148">New Caledonia</collectingCountry>
,
<collectingCountry id="F31382F4FFFB3C20FAFCFDA7FA27FD7C" box="[1399,1436,618,642]" name="Fiji" pageId="1" pageNumber="148">Fiji</collectingCountry>
and
<collectingCountry id="F31382F4FFFB3C20FF48FD43FEC0FD58" box="[195,379,654,678]" name="French Polynesia" pageId="1" pageNumber="148">French Polynesia</collectingCountry>
has a slender shell with bell-shaped aperture and worm-like microsculpture.
<taxonomicName id="4C04B9E7FFFB3C20FB29FD43FE86FD34" authority="Vannozzi, 2019" authorityName="Vannozzi" authorityYear="2019" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="neoguineanum">
<emphasis id="B9701E76FFFB3C20FB29FD43FA27FD5B" box="[1186,1436,654,677]" italics="true" pageId="1" pageNumber="148">Caecum neoguineanum</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FF1CFD7CFE86FD34" author="Vannozzi, A." box="[151,317,689,714]" pageId="1" pageNumber="148" pagination="72 - 104" refId="ref2030" refString="Vannozzi, A. (2019) The family Caecidae (Mollusca: Gastropoda) from northern Papua-New Guinea. Bollettino Malacologico, 55 (1), 72 - 104." type="journal article" year="2019">Vannozzi, 2019</bibRefCitation>
</taxonomicName>
is different due to the more recessed septum and the aperture having a broad swelling lacking the annuli.
<taxonomicName id="4C04B9E7FFFB3C20FF1CFD1BFE62FD13" authority="Vannozzi, 2019" authorityName="Vannozzi" authorityYear="2019" box="[151,473,725,749]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="frugi">
<emphasis id="B9701E76FFFB3C20FF1CFD1BFE90FD13" box="[151,299,725,749]" italics="true" pageId="1" pageNumber="148">Caecum frugi</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FEB9FD18FE62FD13" author="Vannozzi, A." box="[306,473,725,749]" pageId="1" pageNumber="148" pagination="72 - 104" refId="ref2030" refString="Vannozzi, A. (2019) The family Caecidae (Mollusca: Gastropoda) from northern Papua-New Guinea. Bollettino Malacologico, 55 (1), 72 - 104." type="journal article" year="2019">Vannozzi, 2019</bibRefCitation>
</taxonomicName>
from northeastern
<collectingCountry id="F31382F4FFFB3C20FD21FD1BFCC1FD13" box="[682,890,725,749]" name="Papua New Guinea" pageId="1" pageNumber="148">Papua New Guinea</collectingCountry>
has a shorter tube, dome-shaped septum with low mucro and a granulated periostracum.
<taxonomicName id="4C04B9E7FFFB3C20FDA2FD37FC47FCEF" authority="Vannozzi &amp; Renda, 2024" authorityName="Vannozzi &amp; Renda" authorityYear="2024" box="[553,1020,761,785]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="restrictum">
<emphasis id="B9701E76FFFB3C20FDA2FD37FD49FCEF" box="[553,754,762,785]" italics="true" pageId="1" pageNumber="148">Caecum restrictum</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FD7DFD34FC47FCEF" author="Vannozzi, A. &amp; Renda, W." box="[758,1020,761,785]" pageId="1" pageNumber="148" pagination="327 - 347" refId="ref2164" refString="Vannozzi, A. &amp; Renda, W. (2024) Sampling the depth: New data on the Caecidae (Mollusca: Gastropoda) from northeastern Papua New Guinea. Zoosystema, 46 (14), 327 - 347. https: // doi. org / 10.5252 / zoosystema 2024 v 46 a 14" type="journal article" year="2024">Vannozzi &amp; Renda, 2024</bibRefCitation>
</taxonomicName>
has a shorter tube and a swelling before the aperture with very fine and even annuli.
<taxonomicName id="4C04B9E7FFFB3C20FDFBFCD3FBF0FCCB" authority="Vannozzi &amp; Renda, 2024" authorityName="Vannozzi &amp; Renda" authorityYear="2024" box="[624,1099,797,821]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="miliarium">
<emphasis id="B9701E76FFFB3C20FDFBFCD3FC83FCCB" box="[624,824,797,821]" italics="true" pageId="1" pageNumber="148">Caecum miliarium</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FCB5FCD0FBF0FCCB" author="Vannozzi, A. &amp; Renda, W." box="[830,1099,797,821]" pageId="1" pageNumber="148" pagination="327 - 347" refId="ref2164" refString="Vannozzi, A. &amp; Renda, W. (2024) Sampling the depth: New data on the Caecidae (Mollusca: Gastropoda) from northeastern Papua New Guinea. Zoosystema, 46 (14), 327 - 347. https: // doi. org / 10.5252 / zoosystema 2024 v 46 a 14" type="journal article" year="2024">Vannozzi &amp; Renda, 2024</bibRefCitation>
</taxonomicName>
has a similar overall aspect but has a multispiral protoconch and a microsculpture of longitudinal furrows visible throughout the teleoconch.
<taxonomicName id="4C04B9E7FFFB3C20FACEFC8CFE47FC83" authority="Vannozzi &amp; Renda, 2024" authorityName="Vannozzi &amp; Renda" authorityYear="2024" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="lozoueti">
<emphasis id="B9701E76FFFB3C20FACEFC8CFF50FC83" italics="true" pageId="1" pageNumber="148">Caecum lozoueti</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FF7AFCA8FE47FC83" author="Vannozzi, A. &amp; Renda, W." box="[241,508,869,893]" pageId="1" pageNumber="148" pagination="327 - 347" refId="ref2164" refString="Vannozzi, A. &amp; Renda, W. (2024) Sampling the depth: New data on the Caecidae (Mollusca: Gastropoda) from northeastern Papua New Guinea. Zoosystema, 46 (14), 327 - 347. https: // doi. org / 10.5252 / zoosystema 2024 v 46 a 14" type="journal article" year="2024">Vannozzi &amp; Renda, 2024</bibRefCitation>
</taxonomicName>
has dome-shaped, slightly recessed septum, a conspicuous longitudinal microsculpture and a simple aperture.
<taxonomicName id="4C04B9E7FFFB3C20FE01FC44FCDDFC5F" authority="Vannozzi &amp; Renda, 2024" authorityName="Vannozzi &amp; Renda" authorityYear="2024" box="[394,870,905,929]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="temporale">
<emphasis id="B9701E76FFFB3C20FE01FC44FDEFFC5F" box="[394,596,905,929]" italics="true" pageId="1" pageNumber="148">Caecum temporale</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FDD1FC44FCDDFC5F" author="Vannozzi, A. &amp; Renda, W." box="[602,870,905,929]" pageId="1" pageNumber="148" pagination="327 - 347" refId="ref2164" refString="Vannozzi, A. &amp; Renda, W. (2024) Sampling the depth: New data on the Caecidae (Mollusca: Gastropoda) from northeastern Papua New Guinea. Zoosystema, 46 (14), 327 - 347. https: // doi. org / 10.5252 / zoosystema 2024 v 46 a 14" type="journal article" year="2024">Vannozzi &amp; Renda, 2024</bibRefCitation>
</taxonomicName>
is shorter, with a slight but clear swelling just before the aperture and the persistence of parts of the temporary septum.
</paragraph>
<paragraph id="8BBBC264FFFB3C20FF4CFC1DFD12FB3E" blockId="1.[151,1437,151,1216]" pageId="1" pageNumber="148">
Further comparisons can be made with two species never recorded after their description.
<taxonomicName id="4C04B9E7FFFB3C20FB3EFC1CFE83FBF2" authority="de Folin, 1880" authorityName="de Folin" authorityYear="1880" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="crystallinum">
<emphasis id="B9701E76FFFB3C20FB3EFC1CFA27FC16" box="[1205,1436,976,1000]" italics="true" pageId="1" pageNumber="148">Caecum crystallinum</emphasis>
de
<bibRefCitation id="EF95BF95FFFB3C20FF31FC39FE83FBF2" author="Folin, L. de" box="[186,312,1012,1036]" pageId="1" pageNumber="148" pagination="806 - 812" refId="ref1588" refString="Folin, L. de (1880) On the Mollusca of H. M. S. ' Challenger' Expedition. The Caecidae, comprising the genera Parastrophia, Watsonia, and Caecum. Proceedings of the Zoological Society of London, 1879, 806 - 812. https: // doi. org / 10.1111 / j. 1096 - 3642.1879. tb 02720. x" type="book chapter" year="1880">Folin, 1880</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49C00C86FFFB3C20FEF4FC39FE77FBF2" box="[383,460,1012,1036]" country="United States of America" name="Hawaii" pageId="1" pageNumber="148">Hawaii</collectingRegion>
was described from a fragmentary specimen without an apex, therefore the morphology of the septum is unknown. The
<typeStatus id="54BF7CC6FFFB3C20FE64FBD5FDF7FBCE" box="[495,588,1048,1072]" pageId="1" pageNumber="148" type="holotype">holotype</typeStatus>
has a slight swelling before the aperture without annuli and a clear worm-like microsculpture throughout the tube.
<taxonomicName id="4C04B9E7FFFB3C20FDAAFBF1FC78FBAA" authority="Pizzini &amp; Raines, 2006" authorityName="Pizzini &amp; Raines" authorityYear="2006" box="[545,963,1084,1108]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="maori">
<emphasis id="B9701E76FFFB3C20FDAAFBF1FD7BFBAA" box="[545,704,1084,1108]" italics="true" pageId="1" pageNumber="148">Caecum maori</emphasis>
<bibRefCitation id="EF95BF95FFFB3C20FD43FBF1FC78FBAA" author="Pizzini, M. &amp; Raines, B." box="[712,963,1084,1108]" pageId="1" pageNumber="148" pagination="13 - 14" refId="ref1821" refString="Pizzini, M. &amp; Raines, B. (2006) Caecum maori, new name for Caecum solitarium Oliver, 1915 (non Meyer, 1886) (Caenogastropoda: Rissooidea Gray, J. E., 1847). Bollettino Malacologico, 42 (1 - 4), 13 - 14." type="journal article" year="2006">Pizzini &amp; Raines, 2006</bibRefCitation>
</taxonomicName>
described from Kermadec Islands.
<taxonomicName id="4C04B9E7FFFB3C20FACEFBF1FF6CFB86" authorityName="Pizzini &amp; Raines" authorityYear="2006" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="148" phylum="Mollusca" rank="species" species="maori">
<emphasis id="B9701E76FFFB3C20FACEFBF1FF6CFB86" italics="true" pageId="1" pageNumber="148">Caecum maori</emphasis>
</taxonomicName>
has a sculpture of fine, somewhat irregular rings, not visible in the new species. Furthermore, Oliver’s description mentions the presence of a hemispherical septum with abrupt shoulder at the junction of the shell, likely alluding at the occurrence of a mucro, absent in the new species.
</paragraph>
</subSubSection>
</treatment>
</document>

88
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@ -0,0 +1,88 @@
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species.
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Protoconch planispirally coiled. Shell minute; teleoconch slightly curved tube, smooth or crossed by collabral rings, axial ribs, or in combination. Posterior end closed by conical septum.
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<mods:identifier id="D0F4F96EE6F9204845D6593ACC7730B8" type="ZooBank">6707FE34-B41E-428D-B823-ACEAC19085BD</mods:identifier>
<mods:classification id="359247E7A4FFF5B4DBD46E9A650AF12A">journal article</mods:classification>
<mods:identifier id="A71E8C23AB0024995232CE9D2B00806F" type="DOI">10.11646/zootaxa.5493.5.6</mods:identifier>
<mods:identifier id="C6AECD173F50786D767F6222B82EF021" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="454EAC203165E9C744A0F3B077358F11" type="Zenodo-Dep">13330617</mods:identifier>
<mods:identifier id="B4238E118B0745806B7E2D00A220D42F" type="ZooBank">6707FE34-B41E-428D-B823-ACEAC19085BD</mods:identifier>
</mods:mods>
<treatment id="03CEAA3A5272FFAE90DFF95AFBE8FAAC" LSID="urn:lsid:plazi:treatment:03CEAA3A5272FFAE90DFF95AFBE8FAAC" httpUri="http://treatment.plazi.org/id/03CEAA3A5272FFAE90DFF95AFBE8FAAC" lastPageId="5" lastPageNumber="582" pageId="1" pageNumber="578">
<treatment id="03CEAA3A5272FFAE90DFF95AFBE8FAAC" ID-DOI="http://doi.org/10.5281/zenodo.13330567" ID-Zenodo-Dep="13330567" LSID="urn:lsid:plazi:treatment:03CEAA3A5272FFAE90DFF95AFBE8FAAC" httpUri="http://treatment.plazi.org/id/03CEAA3A5272FFAE90DFF95AFBE8FAAC" lastPageId="5" lastPageNumber="582" pageId="1" pageNumber="578">
<subSubSection id="C37D48A75272FFAA90DFF95AFDC4F885" box="[151,620,1789,1816]" pageId="1" pageNumber="578" type="nomenclature">
<paragraph id="8BD81B2C5272FFAA90DFF95AFDC4F885" blockId="1.[151,620,1789,1853]" box="[151,620,1789,1816]" pageId="1" pageNumber="578">
<heading id="D090AC405272FFAA90DFF95AFDC4F885" bold="true" box="[151,620,1789,1816]" fontSize="11" level="1" pageId="1" pageNumber="578" reason="1">
@ -71,7 +72,7 @@
<subSubSection id="C37D48A75272FFAA90DFF885FE88F8A0" box="[151,288,1826,1853]" pageId="1" pageNumber="578" type="description">
<paragraph id="8BD81B2C5272FFAA90DFF885FE88F8A0" blockId="1.[151,620,1789,1853]" box="[151,288,1826,1853]" pageId="1" pageNumber="578">
(
<figureCitation id="135C07A95272FFAA90D7F885FEBFF8A0" box="[159,279,1826,1853]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–15" captionStartId-0="2.[152,270,1365,1389]" captionStartId-1="3.[152,270,1727,1751]" captionTargetBox-0="[167,1430,187,1341]" captionTargetBox-1="[159,1431,190,1704]" captionTargetId-0="figure-29@2.[151,1435,181,1341]" captionTargetId-1="figure-20@3.[151,1436,181,1704]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." captionText-1="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="1" pageNumber="578">Figs. 1–15</figureCitation>
<figureCitation id="135C07A95272FFAA90D7F885FEBFF8A0" box="[159,279,1826,1853]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–15" captionStartId-0="2.[152,270,1365,1389]" captionStartId-1="3.[152,270,1727,1751]" captionTargetBox-0="[167,1430,187,1341]" captionTargetBox-1="[159,1431,190,1704]" captionTargetId-0="figure-29@2.[151,1435,181,1341]" captionTargetId-1="figure-20@3.[151,1436,181,1704]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." captionText-1="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.13330619" figureDoi-1="http://doi.org/10.5281/zenodo.13330621" httpUri-0="https://zenodo.org/record/13330619/files/figure.png" httpUri-1="https://zenodo.org/record/13330621/files/figure.png" pageId="1" pageNumber="578">Figs. 1–15</figureCitation>
)
</paragraph>
</subSubSection>
@ -81,7 +82,7 @@
Mesonotum without obvious stripes. Crossveins r-m and dm-m lighter in color compared to the longitudinal veins, without obvious dark markings on the membrane. Calypter and alula nearly hyaline, with pale grey hairs. Hind femur slightly thickened. Abdominal tergites 2–5 with narrowly darkened apical band.
</paragraph>
</subSubSection>
<caption id="DF184BA45271FFA990D0FAF2FD7EFA0C" pageId="2" pageNumber="579" startId="2.[152,270,1365,1389]" targetBox="[167,1430,187,1341]" targetPageId="2" targetType="figure">
<caption id="DF184BA45271FFA990D0FAF2FD7EFA0C" ID-DOI="http://doi.org/10.5281/zenodo.13330619" ID-Zenodo-Dep="13330619" httpUri="https://zenodo.org/record/13330619/files/figure.png" pageId="2" pageNumber="579" startId="2.[152,270,1365,1389]" targetBox="[167,1430,187,1341]" targetPageId="2" targetType="figure">
<paragraph id="8BD81B2C5271FFA990D0FAF2FD7EFA0C" blockId="2.[151,1436,1365,1426]" pageId="2" pageNumber="579">
<emphasis id="B913C73E5271FFA990D0FAF2FEEAFAF0" bold="true" box="[152,322,1365,1389]" pageId="2" pageNumber="579">FIGURES 1–4.</emphasis>
<taxonomicName id="4C6760AF5271FFA99101FAF1FDD3FAF3" authorityName="Zeng &amp; Ma &amp; Yang" authorityYear="2024" box="[329,635,1366,1390]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="wuyishanensis" status="sp. nov.">
@ -119,7 +120,7 @@ Body length
<paragraph id="8BD81B2C5271FFA9908FF98BFB98F833" blockId="2.[151,1436,1508,1966]" pageId="2" pageNumber="579">
<emphasis id="B913C73E5271FFA9908FF98BFEA3F9DB" bold="true" box="[199,267,1580,1606]" pageId="2" pageNumber="579">Male.</emphasis>
Head (
<figureCitation id="135C07A95271FFA99114F98BFE6FF9DB" box="[348,455,1580,1606]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1365,1389]" captionTargetBox="[167,1430,187,1341]" captionTargetId="figure-29@2.[151,1435,181,1341]" captionTargetPageId="2" captionText="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." pageId="2" pageNumber="579">Figs. 3–4</figureCitation>
<figureCitation id="135C07A95271FFA99114F98BFE6FF9DB" box="[348,455,1580,1606]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1365,1389]" captionTargetBox="[167,1430,187,1341]" captionTargetId="figure-29@2.[151,1435,181,1341]" captionTargetPageId="2" captionText="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330619" httpUri="https://zenodo.org/record/13330619/files/figure.png" pageId="2" pageNumber="579">Figs. 3–4</figureCitation>
) brownish yellow with pale pruinesence; ocellar triangle brown, frons brownish orange with pale dark markings, front half paler than posterior, occiput almost brown; gena pale yellow, but posteriorly slightly brownish yellow. Eyes brownish orange. Hairs and setae on head dark brown. 2 pairs of fronto-orbital setae much shorter than ocellar seta, anterior one 0.75 times as long as posterior one. Gena with 7–9 postgenal setae extended to lower gena. Antenna rounded, brownish yellow except first flagellomere orange yellow to yellow; scape with black short setae dorsoapically; pedicel with 1 black dorsoapical seta and several black ventroapical setae. First flagellomere long and oval with pale grey velvet, 0.29 times as long as head height (exact ratios present in both the
<typeStatus id="54DCA58E5271FFA990DFF88FFF51F8DF" box="[151,249,1832,1858]" pageId="2" pageNumber="579" type="holotype">holotype</typeStatus>
and
@ -130,7 +131,7 @@ and
<typeStatus id="54DCA58E5271FFA99473F8EBFB04F8FB" box="[1083,1196,1868,1894]" pageId="2" pageNumber="579" type="paratype">paratypes</typeStatus>
). Proboscis brownish yellow, with dark brown vibrissa and short thin sub-vibrissa; hairs on proboscis pale grey; palpus brownish yellow with one pair of apical setae and one pair of subapical setae nearly same in length.
</paragraph>
<caption id="DF184BA45270FFA890D0F918FACFF8D9" pageId="3" pageNumber="580" startId="3.[152,270,1727,1751]" targetBox="[159,1431,190,1704]" targetPageId="3" targetType="figure">
<caption id="DF184BA45270FFA890D0F918FACFF8D9" ID-DOI="http://doi.org/10.5281/zenodo.13330621" ID-Zenodo-Dep="13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="3" pageNumber="580" startId="3.[152,270,1727,1751]" targetBox="[159,1431,190,1704]" targetPageId="3" targetType="figure">
<paragraph id="8BD81B2C5270FFA890D0F918FACFF8D9" blockId="3.[151,1437,1727,1860]" pageId="3" pageNumber="580">
<emphasis id="B913C73E5270FFA890D0F918FEFAF94A" bold="true" box="[152,338,1727,1751]" pageId="3" pageNumber="580">FIGURES 5–15.</emphasis>
<taxonomicName id="4C6760AF5270FFA89114F967FD3AF945" authorityName="Zeng &amp; Ma &amp; Yang" authorityYear="2024" box="[348,658,1728,1752]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="wuyishanensis" status="sp. nov.">
@ -142,7 +143,7 @@ and
, male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF184BA45277FFAF90D0F99CFAA7F9EA" pageId="4" pageNumber="581" startId="4.[152,270,1595,1619]" targetBox="[190,1398,181,1572]" targetPageId="4" targetType="figure">
<caption id="DF184BA45277FFAF90D0F99CFAA7F9EA" ID-DOI="http://doi.org/10.5281/zenodo.13330625" ID-Zenodo-Dep="13330625" httpUri="https://zenodo.org/record/13330625/files/figure.png" pageId="4" pageNumber="581" startId="4.[152,270,1595,1619]" targetBox="[190,1398,181,1572]" targetPageId="4" targetType="figure">
<paragraph id="8BD81B2C5277FFAF90D0F99CFAA7F9EA" blockId="4.[151,1436,1595,1656]" pageId="4" pageNumber="581">
<emphasis id="B913C73E5277FFAF90D0F99CFEF3F9C9" bold="true" box="[152,347,1595,1620]" pageId="4" pageNumber="581">FIGURES 16–19.</emphasis>
<taxonomicName id="4C6760AF5277FFAF912AF99BFD0CF9C9" authority="Frey" authorityName="Frey" authorityYear="1934" box="[354,676,1596,1620]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="javanensis">
@ -154,7 +155,7 @@ Frey
</caption>
<paragraph id="8BD81B2C5277FFAF908FF91AFAAFF816" blockId="4.[151,1437,1725,2039]" pageId="4" pageNumber="581">
Thorax (
<figureCitation id="135C07A95277FFAF9163F91AFED9F94A" box="[299,369,1725,1751]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1365,1389]" captionTargetBox="[167,1430,187,1341]" captionTargetId="figure-29@2.[151,1435,181,1341]" captionTargetPageId="2" captionText="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." pageId="4" pageNumber="581">Fig. 2</figureCitation>
<figureCitation id="135C07A95277FFAF9163F91AFED9F94A" box="[299,369,1725,1751]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1365,1389]" captionTargetBox="[167,1430,187,1341]" captionTargetId="figure-29@2.[151,1435,181,1341]" captionTargetPageId="2" captionText="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330619" httpUri="https://zenodo.org/record/13330619/files/figure.png" pageId="4" pageNumber="581">Fig. 2</figureCitation>
) dark brown, mesonotum with pale grey pruinescence. Mesonotum without obvious stripes. Hairs and setae on thorax dark brown. Postpronotum with one pair of setae. 1 well developed postpronotal seta and 3–4 short setae; 3 postsutural dorsocentral setae, anterior one shorter than posterior two, 1
<superScript id="7C12B6645277FFAF94CCF8A2FB27F88E" attach="left" box="[1156,1167,1797,1811]" fontSize="6" pageId="4" pageNumber="581">st</superScript>
dorsocentral seta closer to 2
@ -163,22 +164,22 @@ than to suture, 4 irregular rows of acrostichal setae between them after suture;
</paragraph>
<paragraph id="8BD81B2C5277FFAF908FF832FC81F86A" blockId="4.[151,1437,1725,2039]" pageId="4" pageNumber="581">
Wing (
<figureCitation id="135C07A95277FFAF915AF832FEF9F832" box="[274,337,1941,1967]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1365,1389]" captionTargetBox="[167,1430,187,1341]" captionTargetId="figure-29@2.[151,1435,181,1341]" captionTargetPageId="2" captionText="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." pageId="4" pageNumber="581">Fig. 1</figureCitation>
<figureCitation id="135C07A95277FFAF915AF832FEF9F832" box="[274,337,1941,1967]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1365,1389]" captionTargetBox="[167,1430,187,1341]" captionTargetId="figure-29@2.[151,1435,181,1341]" captionTargetPageId="2" captionText="FIGURES 1–4. Tephrochlamys wuyishanensis sp. nov., male. Habitus, lateral view (1). Mesoscutum, dorsal view (2). Head, lateral view (3) and dorsal view (4). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330619" httpUri="https://zenodo.org/record/13330619/files/figure.png" pageId="4" pageNumber="581">Fig. 1</figureCitation>
) nearly hyaline, slightly brownish, anteriorly tinged with brown; veins brownish yellow, crossveins r-m and dm-m without obvious dark markings on the membrane; costal spines inconspicuous. Calypter and alula nearly hyaline, with pale grey hairs. Halter whitish yellow.
</paragraph>
<paragraph id="8BD81B2C5276FFAE908FFF30FF7FFE80" blockId="5.[151,1437,151,1329]" pageId="5" pageNumber="582">Legs yellowish except femora brownish with pale extreme apex, mid femur paler than fore and hind femora, all tibiae and tarsi darkened towards apex, hind tibia tinged with brownish dorsally. Hairs and setae on legs dark brown. Hind femur slightly thickened. Fore femur with a row of 6 long anterodorsal setae and a row of 4 long anteroventral setae.</paragraph>
<paragraph id="8BD81B2C5276FFAE908FFE80FB83FEF8" blockId="5.[151,1437,151,1329]" pageId="5" pageNumber="582">Abdomen brownish dorsally, but tergites 2–5 with narrowly darkened posterior band. Hairs and setae on abdomen dark brown. Tergites 2–5 with a row of longer setae at posterior margin.</paragraph>
<paragraph id="8BD81B2C5276FFAE908FFEC8FDD9FDA0" blockId="5.[151,1437,151,1329]" pageId="5" pageNumber="582">
Male genitalia (
<figureCitation id="135C07A95276FFAE913FFEC8FE59FE14" box="[375,497,366,393]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 5–11</figureCitation>
<figureCitation id="135C07A95276FFAE913FFEC8FE59FE14" box="[375,497,366,393]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 5–11</figureCitation>
): Epandrium (
<figureCitation id="135C07A95276FFAE92D9FEC8FD53FE14" box="[657,763,366,393]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 5–7</figureCitation>
<figureCitation id="135C07A95276FFAE92D9FEC8FD53FE14" box="[657,763,366,393]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 5–7</figureCitation>
) small, dark yellow, with sparse black thick setae of various lengths. Hypandrium (
<figureCitation id="135C07A95276FFAE91DBFE34FDB0FE30" box="[403,536,403,429]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 10, 11</figureCitation>
<figureCitation id="135C07A95276FFAE91DBFE34FDB0FE30" box="[403,536,403,429]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 10, 11</figureCitation>
) symmetrical. Surstylus (
<figureCitation id="135C07A95276FFAE9370FE34FC0BFE30" box="[824,931,403,429]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 8, 9</figureCitation>
<figureCitation id="135C07A95276FFAE9370FE34FC0BFE30" box="[824,931,403,429]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 8, 9</figureCitation>
) almost trapezoidal in lateral view, about 0.9 times as long as basal width, with distinctly thickened and obtuse process at apex, posterior margin slightly raised, 4 setae along central edge only near apex on outer surface, several irregular sparse rows of setae run along edge on inner surface, with central area nearly bare, and 1 seta near central base. Cercus (
<figureCitation id="135C07A95276FFAE9466FE58FB32FD84" box="[1070,1178,511,537]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 6, 7</figureCitation>
<figureCitation id="135C07A95276FFAE9466FE58FB32FD84" box="[1070,1178,511,537]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 6, 7</figureCitation>
) symmetrical, apically rounded, dark yellow, slightly transparent.
</paragraph>
<paragraph id="8BD81B2C5276FFAE908FFDE0FBCFFDFC" blockId="5.[151,1437,151,1329]" box="[199,1127,582,609]" pageId="5" pageNumber="582">
@ -191,11 +192,11 @@ Externally as in male. Body length 5.0–
</paragraph>
<paragraph id="8BD81B2C5276FFAE908FFDCCFD0EFC88" blockId="5.[151,1437,151,1329]" pageId="5" pageNumber="582">
Female terminalia: Tergite 9 (
<figureCitation id="135C07A95276FFAE9247FDCCFD3CFD18" box="[527,660,619,645]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 12–14</figureCitation>
<figureCitation id="135C07A95276FFAE9247FDCCFD3CFD18" box="[527,660,619,645]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 12–14</figureCitation>
) with 3 short setae in a triangular array. Cerci (
<figureCitation id="135C07A95276FFAE94D2FDCCFAB7FD18" box="[1178,1311,619,645]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 12–14</figureCitation>
<figureCitation id="135C07A95276FFAE94D2FDCCFAB7FD18" box="[1178,1311,619,645]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Figs. 12–14</figureCitation>
) brownish, symmetrical, separated, with consistent width from proximal end to base, rounded at apex, with sparse hairy and several long brown setae at apex. Two spermathecae, dark brown and well sclerotized (
<figureCitation id="135C07A95276FFAE943BFD14FB6EFD50" box="[1139,1222,690,717]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Fig. 15</figureCitation>
<figureCitation id="135C07A95276FFAE943BFD14FB6EFD50" box="[1139,1222,690,717]" captionStart="FIGURES 5–15" captionStartId="3.[152,270,1727,1751]" captionTargetBox="[159,1431,190,1704]" captionTargetId="figure-20@3.[151,1436,181,1704]" captionTargetPageId="3" captionText="FIGURES 5–15. Tephrochlamys wuyishanensis sp. nov., male genitalia and female terminalia. Male genitalia, epandrial complex in lateral view (5). Male genitalia, epandrium and cerci, dorsal view (6), lateral view (7). Male genitalia, outer surface (8) and inner surface (9) of left surstyus. Male genitalia, hypandrial complex in dorsal (10) and ventral (11) view. Female postabdomen, dorsal (12), ventral (13) and lateral (14) view. Spermatheca (15). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330621" httpUri="https://zenodo.org/record/13330621/files/figure.png" pageId="5" pageNumber="582">Fig. 15</figureCitation>
), one of them “8”- shaped, while the other one arranged in a triangular shape with three oblate structures; spermathecal ducts colorless, transparent, or slightly tinged with pale brown.
</paragraph>
</subSubSection>
@ -307,7 +308,7 @@ The new species resembles both
<bibRefCitation id="EFF666DD5276FFAE9303FC50FC6BFB8C" author="Frey, R." box="[843,963,1015,1041]" pageId="5" pageNumber="582" pagination="338 - 339" refId="ref5912" refString="Frey, R. (1934) Diptera Brachycera von den Sunda-Inseln und Nord-Australien. Revue Suisse de Zoologie, 41, 338 - 339. https: // doi. org / 10.5962 / bhl. part. 146005" type="journal article" year="1934">Frey, 1934</bibRefCitation>
</taxonomicName>
(
<figureCitation id="135C07A95276FFAE939AFC50FBF1FB8C" box="[978,1113,1015,1041]" captionStart="FIGURES 16–19" captionStartId="4.[152,270,1595,1619]" captionTargetBox="[190,1398,181,1572]" captionTargetId="figure-29@4.[190,1398,181,1572]" captionTargetPageId="4" captionText="FIGURES 16–19. Tephrochlamys javanensis Frey, female holotype (Naturhistorisches Museum Basel, photo taken by Sarah Müller). Habitus, lateral view (16). Head, lateral view (17) and dorsal view (18). Wing (19). Scale bar = 1.0 mm." pageId="5" pageNumber="582">Figs. 16–19</figureCitation>
<figureCitation id="135C07A95276FFAE939AFC50FBF1FB8C" box="[978,1113,1015,1041]" captionStart="FIGURES 16–19" captionStartId="4.[152,270,1595,1619]" captionTargetBox="[190,1398,181,1572]" captionTargetId="figure-29@4.[190,1398,181,1572]" captionTargetPageId="4" captionText="FIGURES 16–19. Tephrochlamys javanensis Frey, female holotype (Naturhistorisches Museum Basel, photo taken by Sarah Müller). Habitus, lateral view (16). Head, lateral view (17) and dorsal view (18). Wing (19). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330625" httpUri="https://zenodo.org/record/13330625/files/figure.png" pageId="5" pageNumber="582">Figs. 16–19</figureCitation>
) and another new species,
<taxonomicName id="4C6760AF5276FFAE95C0FC50FF7DFBA9" authorityName="Zeng &amp; Ma &amp; Yang" authorityYear="2024" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="pengi" status="sp. nov.">
<emphasis id="B913C73E5276FFAE95C0FC50FF7DFBA9" italics="true" pageId="5" pageNumber="582">T. pengi</emphasis>
@ -320,7 +321,7 @@ However, it can be distinguished from them by the following characteristics: (1)
<emphasis id="B913C73E5276FFAE954CFB98FA3DFBC5" box="[1284,1429,1087,1112]" italics="true" pageId="5" pageNumber="582">T. javanensis</emphasis>
</taxonomicName>
, the anterior pair is at most 0.25 times shorter than the posterior pair (
<figureCitation id="135C07A95276FFAE93E2FBC4FB9BFBE0" box="[938,1075,1123,1149]" captionStart="FIGURES 16–19" captionStartId="4.[152,270,1595,1619]" captionTargetBox="[190,1398,181,1572]" captionTargetId="figure-29@4.[190,1398,181,1572]" captionTargetPageId="4" captionText="FIGURES 16–19. Tephrochlamys javanensis Frey, female holotype (Naturhistorisches Museum Basel, photo taken by Sarah Müller). Habitus, lateral view (16). Head, lateral view (17) and dorsal view (18). Wing (19). Scale bar = 1.0 mm." pageId="5" pageNumber="582">Figs. 17, 18</figureCitation>
<figureCitation id="135C07A95276FFAE93E2FBC4FB9BFBE0" box="[938,1075,1123,1149]" captionStart="FIGURES 16–19" captionStartId="4.[152,270,1595,1619]" captionTargetBox="[190,1398,181,1572]" captionTargetId="figure-29@4.[190,1398,181,1572]" captionTargetPageId="4" captionText="FIGURES 16–19. Tephrochlamys javanensis Frey, female holotype (Naturhistorisches Museum Basel, photo taken by Sarah Müller). Habitus, lateral view (16). Head, lateral view (17) and dorsal view (18). Wing (19). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330625" httpUri="https://zenodo.org/record/13330625/files/figure.png" pageId="5" pageNumber="582">Figs. 17, 18</figureCitation>
)); (2) the antenna is rounded in shape (while in
<taxonomicName id="4C6760AF5276FFAE9103FB20FE0EFB3D" authorityName="Zeng &amp; Ma &amp; Yang" authorityYear="2024" box="[331,422,1159,1184]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="pengi" status="sp. nov.">
<emphasis id="B913C73E5276FFAE9103FB20FE0EFB3D" box="[331,422,1159,1184]" italics="true" pageId="5" pageNumber="582">T. pengi</emphasis>
@ -342,7 +343,7 @@ However, it can be distinguished from them by the following characteristics: (1)
, the abdomen is entirely black, dusted with greenish-gray (
<bibRefCitation id="EFF666DD5276FFAE9108FB54FE12FA90" author="Frey, R." box="[320,442,1267,1293]" pageId="5" pageNumber="582" pagination="338 - 339" refId="ref5912" refString="Frey, R. (1934) Diptera Brachycera von den Sunda-Inseln und Nord-Australien. Revue Suisse de Zoologie, 41, 338 - 339. https: // doi. org / 10.5962 / bhl. part. 146005" type="journal article" year="1934">Frey, 1934</bibRefCitation>
) (
<figureCitation id="135C07A95276FFAE919AFB54FD8CFA90" box="[466,548,1267,1293]" captionStart="FIGURES 16–19" captionStartId="4.[152,270,1595,1619]" captionTargetBox="[190,1398,181,1572]" captionTargetId="figure-29@4.[190,1398,181,1572]" captionTargetPageId="4" captionText="FIGURES 16–19. Tephrochlamys javanensis Frey, female holotype (Naturhistorisches Museum Basel, photo taken by Sarah Müller). Habitus, lateral view (16). Head, lateral view (17) and dorsal view (18). Wing (19). Scale bar = 1.0 mm." pageId="5" pageNumber="582">Fig. 16</figureCitation>
<figureCitation id="135C07A95276FFAE919AFB54FD8CFA90" box="[466,548,1267,1293]" captionStart="FIGURES 16–19" captionStartId="4.[152,270,1595,1619]" captionTargetBox="[190,1398,181,1572]" captionTargetId="figure-29@4.[190,1398,181,1572]" captionTargetPageId="4" captionText="FIGURES 16–19. Tephrochlamys javanensis Frey, female holotype (Naturhistorisches Museum Basel, photo taken by Sarah Müller). Habitus, lateral view (16). Head, lateral view (17) and dorsal view (18). Wing (19). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330625" httpUri="https://zenodo.org/record/13330625/files/figure.png" pageId="5" pageNumber="582">Fig. 16</figureCitation>
)).
</paragraph>
</subSubSection>

103
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<mods:namePart id="33759F2DE5435DF6436CC394C7BF6431">Yang, Ding</mods:namePart>
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<mods:namePart id="3E54487F2207E03F0A80E9D52C488C6B">Yang, Ding</mods:namePart>
<mods:affiliation id="70DE2C56A2F96535F1069B0D4FD4E6C4">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China</mods:affiliation>
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<subSubSection id="C37D48A75276FFAE90DFFA25FD85FA00" box="[151,557,1410,1437]" pageId="5" pageNumber="582" type="nomenclature">
<paragraph id="8BD81B2C5276FFAE90DFFA25FD85FA00" blockId="5.[151,557,1410,1473]" box="[151,557,1410,1437]" pageId="5" pageNumber="582">
<heading id="D090AC405276FFAE90DFFA25FD85FA00" bold="true" box="[151,557,1410,1437]" fontSize="11" level="1" pageId="5" pageNumber="582" reason="1">
@ -71,7 +72,7 @@
<subSubSection id="C37D48A75276FFAE90DFFA00FE86FA5C" box="[151,302,1447,1473]" pageId="5" pageNumber="582" type="description">
<paragraph id="8BD81B2C5276FFAE90DFFA00FE86FA5C" blockId="5.[151,557,1410,1473]" box="[151,302,1447,1473]" pageId="5" pageNumber="582">
(
<figureCitation id="135C07A95276FFAE90D7FA00FE8FFA5C" box="[159,295,1447,1473]" captionStart-0="FIGURES 20–24" captionStart-1="FIGURES 25–27" captionStartId-0="6.[152,270,1721,1745]" captionStartId-1="7.[152,270,1276,1300]" captionTargetBox-0="[163,1414,237,1697]" captionTargetBox-1="[151,1431,777,1253]" captionTargetId-0="figure-60@6.[151,1436,237,1698]" captionTargetId-1="figure-270@7.[151,1435,777,1253]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." captionText-1="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." pageId="5" pageNumber="582">Figs. 20–27</figureCitation>
<figureCitation id="135C07A95276FFAE90D7FA00FE8FFA5C" box="[159,295,1447,1473]" captionStart-0="FIGURES 20–24" captionStart-1="FIGURES 25–27" captionStartId-0="6.[152,270,1721,1745]" captionStartId-1="7.[152,270,1276,1300]" captionTargetBox-0="[163,1414,237,1697]" captionTargetBox-1="[151,1431,777,1253]" captionTargetId-0="figure-60@6.[151,1436,237,1698]" captionTargetId-1="figure-270@7.[151,1435,777,1253]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." captionText-1="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.13330629" figureDoi-1="http://doi.org/10.5281/zenodo.13330633" httpUri-0="https://zenodo.org/record/13330629/files/figure.png" httpUri-1="https://zenodo.org/record/13330633/files/figure.png" pageId="5" pageNumber="582">Figs. 20–27</figureCitation>
)
</paragraph>
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@ -95,10 +96,10 @@ Body length
</paragraph>
<paragraph id="8BD81B2C5276FFAD908FF960FBE2FF48" blockId="5.[151,1437,1518,2013]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="583" pageId="5" pageNumber="582">
Head (
<figureCitation id="135C07A95276FFAE915DF960FE37F97C" box="[277,415,1735,1761]" captionStart="FIGURES 20–24" captionStartId="6.[152,270,1721,1745]" captionTargetBox="[163,1414,237,1697]" captionTargetId="figure-60@6.[151,1436,237,1698]" captionTargetPageId="6" captionText="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." pageId="5" pageNumber="582">Figs. 22–23</figureCitation>
<figureCitation id="135C07A95276FFAE915DF960FE37F97C" box="[277,415,1735,1761]" captionStart="FIGURES 20–24" captionStartId="6.[152,270,1721,1745]" captionTargetBox="[163,1414,237,1697]" captionTargetId="figure-60@6.[151,1436,237,1698]" captionTargetPageId="6" captionText="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330629" httpUri="https://zenodo.org/record/13330629/files/figure.png" pageId="5" pageNumber="582">Figs. 22–23</figureCitation>
) brownish yellow with pale pruinesence; ocellar triangle brown, frons brownish orange without conspicuous dark marking, front half paler than posterior one, occiput almost brown; gena almost yellowish white, but posteriorly slightly brownish yellow. Eyes brownish orange. Hairs and setae on head dark brown. 2 fronto-orbital setae, anterior one point forward straightly and 0.68 times as long as posterior one. Gena with 5 irregular postgenal setae. Antenna brownish orange, scape with black short setae dorsoapically; pedicel with 1 black dorsoapical seta and single row of dense black ventroapical setae. First flagellomere long and oval, with brownish yellow velvet, 0.33 times as long as head height, dorsal side of first flagellomere flat (visible in lateral view); arista pubescent, brownish yellow with pale yellow base segment, 2.84 times longer than first flagellomere. Proboscis brownish yellow, with dark brown vibrissa and short and thin sub-vibrissa; hairs on proboscis pale grey; palpus brownish yellow, with four pairs of apical and subapical setae nearly same in length.
</paragraph>
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<caption id="DF184BA45275FFAD90D0F91EFCCCF968" ID-DOI="http://doi.org/10.5281/zenodo.13330629" ID-Zenodo-Dep="13330629" httpUri="https://zenodo.org/record/13330629/files/figure.png" pageId="6" pageNumber="583" startId="6.[152,270,1721,1745]" targetBox="[163,1414,237,1697]" targetPageId="6" targetType="figure">
<paragraph id="8BD81B2C5275FFAD90D0F91EFCCCF968" blockId="6.[151,1436,1721,1781]" pageId="6" pageNumber="583">
<emphasis id="B913C73E5275FFAD90D0F91EFEF2F94C" bold="true" box="[152,346,1721,1745]" pageId="6" pageNumber="583">FIGURES 20–24.</emphasis>
<taxonomicName id="4C6760AF5275FFAD9128F91EFDF7F94C" authorityName="Zeng &amp; Ma &amp; Yang" authorityYear="2024" box="[352,607,1721,1745]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="variegata" status="sp. nov.">
@ -112,7 +113,7 @@ Head (
</caption>
<paragraph id="8BD81B2C5275FFAC908FF8E0FAC6FF48" blockId="6.[151,1437,1863,2034]" lastBlockId="7.[151,1437,151,753]" lastPageId="7" lastPageNumber="584" pageId="6" pageNumber="583">
Thorax (
<figureCitation id="135C07A95275FFAD9162F8E0FED5F8FC" box="[298,381,1863,1889]" captionStart="FIGURES 20–24" captionStartId="6.[152,270,1721,1745]" captionTargetBox="[163,1414,237,1697]" captionTargetId="figure-60@6.[151,1436,237,1698]" captionTargetPageId="6" captionText="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." pageId="6" pageNumber="583">Fig. 21</figureCitation>
<figureCitation id="135C07A95275FFAD9162F8E0FED5F8FC" box="[298,381,1863,1889]" captionStart="FIGURES 20–24" captionStartId="6.[152,270,1721,1745]" captionTargetBox="[163,1414,237,1697]" captionTargetId="figure-60@6.[151,1436,237,1698]" captionTargetPageId="6" captionText="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330629" httpUri="https://zenodo.org/record/13330629/files/figure.png" pageId="6" pageNumber="583">Fig. 21</figureCitation>
) dark brown, mesonotum with pale grey pruinesence; scutellum brownish yellow with dark brown margin. Scutum with one pair of dark brown stripes in middle discontinuously near 2
<superScript id="7C12B6645275FFAD94D2F8CCFB02F8E4" attach="left" box="[1178,1194,1899,1913]" fontSize="6" pageId="6" pageNumber="583">nd</superScript>
dorsocentral seta and two pairs of short stripes on each side. Hairs and setae on thorax dark brown. Postpronotum with one pair of setae. 1 well developed postpronotal seta and more than 10 short setae; 3 postsutural dorsocentral setae missing due to damage, 1
@ -123,21 +124,21 @@ than to suture, with 6 somewhat irregular rows of acrostichal setae between them
</paragraph>
<paragraph id="8BD81B2C5274FFAC908FFF78FF41FEF8" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="584">
Wing (
<figureCitation id="135C07A95274FFAC915AFF78FECBFF64" box="[274,355,223,249]" captionStart="FIGURES 20–24" captionStartId="6.[152,270,1721,1745]" captionTargetBox="[163,1414,237,1697]" captionTargetId="figure-60@6.[151,1436,237,1698]" captionTargetPageId="6" captionText="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." pageId="7" pageNumber="584">Fig. 24</figureCitation>
<figureCitation id="135C07A95274FFAC915AFF78FECBFF64" box="[274,355,223,249]" captionStart="FIGURES 20–24" captionStartId="6.[152,270,1721,1745]" captionTargetBox="[163,1414,237,1697]" captionTargetId="figure-60@6.[151,1436,237,1698]" captionTargetPageId="6" captionText="FIGURES 20–24. Tephrochlamys variegata sp. nov., female. Habitus, lateral view (20). Mesoscutum, dorsal view (21). Head, lateral view (22) and dorsal view (23). Wing (24). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330629" httpUri="https://zenodo.org/record/13330629/files/figure.png" pageId="7" pageNumber="584">Fig. 24</figureCitation>
) nearly hyaline, slightly brownish; veins brownish yellow, crossveins r-m and dm-m with slightly brownish dark markings on the membrane, CuA, CuA+CuP and bm-m with variegated yellow brown patterns on the veins; costal spines inconspicuous. Calypter and alula nearly hyaline, with brownish yellow hairs. Halter whitish yellow.
</paragraph>
<paragraph id="8BD81B2C5274FFAC908FFEC8FA3DFE68" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="584">Legs predominantly light yellow. Fore femur brownish yellow except apex yellowish, mid femur subapically tinged with paler yellow brownish, hind femur subapically tinged with brownish yellow ventrally; tibiae pale brownish yellow, all tibiae and tarsi darkened towards apex except mid tibia. Hairs and setae on legs dark brown. Fore and hind femora slightly thickened. Fore femur with a row of 7–8 long posterodorsal setae, without anteroventral seta.</paragraph>
<paragraph id="8BD81B2C5274FFAC908FFE58FE1AFDA0" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="584">Abdomen dark brownish dorsally. Hairs and setae on abdomen dark brown. Tergites 2–5 with a row of longer setae at posterior margin.</paragraph>
<paragraph id="8BD81B2C5274FFAC908FFDE0FCF1FD6C" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="584">
Female terminalia: Tergite IX (
<figureCitation id="135C07A95274FFAC9266FDE0FD10FDFC" box="[558,696,582,609]" captionStart="FIGURES 25–27" captionStartId="7.[152,270,1276,1300]" captionTargetBox="[151,1431,777,1253]" captionTargetId="figure-270@7.[151,1435,777,1253]" captionTargetPageId="7" captionText="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." pageId="7" pageNumber="584">Figs. 25–26</figureCitation>
<figureCitation id="135C07A95274FFAC9266FDE0FD10FDFC" box="[558,696,582,609]" captionStart="FIGURES 25–27" captionStartId="7.[152,270,1276,1300]" captionTargetBox="[151,1431,777,1253]" captionTargetId="figure-270@7.[151,1435,777,1253]" captionTargetPageId="7" captionText="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330633" httpUri="https://zenodo.org/record/13330633/files/figure.png" pageId="7" pageNumber="584">Figs. 25–26</figureCitation>
) with 2 short setae. Cerci (
<figureCitation id="135C07A95274FFAC93BEFDE0FB28FDFC" box="[1014,1152,582,609]" captionStart="FIGURES 25–27" captionStartId="7.[152,270,1276,1300]" captionTargetBox="[151,1431,777,1253]" captionTargetId="figure-270@7.[151,1435,777,1253]" captionTargetPageId="7" captionText="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." pageId="7" pageNumber="584">Figs. 25–26</figureCitation>
<figureCitation id="135C07A95274FFAC93BEFDE0FB28FDFC" box="[1014,1152,582,609]" captionStart="FIGURES 25–27" captionStartId="7.[152,270,1276,1300]" captionTargetBox="[151,1431,777,1253]" captionTargetId="figure-270@7.[151,1435,777,1253]" captionTargetPageId="7" captionText="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330633" httpUri="https://zenodo.org/record/13330633/files/figure.png" pageId="7" pageNumber="584">Figs. 25–26</figureCitation>
) brownish, symmetrical, separated, rounded and slightly taper at apex, with sparse hairy and several long brown setae at apex. Two spermathecae, dark brown and well sclerotized (
<figureCitation id="135C07A95274FFAC9288FD28FCBDFD34" box="[704,789,655,681]" captionStart="FIGURES 25–27" captionStartId="7.[152,270,1276,1300]" captionTargetBox="[151,1431,777,1253]" captionTargetId="figure-270@7.[151,1435,777,1253]" captionTargetPageId="7" captionText="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." pageId="7" pageNumber="584">Fig. 27</figureCitation>
<figureCitation id="135C07A95274FFAC9288FD28FCBDFD34" box="[704,789,655,681]" captionStart="FIGURES 25–27" captionStartId="7.[152,270,1276,1300]" captionTargetBox="[151,1431,777,1253]" captionTargetId="figure-270@7.[151,1435,777,1253]" captionTargetPageId="7" captionText="FIGURES 25–27. Tephrochlamys variegata sp. nov., female postabdomen. Female terminalia in dorsal (25) and ventral (26) view. Spermatheca (27). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330633" httpUri="https://zenodo.org/record/13330633/files/figure.png" pageId="7" pageNumber="584">Fig. 27</figureCitation>
), one of them gourd-shaped, while the other arranged in triangular shape resembling Mickey Mouse's head, with smaller bottom sphere and bulging middle part; spermathecal ducts colorless, transparent, or slightly tinged with pale brown.
</paragraph>
<caption id="DF184BA45274FFAC90D0FB5BFC92FAA5" pageId="7" pageNumber="584" startId="7.[152,270,1276,1300]" targetBox="[151,1431,777,1253]" targetPageId="7" targetType="figure">
<caption id="DF184BA45274FFAC90D0FB5BFC92FAA5" ID-DOI="http://doi.org/10.5281/zenodo.13330633" ID-Zenodo-Dep="13330633" httpUri="https://zenodo.org/record/13330633/files/figure.png" pageId="7" pageNumber="584" startId="7.[152,270,1276,1300]" targetBox="[151,1431,777,1253]" targetPageId="7" targetType="figure">
<paragraph id="8BD81B2C5274FFAC90D0FB5BFC92FAA5" blockId="7.[151,1436,1276,1336]" pageId="7" pageNumber="584">
<emphasis id="B913C73E5274FFAC90D0FB5BFEF3FA89" bold="true" box="[152,347,1276,1300]" pageId="7" pageNumber="584">FIGURES 25–27.</emphasis>
<taxonomicName id="4C6760AF5274FFAC912AFB5BFDCBFA89" authorityName="Zeng &amp; Ma &amp; Yang" authorityYear="2024" box="[354,611,1276,1300]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="variegata" status="sp. nov.">

107
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(
<figureCitation id="135C07A9527AFFA290D7FF1BFE8EFF4B" box="[159,294,188,214]" captionStart-0="FIGURES 28–32" captionStart-1="FIGURES 33–39" captionStartId-0="8.[152,270,1943,1967]" captionStartId-1="9.[152,270,1789,1813]" captionTargetBox-0="[164,1410,198,1902]" captionTargetBox-1="[159,1426,865,1761]" captionTargetId-0="figure-29@8.[160,1427,181,1919]" captionTargetId-1="figure-286@9.[151,1435,850,1766]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." captionText-1="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." pageId="9" pageNumber="586">Figs. 28–39</figureCitation>
<figureCitation id="135C07A9527AFFA290D7FF1BFE8EFF4B" box="[159,294,188,214]" captionStart-0="FIGURES 28–32" captionStart-1="FIGURES 33–39" captionStartId-0="8.[152,270,1943,1967]" captionStartId-1="9.[152,270,1789,1813]" captionTargetBox-0="[164,1410,198,1902]" captionTargetBox-1="[159,1426,865,1761]" captionTargetId-0="figure-29@8.[160,1427,181,1919]" captionTargetId-1="figure-286@9.[151,1435,850,1766]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." captionText-1="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.13330636" figureDoi-1="http://doi.org/10.5281/zenodo.13330638" httpUri-0="https://zenodo.org/record/13330636/files/figure.png" httpUri-1="https://zenodo.org/record/13330638/files/figure.png" pageId="9" pageNumber="586">Figs. 28–39</figureCitation>
)
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@ -93,12 +94,12 @@ Body length 5.0 mm, wing length
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Head (
<figureCitation id="135C07A9527AFFA2915AFDA7FECCFD87" box="[274,356,512,538]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." pageId="9" pageNumber="586">Fig. 31</figureCitation>
<figureCitation id="135C07A9527AFFA2915AFDA7FECCFD87" box="[274,356,512,538]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330636" httpUri="https://zenodo.org/record/13330636/files/figure.png" pageId="9" pageNumber="586">Fig. 31</figureCitation>
) brownish yellow with pale pruinesence; ocellar triangle brown, frons brownish orange without conspicuous dark marking, front half paler than posterior one, occiput almost brown; gena almost yellowish white, but posteriorly slightly brownish yellow. Eyes brownish orange. Hairs and setae on head dark brown. 2 fronto-orbital setae and ocellar setae missing due to damage. Gena with 5–6 irregular postgenal setae. Antenna brownish orange, scape with black short setae dorsoapically; pedicel with 1 black dorsoapical seta, a circle of black ventroapical setae. First flagellomere long and oval, with pale brownish yellow velvet, 0.31 times as long as head height, dorsal side of first flagellomere flat, with slight protrusion near base before arista (visible in lateral view,
<figureCitation id="135C07A9527AFFA294BEFD7FFAE2FD6F" box="[1270,1354,728,754]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." pageId="9" pageNumber="586">Fig. 31</figureCitation>
<figureCitation id="135C07A9527AFFA294BEFD7FFAE2FD6F" box="[1270,1354,728,754]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330636" httpUri="https://zenodo.org/record/13330636/files/figure.png" pageId="9" pageNumber="586">Fig. 31</figureCitation>
); arista pubescent, brownish yellow, 3.45 times longer than first flagellomere. Proboscis pale brownish yellow with 1 strong vibrissa, hairs pale grey; palpus with 1 apical seta and 1 subapical seta nearly same in length.
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<emphasis id="B913C73E527AFFA290D0F95AFEF3F888" bold="true" box="[152,347,1789,1813]" pageId="9" pageNumber="586">FIGURES 33–39.</emphasis>
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<paragraph id="8BD81B2C5279FFA190D0F830FBEAF84E" blockId="10.[151,1436,1943,2003]" pageId="10" pageNumber="587">
<emphasis id="B913C73E5279FFA190D0F830FEC9F832" bold="true" box="[152,353,1943,1967]" pageId="10" pageNumber="587">FIGURES 40–41.</emphasis>
<taxonomicName id="4C6760AF5279FFA19126F830FD13F832" authority="Czerny" authorityName="Czerny" authorityYear="1924" box="[366,699,1943,1967]" class="Insecta" family="Heleomyzidae" genus="Tephrochlamys" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="587" phylum="Arthropoda" rank="species" species="fasciata">
@ -122,7 +123,7 @@ Czerny
</caption>
<paragraph id="8BD81B2C5278FFA0908FFF30FAA6FE14" blockId="11.[151,1437,151,1509]" pageId="11" pageNumber="588">
Thorax (
<figureCitation id="135C07A95278FFA0916EFF30FEDFFF2C" box="[294,375,151,177]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." pageId="11" pageNumber="588">Fig. 29</figureCitation>
<figureCitation id="135C07A95278FFA0916EFF30FEDFFF2C" box="[294,375,151,177]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330636" httpUri="https://zenodo.org/record/13330636/files/figure.png" pageId="11" pageNumber="588">Fig. 29</figureCitation>
) dark brown, mesonotum with pale grey pruinesence; scutellum brown yellow with dark brown margin. Scutum with one pairs of stripes in middle discontinuously on presutural scutum and two pairs of short stripes on each side. Hairs and setae on thorax dark brown. Postpronotum with one pair of setae. 1 well developed postpronotal seta and 5 short setae; 3 postsutural dorsocentral seta miss, anterior one shorter than posterior two, 1
<superScript id="7C12B6645278FFA090EDFE80FF18FEA8" attach="left" box="[165,176,295,309]" fontSize="6" pageId="11" pageNumber="588">st</superScript>
dorsocentral seta closer to 2
@ -131,20 +132,20 @@ than to suture, 4 irregular rows of acrostichal setae between them after suture;
</paragraph>
<paragraph id="8BD81B2C5278FFA0908FFE34FC96FE68" blockId="11.[151,1437,151,1509]" pageId="11" pageNumber="588">
Wing (
<figureCitation id="135C07A95278FFA09159FE34FEC8FE30" box="[273,352,403,429]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." pageId="11" pageNumber="588">Fig. 32</figureCitation>
<figureCitation id="135C07A95278FFA09159FE34FEC8FE30" box="[273,352,403,429]" captionStart="FIGURES 28–32" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[164,1410,198,1902]" captionTargetId="figure-29@8.[160,1427,181,1919]" captionTargetPageId="8" captionText="FIGURES 28–32. Tephrochlamys pengi sp. nov., male. Habitus, lateral view (28). Mesoscutum, dorsal view (29). Habitus, dorsal view (30). Head, lateral view (31). Wing (32). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13330636" httpUri="https://zenodo.org/record/13330636/files/figure.png" pageId="11" pageNumber="588">Fig. 32</figureCitation>
) nearly hyaline, slightly brownish, anteriorly tinged with brown; veins brownish yellow, crossveins r-m and dm-m with infuscate markings on the membrane; costal spines inconspicuous. Calypter and alula nearly hyaline, with brownish yellow hairs. Halter yellowish white.
</paragraph>
<paragraph id="8BD81B2C5278FFA0908FFE58FC50FD18" blockId="11.[151,1437,151,1509]" pageId="11" pageNumber="588">Legs almost yellowish. Fore femur brownish yellow except apex yellowish, mid femur subapically tinged with brownish yellow, hind femur subapically tinged with brownish yellow ventrally; all tibiae and tarsi darkened towards apex, hind tibia tinged with brown dorsally. Hairs and setae on legs dark brown. All femora slightly thickened. Fore femur with a row of 8 long anterodorsal setae, but without anteroventral seta.</paragraph>
<paragraph id="8BD81B2C5278FFA0908FFD28FB83FD50" blockId="11.[151,1437,151,1509]" pageId="11" pageNumber="588">Abdomen dark brownish dorsally, but tergites 2–5 with widely darkened posterior band. Hairs and setae on abdomen dark brown. Tergites 2–5 with a row of longer setae at posterior margin.</paragraph>
<paragraph id="8BD81B2C5278FFA0908FFD70FD79FC54" blockId="11.[151,1437,151,1509]" pageId="11" pageNumber="588">
Male genitalia (
<figureCitation id="135C07A95278FFA09137FD70FDA5FD6C" box="[383,525,727,753]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." pageId="11" pageNumber="588">Figs. 33–39</figureCitation>
<figureCitation id="135C07A95278FFA09137FD70FDA5FD6C" box="[383,525,727,753]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330638" httpUri="https://zenodo.org/record/13330638/files/figure.png" pageId="11" pageNumber="588">Figs. 33–39</figureCitation>
): Epandrium (
<figureCitation id="135C07A95278FFA092FBFD70FC96FD6C" box="[691,830,726,753]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." pageId="11" pageNumber="588">Figs. 33–35</figureCitation>
<figureCitation id="135C07A95278FFA092FBFD70FC96FD6C" box="[691,830,726,753]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330638" httpUri="https://zenodo.org/record/13330638/files/figure.png" pageId="11" pageNumber="588">Figs. 33–35</figureCitation>
) small, dark yellow, sparsely with black thick setae different in length. Hypandrium symmetrical. Surstylus (
<figureCitation id="135C07A95278FFA09352FD5CFC0AFC88" box="[794,930,763,789]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." pageId="11" pageNumber="588">Figs. 36, 37</figureCitation>
<figureCitation id="135C07A95278FFA09352FD5CFC0AFC88" box="[794,930,763,789]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330638" httpUri="https://zenodo.org/record/13330638/files/figure.png" pageId="11" pageNumber="588">Figs. 36, 37</figureCitation>
) almost trapezoid in lateral view, about 0.77 times as long as basal width, with distinctly thickened and pointed process at apex, posterior margin straight, 4 widely spaced setae near the apex towards the central area of the edge on outer surface, about 20 setae gather towards the base on one side away from the apex, with several sparse setae along the edge on inner surface, with central area nearly bare, and 2 seta in the middle. Cercus (
<figureCitation id="135C07A95278FFA0935EFC2CFC32FC38" box="[790,922,906,933]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." pageId="11" pageNumber="588">Figs. 34, 35</figureCitation>
<figureCitation id="135C07A95278FFA0935EFC2CFC32FC38" box="[790,922,906,933]" captionStart="FIGURES 33–39" captionStartId="9.[152,270,1789,1813]" captionTargetBox="[159,1426,865,1761]" captionTargetId="figure-286@9.[151,1435,850,1766]" captionTargetPageId="9" captionText="FIGURES 33–39. Tephrochlamys pengi sp. nov., male genitalia. Male genitalia, epandrial complex in lateral view (33). Male genitalia, epandrium and cerci, dorsal view (34), lateral view (35). Male genitalia, outer surface (36) and inner surface (37) of left surstyus. Male genitalia, hypandrial complex in dorsal (38) and ventral (39) view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13330638" httpUri="https://zenodo.org/record/13330638/files/figure.png" pageId="11" pageNumber="588">Figs. 34, 35</figureCitation>
) dark yellow, almost transparent, symmetrical and thin, apically narrow, with a row of long setae.
</paragraph>
<paragraph id="8BD81B2C5278FFA0908FFC74FE0AFC70" blockId="11.[151,1437,151,1509]" box="[199,418,979,1005]" pageId="11" pageNumber="588">

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species:
<taxonomicName id="4C48DE981068FFEBC500F808FD3DF878" authority="Chao, 1984" authorityName="Chao" authorityYear="1984" box="[294,643,1945,1969]" class="Insecta" family="Erebidae" genus="Euproctis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="yunnanpina">
<emphasis id="B93C79091068FFEBC500F808FDB8F878" box="[294,518,1946,1969]" italics="true" pageId="2" pageNumber="103">Euproctis yunnanpina</emphasis>
<bibRefCitation id="EFD9D8EA1068FFEBC62AF80BFD3DF878" author="Chao, C. L." box="[524,643,1945,1969]" pageId="2" pageNumber="103" pagination="449 - 456" refId="ref3726" refString="Chao, C. L. (1984) New species of the genus Euproctis Hubner from China (Lepidoptera: Lymantriidae). Acta Entomologica Sinica, 27 (4), 449 - 456. https: // doi. org / 10.16380 / j. kcxb. 1984.04.017" type="journal article" year="1984">Chao, 1984</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
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<paragraph id="8BF7A51B1069FFEAC4B1F805FED3F81D" blockId="3.[151,1436,1943,2004]" pageId="3" pageNumber="104">
<emphasis id="B93C79091069FFEAC4B1F805FEABF866" bold="true" box="[151,277,1943,1967]" pageId="3" pageNumber="104">FIGURE 1.</emphasis>
Phylogenetic tree inferred from ML analysis of
<emphasis id="B93C79091069FFEAC72CF80AFC88F866" box="[778,822,1944,1967]" italics="true" pageId="3" pageNumber="104">COI</emphasis>
gene of
<taxonomicName id="4C48DE981069FFEAC7B4F80AFC4DF879" box="[914,1011,1944,1968]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="110" phylum="Arthropoda" rank="tribe" tribe="Nygmiini">Nygmiini</taxonomicName>
species. Bootstrap (BS) values are given near the clade nodes.
</paragraph>
</caption>
<subSubSection id="C352F690106EFFEDC4B1FF04FD6EFF30" pageId="4" pageNumber="105" type="diagnosis">
<paragraph id="8BF7A51B106EFFEDC4B1FF04FD6EFF30" blockId="4.[151,1436,150,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4B1FF04FEACFF79" bold="true" box="[151,274,150,176]" pageId="4" pageNumber="105">Diagnosis.</emphasis>
The new genus differs from all other genera including the recently established
<taxonomicName id="4C48DE98106EFFEDC09AFF05FAF7FF79" box="[1212,1353,151,176]" genus="Maeoproctis" pageId="4" pageNumber="110" rank="genus">
<emphasis id="B93C7909106EFFEDC09AFF05FAF7FF79" box="[1212,1353,151,176]" italics="true" pageId="4" pageNumber="105">Maeoproctis</emphasis>
</taxonomicName>
by the genital structures having a triangular uncus, a basally broad valva that narrows towards terminal with two tooth-like processes, and an elongated tongue-shaped saccus.
</paragraph>
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<subSubSection id="C352F690106EFFEDC4E1FE90FE67FDF4" pageId="4" pageNumber="105" type="description">
<paragraph id="8BF7A51B106EFFEDC4E1FE90FF55FE88" blockId="4.[151,1436,150,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4E1FE90FEE7FED5" bold="true" box="[199,345,258,284]" pageId="4" pageNumber="105">Description.</emphasis>
<emphasis id="B93C7909106EFFEDC540FE91FE1DFED4" box="[358,419,259,285]" italics="true" pageId="4" pageNumber="105">Head</emphasis>
. Labial palpus short and porrect. Antenna bipectinate, stronger in male compared to female.
</paragraph>
<paragraph id="8BF7A51B106EFFEDC4E1FED9FCCCFE64" blockId="4.[151,1436,150,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4E1FED9FEA6FEAC" box="[199,280,331,357]" italics="true" pageId="4" pageNumber="105">Thorax</emphasis>
. Wings yellow or light yellow, with variable markings on the forewing. Wing venation: forewing with R
<subScript id="17CCA75E106EFFEDC48CFE13FF7CFE46" attach="left" box="[170,194,385,399]" fontSize="6" pageId="4" pageNumber="105">2–4</subScript>
shortly stalked with R
<subScript id="17CCA75E106EFFEDC5E3FE13FE73FE46" attach="left" box="[453,461,385,399]" fontSize="6" pageId="4" pageNumber="105">5</subScript>
, M
<subScript id="17CCA75E106EFFEDC5D3FE13FE43FE46" attach="left" box="[501,509,385,399]" fontSize="6" pageId="4" pageNumber="105">1</subScript>
arising from upper angle of cell, M
<subScript id="17CCA75E106EFFEDC7B3FE13FC23FE46" attach="left" box="[917,925,385,399]" fontSize="6" pageId="4" pageNumber="105">2</subScript>
and M
<subScript id="17CCA75E106EFFEDC7C9FE13FC49FE46" attach="left" box="[1007,1015,385,399]" fontSize="6" pageId="4" pageNumber="105">3</subScript>
originating from lower angle of cell, CuA
<subScript id="17CCA75E106EFFEDC4EAFE37FF6AFE7A" attach="left" box="[204,212,421,435]" fontSize="6" pageId="4" pageNumber="105">1</subScript>
parallel to CuA
<subScript id="17CCA75E106EFFEDC5AEFE37FE2EFE7A" attach="left" box="[392,400,421,435]" fontSize="6" pageId="4" pageNumber="105">2</subScript>
; hindwing M
<subScript id="17CCA75E106EFFEDC60EFE37FD8EFE7A" attach="left" box="[552,560,421,435]" fontSize="6" pageId="4" pageNumber="105">2</subScript>
present, Rs stalked with M
<subScript id="17CCA75E106EFFEDC745FE37FCD5FE7A" attach="left" box="[867,875,421,435]" fontSize="6" pageId="4" pageNumber="105">1</subScript>
.
</paragraph>
<paragraph id="8BF7A51B106EFFEDC4E1FE25FB5AFE3C" blockId="4.[151,1436,150,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4E1FE25FED0FE18" box="[199,366,439,465]" italics="true" pageId="4" pageNumber="105">Male genitalia</emphasis>
. Uncus nearly triangular. Valva broad at base, narrowed towards terminal, cucullus with two tooth-shaped processes. Saccus well developed, long tongue-shaped. Aedeagus stout, bent basally.
</paragraph>
<paragraph id="8BF7A51B106EFFEDC4E1FE6DFE67FDF4" blockId="4.[151,1436,150,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4E1FE6DFE31FDD0" box="[199,399,511,537]" italics="true" pageId="4" pageNumber="105">Female genitalia.</emphasis>
Papilla analis broad and robust. Ductus long, sclerotized basally. Bursa large, oval-shaped; signum sclerotized medially.
</paragraph>
</subSubSection>
<subSubSection id="C352F690106EFFEDC4E1FDD5FEFEFD4C" pageId="4" pageNumber="105" type="etymology">
<paragraph id="8BF7A51B106EFFEDC4E1FDD5FEFEFD4C" blockId="4.[151,1436,150,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4E1FDD5FEF0FDA8" bold="true" box="[199,334,583,609]" pageId="4" pageNumber="105">Etymology.</emphasis>
The new genus is named after the Latin words &quot;Auri-&quot; and &quot;Valva&quot;, which refer to the ear-like shape of valva.
</paragraph>
</subSubSection>
<subSubSection id="C352F690106EFFEDC4E1FD1CFAB7FD60" box="[199,1289,654,681]" pageId="4" pageNumber="105" type="distribution">
<paragraph id="8BF7A51B106EFFEDC4E1FD1CFAB7FD60" blockId="4.[151,1436,150,681]" box="[199,1289,654,681]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4E1FD1CFEDFFD61" bold="true" box="[199,353,654,680]" pageId="4" pageNumber="105">Distribution.</emphasis>
<collectingCountry id="F35FE58B106EFFEDC541FD1DFE14FD60" box="[359,426,655,681]" name="China" pageId="4" pageNumber="105">China</collectingCountry>
(
<collectingRegion id="498C6BF9106EFFEDC59FFD1DFDAFFD60" box="[441,529,655,681]" country="China" name="Jiangsu" pageId="4" pageNumber="105">Jiangsu</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC63BFD1DFD3AFD60" box="[541,644,655,681]" country="China" name="Zhejiang" pageId="4" pageNumber="105">Zhejiang</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC6A8FD1DFD55FD60" box="[654,747,655,681]" country="China" name="Sichuan" pageId="4" pageNumber="105">Sichuan</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC6D0FD1DFCFFFD60" box="[758,833,655,681]" country="China" name="Fujian" pageId="4" pageNumber="105">Fujian</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC76AFD1DFC27FD60" box="[844,921,655,681]" country="China" name="Hunan" pageId="4" pageNumber="105">Hunan</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC783FD1DFB95FD60" box="[933,1067,655,681]" country="China" name="Guangdong" pageId="4" pageNumber="105">Guangdong</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC013FD1DFB26FD60" box="[1077,1176,655,681]" country="China" name="Guangxi" pageId="4" pageNumber="105">Guangxi</collectingRegion>
,
<collectingRegion id="498C6BF9106EFFEDC085FD1DFB40FD60" box="[1187,1278,655,681]" country="China" name="Yunnan" pageId="4" pageNumber="105">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="B93C7909106CFFEFC4B1FAF5FE3FFA4B" bold="true" box="[151,385,1383,1410]" italics="true" pageId="6" pageNumber="107">Aurivalva conistica</emphasis>
(
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)
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(
<figureCitation id="1373B99E106CFFEFC4B9FA22FF56FA03" box="[159,232,1456,1482]" captionStart="FIGURES 2–7" captionStartId="5.[151,264,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-20@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURES 2–7. Adults. 2–3. Aurivalva yunnanpina comb. nov. (2. male; 3. female); 4–5. Aurivalva telephanes comb. nov. (4. male; 5. female); 6. Aurivalva conistica comb. nov. (male); 7. Nygmia icilia Stoll (type species of Nygmia, holotype, male, copyright from Natural History Museum in London). Scale bars = 10 mm." pageId="6" pageNumber="107">Figs 6</figureCitation>
,
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<emphasis id="B93C7909106CFFEFC4B1FA67FEE4F9C5" box="[151,346,1525,1548]" italics="true" pageId="6" pageNumber="107">Euproctis conistica</emphasis>
<bibRefCitation id="EFD9D8EA106CFFEFC547FA66FD81F9C5" author="Collenette, C. L." box="[353,575,1524,1548]" pageId="6" pageNumber="107" pagination="333 - 337" refId="ref3803" refString="Collenette, C. L. (1936) On a collection of Lymantriidae (Heterocera) from North Yunnan. Annals and Magazine of Natural History, Series 10, 17, 333 - 337. https: // doi. org / 10.1080 / 00222933608655128" type="journal article" year="1936">Collenette, 1936: 336</bibRefCitation>
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;
<treatmentCitation id="0AE9830A106CFFEFC66FFA66FD4AF9C5" author="Chao, C. L." box="[585,756,1524,1548]" page="407" pageId="6" pageNumber="107" year="2003">
<bibRefCitation id="EFD9D8EA106CFFEFC66FFA66FD4AF9C5" author="Chao, C. L." box="[585,756,1524,1548]" pageId="6" pageNumber="107" refId="ref3777" refString="Chao, C. L. (2003) Lepidoptera Lymantriidae. Fauna Sinica. Insecta 30. Science Press, Beijing, 484 pp." type="book" year="2003">Chao, 2003: 407</bibRefCitation>
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.
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</paragraph>
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<emphasis id="B93C7909106CFFEFC4B1F9A8FEFBF99D" bold="true" box="[151,325,1594,1620]" pageId="6" pageNumber="107">Redescription.</emphasis>
Male forewing length
<quantity id="4CB008FE106CFFEFC66CF9A8FD79F99C" box="[586,711,1594,1621]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="6" pageNumber="107" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
. Forewing yellow, scattered with tiny black dots from base near to submargin, denser near costa. Hindwing white. Male genitalia: Tegumen broad. Uncus triangular. Valva broad basally, cucullus with two sclerotized processes, the dorsal one larger than the ventral one. Saccus long tongue-shaped. Aedeagus clavate, curved at basal 1/3, vesica with a piece of tiny spines.
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<emphasis id="B93C7909106CFFEFC4E1F958FE0FF92D" bold="true" box="[199,433,1738,1764]" pageId="6" pageNumber="107">Material examined.</emphasis>
<materialsCitation id="3B20AF46106CFFEFC59EF959FC6AF8C1" collectingDate="2018-07-20" collectorName="Shifang Mo &amp; Zhipeng Miao." country="China" county="Weixi County" location="Tacheng Town" municipality="Diqing Tibetan Autonomous Prefecture" pageId="6" pageNumber="107" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D4E6E92106CFFEFC59EF959FE63F92C" box="[440,477,1739,1765]" count="1" pageId="6" pageNumber="107" type="male">1♂</specimenCount>
, kena village,
<location id="8E97F3C0106CFFEFC6A6F958FC99F92C" LSID="urn:lsid:plazi:treatment:03E1140D106CFFEFC4B1FAF5FD9FF8BC:8E97F3C0106CFFEFC6A6F958FC99F92C" box="[640,807,1738,1765]" country="China" county="Weixi County" municipality="Diqing Tibetan Autonomous Prefecture" name="Tacheng Town" pageId="6" pageNumber="107" stateProvince="Yunnan">Tacheng Town</location>
,
<collectingCounty id="6296DD97106CFFEFC714F958FC6EF92C" box="[818,976,1738,1765]" pageId="6" pageNumber="107">Weixi County</collectingCounty>
,
<collectingMunicipality id="6B933F61106CFFEFC7FDF958FA26F92D" box="[987,1432,1738,1765]" pageId="6" pageNumber="107">Diqing Tibetan Autonomous Prefecture</collectingMunicipality>
,
<collectingRegion id="498C6BF9106CFFEFC4B1F97DFEE3F8C1" box="[151,349,1774,1801]" country="China" name="Yunnan" pageId="6" pageNumber="107">Yunnan Province</collectingRegion>
,
<collectingCountry id="F35FE58B106CFFEFC54AF97CFE0CF8C1" box="[364,434,1774,1800]" name="China" pageId="6" pageNumber="107">China</collectingCountry>
,
<date id="FFF683DB106CFFEFC5E6F97DFDF1F8C0" box="[448,591,1775,1801]" pageId="6" pageNumber="107" value="2018-07-20">
<collectingDate id="EFB27A33106CFFEFC5E6F97DFDF1F8C0" box="[448,591,1775,1801]" pageId="6" pageNumber="107" value="2018-07-20">20-VII-2018</collectingDate>
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, leg.
<collectorName id="26BDC0CD106CFFEFC6B7F97CFCA2F8C0" box="[657,796,1774,1801]" pageId="6" pageNumber="107">Shifang Mo</collectorName>
,
<collectorName id="26BDC0CD106CFFEFC70CF97CFC6AF8C1" box="[810,980,1774,1800]" pageId="6" pageNumber="107">Zhipeng Miao.</collectorName>
</materialsCitation>
<materialsCitation id="3B20AF46106CFFEFC7F8F97DFF5EF898" collectingDate="2004-06-26" collectingDateMax="2004-06-28" collectingDateMin="2004-06-26" collectorName="Houshuai Wang." country="China" county="Diqing Tibetan Autonomous Prefecture" location="Tiger Leaping Gorge Scenic Spot" municipality="Shangri-la City" pageId="6" pageNumber="107" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D4E6E92106CFFEFC7F8F97DFBBCF8C0" box="[990,1026,1775,1801]" count="1" pageId="6" pageNumber="107" type="male">1♂</specimenCount>
,
<location id="8E97F3C0106CFFEFC034F97CFA26F8C0" LSID="urn:lsid:plazi:treatment:03E1140D106CFFEFC4B1FAF5FD9FF8BC:8E97F3C0106CFFEFC034F97CFA26F8C0" box="[1042,1432,1774,1801]" country="China" county="Diqing Tibetan Autonomous Prefecture" municipality="Shangri-la City" name="Tiger Leaping Gorge Scenic Spot" pageId="6" pageNumber="107" stateProvince="Yunnan">Tiger Leaping Gorge Scenic Spot</location>
,
<collectingMunicipality id="6B933F61106CFFEFC4B1F880FEFBF8E5" box="[151,325,1810,1836]" pageId="6" pageNumber="107">Shangri-la City</collectingMunicipality>
,
<collectingCounty id="6296DD97106CFFEFC574F880FCADF8E5" box="[338,787,1810,1837]" pageId="6" pageNumber="107">Diqing Tibetan Autonomous Prefecture</collectingCounty>
,
<collectingRegion id="498C6BF9106CFFEFC738F881FC5CF8E5" box="[798,994,1810,1837]" country="China" name="Yunnan" pageId="6" pageNumber="107">Yunnan Province</collectingRegion>
,
<collectingCountry id="F35FE58B106CFFEFC7C9F880FB8BF8E5" box="[1007,1077,1810,1836]" name="China" pageId="6" pageNumber="107">China</collectingCountry>
,
<date id="FFF683DB106CFFEFC064F880FB4DF8E5" box="[1090,1267,1810,1836]" pageId="6" pageNumber="107" value="2004-06-26" valueMax="2004-06-28" valueMin="2004-06-26">
<collectingDate id="EFB27A33106CFFEFC064F880FB4DF8E5" box="[1090,1267,1810,1836]" pageId="6" pageNumber="107" value="2004-06-26" valueMax="2004-06-28" valueMin="2004-06-26">26–28-VI-2004</collectingDate>
</date>
, leg.
<collectorName id="26BDC0CD106CFFEFC117F880FF5EF898" pageId="6" pageNumber="107">Houshuai Wang.</collectorName>
</materialsCitation>
</paragraph>
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<paragraph id="8BF7A51B106CFFEFC4E1F8C8FD9FF8BC" blockId="6.[151,1437,1594,1909]" box="[199,545,1882,1909]" pageId="6" pageNumber="107">
<emphasis id="B93C7909106CFFEFC4E1F8C8FEDFF8BD" bold="true" box="[199,353,1882,1908]" pageId="6" pageNumber="107">Distribution.</emphasis>
<collectingCountry id="F35FE58B106CFFEFC541F8C8FE14F8BD" box="[359,426,1882,1908]" name="China" pageId="6" pageNumber="107">China</collectingCountry>
(
<collectingRegion id="498C6BF9106CFFEFC59FF8C9FDABF8BC" box="[441,533,1883,1909]" country="China" name="Yunnan" pageId="6" pageNumber="107">Yunnan</collectingRegion>
).
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</document>

254
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<emphasis id="B93C7909106EFFEDC4B1FA57FE28FA29" bold="true" box="[151,406,1477,1504]" italics="true" pageId="4" pageNumber="105">Aurivalva telephanes</emphasis>
(
<bibRefCitation id="EFD9D8EA106EFFEDC580FA57FDCFFA16" author="Collenette, C. L." box="[422,625,1477,1503]" pageId="4" pageNumber="105" pagination="329 - 338" refId="ref3850" refString="Collenette, C. L. (1939) New Lymantriidae (Lepidoptera, Heterocera) from Asia and the East Indies. Annals and Magazine of Natural History, Series 11, 4 (21), 329 - 338. https: // doi. org / 10.1080 / 00222933908526997" type="journal article" year="1939">Collenette, 1939</bibRefCitation>
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(
<figureCitation id="1373B99E106EFFEDC4B9F99FFEBAF9E1" box="[159,260,1549,1576]" captionStart="FIGURES 2–7" captionStartId="5.[151,264,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-20@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURES 2–7. Adults. 2–3. Aurivalva yunnanpina comb. nov. (2. male; 3. female); 4–5. Aurivalva telephanes comb. nov. (4. male; 5. female); 6. Aurivalva conistica comb. nov. (male); 7. Nygmia icilia Stoll (type species of Nygmia, holotype, male, copyright from Natural History Museum in London). Scale bars = 10 mm." pageId="4" pageNumber="105">Figs 4–5</figureCitation>
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<taxonomicName id="4C48DE98106EFFEDC4B1F9C0FDEEF9A3" authority="Collenette, 1939: 335" authorityName="Collenette" authorityPageNumber="335" authorityYear="1939" box="[151,592,1618,1642]" class="Insecta" family="Erebidae" genus="Euproctis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="telephanes">
<emphasis id="B93C7909106EFFEDC4B1F9C0FED5F9A3" box="[151,363,1618,1642]" italics="true" pageId="4" pageNumber="105">Euproctis telephanes</emphasis>
<bibRefCitation id="EFD9D8EA106EFFEDC554F9C0FDEEF9A3" author="Collenette, C. L." box="[370,592,1618,1642]" pageId="4" pageNumber="105" pagination="329 - 338" refId="ref3850" refString="Collenette, C. L. (1939) New Lymantriidae (Lepidoptera, Heterocera) from Asia and the East Indies. Annals and Magazine of Natural History, Series 11, 4 (21), 329 - 338. https: // doi. org / 10.1080 / 00222933908526997" type="journal article" year="1939">Collenette, 1939: 335</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AE9830A106EFFEDC67CF9C0FCBBF9A3" author="Chao, C. L." box="[602,773,1618,1642]" page="391" pageId="4" pageNumber="105" year="2003">
<bibRefCitation id="EFD9D8EA106EFFEDC67CF9C0FCBBF9A3" author="Chao, C. L." box="[602,773,1618,1642]" pageId="4" pageNumber="105" refId="ref3777" refString="Chao, C. L. (2003) Lepidoptera Lymantriidae. Fauna Sinica. Insecta 30. Science Press, Beijing, 484 pp." type="book" year="2003">Chao, 2003: 391</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C352F690106EFFEDC4B1F90AFE0EF843" pageId="4" pageNumber="105" type="description">
<paragraph id="8BF7A51B106EFFEDC4B1F90AFE0EF843" blockId="4.[151,1437,1688,1931]" pageId="4" pageNumber="105">
<emphasis id="B93C7909106EFFEDC4B1F90AFEFBF97B" bold="true" box="[151,325,1688,1714]" pageId="4" pageNumber="105">Redescription.</emphasis>
Forewing length
<quantity id="4CB008FE106EFFEDC632F90AFD2AF97A" box="[532,660,1688,1715]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="4" pageNumber="105" unit="mm" value="18.0" valueMax="20.0" valueMin="16.0">16–20 mm</quantity>
. Forewing yellow, with two yellowish white median lines; anterior median line short and straight, posterior median line elbow-shaped bent. Black scales scattered from lower angle of cell to dorsum, with a black spot at upper angle of cell. Hindwing pale yellow. Male genitalia: Uncus triangular. Valva broad basally, gradually tapering towards terminal, cucullus with two processes; costa broad. Saccus long and robust, tongue-shaped.Aedeagus clavate, with a pointed tip. Female genitalia: Papilla analis broad. Apophysis anterioris longer than apophysis posterioris. Ductus long, sclerotized basally. Bursa broad, signum with a sclerotization medially.
</paragraph>
</subSubSection>
<caption id="DF37F593106FFFECC4B1F9E4FC37F91F" pageId="5" pageNumber="106" startId="5.[151,264,1654,1678]" targetBox="[151,1436,181,1630]" targetPageId="5" targetType="figure">
<paragraph id="8BF7A51B106FFFECC4B1F9E4FC37F91F" blockId="5.[151,1436,1654,1750]" pageId="5" pageNumber="106">
<emphasis id="B93C7909106FFFECC4B1F9E4FE83F947" bold="true" box="[151,317,1654,1678]" pageId="5" pageNumber="106">FIGURES 2–7.</emphasis>
Adults. 2–3.
<taxonomicName id="4C48DE98106FFFECC5EDF9E4FD11F947" authorityName="Li &amp; Xia &amp; Wang" authorityYear="2024" baseAuthorityName="Chao" baseAuthorityYear="1984" box="[459,687,1654,1678]" class="Insecta" family="Erebidae" genus="Aurivalva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="yunnanpina" status="comb. nov.">
<emphasis id="B93C7909106FFFECC5EDF9E4FD11F947" box="[459,687,1654,1678]" italics="true" pageId="5" pageNumber="106">Aurivalva yunnanpina</emphasis>
</taxonomicName>
<emphasis id="B93C7909106FFFECC691F9E4FC93F947" bold="true" box="[695,813,1654,1678]" pageId="5" pageNumber="106">
<taxonomicNameLabel id="A20FC472106FFFECC691F9E4FC93F947" box="[695,813,1654,1678]" pageId="5" pageNumber="106" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(2. male; 3. female); 4–5.
<taxonomicName id="4C48DE98106FFFECC060F9E4FAA0F947" authorityName="Li &amp; Xia &amp; Wang" authorityYear="2024" baseAuthorityName="Collenette" baseAuthorityYear="1939" box="[1094,1310,1654,1678]" class="Insecta" family="Erebidae" genus="Aurivalva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="telephanes" status="comb. nov.">
<emphasis id="B93C7909106FFFECC060F9E4FAA0F947" box="[1094,1310,1654,1678]" italics="true" pageId="5" pageNumber="106">Aurivalva telephanes</emphasis>
</taxonomicName>
<emphasis id="B93C7909106FFFECC100F9E4FA22F947" bold="true" box="[1318,1436,1654,1678]" pageId="5" pageNumber="106">
<taxonomicNameLabel id="A20FC472106FFFECC100F9E4FA22F947" box="[1318,1436,1654,1678]" pageId="5" pageNumber="106" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(4. male; 5. female); 6.
<taxonomicName id="4C48DE98106FFFECC5A0F908FDF2F97B" authorityName="Li &amp; Xia &amp; Wang" authorityYear="2024" baseAuthorityName="Collenette" baseAuthorityYear="1936" box="[390,588,1690,1714]" class="Insecta" family="Erebidae" genus="Aurivalva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="conistica" status="comb. nov.">
<emphasis id="B93C7909106FFFECC5A0F908FDF2F97B" box="[390,588,1690,1714]" italics="true" pageId="5" pageNumber="106">Aurivalva conistica</emphasis>
</taxonomicName>
<emphasis id="B93C7909106FFFECC675F908FD79F97B" bold="true" box="[595,711,1690,1714]" pageId="5" pageNumber="106">
<taxonomicNameLabel id="A20FC472106FFFECC675F908FD79F97B" box="[595,711,1690,1714]" pageId="5" pageNumber="106" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(male); 7.
<taxonomicName id="4C48DE98106FFFECC711F909FC49F97B" authority="Stoll" authorityName="Stoll" authorityYear="1790" box="[823,1015,1690,1714]" class="Insecta" family="Erebidae" genus="Nygmia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="icilia">
<emphasis id="B93C7909106FFFECC711F909FC7EF97B" box="[823,960,1690,1714]" italics="true" pageId="5" pageNumber="106">Nygmia icilia</emphasis>
Stoll
</taxonomicName>
(type species of
<taxonomicName id="4C48DE98106FFFECC085F909FB4CF97B" authorityName="Hubner" authorityYear="1822" box="[1187,1266,1691,1714]" class="Insecta" family="Erebidae" genus="Nygmia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="B93C7909106FFFECC085F909FB4CF97B" box="[1187,1266,1691,1714]" italics="true" pageId="5" pageNumber="106">Nygmia</emphasis>
</taxonomicName>
, holotype, male, copyright from Natural History Museum in London). Scale bars = 10 mm.
</paragraph>
</caption>
<subSubSection id="C352F690106FFFECC4E1F8BAFC1EF81A" pageId="5" pageNumber="106" type="materials_examined">
<paragraph id="8BF7A51B106FFFECC4E1F8BAFC1EF81A" blockId="5.[151,1437,1832,2038]" pageId="5" pageNumber="106">
<emphasis id="B93C7909106FFFECC4E1F8BAFE0CF88B" bold="true" box="[199,434,1832,1858]" pageId="5" pageNumber="106">Material examined.</emphasis>
<materialsCitation id="3B20AF46106FFFECC59CF8BAFE76F8AE" collectingDate="2022-08-17" collectorName="Houshuai Wang." country="China" county="Wuyishan City" location="Sangan" municipality="Wuyishan National Park" pageId="5" pageNumber="106" specimenCount="2" specimenCount-female="2" stateProvince="Fujian">
<specimenCount id="9D4E6E92106FFFECC59CF8BAFE61F88B" box="[442,479,1832,1858]" count="2" pageId="5" pageNumber="106" type="female">2♀</specimenCount>
,
<location id="8E97F3C0106FFFECC5CBF8BAFDFCF88B" LSID="urn:lsid:plazi:treatment:03E1140D106EFFECC4B1FA57FC5EF83F:8E97F3C0106FFFECC5CBF8BAFDFCF88B" box="[493,578,1832,1858]" country="China" county="Wuyishan City" municipality="Wuyishan National Park" name="Sangan" pageId="5" pageNumber="106" stateProvince="Fujian">Sangan</location>
,
<collectingMunicipality id="6B933F61106FFFECC668F8BAFCD8F88B" box="[590,870,1832,1858]" pageId="5" pageNumber="106">Wuyishan National Park</collectingMunicipality>
,
<collectingCounty id="6296DD97106FFFECC754F8BAFBA2F88B" box="[882,1052,1832,1858]" pageId="5" pageNumber="106">Wuyishan City</collectingCounty>
,
<collectingRegion id="498C6BF9106FFFECC00FF8BAFB61F88B" box="[1065,1247,1832,1858]" country="China" name="Fujian" pageId="5" pageNumber="106">Fujian Province</collectingRegion>
,
<collectingCountry id="F35FE58B106FFFECC0CAF8BAFA8CF88B" box="[1260,1330,1832,1858]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
,
<date id="FFF683DB106FFFECC119F8BAFF6CF8AF" pageId="5" pageNumber="106" value="2022-08-17">
<collectingDate id="EFB27A33106FFFECC119F8BAFF6CF8AF" pageId="5" pageNumber="106" value="2022-08-17">17-VIII-2022</collectingDate>
</date>
, leg.
<collectorName id="26BDC0CD106FFFECC52BF8DEFE76F8AE" box="[269,456,1868,1895]" pageId="5" pageNumber="106">Houshuai Wang.</collectorName>
</materialsCitation>
<materialsCitation id="3B20AF46106FFFECC5E8F8DEFE34F843" collectingDate="2021-04-29" collectingDateMax="2021-04-30" collectingDateMin="2021-04-29" collectorName="Min Wang &amp; Yulong Zhang." country="China" location="Tongmu village" municipality="Wuyishan City" pageId="5" pageNumber="106" specimenCount="1" specimenCount-male="1" stateProvince="Fujian">
<specimenCount id="9D4E6E92106FFFECC5E8F8DEFE4CF8AF" box="[462,498,1868,1894]" count="1" pageId="5" pageNumber="106" type="male">1♂</specimenCount>
,
<location id="8E97F3C0106FFFECC5D8F8DFFD0FF8AF" LSID="urn:lsid:plazi:treatment:03E1140D106EFFECC4B1FA57FC5EF83F:8E97F3C0106FFFECC5D8F8DFFD0FF8AF" box="[510,689,1868,1895]" country="China" municipality="Wuyishan City" name="Tongmu village" pageId="5" pageNumber="106" stateProvince="Fujian">Tongmu village</location>
,
<collectingMunicipality id="6B933F61106FFFECC69AF8DEFCDBF8AF" box="[700,869,1868,1894]" pageId="5" pageNumber="106">Wuyishan City</collectingMunicipality>
,
<collectingRegion id="498C6BF9106FFFECC756F8DEFB9BF8AF" box="[880,1061,1868,1894]" country="China" name="Fujian" pageId="5" pageNumber="106">Fujian Province</collectingRegion>
,
<collectingCountry id="F35FE58B106FFFECC016F8DEFBC8F8AF" box="[1072,1142,1868,1894]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
,
<date id="FFF683DB106FFFECC0A7F8DEFA8CF8AF" box="[1153,1330,1868,1894]" pageId="5" pageNumber="106" value="2021-04-29" valueMax="2021-04-30" valueMin="2021-04-29">
<collectingDate id="EFB27A33106FFFECC0A7F8DEFA8CF8AF" box="[1153,1330,1868,1894]" pageId="5" pageNumber="106" value="2021-04-29" valueMax="2021-04-30" valueMin="2021-04-29">29–30-IV-2021</collectingDate>
</date>
, leg.
<collectorName id="26BDC0CD106FFFECC148F8DEFF65F842" pageId="5" pageNumber="106">Min Wang</collectorName>
,
<collectorName id="26BDC0CD106FFFECC4C2F8E2FE34F843" box="[228,394,1904,1930]" pageId="5" pageNumber="106">Yulong Zhang.</collectorName>
</materialsCitation>
<materialsCitation id="3B20AF46106FFFECC5A9F8E2FD1FF866" collectingDate="2004-07-22" collectingDateMax="2004-07-25" collectingDateMin="2004-07-22" collectorName="Houshuai Wang." country="China" county="Ruyuan County" location="Nanling National Nature Reserve" municipality="Shaoguan City" pageId="5" pageNumber="106" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong">
<specimenCount id="9D4E6E92106FFFECC5A9F8E2FE0DF843" box="[399,435,1904,1930]" count="1" pageId="5" pageNumber="106" type="male">1♂</specimenCount>
,
<location id="8E97F3C0106FFFECC599F8E2FC8BF842" LSID="urn:lsid:plazi:treatment:03E1140D106EFFECC4B1FA57FC5EF83F:8E97F3C0106FFFECC599F8E2FC8BF842" box="[447,821,1904,1931]" country="China" county="Ruyuan County" municipality="Shaoguan City" name="Nanling National Nature Reserve" pageId="5" pageNumber="106" stateProvince="Guangdong">Nanling National Nature Reserve</location>
,
<collectingCounty id="6296DD97106FFFECC719F8E3FC4EF843" box="[831,1008,1904,1931]" pageId="5" pageNumber="106">Ruyuan County</collectingCounty>
,
<collectingMunicipality id="6B933F61106FFFECC7DFF8E2FB21F843" box="[1017,1183,1904,1930]" pageId="5" pageNumber="106">Shaoguan City</collectingMunicipality>
,
<collectingRegion id="498C6BF9106FFFECC08FF8E2FA26F843" box="[1193,1432,1904,1930]" country="China" name="Guangdong" pageId="5" pageNumber="106">Guangdong Province</collectingRegion>
,
<collectingCountry id="F35FE58B106FFFECC4B1F806FF63F867" box="[151,221,1940,1966]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
,
<date id="FFF683DB106FFFECC4CDF806FE1AF866" box="[235,420,1940,1967]" pageId="5" pageNumber="106" value="2004-07-22" valueMax="2004-07-25" valueMin="2004-07-22">
<collectingDate id="EFB27A33106FFFECC4CDF806FE1AF866" box="[235,420,1940,1967]" pageId="5" pageNumber="106" value="2004-07-22" valueMax="2004-07-25" valueMin="2004-07-22">22–25-VII-2004</collectingDate>
</date>
, leg.
<collectorName id="26BDC0CD106FFFECC5C3F806FD1FF866" box="[485,673,1940,1967]" pageId="5" pageNumber="106">Houshuai Wang.</collectorName>
</materialsCitation>
<materialsCitation id="3B20AF46106FFFECC68CF806FC1EF81A" collectingDate="2009-08-07" collectorName="Houshuai Wang." country="China" county="Ruyuan County" location="Nanling National Nature Reserve" municipality="Shaoguan City" pageId="5" pageNumber="106" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong">
<specimenCount id="9D4E6E92106FFFECC68CF806FD73F867" box="[682,717,1940,1966]" count="1" pageId="5" pageNumber="106" type="female">1♀</specimenCount>
,
<location id="8E97F3C0106FFFECC6F8F806FBE0F866" LSID="urn:lsid:plazi:treatment:03E1140D106EFFECC4B1FA57FC5EF83F:8E97F3C0106FFFECC6F8F806FBE0F866" box="[734,1118,1940,1967]" country="China" county="Ruyuan County" municipality="Shaoguan City" name="Nanling National Nature Reserve" pageId="5" pageNumber="106" stateProvince="Guangdong">Nanling National Nature Reserve</location>
,
<collectingCounty id="6296DD97106FFFECC04AF807FA9FF867" box="[1132,1313,1940,1967]" pageId="5" pageNumber="106">Ruyuan County</collectingCounty>
,
<collectingMunicipality id="6B933F61106FFFECC108F806FF76F81B" pageId="5" pageNumber="106">Shaoguan City</collectingMunicipality>
,
<collectingRegion id="498C6BF9106FFFECC4F2F82AFE7DF81B" box="[212,451,1976,2002]" country="China" name="Guangdong" pageId="5" pageNumber="106">Guangdong Province</collectingRegion>
,
<collectingCountry id="F35FE58B106FFFECC5E9F82AFDABF81B" box="[463,533,1976,2002]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
,
<date id="FFF683DB106FFFECC606F82AFD14F81B" box="[544,682,1976,2002]" pageId="5" pageNumber="106" value="2009-08-07">
<collectingDate id="EFB27A33106FFFECC606F82AFD14F81B" box="[544,682,1976,2002]" pageId="5" pageNumber="106" value="2009-08-07">7-VIII-2009</collectingDate>
</date>
, leg.
<collectorName id="26BDC0CD106FFFECC6C3F82AFC1EF81A" box="[741,928,1976,2003]" pageId="5" pageNumber="106">Houshuai Wang.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C352F690106FFFECC4E1F84EFC5EF83F" box="[199,992,2012,2038]" pageId="5" pageNumber="106" type="distribution">
<paragraph id="8BF7A51B106FFFECC4E1F84EFC5EF83F" blockId="5.[151,1437,1832,2038]" box="[199,992,2012,2038]" pageId="5" pageNumber="106">
<emphasis id="B93C7909106FFFECC4E1F84EFEDFF83F" bold="true" box="[199,353,2012,2038]" pageId="5" pageNumber="106">Distribution.</emphasis>
<collectingCountry id="F35FE58B106FFFECC541F84EFE14F83F" box="[359,426,2012,2038]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
(
<collectingRegion id="498C6BF9106FFFECC59FF84EFDAFF83F" box="[441,529,2012,2038]" country="China" name="Jiangsu" pageId="5" pageNumber="106">Jiangsu</collectingRegion>
,
<collectingRegion id="498C6BF9106FFFECC63BF84EFD3AF83F" box="[541,644,2012,2038]" country="China" name="Zhejiang" pageId="5" pageNumber="106">Zhejiang</collectingRegion>
,
<collectingRegion id="498C6BF9106FFFECC6A8F84EFD55F83F" box="[654,747,2012,2038]" country="China" name="Sichuan" pageId="5" pageNumber="106">Sichuan</collectingRegion>
,
<collectingRegion id="498C6BF9106FFFECC6D0F84EFCFFF83F" box="[758,833,2012,2038]" country="China" name="Fujian" pageId="5" pageNumber="106">Fujian</collectingRegion>
,
<collectingRegion id="498C6BF9106FFFECC76AF84EFC6AF83F" box="[844,980,2012,2038]" country="China" name="Guangdong" pageId="5" pageNumber="106">Guangdong</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

222
data/03/E1/14/03E1140D106EFFEDC4B1FD68FCC6FABA.xml Normal file
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<mods:title id="0C98B8BDC28D6B81001A1C4CFC46A3AD">DNA barcodes and morphological evidence reveal a new genus of Nygmiini (Lepidoptera: Erebidae: Lymantriinae) from China</mods:title>
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<mods:date id="C97E2874A3C5F17E55DBD5CB4FEC747D">2024</mods:date>
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<mods:number id="D844DD4BD1CFDDEA9A90806C59A454AE">5496</mods:number>
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<taxonomicName id="4C48DE98106EFFEDC4B1FD68FDF1FCDD" authority="(Chao, 1984)" authorityName="Li &amp; Xia &amp; Wang" authorityYear="2024" baseAuthorityName="Chao" baseAuthorityYear="1984" box="[151,591,762,789]" class="Insecta" family="Erebidae" genus="Aurivalva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="yunnanpina" status="comb. nov.">
<emphasis id="B93C7909106EFFEDC4B1FD68FE16FCDC" bold="true" box="[151,424,762,789]" italics="true" pageId="4" pageNumber="105">Aurivalva yunnanpina</emphasis>
(
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)
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(
<figureCitation id="1373B99E106EFFEDC4B9FCD1FEBAFC94" box="[159,260,835,861]" captionStart="FIGURES 2–7" captionStartId="5.[151,264,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-20@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURES 2–7. Adults. 2–3. Aurivalva yunnanpina comb. nov. (2. male; 3. female); 4–5. Aurivalva telephanes comb. nov. (4. male; 5. female); 6. Aurivalva conistica comb. nov. (male); 7. Nygmia icilia Stoll (type species of Nygmia, holotype, male, copyright from Natural History Museum in London). Scale bars = 10 mm." pageId="4" pageNumber="105">Figs 2–3</figureCitation>
,
<figureCitation id="1373B99E106EFFEDC536FCD1FEA1FC94" box="[272,287,835,861]" captionStart="FIGURES 8–11" captionStartId="6.[151,264,1169,1193]" captionTargetBox="[151,1436,181,1146]" captionTargetId="figure-23@6.[151,1436,181,1146]" captionTargetPageId="6" captionText="FIGURES 8–11. Male genitalia. 8. Aurivalva yunnanpina comb. nov.; 9. Aurivalva telephanes comb. nov.; 10. Aurivalva conistica comb. nov.; 11. Nygmia icilia Stoll (type species of Nygmia, holotype, male, copyright from Natural History Museum in London). Scale bars = 1 mm." pageId="4" pageNumber="105">8</figureCitation>
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<figureCitation id="1373B99E106EFFEDC50AFCD1FEF4FC94" box="[300,330,835,861]" captionStart="FIGURES 12–13" captionStartId="7.[151,264,1072,1096]" captionTargetBox="[217,1436,181,1048]" captionTargetId="figure-20@7.[151,1436,181,1048]" captionTargetPageId="7" captionText="FIGURES 12–13. Female genitalia. 12. Aurivalva yunnanpina comb. nov.; 13. Aurivalva telephanes comb. nov. Scale bars = 1 mm." pageId="4" pageNumber="105">12</figureCitation>
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<figureCitation id="1373B99E106EFFEDC570FCD1FECDFC94" box="[342,371,835,861]" captionStart="FIGURE 14" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[165,1422,187,1914]" captionTargetId="figure-23@8.[159,1427,181,1919]" captionTargetPageId="8" captionText="FIGURE 14. Venation of Aurivalva yunnanpina comb. nov., male." pageId="4" pageNumber="105">14</figureCitation>
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</paragraph>
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<taxonomicName id="4C48DE98106EFFEDC4B1FC1AFD96FC56" authority="Chao, 1984: 454" authorityName="Chao" authorityPageNumber="454" authorityYear="1984" box="[151,552,903,927]" class="Insecta" family="Erebidae" genus="Euproctis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="yunnanpina">
<emphasis id="B93C7909106EFFEDC4B1FC1AFEC9FC56" box="[151,375,904,927]" italics="true" pageId="4" pageNumber="105">Euproctis yunnanpina</emphasis>
<bibRefCitation id="EFD9D8EA106EFFEDC55BFC15FD96FC56" author="Chao, C. L." box="[381,552,903,927]" pageId="4" pageNumber="105" pagination="449 - 456" refId="ref3726" refString="Chao, C. L. (1984) New species of the genus Euproctis Hubner from China (Lepidoptera: Lymantriidae). Acta Entomologica Sinica, 27 (4), 449 - 456. https: // doi. org / 10.16380 / j. kcxb. 1984.04.017" type="journal article" year="1984">Chao, 1984: 454</bibRefCitation>
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;
<treatmentCitation id="0AE9830A106EFFEDC615FC15FD60FC56" author="Chao, C. L." box="[563,734,903,927]" page="365" pageId="4" pageNumber="105" year="2003">
<bibRefCitation id="EFD9D8EA106EFFEDC615FC15FD60FC56" author="Chao, C. L." box="[563,734,903,927]" pageId="4" pageNumber="105" refId="ref3777" refString="Chao, C. L. (2003) Lepidoptera Lymantriidae. Fauna Sinica. Insecta 30. Science Press, Beijing, 484 pp." type="book" year="2003">Chao, 2003: 365</bibRefCitation>
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<emphasis id="B93C7909106EFFEDC4B1FC5FFEFBFC2E" bold="true" box="[151,325,973,999]" pageId="4" pageNumber="105">Redescription.</emphasis>
Forewing length
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. Forewing yellow, densely covered with brown scales, with two pale yellow transverse lines medially, and one or two black dots at apex. Hindwing light yellow. Male genitalia: Uncus broad, triangular. Valva broad at base; cucullus concave medially, with two sclerotized processes. Saccus long tongue-shaped. Aedeagus clavate, bent near apex, dorsal process relatively sharp. Female genitalia: Ostium ringshaped. Papilla analis broad. Apophysis anterioris longer than apophysis posterioris. Ductus narrow and long, sclerotized at base. Bursa broad, signum with a transverse sclerotization at middle.
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<emphasis id="B93C7909106EFFEDC4E1FB37FE0CFB76" bold="true" box="[199,434,1189,1215]" pageId="4" pageNumber="105">Material examined.</emphasis>
<materialsCitation id="3B20AF46106EFFEDC59CFB37FE73FB2A" collectingDate="2018-09-06" collectorName="Min Wang &amp; Zhipeng Miao." country="China" location="Shennonggu National Park" municipality="Zhuzhou City" pageId="4" pageNumber="105" specimenCount="1" specimenCount-male="1" stateProvince="Hunan">
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<location id="8E97F3C0106EFFEDC5CAFB37FCA1FB76" LSID="urn:lsid:plazi:treatment:03E1140D106EFFEDC4B1FD68FCC6FABA:8E97F3C0106EFFEDC5CAFB37FCA1FB76" box="[492,799,1189,1215]" country="China" municipality="Zhuzhou City" name="Shennonggu National Park" pageId="4" pageNumber="105" stateProvince="Hunan">Shennonggu National Park</location>
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<collectingMunicipality id="6B933F61106EFFEDC70AFB37FC77FB76" box="[812,969,1189,1215]" pageId="4" pageNumber="105">Zhuzhou City</collectingMunicipality>
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<collectingCountry id="F35FE58B106EFFEDC0BDFB37FB5FFB76" box="[1179,1249,1189,1215]" name="China" pageId="4" pageNumber="105">China</collectingCountry>
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<collectingDate id="EFB27A33106EFFEDC0C8FB37FAD8FB76" box="[1262,1382,1189,1215]" pageId="4" pageNumber="105" value="2018-09-06">6-IX-2018</collectingDate>
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<collectorName id="26BDC0CD106EFFEDC4B1FB5BFEAAFB2D" box="[151,276,1225,1252]" pageId="4" pageNumber="105">Min Wang</collectorName>
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<collectorName id="26BDC0CD106EFFEDC504FB5BFE73FB2A" box="[290,461,1225,1251]" pageId="4" pageNumber="105">Zhipeng Miao.</collectorName>
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<collectingCountry id="F35FE58B106EFFEDC52BFB7FFEEDFACE" box="[269,339,1261,1287]" name="China" pageId="4" pageNumber="105">China</collectingCountry>
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<collectingDate id="EFB27A33106EFFEDC545FB7FFDB7FACE" box="[355,521,1261,1287]" pageId="4" pageNumber="105" value="2008-08-01" valueMax="2008-08-07" valueMin="2008-08-01">1–7-VIII-2008</collectingDate>
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<collectorName id="26BDC0CD106EFFEDC66BFB7FFCB2FAC1" box="[589,780,1261,1288]" pageId="4" pageNumber="105">Houshuai Wang.</collectorName>
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<materialsCitation id="3B20AF46106EFFEDC731FB7FFBB1FAE5" collectingDate="2006-04-31" collectorName="Houshuai Wang." country="China" county="Ruyuan County" location="Nanling National Nature Reserve" municipality="Shaoguan City" pageId="4" pageNumber="105" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong">
<specimenCount id="9D4E6E92106EFFEDC731FB7FFC84FACE" box="[791,826,1261,1287]" count="1" pageId="4" pageNumber="105" type="female">1♀</specimenCount>
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<location id="8E97F3C0106EFFEDC76AFB7FFB6CFAC1" LSID="urn:lsid:plazi:treatment:03E1140D106EFFEDC4B1FD68FCC6FABA:8E97F3C0106EFFEDC76AFB7FFB6CFAC1" box="[844,1234,1261,1288]" country="China" county="Ruyuan County" municipality="Shaoguan City" name="Nanling National Nature Reserve" pageId="4" pageNumber="105" stateProvince="Guangdong">Nanling National Nature Reserve</location>
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<collectingCountry id="F35FE58B106EFFEDC665FA83FD37FAE2" box="[579,649,1297,1323]" name="China" pageId="4" pageNumber="105">China</collectingCountry>
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<date id="FFF683DB106EFFEDC6B5FA83FCA4FAE2" box="[659,794,1297,1323]" pageId="4" pageNumber="105" value="2006-04-31">
<collectingDate id="EFB27A33106EFFEDC6B5FA83FCA4FAE2" box="[659,794,1297,1323]" pageId="4" pageNumber="105" value="2006-04-31">31-IV-2006</collectingDate>
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<collectorName id="26BDC0CD106EFFEDC773FA83FBB1FAE5" box="[853,1039,1297,1324]" pageId="4" pageNumber="105">Houshuai Wang.</collectorName>
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<materialsCitation id="3B20AF46106EFFEDC033FA83FAB3FA99" collectingDate="2018-08-12" collectingDateMax="2018-08-19" collectingDateMin="2018-08-12" collectorName="Tiantian Yu &amp; Shunyun Tan." country="China" county="Jinan County" location="Baiya Mountain" municipality="Yichun City" pageId="4" pageNumber="105" specimenCount="1" specimenCount-female="1" stateProvince="Jiangxi">
<specimenCount id="9D4E6E92106EFFEDC033FA83FB86FAE2" box="[1045,1080,1297,1323]" count="1" pageId="4" pageNumber="105" type="female">1♀</specimenCount>
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<collectingRegion id="498C6BF9106EFFEDC5EDFAA7FD37FA86" box="[459,649,1333,1359]" country="China" name="Jiangxi" pageId="4" pageNumber="105">Jiangxi Province</collectingRegion>
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<collectingCountry id="F35FE58B106EFFEDC6B3FAA7FD65FA86" box="[661,731,1333,1359]" name="China" pageId="4" pageNumber="105">China</collectingCountry>
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<collectingDate id="EFB27A33106EFFEDC6C0FAA7FC16FA86" box="[742,936,1333,1359]" pageId="4" pageNumber="105" value="2018-08-12" valueMax="2018-08-19" valueMin="2018-08-12">12–19-VIII-2018</collectingDate>
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<collectorName id="26BDC0CD106EFFEDC7C5FAA7FBD9FA99" box="[995,1127,1333,1360]" pageId="4" pageNumber="105">Tiantian Yu</collectorName>
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<collectorName id="26BDC0CD106EFFEDC054FAA7FAB3FA99" box="[1138,1293,1333,1360]" pageId="4" pageNumber="105">Shunyun Tan.</collectorName>
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<figureCitation id="66A02FACFFEA7D5EA0C0FA9DFF0FFAD4" box="[159,239,1303,1329]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="2" pageNumber="509">Figs. 2</figureCitation>
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<figureCitation id="66A02FACFFEA7D5EA129FA9DFE4FFAD4" box="[374,431,1303,1329]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="2" pageNumber="509">6a, b</figureCitation>
;
<tableCitation id="B3190692FFEA7D5EA1E5FA9DFDD4FAD4" box="[442,564,1303,1329]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStart-5="TABLE 6" captionStart-6="TABLE 7" captionStartId-0="5.[151,238,476,500]" captionStartId-1="5.[151,239,1153,1177]" captionStartId-2="6.[151,239,440,464]" captionStartId-3="7.[151,237,151,175]" captionStartId-4="7.[151,234,868,892]" captionStartId-5="8.[151,236,151,175]" captionStartId-6="8.[151,237,868,892]" captionText-0="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-1="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-2="TABLE 3. Morphometrics of Rhinogobius rong and R. nami, expressed as percentages of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 2, pelvic fin." captionText-3="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-4="TABLE 5. Longitudinal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-5="TABLE 6. Transverse scale counts of various species of Rhinogobius examined in the present study. *Includes the holotype." captionText-6="TABLE 7. Predorsal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="2" pageNumber="509">Tables 1–7</tableCitation>
)
@ -351,7 +352,7 @@ and
.
</paragraph>
</subSubSection>
<caption id="AAE463A1FFEB7D5FA0C8F8D9FD62F81E" pageId="3" pageNumber="510" startId="3.[151,250,1875,1899]" targetBox="[158,1429,188,1844]" targetPageId="3" targetType="figure">
<caption id="AAE463A1FFEB7D5FA0C8F8D9FD62F81E" ID-DOI="http://doi.org/10.5281/zenodo.13330565" ID-Zenodo-Dep="13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="3" pageNumber="510" startId="3.[151,250,1875,1899]" targetBox="[158,1429,188,1844]" targetPageId="3" targetType="figure">
<paragraph id="FE243329FFEB7D5FA0C8F8D9FD62F81E" blockId="3.[151,1437,1875,2043]" pageId="3" pageNumber="510">
<emphasis id="CCEFEF3BFFEB7D5FA0C8F8D9FEF2F88E" bold="true" box="[151,274,1875,1899]" pageId="3" pageNumber="510">FIGURE 2.</emphasis>
Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of
@ -395,21 +396,21 @@ SL). Pectoral fin with 18–20 rays (usually 19). Second dorsal, and anal fin us
Body nearly cylindrical anteriorly and somewhat compressed posteriorly. Head depressed and larger in large males (&gt;
<quantity id="39639ECCFFEC7D58A1C2FE51FE0BFE10" box="[413,491,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="511" unit="mm" value="40.0">40 mm</quantity>
SL) than females (head length 31.9–35.8 vs. 28.2–31.7% of SL;
<figureCitation id="66A02FACFFEC7D58A4E6FE51FB1CFE10" box="[1209,1276,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="4" pageNumber="511">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFEC7D58A4E6FE51FB1CFE10" box="[1209,1276,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="4" pageNumber="511">Fig. 2</figureCitation>
). Eyes located dorsolaterally. Mouth terminal and oblique with thick upper and lower lips. With mouth closed, anterior tip of lower jaw protruding significantly beyond upper jaw in large males (&gt;
<quantity id="39639ECCFFEC7D58A339FDA9FC54FDD8" box="[870,948,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="511" unit="mm" value="40.0">40 mm</quantity>
SL), and upper and lower jaws almost even in females, and two smaller males (30.4- and 32.6-mm SL). Posterior end of upper jaw exceeding anterior margin of eye. Mouth larger in large males (&gt;
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SL) than females (upper-jaw length 12.2–16.5 vs. 9.9–11.6% of SL;
<figureCitation id="66A02FACFFEC7D58A0C8FD05FF3BFD4C" box="[151,219,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="4" pageNumber="511">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFEC7D58A0C8FD05FF3BFD4C" box="[151,219,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="4" pageNumber="511">Fig. 2</figureCitation>
), and end of upper jaw reaching middle of eye in larger males. Upper and lower jaws usually with 3–4 rows of conical teeth; those on outermost row enlarged and those on inner rows irregularly aligned. Mental flap on chin bilobed (
<figureCitation id="66A02FACFFEC7D58A0A7FD5DFEA6FD14" box="[248,326,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." pageId="4" pageNumber="511">Fig. 3c</figureCitation>
<figureCitation id="66A02FACFFEC7D58A0A7FD5DFEA6FD14" box="[248,326,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330569" httpUri="https://zenodo.org/record/13330569/files/figure.png" pageId="4" pageNumber="511">Fig. 3c</figureCitation>
), rounded rectangular, or trapezoidal (posterior end forming long side). Anterior nostril short, tubular; posterior nostril a pore. Cephalic sensory pore system with B´, C, D(S), E, F, H´, K´, and L´ in oculoscapular canal and M´, N, and O´ in preopercular canal of both left and right sides in nine of
<specimenCount id="E89DF8A0FFED7D59A4D6FF31FAC5FF30" box="[1161,1317,187,213]" count="18" pageId="5" pageNumber="512" type="generic">18 specimens</specimenCount>
examined (including
<typeStatus id="21208D8BFFED7D59A14CFF55FE99FF1C" box="[275,377,223,249]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
;
<figureCitation id="66A02FACFFED7D59A1DAFF55FE12FF1C" box="[389,498,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." pageId="5" pageNumber="512">Fig. 3a–c</figureCitation>
<figureCitation id="66A02FACFFED7D59A1DAFF55FE12FF1C" box="[389,498,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330569" httpUri="https://zenodo.org/record/13330569/files/figure.png" pageId="5" pageNumber="512">Fig. 3a–c</figureCitation>
), but
<specimenCount id="E89DF8A0FFED7D59A26CFF55FD2EFF1C" box="[563,718,223,249]" count="1" pageId="5" pageNumber="512" type="generic">one specimen</specimenCount>
with single Y-shaped pore C and longitudinal groove-like pore D,
@ -423,12 +424,12 @@ lacking pores N in one preopercular canals (either left or right),
lacking pore N in both preopercular canals, and
<specimenCount id="E89DF8A0FFED7D59A296FEC1FC86FE80" box="[713,870,331,357]" count="1" pageId="5" pageNumber="512" type="generic">one specimen</specimenCount>
having only pore O´ in right preopercular canal. Arrangement of cutaneous sensory papillae of head shown in
<figureCitation id="66A02FACFFED7D59A302FEE5FC2BFE6C" box="[861,971,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." pageId="5" pageNumber="512">Fig. 3a–c</figureCitation>
<figureCitation id="66A02FACFFED7D59A302FEE5FC2BFE6C" box="[861,971,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330569" httpUri="https://zenodo.org/record/13330569/files/figure.png" pageId="5" pageNumber="512">Fig. 3a–c</figureCitation>
. Cheek having four longitudinal rows of papillae and no transverse row. Vertebrae 10+15=25 (n=1) or 10+16=26 (n=12;
<tableCitation id="B3190692FFED7D59A449FE19FB87FE48" box="[1046,1127,403,429]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="5" pageNumber="512">Table 1</tableCitation>
), P-V 3/II II I I 0/9 (n=8).
</paragraph>
<caption id="AAE463A1FFEC7D58A0C8F838FA77F80B" pageId="4" pageNumber="511" startId="4.[151,243,1970,1994]" targetBox="[160,1429,786,1939]" targetPageId="4" targetType="figure">
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<paragraph id="FE243329FFEC7D58A0C8F838FA77F80B" blockId="4.[151,1436,1970,2030]" pageId="4" pageNumber="511">
<emphasis id="CCEFEF3BFFEC7D58A0C8F838FEEDF82F" bold="true" box="[151,269,1970,1994]" pageId="4" pageNumber="511">FIGURE 3.</emphasis>
Arrangement of cephalic sensory pores and cutaneous sensory papillae in
@ -864,7 +865,7 @@ and
with 18 segmented rays including 16 branched rays; posterior margin rounded. Large males (&gt;
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SL) having larger second dorsal, anal, and caudal fins than females (second dorsal-fin length 30.3–36.5 vs. 25.9–29.3% of SL, anal-fin length 27.0–30.6 vs. 24.6–26.6% of SL, and caudal-fin length 25.4–28.1 vs. 22.5–25.7% of SL;
<figureCitation id="66A02FACFFEE7D5AA359FEC1FCAAFE80" box="[774,842,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="6" pageNumber="513">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFEE7D5AA359FEC1FCAAFE80" box="[774,842,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="6" pageNumber="513">Fig. 2</figureCitation>
). Pectoral fin with 18–20 rays (usually 19) (
<tableCitation id="B3190692FFEE7D5AA568FEC1FA6FFE80" box="[1335,1423,331,357]" captionStart="TABLE 4" captionStartId="7.[151,237,151,175]" captionText="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="6" pageNumber="513">Table 4</tableCitation>
). Pelvic fin with one spine and five soft rays. Pelvic fins joined to form cuplike disc with fleshy bilobed frenum.
@ -1616,9 +1617,9 @@ examined in the present study. * Includes the holotype.
</paragraph>
<paragraph id="FE243329FFEF7D5BA098F9BAFC33F857" blockId="7.[151,1436,1584,1971]" pageId="7" pageNumber="514">
Color in life and fresh (
<figureCitation id="66A02FACFFEF7D5BA18CF9BAFDC1F9AE" box="[467,545,1584,1611]" captionStart="FIGURE 4" captionStartId="9.[151,250,993,1017]" captionTargetBox="[151,1435,151,977]" captionTargetId="figure-17@9.[151,1435,151,977]" captionTargetPageId="9" captionText="FIGURE 4. Rhinogobius rong immediately after fixation. a, NSMT-P 147886, holotype, male, 45.6 mm SL; b, OISTICH 255, paratype, female, 46.1 mm SL." pageId="7" pageNumber="514">Figs. 4</figureCitation>
<figureCitation id="66A02FACFFEF7D5BA18CF9BAFDC1F9AE" box="[467,545,1584,1611]" captionStart="FIGURE 4" captionStartId="9.[151,250,993,1017]" captionTargetBox="[151,1435,151,977]" captionTargetId="figure-17@9.[151,1435,151,977]" captionTargetPageId="9" captionText="FIGURE 4. Rhinogobius rong immediately after fixation. a, NSMT-P 147886, holotype, male, 45.6 mm SL; b, OISTICH 255, paratype, female, 46.1 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330573" httpUri="https://zenodo.org/record/13330573/files/figure.png" pageId="7" pageNumber="514">Figs. 4</figureCitation>
and
<figureCitation id="66A02FACFFEF7D5BA205F9BAFD8AF9AE" box="[602,618,1584,1611]" captionStart="FIGURE 5" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[151,1435,1077,1933]" captionTargetId="figure-43@9.[151,1435,1077,1933]" captionTargetPageId="9" captionText="FIGURE 5. Live images of Rhinogobius rong in aquarium (a) and in situ underwater footage from Da Nang City, Vietnam (b–d). a, b, males, 12 January 2013; c, female, 12 January 2013; d, female, 1 March 2016 (photo by K. Maeda)." pageId="7" pageNumber="514">5</figureCitation>
<figureCitation id="66A02FACFFEF7D5BA205F9BAFD8AF9AE" box="[602,618,1584,1611]" captionStart="FIGURE 5" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[151,1435,1077,1933]" captionTargetId="figure-43@9.[151,1435,1077,1933]" captionTargetPageId="9" captionText="FIGURE 5. Live images of Rhinogobius rong in aquarium (a) and in situ underwater footage from Da Nang City, Vietnam (b–d). a, b, males, 12 January 2013; c, female, 12 January 2013; d, female, 1 March 2016 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330579" httpUri="https://zenodo.org/record/13330579/files/figure.png" pageId="7" pageNumber="514">5</figureCitation>
): in male, background of head and body greenish or yellowish-grey, but whitish on ventral side. An orange stripe connecting snout tip and anterior margin of eye; this stripe continuing posteriorly from posterior margin of eye to region above gill opening. Faint orange stripe running horizontally below eye to opercle; often intermittent. Broad, diagonally downward orange stripe running from above middle of upper lip to cheek (sometimes intermittent); another short, orange stripe running above posterior end of upper lip parallel to broad stripe; one or two large orange spot(s) on cheek posterior to broad stripe. Broad, orange stripe running almost horizontally (but slightly diagonally upward) along middle of opercle and subopercle; extending posteriorly on pectoral-fin base, where it being darker. Opercular membrane, interopercle, and lower part of subopercle with light orange or yellow markings arranged almost horizontally. Six indistinct dusky-grey blotches and five indistinct cream bands between them sometimes appearing laterally on body from flank to caudal peduncle. Many orange spots scattered on lateral and dorsal sides of body and dorsal side of head.
</paragraph>
<caption id="AAE463A1FFE07D54A0C8FF1DFA73FF4A" box="[151,1427,151,175]" pageId="8" pageNumber="515" startId="8.[151,236,151,175]" targetType="table">
@ -2200,7 +2201,7 @@ examined in the present study. * Includes the holotype.
</table>
</paragraph>
<paragraph id="FE243329FFE07D54A098F9BAFB44F86B" blockId="8.[151,1437,1584,1935]" pageId="8" pageNumber="515">First dorsal fin greyish translucent, with a black spot just below middle on membrane between first and second spines; upper part of this spot bluish. Some orange markings aligned along proximal part of fin and on middle to distal parts of fourth, fifth, and sixth spines. Distal part of fin yellowish, especially on membrane between first and second spines; middle to distal part of membrane between second and third spines light blue. Second dorsal fin greyish translucent, with many orange markings forming four or five indefinite horizontal stripes. Two orange markings on center of caudal-fin base; dorsal one directed upward and ventral one directed downward. Remaining caudal fin greyish translucent, with many orange markings forming five to seven vertical bands. Anal fin greyish translucent proximally, orange distally, with a bluish-white margin. Pectoral fin translucent, but its proximal part cream yellow or yellowish-grey with orange markings aligned vertically along this cream yellow or yellowish-grey part both on proximal and distal sides. Pelvic disc translucent or grey without clear marking.</paragraph>
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<paragraph id="FE243329FFE17D55A0C8FC6BFE32FBF8" blockId="9.[151,1436,993,1053]" pageId="9" pageNumber="516">
<emphasis id="CCEFEF3BFFE17D55A0C8FC6BFEF3FC1C" bold="true" box="[151,275,993,1017]" pageId="9" pageNumber="516">FIGURE 4.</emphasis>
<taxonomicName id="399B48AAFFE17D55A145FC6BFE2FFC1C" authorityName="Maeda &amp; Kobayashi &amp; Iida &amp; Tran" authorityYear="2024" box="[282,463,993,1017]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="516" phylum="Chordata" rank="species" species="rong">
@ -2209,7 +2210,7 @@ examined in the present study. * Includes the holotype.
immediately after fixation. a, NSMT-P 147886, holotype, male, 45.6 mm SL; b, OISTICH 255, paratype, female, 46.1 mm SL.
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<emphasis id="CCEFEF3BFFE17D55A0C8F82FFEF5F85B" bold="true" box="[151,277,1957,1982]" pageId="9" pageNumber="516">FIGURE 5.</emphasis>
Live images of
@ -2221,7 +2222,7 @@ in aquarium (a) and
underwater footage from Da Nang City, Vietnam (b–d). a, b, males, 12 January 2013; c, female, 12 January 2013; d, female, 1 March 2016 (photo by K. Maeda).
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<emphasis id="CCEFEF3BFFE27D56A0C8FB42FEF3FB05" bold="true" box="[151,275,1224,1248]" pageId="10" pageNumber="517">FIGURE 6.</emphasis>
Preserved specimens of
@ -2250,7 +2251,7 @@ Preserved specimens of
<paragraph id="FE243329FFE27D56A098FA15FE9FF989" blockId="10.[151,1437,1438,2041]" pageId="10" pageNumber="517">In female, markings and background color of body and fins similar to those of male, but markings reddish-brown or grey instead of orange and each marking smaller or less distinct. Diagonal stripe from above middle of upper lip usually broken in middle; posterior half of this marking forming part of series of markings horizontally aligned on cheek. First dorsal fin with no or faint black spot between first and second spines; yellowish and bluish color in distal part of fin indistinct. Pectoral fin with a pale-yellow spot on upper-proximal part. Pelvic disc translucent or light yellowish-grey.</paragraph>
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Color in preservative (
<figureCitation id="66A02FACFFE27D56A19EF9FCFDCEF975" box="[449,558,1654,1680]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." pageId="10" pageNumber="517">Fig. 6a, b</figureCitation>
<figureCitation id="66A02FACFFE27D56A19EF9FCFDCEF975" box="[449,558,1654,1680]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="10" pageNumber="517">Fig. 6a, b</figureCitation>
): background of head and body cream and that of fins translucent or light grey. Orange, reddish-brown, and grey markings observed in life becoming brown. Pale orange or yellow patterns on opercular membrane, interopercle, and lower part of subopercle disappeared. All other yellow and bluish markings also disappeared.
</paragraph>
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135
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<paragraph id="FE243329FFF17D45A0C8FF31FD80FF30" blockId="25.[151,608,151,213]" box="[151,608,187,213]" pageId="25" pageNumber="532">
(
<figureCitation id="66A02FACFFF17D45A0C0FF31FF0FFF30" box="[159,239,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="25" pageNumber="532">Figs. 2</figureCitation>
<figureCitation id="66A02FACFFF17D45A0C0FF31FF0FFF30" box="[159,239,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="25" pageNumber="532">Figs. 2</figureCitation>
,
<figureCitation id="66A02FACFFF17D45A0A4FF31FEC9FF30" box="[251,297,187,213]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." pageId="25" pageNumber="532">6i, j</figureCitation>
<figureCitation id="66A02FACFFF17D45A0A4FF31FEC9FF30" box="[251,297,187,213]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="25" pageNumber="532">6i, j</figureCitation>
,
<figureCitation id="66A02FACFFF17D45A16BFF31FE9DFF30" box="[308,381,187,213]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStartId-0="26.[151,250,1943,1967]" captionStartId-1="27.[151,250,1425,1449]" captionStartId-2="28.[151,250,1441,1465]" captionStartId-3="29.[151,250,1124,1148]" captionTargetBox-0="[323,1264,190,1910]" captionTargetBox-1="[151,1435,181,1401]" captionTargetBox-2="[151,1435,561,1417]" captionTargetBox-3="[158,1429,532,1093]" captionTargetId-0="figure-18@26.[314,1273,181,1919]" captionTargetId-1="figure-17@27.[151,1435,181,1401]" captionTargetId-2="figure-182@28.[151,1435,561,1417]" captionTargetId-3="figure-186@29.[151,1436,525,1100]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionText-0="FIGURE 16. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius similis (OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." captionText-1="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." captionText-2="FIGURE 18. In situ underwater photographs of Rhinogobius similis in Da Nang City, Vietnam. a, male, 12 January 2013; b, male, 4 March 2016; c, female, 4 March 2016; d, female, 12 January 2013 (photo by K. Maeda)." captionText-3="FIGURE 19. Relationships of head length and upper-jaw length to standard length (SL) in Rhinogobius similis collected from Da Nang City (light green squares) and Thua Thien Hue Province (orange square) in central Vietnam, Quang Ninh Province in northern Vietnam (red circles), Okinawa Island (light blue circles) and Izu Peninsula (deep blue diamonds) in Japan, holotype and paratypes of Ctenogobius lini Herre, 1934 from Wuzhou, China (deep green x), holotype of Gobius giurinus Rutter, 1897 from Shantou, China (pink x), and holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 from Nagasaki, Japan (deep blue x). Solid symbols represent males, open symbols represent females, and those filled with grey represent specimens of unknown sex." pageId="25" pageNumber="532">16–19</figureCitation>
<figureCitation id="66A02FACFFF17D45A16BFF31FE9DFF30" box="[308,381,187,213]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStartId-0="26.[151,250,1943,1967]" captionStartId-1="27.[151,250,1425,1449]" captionStartId-2="28.[151,250,1441,1465]" captionStartId-3="29.[151,250,1124,1148]" captionTargetBox-0="[323,1264,190,1910]" captionTargetBox-1="[151,1435,181,1401]" captionTargetBox-2="[151,1435,561,1417]" captionTargetBox-3="[158,1429,532,1093]" captionTargetId-0="figure-18@26.[314,1273,181,1919]" captionTargetId-1="figure-17@27.[151,1435,181,1401]" captionTargetId-2="figure-182@28.[151,1435,561,1417]" captionTargetId-3="figure-186@29.[151,1436,525,1100]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionText-0="FIGURE 16. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius similis (OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." captionText-1="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." captionText-2="FIGURE 18. In situ underwater photographs of Rhinogobius similis in Da Nang City, Vietnam. a, male, 12 January 2013; b, male, 4 March 2016; c, female, 4 March 2016; d, female, 12 January 2013 (photo by K. Maeda)." captionText-3="FIGURE 19. Relationships of head length and upper-jaw length to standard length (SL) in Rhinogobius similis collected from Da Nang City (light green squares) and Thua Thien Hue Province (orange square) in central Vietnam, Quang Ninh Province in northern Vietnam (red circles), Okinawa Island (light blue circles) and Izu Peninsula (deep blue diamonds) in Japan, holotype and paratypes of Ctenogobius lini Herre, 1934 from Wuzhou, China (deep green x), holotype of Gobius giurinus Rutter, 1897 from Shantou, China (pink x), and holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 from Nagasaki, Japan (deep blue x). Solid symbols represent males, open symbols represent females, and those filled with grey represent specimens of unknown sex." figureDoi-0="http://doi.org/10.5281/zenodo.13330605" figureDoi-1="http://doi.org/10.5281/zenodo.13330607" figureDoi-2="http://doi.org/10.5281/zenodo.13330609" figureDoi-3="http://doi.org/10.5281/zenodo.13330611" httpUri-0="https://zenodo.org/record/13330605/files/figure.png" httpUri-1="https://zenodo.org/record/13330607/files/figure.png" httpUri-2="https://zenodo.org/record/13330609/files/figure.png" httpUri-3="https://zenodo.org/record/13330611/files/figure.png" pageId="25" pageNumber="532">16–19</figureCitation>
;
<tableCitation id="B3190692FFF17D45A1D6FF31FE06FF30" box="[393,486,187,213]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="25" pageNumber="532">Tables 1</tableCitation>
,
@ -560,7 +561,7 @@ and
<paragraph id="FE243329FFF17D45A098FAC9FD09F90C" blockId="25.[151,1437,663,2022]" pageId="25" pageNumber="532">
<emphasis id="CCEFEF3BFFF17D45A098FAC9FEB9FAB8" bold="true" box="[199,345,1347,1373]" pageId="25" pageNumber="532">Description.</emphasis>
Body nearly cylindrical anteriorly, and somewhat compressed posteriorly. Head depressed and eyes located dorsolaterally. Mouth terminal and oblique, with thick upper and lower lips. When mouth closed, anterior tip of upper jaw slightly protruding beyond lower jaw, or both jaws almost even. Mouth larger in males than females (upper-jaw length 12.6–17.6 vs. 10.9–12.2% of SL). Upper and lower jaws usually with 3–5 rows of conical teeth; those in outermost row enlarged and those in inner rows irregularly aligned. Mental flap on chin usually rectangular, trapezoidal (posterior end forming long side), or emarginate, and often with a small central projection. Anterior nostril short and tubular, posterior nostril a pore. Cephalic sensory pore system usually with B´, C, D(S), E, F, H´, K´, and L´ in oculoscapular canal and M´, N, and O´ in preopercular canal (
<figureCitation id="66A02FACFFF17D45A435F9B5FB5EF9BC" box="[1130,1214,1599,1625]" captionStart="FIGURE 16" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[323,1264,190,1910]" captionTargetId="figure-18@26.[314,1273,181,1919]" captionTargetPageId="26" captionText="FIGURE 16. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius similis (OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." pageId="25" pageNumber="532">Fig. 16</figureCitation>
<figureCitation id="66A02FACFFF17D45A435F9B5FB5EF9BC" box="[1130,1214,1599,1625]" captionStart="FIGURE 16" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[323,1264,190,1910]" captionTargetId="figure-18@26.[314,1273,181,1919]" captionTargetPageId="26" captionText="FIGURE 16. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius similis (OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330605" httpUri="https://zenodo.org/record/13330605/files/figure.png" pageId="25" pageNumber="532">Fig. 16</figureCitation>
), but
<specimenCount id="E89DF8A0FFF17D45A55EF9CAFA7CF9BC" box="[1281,1436,1599,1626]" count="1" pageId="25" pageNumber="532" type="generic">one specimen</specimenCount>
lacking pore C on left side,
@ -568,7 +569,7 @@ lacking pore C on left side,
lacking pore N on right side, and
<specimenCount id="E89DF8A0FFF17D45A3A6F9EEFB74F998" box="[1017,1172,1635,1662]" count="1" pageId="25" pageNumber="532" type="generic">one specimen</specimenCount>
having two pores F on both left and right sides. Arrangement of cutaneous sensory papillae of head shown in
<figureCitation id="66A02FACFFF17D45A40DF90DFB43F944" box="[1106,1187,1671,1697]" captionStart="FIGURE 16" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[323,1264,190,1910]" captionTargetId="figure-18@26.[314,1273,181,1919]" captionTargetPageId="26" captionText="FIGURE 16. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius similis (OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." pageId="25" pageNumber="532">Fig. 16</figureCitation>
<figureCitation id="66A02FACFFF17D45A40DF90DFB43F944" box="[1106,1187,1671,1697]" captionStart="FIGURE 16" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[323,1264,190,1910]" captionTargetId="figure-18@26.[314,1273,181,1919]" captionTargetPageId="26" captionText="FIGURE 16. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius similis (OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330605" httpUri="https://zenodo.org/record/13330605/files/figure.png" pageId="25" pageNumber="532">Fig. 16</figureCitation>
. Cheek having several short transverse rows of papillae below eye and between two longitudinal rows across center of cheek. Vertebrae 10+16=26 (n=10;
<tableCitation id="B3190692FFF17D45A13CF945FE55F90C" box="[355,437,1743,1769]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="25" pageNumber="532">Table 1</tableCitation>
), P-V 3/II II I I 0/9 (n=10).
@ -581,12 +582,12 @@ First dorsal fin with six spines, and second dorsal fin with one spine and eight
. Anal fin with one spine and seven (n=1) or eight (n=23) soft rays (
<tableCitation id="B3190692FFF37D47A568FA76FA6FF9F3" box="[1335,1423,1532,1558]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="27" pageNumber="534">Table 2</tableCitation>
). Caudal fin with 17 segmented rays including 12 (n=3), 13 (n=15), 14 (n=5), or 15 (n=1) branched rays; posterior margin rounded. Males having larger second dorsal and anal fins than females (second dorsal-fin length 31.3–43.7 vs 26.0–30.0% and anal-fin length 26.4–38.5 vs 23.7–28.2% of SL;
<figureCitation id="66A02FACFFF37D47A3CBF9E2FC38F967" box="[916,984,1640,1666]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="27" pageNumber="534">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFF37D47A3CBF9E2FC38F967" box="[916,984,1640,1666]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="27" pageNumber="534">Fig. 2</figureCitation>
). Pectoral fin with 17–20 rays (mode 19 rays) (
<tableCitation id="B3190692FFF37D47A081F906FED6F943" box="[222,310,1676,1702]" captionStart="TABLE 4" captionStartId="7.[151,237,151,175]" captionText="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="27" pageNumber="534">Table 4</tableCitation>
). Pelvic fin with one spine and five soft rays; pelvic fins joined together to form a cuplike disc with fleshy bilobed frenum.
</paragraph>
<caption id="AAE463A1FFF27D46A0C8F81DFC02F831" pageId="26" pageNumber="533" startId="26.[151,250,1943,1967]" targetBox="[323,1264,190,1910]" targetPageId="26" targetType="figure">
<caption id="AAE463A1FFF27D46A0C8F81DFC02F831" ID-DOI="http://doi.org/10.5281/zenodo.13330605" ID-Zenodo-Dep="13330605" httpUri="https://zenodo.org/record/13330605/files/figure.png" pageId="26" pageNumber="533" startId="26.[151,250,1943,1967]" targetBox="[323,1264,190,1910]" targetPageId="26" targetType="figure">
<paragraph id="FE243329FFF27D46A0C8F81DFC02F831" blockId="26.[151,1436,1943,2004]" pageId="26" pageNumber="533">
<emphasis id="CCEFEF3BFFF27D46A0C8F81DFEFFF84A" bold="true" box="[151,287,1943,1967]" pageId="26" pageNumber="533">FIGURE 16.</emphasis>
Arrangement of cephalic sensory pores and cutaneous sensory papillae in
@ -596,7 +597,7 @@ Arrangement of cephalic sensory pores and cutaneous sensory papillae in
(OISTICH 228). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris.
</paragraph>
</caption>
<caption id="AAE463A1FFF37D47A0C8FA1BFAE8FA2B" pageId="27" pageNumber="534" startId="27.[151,250,1425,1449]" targetBox="[151,1435,181,1401]" targetPageId="27" targetType="figure">
<caption id="AAE463A1FFF37D47A0C8FA1BFAE8FA2B" ID-DOI="http://doi.org/10.5281/zenodo.13330607" ID-Zenodo-Dep="13330607" httpUri="https://zenodo.org/record/13330607/files/figure.png" pageId="27" pageNumber="534" startId="27.[151,250,1425,1449]" targetBox="[151,1435,181,1401]" targetPageId="27" targetType="figure">
<paragraph id="FE243329FFF37D47A0C8FA1BFAE8FA2B" blockId="27.[151,1436,1425,1486]" pageId="27" pageNumber="534">
<emphasis id="CCEFEF3BFFF37D47A0C8FA1BFEC2FA4C" bold="true" box="[151,290,1425,1449]" pageId="27" pageNumber="534">FIGURE 17.</emphasis>
<taxonomicName id="399B48AAFFF37D47A175FA1BFE14FA4C" authorityName="Gill" authorityYear="1859" box="[298,500,1425,1449]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="534" phylum="Chordata" rank="species" species="similis">
@ -618,12 +619,12 @@ Suzuki
</paragraph>
<paragraph id="FE243329FFF47D40A098FE89FF02FDFC" blockId="28.[151,1437,151,537]" pageId="28" pageNumber="535">
Color in life and fresh (
<figureCitation id="66A02FACFFF47D40A192FE89FDC8FEF8" box="[461,552,259,285]" captionStart="FIGURE 17" captionStartId="27.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-17@27.[151,1435,181,1401]" captionTargetPageId="27" captionText="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." pageId="28" pageNumber="535">Figs. 17</figureCitation>
<figureCitation id="66A02FACFFF47D40A192FE89FDC8FEF8" box="[461,552,259,285]" captionStart="FIGURE 17" captionStartId="27.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-17@27.[151,1435,181,1401]" captionTargetPageId="27" captionText="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330607" httpUri="https://zenodo.org/record/13330607/files/figure.png" pageId="28" pageNumber="535">Figs. 17</figureCitation>
and
<figureCitation id="66A02FACFFF47D40A201FE89FD9DFEF8" box="[606,637,259,285]" captionStart="FIGURE 18" captionStartId="28.[151,250,1441,1465]" captionTargetBox="[151,1435,561,1417]" captionTargetId="figure-182@28.[151,1435,561,1417]" captionTargetPageId="28" captionText="FIGURE 18. In situ underwater photographs of Rhinogobius similis in Da Nang City, Vietnam. a, male, 12 January 2013; b, male, 4 March 2016; c, female, 4 March 2016; d, female, 12 January 2013 (photo by K. Maeda)." pageId="28" pageNumber="535">18</figureCitation>
<figureCitation id="66A02FACFFF47D40A201FE89FD9DFEF8" box="[606,637,259,285]" captionStart="FIGURE 18" captionStartId="28.[151,250,1441,1465]" captionTargetBox="[151,1435,561,1417]" captionTargetId="figure-182@28.[151,1435,561,1417]" captionTargetPageId="28" captionText="FIGURE 18. In situ underwater photographs of Rhinogobius similis in Da Nang City, Vietnam. a, male, 12 January 2013; b, male, 4 March 2016; c, female, 4 March 2016; d, female, 12 January 2013 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330609" httpUri="https://zenodo.org/record/13330609/files/figure.png" pageId="28" pageNumber="535">18</figureCitation>
): in males, background of head and body grey or yellowish-brown, and whitish ventral side. Head with many reddish-brown dots and lines forming many nearly parallel stripes running diagonally upward from anterior to posterior parts. An indistinct broad blackish band sometimes connecting middle of upper lip and anteroventral margin of eye, and also connecting posteroventral margin of eye and lower part of opercle. Pectoral-fin base with a black blotch on its upper part, and a white, transverse band. Dark-brown markings forming indefinite, bands and blotches on lateral and dorsal sides of body, often irregularly placed and uncountable. Scales on sides of body edged with orange, and each scale having a light blue (lower part) or white (dorsal part) center.
</paragraph>
<caption id="AAE463A1FFF47D40A0C8FA2BFB8AFA38" pageId="28" pageNumber="535" startId="28.[151,250,1441,1465]" targetBox="[151,1435,561,1417]" targetPageId="28" targetType="figure">
<caption id="AAE463A1FFF47D40A0C8FA2BFB8AFA38" ID-DOI="http://doi.org/10.5281/zenodo.13330609" ID-Zenodo-Dep="13330609" httpUri="https://zenodo.org/record/13330609/files/figure.png" pageId="28" pageNumber="535" startId="28.[151,250,1441,1465]" targetBox="[151,1435,561,1417]" targetPageId="28" targetType="figure">
<paragraph id="FE243329FFF47D40A0C8FA2BFB8AFA38" blockId="28.[151,1436,1441,1502]" pageId="28" pageNumber="535">
<emphasis id="CCEFEF3BFFF47D40A0C8FA2BFEC1FA5C" bold="true" box="[151,289,1441,1465]" pageId="28" pageNumber="535">FIGURE 18.</emphasis>
<emphasis id="CCEFEF3BFFF47D40A176FA28FE8AFA5C" box="[297,362,1442,1465]" italics="true" pageId="28" pageNumber="535">In situ</emphasis>
@ -638,7 +639,7 @@ in Da Nang City, Vietnam. a, male, 12 January 2013; b, male, 4 March 2016; c, fe
<paragraph id="FE243329FFF47D40A098F96EFA7CF854" blockId="28.[151,1436,1547,2042]" pageId="28" pageNumber="535">In females, markings and background color of head and body similar to those in males, but females usually appear lighter colored than males Orange margin of scales on sides of body relatively inconspicuous. First and second dorsal fins translucent, with many reddish-brown and bluish-white spots alternately aligned along spines and soft-rays; distal parts reddish-brown translucent. Caudal fin similar to that of males, but without clear orange and yellow, dorsal stripes. Anal fin translucent with faint yellowish markings, and with dark-grey soft rays. Pectoral fin translucent, but its lower proximal part white, with a dorsal reddish-brown band. Pelvic disc yellowish translucent.</paragraph>
<paragraph id="FE243329FFF47D40A098F831FA74F81C" blockId="28.[151,1436,1547,2042]" pageId="28" pageNumber="535">
Color in preservative (
<figureCitation id="66A02FACFFF47D40A1E1F831FDC0F830" box="[446,544,1979,2005]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." pageId="28" pageNumber="535">Fig. 6i, j</figureCitation>
<figureCitation id="66A02FACFFF47D40A1E1F831FDC0F830" box="[446,544,1979,2005]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="28" pageNumber="535">Fig. 6i, j</figureCitation>
): background of head and body cream. Reddish-brown markings observed in life become dark brown. Orange, light blue, white, and yellow markings disappear, and those on fins become pale brown.
</paragraph>
</subSubSection>
@ -654,13 +655,13 @@ have larger head length than those from northern
SL, head length is 29.9–34.0 vs. 28.7–31.2% of SL), but smaller than those from
<collectingCountry id="868C73B9FFF57D41A575FF31FA8AFF30" box="[1322,1386,187,213]" name="Japan" pageId="29" pageNumber="536">Japan</collectingCountry>
(vs. 32.1–35.5% of SL) (
<figureCitation id="66A02FACFFF57D41A126FF55FE2BFF1C" box="[377,459,223,249]" captionStart="FIGURE 19" captionStartId="29.[151,250,1124,1148]" captionTargetBox="[158,1429,532,1093]" captionTargetId="figure-186@29.[151,1436,525,1100]" captionTargetPageId="29" captionText="FIGURE 19. Relationships of head length and upper-jaw length to standard length (SL) in Rhinogobius similis collected from Da Nang City (light green squares) and Thua Thien Hue Province (orange square) in central Vietnam, Quang Ninh Province in northern Vietnam (red circles), Okinawa Island (light blue circles) and Izu Peninsula (deep blue diamonds) in Japan, holotype and paratypes of Ctenogobius lini Herre, 1934 from Wuzhou, China (deep green x), holotype of Gobius giurinus Rutter, 1897 from Shantou, China (pink x), and holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 from Nagasaki, Japan (deep blue x). Solid symbols represent males, open symbols represent females, and those filled with grey represent specimens of unknown sex." pageId="29" pageNumber="536">Fig. 19</figureCitation>
<figureCitation id="66A02FACFFF57D41A126FF55FE2BFF1C" box="[377,459,223,249]" captionStart="FIGURE 19" captionStartId="29.[151,250,1124,1148]" captionTargetBox="[158,1429,532,1093]" captionTargetId="figure-186@29.[151,1436,525,1100]" captionTargetPageId="29" captionText="FIGURE 19. Relationships of head length and upper-jaw length to standard length (SL) in Rhinogobius similis collected from Da Nang City (light green squares) and Thua Thien Hue Province (orange square) in central Vietnam, Quang Ninh Province in northern Vietnam (red circles), Okinawa Island (light blue circles) and Izu Peninsula (deep blue diamonds) in Japan, holotype and paratypes of Ctenogobius lini Herre, 1934 from Wuzhou, China (deep green x), holotype of Gobius giurinus Rutter, 1897 from Shantou, China (pink x), and holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 from Nagasaki, Japan (deep blue x). Solid symbols represent males, open symbols represent females, and those filled with grey represent specimens of unknown sex." figureDoi="http://doi.org/10.5281/zenodo.13330611" httpUri="https://zenodo.org/record/13330611/files/figure.png" pageId="29" pageNumber="536">Fig. 19</figureCitation>
). Although coloration of specimens from central and northern
<collectingCountry id="868C73B9FFF57D41A423FF55FB3BFF1C" box="[1148,1243,223,249]" name="Vietnam" pageId="29" pageNumber="536">Vietnam</collectingCountry>
are similar (
<figureCitation id="66A02FACFFF57D41A53BFF55FF28FEF8" captionStart="FIGURE 9" captionStartId="16.[151,250,207,231]" captionText="FIGURE 9. Comparative material of the genus Rhinogobius from northern Vietnam (a–l) and Japan (m–p) observed in the present study. Photographs were taken immediately after fixation. Specimens on left side are males, right are females. a, b, R. sulcatus (a, OISTICH 184, 30.4 mm SL; b, OISTICH 190, 32.4 mm SL); c, d, R. boa (c, OISTICH 161, 25.4 mm SL; d, OISTICH 160, 28.0 mm SL); e, f, R. virgigena (e, NSMT-P 147872, 36.1 mm SL; f, OISTICH 154, 33.7 mm SL); g, h, R. variolatus (g, OISTICH 169, 31.4 mm SL; h, OISTICH 170, 28.8 mm SL); i, j, R. leavelli (i, OISTICH 209, 38.1 mm SL; j, OISTICH 210, 35.4 mm SL); k, l, R. similis (k, OISTICH 222, 54.4 mm SL; l, NSMT-P 147882, 44.7 mm SL); m, n, R. similis from Okinawa Island (m, URM-P 49149, 48.6 mm SL; n, OISTICH 2, 63.6 mm SL); o, p, R. similis, from Izu Peninsula (o, OISTICH 330, 47.6 mm SL; p, OISTICH 332, 51.0 mm SL)." pageId="29" pageNumber="536">Figs. 9k, l</figureCitation>
and
<figureCitation id="66A02FACFFF57D41A0A3FE89FEFAFEF8" box="[252,282,259,285]" captionStart="FIGURE 17" captionStartId="27.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-17@27.[151,1435,181,1401]" captionTargetPageId="27" captionText="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." pageId="29" pageNumber="536">17</figureCitation>
<figureCitation id="66A02FACFFF57D41A0A3FE89FEFAFEF8" box="[252,282,259,285]" captionStart="FIGURE 17" captionStartId="27.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-17@27.[151,1435,181,1401]" captionTargetPageId="27" captionText="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330607" httpUri="https://zenodo.org/record/13330607/files/figure.png" pageId="29" pageNumber="536">17</figureCitation>
), these differ from Japanese populations. Vietnamese specimens usually lack the large black blotches on lateral side of the body, while Japanese specimens always have five, large black blotches along the lateral midline (
<figureCitation id="66A02FACFFF57D41A0C0FEC1FEFEFE80" box="[159,286,331,357]" captionStart="FIGURE 9" captionStartId="16.[151,250,207,231]" captionText="FIGURE 9. Comparative material of the genus Rhinogobius from northern Vietnam (a–l) and Japan (m–p) observed in the present study. Photographs were taken immediately after fixation. Specimens on left side are males, right are females. a, b, R. sulcatus (a, OISTICH 184, 30.4 mm SL; b, OISTICH 190, 32.4 mm SL); c, d, R. boa (c, OISTICH 161, 25.4 mm SL; d, OISTICH 160, 28.0 mm SL); e, f, R. virgigena (e, NSMT-P 147872, 36.1 mm SL; f, OISTICH 154, 33.7 mm SL); g, h, R. variolatus (g, OISTICH 169, 31.4 mm SL; h, OISTICH 170, 28.8 mm SL); i, j, R. leavelli (i, OISTICH 209, 38.1 mm SL; j, OISTICH 210, 35.4 mm SL); k, l, R. similis (k, OISTICH 222, 54.4 mm SL; l, NSMT-P 147882, 44.7 mm SL); m, n, R. similis from Okinawa Island (m, URM-P 49149, 48.6 mm SL; n, OISTICH 2, 63.6 mm SL); o, p, R. similis, from Izu Peninsula (o, OISTICH 330, 47.6 mm SL; p, OISTICH 332, 51.0 mm SL)." pageId="29" pageNumber="536">
Figs.
@ -668,7 +669,7 @@ Figs.
–p
</figureCitation>
and
<figureCitation id="66A02FACFFF57D41A10DFEC1FE91FE80" box="[338,369,331,357]" captionStart="FIGURE 17" captionStartId="27.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-17@27.[151,1435,181,1401]" captionTargetPageId="27" captionText="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." pageId="29" pageNumber="536">17</figureCitation>
<figureCitation id="66A02FACFFF57D41A10DFEC1FE91FE80" box="[338,369,331,357]" captionStart="FIGURE 17" captionStartId="27.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-17@27.[151,1435,181,1401]" captionTargetPageId="27" captionText="FIGURE 17. Rhinogobius similis from Da Nang City (a, b) and Thua Thien Hue Province (c) immediately after fixation. a, OISTICH 240, male, 56.3 mm SL; b, OISTICH 241, female, 45.2 mm SL; c, OISTICH 243, male, 59.7 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330607" httpUri="https://zenodo.org/record/13330607/files/figure.png" pageId="29" pageNumber="536">17</figureCitation>
). A yellow longitudinal stripe on the second dorsal fin of the males is thinner in Vietnamese than in Japanese specimens. The ventral surface of the head in females is often black in Japanese specimens, especially in gravid individuals, but none of the Vietnamese specimens showed this black color.
<taxonomicName id="399B48AAFFF57D41A422FE19FA7CFE48" authorityName="Jordan &amp; Snyder" authorityYear="1901" box="[1149,1436,403,429]" family="Gobiidae" genus="Ctenogobius" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="536" phylum="Chordata" rank="species" species="hadropterus">
<emphasis id="CCEFEF3BFFF57D41A422FE19FA7CFE48" box="[1149,1436,403,429]" italics="true" pageId="29" pageNumber="536">Ctenogobius hadropterus</emphasis>
@ -688,10 +689,10 @@ collected in
,
<collectingCountry id="868C73B9FFF57D41A4CEFE3DFB31FE34" box="[1169,1233,439,465]" name="Japan" pageId="29" pageNumber="536">Japan</collectingCountry>
before 1901, also has distinct lateral blotches (
<figureCitation id="66A02FACFFF57D41A188FE51FDD9FE10" box="[471,569,475,501]" captionStart="FIGURE 20" captionStartId="30.[151,250,584,608]" captionTargetBox="[151,1435,181,550]" captionTargetId="figure-18@30.[151,1435,181,560]" captionTargetPageId="30" captionText="FIGURE 20. Type specimens of species currently considered synonyms of Rhinogobius similis Gill, 1859. a, b, CAS-SU 29089, possible types of Ctenogobius lini Herre, 1934 (a, 95.4 mm SL; b, 94.5 mm SL; one of them should be the holotype and the other is a paratype); c, CAS-SU 4990, holotype of Gobius giurinus Rutter, 1897 (63.0 mm SL); d, CAS-SU 6449, holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 (45.3 mm SL)." pageId="29" pageNumber="536">Fig. 20d</figureCitation>
<figureCitation id="66A02FACFFF57D41A188FE51FDD9FE10" box="[471,569,475,501]" captionStart="FIGURE 20" captionStartId="30.[151,250,584,608]" captionTargetBox="[151,1435,181,550]" captionTargetId="figure-18@30.[151,1435,181,560]" captionTargetPageId="30" captionText="FIGURE 20. Type specimens of species currently considered synonyms of Rhinogobius similis Gill, 1859. a, b, CAS-SU 29089, possible types of Ctenogobius lini Herre, 1934 (a, 95.4 mm SL; b, 94.5 mm SL; one of them should be the holotype and the other is a paratype); c, CAS-SU 4990, holotype of Gobius giurinus Rutter, 1897 (63.0 mm SL); d, CAS-SU 6449, holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 (45.3 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.13330613" httpUri="https://zenodo.org/record/13330613/files/figure.png" pageId="29" pageNumber="536">Fig. 20d</figureCitation>
).
</paragraph>
<caption id="AAE463A1FFF57D41A0C8FBEEFEC5FAB6" pageId="29" pageNumber="536" startId="29.[151,250,1124,1148]" targetBox="[158,1429,532,1093]" targetPageId="29" targetType="figure">
<caption id="AAE463A1FFF57D41A0C8FBEEFEC5FAB6" ID-DOI="http://doi.org/10.5281/zenodo.13330611" ID-Zenodo-Dep="13330611" httpUri="https://zenodo.org/record/13330611/files/figure.png" pageId="29" pageNumber="536" startId="29.[151,250,1124,1148]" targetBox="[158,1429,532,1093]" targetPageId="29" targetType="figure">
<paragraph id="FE243329FFF57D41A0C8FBEEFEC5FAB6" blockId="29.[151,1437,1124,1364]" pageId="29" pageNumber="536">
<emphasis id="CCEFEF3BFFF57D41A0C8FBEEFEC0FB99" bold="true" box="[151,288,1124,1148]" pageId="29" pageNumber="536">FIGURE 19.</emphasis>
Relationships of head length and upper-jaw length to standard length (SL) in
@ -731,7 +732,7 @@ from
, collected from Wuzhou,
<collectingCountry id="868C73B9FFF57D41A1A3FA2CFDDFFA25" box="[508,575,1446,1472]" name="China" pageId="29" pageNumber="536">China</collectingCountry>
in 1934, retain the distinct markings on head and fins, but lack the lateral blotches on their body (
<figureCitation id="66A02FACFFF57D41A1F1FA40FDD1FA01" box="[430,561,1482,1508]" captionStart="FIGURE 20" captionStartId="30.[151,250,584,608]" captionTargetBox="[151,1435,181,550]" captionTargetId="figure-18@30.[151,1435,181,560]" captionTargetPageId="30" captionText="FIGURE 20. Type specimens of species currently considered synonyms of Rhinogobius similis Gill, 1859. a, b, CAS-SU 29089, possible types of Ctenogobius lini Herre, 1934 (a, 95.4 mm SL; b, 94.5 mm SL; one of them should be the holotype and the other is a paratype); c, CAS-SU 4990, holotype of Gobius giurinus Rutter, 1897 (63.0 mm SL); d, CAS-SU 6449, holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 (45.3 mm SL)." pageId="29" pageNumber="536">Fig. 20a, b</figureCitation>
<figureCitation id="66A02FACFFF57D41A1F1FA40FDD1FA01" box="[430,561,1482,1508]" captionStart="FIGURE 20" captionStartId="30.[151,250,584,608]" captionTargetBox="[151,1435,181,550]" captionTargetId="figure-18@30.[151,1435,181,560]" captionTargetPageId="30" captionText="FIGURE 20. Type specimens of species currently considered synonyms of Rhinogobius similis Gill, 1859. a, b, CAS-SU 29089, possible types of Ctenogobius lini Herre, 1934 (a, 95.4 mm SL; b, 94.5 mm SL; one of them should be the holotype and the other is a paratype); c, CAS-SU 4990, holotype of Gobius giurinus Rutter, 1897 (63.0 mm SL); d, CAS-SU 6449, holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 (45.3 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.13330613" httpUri="https://zenodo.org/record/13330613/files/figure.png" pageId="29" pageNumber="536">Fig. 20a, b</figureCitation>
). In the
<typeStatus id="21208D8BFFF57D41A2C8FA40FD19FA01" box="[663,761,1482,1508]" pageId="29" pageNumber="536" type="holotype">holotype</typeStatus>
of
@ -742,11 +743,11 @@ of
, collected from Shantou,
<collectingCountry id="868C73B9FFF57D41A0C8FA64FF3AF9ED" box="[151,218,1518,1544]" name="China" pageId="29" pageNumber="536">China</collectingCountry>
before 1885, the presence of lateral blotches is unknown (because all markings are unclear although small melanophores are visible) (
<figureCitation id="66A02FACFFF57D41A197F998FDC9F9C9" box="[456,553,1554,1580]" captionStart="FIGURE 20" captionStartId="30.[151,250,584,608]" captionTargetBox="[151,1435,181,550]" captionTargetId="figure-18@30.[151,1435,181,560]" captionTargetPageId="30" captionText="FIGURE 20. Type specimens of species currently considered synonyms of Rhinogobius similis Gill, 1859. a, b, CAS-SU 29089, possible types of Ctenogobius lini Herre, 1934 (a, 95.4 mm SL; b, 94.5 mm SL; one of them should be the holotype and the other is a paratype); c, CAS-SU 4990, holotype of Gobius giurinus Rutter, 1897 (63.0 mm SL); d, CAS-SU 6449, holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 (45.3 mm SL)." pageId="29" pageNumber="536">Fig. 20c</figureCitation>
<figureCitation id="66A02FACFFF57D41A197F998FDC9F9C9" box="[456,553,1554,1580]" captionStart="FIGURE 20" captionStartId="30.[151,250,584,608]" captionTargetBox="[151,1435,181,550]" captionTargetId="figure-18@30.[151,1435,181,560]" captionTargetPageId="30" captionText="FIGURE 20. Type specimens of species currently considered synonyms of Rhinogobius similis Gill, 1859. a, b, CAS-SU 29089, possible types of Ctenogobius lini Herre, 1934 (a, 95.4 mm SL; b, 94.5 mm SL; one of them should be the holotype and the other is a paratype); c, CAS-SU 4990, holotype of Gobius giurinus Rutter, 1897 (63.0 mm SL); d, CAS-SU 6449, holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 (45.3 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.13330613" httpUri="https://zenodo.org/record/13330613/files/figure.png" pageId="29" pageNumber="536">Fig. 20c</figureCitation>
), but the head is as small as in the specimens from northern
<collectingCountry id="868C73B9FFF57D41A492F998FACCF9C9" box="[1229,1324,1554,1580]" name="Vietnam" pageId="29" pageNumber="536">Vietnam</collectingCountry>
(
<figureCitation id="66A02FACFFF57D41A564F998FA6FF9C9" box="[1339,1423,1554,1580]" captionStart="FIGURE 19" captionStartId="29.[151,250,1124,1148]" captionTargetBox="[158,1429,532,1093]" captionTargetId="figure-186@29.[151,1436,525,1100]" captionTargetPageId="29" captionText="FIGURE 19. Relationships of head length and upper-jaw length to standard length (SL) in Rhinogobius similis collected from Da Nang City (light green squares) and Thua Thien Hue Province (orange square) in central Vietnam, Quang Ninh Province in northern Vietnam (red circles), Okinawa Island (light blue circles) and Izu Peninsula (deep blue diamonds) in Japan, holotype and paratypes of Ctenogobius lini Herre, 1934 from Wuzhou, China (deep green x), holotype of Gobius giurinus Rutter, 1897 from Shantou, China (pink x), and holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 from Nagasaki, Japan (deep blue x). Solid symbols represent males, open symbols represent females, and those filled with grey represent specimens of unknown sex." pageId="29" pageNumber="536">Fig. 19</figureCitation>
<figureCitation id="66A02FACFFF57D41A564F998FA6FF9C9" box="[1339,1423,1554,1580]" captionStart="FIGURE 19" captionStartId="29.[151,250,1124,1148]" captionTargetBox="[158,1429,532,1093]" captionTargetId="figure-186@29.[151,1436,525,1100]" captionTargetPageId="29" captionText="FIGURE 19. Relationships of head length and upper-jaw length to standard length (SL) in Rhinogobius similis collected from Da Nang City (light green squares) and Thua Thien Hue Province (orange square) in central Vietnam, Quang Ninh Province in northern Vietnam (red circles), Okinawa Island (light blue circles) and Izu Peninsula (deep blue diamonds) in Japan, holotype and paratypes of Ctenogobius lini Herre, 1934 from Wuzhou, China (deep green x), holotype of Gobius giurinus Rutter, 1897 from Shantou, China (pink x), and holotype of Ctenogobius hadropterus Jordan &amp; Snyder, 1901 from Nagasaki, Japan (deep blue x). Solid symbols represent males, open symbols represent females, and those filled with grey represent specimens of unknown sex." figureDoi="http://doi.org/10.5281/zenodo.13330611" httpUri="https://zenodo.org/record/13330611/files/figure.png" pageId="29" pageNumber="536">Fig. 19</figureCitation>
). These observations suggest that morphology of specimens from
<collectingCountry id="868C73B9FFF57D41A33EF9BCFC44F9B5" box="[865,932,1590,1616]" name="China" pageId="29" pageNumber="536">China</collectingCountry>
is similar to those from

139
data/76/32/82/7632823FFFF87D49A0C8FDBFFEA9FDD8.xml
View file

@ -1,69 +1,70 @@
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<figureCitation id="66A02FACFFF87D4CA0C0FDD3FF0FFD96" box="[159,239,601,627]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="16" pageNumber="523">
Figs.
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<figureCitation id="66A02FACFFF87D4CA0A4FDD3FECFFD96" box="[251,303,601,627]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." pageId="16" pageNumber="523">6e, f</figureCitation>
<figureCitation id="66A02FACFFF87D4CA0A4FDD3FECFFD96" box="[251,303,601,627]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="16" pageNumber="523">6e, f</figureCitation>
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<figureCitation id="66A02FACFFF87D4CA164FDD3FE9FFD96" box="[315,383,601,627]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="16" pageNumber="523">10a–c</figureCitation>
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<figureCitation id="66A02FACFFF87D4CA1D4FDD3FE48FD96" box="[395,424,601,627]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="16" pageNumber="523">11</figureCitation>
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<figureCitation id="66A02FACFFF87D4CA1EBFDD3FE33FD96" box="[436,467,601,627]" captionStart="FIGURE 12" captionStartId="20.[151,250,1061,1085]" captionTargetBox="[151,1435,181,1037]" captionTargetId="figure-18@20.[151,1435,181,1037]" captionTargetPageId="20" captionText="FIGURE 12. In situ underwater photographs of Rhinogobius taenigena in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietnam. a, b, males, 20 June 2019; c, d, females, 20 June 2019 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330593" httpUri="https://zenodo.org/record/13330593/files/figure.png" pageId="16" pageNumber="523">12</figureCitation>
;
<tableCitation id="B3190692FFF87D4CA180FDD3FDDCFD96" box="[479,572,601,627]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="16" pageNumber="523">Tables 1</tableCitation>
,
@ -341,17 +342,17 @@ Song Bo in
<paragraph id="FE243329FFF87D4CA098FAC7FDC3F80A" blockId="16.[151,1437,673,2032]" pageId="16" pageNumber="523">
<emphasis id="CCEFEF3BFFF87D4CA098FAC7FEB9FA82" bold="true" box="[199,345,1357,1383]" pageId="16" pageNumber="523">Description.</emphasis>
Body nearly cylindrical anteriorly and somewhat compressed posteriorly. Anterior tip of head pointed in female, but rounded in male. Head larger in male than female (head length 30.9–33.5 vs. 28.1–30.5% of SL;
<figureCitation id="66A02FACFFF87D4CA0B7FA1FFECDFA4A" box="[232,301,1429,1455]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="16" pageNumber="523">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFF87D4CA0B7FA1FFECDFA4A" box="[232,301,1429,1455]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="16" pageNumber="523">Fig. 2</figureCitation>
,
<tableCitation id="B3190692FFF87D4CA164FA1FFE71FA4A" box="[315,401,1429,1455]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="16" pageNumber="523">Table 9</tableCitation>
). Eyes located dorsolaterally. Mouth terminal and oblique with thick upper and lower lips. When mouth closed, anterior tips of upper and lower jaws almost even, but lower jaw sometimes slightly protruding beyond upper jaw. Upper jaw extending posteriorly to about anterior margin of pupil. Mouth larger in male than female (upper-jaw length 10.2–14.6 vs. 8.5–9.6% of SL;
<figureCitation id="66A02FACFFF87D4CA37FF98BFC84F9FE" box="[800,868,1537,1563]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="16" pageNumber="523">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFF87D4CA37FF98BFC84F9FE" box="[800,868,1537,1563]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="16" pageNumber="523">Fig. 2</figureCitation>
,
<tableCitation id="B3190692FFF87D4CA32EF98BFC27F9FE" box="[881,967,1537,1563]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="16" pageNumber="523">Table 9</tableCitation>
). Upper and lower jaws usually with 2–4 rows of conical teeth; those in outermost row enlarged and those in inner rows irregularly aligned. Mental flap on chin usually rectangular or trapezoidal (posterior end forming long side), often with a small central projection. Anterior nostril short and tubular, posterior nostril a pore. Cephalic sensory pore system with B´, C, D(S), E, F, H´, K´, and L´ in oculoscapular canal and M´, N, and O´ in preopercular canal of both left and right sides in 11 of
<specimenCount id="E89DF8A0FFF87D4CA5DFF91BFEECF92A" count="18" pageId="16" pageNumber="523" type="generic">18 specimens</specimenCount>
examined (
<figureCitation id="66A02FACFFF87D4CA1D1F93FFDE9F92A" box="[398,521,1717,1743]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." pageId="16" pageNumber="523">Fig. 10a–c</figureCitation>
<figureCitation id="66A02FACFFF87D4CA1D1F93FFDE9F92A" box="[398,521,1717,1743]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="16" pageNumber="523">Fig. 10a–c</figureCitation>
), but
<specimenCount id="E89DF8A0FFF87D4CA219F93CFD3FF92A" box="[582,735,1717,1744]" count="1" pageId="16" pageNumber="523" type="generic">one specimen</specimenCount>
having an additional pore behind pore F (not G, but near F) on left side,
@ -367,7 +368,7 @@ having a single heart-shaped pore C,
having two pores aligned longitudinally on one side of C (total three pores at C) and oculoscapular canal of one side without pore M´ and with vertical groove-like N´ instead in one of
<specimenCount id="E89DF8A0FFF87D4CA0C8F8E0FEA1F866" box="[151,321,1897,1924]" count="2" pageId="16" pageNumber="523" type="generic">two specimens</specimenCount>
, and oculoscapular canal of one side lacking N in another one. Arrangement of cutaneous sensory papillae of head shown in
<figureCitation id="66A02FACFFF87D4CA196F807FDA6F842" box="[457,582,1933,1959]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." pageId="16" pageNumber="523">Fig. 10a–c</figureCitation>
<figureCitation id="66A02FACFFF87D4CA196F807FDA6F842" box="[457,582,1933,1959]" captionStart="FIGURE 10" captionStartId="18.[151,250,1352,1376]" captionTargetBox="[160,1427,188,1320]" captionTargetId="figure-18@18.[151,1436,181,1329]" captionTargetPageId="18" captionText="FIGURE 10. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius taenigena (a–c, OISTICH 303) and Rhinogobius leavelli (d–f, NSMT-P 147888). a, d, lateral view; b, e, dorsal view; c, f, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="16" pageNumber="523">Fig. 10a–c</figureCitation>
. Cheek having no significant transverse papilla rows. Vertebrae 10+16=26 (n=1), 10+17=27 (n=11), 10+18=28 (n=1), or 11+17=28 (n=1;
<tableCitation id="B3190692FFF87D4CA300F83BFC52F82E" box="[863,946,1969,1995]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="16" pageNumber="523">Table 1</tableCitation>
), P-V 3/II II I I 0/9 (n=12), P-V 3/II II I I I 0/10 (n=1), or 3/II III 0 I 0/9 (n=1).
@ -625,7 +626,7 @@ with seven spines. Second dorsal fin with one spine and seven (n=4), eight (n=11
). In females, first dorsal fin rounded, almost semi-circular; posterior tip of fin (usually tips of sixth spines) not reaching second dorsal-fin origin. In males, first dorsal fin rounded or parallelogram; posterior tip of fin (usually fourth or fifth spine) extending to, or exceeding base of second dorsal-fin spine, but never exceeding base of second soft ray of second dorsal fin. Anal fin with one spine and six (n=2), seven (n=10), or eight (n=6) soft rays (
<tableCitation id="B3190692FFF97D4DA20FFA16FD48FA53" box="[592,680,1436,1462]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="17" pageNumber="524">Table 2</tableCitation>
). Caudal fin with 17 segmented rays including 12 (n=3), 13 (n=2), 14 (n=12), or 15 (n=1) branched rays; posterior margin rounded. Males having larger second dorsal, anal, and caudal fins, than females (second dorsal-fin length 28.8–36.1 vs. 25.7–30.8% of SL, anal-fin length 25.0–33.0 vs. 22.0–28.4% of SL, and caudal-fin length 24.3–28.7 vs. 23.3–25.9% of SL;
<figureCitation id="66A02FACFFF97D4DA381F982FBC1F9C7" box="[990,1057,1544,1570]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." pageId="17" pageNumber="524">Fig. 2</figureCitation>
<figureCitation id="66A02FACFFF97D4DA381F982FBC1F9C7" box="[990,1057,1544,1570]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="17" pageNumber="524">Fig. 2</figureCitation>
,
<tableCitation id="B3190692FFF97D4DA473F982FB61F9C7" box="[1068,1153,1544,1570]" captionStart="TABLE 9" captionStartId="17.[151,239,151,175]" captionText="TABLE 9. Morphometric data of Rhinogobius taenigena, R. leavelli, and R. similis from central Vietnam, expressed as percentages of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P2, pelvic fin. * Includes five specimens of unknown sex." pageId="17" pageNumber="524">Table 9</tableCitation>
). Pectoral fin with 15–17 rays (mode 16 rays) (
@ -643,7 +644,7 @@ Suzuki
<tableCitation id="B3190692FFF97D4DA1C8F8FAFDF6F86E" box="[407,534,1904,1931]" captionStart-0="TABLE 5" captionStart-1="TABLE 6" captionStart-2="TABLE 7" captionStartId-0="7.[151,234,868,892]" captionStartId-1="8.[151,236,151,175]" captionStartId-2="8.[151,237,868,892]" captionText-0="TABLE 5. Longitudinal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-1="TABLE 6. Transverse scale counts of various species of Rhinogobius examined in the present study. *Includes the holotype." captionText-2="TABLE 7. Predorsal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="17" pageNumber="524">Tables 5–7</tableCitation>
).
</paragraph>
<caption id="AAE463A1FFFA7D4EA0C8FAC2FEBDFA4D" pageId="18" pageNumber="525" startId="18.[151,250,1352,1376]" targetBox="[160,1427,188,1320]" targetPageId="18" targetType="figure">
<caption id="AAE463A1FFFA7D4EA0C8FAC2FEBDFA4D" ID-DOI="http://doi.org/10.5281/zenodo.13330589" ID-Zenodo-Dep="13330589" httpUri="https://zenodo.org/record/13330589/files/figure.png" pageId="18" pageNumber="525" startId="18.[151,250,1352,1376]" targetBox="[160,1427,188,1320]" targetPageId="18" targetType="figure">
<paragraph id="FE243329FFFA7D4EA0C8FAC2FEBDFA4D" blockId="18.[151,1436,1352,1448]" pageId="18" pageNumber="525">
<emphasis id="CCEFEF3BFFFA7D4EA0C8FAC2FEC0FA85" bold="true" box="[151,288,1352,1376]" pageId="18" pageNumber="525">FIGURE 10.</emphasis>
Arrangement of cephalic sensory pores and cutaneous sensory papillae in
@ -659,19 +660,19 @@ Arrangement of cephalic sensory pores and cutaneous sensory papillae in
</caption>
<paragraph id="FE243329FFFA7D4EA098FA5DFDDFF8F4" blockId="18.[151,1437,1495,2025]" pageId="18" pageNumber="525">
Color in life and fresh (
<figureCitation id="66A02FACFFFA7D4EA190FA5DFDC9FA14" box="[463,553,1495,1521]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." pageId="18" pageNumber="525">Figs. 11</figureCitation>
<figureCitation id="66A02FACFFFA7D4EA190FA5DFDC9FA14" box="[463,553,1495,1521]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="18" pageNumber="525">Figs. 11</figureCitation>
and
<figureCitation id="66A02FACFFFA7D4EA23FFA5DFD60FA14" box="[608,640,1495,1521]" captionStart="FIGURE 12" captionStartId="20.[151,250,1061,1085]" captionTargetBox="[151,1435,181,1037]" captionTargetId="figure-18@20.[151,1435,181,1037]" captionTargetPageId="20" captionText="FIGURE 12. In situ underwater photographs of Rhinogobius taenigena in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietnam. a, b, males, 20 June 2019; c, d, females, 20 June 2019 (photo by K. Maeda)." pageId="18" pageNumber="525">12</figureCitation>
<figureCitation id="66A02FACFFFA7D4EA23FFA5DFD60FA14" box="[608,640,1495,1521]" captionStart="FIGURE 12" captionStartId="20.[151,250,1061,1085]" captionTargetBox="[151,1435,181,1037]" captionTargetId="figure-18@20.[151,1435,181,1037]" captionTargetPageId="20" captionText="FIGURE 12. In situ underwater photographs of Rhinogobius taenigena in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietnam. a, b, males, 20 June 2019; c, d, females, 20 June 2019 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330593" httpUri="https://zenodo.org/record/13330593/files/figure.png" pageId="18" pageNumber="525">12</figureCitation>
): in males, background of head and body grey or yellowish-brown, but whitish on ventral side. Reddish-brown stripe connecting posterior margin of center of upper lip and anterior margin of left and right eyes; thinner and unclear stripe continuing posteriorly from posterior margin of eye, but ending immediately and not reaching posterior margin of preopercle. No or faint reddish-brown marking at center of upper lip. Dorsal side of head often darker than lateral side. Lower half of cheek and opercular region often darker than upper half. Opercular membrane, interopercle, and lower part of subopercle light bluish-grey with many small red or orange spots. Six to eight, dark-brown transverse bands regularly aligned along lateral side of body from flank to caudal-fin base, and intervals light blue, but this pattern sometimes irregular and unclear. Many small, orange spots sometimes scattered on sides of body.
</paragraph>
<paragraph id="FE243329FFFA7D48A098F891FD55FAF3" blockId="18.[151,1437,1495,2025]" lastBlockId="20.[151,1437,1168,2022]" lastPageId="20" lastPageNumber="527" pageId="18" pageNumber="525">
First dorsal-fin with series of black or reddish-brown markings on middle parts of membranes between first to fifth spines; anterior two markings (between first and second, and second and third spines) usually black and clear, but posterior ones reddish-brown and often unclear. Distal part of membrane light blue or white between first and second spines, and orange or red between second and fifth spines. Distal parts of spines black or grey. Reddish-brown spots on proximal part of first to fourth spines; spines and membranes light bluish-grey, both above and below reddish-brown spots. Membranes posterior to fifth spine light bluish-grey with some reddish-brown spots. Second dorsal-fin reddish-brown with four to seven light bluish-grey stripes across fin, and with a broad, orange or pale-yellow band at distal margin. A black, vertical rectangular or elliptical spot at middle proximal part of caudal fin, with a yellowish-brown or grey blotch above black spot, but this blotch often unclear. Other part of caudal fin yellowish or greyish translucent with five or six reddish-brown vertical bands with a pale-yellow margin (
<figureCitation id="66A02FACFFFB7D4FA56EFE89FA6FFEF8" box="[1329,1423,259,285]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." pageId="19" pageNumber="526">Fig. 11b</figureCitation>
<figureCitation id="66A02FACFFFB7D4FA56EFE89FA6FFEF8" box="[1329,1423,259,285]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="19" pageNumber="526">Fig. 11b</figureCitation>
), or dusky grey or reddish-brown with a pale-yellow margin without distinct vertical bands (
<figureCitation id="66A02FACFFFB7D4FA42EFEADFB2DFEA4" box="[1137,1229,295,321]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." pageId="19" pageNumber="526">Fig. 11a</figureCitation>
<figureCitation id="66A02FACFFFB7D4FA42EFEADFB2DFEA4" box="[1137,1229,295,321]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="19" pageNumber="526">Fig. 11a</figureCitation>
). Anal fin orange, but dark brown distally with broad, bluish-white margin. Pectoral fin grey or greyish translucent, its proximal part pale yellowish-brown or pale grey with a black spot at middle of base; an additional black spot often appearing at lower part of pectoral-fin base, which is sometimes combined with middle one forming a large black blotch. Pelvic disc dark grey with a white or light blue margin.
</paragraph>
<caption id="AAE463A1FFFB7D4FA0C8F83AFBA0F809" pageId="19" pageNumber="526" startId="19.[151,250,1968,1992]" targetBox="[151,1435,337,1952]" targetPageId="19" targetType="figure">
<caption id="AAE463A1FFFB7D4FA0C8F83AFBA0F809" ID-DOI="http://doi.org/10.5281/zenodo.13330591" ID-Zenodo-Dep="13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="19" pageNumber="526" startId="19.[151,250,1968,1992]" targetBox="[151,1435,337,1952]" targetPageId="19" targetType="figure">
<paragraph id="FE243329FFFB7D4FA0C8F83AFBA0F809" blockId="19.[151,1436,1968,2028]" pageId="19" pageNumber="526">
<emphasis id="CCEFEF3BFFFB7D4FA0C8F83AFEFFF82D" bold="true" box="[151,287,1968,1992]" pageId="19" pageNumber="526">FIGURE 11.</emphasis>
<taxonomicName id="399B48AAFFFB7D4FA17AF83AFDF0F82D" authorityName="Chen, Kottelat &amp; Miller" authorityYear="1999" box="[293,528,1968,1992]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="526" phylum="Chordata" rank="species" species="taenigena">
@ -680,7 +681,7 @@ First dorsal-fin with series of black or reddish-brown markings on middle parts
immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL.
</paragraph>
</caption>
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<paragraph id="FE243329FFFC7D48A0C8FBAFFAEFFB87" blockId="20.[151,1437,1061,1122]" pageId="20" pageNumber="527">
<emphasis id="CCEFEF3BFFFC7D48A0C8FBAFFEC1FBD8" bold="true" box="[151,289,1061,1085]" pageId="20" pageNumber="527">FIGURE 12.</emphasis>
<emphasis id="CCEFEF3BFFFC7D48A176FBACFE8BFBD8" box="[297,363,1062,1085]" italics="true" pageId="20" pageNumber="527">In situ</emphasis>
@ -694,7 +695,7 @@ in the upper reaches of Xe Kong (Mekong basin) in Thua Thien Hue Province, Vietn
<paragraph id="FE243329FFFC7D48A098FAAAFDB8F9D3" blockId="20.[151,1437,1168,2022]" pageId="20" pageNumber="527">In females, markings and background color of head and body similar to those of males, but opercular membrane, interopercle, and lower part of subopercle yellowish-brown or dark grey without red spots. First dorsal fin having two to four, indefinite brown stripes; intervals translucent or white; distal part of fin yellow or bluish-white. Second dorsal fin with four to five, brown stripes, intervals white or translucent; with a yellow or bluish-white margin. Caudal fin translucent or white with five or six brown bands with a thin, pale-yellow margin; color and markings of proximal part of fin similar to those in males. Anal fin translucent, but reddish-brown or dark brown distally with a bluish-white margin. Pectoral fin translucent with a pattern on its base similar to that in males. Pelvic disc translucent or grey with a white margin.</paragraph>
<paragraph id="FE243329FFFC7D48A098F9CAFD67F90F" blockId="20.[151,1437,1168,2022]" pageId="20" pageNumber="527">
Color in preservative (
<figureCitation id="66A02FACFFFC7D48A196F9CAFDD8F9BF" box="[457,568,1600,1626]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." pageId="20" pageNumber="527">Fig. 6e, f</figureCitation>
<figureCitation id="66A02FACFFFC7D48A196F9CAFDD8F9BF" box="[457,568,1600,1626]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="20" pageNumber="527">Fig. 6e, f</figureCitation>
): background of head and body cream and those of fins translucent or light brown. Reddish-brown markings observed in life become dark brown, and dusky-brown markings become black. Light bluish-grey color on opercular membrane, interopercle, and lower part of subopercle of male become dark brown, and red and orange spots become light brown (dark and pale pattern opposite from that in life). Red, orange, white, and bluish colors of fins disappeared.
</paragraph>
</subSubSection>
@ -718,11 +719,11 @@ Chen
, it is close to the border with
<collectingCountry id="868C73B9FFFC7D48A405F892FB59F8D7" box="[1114,1209,1816,1842]" name="Vietnam" pageId="20" pageNumber="527">Vietnam</collectingCountry>
(a little downstream from the Xe Pon reaches shown in
<figureCitation id="66A02FACFFFC7D48A274F8B6FD8DF8B3" box="[555,621,1852,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." pageId="20" pageNumber="527">Fig. 1</figureCitation>
<figureCitation id="66A02FACFFFC7D48A274F8B6FD8DF8B3" box="[555,621,1852,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." figureDoi="http://doi.org/10.5281/zenodo.13330563" httpUri="https://zenodo.org/record/13330563/files/figure.png" pageId="20" pageNumber="527">Fig. 1</figureCitation>
). This species was also reported from
<collectingRegion id="3C5FFDCBFFFC7D48A475F8B6FAC0F8B3" box="[1066,1312,1852,1878]" country="Vietnam" name="Quang Binh" pageId="20" pageNumber="527">Quang Binh Province</collectingRegion>
(
<figureCitation id="66A02FACFFFC7D48A56FF8B6FA92F8B3" box="[1328,1394,1852,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." pageId="20" pageNumber="527">Fig. 1</figureCitation>
<figureCitation id="66A02FACFFFC7D48A56FF8B6FA92F8B3" box="[1328,1394,1852,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." figureDoi="http://doi.org/10.5281/zenodo.13330563" httpUri="https://zenodo.org/record/13330563/files/figure.png" pageId="20" pageNumber="527">Fig. 1</figureCitation>
) in
<collectingCountry id="868C73B9FFFC7D48A0C8F8EAFF16F89F" box="[151,246,1888,1914]" name="Vietnam" pageId="20" pageNumber="527">Vietnam</collectingCountry>
from the Gianh River basin to Phong Nha River (
@ -734,7 +735,7 @@ Serov
). We found this species in two river systems in Thua Thien Hue Province of
<collectingCountry id="868C73B9FFFC7D48A23CF80EFD22F87B" box="[611,706,1924,1950]" name="Vietnam" pageId="20" pageNumber="527">Vietnam</collectingCountry>
(
<figureCitation id="66A02FACFFFC7D48A28CF80EFCF5F87B" box="[723,789,1924,1950]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." pageId="20" pageNumber="527">Fig. 1</figureCitation>
<figureCitation id="66A02FACFFFC7D48A28CF80EFCF5F87B" box="[723,789,1924,1950]" captionStart="FIGURE 1" captionStartId="1.[151,250,669,693]" captionTargetBox="[151,1435,181,645]" captionTargetId="figure-17@1.[151,1435,181,645]" captionTargetPageId="1" captionText="FIGURE 1. Map of South East Asia showing Vietnam, the Mekong and the Red River basins (left), and collection sites for Rhinogobius in Da Nang City and Thua Thien Hue Province in central Vietnam (orange dashed circles, right). Symbols below or above orange dashed circles indicate the occurrence of R. rong (red circles), R. nami (light blue circle), R. taenigena (deep blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) at their respective locations." figureDoi="http://doi.org/10.5281/zenodo.13330563" httpUri="https://zenodo.org/record/13330563/files/figure.png" pageId="20" pageNumber="527">Fig. 1</figureCitation>
). One of them, Xe Kong (Se Kong), is a tributary of the Mekong, originating in Thua Thien Hue Province and flowing through southern
<collectingCountry id="868C73B9FFFC7D48A450F822FBA5F827" box="[1039,1093,1960,1986]" name="Laos" pageId="20" pageNumber="527">Laos</collectingCountry>
and eastern

661
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<taxonomicName id="388A73D1E9014A4FFF3CFDB9FDCFFD1C" ID-CoL="7BBLJ" authority="(Walker, 1874)" baseAuthorityName="Walker" baseAuthorityYear="1874" box="[151,561,637,663]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="apicifer">
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<emphasis id="CDFED440E9014A4FFF3CFDB9FE8CFD1C" bold="true" box="[151,370,637,663]" italics="true" pageId="3" pageNumber="38">Syzeuctus apicifer</emphasis>
(
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<subSubSection id="B7905BD9E9014A4FFF3CFD65FE15FD30" box="[151,491,673,699]" pageId="3" pageNumber="38" type="description">
<paragraph id="FF350852E9014A4FFF3CFD65FE15FD30" blockId="3.[151,561,637,699]" box="[151,491,673,699]" pageId="3" pageNumber="38">
(
<figureCitation id="67B114D7E9014A4FFF34FD65FEFCFD30" box="[159,258,673,699]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="3" pageNumber="38">Figs 1 A</figureCitation>
,
<figureCitation id="67B114D7E9014A4FFEA6FD65FEC6FD30" box="[269,312,673,699]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="3" pageNumber="38">2 A</figureCitation>
,
<figureCitation id="67B114D7E9014A4FFEE8FD65FE73FD30" box="[323,397,673,699]" captionStart="FIGURE 3" captionStartId="6.[151,250,1534,1558]" captionTargetBox="[151,1436,181,1510]" captionTargetId="figure-19@6.[151,1436,181,1510]" captionTargetPageId="6" captionText="FIGURE 3. Syzeuctus apicifer (Walker, 1874), female from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="3" pageNumber="38">3 A–E</figureCitation>
,
<figureCitation id="67B114D7E9014A4FFE33FD65FE1DFD30" box="[408,483,673,699]" captionStart="FIGURE 4" captionStartId="7.[151,250,1621,1645]" captionTargetBox="[151,1436,345,1597]" captionTargetId="figure-103@7.[151,1436,345,1597]" captionTargetPageId="7" captionText="FIGURE 4. Syzeuctus apicifer (Walker, 1874), male from Japan—A: lateral habitus; B: dorsal habitus; C, D: head, frontal view." pageId="3" pageNumber="38">4 A–D</figureCitation>
)
</paragraph>
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<subSubSection id="B7905BD9E9014A4FFF3CFD22FDFFFD76" box="[151,513,741,766]" pageId="3" pageNumber="38" type="reference_group">
<paragraph id="FF350852E9014A4FFF3CFD22FDFFFD76" blockId="3.[151,513,741,766]" box="[151,513,741,766]" pageId="3" pageNumber="38">
<treatmentCitationGroup id="DF9A2F7CE9014A4FFF3CFD22FDFFFD76" box="[151,513,741,766]" pageId="3" pageNumber="38">
<treatmentCitation id="7E2B2E43E9014A4FFF3CFD22FE03FD76" author="Walker, F." box="[151,509,741,766]" page="305" pageId="3" pageNumber="38" year="1874">
<taxonomicName id="388A73D1E9014A4FFF3CFD22FE03FD76" authority="Walker, 1874: 305" authorityName="Walker" authorityPageNumber="305" authorityYear="1874" box="[151,509,741,766]" class="Insecta" family="Ichneumonidae" genus="Macrus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="apicifer">
<emphasis id="CDFED440E9014A4FFF3CFD22FEC4FD75" box="[151,314,742,766]" italics="true" pageId="3" pageNumber="38">Macrus apicifer</emphasis>
<bibRefCitation id="9B1B75A3E9014A4FFEEBFD22FE03FD76" author="Walker, F." box="[320,509,741,766]" pageId="3" pageNumber="38" pagination="301 - 310" refId="ref23546" refString="Walker, F. (1874) Descriptions of some Japanese Hymenoptera. Cistula Entomologica, 1, 301 - 310." type="journal article" year="1874">Walker, 1874: 305</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9014A4FFF3CFCE8FE3CFA21" pageId="3" pageNumber="38" type="materials_examined">
<paragraph id="FF350852E9014A4FFF3CFCE8FE3CFA21" blockId="3.[151,1437,812,2027]" pageId="3" pageNumber="38">
<emphasis id="CDFED440E9014A4FFF3CFCE8FE7DFCCD" bold="true" box="[151,387,812,838]" pageId="3" pageNumber="38">Materials examined</emphasis>
.
<materialsCitation id="4FE2020FE9014A4FFE24FCE8FAF4FCCD" box="[399,1290,812,838]" collectionCode="NHMUK" country="Japan" location="JAPAN" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Hokkaido">
<emphasis id="CDFED440E9014A4FFE24FCE8FE17FCCD" bold="true" box="[399,489,812,838]" pageId="3" pageNumber="38">
<collectingCountry id="879D48C2E9014A4FFE24FCE8FE17FCCD" box="[399,489,812,838]" name="Japan" pageId="3" pageNumber="38">JAPAN</collectingCountry>
</emphasis>
: 1 F (
<typeStatus id="2031B6F0E9014A4FFD8DFCE8FDA6FCCD" box="[550,600,812,838]" pageId="3" pageNumber="38">type</typeStatus>
of
<taxonomicName id="388A73D1E9014A4FFDD1FCE9FD0BFCCD" authorityName="Walker" authorityYear="1874" box="[634,757,812,838]" class="Insecta" family="Ichneumonidae" genus="Macrus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="apicifer">
<emphasis id="CDFED440E9014A4FFDD1FCE9FD0BFCCD" box="[634,757,812,838]" italics="true" pageId="3" pageNumber="38">M. apicifer</emphasis>
</taxonomicName>
), “Japan. Coll. F. Walker. 1913-71.” (
<collectionCode id="999B9097E9014A4FFB3AFCE8FAFCFCCD" box="[1169,1282,812,838]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="38" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFAB8FCE8FC6DFCE1" collectingDate="1968-08-26" collectionCode="SEHU" collectorName="K. Kusigemati" country="Japan" location="Mt. Muine" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Hokkaido">
[
<emphasis id="CDFED440E9014A4FFAB6FCE8FA6DFCCD" bold="true" box="[1309,1427,812,838]" pageId="3" pageNumber="38">
<collectingRegion id="3D4EC6B0E9014A4FFAB6FCE8FA6DFCCD" box="[1309,1427,812,838]" country="Japan" name="Hokkaido" pageId="3" pageNumber="38">Hokkaido</collectingRegion>
</emphasis>
] 2 F &amp; 2 M,
<location id="FA555E89E9014A4FFEB4FC94FE65FCE1" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFEB4FC94FE65FCE1" box="[287,411,848,874]" country="Japan" name="Mt. Muine" pageId="3" pageNumber="38" stateProvince="Hokkaido">Mt. Muine</location>
,
<date id="8B342E92E9014A4FFE0CFC94FDB5FCE1" box="[423,587,848,874]" pageId="3" pageNumber="38" value="1968-08-26">
<collectingDate id="9B70D77AE9014A4FFE0CFC94FDB5FCE1" box="[423,587,848,874]" pageId="3" pageNumber="38" value="1968-08-26">26. VIII. 1968</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFDFCFC94FD05FCE1" box="[599,763,848,874]" pageId="3" pageNumber="38">K. Kusigemati</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFC97FC94FC75FCE1" box="[828,907,848,874]" pageId="3" pageNumber="38">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFC34FC94FE1DFC05" collectingDate="1979-08-12" collectionCode="EUM" collectorName="Y. Seiyama" country="Japan" county="Abuta County" location="Nakayama-pass" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Hokkaido">
2 F,
<collectingCounty id="165470DEE9014A4FFC64FC94FB91FCE1" box="[975,1135,848,874]" pageId="3" pageNumber="38">Abuta County</collectingCounty>
,
<location id="FA555E89E9014A4FFBD0FC94FAD1FCE1" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFBD0FC94FAD1FCE1" box="[1147,1327,848,874]" country="Japan" county="Abuta County" name="Nakayama-pass" pageId="3" pageNumber="38" stateProvince="Hokkaido">Nakayama-pass</location>
,
<date id="8B342E92E9014A4FFA90FC94FF2CFC05" pageId="3" pageNumber="38" value="1979-08-12">
<collectingDate id="9B70D77AE9014A4FFA90FC94FF2CFC05" pageId="3" pageNumber="38" value="1979-08-12">12. VIII. 1979</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFF70FCB0FEA7FC05" box="[219,345,884,910]" pageId="3" pageNumber="38">Y. Seiyama</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFE3CFCB0FE25FC05" box="[407,475,884,910]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFE45FCB0FEEDFC39" collectingDate="2016-09-10" collectionCode="KPMNH" collectorName="J. Kobayashi" country="Japan" location="Tonosawa" municipality="Mitane Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Akita">
[
<emphasis id="CDFED440E9014A4FFE5CFCB0FDAAFC05" bold="true" box="[503,596,884,910]" pageId="3" pageNumber="38">Honshu</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9014A4FFD36FCB0FCE4FC05" box="[669,794,884,910]" country="Japan" name="Akita" pageId="3" pageNumber="38">Akita Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFC8FFCB0FC45FC05" box="[804,955,884,910]" pageId="3" pageNumber="38">Mitane Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFC6EFCB0FBC7FC05" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFC6EFCB0FBC7FC05" box="[965,1081,884,910]" country="Japan" municipality="Mitane Town" name="Tonosawa" pageId="3" pageNumber="38" stateProvince="Akita">Tonosawa</location>
,
<date id="8B342E92E9014A4FFBEFFCB0FB2DFC05" box="[1092,1235,884,910]" pageId="3" pageNumber="38" value="2016-09-10">
<collectingDate id="9B70D77AE9014A4FFBEFFCB0FB2DFC05" box="[1092,1235,884,910]" pageId="3" pageNumber="38" value="2016-09-10">10. IX. 2016</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFB76FCB0FA93FC05" box="[1245,1389,884,910]" pageId="3" pageNumber="38">J. Kobayashi</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFF34FC5CFEF5FC39" box="[159,267,920,946]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFEB4FC5CFB38FC39" box="[287,1222,920,946]" collectingDate="2007-09-27" collectionCode="KPMNH" country="Japan" location="Seabura-toge" municipality="Aizuwakamatsu City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Fukushima">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFEFBFC5CFDF0FC39" box="[336,526,920,946]" country="Japan" name="Fukushima" pageId="3" pageNumber="38">Fukushima Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFDBCFC5CFCFBFC39" box="[535,773,920,946]" pageId="3" pageNumber="38">Aizuwakamatsu City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFCBAFC5CFC56FC39" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFCBAFC5CFC56FC39" box="[785,936,920,946]" country="Japan" municipality="Aizuwakamatsu City" name="Seabura-toge" pageId="3" pageNumber="38" stateProvince="Fukushima">Seabura-toge</location>
,
<date id="8B342E92E9014A4FFC18FC5CFBBCFC39" box="[947,1090,920,946]" pageId="3" pageNumber="38" value="2007-09-27">
<collectingDate id="9B70D77AE9014A4FFC18FC5CFBBCFC39" box="[947,1090,920,946]" pageId="3" pageNumber="38" value="2007-09-27">27. IX. 2007</collectingDate>
</date>
(
<collectionCode id="999B9097E9014A4FFBF9FC5CFB40FC39" box="[1106,1214,920,946]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFB7AFC5CFC6DFC5D" collectingDate="2000-07-13" collectionCode="KPMNH" collectorName="E. Katayama" country="Japan" location="Enna-skyline" municipality="Kuroiso" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Tochigi">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFAA8FC5CFA67FC39" box="[1283,1433,920,946]" country="Japan" name="Tochigi" pageId="3" pageNumber="38">Tochigi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFF3CFC78FF0AFC5D" box="[151,244,956,982]" pageId="3" pageNumber="38">Kuroiso</collectingMunicipality>
,
<location id="FA555E89E9014A4FFEAAFC78FE66FC5D" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFEAAFC78FE66FC5D" box="[257,408,956,982]" country="Japan" municipality="Kuroiso" name="Enna-skyline" pageId="3" pageNumber="38" stateProvince="Tochigi">Enna-skyline</location>
,
<date id="8B342E92E9014A4FFE0FFC78FDC1FC5D" box="[420,575,956,982]" pageId="3" pageNumber="38" value="2000-07-13">
<collectingDate id="9B70D77AE9014A4FFE0FFC78FDC1FC5D" box="[420,575,956,982]" pageId="3" pageNumber="38" value="2000-07-13">13. VII. 2000</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFDE7FC78FD20FC5D" box="[588,734,956,982]" pageId="3" pageNumber="38">E. Katayama</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFCB4FC78FC75FC5D" box="[799,907,956,982]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFC34FC78FDA8FC71" collectingDate="2000-08-04" collectionCode="KPMNH" collectorName="E. Katayama" country="Japan" location="Morura" municipality="Kuroiso Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Tochigi">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFC7AFC78FB96FC5D" box="[977,1128,956,982]" country="Japan" name="Tochigi" pageId="3" pageNumber="38">Tochigi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFBD8FC78FAE9FC5D" box="[1139,1303,956,982]" pageId="3" pageNumber="38">Kuroiso Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFA8FFC78FA85FC5D" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFA8FFC78FA85FC5D" box="[1316,1403,956,982]" country="Japan" municipality="Kuroiso Town" name="Morura" pageId="3" pageNumber="38" stateProvince="Tochigi">Morura</location>
,
<date id="8B342E92E9014A4FFA2CFC78FEF1FC71" pageId="3" pageNumber="38" value="2000-08-04">
<collectingDate id="9B70D77AE9014A4FFA2CFC78FEF1FC71" pageId="3" pageNumber="38" value="2000-08-04">4. VIII. 2000</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFEB3FC24FE59FC71" box="[280,423,992,1018]" pageId="3" pageNumber="38">E. Katayama</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFE49FC24FDB0FC71" box="[482,590,992,1018]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFDF4FC24FF15FB95" collectingDate="2015-09-22" collectionCode="EUM" collectorName="J. Okayasu" country="Japan" location="Yumoto" municipality="Nikko City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Tochigi">
2 F &amp; 2 M,
<collectingRegion id="3D4EC6B0E9014A4FFD72FC24FC92FC71" box="[729,876,992,1018]" country="Japan" name="Tochigi" pageId="3" pageNumber="38">Tochigi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFCDEFC24FC0FFC71" box="[885,1009,992,1018]" pageId="3" pageNumber="38">Nikko City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFC51FC24FBA8FC71" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFC51FC24FBA8FC71" box="[1018,1110,992,1018]" country="Japan" municipality="Nikko City" name="Yumoto" pageId="3" pageNumber="38" stateProvince="Tochigi">Yumoto</location>
,
<date id="8B342E92E9014A4FFBF4FC24FB12FC70" box="[1119,1260,992,1019]" pageId="3" pageNumber="38" value="2015-09-22">
<collectingDate id="9B70D77AE9014A4FFBF4FC24FB12FC70" box="[1119,1260,992,1019]" pageId="3" pageNumber="38" value="2015-09-22">22. IX. 2015</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFB5EFC24FA90FC71" box="[1269,1390,992,1018]" pageId="3" pageNumber="38">J. Okayasu</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFF34FBC0FF1DFB95" box="[159,227,1028,1054]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFF5DFBC0FB4CFB95" box="[246,1202,1028,1055]" collectingDate="2005-09-08" collectionCode="KPMNH" collectorName="M. Uchida" country="Japan" location="Nagatoro" municipality="Nagatoro Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Saitama">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFE8DFBC0FE3EFB95" box="[294,448,1028,1054]" country="Japan" name="Saitama" pageId="3" pageNumber="38">Saitama Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFE61FBC0FD85FB95" box="[458,635,1028,1054]" pageId="3" pageNumber="38">Nagatoro Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFD2DFBC0FD0FFB95" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFD2DFBC0FD0FFB95" box="[646,753,1028,1054]" country="Japan" municipality="Nagatoro Town" name="Nagatoro" pageId="3" pageNumber="38" stateProvince="Saitama">Nagatoro</location>
,
<date id="8B342E92E9014A4FFD57FBC0FC83FB94" box="[764,893,1028,1055]" pageId="3" pageNumber="38" value="2005-09-08">
<collectingDate id="9B70D77AE9014A4FFD57FBC0FC83FB94" box="[764,893,1028,1055]" pageId="3" pageNumber="38" value="2005-09-08">8. IX. 2005</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFC23FBC0FBFEFB95" box="[904,1024,1028,1054]" pageId="3" pageNumber="38">M. Uchida</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFB95FBC0FB54FB95" box="[1086,1194,1028,1054]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFB17FBC0FCD3FBC9" collectingDate="2009-09-01" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Ninomiya" municipality="Akiruno City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Tokyo">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFB47FBC0FAC8FB95" box="[1260,1334,1028,1054]" country="Japan" name="Tokyo" pageId="3" pageNumber="38">Tokyo</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFA94FBC0FF36FBC9" pageId="3" pageNumber="38">Akiruno City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFF7EFBECFEB6FBC9" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFF7EFBECFEB6FBC9" box="[213,328,1064,1090]" country="Japan" municipality="Akiruno City" name="Ninomiya" pageId="3" pageNumber="38" stateProvince="Tokyo">Ninomiya</location>
,
<date id="8B342E92E9014A4FFEFFFBECFE24FBC9" box="[340,474,1064,1090]" pageId="3" pageNumber="38" value="2009-09-01">
<collectingDate id="9B70D77AE9014A4FFEFFFBECFE24FBC9" box="[340,474,1064,1090]" pageId="3" pageNumber="38" value="2009-09-01">1. IX. 2009</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFE4DFBECFD89FBC9" box="[486,631,1064,1090]" pageId="3" pageNumber="38">K. Watanabe</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFD12FBECFCDBFBC9" box="[697,805,1064,1090]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFC92FBECFE15FBED" collectingDate="1997-06-19" collectionCode="KPMNH" collectorName="M. Enju" country="Japan" location="Nanasawa" municipality="Atsugi City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Kanagawa">
1 M,
<collectingRegion id="3D4EC6B0E9014A4FFCD3FBECFBD1FBC9" box="[888,1071,1064,1090]" country="Japan" name="Kanagawa" pageId="3" pageNumber="38">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFB91FBECFB40FBC9" box="[1082,1214,1064,1090]" pageId="3" pageNumber="38">Atsugi City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFB61FBECFABEFBC9" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFB61FBECFABEFBC9" box="[1226,1344,1064,1090]" country="Japan" municipality="Atsugi City" name="Nanasawa" pageId="3" pageNumber="38" stateProvince="Kanagawa">Nanasawa</location>
,
<date id="8B342E92E9014A4FFAE6FBECFF2CFBED" pageId="3" pageNumber="38" value="1997-06-19">
<collectingDate id="9B70D77AE9014A4FFAE6FBECFF2CFBED" pageId="3" pageNumber="38" value="1997-06-19">19. VI. 1997</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFF76FB88FEC7FBED" box="[221,313,1100,1126]" pageId="3" pageNumber="38">M. Enju</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFED3FB88FE1AFBED" box="[376,484,1100,1126]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFE5DFB88FEEDFB01" collectingDate="2009-09-13" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Mt. Jinba-yama" municipality="Fujino Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Kanagawa">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFD8DFB88FD23FBED" box="[550,733,1100,1126]" country="Japan" name="Kanagawa" pageId="3" pageNumber="38">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFD4CFB88FC87FBED" box="[743,889,1100,1126]" pageId="3" pageNumber="38">Fujino Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFC2FFB88FBC9FBED" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFC2FFB88FBC9FBED" box="[900,1079,1100,1126]" country="Japan" municipality="Fujino Town" name="Mt. Jinba-yama" pageId="3" pageNumber="38" stateProvince="Kanagawa">Mt. Jinba-yama</location>
,
<date id="8B342E92E9014A4FFBE9FB88FB2CFBED" box="[1090,1234,1100,1126]" pageId="3" pageNumber="38" value="2009-09-13">
<collectingDate id="9B70D77AE9014A4FFBE9FB88FB2CFBED" box="[1090,1234,1100,1126]" pageId="3" pageNumber="38" value="2009-09-13">13. IX. 2009</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFB76FB88FA92FBED" box="[1245,1388,1100,1126]" pageId="3" pageNumber="38">K. Watanabe</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFF34FBB4FEF5FB01" box="[159,267,1136,1162]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFEB4FBB4FB3FFB01" box="[287,1217,1136,1163]" collectingDate="2004-08-05" collectingDateMax="2004-08-07" collectingDateMin="2004-08-05" collectionCode="KPMNH" collectorName="S. Motoi" country="Japan" location="Shinkai" municipality="Kiso Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Nagano">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFEF9FBB4FE15FB01" box="[338,491,1136,1162]" country="Japan" name="Nagano" pageId="3" pageNumber="38">Nagano Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFE5CFBB4FD88FB01" box="[503,630,1136,1162]" pageId="3" pageNumber="38">Kiso Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFD28FBB4FD25FB01" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFD28FBB4FD25FB01" box="[643,731,1136,1162]" country="Japan" municipality="Kiso Town" name="Shinkai" pageId="3" pageNumber="38" stateProvince="Nagano">Shinkai</location>
,
<date id="8B342E92E9014A4FFD4CFBB4FC65FB01" box="[743,923,1136,1163]" pageId="3" pageNumber="38" value="2004-08-05" valueMax="2004-08-07" valueMin="2004-08-05">
<collectingDate id="9B70D77AE9014A4FFD4CFBB4FC65FB01" box="[743,923,1136,1163]" pageId="3" pageNumber="38" value="2004-08-05" valueMax="2004-08-07" valueMin="2004-08-05">5–7. VIII. 2004</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFC0CFBB4FBF4FB01" box="[935,1034,1136,1162]" pageId="3" pageNumber="38">S. Motoi</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFBE6FBB4FB47FB01" box="[1101,1209,1136,1162]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFB66FBB4FC7CFB25" collectingDate="2022-08-25" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Sugadaira" municipality="Ueda City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Nagano">
8 F,
<collectingRegion id="3D4EC6B0E9014A4FFAABFBB4FA67FB01" box="[1280,1433,1136,1162]" country="Japan" name="Nagano" pageId="3" pageNumber="38">Nagano Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFF3CFB50FEF4FB25" box="[151,266,1172,1198]" pageId="3" pageNumber="38">Ueda City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFEBEFB50FE76FB25" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFEBEFB50FE76FB25" box="[277,392,1172,1198]" country="Japan" municipality="Ueda City" name="Sugadaira" pageId="3" pageNumber="38" stateProvince="Nagano">Sugadaira</location>
,
<date id="8B342E92E9014A4FFE38FB50FDCBFB25" box="[403,565,1172,1199]" pageId="3" pageNumber="38" value="2022-08-25">
<collectingDate id="9B70D77AE9014A4FFE38FB50FDCBFB25" box="[403,565,1172,1199]" pageId="3" pageNumber="38" value="2022-08-25">25. VIII. 2022</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFDEBFB50FD2EFB25" box="[576,720,1172,1198]" pageId="3" pageNumber="38">K. Watanabe</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFCA5FB50FC84FB25" box="[782,890,1172,1198]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFC26FB50FD55FB59" collectingDate="2000-08-22" collectionCode="MU" collectorName="C. Mizuno &amp; M. Suzuki" country="Japan" location="Takagi" municipality="Kasugai City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Aichi">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFC10FB50FBC7FB25" box="[955,1081,1172,1198]" country="Japan" name="Aichi" pageId="3" pageNumber="38">Aichi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFBEFFB50FB29FB25" box="[1092,1239,1172,1198]" pageId="3" pageNumber="38">Kasugai City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFB49FB50FACFFB25" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFB49FB50FACFFB25" box="[1250,1329,1172,1198]" country="Japan" municipality="Kasugai City" name="Takagi" pageId="3" pageNumber="38" stateProvince="Aichi">Takagi</location>
,
<date id="8B342E92E9014A4FFA97FB50FF2CFB59" pageId="3" pageNumber="38" value="2000-08-22">
<collectingDate id="9B70D77AE9014A4FFA97FB50FF2CFB59" pageId="3" pageNumber="38" value="2000-08-22">22. VIII. 2000</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFF76FB7CFEABFB59" box="[221,341,1208,1234]" pageId="3" pageNumber="38">C. Mizuno</collectorName>
&amp;
<collectorName id="527F6D84E9014A4FFED2FB7CFE10FB59" box="[377,494,1208,1234]" pageId="3" pageNumber="38">M. Suzuki</collectorName>
leg., MsT (
<collectionCode id="999B9097E9014A4FFDDBFB7CFD5DFB59" box="[624,675,1208,1234]" country="USA" name="Midwestern University" pageId="3" pageNumber="38">MU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFD1CFB7CFE5BFB7D" collectingDate="2000-09-20" collectionCode="MU" collectorName="C. Mizuno &amp; M. Suzuki" country="Japan" location="Johkoji" municipality="Seto City" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Aichi">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFD4DFB7CFC9AFB59" box="[742,868,1208,1234]" country="Japan" name="Aichi" pageId="3" pageNumber="38">Aichi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFCC4FB7CFC27FB59" box="[879,985,1208,1234]" pageId="3" pageNumber="38">Seto City</collectingMunicipality>
,
<location id="FA555E89E9014A4FFC4FFB7CFBC7FB59" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFC4FFB7CFBC7FB59" box="[996,1081,1208,1234]" country="Japan" municipality="Seto City" name="Johkoji" pageId="3" pageNumber="38" stateProvince="Aichi">Johkoji</location>
,
<date id="8B342E92E9014A4FFBEEFB7CFB2BFB59" box="[1093,1237,1208,1234]" pageId="3" pageNumber="38" value="2000-09-20">
<collectingDate id="9B70D77AE9014A4FFBEEFB7CFB2BFB59" box="[1093,1237,1208,1234]" pageId="3" pageNumber="38" value="2000-09-20">20. IX. 2000</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFB4AFB7CFAA6FB59" box="[1249,1368,1208,1234]" pageId="3" pageNumber="38">C. Mizuno</collectorName>
&amp;
<collectorName id="527F6D84E9014A4FFAD7FB7CFF1BFB7D" pageId="3" pageNumber="38">M. Suzuki</collectorName>
leg., MsT (
<collectionCode id="999B9097E9014A4FFEC0FB18FE60FB7D" box="[363,414,1244,1270]" country="USA" name="Midwestern University" pageId="3" pageNumber="38">MU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFE19FB18FA66FB7D" box="[434,1432,1244,1270]" collectingDate="2007-09-18" collectionCode="KPMNH" collectorName="T. Mita" country="Japan" location="Kuzunohama" municipality="Kumihama Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Kyoto">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFE4DFB18FD93FB7D" box="[486,621,1244,1270]" country="Japan" name="Kyoto" pageId="3" pageNumber="38">Kyoto Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFDD2FB18FCC1FB7D" box="[633,831,1244,1270]" pageId="3" pageNumber="38">Kumihama Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFCE7FB18FC1AFB7D" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFCE7FB18FC1AFB7D" box="[844,996,1244,1270]" country="Japan" municipality="Kumihama Town" name="Kuzunohama" pageId="3" pageNumber="38" stateProvince="Kyoto">Kuzunohama</location>
,
<date id="8B342E92E9014A4FFC5BFB18FB7AFB7D" box="[1008,1156,1244,1270]" pageId="3" pageNumber="38" value="2007-09-18">
<collectingDate id="9B70D77AE9014A4FFC5BFB18FB7AFB7D" box="[1008,1156,1244,1270]" pageId="3" pageNumber="38" value="2007-09-18">18. IX. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFB24FB18FB1DFB7D" box="[1167,1251,1244,1270]" pageId="3" pageNumber="38">T. Mita</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFA8FFB18FA6EFB7D" box="[1316,1424,1244,1270]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFF3CFAC4FBC5FA91" box="[151,1083,1280,1307]" collectingDate="1995-08-14" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Oda Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Ehime">
[
<emphasis id="CDFED440E9014A4FFF0AFAC4FEFBFA91" bold="true" box="[161,261,1280,1306]" pageId="3" pageNumber="38">Shikoku</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9014A4FFEFAFAC4FE25FA91" box="[337,475,1280,1306]" country="Japan" name="Ehime" pageId="3" pageNumber="38">Ehime Pref.</collectingRegion>
,
<location id="FA555E89E9014A4FFE4DFAC4FDA3FA91" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFE4DFAC4FDA3FA91" box="[486,605,1280,1306]" country="Japan" name="Oda Town" pageId="3" pageNumber="38" stateProvince="Ehime">Oda Town</location>
,
<date id="8B342E92E9014A4FFDC2FAC4FCF2FA90" box="[617,780,1280,1307]" pageId="3" pageNumber="38" value="1995-08-14">
<collectingDate id="9B70D77AE9014A4FFDC2FAC4FCF2FA90" box="[617,780,1280,1307]" pageId="3" pageNumber="38" value="1995-08-14">14. VIII. 1995</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFCBCFAC4FC4EFA91" box="[791,944,1280,1306]" pageId="3" pageNumber="38">E. Yamamoto</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFC44FAC4FBCDFA91" box="[1007,1075,1280,1306]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFBECFAC4FA66FA91" box="[1095,1432,1280,1307]" collectingDate="1991-10-05" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Oda Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Hokkaido">
1 M, ditto,
<date id="8B342E92E9014A4FFB6EFAC4FAC2FA91" box="[1221,1340,1280,1307]" pageId="3" pageNumber="38" value="1991-10-05">
<collectingDate id="9B70D77AE9014A4FFB6EFAC4FAC2FA91" box="[1221,1340,1280,1307]" pageId="3" pageNumber="38" value="1991-10-05">5. X. 1991</collectingDate>
</date>
(
<collectionCode id="999B9097E9014A4FFAE0FAC4FA71FA91" box="[1355,1423,1280,1306]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFF3CFAE0FBDAFAB5" box="[151,1060,1316,1342]" collectingDate="2006-08-02" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Yurano" municipality="Kumakogen" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Ehime">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFF61FAE0FEABFAB5" box="[202,341,1316,1342]" country="Japan" name="Ehime" pageId="3" pageNumber="38">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFECAFAE0FE13FAB5" box="[353,493,1316,1342]" pageId="3" pageNumber="38">Kumakogen</collectingMunicipality>
,
<location id="FA555E89E9014A4FFE53FAE0FDB2FAB5" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFE53FAE0FDB2FAB5" box="[504,588,1316,1342]" country="Japan" municipality="Kumakogen" name="Yurano" pageId="3" pageNumber="38" stateProvince="Ehime">Yurano</location>
,
<date id="8B342E92E9014A4FFDF3FAE0FD0EFAB5" box="[600,752,1316,1342]" pageId="3" pageNumber="38" value="2006-08-02">
<collectingDate id="9B70D77AE9014A4FFDF3FAE0FD0EFAB5" box="[600,752,1316,1342]" pageId="3" pageNumber="38" value="2006-08-02">2. VIII. 2006</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFD57FAE0FC68FAB5" box="[764,918,1316,1342]" pageId="3" pageNumber="38">E. Yamamoto</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFC7CFAE0FBE5FAB5" box="[983,1051,1316,1342]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFB9BFAE0FCFAFAE9" collectingDate="2006-09-27" collectionCode="EUM" collectorName="Y. Kawakami" country="Japan" location="Okufujikawa" municipality="Kihoku Town" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Ehime">
1 F,
<collectingRegion id="3D4EC6B0E9014A4FFBC8FAE0FB10FAB5" box="[1123,1262,1316,1342]" country="Japan" name="Ehime" pageId="3" pageNumber="38">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFB51FAE0FA66FAB5" box="[1274,1432,1316,1342]" pageId="3" pageNumber="38">Kihoku Town</collectingMunicipality>
,
<location id="FA555E89E9014A4FFF3CFA8CFED2FAE9" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFF3CFA8CFED2FAE9" box="[151,300,1352,1378]" country="Japan" municipality="Kihoku Town" name="Okufujikawa" pageId="3" pageNumber="38" stateProvince="Ehime">Okufujikawa</location>
,
<date id="8B342E92E9014A4FFE92FA8CFE30FAE9" box="[313,462,1352,1378]" pageId="3" pageNumber="38" value="2006-09-27">
<collectingDate id="9B70D77AE9014A4FFE92FA8CFE30FAE9" box="[313,462,1352,1378]" pageId="3" pageNumber="38" value="2006-09-27">27. IX. 2006</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFE71FA8CFD8AFAE9" box="[474,628,1352,1378]" pageId="3" pageNumber="38">Y. Kawakami</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFD13FA8CFD02FAE9" box="[696,764,1352,1378]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFCBAFA8CFDFEFA0D" collectingDate="2002-09-22" collectionCode="EUM" collectorName="Y. Kawakami" country="Japan" location="Chikaraishi" municipality="Higashitsuno Vil." pageId="3" pageNumber="38" specimenCount="1" stateProvince="Kochi">
1 M,
<collectingRegion id="3D4EC6B0E9014A4FFCF9FA8CFC26FAE9" box="[850,984,1352,1378]" country="Japan" name="Kochi" pageId="3" pageNumber="38">Kochi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFC4EFA8CFB53FAE9" box="[997,1197,1352,1378]" pageId="3" pageNumber="38">Higashitsuno Vil.</collectingMunicipality>
,
<location id="FA555E89E9014A4FFB11FA8CFAC0FAE9" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFB11FA8CFAC0FAE9" box="[1210,1342,1352,1378]" country="Japan" municipality="Higashitsuno Vil." name="Chikaraishi" pageId="3" pageNumber="38" stateProvince="Kochi">Chikaraishi</location>
,
<date id="8B342E92E9014A4FFAE7FA8CFF2CFA0D" pageId="3" pageNumber="38" value="2002-09-22">
<collectingDate id="9B70D77AE9014A4FFAE7FA8CFF2CFA0D" pageId="3" pageNumber="38" value="2002-09-22">22. IX. 2002</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFF77FAA8FE8AFA0D" box="[220,372,1388,1414]" pageId="3" pageNumber="38">Y. Kawakami</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFE1FFAA8FE06FA0D" box="[436,504,1388,1414]" pageId="3" pageNumber="38">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9014A4FFDA7FAA8FE41FA21" collectingDate="1986-11-18" collectionCode="KPMNH" collectorName="Tanegashima Is. &amp; H. Nagase" country="Japan" location="Ohirono" municipality="Nishinoomote" pageId="3" pageNumber="38" specimenCount="1" stateProvince="Kagoshima">
[
<emphasis id="CDFED440E9014A4FFDBEFAA8FD29FA0D" bold="true" box="[533,727,1388,1414]" pageId="3" pageNumber="38">
<collectorName id="527F6D84E9014A4FFDBEFAA8FD29FA0D" box="[533,727,1388,1414]" pageId="3" pageNumber="38">Tanegashima Is.</collectorName>
</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9014A4FFC8FFAA8FC1AFA0D" box="[804,996,1388,1414]" country="Japan" name="Kagoshima" pageId="3" pageNumber="38">Kagoshima Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9014A4FFC44FAA8FB6FFA0D" box="[1007,1169,1388,1414]" pageId="3" pageNumber="38">Nishinoomote</collectingMunicipality>
,
<location id="FA555E89E9014A4FFB37FAA8FB02FA0D" LSID="urn:lsid:plazi:treatment:7723B944E9014A49FF3CFDB9FACEFEEE:FA555E89E9014A4FFB37FAA8FB02FA0D" box="[1180,1276,1388,1414]" country="Japan" municipality="Nishinoomote" name="Ohirono" pageId="3" pageNumber="38" stateProvince="Kagoshima">Ohirono</location>
,
<date id="8B342E92E9014A4FFAACFAA8FA66FA0D" box="[1287,1432,1388,1414]" pageId="3" pageNumber="38" value="1986-11-18">
<collectingDate id="9B70D77AE9014A4FFAACFAA8FA66FA0D" box="[1287,1432,1388,1414]" pageId="3" pageNumber="38" value="1986-11-18">18. XI. 1986</collectingDate>
</date>
,
<collectorName id="527F6D84E9014A4FFF3CFA54FEF5FA21" box="[151,267,1424,1450]" pageId="3" pageNumber="38">H. Nagase</collectorName>
leg. (
<collectionCode id="999B9097E9014A4FFEE0FA54FE49FA21" box="[331,439,1424,1450]" pageId="3" pageNumber="38">KPMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9014A49FF6CFA70FE81FF5E" lastPageId="5" lastPageNumber="40" pageId="3" pageNumber="38" type="description">
<paragraph id="FF350852E9014A4FFF6CFA70FC71FA79" blockId="3.[151,1437,812,2027]" pageId="3" pageNumber="38">
<emphasis id="CDFED440E9014A4FFF6CFA70FEAFFA45" bold="true" box="[199,337,1460,1486]" pageId="3" pageNumber="38">Description</emphasis>
.
<emphasis id="CDFED440E9014A4FFECAFA70FE46FA45" bold="true" box="[353,440,1460,1486]" pageId="3" pageNumber="38">Female</emphasis>
(n = 29). Body polished and covered with punctures and silvery setae; length 7.0–
<quantity id="3872A5B7E9014A4FFAC6FA70FF38FA79" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="38" unit="mm" value="10.5">10.5 mm</quantity>
. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E9014A4FFF6CFA38FCEFF899" blockId="3.[151,1437,812,2027]" pageId="3" pageNumber="38">
Head 0.6–0.7 × as long as wide. Combined length of clypeus and face 0.95–1.0 × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.65–0.7 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.45–1.5 × as long as OD. OOL 1.15–1.5 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9014A4FFB20F974FB11F941" box="[1163,1263,1712,1738]" captionStart="FIGURE 3" captionStartId="6.[151,250,1534,1558]" captionTargetBox="[151,1436,181,1510]" captionTargetId="figure-19@6.[151,1436,181,1510]" captionTargetPageId="6" captionText="FIGURE 3. Syzeuctus apicifer (Walker, 1874), female from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="3" pageNumber="38">Fig. 3 D</figureCitation>
). Antenna with 42–44 flagellomeres. FL I 4.3–5.0 × as long as maximum depth in lateral view and 1.6–1.75 × as long as FL II. Preapical flagellomere 1.2 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E9014A48FF6CF8D8FCAAFE96" blockId="3.[151,1437,812,2027]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="39" pageId="3" pageNumber="38">
Mesosoma coarsely and densely punctate, 1.4–1.45 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate; speculum with a small smooth area (
<figureCitation id="67B114D7E9014A4FFD31F8A0FD09F8F5" box="[666,759,1892,1918]" captionStart="FIGURE 3" captionStartId="6.[151,250,1534,1558]" captionTargetBox="[151,1436,181,1510]" captionTargetId="figure-19@6.[151,1436,181,1510]" captionTargetPageId="6" captionText="FIGURE 3. Syzeuctus apicifer (Walker, 1874), female from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="3" pageNumber="38">Fig. 3 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E9014A4FFC41F84CFBB9F829" box="[1002,1095,1928,1954]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="3" pageNumber="38">Fig. 1 A</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum absent; sometimes obtusely present medially. Fore wing length
<quantity id="3872A5B7E9014A4FFEF7F814FE0DF861" box="[348,499,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="6.425" metricValueMax="7.25" metricValueMin="5.6" pageId="3" pageNumber="38" unit="mm" value="6.425" valueMax="7.25" valueMin="5.6">5.6–7.25 mm</quantity>
. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole longer than areolet (
<figureCitation id="67B114D7E9064A48FE6CFF53FDDDFF3A" box="[455,547,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1534,1558]" captionTargetBox="[151,1436,181,1510]" captionTargetId="figure-19@6.[151,1436,181,1510]" captionTargetPageId="6" captionText="FIGURE 3. Syzeuctus apicifer (Walker, 1874), female from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="4" pageNumber="39">Fig. 3 B</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.75–6.4 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.0–4.1: 2.0: 1.4: 0.7–0.75: 0.8–0.85. Hind tarsal claws pectinate by short teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9064A48FD43FEC7FCB6FE96" box="[744,840,259,285]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="4" pageNumber="39">Fig. 2 A</figureCitation>
).
</paragraph>
<caption id="ABF558DAE9064A48FF3CFC1CFB9EFBD7" pageId="4" pageNumber="39" startId="4.[151,250,984,1008]" targetBox="[159,1432,313,958]" targetPageId="4" targetType="figure">
<paragraph id="FF350852E9064A48FF3CFC1CFB9EFBD7" blockId="4.[151,1437,984,1116]" pageId="4" pageNumber="39">
<emphasis id="CDFED440E9064A48FF3CFC1CFEECFC7B" bold="true" box="[151,274,984,1008]" pageId="4" pageNumber="39">FIGURE 1.</emphasis>
Epicnemial carina of Japanese species of
<taxonomicName id="388A73D1E9064A48FD1CFC1CFCB5FC64" box="[695,843,984,1007]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="CDFED440E9064A48FD1CFC1CFCE7FC64" box="[695,793,984,1007]" italics="true" pageId="4" pageNumber="39">Syzeuctus</emphasis>
spp.
</taxonomicName>
, females—A:
<taxonomicName id="388A73D1E9064A48FC70FC1CFB1CFC7B" authority="(Walker, 1874)" baseAuthorityName="Walker" baseAuthorityYear="1874" box="[987,1250,984,1008]" class="Insecta" family="Ichneumonidae" genus="Macrus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="apicifer">
<emphasis id="CDFED440E9064A48FC70FC1CFBBDFC7B" box="[987,1091,984,1008]" italics="true" pageId="4" pageNumber="39">S. apicifer</emphasis>
(
<bibRefCitation id="9B1B75A3E9064A48FBE4FC1CFB25FC7B" author="Walker, F." box="[1103,1243,984,1008]" pageId="4" pageNumber="39" pagination="301 - 310" refId="ref23546" refString="Walker, F. (1874) Descriptions of some Japanese Hymenoptera. Cistula Entomologica, 1, 301 - 310." type="journal article" year="1874">Walker, 1874</bibRefCitation>
)
</taxonomicName>
; B:
<taxonomicName id="388A73D1E9064A48FAA2FC1CFA62FC7B" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[1289,1436,984,1008]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="brevicaudus" status="sp. nov.">
<emphasis id="CDFED440E9064A48FAA2FC1CFA62FC7B" box="[1289,1436,984,1008]" italics="true" pageId="4" pageNumber="39">S. brevicaudus</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9064A48FF3CFC38FF16FB9F" bold="true" box="[151,232,1020,1044]" pageId="4" pageNumber="39">
<taxonomicNameLabel id="D6CD693BE9064A48FF3CFC38FF16FB9F" box="[151,232,1020,1044]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); C:
<taxonomicName id="388A73D1E9064A48FE28FC38FD74FB9F" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[387,650,1020,1044]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="CDFED440E9064A48FE28FC38FE06FB9F" box="[387,504,1020,1044]" italics="true" pageId="4" pageNumber="39">S. coreanus</emphasis>
<bibRefCitation id="9B1B75A3E9064A48FE55FC38FD74FB9F" author="Uchida, T." box="[510,650,1020,1044]" pageId="4" pageNumber="39" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
; D:
<taxonomicName id="388A73D1E9064A48FD18FC38FCCCFB9F" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[691,818,1020,1044]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="flavitarsis" status="sp. nov.">
<emphasis id="CDFED440E9064A48FD18FC38FCCCFB9F" box="[691,818,1020,1044]" italics="true" pageId="4" pageNumber="39">S. flavitarsis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9064A48FC93FC38FC77FB9F" bold="true" box="[824,905,1020,1044]" pageId="4" pageNumber="39">
<taxonomicNameLabel id="D6CD693BE9064A48FC93FC38FC77FB9F" box="[824,905,1020,1044]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype); E:
<taxonomicName id="388A73D1E9064A48FB8AFC38FA83FB9F" authority="Kang &amp; Lee, 2020" authorityName="Kang &amp; Lee" authorityYear="2020" box="[1057,1405,1020,1044]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="flavofacialis">
<emphasis id="CDFED440E9064A48FB8AFC38FB49FB9F" box="[1057,1207,1020,1044]" italics="true" pageId="4" pageNumber="39">S. flavofacialis</emphasis>
<bibRefCitation id="9B1B75A3E9064A48FB15FC38FA83FB9F" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[1214,1405,1020,1044]" pageId="4" pageNumber="39" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">Kang &amp; Lee, 2020</bibRefCitation>
</taxonomicName>
; F:
<taxonomicName id="388A73D1E9064A48FF3CFBE4FE75FBB3" authority="Riedel, 2022" authorityName="Riedel" authorityYear="2022" box="[151,395,1056,1080]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="CDFED440E9064A48FF3CFBE4FEFEFBB3" box="[151,256,1056,1080]" italics="true" pageId="4" pageNumber="39">S. laoticus</emphasis>
<bibRefCitation id="9B1B75A3E9064A48FEACFBE4FE75FBB3" author="Riedel, M." box="[263,395,1056,1080]" pageId="4" pageNumber="39" pagination="257 - 273" refId="ref23331" refString="Riedel, M. (2022) Contribution to the knowledge of Oriental Syzeuctus Forster (Hymenoptera, Ichneumonidae, Banchinae), with description of six new species. Linzer biologische Beitrage, 54 (1), 257 - 273." type="journal article" year="2022">Riedel, 2022</bibRefCitation>
</taxonomicName>
; G:
<taxonomicName id="388A73D1E9064A48FE1FFBE4FD40FBB3" authority="Sheng, 2009" authorityName="Sheng" authorityYear="2009" box="[436,702,1056,1080]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="CDFED440E9064A48FE1FFBE4FDC8FBB3" box="[436,566,1056,1080]" italics="true" pageId="4" pageNumber="39">S. maculatus</emphasis>
<bibRefCitation id="9B1B75A3E9064A48FD97FBE4FD40FBB3" author="Sheng, M-L. &amp; Sun, S." box="[572,702,1056,1080]" pageId="4" pageNumber="39" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">Sheng, 2009</bibRefCitation>
</taxonomicName>
; H:
<taxonomicName id="388A73D1E9064A48FD4CFBE4FCBFFBB3" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[743,833,1056,1080]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrus" status="sp. nov.">
<emphasis id="CDFED440E9064A48FD4CFBE4FCBFFBB3" box="[743,833,1056,1080]" italics="true" pageId="4" pageNumber="39">S. nigrus</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9064A48FCECFBE4FC69FBB3" bold="true" box="[839,919,1056,1080]" pageId="4" pageNumber="39">
<taxonomicNameLabel id="D6CD693BE9064A48FCECFBE4FC69FBB3" box="[839,919,1056,1080]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); I:
<taxonomicName id="388A73D1E9064A48FB81FBE4FB46FBB3" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[1066,1208,1056,1080]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="rufiapicalis" status="sp. nov.">
<emphasis id="CDFED440E9064A48FB81FBE4FB46FBB3" box="[1066,1208,1056,1080]" italics="true" pageId="4" pageNumber="39">S. rufiapicalis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9064A48FB15FBE4FAF0FBB3" bold="true" box="[1214,1294,1056,1080]" pageId="4" pageNumber="39">
<taxonomicNameLabel id="D6CD693BE9064A48FB15FBE4FAF0FBB3" box="[1214,1294,1056,1080]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); J:
<taxonomicName id="388A73D1E9064A48FF3CFB80FE5CFBD7" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[151,418,1092,1116]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="sambonis">
<emphasis id="CDFED440E9064A48FF3CFB80FEEFFBD7" box="[151,273,1092,1116]" italics="true" pageId="4" pageNumber="39">S. sambonis</emphasis>
<bibRefCitation id="9B1B75A3E9064A48FEBCFB80FE5CFBD7" author="Uchida, T." box="[279,418,1092,1116]" pageId="4" pageNumber="39" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
; K:
<taxonomicName id="388A73D1E9064A48FE67FB80FD14FBD7" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[460,746,1092,1116]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E9064A48FE67FB80FDA9FBD7" box="[460,599,1092,1116]" italics="true" pageId="4" pageNumber="39">S. takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E9064A48FDF5FB80FD14FBD7" author="Uchida, T." box="[606,746,1092,1116]" pageId="4" pageNumber="39" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
; L:
<taxonomicName id="388A73D1E9064A48FCBBFB80FC6CFBD7" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[784,914,1092,1116]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9064A48FCBBFB80FC6CFBD7" box="[784,914,1092,1116]" italics="true" pageId="4" pageNumber="39">S. yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9064A48FC33FB80FC17FBD7" bold="true" box="[920,1001,1092,1116]" pageId="4" pageNumber="39">
<taxonomicNameLabel id="D6CD693BE9064A48FC33FB80FC17FBD7" box="[920,1001,1092,1116]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype).
</paragraph>
</caption>
<paragraph id="FF350852E9064A48FF6CFB4FFCEEFA9B" blockId="4.[151,1437,1162,2017]" pageId="4" pageNumber="39">Metasoma. T I 2.0–2.35 × as long as maximum width, finely punctate except for smooth areas on median longitudinal area, area along posterior margin and lateral sides. T II 1.2–1.4 × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 2.5–2.8 × as long as hind tibia, ca. 1.2 × as long as fore wing, and shorter than body length.</paragraph>
<paragraph id="FF350852E9064A48FF6CFADEFE1AF8DA" blockId="4.[151,1437,1162,2017]" pageId="4" pageNumber="39">
Colouration (
<figureCitation id="67B114D7E9064A48FEF7FADEFE1EFABF" box="[348,480,1306,1332]" captionStart="FIGURE 3" captionStartId="6.[151,250,1534,1558]" captionTargetBox="[151,1436,181,1510]" captionTargetId="figure-19@6.[151,1436,181,1510]" captionTargetPageId="6" captionText="FIGURE 3. Syzeuctus apicifer (Walker, 1874), female from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="4" pageNumber="39">Figs 3 A–E</figureCitation>
). Body (excluding wings and legs) black. Clypeus yellow, usually with black area(s). Mandible except for teeth and base, malar space, inner orbit on frons and vertex, outer orbit except for dorsal part yellow. Scape partly narrowly tinged with yellow. Flagellum and palpi largely tinged with reddish-brown. Postero-dorsal corner of pronotum tinged with yellow. Subtegular ridge, tegula, scutellum except for baso-median black area, postscutellum, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of propodeum yellow. Lower part of pronotum and antero-lateral area of mesoscutum usually with a pair of small yellow spots. Metasomal tergites red to reddish-brown except for posterior segments tinged with black; anterior and posterior parts of T I yellow; posterior part of T II with yellow; posterior margin of T III narrowly tinged with yellow. Red area of metasomal tergites sometimes partly darkened. Membranous parts of metasomal sternites largely yellow. Subgenital plate blackish-brown except of reddish-brown apex. Ovipositor reddish-brown. Wings hyaline except for apical clouded area of fore wing. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow; coxae with small black spots ventrally; apex of middle tarsus tinged with yellowish-brown. Hind coxa and trochanter black, each with yellow apical margin. Hind trochantellus yellow. Hind femur reddish-brown. Hind tibia yellowish-brown to reddish-brown, usually more or less darkened apically. Hind tibial spurs yellowish-brown. Hind tarsus brown to blackish-brown, base of first tarsomere and apices of each tarsomere usually paler than other parts.
</paragraph>
<paragraph id="FF350852E9064A49FF6CF89EFE81FF5E" blockId="4.[151,1437,1162,2017]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="40" pageId="4" pageNumber="39">
<emphasis id="CDFED440E9064A48FF6CF89EFEFAF8FF" bold="true" box="[199,260,1882,1908]" pageId="4" pageNumber="39">Male</emphasis>
(n = 13). Similar to female (
<figureCitation id="67B114D7E9064A48FDF1F89EFD19F8FF" box="[602,743,1882,1908]" captionStart="FIGURE 4" captionStartId="7.[151,250,1621,1645]" captionTargetBox="[151,1436,345,1597]" captionTargetId="figure-103@7.[151,1436,345,1597]" captionTargetPageId="7" captionText="FIGURE 4. Syzeuctus apicifer (Walker, 1874), male from Japan—A: lateral habitus; B: dorsal habitus; C, D: head, frontal view." pageId="4" pageNumber="39">Figs 4 A–D</figureCitation>
). Malar space 0.9–1.0 × as long as basal mandibular width. Antenna with 43–45 flagellomeres. FL I 5.3–5.7 × as long as maximum depth in lateral view and 1.85 × as long as FL II. Tergal spiracles not enlarged. T I 2.4–2.65 × as long as maximum width. T II 1.4–1.55 × as long as maximum width. Yellow area of body more extended than in female. Clypeus, malar space, inner orbit on face, palpi, and postero-ventral spot of mesopleuron yellow. Face usually with yellow longitudinal markings; size of black area varied (
<figureCitation id="67B114D7E9074A49FF47FF7FFE8CFF5E" box="[236,370,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1621,1645]" captionTargetBox="[151,1436,345,1597]" captionTargetId="figure-103@7.[151,1436,345,1597]" captionTargetPageId="7" captionText="FIGURE 4. Syzeuctus apicifer (Walker, 1874), male from Japan—A: lateral habitus; B: dorsal habitus; C, D: head, frontal view." pageId="5" pageNumber="40">Figs 4 C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9074A49FF6CFF1AFACEFEEE" pageId="5" pageNumber="40" type="distribution">
<paragraph id="FF350852E9074A49FF6CFF1AFE52FE96" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="40">
<emphasis id="CDFED440E9074A49FF6CFF1AFEA7FF73" bold="true" box="[199,345,222,248]" pageId="5" pageNumber="40">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9074A49FEC3FF1BFE56FF72" box="[360,424,223,249]" name="Japan" pageId="5" pageNumber="40">Japan</collectingCountry>
(
<collectingRegion id="3D4EC6B0E9074A49FE1CFF1BFDD4FF72" box="[439,554,223,249]" country="Japan" name="Hokkaido" pageId="5" pageNumber="40">Hokkaido</collectingRegion>
, Honshu, Shikoku, and Tanegashima Is.),
<collectingCountry id="879D48C2E9074A49FBADFF1BFBB7FF72" box="[1030,1097,223,249]" name="China" pageId="5" pageNumber="40">China</collectingCountry>
and
<collectingCountry id="879D48C2E9074A49FB2AFF1BFB37FF72" box="[1153,1225,223,249]" name="South Korea" pageId="5" pageNumber="40">Korea</collectingCountry>
. New to Hokkaido and Tanegashima Island.
</paragraph>
<paragraph id="FF350852E9074A49FF6CFEE2FACEFEEE" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="40">
<emphasis id="CDFED440E9074A49FF6CFEE2FEBDFECB" bold="true" box="[199,323,294,320]" pageId="5" pageNumber="40">Bionomics</emphasis>
. Host is unknown. Adults usually collected in grassland. In Sugadaira,
<collectingRegion id="3D4EC6B0E9074A49FBD1FEE3FAAFFECA" box="[1146,1361,295,321]" country="Japan" name="Nagano" pageId="5" pageNumber="40">Nagano Prefecture</collectingRegion>
, many females (including above
<specimenCount id="E98CC3DBE9074A49FE12FE8FFDAFFEEE" box="[441,593,331,357]" count="8" pageId="5" pageNumber="40" type="female">eight females</specimenCount>
) came to the flower of
<taxonomicName id="388A73D1E9074A49FCFDFE8FFB59FEEE" authority="Maxim." authorityName="Maxim." box="[854,1191,331,357]" class="Insecta" family="Braconidae" genus="Angelica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="polymorpha">
<emphasis id="CDFED440E9074A49FCFDFE8FFBB6FEEE" box="[854,1096,331,357]" italics="true" pageId="5" pageNumber="40">Angelica polymorpha</emphasis>
Maxim.
</taxonomicName>
(
<taxonomicName id="388A73D1E9074A49FB1EFE8FFADAFEEE" box="[1205,1316,331,357]" class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" order="Apiales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="family">Apiaceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

199
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@ -0,0 +1,199 @@
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<mods:title id="B940E0D5755D87C8C2D254F3113AC254">Revision of the genus Syzeuctus Förster, 1869 (Hymenoptera, Ichneumonidae, Banchinae) from Japan</mods:title>
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<mods:date id="4795C338A7D9CD680A3E8BB323BF7723">2024</mods:date>
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<subSubSection id="B7905BD9E9044A4AFF3CF949FDF3F92C" box="[151,525,1676,1703]" pageId="6" pageNumber="41" type="nomenclature">
<paragraph id="FF350852E9044A4AFF3CF949FDF3F92C" blockId="6.[151,525,1676,1739]" box="[151,525,1676,1703]" pageId="6" pageNumber="41">
<heading id="A47DBF3EE9044A4AFF3CF949FDF3F92C" bold="true" box="[151,525,1676,1703]" fontSize="11" level="1" pageId="6" pageNumber="41" reason="1">
<emphasis id="CDFED440E9044A4AFF3CF949FDF3F92C" bold="true" box="[151,525,1676,1703]" pageId="6" pageNumber="41">
<taxonomicName id="388A73D1E9044A4AFF3CF949FE56F92C" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[151,424,1676,1703]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="brevicaudus" status="sp. nov.">
<emphasis id="CDFED440E9044A4AFF3CF949FE56F92C" bold="true" box="[151,424,1676,1703]" italics="true" pageId="6" pageNumber="41">Syzeuctus brevicaudus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6CD693BE9044A4AFE04F949FDF3F92C" box="[431,525,1677,1703]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9044A4AFF3CF975FE6CF940" box="[151,402,1712,1739]" pageId="6" pageNumber="41" type="description">
<paragraph id="FF350852E9044A4AFF3CF975FE6CF940" blockId="6.[151,525,1676,1739]" box="[151,402,1712,1739]" pageId="6" pageNumber="41">
(
<figureCitation id="67B114D7E9044A4AFF34F975FEFCF940" box="[159,258,1713,1739]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="6" pageNumber="41">Figs 1 B</figureCitation>
,
<figureCitation id="67B114D7E9044A4AFEA6F975FEC9F940" box="[269,311,1713,1739]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="6" pageNumber="41">2 B</figureCitation>
,
<figureCitation id="67B114D7E9044A4AFEE8F974FE74F940" box="[323,394,1712,1739]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[151,1436,273,1843]" captionTargetId="figure-76@8.[151,1436,273,1843]" captionTargetPageId="8" captionText="FIGURE 5. Syzeuctus brevicaudus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="6" pageNumber="41">5 A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9044A4AFF3CF93DFC3DF8BC" pageId="6" pageNumber="41" type="materials_examined">
<paragraph id="FF350852E9044A4AFF3CF93DFC3DF8BC" blockId="6.[151,1437,1784,2027]" pageId="6" pageNumber="41">
<emphasis id="CDFED440E9044A4AFF3CF93DFEE6F899" bold="true" box="[151,280,1784,1811]" pageId="6" pageNumber="41">Type series</emphasis>
.
<materialsCitation id="4FE2020FE9044A4AFE88F93DFC3DF8BC" collectingDate="2020-08-13" collectingDateMax="2020-09-13" collectingDateMin="2020-08-13" collectionCode="EUM" collectorName="Y. Okatsu &amp; Y. Nakano" country="Japan" county="Honshu" location="Mt. Asakusa-dake" municipality="Tadami" pageId="6" pageNumber="41" specimenCount="1" stateProvince="Fukushima" typeStatus="holotype">
<emphasis id="CDFED440E9044A4AFE88F93DFE6EF898" bold="true" box="[291,400,1785,1811]" pageId="6" pageNumber="41">
<typeStatus id="2031B6F0E9044A4AFE88F93DFE6EF898" box="[291,400,1785,1811]" pageId="6" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
:F,
<emphasis id="CDFED440E9044A4AFE18F93CFDF3F899" bold="true" box="[435,525,1784,1810]" pageId="6" pageNumber="41">
<collectingCountry id="879D48C2E9044A4AFE18F93CFDF3F899" box="[435,525,1784,1810]" name="Japan" pageId="6" pageNumber="41">JAPAN</collectingCountry>
</emphasis>
,
<emphasis id="CDFED440E9044A4AFDBCF93DFD8AF898" bold="true" box="[535,628,1785,1811]" pageId="6" pageNumber="41">
<collectingCounty id="165470DEE9044A4AFDBCF93DFD8AF898" box="[535,628,1785,1811]" pageId="6" pageNumber="41">Honshu</collectingCounty>
</emphasis>
,
<collectingRegion id="3D4EC6B0E9044A4AFDD4F93DFCC6F898" box="[639,824,1785,1811]" country="Japan" name="Fukushima" pageId="6" pageNumber="41">Fukushima Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9044A4AFC94F93DFC68F898" box="[831,918,1785,1811]" pageId="6" pageNumber="41">Tadami</collectingMunicipality>
,
<location id="FA555E89E9044A4AFC36F93DFB99F898" LSID="urn:lsid:plazi:treatment:7723B944E9044A45FF3CF949FE6BFDB6:FA555E89E9044A4AFC36F93DFB99F898" box="[925,1127,1785,1811]" country="Japan" county="Honshu" municipality="Tadami" name="Mt. Asakusa-dake" pageId="6" pageNumber="41" stateProvince="Fukushima">Mt.Asakusa-dake</location>
,
<location id="FA555E89E9044A4AFBC4F93DFAFAF898" LSID="urn:lsid:plazi:treatment:7723B944E9044A45FF3CF949FE6BFDB6:FA555E89E9044A4AFBC4F93DFAFAF898" box="[1135,1284,1785,1811]" country="Japan" county="Honshu" municipality="Tadami" name="Numanotaira" pageId="6" pageNumber="41" stateProvince="Fukushima">Numanotaira</location>
,
<location id="FA555E89E9044A4AFAA7F93DFF28F8BC" LSID="urn:lsid:plazi:treatment:7723B944E9044A45FF3CF949FE6BFDB6:FA555E89E9044A4AFAA7F93DFF28F8BC" country="Japan" county="Honshu" municipality="Tadami" name="Gonzaemonyachi" pageId="6" pageNumber="41" stateProvince="Fukushima">Gonzaemonyachi</location>
,
<date id="8B342E92E9044A4AFF49F8D9FE1FF8BC" box="[226,481,1821,1847]" pageId="6" pageNumber="41" value="2020-08-13" valueMax="2020-09-13" valueMin="2020-08-13">
<collectingDate id="9B70D77AE9044A4AFF49F8D9FE1FF8BC" box="[226,481,1821,1847]" pageId="6" pageNumber="41" value="2020-08-13" valueMax="2020-09-13" valueMin="2020-08-13">13. VIII.–13. IX. 2020</collectingDate>
</date>
,
<collectorName id="527F6D84E9044A4AFE40F8D9FDA7F8BC" box="[491,601,1821,1847]" pageId="6" pageNumber="41">Y. Okatsu</collectorName>
&amp;
<collectorName id="527F6D84E9044A4AFDD7F8D9FD0FF8BC" box="[636,753,1821,1847]" pageId="6" pageNumber="41">Y. Nakano</collectorName>
leg., MsT (
<collectionCode id="999B9097E9044A4AFCD8F8D9FC49F8BC" box="[883,951,1821,1847]" pageId="6" pageNumber="41">EUM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9044A45FF6CF884FE76FE02" lastPageId="9" lastPageNumber="44" pageId="6" pageNumber="41" type="description">
<paragraph id="FF350852E9044A4AFF6CF884FCABF8F4" blockId="6.[151,1437,1784,2027]" pageId="6" pageNumber="41">
<emphasis id="CDFED440E9044A4AFF6CF884FEAFF8D1" bold="true" box="[199,337,1856,1882]" pageId="6" pageNumber="41">Description</emphasis>
.
<emphasis id="CDFED440E9044A4AFECAF885FE46F8D0" bold="true" box="[353,440,1857,1883]" pageId="6" pageNumber="41">Female</emphasis>
(n = 1). Body polished and covered with punctures and silvery setae; length
<quantity id="3872A5B7E9044A4AFA9BF884FA67F8D0" box="[1328,1433,1856,1883]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" pageId="6" pageNumber="41" unit="mm" value="9.75">9.75 mm</quantity>
. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E9044A4BFF6CF84DFF21FECA" blockId="6.[151,1437,1784,2027]" lastBlockId="7.[151,1437,150,321]" lastPageId="7" lastPageNumber="42" pageId="6" pageNumber="41">
Head 0.65 × as long as wide. Combined length of clypeus and face 0.8 × facial width. Clypeus smooth ventrally, punctate dorsally, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, with a pair of slight convexities just above of each anterior tentorial pit. Frons concave medially, with a pair of distinct convexities. Malar space 0.85 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.1 × as long as OD. OOL 1.6 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9054A4BFD59FF1BFCAAFF72" box="[754,852,222,249]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[151,1436,273,1843]" captionTargetId="figure-76@8.[151,1436,273,1843]" captionTargetPageId="8" captionText="FIGURE 5. Syzeuctus brevicaudus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="7" pageNumber="42">Fig. 5 D</figureCitation>
). Antenna with 48 flagellomeres. FL I 3.3 × as long as maximum depth in lateral view and 1.5 × as long as FL II. Preapical flagellomere 1.2 × as long as maximum width.
</paragraph>
<caption id="ABF558DAE9054A4BFF3CF991FF30F91A" pageId="7" pageNumber="42" startId="7.[151,250,1621,1645]" targetBox="[151,1436,345,1597]" targetPageId="7" targetType="figure">
<paragraph id="FF350852E9054A4BFF3CF991FF30F91A" blockId="7.[151,1437,1621,1681]" pageId="7" pageNumber="42">
<emphasis id="CDFED440E9054A4BFF3CF991FEE8F9E6" bold="true" box="[151,278,1621,1645]" pageId="7" pageNumber="42">FIGURE 4.</emphasis>
<taxonomicName id="388A73D1E9054A4BFEB4F991FD80F9E6" authority="(Walker, 1874)" baseAuthorityName="Walker" baseAuthorityYear="1874" box="[287,638,1621,1645]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="apicifer">
<emphasis id="CDFED440E9054A4BFEB4F991FE27F9E6" box="[287,473,1621,1645]" italics="true" pageId="7" pageNumber="42">Syzeuctus apicifer</emphasis>
(
<bibRefCitation id="9B1B75A3E9054A4BFE43F991FD89F9E6" author="Walker, F." box="[488,631,1621,1645]" pageId="7" pageNumber="42" pagination="301 - 310" refId="ref23546" refString="Walker, F. (1874) Descriptions of some Japanese Hymenoptera. Cistula Entomologica, 1, 301 - 310." type="journal article" year="1874">Walker, 1874</bibRefCitation>
)
</taxonomicName>
, male from Japan—A: lateral habitus; B: dorsal habitus; C, D: head, frontal view.
</paragraph>
</caption>
<paragraph id="FF350852E9054A4BFF6CF904FB53F872" blockId="7.[151,1437,1727,2041]" pageId="7" pageNumber="42">
Mesosoma coarsely and densely punctate, 1.6 × as long as maximum height. Pronotum with small smooth area on ventral part. Epomia present, its dorsal end not reaching dorsal margin (
<figureCitation id="67B114D7E9054A4BFB9BF927FB6DF976" box="[1072,1171,1763,1789]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="7" pageNumber="42">Fig. 1 B</figureCitation>
). Mesopleuron densely punctate; speculum with a small smooth area (
<figureCitation id="67B114D7E9054A4BFD31F8C3FD09F8AA" box="[666,759,1799,1826]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[151,1436,273,1843]" captionTargetId="figure-76@8.[151,1436,273,1843]" captionTargetPageId="8" captionText="FIGURE 5. Syzeuctus brevicaudus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="7" pageNumber="42">Fig. 5 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation. Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum absent. Fore wing length 7.0 mm. Areolet receiving vein 2m-cu near the middle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E9054A4BFC1EF8B7FBECF806" box="[949,1042,1907,1934]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[151,1436,273,1843]" captionTargetId="figure-76@8.[151,1436,273,1843]" captionTargetPageId="8" captionText="FIGURE 5. Syzeuctus brevicaudus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="7" pageNumber="42">Fig. 5 F</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted slightly posterior to the middle. Hind femur 5.3 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.2: 2.0: 1.5: 0.75: 0.95. Hind tarsal claws pectinate by short teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9054A4BFBEBF81BFB61F872" box="[1088,1183,2015,2041]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="7" pageNumber="42">Fig. 2 B</figureCitation>
).
</paragraph>
<paragraph id="FF350852E90A4A44FF6CFF53FCEEFF72" blockId="8.[151,1437,151,249]" pageId="8" pageNumber="43">Metasoma. T I 2.3 × as long as maximum width, smooth except for subapical part finely and sparsely punctate. T II 1.3 × as long as maximum width. T II to T V finely punctate. Ovipositor sheath 0.85 × as long as hind tibia, ca. 0.3 × as long as fore wing, and shorter than body length.</paragraph>
<caption id="ABF558DAE90A4A44FF3CF88FFB1FF80C" pageId="8" pageNumber="43" startId="8.[151,250,1867,1891]" targetBox="[151,1436,273,1843]" targetPageId="8" targetType="figure">
<paragraph id="FF350852E90A4A44FF3CF88FFB1FF80C" blockId="8.[151,1436,1867,1927]" pageId="8" pageNumber="43">
<emphasis id="CDFED440E90A4A44FF3CF88FFEEAF8E8" bold="true" box="[151,276,1867,1891]" pageId="8" pageNumber="43">FIGURE 5.</emphasis>
<taxonomicName id="388A73D1E90A4A44FEB0F888FDFEF8E8" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[283,512,1867,1891]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="brevicaudus" status="sp. nov.">
<emphasis id="CDFED440E90A4A44FEB0F888FDFEF8E8" box="[283,512,1867,1891]" italics="true" pageId="8" pageNumber="43">Syzeuctus brevicaudus</emphasis>
</taxonomicName>
<emphasis id="CDFED440E90A4A44FDACF88FFDA6F8E8" bold="true" box="[519,600,1867,1891]" pageId="8" pageNumber="43">
<taxonomicNameLabel id="D6CD693BE90A4A44FDACF88FFDA6F8E8" box="[519,600,1867,1891]" pageId="8" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing.
</paragraph>
</caption>
<paragraph id="FF350852E90A4A45FF6CF871FBEAFEEE" blockId="8.[151,1437,1973,2036]" lastBlockId="9.[151,1436,151,573]" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
Colouration (
<figureCitation id="67B114D7E90A4A44FEF6F871FE1FF85B" box="[349,481,1973,2000]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[151,1436,273,1843]" captionTargetId="figure-76@8.[151,1436,273,1843]" captionTargetPageId="8" captionText="FIGURE 5. Syzeuctus brevicaudus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="8" pageNumber="43">Figs 5 A–F</figureCitation>
). Body (excluding wings and legs) reddish-yellow. Clypeus, face, malar space, inner and outer orbits, mandible except for teeth, palpi, propleuron, fore and mid legs, membranous part of metasomal sternites, and subgenital plate yellow. Ventral surface of scape, postero-dorsal corner and ventral part of pronotum, tegula, subtegular ridge, median longitudinal area of mesopleurou, apices of scutellum and postscutellum, postero-ventral part of metapleuron, and posterior part of propodeum tinged with yellow. Antenna largely brown. An anterior median longitudinal stripe of mesoscutum, an anterior longitudinal stripe of mesopleuron, lateral sides of laterotergites of T I to T III blackish-brown to black. Wings hyaline except for weakly clouded area of apical part of fore wing. Veins and pterostigma brown except for yellowish-brown wing base.
</paragraph>
<paragraph id="FF350852E90B4A45FF6CFEABFE76FE02" blockId="9.[151,1436,151,573]" box="[199,392,367,393]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF6CFEABFEFAFE02" bold="true" box="[199,260,367,393]" pageId="9" pageNumber="44">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90B4A45FF6CFE56FDDEFE26" box="[199,544,402,429]" pageId="9" pageNumber="44" type="distribution">
<paragraph id="FF350852E90B4A45FF6CFE56FDDEFE26" blockId="9.[151,1436,151,573]" box="[199,544,402,429]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF6CFE56FEA7FE27" bold="true" box="[199,345,402,428]" pageId="9" pageNumber="44">Distribution</emphasis>
.
<collectingCountry id="879D48C2E90B4A45FECCFE57FE59FE26" box="[359,423,403,429]" name="Japan" pageId="9" pageNumber="44">Japan</collectingCountry>
(Honshu).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90B4A45FF6CFE73FE36FE7E" pageId="9" pageNumber="44" type="etymology">
<paragraph id="FF350852E90B4A45FF6CFE73FC93FE5A" blockId="9.[151,1436,151,573]" box="[199,877,439,465]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF6CFE73FEB6FE5A" bold="true" box="[199,328,439,465]" pageId="9" pageNumber="44">Etymology</emphasis>
. The specific name refers to the short ovipositor.
</paragraph>
<paragraph id="FF350852E90B4A45FF6CFE1EFE36FE7E" blockId="9.[151,1436,151,573]" box="[199,456,474,501]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF6CFE1EFEBDFE7F" bold="true" box="[199,323,474,500]" pageId="9" pageNumber="44">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90B4A45FF6CFE3BFE6BFDB6" pageId="9" pageNumber="44" type="discussion">
<paragraph id="FF350852E90B4A45FF6CFE3BFE6BFDB6" blockId="9.[151,1436,151,573]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF6CFE3BFECAFD92" bold="true" box="[199,308,511,537]" pageId="9" pageNumber="44">Remarks</emphasis>
. This species can be easily distinguished from all other Japanese species by the ovipositor distinctly shorter than hind tibia.
</paragraph>
</subSubSection>
</treatment>
</document>

339
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<paragraph id="FF350852E90B4A45FF3CFD4BFDD1FD23" blockId="9.[151,559,654,717]" box="[151,559,654,681]" pageId="9" pageNumber="44">
<heading id="A47DBF3EE90B4A45FF3CFD4BFDD1FD23" bold="true" box="[151,559,654,681]" fontSize="11" level="1" pageId="9" pageNumber="44" reason="1">
<taxonomicName id="388A73D1E90B4A45FF3CFD4BFDD1FD23" ID-CoL="545FD" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[151,559,654,681]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="coreanus">
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<emphasis id="CDFED440E90B4A45FF3CFD4BFE7AFD22" bold="true" box="[151,388,655,681]" italics="true" pageId="9" pageNumber="44">Syzeuctus coreanus</emphasis>
<bibRefCitation id="9B1B75A3E90B4A45FE27FD4AFDD1FD23" author="Uchida, T." box="[396,559,654,680]" pageId="9" pageNumber="44" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</emphasis>
</taxonomicName>
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<subSubSection id="B7905BD9E90B4A45FF3CFD77FE11FD46" box="[151,495,691,717]" pageId="9" pageNumber="44" type="description">
<paragraph id="FF350852E90B4A45FF3CFD77FE11FD46" blockId="9.[151,559,654,717]" box="[151,495,691,717]" pageId="9" pageNumber="44">
(
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,
<figureCitation id="67B114D7E90B4A45FEA6FD77FEC9FD46" box="[269,311,691,717]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="9" pageNumber="44">2 C</figureCitation>
,
<figureCitation id="67B114D7E90B4A45FEE8FD77FE6EFD46" box="[323,400,691,717]" captionStart="FIGURE 6" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-19@10.[151,1436,181,1723]" captionTargetPageId="10" captionText="FIGURE 6. Syzeuctus coreanus Uchida, 1928, female—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view." pageId="9" pageNumber="44">6 A–D</figureCitation>
,
<figureCitation id="67B114D7E90B4A45FE30FD77FE18FD46" box="[411,486,691,717]" captionStart="FIGURE 7" captionStartId="11.[151,250,1617,1641]" captionTargetBox="[151,1436,453,1593]" captionTargetId="figure-157@11.[151,1436,453,1593]" captionTargetPageId="11" captionText="FIGURE 7. Syzeuctus coreanus Uchida, 1928, male—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: propodeum, dorsal view." pageId="9" pageNumber="44">7 A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90B4A45FF3CFD33FDE6FC84" box="[151,536,759,783]" pageId="9" pageNumber="44" type="reference_group">
<paragraph id="FF350852E90B4A45FF3CFD33FDE6FC84" blockId="9.[151,536,759,783]" box="[151,536,759,783]" pageId="9" pageNumber="44">
<treatmentCitationGroup id="DF9A2F7CE90B4A45FF3CFD33FDE6FC84" box="[151,536,759,783]" pageId="9" pageNumber="44">
<treatmentCitation id="7E2B2E43E90B4A45FF3CFD33FDEAFC84" author="Uchida, T." box="[151,532,759,783]" page="93" pageId="9" pageNumber="44" year="1928">
<taxonomicName id="388A73D1E90B4A45FF3CFD33FDEAFC84" ID-CoL="545FD" authority="Uchida, 1928: 93" authorityName="Uchida" authorityPageNumber="93" authorityYear="1928" box="[151,532,759,783]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="CDFED440E90B4A45FF3CFD33FEA2FC84" box="[151,348,759,783]" italics="true" pageId="9" pageNumber="44">Syzeuctus coreanus</emphasis>
<bibRefCitation id="9B1B75A3E90B4A45FEC9FD33FDEAFC84" author="Uchida, T." box="[354,532,759,783]" pageId="9" pageNumber="44" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928: 93</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90B4A45FF3CFCF9FED5FBA4" pageId="9" pageNumber="44" type="materials_examined">
<paragraph id="FF350852E90B4A45FF3CFCF9FED5FBA4" blockId="9.[151,1437,829,1936]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF3CFCF9FE7AFCDC" bold="true" box="[151,388,829,855]" pageId="9" pageNumber="44">Materials examined</emphasis>
.
<materialsCitation id="4FE2020FE90B4A45FE39FCF9FA66FCDC" box="[402,1432,829,856]" collectingDate="2000-08-25" collectionCode="EUM" collectorName="H. Makihara" country="Japan" location="Nanakai Vil." pageId="9" pageNumber="44" specimenCount="1" stateProvince="Ibaraki">
<emphasis id="CDFED440E90B4A45FE39FCF9FE12FCDC" bold="true" box="[402,492,829,855]" pageId="9" pageNumber="44">
<collectingCountry id="879D48C2E90B4A45FE39FCF9FE12FCDC" box="[402,492,829,855]" name="Japan" pageId="9" pageNumber="44">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E90B4A45FDA8FCF9FD9EFCDC" bold="true" box="[515,608,829,855]" pageId="9" pageNumber="44">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E90B4A45FD0BFCF9FCD1FCDC" box="[672,815,829,855]" country="Japan" name="Ibaraki" pageId="9" pageNumber="44">Ibaraki Pref.</collectingRegion>
,
<location id="FA555E89E90B4A45FC92FCF9FC37FCDC" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FC92FCF9FC37FCDC" box="[825,969,829,855]" country="Japan" name="Nanakai Vil." pageId="9" pageNumber="44" stateProvince="Ibaraki">Nanakai Vil.</location>
,
<date id="8B342E92E90B4A45FC78FCF9FB8BFCDC" box="[979,1141,829,856]" pageId="9" pageNumber="44" value="2000-08-25">
<collectingDate id="9B70D77AE90B4A45FC78FCF9FB8BFCDC" box="[979,1141,829,856]" pageId="9" pageNumber="44" value="2000-08-25">25. VIII. 2000</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FB2BFCFAFAF3FCDC" box="[1152,1293,829,856]" pageId="9" pageNumber="44">H. Makihara</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FAE7FCF9FA6EFCDC" box="[1356,1424,829,855]" pageId="9" pageNumber="44">EUM</collectionCode>
)
</materialsCitation>
; 1 M,
<materialsCitation id="4FE2020FE90B4A45FF70FCA5FBE3FCF0" box="[219,1053,865,892]" collectingDate="1966-08-07" collectionCode="MNHAH" collectorName="T. Nambu" country="Japan" location="Urayamaguchi" pageId="9" pageNumber="44" specimenCount="1" stateProvince="Saitama">
<collectingRegion id="3D4EC6B0E90B4A45FF70FCA5FE87FCF0" box="[219,377,865,891]" country="Japan" name="Saitama" pageId="9" pageNumber="44">Saitama Pref.</collectingRegion>
,
<location id="FA555E89E90B4A45FE23FCA5FDCEFCF0" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FE23FCA5FDCEFCF0" box="[392,560,865,891]" country="Japan" name="Urayamaguchi" pageId="9" pageNumber="44" stateProvince="Saitama">Urayamaguchi</location>
,
<date id="8B342E92E90B4A45FD94FCA6FD25FCF0" box="[575,731,865,892]" pageId="9" pageNumber="44" value="1966-08-07">
<collectingDate id="9B70D77AE90B4A45FD94FCA6FD25FCF0" box="[575,731,865,892]" pageId="9" pageNumber="44" value="1966-08-07">7. VIII. 1966</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FD41FCA6FCA3FCF0" box="[746,861,865,892]" pageId="9" pageNumber="44">T. Nambu</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FC0FFCA5FBEBFCF0" box="[932,1045,865,891]" pageId="9" pageNumber="44">MNHAH</collectionCode>
)
</materialsCitation>
; 1 F,
<materialsCitation id="4FE2020FE90B4A45FBCFFCA5FD20FC14" collectingDate="2010-08-28" collectionCode="KPMNH" collectorName="Y. Saito" country="Japan" location="Nishiagano" municipality="Hanno City" pageId="9" pageNumber="44" specimenCount="1" stateProvince="Saitama">
<collectingRegion id="3D4EC6B0E90B4A45FBCFFCA5FAFCFCF0" box="[1124,1282,865,891]" country="Japan" name="Saitama" pageId="9" pageNumber="44">Saitama Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90B4A45FABAFCA6FA66FCF0" box="[1297,1432,865,892]" pageId="9" pageNumber="44">Hanno City</collectingMunicipality>
,
<location id="FA555E89E90B4A45FF3CFC41FEE7FC14" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FF3CFC41FEE7FC14" box="[151,281,901,927]" country="Japan" municipality="Hanno City" name="Nishiagano" pageId="9" pageNumber="44" stateProvince="Saitama">Nishiagano</location>
,
<date id="8B342E92E90B4A45FE8EFC41FE36FC14" box="[293,456,901,928]" pageId="9" pageNumber="44" value="2010-08-28">
<collectingDate id="9B70D77AE90B4A45FE8EFC41FE36FC14" box="[293,456,901,928]" pageId="9" pageNumber="44" value="2010-08-28">28. VIII. 2010</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FE78FC42FDD5FC14" box="[467,555,901,928]" pageId="9" pageNumber="44">Y. Saito</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FDC0FC41FD29FC14" box="[619,727,901,927]" pageId="9" pageNumber="44">KPMNH</collectionCode>
)
</materialsCitation>
; 1 F,
<materialsCitation id="4FE2020FE90B4A45FCB0FC42FE78FC48" collectingDate="2002-08-01" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" location="Mine" municipality="Yokohama City" pageId="9" pageNumber="44" specimenCount="1" stateProvince="Kanagawa">
<collectingRegion id="3D4EC6B0E90B4A45FCB0FC42FC2FFC14" box="[795,977,901,928]" country="Japan" name="Kanagawa" pageId="9" pageNumber="44">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90B4A45FC70FC41FB75FC14" box="[987,1163,901,927]" pageId="9" pageNumber="44">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E90B4A45FB3CFC41FB2BFC14" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FB3CFC41FB2BFC14" box="[1175,1237,901,927]" country="Japan" municipality="Yokohama City" name="Mine" pageId="9" pageNumber="44" stateProvince="Kanagawa">Mine</location>
,
<date id="8B342E92E90B4A45FB4BFC41FA88FC14" box="[1248,1398,901,928]" pageId="9" pageNumber="44" value="2002-08-01">
<collectingDate id="9B70D77AE90B4A45FB4BFC41FA88FC14" box="[1248,1398,901,928]" pageId="9" pageNumber="44" value="2002-08-01">1. VIII. 2002</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FA2AFC42FF2BFC48" pageId="9" pageNumber="44">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FEB9FC6DFE80FC48" box="[274,382,937,963]" pageId="9" pageNumber="44">KPMNH</collectionCode>
)
</materialsCitation>
; 1 F,
<materialsCitation id="4FE2020FE90B4A45FE15FC6EFACEFC48" box="[446,1328,937,964]" collectingDate="2008-08-03" collectionCode="EUM" collectorName="K. Kubo" country="Japan" location="Enkaizan" municipality="Yokohama City" pageId="9" pageNumber="44" specimenCount="1" stateProvince="Kanagawa">
<collectingRegion id="3D4EC6B0E90B4A45FE15FC6EFD8AFC48" box="[446,628,937,964]" country="Japan" name="Kanagawa" pageId="9" pageNumber="44">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90B4A45FDD7FC6DFCD5FC48" box="[636,811,937,963]" pageId="9" pageNumber="44">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E90B4A45FC9DFC6DFC5FFC48" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FC9DFC6DFC5FFC48" box="[822,929,937,963]" country="Japan" municipality="Yokohama City" name="Enkaizan" pageId="9" pageNumber="44" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E90B4A45FC00FC6DFBC3FC48" box="[939,1085,937,964]" pageId="9" pageNumber="44" value="2008-08-03">
<collectingDate id="9B70D77AE90B4A45FC00FC6DFBC3FC48" box="[939,1085,937,964]" pageId="9" pageNumber="44" value="2008-08-03">3. VIII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FBECFC6EFB59FC48" box="[1095,1191,937,964]" pageId="9" pageNumber="44">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FB4FFC6DFAD6FC48" box="[1252,1320,937,963]" pageId="9" pageNumber="44">EUM</collectionCode>
)
</materialsCitation>
; 1 F,
<materialsCitation id="4FE2020FE90B4A45FAC2FC6DFC3CFC6C" collectingDate="2004-08-20" collectingDateMax="2004-08-26" collectingDateMin="2004-08-20" collectionCode="MU" collectorName="K. Yamagishi" country="Japan" location="Katabira" municipality="Kani" pageId="9" pageNumber="44" specimenCount="1" stateProvince="Gifu">
<collectingRegion id="3D4EC6B0E90B4A45FAC2FC6DFF2EFC6C" country="Japan" name="Gifu" pageId="9" pageNumber="44">Gifu Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90B4A45FF77FC09FEEBFC6C" box="[220,277,973,999]" pageId="9" pageNumber="44">Kani</collectingMunicipality>
,
<location id="FA555E89E90B4A45FE8AFC09FE7AFC6C" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FE8AFC09FE7AFC6C" box="[289,388,973,999]" country="Japan" municipality="Kani" name="Katabira" pageId="9" pageNumber="44" stateProvince="Gifu">Katabira</location>
,
<date id="8B342E92E90B4A45FE3BFC09FDA1FC6C" box="[400,607,973,1000]" pageId="9" pageNumber="44" value="2004-08-20" valueMax="2004-08-26" valueMin="2004-08-20">
<collectingDate id="9B70D77AE90B4A45FE3BFC09FDA1FC6C" box="[400,607,973,1000]" pageId="9" pageNumber="44" value="2004-08-20" valueMax="2004-08-26" valueMin="2004-08-20">20–26. VIII. 2004</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FDC1FC0AFCFAFC6C" box="[618,772,973,1000]" pageId="9" pageNumber="44">K. Yamagishi</collectorName>
leg., MsT (
<collectionCode id="999B9097E90B4A45FC2CFC09FC44FC6C" box="[903,954,973,999]" country="USA" name="Midwestern University" pageId="9" pageNumber="44">MU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90B4A45FC64FC0AFB95FC63" box="[975,1131,973,1000]" country="Japan" location="Shikoku" pageId="9" pageNumber="44" specimenCount="1">
[
<emphasis id="CDFED440E90B4A45FC73FC09FBC2FC6C" bold="true" box="[984,1084,973,999]" pageId="9" pageNumber="44">
<location id="FA555E89E90B4A45FC73FC09FBC2FC6C" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FC73FC09FBC2FC6C" box="[984,1084,973,999]" country="Japan" name="Shikoku" pageId="9" pageNumber="44">Shikoku</location>
</emphasis>
]1 F
</materialsCitation>
,
<materialsCitation id="4FE2020FE90B4A45FBDCFC09FCFCFB80" collectingDate="1998-06-28" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Namakusadani" municipality="Odamiyama" pageId="9" pageNumber="44" specimenCount="1" stateProvince="Ehime">
<collectingRegion id="3D4EC6B0E90B4A45FBDCFC09FAFFFC6C" box="[1143,1281,973,999]" country="Japan" name="Ehime" pageId="9" pageNumber="44">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90B4A45FAA7FC09FA66FC6C" box="[1292,1432,973,999]" pageId="9" pageNumber="44">Odamiyama</collectingMunicipality>
,
<location id="FA555E89E90B4A45FF3CFC35FEBEFB80" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FF3CFC35FEBEFB80" box="[151,320,1009,1035]" country="Japan" municipality="Odamiyama" name="Namakusadani" pageId="9" pageNumber="44" stateProvince="Ehime">Namakusadani</location>
,
<date id="8B342E92E90B4A45FEE1FC35FE26FB80" box="[330,472,1009,1036]" pageId="9" pageNumber="44" value="1998-06-28">
<collectingDate id="9B70D77AE90B4A45FEE1FC35FE26FB80" box="[330,472,1009,1036]" pageId="9" pageNumber="44" value="1998-06-28">28. VI. 1998</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FE49FC36FD87FB87" box="[482,633,1010,1036]" pageId="9" pageNumber="44">E. Yamamoto</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FD1DFC35FD04FB80" box="[694,762,1009,1035]" pageId="9" pageNumber="44">EUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE90B4A45FCA7FC35FED9FBA4" collectingDate="1924-07-25" collectionCode="SEHU" collectorName="T. Kurisue" country="South Korea" location="Onseri" pageId="9" pageNumber="44" specimenCount="1" typeStatus="lectotype">
<emphasis id="CDFED440E90B4A45FCA7FC35FC8DFB80" bold="true" box="[780,883,1009,1035]" pageId="9" pageNumber="44">
<collectingCountry id="879D48C2E90B4A45FCA7FC35FC8DFB80" box="[780,883,1009,1035]" name="South Korea" pageId="9" pageNumber="44">KOREA</collectingCountry>
</emphasis>
: 1 F (
<typeStatus id="2031B6F0E90B4A45FC1AFC35FBE0FB80" box="[945,1054,1009,1035]" pageId="9" pageNumber="44" type="lectotype">lectotype</typeStatus>
),
<location id="FA555E89E90B4A45FB9BFC35FB83FB80" LSID="urn:lsid:plazi:treatment:7723B944E90B4A47FF3CFD4BFD2AF8CA:FA555E89E90B4A45FB9BFC35FB83FB80" box="[1072,1149,1009,1035]" country="South Korea" name="Onseri" pageId="9" pageNumber="44">Onseri</location>
,
<date id="8B342E92E90B4A45FB2CFC35FAE0FB80" box="[1159,1310,1009,1036]" pageId="9" pageNumber="44" value="1924-07-25">
<collectingDate id="9B70D77AE90B4A45FB2CFC35FAE0FB80" box="[1159,1310,1009,1036]" pageId="9" pageNumber="44" value="1924-07-25">25. VII. 1924</collectingDate>
</date>
,
<collectorName id="527F6D84E90B4A45FA83FC36FA63FB80" box="[1320,1437,1009,1036]" pageId="9" pageNumber="44">T. Kurisue</collectorName>
leg. (
<collectionCode id="999B9097E90B4A45FF64FBD1FEE0FBA4" box="[207,286,1045,1071]" pageId="9" pageNumber="44">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90B4A47FF6CFBFDFAE5FE26" lastPageId="11" lastPageNumber="46" pageId="9" pageNumber="44" type="description">
<paragraph id="FF350852E90B4A45FF6CFBFDFCABFBF3" blockId="9.[151,1437,829,1936]" pageId="9" pageNumber="44">
<emphasis id="CDFED440E90B4A45FF6CFBFDFEAFFBD8" bold="true" box="[199,337,1081,1107]" pageId="9" pageNumber="44">Description</emphasis>
.
<emphasis id="CDFED440E90B4A45FEF4FBFDFE48FBD8" bold="true" box="[351,438,1081,1107]" pageId="9" pageNumber="44">Female</emphasis>
(n = 7). Body polished and covered with punctures and silvery setae; length
<quantity id="3872A5B7E90B4A45FAA4FBFDFA67FBDF" box="[1295,1433,1081,1108]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="8.4" metricValueMin="6.7" pageId="9" pageNumber="44" unit="mm" value="7.55" valueMax="8.4" valueMin="6.7">6.7–8.4 mm</quantity>
. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E90B4A45FF6CFB45FE1DFA1C" blockId="9.[151,1437,829,1936]" pageId="9" pageNumber="44">
Head 0.6–0.65 × as long as wide. Combined length of clypeus and face 1.0 × facial width. Clypeus matt or polished, dorsally punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.8–0.85 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.5–1.65 × as long as OD. OOL 0.95–1.1 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E90B4A45FCF3FAF1FC43FAC4" box="[856,957,1333,1359]" captionStart="FIGURE 6" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-19@10.[151,1436,181,1723]" captionTargetPageId="10" captionText="FIGURE 6. Syzeuctus coreanus Uchida, 1928, female—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view." pageId="9" pageNumber="44">Fig. 6 D</figureCitation>
). Antenna with 38–40 flagellomeres. FL I 4.15–4.4 × as long as maximum depth in lateral view and 1.65–1.75 × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E90B4A45FF6CFA66FD2FF974" blockId="9.[151,1437,829,1936]" pageId="9" pageNumber="44">
Mesosoma coarsely and densely punctate, 1.5 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, short, its dorsal part largely absent. Mesopleuron densely punctate; speculum with a large smooth, weakly concave area (
<figureCitation id="67B114D7E90B4A45FCA2FA2DFC8FF988" box="[777,881,1513,1539]" captionStart="FIGURE 6" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-19@10.[151,1436,181,1723]" captionTargetPageId="10" captionText="FIGURE 6. Syzeuctus coreanus Uchida, 1928, female—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view." pageId="9" pageNumber="44">Fig. 6 D</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E90B4A45FB2CF9C9FB18F9AC" box="[1159,1254,1549,1575]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="9" pageNumber="44">Fig. 1 C</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum present and complete. Fore wing length 5.45–6.0 mm. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E90B4A45FEE2F9BDFE58F918" box="[329,422,1657,1683]" captionStart="FIGURE 6" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-19@10.[151,1436,181,1723]" captionTargetPageId="10" captionText="FIGURE 6. Syzeuctus coreanus Uchida, 1928, female—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view." pageId="9" pageNumber="44">Fig. 6 B</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.5–6.6 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 3.9–4.2: 2.0: 1.35–1.4: 0.7: 0.8. Hind tarsal claws pectinate by short teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E90B4A45FDCFF921FD3AF974" box="[612,708,1765,1791]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="9" pageNumber="44">Fig. 2 C</figureCitation>
).
</paragraph>
<paragraph id="FF350852E90B4A45FF6CF8CEFCC8F804" blockId="9.[151,1437,829,1936]" pageId="9" pageNumber="44">Metasoma. T I 2.1–2.5 × as long as maximum width, punctate except for smooth areas on median longitudinal smooth area, area along posterior margin and lateral sides. T II 1.45–1.65 × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 2.35–2.55 × as long as hind tibia, ca. 1.0 × as long as fore wing, and shorter than body length.</paragraph>
<caption id="ABF558DAE9084A46FF3CF917FDC3F884" pageId="10" pageNumber="45" startId="10.[151,250,1747,1771]" targetBox="[151,1436,181,1723]" targetPageId="10" targetType="figure">
<paragraph id="FF350852E9084A46FF3CF917FDC3F884" blockId="10.[151,1436,1747,1807]" pageId="10" pageNumber="45">
<emphasis id="CDFED440E9084A46FF3CF917FEEAF960" bold="true" box="[151,276,1747,1771]" pageId="10" pageNumber="45">FIGURE 6.</emphasis>
<taxonomicName id="388A73D1E9084A46FEB7F910FD88F960" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[284,630,1747,1771]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="CDFED440E9084A46FEB7F910FE1CF960" box="[284,482,1748,1771]" italics="true" pageId="10" pageNumber="45">Syzeuctus coreanus</emphasis>
<bibRefCitation id="9B1B75A3E9084A46FE41F917FD88F960" author="Uchida, T." box="[490,630,1747,1771]" pageId="10" pageNumber="45" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
, female—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view.
</paragraph>
</caption>
<paragraph id="FF350852E9084A47FF6CF8F9FAD0FECA" blockId="10.[151,1437,1853,2024]" lastBlockId="11.[151,1437,151,429]" lastPageId="11" lastPageNumber="46" pageId="10" pageNumber="45">
Colouration (
<figureCitation id="67B114D7E9084A46FEF5F8F9FE17F8D3" box="[350,489,1853,1880]" captionStart="FIGURE 6" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-19@10.[151,1436,181,1723]" captionTargetPageId="10" captionText="FIGURE 6. Syzeuctus coreanus Uchida, 1928, female—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view." pageId="10" pageNumber="45">Figs 6 A–D</figureCitation>
). Body (excluding wings) yellow. Frons, vertex, and gena except for broad yellow area long orbits, occiput, mandibular teeth, dorso-median black spot and postero-lateral triangular spot of pronotum, median longitudinal spot and U-shaped posterior spot of mesoscutum, axillae, baso-median small spot of scutellum, antero-ventral oblique spot and postero-dorsal curved spot of mesopleuron, W-shaped black marking of propodeum, median small spot of posterior margin of propodeum, base of hind coxa, hind trochanter, T I to T IV except for anterior and posterior margins of each segment, and T V to T VII except for each posterior margin black to blackish-brown. Black spot of scutellum sometimes absent. Scape and pedicel blackish-brown dorsally, yellow ventrally. Flagellum and ovipositor brown to reddish-brown. Yellow parts of metasomal tergites, femora, and tibiae sometimes more or less tinged with reddish-brown. Hind tarsus tinged with brown. Wings hyaline except for weak apical clouded area of fore wing. Veins and pterostigma blackish-brown except for yellowish-brown wing base.
</paragraph>
<paragraph id="FF350852E9094A47FF6CFE8FFAE5FE26" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="46">
<emphasis id="CDFED440E9094A47FF6CFE8FFEFAFEEE" bold="true" box="[199,260,331,357]" pageId="11" pageNumber="46">Male</emphasis>
(n = 1). Similar to female (
<figureCitation id="67B114D7E9094A47FD9DFE8FFD43FEEE" box="[566,701,331,357]" captionStart="FIGURE 7" captionStartId="11.[151,250,1617,1641]" captionTargetBox="[151,1436,453,1593]" captionTargetId="figure-157@11.[151,1436,453,1593]" captionTargetPageId="11" captionText="FIGURE 7. Syzeuctus coreanus Uchida, 1928, male—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: propodeum, dorsal view." pageId="11" pageNumber="46">Figs 7 A–D</figureCitation>
). Combined length of clypeus and face 0.85 × facial width. Malar space 0.95 × as long as basal mandibular width. Antenna with 42 flagellomeres. Hind femur 5.35 × as long as maximum depth in lateral view. Tergal spiracles not enlarged. Black areas of body smaller than female.
</paragraph>
</subSubSection>
<caption id="ABF558DAE9094A47FF3CF995FD89F906" pageId="11" pageNumber="46" startId="11.[151,250,1617,1641]" targetBox="[151,1436,453,1593]" targetPageId="11" targetType="figure">
<paragraph id="FF350852E9094A47FF3CF995FD89F906" blockId="11.[151,1437,1617,1677]" pageId="11" pageNumber="46">
<emphasis id="CDFED440E9094A47FF3CF995FEEAF9E2" bold="true" box="[151,276,1617,1641]" pageId="11" pageNumber="46">FIGURE 7.</emphasis>
<taxonomicName id="388A73D1E9094A47FEB0F996FD8BF9E2" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[283,629,1617,1641]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="CDFED440E9094A47FEB0F996FE1FF9E2" box="[283,481,1618,1641]" italics="true" pageId="11" pageNumber="46">Syzeuctus coreanus</emphasis>
<bibRefCitation id="9B1B75A3E9094A47FE42F995FD8BF9E2" author="Uchida, T." box="[489,629,1617,1641]" pageId="11" pageNumber="46" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
, male—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: propodeum, dorsal view.
</paragraph>
</caption>
<subSubSection id="B7905BD9E9094A47FF6CF97FFE36F972" pageId="11" pageNumber="46" type="distribution">
<paragraph id="FF350852E9094A47FF6CF97FFBBDF95E" blockId="11.[151,1436,1723,1858]" box="[199,1091,1723,1750]" pageId="11" pageNumber="46">
<emphasis id="CDFED440E9094A47FF6CF97FFEA7F95E" bold="true" box="[199,345,1723,1749]" pageId="11" pageNumber="46">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9094A47FECCF978FE59F95D" box="[359,423,1724,1750]" name="Japan" pageId="11" pageNumber="46">Japan</collectingCountry>
(Honshu and Shikoku),
<collectingCountry id="879D48C2E9094A47FD11F97FFD03F95E" box="[698,765,1723,1749]" name="China" pageId="11" pageNumber="46">China</collectingCountry>
and
<collectingCountry id="879D48C2E9094A47FC98F978FC85F95D" box="[819,891,1724,1750]" name="South Korea" pageId="11" pageNumber="46">Korea</collectingCountry>
. New to Shikoku.
</paragraph>
<paragraph id="FF350852E9094A47FF6CF91BFE36F972" blockId="11.[151,1436,1723,1858]" box="[199,456,1759,1786]" pageId="11" pageNumber="46">
<emphasis id="CDFED440E9094A47FF6CF91BFEBDF972" bold="true" box="[199,323,1759,1785]" pageId="11" pageNumber="46">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9094A47FF6CF8C0FD2AF8CA" pageId="11" pageNumber="46" type="discussion">
<paragraph id="FF350852E9094A47FF6CF8C0FD2AF8CA" blockId="11.[151,1436,1723,1858]" pageId="11" pageNumber="46">
<emphasis id="CDFED440E9094A47FF6CF8C0FECAF895" bold="true" box="[199,308,1796,1822]" pageId="11" pageNumber="46">Remarks</emphasis>
. The photo of this species in
<bibRefCitation id="9B1B75A3E9094A47FD2AF8C0FCB7F895" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[641,841,1795,1822]" pageId="11" pageNumber="46" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang
<emphasis id="CDFED440E9094A47FD6EF8C0FD00F896" box="[709,766,1795,1821]" italics="true" pageId="11" pageNumber="46">et al.</emphasis>
(2020
</bibRefCitation>
,
<figureCitation id="67B114D7E9094A47FCFEF8C7FC4CF895" box="[853,946,1795,1822]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="11" pageNumber="46">Fig. 2 B</figureCitation>
) probably shows
<taxonomicName id="388A73D1E9094A47FBD0F8C0FAF5F896" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[1147,1291,1796,1822]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9094A47FBD0F8C0FAF5F896" box="[1147,1291,1796,1822]" italics="true" pageId="11" pageNumber="46">S. yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9094A47FAB9F8C0FA92F895" bold="true" box="[1298,1388,1796,1822]" pageId="11" pageNumber="46">
<taxonomicNameLabel id="D6CD693BE9094A47FAB9F8C0FA92F895" box="[1298,1388,1796,1822]" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
due to a different coloration of mesosoma and hind leg.
</paragraph>
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<paragraph id="FF350852E90C4A42FF3CFAEBFD6AFAC2" blockId="14.[151,660,1326,1389]" box="[151,660,1326,1353]" pageId="14" pageNumber="49">
<heading id="A47DBF3EE90C4A42FF3CFAEBFD6AFAC2" bold="true" box="[151,660,1326,1353]" fontSize="11" level="1" pageId="14" pageNumber="49" reason="1">
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<emphasis id="CDFED440E90C4A42FF3CFAEBFE57FAC2" bold="true" box="[151,425,1326,1353]" italics="true" pageId="14" pageNumber="49">Syzeuctus flavofacialis</emphasis>
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<paragraph id="FF350852E90C4A42FF3CFA97FDDEFAE6" blockId="14.[151,660,1326,1389]" box="[151,544,1363,1389]" pageId="14" pageNumber="49">
(
<figureCitation id="67B114D7E90C4A42FF34FA97FEFEFAE6" box="[159,256,1363,1389]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="14" pageNumber="49">Figs 1 E</figureCitation>
,
<figureCitation id="67B114D7E90C4A42FEA7FA97FEA9FAE6" box="[268,343,1363,1389]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="14" pageNumber="49">2 E–G</figureCitation>
,
<figureCitation id="67B114D7E90C4A42FEC9FA97FE49FAE6" box="[354,439,1363,1389]" captionStart="FIGURE 10" captionStartId="15.[151,250,1813,1837]" captionTargetBox="[151,1436,181,1789]" captionTargetId="figure-19@15.[151,1436,181,1789]" captionTargetPageId="15" captionText="FIGURE 10. Syzeuctus flavofacialis Kang &amp; Lee, 2020, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view; E: fore wing; F: apical part of hind tibia and hind tarsus." pageId="14" pageNumber="49">10 A–F</figureCitation>
,
<figureCitation id="67B114D7E90C4A42FE6AFA97FDE6FAE6" box="[449,536,1363,1389]" captionStart="FIGURE 11" captionStartId="16.[151,250,1518,1542]" captionTargetBox="[151,1436,633,1494]" captionTargetId="figure-233@16.[151,1436,633,1494]" captionTargetPageId="16" captionText="FIGURE 11. Syzeuctus flavofacialis Kang &amp; Lee, 2020, male from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view." pageId="14" pageNumber="49">11 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90C4A42FF3CFA5CFCB6FA24" box="[151,840,1431,1456]" pageId="14" pageNumber="49" type="reference_group">
<paragraph id="FF350852E90C4A42FF3CFA5CFCB6FA24" blockId="14.[151,840,1431,1456]" box="[151,840,1431,1456]" pageId="14" pageNumber="49">
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<taxonomicName id="388A73D1E90C4A42FF3CFA5CFDBEFA24" authority="Kang &amp; Lee, 2020" authorityName="Kang &amp; Lee" authorityYear="2020" box="[151,576,1431,1456]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="flavofacialis">
<emphasis id="CDFED440E90C4A42FF3CFA5CFE83FA24" box="[151,381,1431,1455]" italics="true" pageId="14" pageNumber="49">Syzeuctus flavofacialis</emphasis>
<bibRefCitation id="9B1B75A3E90C4A42FE2FFA5CFDBEFA24" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[388,576,1431,1456]" pageId="14" pageNumber="49" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang &amp; Lee,
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in
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<bibRefCitation id="9B1B75A3E90C4A42FDCAFA5CFCE0FA24" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[609,798,1431,1456]" pageId="14" pageNumber="49" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang
<emphasis id="CDFED440E90C4A42FD35FA5CFD2FFA24" box="[670,721,1431,1455]" italics="true" pageId="14" pageNumber="49">et al.</emphasis>
(2020)
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: 24
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.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90C4A42FF3CFA19FE37F903" pageId="14" pageNumber="49" type="materials_examined">
<paragraph id="FF350852E90C4A42FF3CFA19FE37F903" blockId="14.[151,1437,1501,2032]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF3CFA19FE7BFA7C" bold="true" box="[151,389,1501,1527]" pageId="14" pageNumber="49">
<collectingRegion id="3D4EC6B0E90C4A42FF3CFA19FEF4FA7C" box="[151,266,1501,1527]" pageId="14" pageNumber="49">Materials</collectingRegion>
examined
</emphasis>
.
<materialsCitation id="4FE2020FE90C4A42FE3FFA19FF03F997" collectingDate="1997-08-07" collectionCode="NARO" collectorName="T. Nambu" country="Japan" location="Sugaya" municipality="Ranzan Town" pageId="14" pageNumber="49" specimenCount="1" stateProvince="Saitama">
<emphasis id="CDFED440E90C4A42FE3FFA19FE10FA7C" bold="true" box="[404,494,1501,1527]" pageId="14" pageNumber="49">
<collectingCountry id="879D48C2E90C4A42FE3FFA19FE10FA7C" box="[404,494,1501,1527]" name="Japan" pageId="14" pageNumber="49">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E90C4A42FDACFA1AFD9AFA73" bold="true" box="[519,612,1502,1528]" pageId="14" pageNumber="49">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E90C4A42FD0CFA19FCBCFA7C" box="[679,834,1501,1527]" country="Japan" name="Saitama" pageId="14" pageNumber="49">Saitama Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90C4A42FCE5FA1AFC15FA73" box="[846,1003,1502,1528]" pageId="14" pageNumber="49">Ranzan Town</collectingMunicipality>
,
<location id="FA555E89E90C4A42FC53FA1AFBB3FA73" LSID="urn:lsid:plazi:treatment:7723B944E90C4A5DFF3CFAEBFD66FD46:FA555E89E90C4A42FC53FA1AFBB3FA73" box="[1016,1101,1502,1528]" country="Japan" municipality="Ranzan Town" name="Sugaya" pageId="14" pageNumber="49" stateProvince="Saitama">Sugaya</location>
,
<date id="8B342E92E90C4A42FBF2FA1AFB11FA73" box="[1113,1263,1502,1528]" pageId="14" pageNumber="49" value="1997-08-07">
<collectingDate id="9B70D77AE90C4A42FBF2FA1AFB11FA73" box="[1113,1263,1502,1528]" pageId="14" pageNumber="49" value="1997-08-07">7. VIII. 1997</collectingDate>
</date>
,
<collectorName id="527F6D84E90C4A42FB50FA1AFA95FA7C" box="[1275,1387,1501,1528]" pageId="14" pageNumber="49">T. Nambu</collectorName>
leg. (
<collectionCode id="999B9097E90C4A42FF34F9C6FF0BF997" box="[159,245,1538,1564]" pageId="14" pageNumber="49">NARO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90C4A42FEA7F9C6FA9EF997" box="[268,1376,1537,1564]" collectingDate="2011-07-17" collectionCode="KPMNH" collectorName="N. Kikuchi" country="Japan" location="Shimoshingashi" municipality="Kawagoe City" pageId="14" pageNumber="49" specimenCount="1" stateProvince="Saitama">
1 M,
<collectingRegion id="3D4EC6B0E90C4A42FEFBF9C5FE10F990" box="[336,494,1537,1563]" country="Japan" name="Saitama" pageId="14" pageNumber="49">Saitama Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90C4A42FE57F9C6FD5CF990" box="[508,674,1537,1564]" pageId="14" pageNumber="49">Kawagoe City</collectingMunicipality>
,
<location id="FA555E89E90C4A42FD1AF9C5FC99F990" LSID="urn:lsid:plazi:treatment:7723B944E90C4A5DFF3CFAEBFD66FD46:FA555E89E90C4A42FD1AF9C5FC99F990" box="[689,871,1537,1563]" country="Japan" municipality="Kawagoe City" name="Shimoshingashi" pageId="14" pageNumber="49" stateProvince="Saitama">Shimoshingashi</location>
,
<date id="8B342E92E90C4A42FCDDF9C6FBE8F997" box="[886,1046,1538,1564]" pageId="14" pageNumber="49" value="2011-07-17">
<collectingDate id="9B70D77AE90C4A42FCDDF9C6FBE8F997" box="[886,1046,1538,1564]" pageId="14" pageNumber="49" value="2011-07-17">17. VII. 2011</collectingDate>
</date>
,
<collectorName id="527F6D84E90C4A42FB8EF9C6FB5BF990" box="[1061,1189,1537,1564]" pageId="14" pageNumber="49">N. Kikuchi</collectorName>
leg. (
<collectionCode id="999B9097E90C4A42FB47F9C6FAA6F997" box="[1260,1368,1538,1564]" pageId="14" pageNumber="49">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90C4A42FAC4F9C6FC1DF9CB" collectingDate="1987-08-13" collectionCode="NARO" collectorName="S. Saito" country="Japan" location="Shichikokuyama" municipality="Machida" pageId="14" pageNumber="49" specimenCount="1" stateProvince="Tokyo">
1 F,
<collectingRegion id="3D4EC6B0E90C4A42FF3CF9E1FF1FF9B4" box="[151,225,1573,1599]" country="Japan" name="Tokyo" pageId="14" pageNumber="49">Tokyo</collectingRegion>
,
<collectingMunicipality id="1F519228E90C4A42FF44F9E1FEAAF9B4" box="[239,340,1573,1599]" pageId="14" pageNumber="49">Machida</collectingMunicipality>
,
<location id="FA555E89E90C4A42FEC9F9E1FDDFF9B4" LSID="urn:lsid:plazi:treatment:7723B944E90C4A5DFF3CFAEBFD66FD46:FA555E89E90C4A42FEC9F9E1FDDFF9B4" box="[354,545,1573,1599]" country="Japan" municipality="Machida" name="Shichikokuyama" pageId="14" pageNumber="49" stateProvince="Tokyo">Shichikokuyama</location>
,
<date id="8B342E92E90C4A42FD84F9E2FD26F9CB" box="[559,728,1574,1600]" pageId="14" pageNumber="49" value="1987-08-13">
<collectingDate id="9B70D77AE90C4A42FD84F9E2FD26F9CB" box="[559,728,1574,1600]" pageId="14" pageNumber="49" value="1987-08-13">13. VIII. 1987</collectingDate>
</date>
,
<collectorName id="527F6D84E90C4A42FD4DF9E2FCBFF9B4" box="[742,833,1573,1600]" pageId="14" pageNumber="49">S. Saito</collectorName>
leg. (
<collectionCode id="999B9097E90C4A42FC2EF9E2FC25F9CB" box="[901,987,1574,1600]" pageId="14" pageNumber="49">NARO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90C4A42FC5AF9E2FCFAF9EF" collectingDate="2002-08-30" collectionCode="KPMNH" collectorName="I. Waki" country="Japan" location="Ikutaryokuchi" municipality="Kawasaki City" pageId="14" pageNumber="49" specimenCount="1" stateProvince="Kanagawa">
1 F,
<collectingRegion id="3D4EC6B0E90C4A42FB83F9E2FB1FF9B4" box="[1064,1249,1573,1600]" country="Japan" name="Kanagawa" pageId="14" pageNumber="49">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90C4A42FB45F9E1FA66F9B4" box="[1262,1432,1573,1599]" pageId="14" pageNumber="49">Kawasaki City</collectingMunicipality>
,
<location id="FA555E89E90C4A42FF3CF98DFEC9F9E8" LSID="urn:lsid:plazi:treatment:7723B944E90C4A5DFF3CFAEBFD66FD46:FA555E89E90C4A42FF3CF98DFEC9F9E8" box="[151,311,1609,1635]" country="Japan" municipality="Kawasaki City" name="Ikutaryokuchi" pageId="14" pageNumber="49" stateProvince="Kanagawa">Ikutaryokuchi</location>
,
<date id="8B342E92E90C4A42FEEEF98EFE12F9EF" box="[325,492,1610,1636]" pageId="14" pageNumber="49" value="2002-08-30">
<collectingDate id="9B70D77AE90C4A42FEEEF98EFE12F9EF" box="[325,492,1610,1636]" pageId="14" pageNumber="49" value="2002-08-30">30. VIII. 2002</collectingDate>
</date>
,
<collectorName id="527F6D84E90C4A42FE52F98EFDB2F9E8" box="[505,588,1609,1636]" pageId="14" pageNumber="49">I. Waki</collectorName>
leg. (
<collectionCode id="999B9097E90C4A42FD3BF98EFD02F9EF" box="[656,764,1610,1636]" pageId="14" pageNumber="49">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90C4A42FCBAF98EFE3AF903" collectingDate="2010-07-28" collectionCode="ZSM" collectorName="K. Kubo" country="Japan" location="Enkaizan" municipality="Yokohama City" pageId="14" pageNumber="49" specimenCount="1" stateProvince="Kanagawa">
1 F,
<collectingRegion id="3D4EC6B0E90C4A42FCEDF98EFC00F9E8" box="[838,1022,1609,1636]" country="Japan" name="Kanagawa" pageId="14" pageNumber="49">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90C4A42FBA1F98DFB42F9E8" box="[1034,1212,1609,1635]" pageId="14" pageNumber="49">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E90C4A42FB61F98DFACBF9E8" LSID="urn:lsid:plazi:treatment:7723B944E90C4A5DFF3CFAEBFD66FD46:FA555E89E90C4A42FB61F98DFACBF9E8" box="[1226,1333,1609,1635]" country="Japan" municipality="Yokohama City" name="Enkaizan" pageId="14" pageNumber="49" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E90C4A42FAE8F98EFF2CF903" pageId="14" pageNumber="49" value="2010-07-28">
<collectingDate id="9B70D77AE90C4A42FAE8F98EFF2CF903" pageId="14" pageNumber="49" value="2010-07-28">28. VII. 2010</collectingDate>
</date>
,
<collectorName id="527F6D84E90C4A42FF76F9AAFEC3F90C" box="[221,317,1645,1672]" pageId="14" pageNumber="49">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E90C4A42FED6F9AAFE42F903" box="[381,444,1646,1672]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="49" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90C4A5DFF6CF955FC55FE7E" lastPageId="17" lastPageNumber="52" pageId="14" pageNumber="49" type="description">
<paragraph id="FF350852E90C4A42FF6CF955FCABF95B" blockId="14.[151,1437,1501,2032]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF6CF955FEAFF920" bold="true" box="[199,337,1681,1707]" pageId="14" pageNumber="49">Description</emphasis>
.
<emphasis id="CDFED440E90C4A42FEF4F956FE48F927" bold="true" box="[351,438,1682,1708]" pageId="14" pageNumber="49">Female</emphasis>
(n = 4). Body polished and covered with punctures and silvery setae; length 8.0–9.0 mm. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E90C4A42FF6CF91DFD5CF864" blockId="14.[151,1437,1501,2032]" pageId="14" pageNumber="49">
Head 0.55–0.6 × as long as wide. Combined length of clypeus and face 1.05–1.15 × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, with a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns, with a pair of slight convexities. Malar space 0.5 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.6–1.75 × as long as OD. OOL 1.0 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E90C4A42FB9DF849FB5BF823" box="[1078,1189,1933,1960]" captionStart="FIGURE 10" captionStartId="15.[151,250,1813,1837]" captionTargetBox="[151,1436,181,1789]" captionTargetId="figure-19@15.[151,1436,181,1789]" captionTargetPageId="15" captionText="FIGURE 10. Syzeuctus flavofacialis Kang &amp; Lee, 2020, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view; E: fore wing; F: apical part of hind tibia and hind tarsus." pageId="14" pageNumber="49">Fig. 10 D</figureCitation>
). Antenna with 40–44 flagellomeres. FL I 4.25–4.65 × as long as maximum depth in lateral view and 1.7–1.85 × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<caption id="ABF558DAE90D4A43FF3CF8D1FE44F8FE" pageId="15" pageNumber="50" startId="15.[151,250,1813,1837]" targetBox="[151,1436,181,1789]" targetPageId="15" targetType="figure">
<paragraph id="FF350852E90D4A43FF3CF8D1FE44F8FE" blockId="15.[151,1437,1813,1909]" pageId="15" pageNumber="50">
<emphasis id="CDFED440E90D4A43FF3CF8D1FEDDF8A6" bold="true" box="[151,291,1813,1837]" pageId="15" pageNumber="50">FIGURE 10.</emphasis>
<taxonomicName id="388A73D1E90D4A43FE86F8D1FD14F8A6" authority="Kang &amp; Lee, 2020" authorityName="Kang &amp; Lee" authorityYear="2020" box="[301,746,1813,1838]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="50" phylum="Arthropoda" rank="species" species="flavofacialis">
<emphasis id="CDFED440E90D4A43FE86F8D1FDE8F8A6" box="[301,534,1813,1837]" italics="true" pageId="15" pageNumber="50">Syzeuctus flavofacialis</emphasis>
<bibRefCitation id="9B1B75A3E90D4A43FD8BF8D2FD14F8A6" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[544,746,1813,1838]" pageId="15" pageNumber="50" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">Kang &amp; Lee, 2020</bibRefCitation>
</taxonomicName>
, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view; E: fore wing; F: apical part of hind tibia and hind tarsus.
</paragraph>
</caption>
<paragraph id="FF350852E90D4A5CFF6CF860FC2CFE5A" blockId="15.[151,1436,1955,2018]" lastBlockId="16.[151,1437,151,609]" lastPageId="16" lastPageNumber="51" pageId="15" pageNumber="50">
Mesosoma coarsely and densely punctate, 1.4–1.45 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, short, its dorsal part largely absent. Mesopleuron densely punctate; speculum with a large smooth area (
<figureCitation id="67B114D7E9124A5CFD9BFF53FD5FFF3A" box="[560,673,151,177]" captionStart="FIGURE 10" captionStartId="15.[151,250,1813,1837]" captionTargetBox="[151,1436,181,1789]" captionTargetId="figure-19@15.[151,1436,181,1789]" captionTargetPageId="15" captionText="FIGURE 10. Syzeuctus flavofacialis Kang &amp; Lee, 2020, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view; E: fore wing; F: apical part of hind tibia and hind tarsus." pageId="16" pageNumber="51">Fig. 10 D</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E9124A5CFC00FF7FFBF7FF5E" box="[939,1033,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="16" pageNumber="51">Fig. 1 E</figureCitation>
). Dorsal surface of propodeum with transverse rugae just in front of posterior transverse carina. Posterior transverse carina of propodeum present and complete. Fore wing length
<quantity id="3872A5B7E9124A5CFE7EFEC7FD9EFE96" box="[469,608,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.3" metricValueMin="6.6" pageId="16" pageNumber="51" unit="mm" value="6.949999999999999" valueMax="7.3" valueMin="6.6">6.6–7.3 mm</quantity>
. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole longer than areolet (
<figureCitation id="67B114D7E9124A5CFDF2FEE3FD32FECA" box="[601,716,295,321]" captionStart="FIGURE 10" captionStartId="15.[151,250,1813,1837]" captionTargetBox="[151,1436,181,1789]" captionTargetId="figure-19@15.[151,1436,181,1789]" captionTargetPageId="15" captionText="FIGURE 10. Syzeuctus flavofacialis Kang &amp; Lee, 2020, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view; E: fore wing; F: apical part of hind tibia and hind tarsus." pageId="16" pageNumber="51">Fig. 10 E</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.2–5.7 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.0: 2.0: 1.4–1.5: 0.7: 0.8–0.85. Hind tarsal claws simple except for minute subbasal teeth, sometimes with a small tooth, strongly bent apically in outer tooth (ca. 90°), moderately bent apically in inner tooth (less than 90°) (
<figureCitation id="67B114D7E9124A5CFCEBFE73FC39FE5A" box="[832,967,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="16" pageNumber="51">Figs 2 E–G</figureCitation>
).
</paragraph>
<paragraph id="FF350852E9124A5CFF6CFE1FFF1BFDEA" blockId="16.[151,1437,151,609]" pageId="16" pageNumber="51">Metasoma. T I 1.5 × as long as maximum width, nearly entirely punctate except for small median longitudinal smooth area. T II 1.05–1.1 × as long as maximum width, densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 2.85–3.15 × as long as hind tibia, ca. 1.1 × as long as fore wing, and shorter than body length.</paragraph>
<caption id="ABF558DAE9124A5CFF3CFA2AFD97F9A1" pageId="16" pageNumber="51" startId="16.[151,250,1518,1542]" targetBox="[151,1436,633,1494]" targetPageId="16" targetType="figure">
<paragraph id="FF350852E9124A5CFF3CFA2AFD97F9A1" blockId="16.[151,1437,1518,1578]" pageId="16" pageNumber="51">
<emphasis id="CDFED440E9124A5CFF3CFA2AFEDDF98D" bold="true" box="[151,291,1518,1542]" pageId="16" pageNumber="51">FIGURE 11.</emphasis>
<taxonomicName id="388A73D1E9124A5CFE85FA2AFD0CF98D" authority="Kang &amp; Lee, 2020" authorityName="Kang &amp; Lee" authorityYear="2020" box="[302,754,1518,1543]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="flavofacialis">
<emphasis id="CDFED440E9124A5CFE85FA2AFDE7F98D" box="[302,537,1518,1542]" italics="true" pageId="16" pageNumber="51">Syzeuctus flavofacialis</emphasis>
<bibRefCitation id="9B1B75A3E9124A5CFD8FFA2BFD0CF98D" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[548,754,1518,1543]" pageId="16" pageNumber="51" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">Kang &amp; Lee, 2020</bibRefCitation>
</taxonomicName>
, male from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view.
</paragraph>
</caption>
<paragraph id="FF350852E9124A5DFF6CF99CFC80FE02" blockId="16.[151,1437,1624,2011]" lastBlockId="17.[151,1437,151,717]" lastPageId="17" lastPageNumber="52" pageId="16" pageNumber="51">
Colouration (
<figureCitation id="67B114D7E9124A5CFEF0F99CFE17F9F9" box="[347,489,1624,1650]" captionStart="FIGURE 10" captionStartId="15.[151,250,1813,1837]" captionTargetBox="[151,1436,181,1789]" captionTargetId="figure-19@15.[151,1436,181,1789]" captionTargetPageId="15" captionText="FIGURE 10. Syzeuctus flavofacialis Kang &amp; Lee, 2020, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, pronotum, and mesopleuron, lateral view; E: fore wing; F: apical part of hind tibia and hind tarsus." pageId="16" pageNumber="51">Figs 10 A–F</figureCitation>
). Body (excluding wings and legs) black. Clypeus yellow, usually darkened ventrally. Mandible except for teeth, face except for a median longitudinal black stripe and area around anterior tentorial pits, malar space, inner orbit on frons and vertex, and outer orbit except for dorsal part yellow. Scape and pedicel partly narrowly tinged with yellow. Apical part of flagellum brown. Ventral part of flagellum and palpi largely tinged with reddish-brown. Pronotum except for median longitudinal black stripe yellow. Propleuron except for anterior black marking yellow. Mesoscutum with large M-shaped yellow marking. Subtegular ridge, antero-dorsal and small postero-ventral spots of mesopleuron, tegula, scutellum, postscutellum, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of propodeum yellow.A pair of antero-lateral yellow spots of propodeum sometimes united into a posterior triangle yellow marking of propodeum. T I yellow except for a subapical black stripe. T II to T IV each with yellow or reddish-brown base and yellow posterior margin. Reddish-brown area of metasomal tergites sometimes partly darkened. Membranous parts of metasomal sternites largely yellow. Subgenital plate blackish-brown except of reddish-brown apex. Ovipositor reddish-brown. Wings hyaline. Apical clouded area of fore wing slightly present. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow except for mid fifth tarsomere black. Hind coxa black with yellow apical part and a dorsal small yellow spot. Hind trochanter black except for dorsal and apical yellow parts. Hind trochantellus yellow. Hind femur reddish-brown, weakly tinged with yellow apically. Hind tibia yellow basally, blackish-brown apically. Hind tibial spurs blackish-brown. Hind tarsus blackish-brown except for median part of first tarsomere, basal parts of second and third tarsomeres, and fourth tarsomere yellowish-brown to yellow.
</paragraph>
<paragraph id="FF350852E9134A5DFF6CFE57FC55FE7E" blockId="17.[151,1437,151,717]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF6CFE57FEFAFE26" bold="true" box="[199,260,403,429]" pageId="17" pageNumber="52">Male</emphasis>
(n = 1). Similar to female (
<figureCitation id="67B114D7E9134A5DFD9EFE57FD38FE26" box="[565,710,403,429]" captionStart="FIGURE 11" captionStartId="16.[151,250,1518,1542]" captionTargetBox="[151,1436,633,1494]" captionTargetId="figure-233@16.[151,1436,633,1494]" captionTargetPageId="16" captionText="FIGURE 11. Syzeuctus flavofacialis Kang &amp; Lee, 2020, male from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view." pageId="17" pageNumber="52">Figs 11 A–C</figureCitation>
). Malar space 0.65 × as long as basal mandibular width. POL 1.8 × as long as OD. OOL 0.8 × as long as OD. FL I 4.8 × as long as maximum depth in lateral view. Tergal spiracles not enlarged. Face entirely yellow. T III to T V largely reddish-yellow.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9134A5DFF6CFE3AFE36FDB6" pageId="17" pageNumber="52" type="distribution">
<paragraph id="FF350852E9134A5DFF6CFE3AFCBFFD92" blockId="17.[151,1437,151,717]" box="[199,833,510,537]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF6CFE3AFEA7FD93" bold="true" box="[199,345,510,536]" pageId="17" pageNumber="52">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9134A5DFECCFE3BFE59FD92" box="[359,423,511,537]" name="Japan" pageId="17" pageNumber="52">Japan</collectingCountry>
(Honshu) and
<collectingCountry id="879D48C2E9134A5DFDE5FE3BFD68FD92" box="[590,662,511,537]" name="South Korea" pageId="17" pageNumber="52">Korea</collectingCountry>
. New to
<collectingCountry id="879D48C2E9134A5DFD52FE3BFCC2FD92" box="[761,828,511,537]" name="Japan" pageId="17" pageNumber="52">Japan</collectingCountry>
.
</paragraph>
<paragraph id="FF350852E9134A5DFF6CFDE6FE36FDB6" blockId="17.[151,1437,151,717]" box="[199,456,546,573]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF6CFDE6FEBDFDB7" bold="true" box="[199,323,546,572]" pageId="17" pageNumber="52">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9134A5DFF6CFD83FD66FD46" pageId="17" pageNumber="52" type="discussion">
<paragraph id="FF350852E9134A5DFF6CFD83FD66FD46" blockId="17.[151,1437,151,717]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF6CFD83FECAFDEA" bold="true" box="[199,308,583,609]" pageId="17" pageNumber="52">Remarks</emphasis>
. Japanese specimens differ from the original description based on Korean specimens (
<bibRefCitation id="9B1B75A3E9134A5DFAB1FD83FF2FFD0E" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." pageId="17" pageNumber="52" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang
<emphasis id="CDFED440E9134A5DFAC9FD83FA63FDEA" box="[1378,1437,583,609]" italics="true" pageId="17" pageNumber="52">et al.</emphasis>
2020
</bibRefCitation>
) in the body size (Japanese at most 9.0 mm, Korean specimen is
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), while we conclude that it is the intraspecific variation. Since
<bibRefCitation id="9B1B75A3E9134A5DFE72FD4BFD5EFD22" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[473,672,655,681]" pageId="17" pageNumber="52" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang
<emphasis id="CDFED440E9134A5DFDB0FD4BFDAFFD22" box="[539,593,655,681]" italics="true" pageId="17" pageNumber="52">et al.</emphasis>
(2020)
</bibRefCitation>
recorded the males of this species with no morphological description, this is the first formal description of the male.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="FF350852E90E4A40FF3CFF53FE0FFF3A" blockId="12.[151,530,151,214]" box="[151,497,151,178]" pageId="12" pageNumber="47">
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<emphasis id="CDFED440E90E4A40FF3CFF53FE72FF39" bold="true" box="[151,396,151,178]" italics="true" pageId="12" pageNumber="47">Syzeuctus flavitarsis</emphasis>
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<paragraph id="FF350852E90E4A40FF3CFF7FFDECFF5E" blockId="12.[151,530,151,214]" box="[151,530,187,214]" pageId="12" pageNumber="47">
(
<figureCitation id="67B114D7E90E4A40FF34FF7FFEFAFF5D" box="[159,260,187,214]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="12" pageNumber="47">Figs 1 D</figureCitation>
,
<figureCitation id="67B114D7E90E4A40FEA4FF7FFEA2FF5E" box="[271,348,187,214]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="12" pageNumber="47">2 D, O</figureCitation>
,
<figureCitation id="67B114D7E90E4A40FEC3FF7FFE4BFF5E" box="[360,437,187,213]" captionStart="FIGURE 8" captionStartId="13.[151,250,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@13.[151,1436,181,1821]" captionTargetPageId="13" captionText="FIGURE 8. Syzeuctus flavitarsis sp. nov., females, holotype (A, C, E–G) and paratype (B, D)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F: pronotum and mesopleuron, lateral view; G: fore wing; H: hind tibia and tarsus, lateral view." pageId="12" pageNumber="47">8 A–H</figureCitation>
,
<figureCitation id="67B114D7E90E4A40FE6BFF7FFDF4FF5E" box="[448,522,187,213]" captionStart="FIGURE 9" captionStartId="14.[151,250,860,884]" captionTargetBox="[151,1436,166,837]" captionTargetId="figure-19@14.[151,1436,166,837]" captionTargetPageId="14" captionText="FIGURE 9. Syzeuctus flavitarsis sp. nov., male, paratype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view." pageId="12" pageNumber="47">9 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90E4A40FF3CFEC7FB19FE5A" pageId="12" pageNumber="47" type="materials_examined">
<paragraph id="FF350852E90E4A40FF3CFEC7FB19FE5A" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">
<emphasis id="CDFED440E90E4A40FF3CFEC7FEE5FE96" bold="true" box="[151,283,259,285]" pageId="12" pageNumber="47">Type series</emphasis>
.
<materialsCitation id="4FE2020FE90E4A40FE83FEC7FDFFFECA" collectingDate="2010-08-11" collectingDateMax="2010-08-25" collectingDateMin="2010-08-11" collectionCode="KPMNH" collectorName="T. Nakayama" country="Japan" county="Honshu" location="Ohkawa-rindo" municipality="Nasushiobara City" pageId="12" pageNumber="47" specimenCount="1" stateProvince="Tochigi" typeStatus="holotype">
<emphasis id="CDFED440E90E4A40FE83FEC7FE6BFE96" bold="true" box="[296,405,259,285]" pageId="12" pageNumber="47">
<typeStatus id="2031B6F0E90E4A40FE83FEC7FE6BFE96" box="[296,405,259,285]" pageId="12" pageNumber="47" type="holotype">Holotype</typeStatus>
</emphasis>
: F,
<emphasis id="CDFED440E90E4A40FE17FEC7FDE8FE96" bold="true" box="[444,534,259,285]" pageId="12" pageNumber="47">
<collectingCountry id="879D48C2E90E4A40FE17FEC7FDE8FE96" box="[444,534,259,285]" name="Japan" pageId="12" pageNumber="47">JAPAN</collectingCountry>
</emphasis>
,
<emphasis id="CDFED440E90E4A40FD88FEC7FD7EFE96" bold="true" box="[547,640,259,285]" pageId="12" pageNumber="47">
<collectingCounty id="165470DEE90E4A40FD88FEC7FD7EFE96" box="[547,640,259,285]" pageId="12" pageNumber="47">Honshu</collectingCounty>
</emphasis>
,
<collectingRegion id="3D4EC6B0E90E4A40FD26FEC7FCDFFE96" box="[653,801,259,285]" country="Japan" name="Tochigi" pageId="12" pageNumber="47">Tochigi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90E4A40FC80FEC7FC04FE96" box="[811,1018,259,285]" pageId="12" pageNumber="47">Nasushiobara City</collectingMunicipality>
,
<location id="FA555E89E90E4A40FBAFFEC7FB57FE96" LSID="urn:lsid:plazi:treatment:7723B944E90E4A42FF3CFF53FB6AFB56:FA555E89E90E4A40FBAFFEC7FB57FE96" box="[1028,1193,259,285]" country="Japan" county="Honshu" municipality="Nasushiobara City" name="Ohkawa-rindo" pageId="12" pageNumber="47" stateProvince="Tochigi">Ohkawa-rindo</location>
,
<date id="8B342E92E90E4A40FB18FEC7FA83FE96" box="[1203,1405,259,286]" pageId="12" pageNumber="47" value="2010-08-11" valueMax="2010-08-25" valueMin="2010-08-11">
<collectingDate id="9B70D77AE90E4A40FB18FEC7FA83FE96" box="[1203,1405,259,286]" pageId="12" pageNumber="47" value="2010-08-11" valueMax="2010-08-25" valueMin="2010-08-11">11–25. VIII. 2010</collectingDate>
</date>
,
<collectorName id="527F6D84E90E4A40FA2DFEC0FEF1FECA" pageId="12" pageNumber="47">T. Nakayama</collectorName>
leg., MsT (
<collectionCode id="999B9097E90E4A40FE26FEE3FE07FECA" box="[397,505,295,321]" pageId="12" pageNumber="47">KPMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE90E4A40FDA2FEE3FD31FEEE" collectingDate="2012-08-21" collectionCode="ZSM" collectorName="Y. Matsubara &amp; K. Fukuda" country="Japan" location="Mamurogawa Town" pageId="12" pageNumber="47" specimenCount="1" stateProvince="Yamagata" typeStatus="paratype">
<emphasis id="CDFED440E90E4A40FDA2FEE3FD7FFECA" bold="true" box="[521,641,295,321]" pageId="12" pageNumber="47">
<typeStatus id="2031B6F0E90E4A40FDA2FEE3FD7FFECA" box="[521,641,295,321]" pageId="12" pageNumber="47" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="CDFED440E90E4A40FD24FEE3FD17FECA" bold="true" box="[655,745,295,321]" pageId="12" pageNumber="47">
<collectingCountry id="879D48C2E90E4A40FD24FEE3FD17FECA" box="[655,745,295,321]" name="Japan" pageId="12" pageNumber="47">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E90E4A40FD54FEE3FCA2FECA" bold="true" box="[767,860,295,321]" pageId="12" pageNumber="47">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E90E4A40FC32FEECFBB8FECA" box="[921,1094,295,322]" country="Japan" name="Yamagata" pageId="12" pageNumber="47">Yamagata Pref.</collectingRegion>
,
<location id="FA555E89E90E4A40FBFBFEECFACDFEC9" LSID="urn:lsid:plazi:treatment:7723B944E90E4A42FF3CFF53FB6AFB56:FA555E89E90E4A40FBFBFEECFACDFEC9" box="[1104,1331,296,322]" country="Japan" name="Mamurogawa Town" pageId="12" pageNumber="47" stateProvince="Yamagata">Mamurogawa Town</location>
,
<date id="8B342E92E90E4A40FA96FEE3FF2CFEEE" pageId="12" pageNumber="47" value="2012-08-21">
<collectingDate id="9B70D77AE90E4A40FA96FEE3FF2CFEEE" pageId="12" pageNumber="47" value="2012-08-21">21. VIII. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E90E4A40FF77FE88FE8EFEEE" box="[220,368,331,358]" pageId="12" pageNumber="47">Y. Matsubara</collectorName>
&amp;
<collectorName id="527F6D84E90E4A40FE39FE88FDF6FEEE" box="[402,520,331,358]" pageId="12" pageNumber="47">K. Fukuda</collectorName>
leg., MsT (
<collectionCode id="999B9097E90E4A40FD23FE8FFD39FEEE" box="[648,711,331,357]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="47" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90E4A40FD71FE8FFEBCFE02" collectingDate="2011-06-29" collectionCode="KPMNH" collectorName="M. Ito" country="Japan" location="Biwako-Ballay" municipality="Otsu City" pageId="12" pageNumber="47" specimenCount="1" stateProvince="Shiga" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E90E4A40FCA2FE8FFC76FEEE" box="[777,904,331,357]" country="Japan" name="Shiga" pageId="12" pageNumber="47">Shiga Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90E4A40FC39FE8FFC01FEEE" box="[914,1023,331,357]" pageId="12" pageNumber="47">Otsu City</collectingMunicipality>
,
<location id="FA555E89E90E4A40FBA2FE8FFB4AFEEE" LSID="urn:lsid:plazi:treatment:7723B944E90E4A42FF3CFF53FB6AFB56:FA555E89E90E4A40FBA2FE8FFB4AFEEE" box="[1033,1204,331,357]" country="Japan" municipality="Otsu City" name="Biwako-Ballay" pageId="12" pageNumber="47" stateProvince="Shiga">Biwako-Ballay</location>
,
<date id="8B342E92E90E4A40FB14FE8FFAB2FEEE" box="[1215,1356,331,358]" pageId="12" pageNumber="47" value="2011-06-29">
<collectingDate id="9B70D77AE90E4A40FB14FE8FFAB2FEEE" box="[1215,1356,331,358]" pageId="12" pageNumber="47" value="2011-06-29">29. VI. 2011</collectingDate>
</date>
,
<collectorName id="527F6D84E90E4A40FAFCFE88FA62FEED" box="[1367,1436,332,358]" pageId="12" pageNumber="47">M. Ito</collectorName>
leg. (
<collectionCode id="999B9097E90E4A40FF65FEABFEC4FE02" box="[206,314,367,393]" pageId="12" pageNumber="47">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90E4A40FEE0FEABFEFDFE26" collectingDate="2013-07-20" collectingDateMax="2013-08-11" collectingDateMin="2013-07-20" collectionCode="OMNH" collectorName="S. Fujie &amp; T. Hirooka" country="Japan" location="Mt. Shakagatake" municipality="Totsukawa Vil." pageId="12" pageNumber="47" specimenCount="1" stateProvince="Nara" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E90E4A40FED2FEB4FE10FE02" box="[377,494,367,394]" country="Japan" name="Nara" pageId="12" pageNumber="47">Nara Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90E4A40FE5CFEABFD5FFE02" box="[503,673,367,393]" pageId="12" pageNumber="47">Totsukawa Vil.</collectingMunicipality>
,
<location id="FA555E89E90E4A40FD00FEB4FC96FE02" LSID="urn:lsid:plazi:treatment:7723B944E90E4A42FF3CFF53FB6AFB56:FA555E89E90E4A40FD00FEB4FC96FE02" box="[683,872,367,394]" country="Japan" municipality="Totsukawa Vil." name="Mt. Shakagatake" pageId="12" pageNumber="47" stateProvince="Nara">Mt. Shakagatake</location>
,
<date id="8B342E92E90E4A40FCD9FEABFB88FE02" box="[882,1142,367,394]" pageId="12" pageNumber="47" value="2013-07-20" valueMax="2013-08-11" valueMin="2013-07-20">
<collectingDate id="9B70D77AE90E4A40FCD9FEABFB88FE02" box="[882,1142,367,394]" pageId="12" pageNumber="47" value="2013-07-20" valueMax="2013-08-11" valueMin="2013-07-20">20. VII.–11. VIII. 2013</collectingDate>
</date>
,
<collectorName id="527F6D84E90E4A40FBD4FEABFB28FE02" box="[1151,1238,367,393]" pageId="12" pageNumber="47">S. Fujie</collectorName>
&amp;
<collectorName id="527F6D84E90E4A40FB5DFEB4FA90FE02" box="[1270,1390,367,394]" pageId="12" pageNumber="47">T. Hirooka</collectorName>
leg. (
<collectionCode id="999B9097E90E4A40FF34FE57FF05FE26" box="[159,251,403,429]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="12" pageNumber="47" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90E4A40FEBBFE57FA66FE26" box="[272,1432,403,430]" collectingDate="2011-08-06" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Mt. Hyonosen" municipality="Wakasa Town" pageId="12" pageNumber="47" specimenCount="1" stateProvince="Tottori" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E90E4A40FEE9FE57FE31FE26" box="[322,463,403,429]" country="Japan" name="Tottori" pageId="12" pageNumber="47">Tottori Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90E4A40FE70FE57FD85FE25" box="[475,635,403,430]" pageId="12" pageNumber="47">Wakasa Town</collectingMunicipality>
,
<location id="FA555E89E90E4A40FD23FE50FCD2FE25" LSID="urn:lsid:plazi:treatment:7723B944E90E4A42FF3CFF53FB6AFB56:FA555E89E90E4A40FD23FE50FCD2FE25" box="[648,812,404,430]" country="Japan" municipality="Wakasa Town" name="Mt. Hyonosen" pageId="12" pageNumber="47" stateProvince="Tottori">Mt. Hyonosen</location>
,
<date id="8B342E92E90E4A40FC92FE57FC31FE26" box="[825,975,403,430]" pageId="12" pageNumber="47" value="2011-08-06">
<collectingDate id="9B70D77AE90E4A40FC92FE57FC31FE26" box="[825,975,403,430]" pageId="12" pageNumber="47" value="2011-08-06">6. VIII. 2011</collectingDate>
</date>
,
<collectorName id="527F6D84E90E4A40FC70FE50FB93FE26" box="[987,1133,403,430]" pageId="12" pageNumber="47">K. Watanabe</collectorName>
leg.,
<collectingMethod id="26CB7045E90E4A40FB06FE57FAEBFE25" box="[1197,1301,403,430]" pageId="12" pageNumber="47">light trap</collectingMethod>
(
<collectionCode id="999B9097E90E4A40FA8FFE57FA6EFE26" box="[1316,1424,403,429]" pageId="12" pageNumber="47">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE90E4A40FF3CFE7CFB1DFE5A" box="[151,1251,439,466]" collectingDate="2012-06-30" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Mt. Kasatori" municipality="Uchico Town" pageId="12" pageNumber="47" specimenCount="1" stateProvince="Ehime" typeStatus="paratype">
[
<emphasis id="CDFED440E90E4A40FF0AFE73FEFBFE5A" bold="true" box="[161,261,439,465]" pageId="12" pageNumber="47">Shikoku</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E90E4A40FEFBFE73FE27FE5A" box="[336,473,439,465]" country="Japan" name="Ehime" pageId="12" pageNumber="47">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E90E4A40FE4FFE73FD83FE59" box="[484,637,439,466]" pageId="12" pageNumber="47">Uchico Town</collectingMunicipality>
,
<location id="FA555E89E90E4A40FD22FE7CFCE7FE5A" LSID="urn:lsid:plazi:treatment:7723B944E90E4A42FF3CFF53FB6AFB56:FA555E89E90E4A40FD22FE7CFCE7FE5A" box="[649,793,439,466]" country="Japan" municipality="Uchico Town" name="Mt. Kasatori" pageId="12" pageNumber="47" stateProvince="Ehime">Mt. Kasatori</location>
,
<date id="8B342E92E90E4A40FC8FFE73FC4AFE5A" box="[804,948,439,466]" pageId="12" pageNumber="47" value="2012-06-30">
<collectingDate id="9B70D77AE90E4A40FC8FFE73FC4AFE5A" box="[804,948,439,466]" pageId="12" pageNumber="47" value="2012-06-30">30. VI. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E90E4A40FC14FE7CFBA9FE59" box="[959,1111,440,466]" pageId="12" pageNumber="47">E. Yamamoto</collectorName>
leg. (
<collectionCode id="999B9097E90E4A40FB3DFE73FB24FE5A" box="[1174,1242,439,465]" pageId="12" pageNumber="47">EUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90E4A40FF6CFE1FFC99F856" pageId="12" pageNumber="47" type="description">
<paragraph id="FF350852E90E4A40FF6CFE1FFC3CFD91" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">
<emphasis id="CDFED440E90E4A40FF6CFE1FFEAFFE7E" bold="true" box="[199,337,475,501]" pageId="12" pageNumber="47">Description</emphasis>
.
<emphasis id="CDFED440E90E4A40FEF4FE1FFE48FE7E" bold="true" box="[351,438,475,501]" pageId="12" pageNumber="47">Female</emphasis>
(n = 5). Body polished and covered with punctures and silvery setae; length 8.1–9.2 (HT: 9.1) mm. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E90E4A40FF6CFDE7FB20FCB2" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">
Head 0.6 × as long as wide. Combined length of clypeus and face 1.0–1.05 (HT: 1.05) × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.5 × as long as basal mandibular width. Genal carina lamella-like ventrally, reaching mandibular base. POL 1.5–1.7 (HT: 1.7) × as long as OD. OOL 1.2–1.4 (HT: 1.2) × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E90E4A40FAF1FD13FF52FC9E" captionStart="FIGURE 8" captionStartId="13.[151,250,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@13.[151,1436,181,1821]" captionTargetPageId="13" captionText="FIGURE 8. Syzeuctus flavitarsis sp. nov., females, holotype (A, C, E–G) and paratype (B, D)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F: pronotum and mesopleuron, lateral view; G: fore wing; H: hind tibia and tarsus, lateral view." pageId="12" pageNumber="47">Fig. 8 E</figureCitation>
). Antenna with 42–45 (HT: 42) flagellomeres. FL I 4.3–4.6 (HT: 4.6) × as long as maximum depth in lateral view and 1.5–1.6 (HT: 1.6) × as long as FL II. Preapical flagellomere 1.1 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E90E4A40FF6CFC80FD91FB4E" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">
Mesosoma coarsely and densely punctate, 1.45 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate; speculum with a large smooth area (
<figureCitation id="67B114D7E90E4A40FD9EFC4FFD6AFC2E" box="[565,660,907,933]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="12" pageNumber="47">Fig. 2 F</figureCitation>
). Punctures on upper part of mesopleuron irregular and sparser than on lower part. Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E90E4A40FD95FC17FD5AFC66" box="[574,676,979,1005]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="12" pageNumber="47">Fig. 1 D</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum largely absent medially, obsolete laterally. Fore wing length 7.0–7.3 (HT: 7.0) mm. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E90E4A40FAC4FBDFFF3DFBD2" captionStart="FIGURE 8" captionStartId="13.[151,250,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@13.[151,1436,181,1821]" captionTargetPageId="13" captionText="FIGURE 8. Syzeuctus flavitarsis sp. nov., females, holotype (A, C, E–G) and paratype (B, D)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F: pronotum and mesopleuron, lateral view; G: fore wing; H: hind tibia and tarsus, lateral view." pageId="12" pageNumber="47">Fig. 8 G</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.5–6.0 (HT: 5.8) × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.2–4.3 (HT: 4.2): 2.0: 1.4: 0.8: 0.9. Hind tarsal claws simple except for minute subbasal teeth, strongly bent apically (ca. 90°) (
<figureCitation id="67B114D7E90E4A40FDAAFB6FFD9CFB4E" box="[513,610,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="12" pageNumber="47">Fig. 2 D</figureCitation>
).
</paragraph>
<paragraph id="FF350852E90E4A40FF6CFB14FF1BFADE" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">Metasoma. T I 2.2–2.4 (HT: 2.4) × as long as maximum width, finely punctate laterally, smooth medially. T II 1.25–1.5 (HT: 1.5) × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine and dense punctures. Ovipositor sheath 2.0 × as long as hind tibia, ca. 0.8 × as long as fore wing, and shorter than body length.</paragraph>
<paragraph id="FF350852E90E4A40FF6CFA9BFC2CF8F9" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">
Colouration (
<figureCitation id="67B114D7E90E4A40FEF6FA9BFE18FAF2" box="[349,486,1375,1401]" captionStart="FIGURE 8" captionStartId="13.[151,250,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@13.[151,1436,181,1821]" captionTargetPageId="13" captionText="FIGURE 8. Syzeuctus flavitarsis sp. nov., females, holotype (A, C, E–G) and paratype (B, D)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F: pronotum and mesopleuron, lateral view; G: fore wing; H: hind tibia and tarsus, lateral view." pageId="12" pageNumber="47">Figs 8 A–H</figureCitation>
). Body (excluding wings and legs) black. Clypeus, mandible except for teeth, malar space, a pair of small yellow spots below antennal sockets (sometimes absent as
<figureCitation id="67B114D7E90E4A40FB9DFA47FB67FA16" box="[1078,1177,1411,1437]" captionStart="FIGURE 8" captionStartId="13.[151,250,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@13.[151,1436,181,1821]" captionTargetPageId="13" captionText="FIGURE 8. Syzeuctus flavitarsis sp. nov., females, holotype (A, C, E–G) and paratype (B, D)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F: pronotum and mesopleuron, lateral view; G: fore wing; H: hind tibia and tarsus, lateral view." pageId="12" pageNumber="47">Fig. 8 D</figureCitation>
), orbits on face, frons, vertex and gena, and palpi yellow. Antenna blackish-brown to reddish-brown. Scape and pedicel usually partly tinged with yellow ventrally. Two spots of collar, a L-shape marking of dorsal margin of pronotum, a pair of antero-lateral markings and a median marking of mesoscutum, subtegular ridge, antero-dorsal and postero-ventral spots of mesopleuron, scutellum except for a median black spot, postscutellum, posterior spot of metapleuron, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of propodeum yellow. Scutellum sometimes entirely yellow. Metasomal tergites black except for a basal band of T I and posterior margins of T I to T V yellow. Membranous parts of metasomal sternites largely yellow. Subgenital plate blackish-brown, usually partly tinged with yellowish-brown. Ovipositor reddish-brown. Wings hyaline. Apical clouded area of fore wing slightly to weakly present. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow; mid coxa partly tinged with blackish-brown basally. Hind coxa black except for a dorsal spot and apex yellow to yellowish-brown. Hind trochanter black to blackish-brown. Hind trochantellus yellow. Hind femur reddish-brown except for apex tinged with yellow. Hind tibia and tibial spurs ivory except for apical part of tibia blackish-brown. Hind tarsus ivory to yellowish white part except for black fifth tarsomere.
</paragraph>
<paragraph id="FF350852E90E4A40FF6CF8BFFC99F856" blockId="12.[151,1437,259,2014]" pageId="12" pageNumber="47">
<emphasis id="CDFED440E90E4A40FF6CF8BFFEFAF81E" bold="true" box="[199,260,1915,1941]" pageId="12" pageNumber="47">Male</emphasis>
(n = 1). Similar to female (
<figureCitation id="67B114D7E90E4A40FD83F8BFFD54F81E" box="[552,682,1915,1941]" captionStart="FIGURE 9" captionStartId="14.[151,250,860,884]" captionTargetBox="[151,1436,166,837]" captionTargetId="figure-19@14.[151,1436,166,837]" captionTargetPageId="14" captionText="FIGURE 9. Syzeuctus flavitarsis sp. nov., male, paratype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view." pageId="12" pageNumber="47">Figs 9 A–C</figureCitation>
). Body length
<quantity id="3872A5B7E90E4A40FCECF8BFFC60F81D" box="[839,926,1915,1942]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="12" pageNumber="47" unit="mm" value="7.8">7.8 mm</quantity>
. Malar space 0.8 × as long as basal mandibular width. Tergal spiracles not enlarged (
<figureCitation id="67B114D7E90E4A40FD97F85BFD5EF832" box="[572,672,1951,1977]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="12" pageNumber="47">Fig. 2 O</figureCitation>
). Ratio of length of hind first and second tarsomeres 4.0: 2.0. Face entirely yellow. Bases of T II to T IV each with yellow marking.
</paragraph>
</subSubSection>
<caption id="ABF558DAE90F4A41FF3CF8F1FD24F81E" pageId="13" pageNumber="48" startId="13.[151,250,1845,1869]" targetBox="[151,1436,181,1821]" targetPageId="13" targetType="figure">
<paragraph id="FF350852E90F4A41FF3CF8F1FD24F81E" blockId="13.[151,1436,1845,1941]" pageId="13" pageNumber="48">
<emphasis id="CDFED440E90F4A41FF3CF8F1FEEBF8C6" bold="true" box="[151,277,1845,1869]" pageId="13" pageNumber="48">FIGURE 8.</emphasis>
<taxonomicName id="388A73D1E90F4A41FEB6F8F1FE13F8C6" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[285,493,1845,1869]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="flavitarsis" status="sp. nov.">
<emphasis id="CDFED440E90F4A41FEB6F8F1FE13F8C6" box="[285,493,1845,1869]" italics="true" pageId="13" pageNumber="48">Syzeuctus flavitarsis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E90F4A41FE5EF8F1FDB6F8C6" bold="true" box="[501,584,1845,1869]" pageId="13" pageNumber="48">
<taxonomicNameLabel id="D6CD693BE90F4A41FE5EF8F1FDB6F8C6" box="[501,584,1845,1869]" pageId="13" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, females, holotype (A, C, E–G) and paratype (B, D)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F: pronotum and mesopleuron, lateral view; G: fore wing; H: hind tibia and tarsus, lateral view.
</paragraph>
</caption>
<caption id="ABF558DAE90C4A42FF3CFC98FE8DFC12" pageId="14" pageNumber="49" startId="14.[151,250,860,884]" targetBox="[151,1436,166,837]" targetPageId="14" targetType="figure">
<paragraph id="FF350852E90C4A42FF3CFC98FE8DFC12" blockId="14.[151,1436,860,921]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF3CFC98FEECFCFF" bold="true" box="[151,274,860,884]" pageId="14" pageNumber="49">FIGURE 9.</emphasis>
<taxonomicName id="388A73D1E90C4A42FEB2FC99FE18FCFE" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[281,486,861,885]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="flavitarsis" status="sp. nov.">
<emphasis id="CDFED440E90C4A42FEB2FC99FE18FCFE" box="[281,486,861,885]" italics="true" pageId="14" pageNumber="49">Syzeuctus flavitarsis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E90C4A42FE47FC99FDC2FCFE" bold="true" box="[492,572,861,885]" pageId="14" pageNumber="49">
<taxonomicNameLabel id="D6CD693BE90C4A42FE47FC99FDC2FCFE" box="[492,572,861,885]" pageId="14" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, paratype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view.
</paragraph>
</caption>
<subSubSection id="B7905BD9E90C4A42FF6CFC03FD4DFC6A" box="[199,691,967,993]" pageId="14" pageNumber="49" type="distribution">
<paragraph id="FF350852E90C4A42FF6CFC03FD4DFC6A" blockId="14.[151,1437,967,1245]" box="[199,691,967,993]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF6CFC03FEA7FC6A" bold="true" box="[199,345,967,993]" pageId="14" pageNumber="49">Distribution</emphasis>
.
<collectingCountry id="879D48C2E90C4A42FECCFC03FE59FC6A" box="[359,423,967,993]" name="Japan" pageId="14" pageNumber="49">Japan</collectingCountry>
(Honshu and Shikoku).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90C4A42FF6CFC2FFE36FBA2" pageId="14" pageNumber="49" type="etymology">
<paragraph id="FF350852E90C4A42FF6CFC2FFAD4FB8E" blockId="14.[151,1437,967,1245]" box="[199,1322,1003,1029]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF6CFC2FFEB6FB8E" bold="true" box="[199,328,1003,1029]" pageId="14" pageNumber="49">Etymology</emphasis>
. The specific refers to the yellowish (flavi-) coloration of first to fourth hind tarsomeres.
</paragraph>
<paragraph id="FF350852E90C4A42FF6CFBCBFE36FBA2" blockId="14.[151,1437,967,1245]" box="[199,456,1039,1065]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF6CFBCBFEBDFBA2" bold="true" box="[199,323,1039,1065]" pageId="14" pageNumber="49">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E90C4A42FF6CFBF7FB6AFB56" pageId="14" pageNumber="49" type="discussion">
<paragraph id="FF350852E90C4A42FF6CFBF7FB6AFB56" blockId="14.[151,1437,967,1245]" pageId="14" pageNumber="49">
<emphasis id="CDFED440E90C4A42FF6CFBF7FECAFBC6" bold="true" box="[199,308,1075,1101]" pageId="14" pageNumber="49">Remarks</emphasis>
. In the key provided by
<bibRefCitation id="9B1B75A3E90C4A42FDE0FBF7FD16FBC6" author="Riedel, M." box="[587,744,1075,1101]" pageId="14" pageNumber="49" pagination="1013 - 1098" refId="ref23284" refString="Riedel, M. (2021) The genus Syzeuctus Forster in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species. Linzer biologische Beitrage, 52 (2), 1013 - 1098." type="journal article" year="2021">Riedel (2021)</bibRefCitation>
, the female runs to couplet 37 (
<taxonomicName id="388A73D1E90C4A42FBE4FBF7FAE9FBC6" box="[1103,1303,1075,1101]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="flavomaculatus">
<emphasis id="CDFED440E90C4A42FBE4FBF7FAE9FBC6" box="[1103,1303,1075,1101]" italics="true" pageId="14" pageNumber="49">S. flavomaculatus</emphasis>
</taxonomicName>
) but differs by the T I 2.2–2.4 × as long as maximum width (1.95 × in
<taxonomicName id="388A73D1E90C4A42FC8EFB93FC12FBFA" box="[805,1004,1111,1137]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="flavomaculatus">
<emphasis id="CDFED440E90C4A42FC8EFB93FC12FBFA" box="[805,1004,1111,1137]" italics="true" pageId="14" pageNumber="49">S. flavomaculatus</emphasis>
</taxonomicName>
), the ovipositor shorter than fore wing (longer than fore wing in
<taxonomicName id="388A73D1E90C4A42FE6DFBBFFD6EFB1E" box="[454,656,1147,1173]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="flavomaculatus">
<emphasis id="CDFED440E90C4A42FE6DFBBFFD6EFB1E" box="[454,656,1147,1173]" italics="true" pageId="14" pageNumber="49">S. flavomaculatus</emphasis>
</taxonomicName>
), and hind tarsus ivory except fifth tarsomere (reddish-brown in
<taxonomicName id="388A73D1E90C4A42FA2CFBBFFEBCFB32" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="flavomaculatus">
<emphasis id="CDFED440E90C4A42FA2CFBBFFEBCFB32" italics="true" pageId="14" pageNumber="49">S. flavomaculatus</emphasis>
</taxonomicName>
). This species closely resembles
<taxonomicName id="388A73D1E90C4A42FD11FB5BFCB7FB32" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[698,841,1183,1209]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E90C4A42FD11FB5BFCB7FB32" box="[698,841,1183,1209]" italics="true" pageId="14" pageNumber="49">S. yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E90C4A42FCFAFB5BFC55FB32" bold="true" box="[849,939,1183,1209]" pageId="14" pageNumber="49">
<taxonomicNameLabel id="D6CD693BE90C4A42FCFAFB5BFC55FB32" box="[849,939,1183,1209]" pageId="14" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but can be distinguished by the punctuation of mesopleuron and the colour patterns of hind leg and metasomal tergites (see key above).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:url id="A0886947AA2259AD6D69613776489A58">http://dx.doi.org/10.11646/zootaxa.5496.1.2</mods:url>
</mods:location>
<mods:classification id="6FE7A83133AA727843A13F1DB589182F">journal article</mods:classification>
<mods:identifier id="311562B168FDF4558905EDADCE37C614" type="DOI">10.11646/zootaxa.5496.1.2</mods:identifier>
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</mods:mods>
<treatment id="7723B944E9114A5AFF3CFD77FAF2FCD6" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6" httpUri="http://treatment.plazi.org/id/7723B944E9114A5AFF3CFD77FAF2FCD6" lastPageId="22" lastPageNumber="57" pageId="19" pageNumber="54">
<subSubSection id="B7905BD9E9114A5FFF3CFD77FDCFFD47" box="[151,561,690,717]" pageId="19" pageNumber="54" type="nomenclature">
<paragraph id="FF350852E9114A5FFF3CFD77FDCFFD47" blockId="19.[151,561,690,753]" box="[151,561,690,717]" pageId="19" pageNumber="54">
<heading id="A47DBF3EE9114A5FFF3CFD77FDCFFD47" bold="true" box="[151,561,690,717]" fontSize="11" level="1" pageId="19" pageNumber="54" reason="1">
<taxonomicName id="388A73D1E9114A5FFF3CFD77FDCFFD47" authority="Sheng, 2009" authorityName="Sheng" authorityYear="2009" box="[151,561,690,717]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="54" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="CDFED440E9114A5FFF3CFD77FDCFFD47" bold="true" box="[151,561,690,717]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF3CFD77FE6DFD46" bold="true" box="[151,403,690,717]" italics="true" pageId="19" pageNumber="54">Syzeuctus maculatus</emphasis>
<bibRefCitation id="9B1B75A3E9114A5FFE31FD76FDCFFD47" author="Sheng, M-L. &amp; Sun, S." box="[410,561,690,716]" pageId="19" pageNumber="54" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">Sheng, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9114A5FFF3CFD13FE04FD7A" box="[151,506,727,753]" pageId="19" pageNumber="54" type="description">
<paragraph id="FF350852E9114A5FFF3CFD13FE04FD7A" blockId="19.[151,561,690,753]" box="[151,506,727,753]" pageId="19" pageNumber="54">
(
<figureCitation id="67B114D7E9114A5FFF34FD13FEFAFD7A" box="[159,260,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="19" pageNumber="54">Figs 1 G</figureCitation>
,
<figureCitation id="67B114D7E9114A5FFEA4FD13FED1FD7A" box="[271,303,727,753]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="19" pageNumber="54">2 I</figureCitation>
,
<figureCitation id="67B114D7E9114A5FFE90FD13FE6EFD7A" box="[315,400,727,753]" captionStart="FIGURE 13" captionStartId="20.[151,250,1828,1852]" captionTargetBox="[151,1436,199,1805]" captionTargetId="figure-19@20.[151,1436,199,1805]" captionTargetPageId="20" captionText="FIGURE 13. Syzeuctus maculatus Sheng, 2009, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="19" pageNumber="54">13 A–F</figureCitation>
,
<figureCitation id="67B114D7E9114A5FFE31FD13FE0CFD7A" box="[410,498,727,753]" captionStart="FIGURE 14" captionStartId="21.[151,250,1635,1659]" captionTargetBox="[151,1436,181,1611]" captionTargetId="figure-19@21.[151,1436,181,1611]" captionTargetPageId="21" captionText="FIGURE 14. Syzeuctus maculatus Sheng, 2009, male from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view." pageId="19" pageNumber="54">14 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9114A5FFF3CFCDFFCF2FCB8" box="[151,780,795,819]" pageId="19" pageNumber="54" type="reference_group">
<paragraph id="FF350852E9114A5FFF3CFCDFFCF2FCB8" blockId="19.[151,780,795,819]" box="[151,780,795,819]" pageId="19" pageNumber="54">
<treatmentCitationGroup id="DF9A2F7CE9114A5FFF3CFCDFFCF2FCB8" box="[151,780,795,819]" pageId="19" pageNumber="54">
<taxonomicName id="388A73D1E9114A5FFF3CFCDFFE13FCB8" authority="Sheng, 2009" authorityName="Sheng" authorityYear="2009" box="[151,493,795,819]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="54" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="CDFED440E9114A5FFF3CFCDFFE96FCB8" box="[151,360,795,819]" italics="true" pageId="19" pageNumber="54">Syzeuctus maculatus</emphasis>
<bibRefCitation id="9B1B75A3E9114A5FFEC5FCDFFE13FCB8" author="Sheng, M-L. &amp; Sun, S." box="[366,493,795,819]" pageId="19" pageNumber="54" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">
Sheng,
<quantity id="3872A5B7E9114A5FFE11FCDFFE13FCB8" box="[442,493,795,819]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="19" pageNumber="54" unit="in" value="2009.0">2009</quantity>
</bibRefCitation>
</taxonomicName>
in
<treatmentCitation id="7E2B2E43E9114A5FFDA5FCDFFCF6FCB8" author="Sheng, M-L. &amp; Sun, S." box="[526,776,795,819]" page="42" pageId="19" pageNumber="54" year="2009">
<bibRefCitation id="9B1B75A3E9114A5FFDA5FCDFFD1CFCB8" author="Sheng, M-L. &amp; Sun, S." box="[526,738,795,819]" pageId="19" pageNumber="54" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">Sheng &amp; Sun (2009)</bibRefCitation>
: 42
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9114A5FFF3CFCA5FA95FA8C" pageId="19" pageNumber="54" type="materials_examined">
<paragraph id="FF350852E9114A5FFF3CFCA5FA95FA8C" blockId="19.[151,1437,865,2007]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF3CFCA5FE76FCF0" bold="true" box="[151,392,865,891]" pageId="19" pageNumber="54">Materials examined</emphasis>
.
<materialsCitation id="4FE2020FE9114A5FFE31FCA5FE39FC14" collectingDate="1999-07-15" collectionCode="MU" collectorName="T. Naito" country="Japan" location="Higashiyama" municipality="Nagoya City" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Aichi">
<emphasis id="CDFED440E9114A5FFE31FCA5FE0AFCF0" bold="true" box="[410,500,865,891]" pageId="19" pageNumber="54">
<collectingCountry id="879D48C2E9114A5FFE31FCA5FE0AFCF0" box="[410,500,865,891]" name="Japan" pageId="19" pageNumber="54">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E9114A5FFDA4FCA5FD92FCF0" bold="true" box="[527,620,865,891]" pageId="19" pageNumber="54">Honshu</emphasis>
]1 M,
<collectingRegion id="3D4EC6B0E9114A5FFD1CFCA5FCC7FCF0" box="[695,825,865,891]" country="Japan" name="Aichi" pageId="19" pageNumber="54">Aichi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9114A5FFCECFCA6FC24FCF0" box="[839,986,865,892]" pageId="19" pageNumber="54">Nagoya City</collectingMunicipality>
,
<location id="FA555E89E9114A5FFC42FCA5FB7EFCF0" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFC42FCA5FB7EFCF0" box="[1001,1152,865,891]" country="Japan" municipality="Nagoya City" name="Higashiyama" pageId="19" pageNumber="54" stateProvince="Aichi">Higashiyama</location>
,
<date id="8B342E92E9114A5FFB24FCA5FAD1FCF0" box="[1167,1327,865,892]" pageId="19" pageNumber="54" value="1999-07-15">
<collectingDate id="9B70D77AE9114A5FFB24FCA5FAD1FCF0" box="[1167,1327,865,892]" pageId="19" pageNumber="54" value="1999-07-15">15. VII. 1999</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFA96FCA6FA62FCF0" box="[1341,1436,865,892]" pageId="19" pageNumber="54">T. Naito</collectorName>
leg.,
<collectingMethod id="26CB7045E9114A5FFF65FC41FE83FC2B" box="[206,381,901,928]" pageId="19" pageNumber="54">yellow pan trap</collectingMethod>
(
<collectionCode id="999B9097E9114A5FFE27FC41FE41FC14" box="[396,447,901,927]" country="USA" name="Midwestern University" pageId="19" pageNumber="54">MU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFE78FC42FE31FC48" collectingDate="2012-08-17" collectingDateMax="2012-10-05" collectingDateMin="2012-08-17" collectionCode="KPMNH" collectorName="Izuoshima Is. &amp; K. Tsujii" country="Japan" location="Mt. Omaru" municipality="Motomachi" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
[
<emphasis id="CDFED440E9114A5FFE77FC41FDC7FC14" bold="true" box="[476,569,901,927]" pageId="19" pageNumber="54">Izu Isls.</emphasis>
] 4 F,
<collectingRegion id="3D4EC6B0E9114A5FFDD1FC41FD3AFC14" box="[634,708,901,927]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFD64FC41FC99FC2B" box="[719,871,901,928]" pageId="19" pageNumber="54">Izuoshima Is.</collectorName>
,
<collectingMunicipality id="1F519228E9114A5FFCDFFC41FC06FC14" box="[884,1016,901,927]" pageId="19" pageNumber="54">Motomachi</collectingMunicipality>
,
<location id="FA555E89E9114A5FFBA8FC42FB7DFC14" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFBA8FC42FB7DFC14" box="[1027,1155,901,928]" country="Japan" municipality="Motomachi" name="Mt. Omaru" pageId="19" pageNumber="54" stateProvince="Tokyo">Mt. Omaru</location>
,
<date id="8B342E92E9114A5FFB25FC41FA88FC14" box="[1166,1398,901,928]" pageId="19" pageNumber="54" value="2012-08-17" valueMax="2012-10-05" valueMin="2012-08-17">
<collectingDate id="9B70D77AE9114A5FFB25FC41FA88FC14" box="[1166,1398,901,928]" pageId="19" pageNumber="54" value="2012-08-17" valueMax="2012-10-05" valueMin="2012-08-17">17. VIII.–5. X. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFA2AFC42FF28FC48" pageId="19" pageNumber="54">K. Tsujii</collectorName>
leg., MsT (
<collectionCode id="999B9097E9114A5FFEF0FC6DFE39FC48" box="[347,455,937,963]" pageId="19" pageNumber="54">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFE70FC6DFAFBFC48" box="[475,1285,937,964]" collectingDate="2007-07-01" collectionCode="KPMNH" collectorName="Shikinejima Is. &amp; S. Fukasawa" country="Japan" location="Tokyo" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
1 M,
<collectingRegion id="3D4EC6B0E9114A5FFDB3FC6DFD9CFC48" box="[536,610,937,963]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFDC5FC6DFCE4FC4F" box="[622,794,937,964]" pageId="19" pageNumber="54">Shikinejima Is.</collectorName>
,
<date id="8B342E92E9114A5FFC83FC6DFC4BFC48" box="[808,949,937,964]" pageId="19" pageNumber="54" value="2007-07-01">
<collectingDate id="9B70D77AE9114A5FFC83FC6DFC4BFC48" box="[808,949,937,964]" pageId="19" pageNumber="54" value="2007-07-01">1. VII. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFC6AFC6DFBAEFC48" box="[961,1104,937,963]" pageId="19" pageNumber="54">S. Fukasawa</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFB3AFC6DFB03FC48" box="[1169,1277,937,963]" pageId="19" pageNumber="54">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFABAFC6DFBB4FC6C" collectingDate="2003-07-06" collectionCode="OMNH" collectorName="Kozushima Is. &amp; O. Tominaga" country="Japan" location="Koube-yama" municipality="Kozuhondou" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
2 M,
<collectingRegion id="3D4EC6B0E9114A5FFAE5FC6DFA66FC48" box="[1358,1432,937,963]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFF3CFC09FEC6FC63" box="[151,312,973,1000]" pageId="19" pageNumber="54">Kozushima Is.</collectorName>
,
<collectingMunicipality id="1F519228E9114A5FFEE9FC09FE2DFC6C" box="[322,467,973,999]" pageId="19" pageNumber="54">Kozuhondou</collectingMunicipality>
to
<location id="FA555E89E9114A5FFE59FC09FD78FC6C" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFE59FC09FD78FC6C" box="[498,646,973,999]" country="Japan" municipality="Kozuhondou" name="Koube-yama" pageId="19" pageNumber="54" stateProvince="Tokyo">Koube-yama</location>
,
<date id="8B342E92E9114A5FFD25FC09FCEAFC6C" box="[654,788,973,1000]" pageId="19" pageNumber="54" value="2003-07-06">
<collectingDate id="9B70D77AE9114A5FFD25FC09FCEAFC6C" box="[654,788,973,1000]" pageId="19" pageNumber="54" value="2003-07-06">6. VII. 2003</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFCB6FC09FC52FC6C" box="[797,940,973,999]" pageId="19" pageNumber="54">O. Tominaga</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFC4DFC09FBBCFC6C" box="[998,1090,973,999]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="19" pageNumber="54" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFBF9FC09FC33FB80" collectingDate="2003-07-07" collectionCode="OMNH" collectorName="Kozushima Is. &amp; O. Tominaga" country="Japan" location="Maehama" municipality="Chichibuyama" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
1 F &amp; 1 M,
<collectingRegion id="3D4EC6B0E9114A5FFB61FC09FAEAFC6C" box="[1226,1300,973,999]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFAB6FC09FF4AFB87" pageId="19" pageNumber="54">Kozushima Is.</collectorName>
,
<collectingMunicipality id="1F519228E9114A5FFF6BFC35FE9DFB80" box="[192,355,1009,1035]" pageId="19" pageNumber="54">Chichibuyama</collectingMunicipality>
to
<location id="FA555E89E9114A5FFE2CFC35FE06FB80" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFE2CFC35FE06FB80" box="[391,504,1009,1035]" country="Japan" municipality="Chichibuyama" name="Maehama" pageId="19" pageNumber="54" stateProvince="Tokyo">Maehama</location>
,
<date id="8B342E92E9114A5FFDA8FC36FD70FB80" box="[515,654,1009,1036]" pageId="19" pageNumber="54" value="2003-07-07">
<collectingDate id="9B70D77AE9114A5FFDA8FC36FD70FB80" box="[515,654,1009,1036]" pageId="19" pageNumber="54" value="2003-07-07">7. VII. 2003</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFD32FC35FCD4FB80" box="[665,810,1009,1035]" pageId="19" pageNumber="54">O. Tominaga</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFCC2FC35FC3BFB80" box="[873,965,1009,1035]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="19" pageNumber="54" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFC73FC35FD41FBA4" collectingDate="2008-07-07" collectingDateMax="2008-07-30" collectingDateMin="2008-07-07" collectionCode="EUM" collectorName="Miyakejima Is. &amp; Ako &amp; H. Makihara" country="Japan" location="Tokyo" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
1 F &amp; 5 M,
<collectingRegion id="3D4EC6B0E9114A5FFBF1FC35FB5AFB80" box="[1114,1188,1009,1035]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFB04FC35FAA4FB87" box="[1199,1370,1009,1036]" pageId="19" pageNumber="54">Miyakejima Is.</collectorName>
,
<collectorName id="527F6D84E9114A5FFACEFC35FA66FB80" box="[1381,1432,1009,1035]" pageId="19" pageNumber="54">Ako</collectorName>
,
<date id="8B342E92E9114A5FFF3CFBD1FEAEFBA4" box="[151,336,1045,1072]" pageId="19" pageNumber="54" value="2008-07-07" valueMax="2008-07-30" valueMin="2008-07-07">
<collectingDate id="9B70D77AE9114A5FFF3CFBD1FEAEFBA4" box="[151,336,1045,1072]" pageId="19" pageNumber="54" value="2008-07-07" valueMax="2008-07-30" valueMin="2008-07-07">7–30. VII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFEF6FBD2FE15FBA4" box="[349,491,1045,1072]" pageId="19" pageNumber="54">H. Makihara</collectorName>
leg., MsT (
<collectionCode id="999B9097E9114A5FFDD9FBD1FD48FBA4" box="[626,694,1045,1071]" pageId="19" pageNumber="54">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFD67FBD1FDC6FBD8" collectingDate="2008-07-07" collectingDateMax="2008-07-30" collectingDateMin="2008-07-07" collectionCode="EUM" collectorName="Miyakejima Is. &amp; Hinoyama-toge &amp; H. Makihara" country="Japan" location="Tokyo" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
1 F &amp; 3 M,
<collectingRegion id="3D4EC6B0E9114A5FFCF3FBD1FC5CFBA4" box="[856,930,1045,1071]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFC05FBD1FBA5FBBB" box="[942,1115,1045,1072]" pageId="19" pageNumber="54">Miyakejima Is.</collectorName>
,
<collectorName id="527F6D84E9114A5FFBC2FBD1FAE4FBA4" box="[1129,1306,1045,1071]" pageId="19" pageNumber="54">Hinoyama-toge</collectorName>
,
<date id="8B342E92E9114A5FFA8CFBD1FF2CFBD8" pageId="19" pageNumber="54" value="2008-07-07" valueMax="2008-07-30" valueMin="2008-07-07">
<collectingDate id="9B70D77AE9114A5FFA8CFBD1FF2CFBD8" pageId="19" pageNumber="54" value="2008-07-07" valueMax="2008-07-30" valueMin="2008-07-07">7–30. VII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFF76FBFEFE94FBD8" box="[221,362,1081,1108]" pageId="19" pageNumber="54">H. Makihara</collectorName>
leg., MsT (
<collectionCode id="999B9097E9114A5FFE40FBFDFDD1FBD8" box="[491,559,1081,1107]" pageId="19" pageNumber="54">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFDE8FBFDFEAEFBFC" collectingDate="2008-07-30" collectionCode="KPMNH" collectorName="Miyakejima Is. &amp; Tsubota-rindo &amp; H. Makihara" country="Japan" location="Tokyo" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Tokyo">
5 M,
<collectingRegion id="3D4EC6B0E9114A5FFDD5FBFDFD36FBD8" box="[638,712,1081,1107]" country="Japan" name="Tokyo" pageId="19" pageNumber="54">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFD78FBFDFC80FBDF" box="[723,894,1081,1108]" pageId="19" pageNumber="54">Miyakejima Is.</collectorName>
,
<collectorName id="527F6D84E9114A5FFC21FBFDFBD4FBD8" box="[906,1066,1081,1107]" pageId="19" pageNumber="54">Tsubota-rindo</collectorName>
,
<date id="8B342E92E9114A5FFB9FFBFDFB33FBD8" box="[1076,1229,1081,1108]" pageId="19" pageNumber="54" value="2008-07-30">
<collectingDate id="9B70D77AE9114A5FFB9FFBFDFB33FBD8" box="[1076,1229,1081,1108]" pageId="19" pageNumber="54" value="2008-07-30">30. VII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFB73FBFEFA9BFBD8" box="[1240,1381,1081,1108]" pageId="19" pageNumber="54">H. Makihara</collectorName>
leg., MsT (
<collectionCode id="999B9097E9114A5FFF77FB99FEB6FBFC" box="[220,328,1117,1143]" pageId="19" pageNumber="54">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFEF6FB9AFEE5FB10" collectingDate="2001-08-16" collectionCode="EUM" collectorName="Y. Kawakami" country="Japan" location="Mt. Oshibayama" municipality="Tosashimizu City" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Kochi">
[
<emphasis id="CDFED440E9114A5FFECDFB99FE34FBFC" bold="true" box="[358,458,1117,1143]" pageId="19" pageNumber="54">Shikoku</emphasis>
] 2 M,
<collectingRegion id="3D4EC6B0E9114A5FFDB7FB99FD5DFBFC" box="[540,675,1117,1143]" country="Japan" name="Kochi" pageId="19" pageNumber="54">Kochi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9114A5FFD04FB99FC88FBFC" box="[687,886,1117,1143]" pageId="19" pageNumber="54">Tosashimizu City</collectingMunicipality>
,
<location id="FA555E89E9114A5FFC28FB9AFBBDFBFC" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFC28FB9AFBBDFBFC" box="[899,1091,1117,1144]" country="Japan" municipality="Tosashimizu City" name="Mt. Oshibayama" pageId="19" pageNumber="54" stateProvince="Kochi">Mt. Oshibayama</location>
,
<date id="8B342E92E9114A5FFBFBFB99FB08FBFC" box="[1104,1270,1117,1144]" pageId="19" pageNumber="54" value="2001-08-16">
<collectingDate id="9B70D77AE9114A5FFBFBFB99FB08FBFC" box="[1104,1270,1117,1144]" pageId="19" pageNumber="54" value="2001-08-16">16. VIII. 2001</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFAA9FB9AFA62FBFC" box="[1282,1436,1117,1144]" pageId="19" pageNumber="54">Y. Kawakami</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFF64FB45FEEDFB10" box="[207,275,1153,1179]" pageId="19" pageNumber="54">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFE8EFB46FAF5FB10" box="[293,1291,1153,1180]" collectingDate="1959-07-18" collectionCode="EUM" collectorName="Oomishima Is. &amp; T. Ishihara" country="Japan" location="Oomishima Is." pageId="19" pageNumber="54" specimenCount="1" stateProvince="Ehime">
[
<emphasis id="CDFED440E9114A5FFE84FB45FE22FB10" bold="true" box="[303,476,1153,1179]" pageId="19" pageNumber="54">
<location id="FA555E89E9114A5FFE84FB45FE22FB10" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFE84FB45FE22FB10" box="[303,476,1153,1179]" country="Japan" name="Oomishima Is." pageId="19" pageNumber="54" stateProvince="Ehime">Oomishima Is.</location>
</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9114A5FFD8DFB45FD50FB10" box="[550,686,1153,1179]" country="Japan" name="Ehime" pageId="19" pageNumber="54">Ehime Pref.</collectingRegion>
,
<collectorName id="527F6D84E9114A5FFD13FB45FCA0FB17" box="[696,862,1153,1180]" pageId="19" pageNumber="54">Oomishima Is.</collectorName>
,
<date id="8B342E92E9114A5FFCC2FB45FBFFFB17" box="[873,1025,1153,1180]" pageId="19" pageNumber="54" value="1959-07-18">
<collectingDate id="9B70D77AE9114A5FFCC2FB45FBFFFB17" box="[873,1025,1153,1180]" pageId="19" pageNumber="54" value="1959-07-18">18. VII. 1959</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFBA0FB46FB7FFB10" box="[1035,1153,1153,1180]" pageId="19" pageNumber="54">T. Ishihara</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFB14FB45FAFDFB10" box="[1215,1283,1153,1179]" pageId="19" pageNumber="54">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFABDFB46FAFDFB34" collectingDate="2021-07-17" collectionCode="KPMNH" collectorName="Iki Is. &amp; K. Otsui" country="Japan" county="Iki City" location="Hakozakihonmurafure" municipality="Ashibe Town" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Nagasaki">
[
<emphasis id="CDFED440E9114A5FFAB4FB45FA9AFB10" bold="true" box="[1311,1380,1153,1179]" pageId="19" pageNumber="54">
<collectorName id="527F6D84E9114A5FFAB4FB45FA9AFB10" box="[1311,1380,1153,1179]" pageId="19" pageNumber="54">Iki Is.</collectorName>
</emphasis>
] 2 F,
<collectingRegion id="3D4EC6B0E9114A5FFF3CFB61FEC0FB34" box="[151,318,1189,1215]" country="Japan" name="Nagasaki" pageId="19" pageNumber="54">Nagasaki Pref.</collectingRegion>
,
<collectingCounty id="165470DEE9114A5FFEE3FB61FE61FB34" box="[328,415,1189,1215]" pageId="19" pageNumber="54">Iki City</collectingCounty>
,
<collectingMunicipality id="1F519228E9114A5FFE03FB61FDC0FB4B" box="[424,574,1189,1216]" pageId="19" pageNumber="54">Ashibe Town</collectingMunicipality>
,
<location id="FA555E89E9114A5FFDE2FB61FCB6FB34" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFDE2FB61FCB6FB34" box="[585,840,1189,1215]" country="Japan" county="Iki City" municipality="Ashibe Town" name="Hakozakihonmurafure" pageId="19" pageNumber="54" stateProvince="Nagasaki">Hakozakihonmurafure</location>
,
<date id="8B342E92E9114A5FFCF9FB61FC14FB34" box="[850,1002,1189,1216]" pageId="19" pageNumber="54" value="2021-07-17">
<collectingDate id="9B70D77AE9114A5FFCF9FB61FC14FB34" box="[850,1002,1189,1216]" pageId="19" pageNumber="54" value="2021-07-17">17. VII. 2021</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFC5FFB62FBACFB34" box="[1012,1106,1189,1216]" pageId="19" pageNumber="54">K. Otsui</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFB24FB61FB05FB34" box="[1167,1275,1189,1215]" pageId="19" pageNumber="54">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFAA6FB62FBA9FB68" collectingDate="1999-07-12" collectingDateMax="1999-08-05" collectingDateMin="1999-07-12" collectionCode="MU" collectorName="Yakushima Is. &amp; T. Murata" country="Japan" location="Miyanoura" municipality="Is." pageId="19" pageNumber="54" specimenCount="1" stateProvince="Kagoshima">
[
<emphasis id="CDFED440E9114A5FFABDFB61FF4AFB68" bold="true" pageId="19" pageNumber="54">
<collectorName id="527F6D84E9114A5FFABDFB61FF4AFB68" pageId="19" pageNumber="54">Yakushima Is.</collectorName>
</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9114A5FFF56FB0DFE83FB68" box="[253,381,1225,1251]" country="Japan" name="Kagoshima" pageId="19" pageNumber="54">Kagoshima</collectingRegion>
<collectingMunicipality id="1F519228E9114A5FFE29FB0EFE61FB6F" box="[386,415,1226,1252]" pageId="19" pageNumber="54">Is.</collectingMunicipality>
,
<location id="FA555E89E9114A5FFE01FB0DFDD6FB68" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFE01FB0DFDD6FB68" box="[426,552,1225,1251]" country="Japan" municipality="Is." name="Miyanoura" pageId="19" pageNumber="54" stateProvince="Kagoshima">Miyanoura</location>
,
<date id="8B342E92E9114A5FFD99FB0DFCD6FB68" box="[562,808,1225,1252]" pageId="19" pageNumber="54" value="1999-07-12" valueMax="1999-08-05" valueMin="1999-07-12">
<collectingDate id="9B70D77AE9114A5FFD99FB0DFCD6FB68" box="[562,808,1225,1252]" pageId="19" pageNumber="54" value="1999-07-12" valueMax="1999-08-05" valueMin="1999-07-12">12. VII.–5. VIII. 1999</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFC99FB0EFC61FB6F" box="[818,927,1226,1252]" pageId="19" pageNumber="54">T. Murata</collectorName>
leg., MsT (
<collectionCode id="999B9097E9114A5FFBB6FB0DFBAEFB68" box="[1053,1104,1225,1251]" country="USA" name="Midwestern University" pageId="19" pageNumber="54">MU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9114A5FFBC9FB0DFEE7FA8C" collectingDate="1999-08-06" collectingDateMax="1999-08-27" collectingDateMin="1999-08-06" collectionCode="MU" collectorName="Yakushima Is. &amp; T. Murata" country="Japan" location="Miyanoura" municipality="Is." pageId="19" pageNumber="54" specimenCount="1" stateProvince="Kagoshima">
1 F &amp; 1 M, ditto,
<date id="8B342E92E9114A5FFA8AFB0DFF31FA8C" pageId="19" pageNumber="54" value="1999-08-06" valueMax="1999-08-27" valueMin="1999-08-06">
<collectingDate id="9B70D77AE9114A5FFA8AFB0DFF31FA8C" pageId="19" pageNumber="54" value="1999-08-06" valueMax="1999-08-27" valueMin="1999-08-06">6–27. VIII. 1999</collectingDate>
</date>
(
<collectionCode id="999B9097E9114A5FFF75FB29FEEFFA8C" box="[222,273,1261,1287]" country="USA" name="Midwestern University" pageId="19" pageNumber="54">MU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE9114A5FFE8FFB29FA99FA8C" box="[292,1383,1261,1288]" collectingDate="2007-07-20" collectionCode="GSFGPM" collectorName="Jiang Fengchen" country="China" location="Jingshan" municipality="Sheng" pageId="19" pageNumber="54" specimenCount="1" stateProvince="Hubei">
<emphasis id="CDFED440E9114A5FFE8FFB29FE7FFA8C" bold="true" box="[292,385,1261,1287]" pageId="19" pageNumber="54">
<collectingCountry id="879D48C2E9114A5FFE8FFB29FE7FFA8C" box="[292,385,1261,1287]" name="China" pageId="19" pageNumber="54">CHINA</collectingCountry>
</emphasis>
: 1 F (det. M-L.
<collectingMunicipality id="1F519228E9114A5FFD9CFB29FD7FFA8C" box="[567,641,1261,1287]" pageId="19" pageNumber="54">Sheng</collectingMunicipality>
),
<collectingRegion id="3D4EC6B0E9114A5FFD3FFB29FD25FA8C" box="[660,731,1261,1287]" country="China" name="Hubei" pageId="19" pageNumber="54">Hubei</collectingRegion>
,
<location id="FA555E89E9114A5FFD4DFB29FCB5FA8C" LSID="urn:lsid:plazi:treatment:7723B944E9114A5AFF3CFD77FAF2FCD6:FA555E89E9114A5FFD4DFB29FCB5FA8C" box="[742,843,1261,1287]" country="China" municipality="Sheng" name="Jingshan" pageId="19" pageNumber="54" stateProvince="Hubei">Jingshan</location>
,
<date id="8B342E92E9114A5FFCFDFB29FC0EFA8C" box="[854,1008,1261,1288]" pageId="19" pageNumber="54" value="2007-07-20">
<collectingDate id="9B70D77AE9114A5FFCFDFB29FC0EFA8C" box="[854,1008,1261,1288]" pageId="19" pageNumber="54" value="2007-07-20">20. VII. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E9114A5FFC50FB29FB54FA8C" box="[1019,1194,1261,1287]" pageId="19" pageNumber="54">Jiang Fengchen</collectorName>
leg. (
<collectionCode id="999B9097E9114A5FFB43FB29FAA1FA8C" box="[1256,1375,1261,1287]" pageId="19" pageNumber="54">GSFGPM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9114A5AFF6CFAD5FDC6FD46" lastPageId="22" lastPageNumber="57" pageId="19" pageNumber="54" type="description">
<paragraph id="FF350852E9114A5FFF6CFAD5FC9AFADB" blockId="19.[151,1437,865,2007]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF6CFAD5FEAFFAA0" bold="true" box="[199,337,1297,1323]" pageId="19" pageNumber="54">Description</emphasis>
.
<emphasis id="CDFED440E9114A5FFECBFAD5FE49FAA0" bold="true" box="[352,439,1297,1323]" pageId="19" pageNumber="54">Female</emphasis>
(n = 10). Body polished and covered with punctures and silvery setae; length 11.7–15.0 mm. Setae on head and mesosoma exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E9114A5FFF6CFA9DFE1DF9E4" blockId="19.[151,1437,865,2007]" pageId="19" pageNumber="54">
Head 0.55–0.6 × as long as wide. Combined length of clypeus and face 1.0 × facial width. Clypeus smooth ventrally, densely punctate dorsally, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons concave, without horns or convexities. Malar space 0.55–0.6 × as long as basal mandibular width. Genal carina lamella-like ventrally, reaching mandibular base. POL 1.5–1.65 × as long as OD. OOL 1.5–1.7 × as long as OD. Temple rather linearly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9114A5FFCCAF9C9FC31F9AC" box="[865,975,1549,1575]" captionStart="FIGURE 13" captionStartId="20.[151,250,1828,1852]" captionTargetBox="[151,1436,199,1805]" captionTargetId="figure-19@20.[151,1436,199,1805]" captionTargetPageId="20" captionText="FIGURE 13. Syzeuctus maculatus Sheng, 2009, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="19" pageNumber="54">Fig. 13 D</figureCitation>
). Antenna with 46–49 flagellomeres. FL I 4.8–5.0 × as long as maximum depth in lateral view and 2.0–2.15 × as long as FL II. Preapical flagellomere 1.1–1.2 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E9114A5FFF6CF9BEFBC1F85C" blockId="19.[151,1437,865,2007]" pageId="19" pageNumber="54">
Mesosoma coarsely and densely punctate, 1.4 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, short, its dorsal part largely absent. Mesopleuron densely punctate; speculum with a large smooth area, its lower area concave (
<figureCitation id="67B114D7E9114A5FFC89F905FC75F950" box="[802,907,1729,1755]" captionStart="FIGURE 13" captionStartId="20.[151,250,1828,1852]" captionTargetBox="[151,1436,199,1805]" captionTargetId="figure-19@20.[151,1436,199,1805]" captionTargetPageId="20" captionText="FIGURE 13. Syzeuctus maculatus Sheng, 2009, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="19" pageNumber="54">Fig. 13 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E9114A5FFB2DF921FB19F974" box="[1158,1255,1765,1791]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="19" pageNumber="54">Fig. 1 G</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum present but sometimes partly obsolete medially and/or laterally. Fore wing length
<quantity id="3872A5B7E9114A5FFD40F8E9FC7AF8C3" box="[747,900,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="9.899999999999999" metricValueMax="10.1" metricValueMin="9.7" pageId="19" pageNumber="54" unit="mm" value="9.899999999999999" valueMax="10.1" valueMin="9.7">9.7–10.1 mm</quantity>
. Areolet receiving vein 2m-cu near the middle, petiolated anteriorly; petiole longer than areolet (
<figureCitation id="67B114D7E9114A5FFD11F895FCDAF8E0" box="[698,804,1873,1899]" captionStart="FIGURE 13" captionStartId="20.[151,250,1828,1852]" captionTargetBox="[151,1436,199,1805]" captionTargetId="figure-19@20.[151,1436,199,1805]" captionTargetPageId="20" captionText="FIGURE 13. Syzeuctus maculatus Sheng, 2009, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="19" pageNumber="54">Fig. 13 F</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal or interstitial to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 6.25–6.5 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 3.8–4.0: 2.0: 1.4: 0.55–0.65: 0.7. Hind tarsal claws pectinate by short teeth, strongly bent apically (ca. 90°) (
<figureCitation id="67B114D7E9114A5FFC76F879FBCFF85C" box="[989,1073,1981,2007]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="19" pageNumber="54">Fig. 2 I</figureCitation>
).
</paragraph>
<caption id="ABF558DAE9164A58FF3CF8E0FADBF8EB" pageId="20" pageNumber="55" startId="20.[151,250,1828,1852]" targetBox="[151,1436,199,1805]" targetPageId="20" targetType="figure">
<paragraph id="FF350852E9164A58FF3CF8E0FADBF8EB" blockId="20.[151,1437,1828,1889]" pageId="20" pageNumber="55">
<emphasis id="CDFED440E9164A58FF3CF8E0FEDFF8B7" bold="true" box="[151,289,1828,1852]" pageId="20" pageNumber="55">FIGURE 13.</emphasis>
<taxonomicName id="388A73D1E9164A58FE82F8E1FD78F8B6" authority="Sheng, 2009" authorityName="Sheng" authorityYear="2009" box="[297,646,1828,1853]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="55" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="CDFED440E9164A58FE82F8E1FE02F8B7" box="[297,508,1828,1852]" italics="true" pageId="20" pageNumber="55">Syzeuctus maculatus</emphasis>
<bibRefCitation id="9B1B75A3E9164A58FDAFF8E0FD78F8B6" author="Sheng, M-L. &amp; Sun, S." box="[516,646,1828,1853]" pageId="20" pageNumber="55" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">Sheng, 2009</bibRefCitation>
</taxonomicName>
, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing.
</paragraph>
</caption>
<caption id="ABF558DAE9174A59FF3CF9A7FA65F9F0" box="[151,1435,1635,1659]" pageId="21" pageNumber="56" startId="21.[151,250,1635,1659]" targetBox="[151,1436,181,1611]" targetPageId="21" targetType="figure">
<paragraph id="FF350852E9174A59FF3CF9A7FA65F9F0" blockId="21.[151,1435,1635,1659]" box="[151,1435,1635,1659]" pageId="21" pageNumber="56">
<emphasis id="CDFED440E9174A59FF3CF9A7FEDEF9F0" bold="true" box="[151,288,1635,1659]" pageId="21" pageNumber="56">FIGURE 14.</emphasis>
<taxonomicName id="388A73D1E9174A59FE8DF9A7FD80F9F0" authority="Sheng, 2009" authorityName="Sheng" authorityYear="2009" box="[294,638,1635,1659]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="56" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="CDFED440E9174A59FE8DF9A7FE09F9F0" box="[294,503,1635,1659]" italics="true" pageId="21" pageNumber="56">Syzeuctus maculatus</emphasis>
<bibRefCitation id="9B1B75A3E9174A59FE56F9A7FD80F9F0" author="Sheng, M-L. &amp; Sun, S." box="[509,638,1635,1659]" pageId="21" pageNumber="56" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">Sheng, 2009</bibRefCitation>
</taxonomicName>
, male from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view.
</paragraph>
</caption>
<paragraph id="FF350852E9174A59FF6CF96EFD86F8A4" blockId="21.[151,1436,1705,2020]" pageId="21" pageNumber="56">Metasoma. T I 2.3–2.6 × as long as maximum width, densely punctate except for smooth areas on median longitudinal area and area along posterior margin. T II 1.8 × as long as maximum width, densely punctate. T III to T V coriaceous with fine and dense punctures. Ovipositor sheath 2.1–2.2 × as long as hind tibia, ca. 1.05 × as long as fore wing, and shorter than body length.</paragraph>
<paragraph id="FF350852E9174A5AFF6CF8FDFC78FD92" blockId="21.[151,1436,1705,2020]" lastBlockId="22.[151,1437,151,861]" lastPageId="22" lastPageNumber="57" pageId="21" pageNumber="56">
Colouration (
<figureCitation id="67B114D7E9174A59FEF6F8FDFE0EF8D8" box="[349,496,1849,1875]" captionStart="FIGURE 13" captionStartId="20.[151,250,1828,1852]" captionTargetBox="[151,1436,199,1805]" captionTargetId="figure-19@20.[151,1436,199,1805]" captionTargetPageId="20" captionText="FIGURE 13. Syzeuctus maculatus Sheng, 2009, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="21" pageNumber="56">Figs 13 A–F</figureCitation>
). Body (excluding wings and legs) black. Clypeus, mandible except for teeth, face, malar space, inner orbit on frons and vertex, and outer orbit yellow. Scape and pedicel tinged with yellow to reddish-yellow ventrally. Flagellum brown except for basal segments tinged with reddish-yellow. Pronotum with a pair of dorsal spots and collar yellow. Propleuron except for anterior black marking yellow. Mesoscutum with a pair of antero-lateral yellow markings and a median yellow marking. Subtegular ridge, antero-dorsal and postero-ventral spots of mesopleuron, posterior part of mesosternum, tegula, scutellum except a median small black spot, postscutellum, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of metapleuron and propodeum yellow. A pair of antero-lateral yellow spots of propodeum sometimes united into a posterior triangle yellow marking of propodeum. T I yellow except for a subapical black stripe. T II to T VII reddish-yellow each with yellow posterior margin. T II and T III sometimes partly indistinctly darkened. Membranous parts of metasomal sternites largely yellow. Subgenital plate reddish-yellow. Ovipositor reddish-brown. Wings hyaline. Apical clouded area of fore wing present and distinct. Veins and pterostigma brown except for yellowish-brown wing base and median part of pterostigma. Fore and mid legs yellow except for mid fifth tarsomere black. Hind coxa blackish-brown basally, yellow apically. Hind trochanter and femur reddish-yellow except for each apex tinged with yellow. Hind trochantellus yellow. Hind tibia yellow basally, reddish-yellow apically. Hind tibial spurs reddish-yellow to reddish-brown. Hind tarsus dark reddish-brown to blackish-brown.
</paragraph>
<paragraph id="FF350852E9144A5AFF6CFDE7FDC6FD46" blockId="22.[151,1437,151,861]" pageId="22" pageNumber="57">
<emphasis id="CDFED440E9144A5AFF6CFDE7FEFAFDB6" bold="true" box="[199,260,547,573]" pageId="22" pageNumber="57">Male</emphasis>
(n = 23). Similar to female (
<figureCitation id="67B114D7E9144A5AFDEEFDE7FD27FDB6" box="[581,729,547,573]" captionStart="FIGURE 14" captionStartId="21.[151,250,1635,1659]" captionTargetBox="[151,1436,181,1611]" captionTargetId="figure-19@21.[151,1436,181,1611]" captionTargetPageId="21" captionText="FIGURE 14. Syzeuctus maculatus Sheng, 2009, male from Japan—A: lateral habitus; B: dorsal habitus; C: head, frontal view." pageId="22" pageNumber="57">Figs 14 A–C</figureCitation>
). POL 1.6–1.8 × as long as OD. OOL 1.3–1.6 × as long as OD. Antenna with 50–52 flagellomeres. Tergal spiracles not enlarged. Hind femur 5.75–6.2 × as long as maximum depth in lateral view. T I 2.75–3.05 × as long as maximum width. T II 2.0–2.25 × as long as maximum width. Yellow area of body larger than female. Yellow spots of mesopleuron united into a large yellow area with yellow mesosternum. Apex of metasoma weakly darkened.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9144A5AFF6CFD12FE36FCB2" pageId="22" pageNumber="57" type="distribution">
<paragraph id="FF350852E9144A5AFF6CFD12FC69FC9E" blockId="22.[151,1437,151,861]" pageId="22" pageNumber="57">
<emphasis id="CDFED440E9144A5AFF6CFD12FEA7FD7B" bold="true" box="[199,345,726,752]" pageId="22" pageNumber="57">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9144A5AFEC4FD13FE51FD7A" box="[367,431,727,753]" name="Japan" pageId="22" pageNumber="57">Japan</collectingCountry>
(Honshu, Izuoshima Is., Shikinejima Is., Kozushima Is., Miyakejima Is., Shikoku, Oomishima Is., Iki Is., and Yakushima Is.) and
<collectingCountry id="879D48C2E9144A5AFD0DFD3FFD13FC9E" box="[678,749,763,789]" name="China" pageId="22" pageNumber="57">China</collectingCountry>
. New to
<collectingCountry id="879D48C2E9144A5AFCFBFD3FFC6DFC9E" box="[848,915,763,789]" name="Japan" pageId="22" pageNumber="57">Japan</collectingCountry>
.
</paragraph>
<paragraph id="FF350852E9144A5AFF6CFCDAFE36FCB2" blockId="22.[151,1437,151,861]" box="[199,456,798,825]" pageId="22" pageNumber="57">
<emphasis id="CDFED440E9144A5AFF6CFCDAFEBDFCB3" bold="true" box="[199,323,798,824]" pageId="22" pageNumber="57">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9144A5AFF6CFC87FAF2FCD6" box="[199,1292,835,861]" pageId="22" pageNumber="57" type="discussion">
<paragraph id="FF350852E9144A5AFF6CFC87FAF2FCD6" blockId="22.[151,1437,151,861]" box="[199,1292,835,861]" pageId="22" pageNumber="57">
<emphasis id="CDFED440E9144A5AFF6CFC87FECAFCD6" bold="true" box="[199,308,835,861]" pageId="22" pageNumber="57">Remarks</emphasis>
. The distribution of this species is limited to island areas or dry areas close to the coast.
</paragraph>
</subSubSection>
</treatment>
</document>

214
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<paragraph id="FF350852E9134A5DFF3CFCDBFDEBFCB3" blockId="17.[151,533,798,861]" box="[151,533,798,825]" pageId="17" pageNumber="52">
<heading id="A47DBF3EE9134A5DFF3CFCDBFDEBFCB3" bold="true" box="[151,533,798,825]" fontSize="11" level="1" pageId="17" pageNumber="52" reason="1">
<taxonomicName id="388A73D1E9134A5DFF3CFCDBFDEBFCB3" authority="Riedel, 2022" authorityName="Riedel" authorityYear="2022" box="[151,533,798,825]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="CDFED440E9134A5DFF3CFCDBFDEBFCB3" bold="true" box="[151,533,798,825]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF3CFCDBFE8AFCB2" bold="true" box="[151,372,798,825]" italics="true" pageId="17" pageNumber="52">Syzeuctus laoticus</emphasis>
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</emphasis>
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<paragraph id="FF350852E9134A5DFF3CFC87FE62FCD6" blockId="17.[151,533,798,861]" box="[151,412,835,861]" pageId="17" pageNumber="52">
(
<figureCitation id="67B114D7E9134A5DFF34FC87FF03FCD6" box="[159,253,835,861]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="17" pageNumber="52">Figs 1 F</figureCitation>
,
<figureCitation id="67B114D7E9134A5DFEA3FC87FECAFCD6" box="[264,308,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="17" pageNumber="52">2 H</figureCitation>
,
<figureCitation id="67B114D7E9134A5DFE94FC87FE6AFCD6" box="[319,404,835,861]" captionStart="FIGURE 12" captionStartId="18.[151,250,1596,1620]" captionTargetBox="[151,1436,181,1571]" captionTargetId="figure-19@18.[151,1436,181,1571]" captionTargetPageId="18" captionText="FIGURE 12. Syzeuctus laoticus Riedel, 2022, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="17" pageNumber="52">12 A–F</figureCitation>
)
</paragraph>
</subSubSection>
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<treatmentCitationGroup id="DF9A2F7CE9134A5DFF3CFC43FDEDFC14" box="[151,531,903,927]" pageId="17" pageNumber="52">
<treatmentCitation id="7E2B2E43E9134A5DFF3CFC43FDF0FC14" author="Riedel, M." box="[151,526,903,927]" page="267" pageId="17" pageNumber="52" year="2022">
<taxonomicName id="388A73D1E9134A5DFF3CFC43FDF0FC14" authority="Riedel, 2022: 267" authorityName="Riedel" authorityPageNumber="267" authorityYear="2022" box="[151,526,903,927]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="CDFED440E9134A5DFF3CFC43FEAEFC14" box="[151,336,903,927]" italics="true" pageId="17" pageNumber="52">Syzeuctus laoticus</emphasis>
<bibRefCitation id="9B1B75A3E9134A5DFEFCFC43FDF0FC14" author="Riedel, M." box="[343,526,903,927]" pageId="17" pageNumber="52" pagination="257 - 273" refId="ref23331" refString="Riedel, M. (2022) Contribution to the knowledge of Oriental Syzeuctus Forster (Hymenoptera, Ichneumonidae, Banchinae), with description of six new species. Linzer biologische Beitrage, 54 (1), 257 - 273." type="journal article" year="2022">Riedel, 2022: 267</bibRefCitation>
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.
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<paragraph id="FF350852E9134A5DFF3CFC09FD02FBA4" blockId="17.[151,1437,973,1936]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF3CFC09FE7DFC6C" bold="true" box="[151,387,973,999]" pageId="17" pageNumber="52">Materials examined</emphasis>
.
<materialsCitation id="4FE2020FE9134A5DFE3AFC09FDEEFB80" collectingDate="2013-07-20" collectingDateMax="2013-07-21" collectingDateMin="2013-07-20" collectionCode="KPMNH" collectorName="N. Kikuchi" country="Japan" county="Chichibu City" location="Irikawa forest" municipality="Otaki" pageId="17" pageNumber="52" specimenCount="1" stateProvince="Saitama">
<emphasis id="CDFED440E9134A5DFE3AFC09FE15FC6C" bold="true" box="[401,491,973,999]" pageId="17" pageNumber="52">
<collectingCountry id="879D48C2E9134A5DFE3AFC09FE15FC6C" box="[401,491,973,999]" name="Japan" pageId="17" pageNumber="52">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E9134A5DFDA9FC09FD97FC6C" bold="true" box="[514,617,973,999]" pageId="17" pageNumber="52">Honshu]</emphasis>
, 1 F,
<collectingRegion id="3D4EC6B0E9134A5DFD0FFC09FCC3FC6C" box="[676,829,973,999]" country="Japan" name="Saitama" pageId="17" pageNumber="52">Saitama Pref.</collectingRegion>
,
<collectingCounty id="165470DEE9134A5DFCECFC09FC1BFC6C" box="[839,997,973,999]" pageId="17" pageNumber="52">Chichibu City</collectingCounty>
,
<collectingMunicipality id="1F519228E9134A5DFC5BFC09FBCFFC6C" box="[1008,1073,973,999]" pageId="17" pageNumber="52">Otaki</collectingMunicipality>
,
<location id="FA555E89E9134A5DFB90FC09FB29FC6C" LSID="urn:lsid:plazi:treatment:7723B944E9134A5FFF3CFCDBFE43FDEA:FA555E89E9134A5DFB90FC09FB29FC6C" box="[1083,1239,973,999]" country="Japan" county="Chichibu City" municipality="Otaki" name="Irikawa forest" pageId="17" pageNumber="52" stateProvince="Saitama">Irikawa forest</location>
road,
<date id="8B342E92E9134A5DFAB7FC09FF2CFB80" pageId="17" pageNumber="52" value="2013-07-20" valueMax="2013-07-21" valueMin="2013-07-20">
<collectingDate id="9B70D77AE9134A5DFAB7FC09FF2CFB80" pageId="17" pageNumber="52" value="2013-07-20" valueMax="2013-07-21" valueMin="2013-07-20">20–21. VII. 2013</collectingDate>
</date>
,
<collectorName id="527F6D84E9134A5DFF75FC36FEA5FB80" box="[222,347,1009,1036]" pageId="17" pageNumber="52">N. Kikuchi</collectorName>
leg. (
<collectionCode id="999B9097E9134A5DFE37FC35FDF6FB80" box="[412,520,1009,1035]" pageId="17" pageNumber="52">KPMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE9134A5DFDB0FC35FD06FBA4" collectingDate="2013-06-20" collectorName="C. Holzschuh" country="Laos" location="Phon-Pan" pageId="17" pageNumber="52" specimenCount="1" stateProvince="Houaphan" typeStatus="holotype">
<emphasis id="CDFED440E9134A5DFDB0FC35FD99FB80" bold="true" box="[539,615,1009,1035]" pageId="17" pageNumber="52">
<collectingCountry id="879D48C2E9134A5DFDB0FC35FD99FB80" box="[539,615,1009,1035]" name="Laos" pageId="17" pageNumber="52">LAOS</collectingCountry>
</emphasis>
: 1 F (
<typeStatus id="2031B6F0E9134A5DFD06FC35FCEBFB80" box="[685,789,1009,1035]" pageId="17" pageNumber="52" type="holotype">holotype</typeStatus>
), Prov.
<collectingRegion id="3D4EC6B0E9134A5DFCC1FC36FBE0FB87" box="[874,1054,1009,1036]" country="Laos" name="Houaphan" pageId="17" pageNumber="52">Hua Phan Prov.</collectingRegion>
,
<location id="FA555E89E9134A5DFB81FC35FB64FB80" LSID="urn:lsid:plazi:treatment:7723B944E9134A5FFF3CFCDBFE43FDEA:FA555E89E9134A5DFB81FC35FB64FB80" box="[1066,1178,1009,1035]" country="Laos" name="Phon-Pan" pageId="17" pageNumber="52" stateProvince="Houaphan">Phon-Pan</location>
, Umg. Ort Ban Saleui,
<date id="8B342E92E9134A5DFF3CFBD1FED9FBA4" box="[151,295,1045,1072]" pageId="17" pageNumber="52" value="2013-06-20">
<collectingDate id="9B70D77AE9134A5DFF3CFBD1FED9FBA4" box="[151,295,1045,1072]" pageId="17" pageNumber="52" value="2013-06-20">20. VI. 2013</collectingDate>
</date>
, KJa, leg.
<collectorName id="527F6D84E9134A5DFE37FBD1FDCBFBA4" box="[412,565,1045,1071]" pageId="17" pageNumber="52">C. Holzschuh</collectorName>
&amp; natives (Linz)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9134A5FFF6CFBFDFE76FE26" lastPageId="19" lastPageNumber="54" pageId="17" pageNumber="52" type="description">
<paragraph id="FF350852E9134A5DFF6CFBFDFB5EFBF3" blockId="17.[151,1437,973,1936]" pageId="17" pageNumber="52">
<emphasis id="CDFED440E9134A5DFF6CFBFDFD4CFBD8" bold="true" box="[199,690,1081,1107]" pageId="17" pageNumber="52">Description based on Japanese specimen</emphasis>
.
<emphasis id="CDFED440E9134A5DFD68FBFDFCE4FBD8" bold="true" box="[707,794,1081,1107]" pageId="17" pageNumber="52">Female</emphasis>
(n = 1). Body polished and covered with punctures and silvery setae; length
<quantity id="3872A5B7E9134A5DFED5FB99FE29FBF3" box="[382,471,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="17" pageNumber="52" unit="mm" value="8.7">8.7 mm</quantity>
. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E9134A5DFF6CFB45FF21FA1C" blockId="17.[151,1437,973,1936]" pageId="17" pageNumber="52">
Head 0.57 × as long as wide. Combined length of clypeus and face 1.0 × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons almost flat, without horns or convexities. Malar space 0.7 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.75 × as long as OD. OOL 1.0 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9134A5DFD55FAF1FC8CFAC4" box="[766,882,1333,1359]" captionStart="FIGURE 12" captionStartId="18.[151,250,1596,1620]" captionTargetBox="[151,1436,181,1571]" captionTargetId="figure-19@18.[151,1436,181,1571]" captionTargetPageId="18" captionText="FIGURE 12. Syzeuctus laoticus Riedel, 2022, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="17" pageNumber="52">Fig. 12 D</figureCitation>
). Antenna with 42 flagellomeres. FL I 3.75 × as long as maximum depth in lateral view and 1.5 × as long as FL II. Preapical flagellomere 1.2 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E9134A5DFF6CFA66FCE9F974" blockId="17.[151,1437,973,1936]" pageId="17" pageNumber="52">
Mesosoma coarsely and densely punctate, 1.4 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron punctate; speculum with a large smooth area (
<figureCitation id="67B114D7E9134A5DFE1DFA2DFDDFF988" box="[438,545,1513,1539]" captionStart="FIGURE 12" captionStartId="18.[151,250,1596,1620]" captionTargetBox="[151,1436,181,1571]" captionTargetId="figure-19@18.[151,1436,181,1571]" captionTargetPageId="18" captionText="FIGURE 12. Syzeuctus laoticus Riedel, 2022, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="17" pageNumber="52">Fig. 12 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end close to anterior margin of mesopleuron, with weak angulation (
<figureCitation id="67B114D7E9134A5DFD55F9C9FCA6F9AC" box="[766,856,1549,1575]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="17" pageNumber="52">Fig. 1 F</figureCitation>
). Dorsal surface of propodeum with weak transverse rugae posteriorly. Posterior transverse carina of propodeum absent laterally, shortly and obsoletely present medially. Fore wing length
<quantity id="3872A5B7E9134A5DFEC3F991FE3AF9FB" box="[360,452,1621,1648]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="17" pageNumber="52" unit="mm" value="6.6">6.6 mm</quantity>
. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E9134A5DFE1CF9BDFDDEF918" box="[439,544,1657,1683]" captionStart="FIGURE 12" captionStartId="18.[151,250,1596,1620]" captionTargetBox="[151,1436,181,1571]" captionTargetId="figure-19@18.[151,1436,181,1571]" captionTargetPageId="18" captionText="FIGURE 12. Syzeuctus laoticus Riedel, 2022, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="17" pageNumber="52">Fig. 12 F</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.75 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 3.9: 2.0: 1.4: 0.7: 0.8. Hind tarsal claws simple except for minute subbasal teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9134A5DFD02F921FCF4F974" box="[681,778,1765,1791]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="17" pageNumber="52">Fig. 2 H</figureCitation>
).
</paragraph>
<paragraph id="FF350852E9134A5DFF6CF8CEFD86F804" blockId="17.[151,1437,973,1936]" pageId="17" pageNumber="52">Metasoma. T I 2.0 × as long as maximum width, finely punctate except for smooth areas on median longitudinal area, area along posterior margin and lateral sides. T II 1.2–1.4 × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 3.35 × as long as hind tibia, ca. 1.2 × as long as fore wing, and shorter than body length.</paragraph>
<caption id="ABF558DAE9104A5EFF3CF9F8FADBF9F3" pageId="18" pageNumber="53" startId="18.[151,250,1596,1620]" targetBox="[151,1436,181,1571]" targetPageId="18" targetType="figure">
<paragraph id="FF350852E9104A5EFF3CF9F8FADBF9F3" blockId="18.[151,1437,1596,1656]" pageId="18" pageNumber="53">
<emphasis id="CDFED440E9104A5EFF3CF9F8FEDDF9DF" bold="true" box="[151,291,1596,1620]" pageId="18" pageNumber="53">FIGURE 12.</emphasis>
<taxonomicName id="388A73D1E9104A5EFE87F9F8FD86F9DF" authority="Riedel, 2022" authorityName="Riedel" authorityYear="2022" box="[300,632,1596,1620]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="CDFED440E9104A5EFE87F9F8FE16F9DF" box="[300,488,1596,1620]" italics="true" pageId="18" pageNumber="53">Syzeuctus laoticus</emphasis>
<bibRefCitation id="9B1B75A3E9104A5EFE5AF9F8FD86F9DF" author="Riedel, M." box="[497,632,1596,1620]" pageId="18" pageNumber="53" pagination="257 - 273" refId="ref23331" refString="Riedel, M. (2022) Contribution to the knowledge of Oriental Syzeuctus Forster (Hymenoptera, Ichneumonidae, Banchinae), with description of six new species. Linzer biologische Beitrage, 54 (1), 257 - 273." type="journal article" year="2022">Riedel, 2022</bibRefCitation>
</taxonomicName>
, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing.
</paragraph>
</caption>
<paragraph id="FF350852E9104A5FFF6CF962FC71FE02" blockId="18.[151,1437,1702,2017]" lastBlockId="19.[151,1437,151,609]" lastPageId="19" lastPageNumber="54" pageId="18" pageNumber="53">
Colouration (
<figureCitation id="67B114D7E9104A5EFEC8F962FDFFF94B" box="[355,513,1702,1728]" captionStart="FIGURE 12" captionStartId="18.[151,250,1596,1620]" captionTargetBox="[151,1436,181,1571]" captionTargetId="figure-19@18.[151,1436,181,1571]" captionTargetPageId="18" captionText="FIGURE 12. Syzeuctus laoticus Riedel, 2022, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: fore wing." pageId="18" pageNumber="53">Figs 12 A–F</figureCitation>
). Body (excluding wings and legs) black. Clypeus, face except for a dorso-median longitudinal black stripe, malar space, inner and outer orbits, ventral surfaces of scape and pedicel, and palpi yellow. Flagellum largely reddish-brown to brown. Pronotum except for antero-median part and area along posterior margin, posterior part of propleuron, a pair of antero-lateral triangular markings and a median marking of mesoscutum, scutellum except for an antero-median black spot, postscutellum, subtegular ridge, large C-shaped marking of mesopleuron, upper division of metapleuron, posterior part of metapleuron, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of propodeum yellow. Base of T I with a pair of yellow markings. Posterior margins of T I to T IV each with yellow to ivory band. Posterior margins of T V to T VII largely tinged with red to reddish-brown. Metasomal tergites more or less tinged with red to reddish-brown on T III to T VII. Membranous parts of metasomal sternites and subgenital plate except for an apical blackish spot yellow. Ovipositor reddish-brown. Wings hyaline, apical part with slightly darkened clouded area. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow. Hind coxa black laterally, yellow dorsally and ventrally. Hind trochanter black with yellow apical margin. Hind trochantellus yellow. Hind femur dark brown except for yellowish-brown apically. Hind tibia yellowish-brown, its apical part darkened. Hind tibial spurs pale yellowish-brown. Hind tarsus brown except for blackish-brown fifth tarsomere; bases of first to third tarsomeres, apex of third tarsomere, and fourth tarsomere paler than other parts.
</paragraph>
<paragraph id="FF350852E9114A5FFF6CFE57FE76FE26" blockId="19.[151,1437,151,609]" box="[199,392,403,429]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF6CFE57FEFAFE26" bold="true" box="[199,260,403,429]" pageId="19" pageNumber="54">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9114A5FFF6CFE72FE36FE7E" pageId="19" pageNumber="54" type="distribution">
<paragraph id="FF350852E9114A5FFF6CFE72FCCDFE5A" blockId="19.[151,1437,151,609]" box="[199,819,438,465]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF6CFE72FEA7FE5B" bold="true" box="[199,345,438,464]" pageId="19" pageNumber="54">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9114A5FFECCFE73FE59FE5A" box="[359,423,439,465]" name="Japan" pageId="19" pageNumber="54">Japan</collectingCountry>
(Honshu) and
<collectingCountry id="879D48C2E9114A5FFDE5FE73FD76FE5A" box="[590,648,439,465]" name="Laos" pageId="19" pageNumber="54">Laos</collectingCountry>
. New to
<collectingCountry id="879D48C2E9114A5FFD40FE73FCD0FE5A" box="[747,814,439,465]" name="Japan" pageId="19" pageNumber="54">Japan</collectingCountry>
.
</paragraph>
<paragraph id="FF350852E9114A5FFF6CFE1EFE36FE7E" blockId="19.[151,1437,151,609]" box="[199,456,474,501]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF6CFE1EFEBDFE7F" bold="true" box="[199,323,474,500]" pageId="19" pageNumber="54">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9114A5FFF6CFE3BFE43FDEA" pageId="19" pageNumber="54" type="discussion">
<paragraph id="FF350852E9114A5FFF6CFE3BFE43FDEA" blockId="19.[151,1437,151,609]" pageId="19" pageNumber="54">
<emphasis id="CDFED440E9114A5FFF6CFE3BFECAFD92" bold="true" box="[199,308,511,537]" pageId="19" pageNumber="54">Remarks</emphasis>
. The Japanese specimen differs from
<typeStatus id="2031B6F0E9114A5FFD72FE3BFCC5FD92" box="[729,827,511,537]" pageId="19" pageNumber="54" type="holotype">holotype</typeStatus>
by the smaller body, the T I 2.0 × as long as maximum width (2.6 × in
<typeStatus id="2031B6F0E9114A5FFE96FDE7FE5DFDB6" box="[317,419,547,573]" pageId="19" pageNumber="54" type="holotype">holotype</typeStatus>
), and ovipositor sheath slightly shorter than
<typeStatus id="2031B6F0E9114A5FFC2CFDE7FC12FDB6" box="[903,1004,547,573]" pageId="19" pageNumber="54" type="holotype">holotype</typeStatus>
, But we conclude that these differences are intraspecific variation.
</paragraph>
</subSubSection>
</treatment>
</document>

422
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<mods:location id="579D5A1330D0B9E5D4258E848606EAFF">
<mods:url id="9FEF625F987AED6925F91F8FD6FEC614">http://dx.doi.org/10.11646/zootaxa.5496.1.2</mods:url>
</mods:location>
<mods:classification id="67DEC654F54F01F1D801CBBCE51A19BF">journal article</mods:classification>
<mods:identifier id="99A5C886AC2AA67A158521C85C230F1B" type="DOI">10.11646/zootaxa.5496.1.2</mods:identifier>
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<mods:identifier id="BF5DD8992D12CC9FFE9BA1A7F1BBC31D" type="ZooBank">D993FA3C-03B0-4232-B762-0F24CA6EBF08</mods:identifier>
</mods:mods>
<treatment id="7723B944E9144A55FF3CFC6BFD46FBD9" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9" httpUri="http://treatment.plazi.org/id/7723B944E9144A55FF3CFC6BFD46FBD9" lastPageId="25" lastPageNumber="60" pageId="22" pageNumber="57">
<subSubSection id="B7905BD9E9144A5AFF3CFC6BFE37FC42" box="[151,457,943,969]" pageId="22" pageNumber="57" type="nomenclature">
<paragraph id="FF350852E9144A5AFF3CFC6BFE37FC42" blockId="22.[151,515,943,1005]" box="[151,457,943,969]" pageId="22" pageNumber="57">
<heading id="A47DBF3EE9144A5AFF3CFC6BFE37FC42" bold="true" box="[151,457,943,969]" fontSize="11" level="1" pageId="22" pageNumber="57" reason="1">
<emphasis id="CDFED440E9144A5AFF3CFC6BFE37FC42" bold="true" box="[151,457,943,969]" pageId="22" pageNumber="57">
<taxonomicName id="388A73D1E9144A5AFF3CFC6BFE9DFC42" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[151,355,943,969]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="57" phylum="Arthropoda" rank="species" species="nigrus" status="sp. nov.">
<emphasis id="CDFED440E9144A5AFF3CFC6BFE9DFC42" bold="true" box="[151,355,943,969]" italics="true" pageId="22" pageNumber="57">Syzeuctus nigrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6CD693BE9144A5AFEC0FC6BFE37FC42" box="[363,457,943,969]" pageId="22" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9144A5AFF3CFC17FDFDFC66" box="[151,515,978,1005]" pageId="22" pageNumber="57" type="description">
<paragraph id="FF350852E9144A5AFF3CFC17FDFDFC66" blockId="22.[151,515,943,1005]" box="[151,515,978,1005]" pageId="22" pageNumber="57">
(
<figureCitation id="67B114D7E9144A5AFF34FC17FEFAFC66" box="[159,260,979,1005]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="22" pageNumber="57">Figs 1 H</figureCitation>
,
<figureCitation id="67B114D7E9144A5AFEA4FC17FECFFC66" box="[271,305,979,1005]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="22" pageNumber="57">2 J</figureCitation>
,
<figureCitation id="67B114D7E9144A5AFE96FC16FE66FC66" box="[317,408,978,1005]" captionStart="FIGURE 15" captionStartId="23.[151,250,1925,1949]" captionTargetBox="[151,1436,381,1901]" captionTargetId="figure-122@23.[151,1436,381,1901]" captionTargetPageId="23" captionText="FIGURE 15. Syzeuctus nigrus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing." pageId="22" pageNumber="57">15 A–G</figureCitation>
,
<figureCitation id="67B114D7E9144A5AFE08FC17FE04FC66" box="[419,506,979,1005]" captionStart="FIGURE 16" captionStartId="24.[151,250,1654,1678]" captionTargetBox="[151,1436,151,1638]" captionTargetId="figure-19@24.[151,1436,151,1638]" captionTargetPageId="24" captionText="FIGURE 16. Syzeuctus nigrus sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view." pageId="22" pageNumber="57">16 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9144A5AFF3CFBDFFEE9FA16" pageId="22" pageNumber="57" type="materials_examined">
<paragraph id="FF350852E9144A5AFF3CFBDFFEE9FA16" blockId="22.[151,1437,1050,2013]" pageId="22" pageNumber="57">
<emphasis id="CDFED440E9144A5AFF3CFBDFFEE5FBBF" bold="true" box="[151,283,1050,1077]" pageId="22" pageNumber="57">Type series</emphasis>
.
<materialsCitation id="4FE2020FE9144A5AFE83FBDFFE71FBD2" collectingDate="2004-07-15" collectionCode="KPMNH" collectorName="Amamioshima Is. &amp; H. Makihara" country="Japan" location="Mt. Yui-dake" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<emphasis id="CDFED440E9144A5AFE83FBDFFE6BFBBE" bold="true" box="[296,405,1051,1077]" pageId="22" pageNumber="57">
<typeStatus id="2031B6F0E9144A5AFE83FBDFFE6BFBBE" box="[296,405,1051,1077]" pageId="22" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
: F,
<emphasis id="CDFED440E9144A5AFE15FBDEFDE6FBBF" bold="true" box="[446,536,1050,1076]" pageId="22" pageNumber="57">
<collectingCountry id="879D48C2E9144A5AFE15FBDEFDE6FBBF" box="[446,536,1050,1076]" name="Japan" pageId="22" pageNumber="57">JAPAN</collectingCountry>
</emphasis>
,
<emphasis id="CDFED440E9144A5AFD8EFBDEFD08FBBE" bold="true" box="[549,758,1050,1077]" pageId="22" pageNumber="57">
<collectorName id="527F6D84E9144A5AFD8EFBDEFD08FBBE" box="[549,758,1050,1077]" pageId="22" pageNumber="57">Amamioshima Is.</collectorName>
</emphasis>
,
<collectingRegion id="3D4EC6B0E9144A5AFCAFFBDFFC3DFBBE" box="[772,963,1051,1077]" country="Japan" name="Kagoshima" pageId="22" pageNumber="57">Kagoshima Pref.</collectingRegion>
,
<location id="FA555E89E9144A5AFC66FBDFFB9CFBBE" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFC66FBDFFB9CFBBE" box="[973,1122,1051,1077]" country="Japan" name="Mt. Yui-dake" pageId="22" pageNumber="57" stateProvince="Kagoshima">Mt. Yui-dake</location>
,
<date id="8B342E92E9144A5AFBC6FBDEFAFBFBBE" box="[1133,1285,1050,1077]" pageId="22" pageNumber="57" value="2004-07-15">
<collectingDate id="9B70D77AE9144A5AFBC6FBDEFAFBFBBE" box="[1133,1285,1050,1077]" pageId="22" pageNumber="57" value="2004-07-15">15. VII. 2004</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFAA4FBDFFA62FBBE" box="[1295,1436,1051,1077]" pageId="22" pageNumber="57">H. Makihara</collectorName>
leg., MsT (
<collectionCode id="999B9097E9144A5AFEB0FBFBFE79FBD2" box="[283,391,1087,1113]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE9144A5AFE36FBFBFDDDFBF6" collectingDate="2012-09-30" collectionCode="KPMNH" collectorName="Y. Saito" country="Japan" location="Shiroi City" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Chiba" typeStatus="paratype">
<emphasis id="CDFED440E9144A5AFE36FBFBFDEBFBD2" bold="true" box="[413,533,1087,1113]" pageId="22" pageNumber="57">
<typeStatus id="2031B6F0E9144A5AFE36FBFBFDEBFBD2" box="[413,533,1087,1113]" pageId="22" pageNumber="57" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="CDFED440E9144A5AFD83FBFAFD7CFBD3" bold="true" box="[552,642,1086,1112]" pageId="22" pageNumber="57">
<collectingCountry id="879D48C2E9144A5AFD83FBFAFD7CFBD3" box="[552,642,1086,1112]" name="Japan" pageId="22" pageNumber="57">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E9144A5AFD34FBFBFD02FBD2" bold="true" box="[671,764,1087,1113]" pageId="22" pageNumber="57">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9144A5AFCE0FBFBFC2CFBD2" box="[843,978,1087,1113]" country="Japan" name="Chiba" pageId="22" pageNumber="57">Chiba Pref.</collectingRegion>
,
<location id="FA555E89E9144A5AFC4AFBFBFB9CFBD2" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFC4AFBFBFB9CFBD2" box="[993,1122,1087,1113]" country="Japan" name="Shiroi City" pageId="22" pageNumber="57" stateProvince="Chiba">Shiroi City</location>
, Shiroishi-undou-koen,
<date id="8B342E92E9144A5AFAD2FBFBFEFFFBF6" pageId="22" pageNumber="57" value="2012-09-30">
<collectingDate id="9B70D77AE9144A5AFAD2FBFBFEFFFBF6" pageId="22" pageNumber="57" value="2012-09-30">30. IX. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFEA5FBA7FE94FBF6" box="[270,362,1123,1149]" pageId="22" pageNumber="57">Y. Saito</collectorName>
leg. (
<collectionCode id="999B9097E9144A5AFE1BFBA7FDE2FBF6" box="[432,540,1123,1149]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFD99FBA7FEEDFB2A" collectingDate="2008-10-05" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" location="Enkaizan" municipality="Yokohama City" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Kanagawa" typeStatus="paratype">
1 M,
<collectingRegion id="3D4EC6B0E9144A5AFDDFFBA7FCD0FBF6" box="[628,814,1123,1149]" country="Japan" name="Kanagawa" pageId="22" pageNumber="57">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9144A5AFC90FBA7FC10FBF6" box="[827,1006,1123,1149]" pageId="22" pageNumber="57">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E9144A5AFC56FBA7FB96FBF6" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFC56FBA7FB96FBF6" box="[1021,1128,1123,1149]" country="Japan" municipality="Yokohama City" name="Enkaizan" pageId="22" pageNumber="57" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E9144A5AFBDCFBA6FB09FBF6" box="[1143,1271,1122,1149]" pageId="22" pageNumber="57" value="2008-10-05">
<collectingDate id="9B70D77AE9144A5AFBDCFBA6FB09FBF6" box="[1143,1271,1122,1149]" pageId="22" pageNumber="57" value="2008-10-05">5. X. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFAAEFBA7FA96FBF6" box="[1285,1384,1123,1149]" pageId="22" pageNumber="57">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E9144A5AFF34FB43FEF5FB2A" box="[159,267,1159,1185]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFEB6FB43FEAEFB4E" collectingDate="2012-08-17" collectingDateMax="2012-10-05" collectingDateMin="2012-08-17" collectionCode="KPMNH" collectorName="Izuoshima Is. &amp; K. Tsujii" country="Japan" location="Mt. Omaru-yama" municipality="Motomachi" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Tokyo" typeStatus="paratype">
[Izu Isls.] 1 F,
<collectingRegion id="3D4EC6B0E9144A5AFE16FB43FDF9FB2A" box="[445,519,1159,1185]" country="Japan" name="Tokyo" pageId="22" pageNumber="57">Tokyo</collectingRegion>
,
<collectorName id="527F6D84E9144A5AFDB9FB43FD54FB2A" box="[530,682,1159,1185]" pageId="22" pageNumber="57">Izuoshima Is.</collectorName>
,
<collectingMunicipality id="1F519228E9144A5AFD1DFB43FCC4FB2A" box="[694,826,1159,1185]" pageId="22" pageNumber="57">Motomachi</collectingMunicipality>
,
<location id="FA555E89E9144A5AFCEEFB43FBF4FB2A" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFCEEFB43FBF4FB2A" box="[837,1034,1159,1185]" country="Japan" municipality="Motomachi" name="Mt. Omaru-yama" pageId="22" pageNumber="57" stateProvince="Tokyo">Mt. Omaru-yama</location>
,
<date id="8B342E92E9144A5AFBBFFB43FB04FB2A" box="[1044,1274,1158,1185]" pageId="22" pageNumber="57" value="2012-08-17" valueMax="2012-10-05" valueMin="2012-08-17">
<collectingDate id="9B70D77AE9144A5AFBBFFB43FB04FB2A" box="[1044,1274,1158,1185]" pageId="22" pageNumber="57" value="2012-08-17" valueMax="2012-10-05" valueMin="2012-08-17">17. VIII.–5. X. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFAAEFB43FA9BFB2A" box="[1285,1381,1159,1185]" pageId="22" pageNumber="57">K. Tsujii</collectorName>
leg., MsT (
<collectionCode id="999B9097E9144A5AFF77FB6FFEB6FB4E" box="[220,328,1195,1221]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFEF6FB6FFA66FB4E" box="[349,1432,1194,1221]" collectingDate="2002-09-22" collectionCode="KPMNH" collectorName="S. Ohkusa" country="Japan" location="Uchida" municipality="Sumoto" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Hyogo" typeStatus="paratype">
[
<emphasis id="CDFED440E9144A5AFECDFB6EFDE6FB4E" bold="true" box="[358,536,1194,1221]" pageId="22" pageNumber="57">Awajishima Is.</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9144A5AFDC1FB6FFD06FB4E" box="[618,760,1195,1221]" country="Japan" name="Hyogo" pageId="22" pageNumber="57">Hyogo Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9144A5AFCAEFB6FFCA1FB4E" box="[773,863,1195,1221]" pageId="22" pageNumber="57">Sumoto</collectingMunicipality>
,
<location id="FA555E89E9144A5AFCC0FB6FFC41FB4E" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFCC0FB6FFC41FB4E" box="[875,959,1195,1221]" country="Japan" municipality="Sumoto" name="Uchida" pageId="22" pageNumber="57" stateProvince="Hyogo">Uchida</location>
,
<date id="8B342E92E9144A5AFC67FB6FFB9EFB4E" box="[972,1120,1195,1221]" pageId="22" pageNumber="57" value="2002-09-22">
<collectingDate id="9B70D77AE9144A5AFC67FB6FFB9EFB4E" box="[972,1120,1195,1221]" pageId="22" pageNumber="57" value="2002-09-22">22. IX. 2002</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFBC6FB6FFB1CFB4E" box="[1133,1250,1195,1221]" pageId="22" pageNumber="57">S. Ohkusa</collectorName>
leg. (
<collectionCode id="999B9097E9144A5AFA8FFB6FFA6EFB4E" box="[1316,1424,1195,1221]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFF3CFB0BFB29FB62" box="[151,1239,1230,1257]" collectingDate="2007-08-20" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Yurano" municipality="Kumakogen" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Ehime" typeStatus="paratype">
[
<emphasis id="CDFED440E9144A5AFF0AFB0AFEFBFB63" bold="true" box="[161,261,1230,1256]" pageId="22" pageNumber="57">Shikoku</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9144A5AFEE8FB0BFE35FB62" box="[323,459,1231,1257]" country="Japan" name="Ehime" pageId="22" pageNumber="57">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9144A5AFE7EFB0BFD9FFB62" box="[469,609,1231,1257]" pageId="22" pageNumber="57">Kumakogen</collectingMunicipality>
,
<location id="FA555E89E9144A5AFDC1FB0BFD40FB62" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFDC1FB0BFD40FB62" box="[618,702,1231,1257]" country="Japan" municipality="Kumakogen" name="Yurano" pageId="22" pageNumber="57" stateProvince="Ehime">Yurano</location>
,
<date id="8B342E92E9144A5AFD63FB0BFC94FB62" box="[712,874,1231,1257]" pageId="22" pageNumber="57" value="2007-08-20">
<collectingDate id="9B70D77AE9144A5AFD63FB0BFC94FB62" box="[712,874,1231,1257]" pageId="22" pageNumber="57" value="2007-08-20">20. VIII. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFCDFFB0BFBF5FB62" box="[884,1035,1231,1257]" pageId="22" pageNumber="57">E. Yamamoto</collectorName>
leg., MsT (
<collectionCode id="999B9097E9144A5AFB21FB0BFB30FB62" box="[1162,1230,1231,1257]" pageId="22" pageNumber="57">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFB4AFB0BFBBDFA86" collectingDate="2006-07-15" collectionCode="EUM" collectorName="M. Nishikawa" country="Japan" location="Narukawa valley" municipality="Kihoku Town" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Ehime" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9144A5AFABBFB0BFA67FB62" box="[1296,1433,1231,1257]" country="Japan" name="Ehime" pageId="22" pageNumber="57">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9144A5AFF3CFB37FECBFA86" box="[151,309,1267,1293]" pageId="22" pageNumber="57">Kihoku Town</collectingMunicipality>
,
<location id="FA555E89E9144A5AFEEAFB37FDFCFA86" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFEEAFB37FDFCFA86" box="[321,514,1267,1293]" country="Japan" municipality="Kihoku Town" name="Narukawa valley" pageId="22" pageNumber="57" stateProvince="Ehime">Narukawa valley</location>
,
<date id="8B342E92E9144A5AFDA5FB36FD54FA86" box="[526,682,1266,1293]" pageId="22" pageNumber="57" value="2006-07-15">
<collectingDate id="9B70D77AE9144A5AFDA5FB36FD54FA86" box="[526,682,1266,1293]" pageId="22" pageNumber="57" value="2006-07-15">15. VII. 2006</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFD1DFB37FCA8FA86" box="[694,854,1267,1293]" pageId="22" pageNumber="57">M. Nishikawa</collectorName>
leg., at light (
<collectionCode id="999B9097E9144A5AFC5CFB37FBC5FA86" box="[1015,1083,1267,1293]" pageId="22" pageNumber="57">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFBE4FB37FC07FABA" collectingDate="1979-07-22" collectingDateMax="1979-07-23" collectingDateMin="1979-07-22" collectionCode="NARO" collectorName="I. Kanazawa" country="Japan" location="Mt. Ariake" municipality="Izuhara Town" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Nagasaki" typeStatus="paratype">
[
<emphasis id="CDFED440E9144A5AFBF2FB36FB11FA86" bold="true" box="[1113,1263,1266,1293]" pageId="22" pageNumber="57">Tsushima Is.</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9144A5AFA9FFB37FF2EFABA" country="Japan" name="Nagasaki" pageId="22" pageNumber="57">Nagasaki Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9144A5AFF71FAD3FE88FABA" box="[218,374,1303,1329]" pageId="22" pageNumber="57">Izuhara Town</collectingMunicipality>
,
<location id="FA555E89E9144A5AFE2BFAD3FE02FABA" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFE2BFAD3FE02FABA" box="[384,508,1303,1329]" country="Japan" municipality="Izuhara Town" name="Mt. Ariake" pageId="22" pageNumber="57" stateProvince="Nagasaki">Mt. Ariake</location>
,
<date id="8B342E92E9144A5AFDADFAD3FD36FABA" box="[518,712,1303,1329]" pageId="22" pageNumber="57" value="1979-07-22" valueMax="1979-07-23" valueMin="1979-07-22">
<collectingDate id="9B70D77AE9144A5AFDADFAD3FD36FABA" box="[518,712,1303,1329]" pageId="22" pageNumber="57" value="1979-07-22" valueMax="1979-07-23" valueMin="1979-07-22">22–23. VII. 1979</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFD78FAD3FCA0FABA" box="[723,862,1303,1329]" pageId="22" pageNumber="57">I. Kanazawa</collectorName>
leg. (
<collectionCode id="999B9097E9144A5AFC30FAD3FC0FFABA" box="[923,1009,1303,1329]" pageId="22" pageNumber="57">NARO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFBAFFAD3FBDEFADE" collectingDate="2007-06-04" collectionCode="KPMNH" collectorName="Nr. Santaro-toge &amp; K. Watanabe" country="Japan" location="Sumiyou Town" pageId="22" pageNumber="57" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
[
<emphasis id="CDFED440E9144A5AFBA6FAD2FB20FABA" bold="true" box="[1037,1246,1302,1329]" pageId="22" pageNumber="57">Amamioshima Is.</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9144A5AFAB6FAD3FF2EFADE" country="Japan" name="Kagoshima" pageId="22" pageNumber="57">Kagoshima Pref.</collectingRegion>
,
<location id="FA555E89E9144A5AFF71FAFFFE79FADE" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFF71FAFFFE79FADE" box="[218,391,1339,1365]" country="Japan" name="Sumiyou Town" pageId="22" pageNumber="57" stateProvince="Kagoshima">Sumiyou Town</location>
,
<collectorName id="527F6D84E9144A5AFE3AFAFFFDB2FADE" box="[401,588,1339,1365]" pageId="22" pageNumber="57">Nr. Santaro-toge</collectorName>
,
<date id="8B342E92E9144A5AFDFDFAFFFD28FADE" box="[598,726,1339,1365]" pageId="22" pageNumber="57" value="2007-06-04">
<collectingDate id="9B70D77AE9144A5AFDFDFAFFFD28FADE" box="[598,726,1339,1365]" pageId="22" pageNumber="57" value="2007-06-04">4. VI. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFD4BFAFFFC91FADE" box="[736,879,1339,1365]" pageId="22" pageNumber="57">K. Watanabe</collectorName>
leg. (
<collectionCode id="999B9097E9144A5AFC07FAFFFBE6FADE" box="[940,1048,1339,1365]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFB81FAFFFB72FAF2" collectingDate="2003-06-13" collectionCode="KPMNH, ZSM" collectorName="H. Makihara" country="Japan" location="Oku" municipality="Kunigami Vil." pageId="22" pageNumber="57" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
[
<emphasis id="CDFED440E9144A5AFB98FAFEFB07FADE" bold="true" box="[1075,1273,1338,1365]" pageId="22" pageNumber="57">Okinawajima Is.</emphasis>
] 2 F,
<collectingRegion id="3D4EC6B0E9144A5AFA9CFAFFFF2EFAF2" country="Japan" name="Okinawa" pageId="22" pageNumber="57">Okinawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9144A5AFF71FA9BFE82FAF2" box="[218,380,1375,1401]" pageId="22" pageNumber="57">Kunigami Vil.</collectingMunicipality>
,
<location id="FA555E89E9144A5AFE2CFA9BFE44FAF2" LSID="urn:lsid:plazi:treatment:7723B944E9144A55FF3CFC6BFD46FBD9:FA555E89E9144A5AFE2CFA9BFE44FAF2" box="[391,442,1375,1401]" country="Japan" municipality="Kunigami Vil." name="Oku" pageId="22" pageNumber="57" stateProvince="Okinawa">Oku</location>
,
<date id="8B342E92E9144A5AFE6EFA9BFDAAFAF2" box="[453,596,1375,1401]" pageId="22" pageNumber="57" value="2003-06-13">
<collectingDate id="9B70D77AE9144A5AFE6EFA9BFDAAFAF2" box="[453,596,1375,1401]" pageId="22" pageNumber="57" value="2003-06-13">13. VI. 2003</collectingDate>
</date>
,
<collectorName id="527F6D84E9144A5AFDF5FA9BFD15FAF2" box="[606,747,1375,1401]" pageId="22" pageNumber="57">H. Makihara</collectorName>
leg., MsT (1 F:
<collectionCode id="999B9097E9144A5AFC34FA9BFBF6FAF2" box="[927,1032,1375,1401]" pageId="22" pageNumber="57">KPMNH</collectionCode>
; 1 F:
<collectionCode id="999B9097E9144A5AFBEEFA9BFB7DFAF2" box="[1093,1155,1375,1401]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="22" pageNumber="57" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9144A5AFB3DFA9BFEEDFA16" collectingDate="2003-07-10" collectionCode="KPMNH" collectorName="H. Makihara" country="Japan" location="Oku" municipality="Kunigami Vil." pageId="22" pageNumber="57" specimenCount="1" stateProvince="Okinawajima Is." typeStatus="paratype">
1 F, ditto,
<date id="8B342E92E9144A5AFAACFA9BFA62FAF2" box="[1287,1436,1375,1401]" pageId="22" pageNumber="57" value="2003-07-10">
<collectingDate id="9B70D77AE9144A5AFAACFA9BFA62FAF2" box="[1287,1436,1375,1401]" pageId="22" pageNumber="57" value="2003-07-10">10. VII. 2003</collectingDate>
</date>
(
<collectionCode id="999B9097E9144A5AFF34FA47FEF5FA16" box="[159,267,1411,1437]" pageId="22" pageNumber="57">KPMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9144A55FF6CFA62FE37FD42" lastPageId="25" lastPageNumber="60" pageId="22" pageNumber="57" type="description">
<paragraph id="FF350852E9144A5AFF6CFA62FBF2FA6E" blockId="22.[151,1437,1050,2013]" pageId="22" pageNumber="57">
<emphasis id="CDFED440E9144A5AFF6CFA62FEAFFA4B" bold="true" box="[199,337,1446,1472]" pageId="22" pageNumber="57">Description</emphasis>
.
<emphasis id="CDFED440E9144A5AFECAFA63FE46FA4A" bold="true" box="[353,440,1447,1473]" pageId="22" pageNumber="57">Female</emphasis>
(n = 10). Body polished and covered with punctures and silvery setae; length 8.5–10.5 (HT: 10.4) mm. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E9144A5AFF6CFA2BFF21F8A2" blockId="22.[151,1437,1050,2013]" pageId="22" pageNumber="57">
Head 0.55 × as long as wide. Combined length of clypeus and face 1.05–1.15 (HT: 1.05) × facial width. Clypeus sparsely punctate ventrally, densely punctate dorsally, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.6–0.7 (HT: 0.7) × as long as basal mandibular width. Genal carina lamella-like ventrally, reaching mandibular base. POL 1.2–1.5 (HT: 1.3) × as long as OD. OOL 1.0–1.1 (HT: 1.0) × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9144A5AFE8DF903FE6AF96A" box="[294,404,1734,1761]" captionStart="FIGURE 15" captionStartId="23.[151,250,1925,1949]" captionTargetBox="[151,1436,381,1901]" captionTargetId="figure-122@23.[151,1436,381,1901]" captionTargetPageId="23" captionText="FIGURE 15. Syzeuctus nigrus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing." pageId="22" pageNumber="57">Fig. 15 D</figureCitation>
). Antenna with 38–40 (HT: 40) flagellomeres. FL I 4.0–4.25 (HT: 4.1) × as long as maximum depth in lateral view and 1.8–1.85 (HT: 1.85) × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E9144A5BFF6CF8F7FCF3FEEE" blockId="22.[151,1437,1050,2013]" lastBlockId="23.[151,1437,150,357]" lastPageId="23" lastPageNumber="58" pageId="22" pageNumber="57">
Mesosoma coarsely and densely punctate, 1.4–1.45 (HT: 1.45) × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate ventrally, sparsely punctate dorsally; speculum with a large smooth area, its lower area concave (
<figureCitation id="67B114D7E9144A5AFF34F85BFEF3F832" box="[159,269,1950,1977]" captionStart="FIGURE 15" captionStartId="23.[151,250,1925,1949]" captionTargetBox="[151,1436,381,1901]" captionTargetId="figure-122@23.[151,1436,381,1901]" captionTargetPageId="23" captionText="FIGURE 15. Syzeuctus nigrus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing." pageId="22" pageNumber="57">Fig. 15 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E9144A5AFDB9F807FD8EF856" box="[530,624,1987,2013]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="22" pageNumber="57">Fig. 1 H</figureCitation>
). Dorsal surface of propodeum with transverse rugae. Posterior transverse carina of propodeum present and complete (
<figureCitation id="67B114D7E9154A5BFD22FF53FD0BFF3A" box="[649,757,150,177]" captionStart="FIGURE 15" captionStartId="23.[151,250,1925,1949]" captionTargetBox="[151,1436,381,1901]" captionTargetId="figure-122@23.[151,1436,381,1901]" captionTargetPageId="23" captionText="FIGURE 15. Syzeuctus nigrus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing." pageId="23" pageNumber="58">Fig. 15 F</figureCitation>
). Fore wing length 7.0–8.0 (HT: 7.5) mm. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E9154A5BFB0EFF7FFAEBFF5E" box="[1189,1301,186,213]" captionStart="FIGURE 15" captionStartId="23.[151,250,1925,1949]" captionTargetBox="[151,1436,381,1901]" captionTargetId="figure-122@23.[151,1436,381,1901]" captionTargetPageId="23" captionText="FIGURE 15. Syzeuctus nigrus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing." pageId="23" pageNumber="58">Fig. 15 G</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.25–5.7 (HT: 5.4) × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.0–4.2 (HT: 4.2): 2.0: 1.35–1.4 (HT: 1.4): 0.6: 0.7–0.8 (HT: 0.7). Hind tarsal claws pectinate by short teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9154A5BFD02FE8FFD01FEEE" box="[681,767,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="23" pageNumber="58">Fig. 2 J</figureCitation>
).
</paragraph>
<caption id="ABF558DAE9154A5BFF3CF841FF2FF86E" pageId="23" pageNumber="58" startId="23.[151,250,1925,1949]" targetBox="[151,1436,381,1901]" targetPageId="23" targetType="figure">
<paragraph id="FF350852E9154A5BFF3CF841FF2FF86E" blockId="23.[151,1437,1925,2021]" pageId="23" pageNumber="58">
<emphasis id="CDFED440E9154A5BFF3CF841FEE1F816" bold="true" box="[151,287,1925,1949]" pageId="23" pageNumber="58">FIGURE 15.</emphasis>
<taxonomicName id="388A73D1E9154A5BFE8DF841FE31F817" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[294,463,1925,1948]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="58" phylum="Arthropoda" rank="species" species="nigrus" status="sp. nov.">
<emphasis id="CDFED440E9154A5BFE8DF841FE31F817" box="[294,463,1925,1948]" italics="true" pageId="23" pageNumber="58">Syzeuctus nigrus</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9154A5BFE7EF841FDDBF816" bold="true" box="[469,549,1925,1949]" pageId="23" pageNumber="58">
<taxonomicNameLabel id="D6CD693BE9154A5BFE7EF841FDDBF816" box="[469,549,1925,1949]" pageId="23" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing.
</paragraph>
</caption>
<caption id="ABF558DAE91A4A54FF3CF9B2FAB8F904" box="[151,1350,1654,1679]" pageId="24" pageNumber="59" startId="24.[151,250,1654,1678]" targetBox="[151,1436,151,1638]" targetPageId="24" targetType="figure">
<paragraph id="FF350852E91A4A54FF3CF9B2FAB8F904" blockId="24.[151,1350,1654,1679]" box="[151,1350,1654,1679]" pageId="24" pageNumber="59">
<emphasis id="CDFED440E91A4A54FF3CF9B2FEDEF905" bold="true" box="[151,288,1654,1678]" pageId="24" pageNumber="59">FIGURE 16.</emphasis>
<taxonomicName id="388A73D1E91A4A54FE8DF9B3FE2EF905" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[294,464,1655,1678]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrus" status="sp. nov.">
<emphasis id="CDFED440E91A4A54FE8DF9B3FE2EF905" box="[294,464,1655,1678]" italics="true" pageId="24" pageNumber="59">Syzeuctus nigrus</emphasis>
</taxonomicName>
<emphasis id="CDFED440E91A4A54FE7DF9B3FDD8F904" bold="true" box="[470,550,1655,1679]" pageId="24" pageNumber="59">
<taxonomicNameLabel id="D6CD693BE91A4A54FE7DF9B3FDD8F904" box="[470,550,1655,1679]" pageId="24" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view.
</paragraph>
</caption>
<paragraph id="FF350852E91A4A54FF6CF979FD4EF8C8" blockId="24.[151,1437,1725,2039]" pageId="24" pageNumber="59">Metasoma. T I 1.6–2.0 (HT: 2.0) × as long as maximum width, densely punctate laterally, largely smooth medially. T II 1.35–1.5 (HT: 1.5) × as long as maximum width T II and T III coriaceous with dense punctures. T IV to T V coriaceous with fine punctures. Ovipositor sheath 2.55–3.0 (HT: 2.75) × as long as hind tibia, ca. 1.1 × as long as fore wing, and shorter than body length.</paragraph>
<paragraph id="FF350852E91A4A55FF6CF889FC45FE45" blockId="24.[151,1437,1725,2039]" lastBlockId="25.[151,1437,151,1106]" lastPageId="25" lastPageNumber="60" pageId="24" pageNumber="59">
Colouration (
<figureCitation id="67B114D7E91A4A54FEF0F889FE10F8EC" box="[347,494,1869,1896]" captionStart="FIGURE 15" captionStartId="23.[151,250,1925,1949]" captionTargetBox="[151,1436,381,1901]" captionTargetId="figure-122@23.[151,1436,381,1901]" captionTargetPageId="23" captionText="FIGURE 15. Syzeuctus nigrus sp. nov., female, holotype—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: propodeum, dorsal view; G: fore wing." pageId="24" pageNumber="59">Figs 15 A–G</figureCitation>
). Body (excluding wings and legs) black. Lower part of clypeus, mandible except for teeth, malar space, W-shaped marking of face, inner orbit on frons and vertex, palpi, and outer orbit except for dorsal part yellow to ivory. Antenna blackish-brown. Collar and dorsal longitudinal stipe of pronotum yellow to ivory. A pair of antero-lateral markings and a median marking of mesoscutum, subtegular ridge, antero-dorsal and postero-ventral spots of mesopleuron, a pair of lateral spots of scutellum, postscutellum, posterior spot of metapleuron, and a posterior transverse marking of propodeum yellow to ivory. Metasomal tergites black except for a pair of basal spots of T I and posterior margins of T I to T III yellow to ivory. Posterior margins of T IV to T VII usually narrowly tinged with reddish-brown. Membranous parts of metasomal sternites ivory. Subgenital plate sometimes reddish-brown. Ovipositor reddish-brown. Wings hyaline. Apical clouded area of fore wing weakly present. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid coxae ivory apically, blackish-brown basally. Fore and mid trochanters ivory. Fore and mid legs except for coxae and trochanters reddish-brown to reddish-yellow. Mid trochantellus and tarsus more or less darkened. Hind leg blackish-brown to dark reddish-brown.Apices of coxa, trochanter, trochantellus, and femur tinged with yellowish-brown. Basal part of hind tibia and tibial spurs yellowish-brown. Apex of hind coxa sometimes with a yellow marking.
</paragraph>
<paragraph id="FF350852E91B4A55FF6CFE1CFE37FD42" blockId="25.[151,1437,151,1106]" pageId="25" pageNumber="60">
<emphasis id="CDFED440E91B4A55FF6CFE1CFEFAFE79" bold="true" box="[199,260,472,498]" pageId="25" pageNumber="60">Male</emphasis>
(n = 2). Similar to female (
<figureCitation id="67B114D7E91B4A55FD9CFE1CFD34FE79" box="[567,714,472,498]" captionStart="FIGURE 16" captionStartId="24.[151,250,1654,1678]" captionTargetBox="[151,1436,151,1638]" captionTargetId="figure-19@24.[151,1436,151,1638]" captionTargetPageId="24" captionText="FIGURE 16. Syzeuctus nigrus sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view." pageId="25" pageNumber="60">Figs 16 A–C</figureCitation>
). Malar space 0.8–0.9 × as long as basal mandibular width. POL 1.0–1.5 × as long as OD. OOL 1.0–1.2 × as long as OD. Antenna with 41–42 flagellomeres. FL I 3.55–4.5 × as long as maximum depth in lateral view and 1.6–1.7 × as long as FL II. Hind femur 6.0–6.1× as long as maximum depth in lateral view. Tergal spiracles not enlarged. T I 2.2–2.4 × as long as maximum width. T II 1.4–1.63 × as long as maximum width. Yellow area of body larger than female. Face yellow except for a small median longitudinal black stripe. Antenna reddish-brown apically. Fore and mid legs largely yellow. Yellow area of posterior margins of T I to T III narrower than female.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91B4A55FF6CFD16FE9EFC9B" pageId="25" pageNumber="60" type="distribution">
<paragraph id="FF350852E91B4A55FF6CFD16FE9EFC9B" blockId="25.[151,1437,151,1106]" pageId="25" pageNumber="60">
<emphasis id="CDFED440E91B4A55FF6CFD16FEA7FD67" bold="true" box="[199,345,722,748]" pageId="25" pageNumber="60">Distribution</emphasis>
.
<collectingCountry id="879D48C2E91B4A55FEC0FD17FE55FD66" box="[363,427,723,749]" name="Japan" pageId="25" pageNumber="60">Japan</collectingCountry>
(Honshu, Izuoshima Is., Awajishima Is., Shikoku, Tsushima Is., Amamioshima Is., and Okinawajima Is.).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91B4A55FF6CFCDEFE36FCDC" pageId="25" pageNumber="60" type="etymology">
<paragraph id="FF350852E91B4A55FF6CFCDEFCCFFCBF" blockId="25.[151,1437,151,1106]" box="[199,817,794,820]" pageId="25" pageNumber="60">
<emphasis id="CDFED440E91B4A55FF6CFCDEFEB6FCBF" bold="true" box="[199,328,794,820]" pageId="25" pageNumber="60">Etymology</emphasis>
. The specific name refers to its black body.
</paragraph>
<paragraph id="FF350852E91B4A55FF6CFCF9FE36FCDC" blockId="25.[151,1437,151,1106]" box="[199,456,829,856]" pageId="25" pageNumber="60">
<emphasis id="CDFED440E91B4A55FF6CFCF9FEBDFCDC" bold="true" box="[199,323,829,855]" pageId="25" pageNumber="60">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91B4A55FF6CFCA5FD46FBD9" pageId="25" pageNumber="60" type="discussion">
<paragraph id="FF350852E91B4A55FF6CFCA5FD46FBD9" blockId="25.[151,1437,151,1106]" pageId="25" pageNumber="60">
<emphasis id="CDFED440E91B4A55FF6CFCA5FECAFCF0" bold="true" box="[199,308,865,891]" pageId="25" pageNumber="60">Remarks</emphasis>
. This is the first record of this genus from Ryukyu Islands (Oriental part of
<collectingCountry id="879D48C2E91B4A55FBDBFCA5FB4BFCF0" box="[1136,1205,865,891]" name="Japan" pageId="25" pageNumber="60">Japan</collectingCountry>
). In the key provided by
<bibRefCitation id="9B1B75A3E91B4A55FF11FC41FEA6FC14" author="Riedel, M." box="[186,344,901,927]" pageId="25" pageNumber="60" pagination="1013 - 1098" refId="ref23284" refString="Riedel, M. (2021) The genus Syzeuctus Forster in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species. Linzer biologische Beitrage, 52 (2), 1013 - 1098." type="journal article" year="2021">Riedel (2021)</bibRefCitation>
, the female runs to couplet 27 (
<taxonomicName id="388A73D1E91B4A55FD17FC41FC12FC14" authority="(Panzer, 1809))" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" box="[700,1004,901,927]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FD17FC41FCCAFC14" box="[700,820,901,927]" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
(Panzer, 1809))
</taxonomicName>
, but differs by the mesopleuron with a postero-ventral yellow marking (absent in
<taxonomicName id="388A73D1E91B4A55FDD0FC6DFD0BFC48" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" box="[635,757,937,963]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FDD0FC6DFD0BFC48" box="[635,757,937,963]" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
</taxonomicName>
), the dorsal margin of pronotum with a longitudinal yellow marking (with a triangular yellow marking in
<taxonomicName id="388A73D1E91B4A55FD05FC09FCD7FC6C" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" box="[686,809,973,999]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FD05FC09FCD7FC6C" box="[686,809,973,999]" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
</taxonomicName>
), the apical clouded area of fore wing weaker than
<taxonomicName id="388A73D1E91B4A55FA2CFC09FF0DFB81" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FA2CFC09FF0DFB81" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
</taxonomicName>
, and the punctures of T II sparser than
<taxonomicName id="388A73D1E91B4A55FD05FC35FCD9FB81" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" box="[686,807,1008,1035]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FD05FC35FCD9FB81" box="[686,807,1008,1035]" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
</taxonomicName>
. The male of this species also resembles
<taxonomicName id="388A73D1E91B4A55FB52FC35FA8FFB81" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" box="[1273,1393,1008,1035]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FB52FC35FA8FFB81" box="[1273,1393,1008,1035]" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
</taxonomicName>
but can be distinguished by the shape of yellow markings of pronotum and mesopleuron as females and the yellow margins of T I to T IV narrower than
<taxonomicName id="388A73D1E91B4A55FD92FBFCFD4FFBD9" authorityName="" baseAuthorityName="Panzer" baseAuthorityYear="1809" box="[569,689,1080,1106]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="fuscator">
<emphasis id="CDFED440E91B4A55FD92FBFCFD4FFBD9" box="[569,689,1080,1106]" italics="true" pageId="25" pageNumber="60">S. fuscator</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

338
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<mods:title id="1747581A1EA7A75B60D18AD5E83C3686">Revision of the genus Syzeuctus Förster, 1869 (Hymenoptera, Ichneumonidae, Banchinae) from Japan</mods:title>
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<heading id="A47DBF3EE91B4A55FF3CFB60FDFBFB35" bold="true" box="[151,517,1187,1214]" fontSize="11" level="1" pageId="25" pageNumber="60" reason="1">
<emphasis id="CDFED440E91B4A55FF3CFB60FDFBFB35" bold="true" box="[151,517,1187,1214]" pageId="25" pageNumber="60">
<taxonomicName id="388A73D1E91B4A55FF3CFB60FE5EFB35" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[151,416,1187,1214]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="60" phylum="Arthropoda" rank="species" species="rufiapicalis" status="sp. nov.">
<emphasis id="CDFED440E91B4A55FF3CFB60FE5EFB35" bold="true" box="[151,416,1187,1214]" italics="true" pageId="25" pageNumber="60">Syzeuctus rufiapicalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6CD693BE91B4A55FE0CFB60FDFBFB35" box="[423,517,1188,1214]" pageId="25" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="FF350852E91B4A55FF3CFB0CFDC0FB69" blockId="25.[151,574,1187,1250]" box="[151,574,1224,1250]" pageId="25" pageNumber="60">
(
<figureCitation id="67B114D7E91B4A55FF34FB0CFF06FB69" box="[159,248,1224,1250]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="25" pageNumber="60">Figs 1 I</figureCitation>
,
<figureCitation id="67B114D7E91B4A55FEAFFB0CFE93FB69" box="[260,365,1224,1250]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="25" pageNumber="60">2 K, P, Q</figureCitation>
,
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,
<figureCitation id="67B114D7E91B4A55FE75FB0CFDCBFB69" box="[478,565,1224,1250]" captionStart="FIGURE 18" captionStartId="27.[151,246,746,770]" captionTargetBox="[151,1436,181,722]" captionTargetId="figure-19@27.[151,1436,181,722]" captionTargetPageId="27" captionText="FIGURE 18. Syzeuctus rufiapicalis sp. nov., male, paratype—A: lateral habitus; B: head, frontal view; C: T I and T II, lateral view." pageId="25" pageNumber="60">18 A–C</figureCitation>
)
</paragraph>
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<emphasis id="CDFED440E91B4A55FF3CFAD4FEE2FAA2" bold="true" box="[151,284,1295,1322]" pageId="25" pageNumber="60">Type series</emphasis>
.
<materialsCitation id="4FE2020FE91B4A55FE80FAD4FE2DFAC5" collectingDate="2022-07-19" collectingDateMax="2022-07-31" collectingDateMin="2022-07-19" collectionCode="KPMNH" collectorName="K. Esaki" country="Japan" county="Honshu" location="Mt. Kariyasu-yama" municipality="Kaga City" pageId="25" pageNumber="60" specimenCount="1" stateProvince="Ishikawa" typeStatus="holotype">
<emphasis id="CDFED440E91B4A55FE80FAD4FE66FAA1" bold="true" box="[299,408,1296,1322]" pageId="25" pageNumber="60">
<typeStatus id="2031B6F0E91B4A55FE80FAD4FE66FAA1" box="[299,408,1296,1322]" pageId="25" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
: F,
<emphasis id="CDFED440E91B4A55FE68FACBFDE3FAA2" bold="true" box="[451,541,1295,1321]" pageId="25" pageNumber="60">
<collectingCountry id="879D48C2E91B4A55FE68FACBFDE3FAA2" box="[451,541,1295,1321]" name="Japan" pageId="25" pageNumber="60">JAPAN</collectingCountry>
</emphasis>
,
<emphasis id="CDFED440E91B4A55FD80FAD4FD76FAA1" bold="true" box="[555,648,1296,1322]" pageId="25" pageNumber="60">
<collectingCounty id="165470DEE91B4A55FD80FAD4FD76FAA1" box="[555,648,1296,1322]" pageId="25" pageNumber="60">Honshu</collectingCounty>
</emphasis>
,
<collectingRegion id="3D4EC6B0E91B4A55FD3CFAD4FCC3FAA1" box="[663,829,1296,1322]" country="Japan" name="Ishikawa" pageId="25" pageNumber="60">Ishikawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91B4A55FCE3FAD4FC42FAA1" box="[840,956,1296,1322]" pageId="25" pageNumber="60">Kaga City</collectingMunicipality>
,
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,
<date id="8B342E92E91B4A55FB1AFAD4FA8BFAA1" box="[1201,1397,1296,1322]" pageId="25" pageNumber="60" value="2022-07-19" valueMax="2022-07-31" valueMin="2022-07-19">
<collectingDate id="9B70D77AE91B4A55FB1AFAD4FA8BFAA1" box="[1201,1397,1296,1322]" pageId="25" pageNumber="60" value="2022-07-19" valueMax="2022-07-31" valueMin="2022-07-19">19–31. VII. 2022</collectingDate>
</date>
,
<collectorName id="527F6D84E91B4A55FA2AFAD4FF2BFAC5" pageId="25" pageNumber="60">K. Esaki</collectorName>
leg., MsT (
<collectionCode id="999B9097E91B4A55FEF4FAF0FE35FAC5" box="[351,459,1332,1358]" pageId="25" pageNumber="60">KPMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE91B4A55FE4BFAF0FD07FAF9" collectingDate="2012-08-18" collectionCode="KPMNH" collectorName="Y. Matsubara &amp; K. Fukuda" country="Japan" location="Mamurogawa Town" pageId="25" pageNumber="60" specimenCount="1" stateProvince="Yamagata" typeStatus="paratype">
<emphasis id="CDFED440E91B4A55FE4BFAF0FDA6FAC5" bold="true" box="[480,600,1332,1358]" pageId="25" pageNumber="60">
<typeStatus id="2031B6F0E91B4A55FE4BFAF0FDA6FAC5" box="[480,600,1332,1358]" pageId="25" pageNumber="60" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="CDFED440E91B4A55FDC1FAF7FD3AFAC6" bold="true" box="[618,708,1331,1357]" pageId="25" pageNumber="60">
<collectingCountry id="879D48C2E91B4A55FDC1FAF7FD3AFAC6" box="[618,708,1331,1357]" name="Japan" pageId="25" pageNumber="60">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E91B4A55FD74FAF0FCC2FAC5" bold="true" box="[735,828,1332,1358]" pageId="25" pageNumber="60">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E91B4A55FC2FFAF0FBC8FAC5" box="[900,1078,1332,1358]" country="Japan" name="Yamagata" pageId="25" pageNumber="60">Yamagata Pref.</collectingRegion>
,
<location id="FA555E89E91B4A55FBE8FAF0FAD4FAC5" LSID="urn:lsid:plazi:treatment:7723B944E91B4A57FF3CFB60FAFEF865:FA555E89E91B4A55FBE8FAF0FAD4FAC5" box="[1091,1322,1332,1358]" country="Japan" name="Mamurogawa Town" pageId="25" pageNumber="60" stateProvince="Yamagata">Mamurogawa Town</location>
,
<date id="8B342E92E91B4A55FA92FAF0FF2CFAF9" pageId="25" pageNumber="60" value="2012-08-18">
<collectingDate id="9B70D77AE91B4A55FA92FAF0FF2CFAF9" pageId="25" pageNumber="60" value="2012-08-18">18. VIII. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E91B4A55FF70FA9CFE91FAF9" box="[219,367,1368,1394]" pageId="25" pageNumber="60">Y. Matsubara</collectorName>
&amp;
<collectorName id="527F6D84E91B4A55FE3AFA9CFDF9FAF9" box="[401,519,1368,1394]" pageId="25" pageNumber="60">K. Fukuda</collectorName>
leg., MsT (
<collectionCode id="999B9097E91B4A55FD2EFA9CFD0FFAF9" box="[645,753,1368,1394]" pageId="25" pageNumber="60">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91B4A55FCA8FA9CFE3FFA1D" collectingDate="1995-09-23" collectionCode="KPMNH" collectorName="M. Uchida" country="Japan" location="Kinshouzan" municipality="Ogawa Town" pageId="25" pageNumber="60" specimenCount="1" stateProvince="Saitama" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E91B4A55FC99FA9CFC35FAF9" box="[818,971,1368,1394]" country="Japan" name="Saitama" pageId="25" pageNumber="60">Saitama Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91B4A55FC7EFA9CFB92FAF9" box="[981,1132,1368,1394]" pageId="25" pageNumber="60">Ogawa Town</collectingMunicipality>
,
<location id="FA555E89E91B4A55FBDDFA9CFB01FAF9" LSID="urn:lsid:plazi:treatment:7723B944E91B4A57FF3CFB60FAFEF865:FA555E89E91B4A55FBDDFA9CFB01FAF9" box="[1142,1279,1368,1394]" country="Japan" municipality="Ogawa Town" name="Kinshouzan" pageId="25" pageNumber="60" stateProvince="Saitama">Kinshouzan</location>
,
<date id="8B342E92E91B4A55FAA2FA9CFA66FAF9" box="[1289,1432,1367,1394]" pageId="25" pageNumber="60" value="1995-09-23">
<collectingDate id="9B70D77AE91B4A55FAA2FA9CFA66FAF9" box="[1289,1432,1367,1394]" pageId="25" pageNumber="60" value="1995-09-23">23. IX. 1995</collectingDate>
</date>
,
<collectorName id="527F6D84E91B4A55FF3CFAB8FEF1FA1D" box="[151,271,1404,1430]" pageId="25" pageNumber="60">M. Uchida</collectorName>
leg. (
<collectionCode id="999B9097E91B4A55FEE6FAB8FE47FA1D" box="[333,441,1404,1430]" pageId="25" pageNumber="60">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91B4A55FE67FABFFB2DFA1D" box="[460,1235,1403,1430]" collectingDate="1995-09-23" collectionCode="KPMNH, ZSM, OMNH" collectorName="M. Uchida" country="Japan" location="Kinshouzan" municipality="Ogawa Town" pageId="25" pageNumber="60" specimenCount="1" stateProvince="Honshu" typeStatus="paratype">
15 F, same data of
<typeStatus id="2031B6F0E91B4A55FD35FAB8FCFEFA1D" box="[670,768,1404,1430]" pageId="25" pageNumber="60" type="holotype">holotype</typeStatus>
(10 F:
<collectionCode id="999B9097E91B4A55FCF9FAB8FC45FA1D" box="[850,955,1404,1430]" pageId="25" pageNumber="60">KPMNH</collectionCode>
; 4 F:
<collectionCode id="999B9097E91B4A55FC53FAB8FBCBFA1D" box="[1016,1077,1404,1430]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="25" pageNumber="60" type="Museum">ZSM</collectionCode>
; 1 F,
<collectionCode id="999B9097E91B4A55FBDAFAB8FB32FA1D" box="[1137,1228,1404,1430]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="25" pageNumber="60" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91B4A55FB75FAB8FDEFFA31" collectingDate="2002-06-28" collectingDateMax="2002-07-18" collectingDateMin="2002-06-28" collectionCode="KPMNH" collectorName="M. Uchida" country="Japan" location="Kinshouzan" municipality="Ogawa Town" pageId="25" pageNumber="60" specimenCount="1" stateProvince="Honshu" typeStatus="paratype">
1 F &amp; 1 M, ditto,
<date id="8B342E92E91B4A55FF3CFA64FE73FA31" box="[151,397,1440,1466]" pageId="25" pageNumber="60" value="2002-06-28" valueMax="2002-07-18" valueMin="2002-06-28">
<collectingDate id="9B70D77AE91B4A55FF3CFA64FE73FA31" box="[151,397,1440,1466]" pageId="25" pageNumber="60" value="2002-06-28" valueMax="2002-07-18" valueMin="2002-06-28">28. VI.–18. VII. 2002</collectingDate>
</date>
(
<collectionCode id="999B9097E91B4A55FE36FA64FDF7FA31" box="[413,521,1440,1466]" pageId="25" pageNumber="60">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91B4A55FDB6FA64FEA0FA55" collectingDate="2007-07-31" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Yurano" municipality="Kumakogen" pageId="25" pageNumber="60" specimenCount="1" stateProvince="Ehime" typeStatus="paratype">
[
<emphasis id="CDFED440E91B4A55FD8DFA5BFD74FA32" bold="true" box="[550,650,1439,1465]" pageId="25" pageNumber="60">Shikoku</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E91B4A55FD65FA64FCA6FA31" box="[718,856,1440,1466]" country="Japan" name="Ehime" pageId="25" pageNumber="60">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91B4A55FCCFFA64FC0EFA31" box="[868,1008,1440,1466]" pageId="25" pageNumber="60">Kumakogen</collectingMunicipality>
,
<location id="FA555E89E91B4A55FC50FA64FBB1FA31" LSID="urn:lsid:plazi:treatment:7723B944E91B4A57FF3CFB60FAFEF865:FA555E89E91B4A55FC50FA64FBB1FA31" box="[1019,1103,1440,1466]" country="Japan" municipality="Kumakogen" name="Yurano" pageId="25" pageNumber="60" stateProvince="Ehime">Yurano</location>
,
<date id="8B342E92E91B4A55FBF0FA64FB09FA31" box="[1115,1271,1440,1466]" pageId="25" pageNumber="60" value="2007-07-31">
<collectingDate id="9B70D77AE91B4A55FBF0FA64FB09FA31" box="[1115,1271,1440,1466]" pageId="25" pageNumber="60" value="2007-07-31">31. VII. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E91B4A55FAA8FA64FA63FA31" box="[1283,1437,1440,1466]" pageId="25" pageNumber="60">E. Yamamoto</collectorName>
leg., MsT (
<collectionCode id="999B9097E91B4A55FEB9FA00FEA8FA55" box="[274,342,1476,1502]" pageId="25" pageNumber="60">EUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91B4A57FF6CFA23FDB5F9E8" lastPageId="27" lastPageNumber="62" pageId="25" pageNumber="60" type="description">
<paragraph id="FF350852E91B4A55FF6CFA23FC00F9AD" blockId="25.[151,1437,1295,2042]" pageId="25" pageNumber="60">
<emphasis id="CDFED440E91B4A55FF6CFA23FEAFF98A" bold="true" box="[199,337,1511,1537]" pageId="25" pageNumber="60">Description</emphasis>
.
<emphasis id="CDFED440E91B4A55FECBFA2CFE49F989" bold="true" box="[352,439,1512,1538]" pageId="25" pageNumber="60">Female</emphasis>
(n = 20). Body polished and covered with punctures and silvery setae; length 9.25–12.2 (HT: 9.8) mm. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E91B4A55FF6CF9F4FA78F8CD" blockId="25.[151,1437,1295,2042]" pageId="25" pageNumber="60">
Head 0.55 × as long as wide. Combined length of clypeus and face 0.9–1.0 (HT: 0.9) × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, with a pair of indistinct convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.75–0.8 (HT: 0.75) × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.5–1.9 (HT: 1.8) × as long as OD. OOL 1.25–1.7 (HT: 1.6) × as long as OD. Temple slightly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E91B4A55FF77F8CCFEB6F8A9" box="[220,328,1800,1826]" captionStart="FIGURE 17" captionStartId="26.[151,250,1938,1962]" captionTargetBox="[151,1436,402,1922]" captionTargetId="figure-131@26.[151,1436,402,1922]" captionTargetPageId="26" captionText="FIGURE 17. Syzeuctus rufiapicalis sp. nov., females, holotype (A-C, E, F, H) and paratype (D, G)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F, G: pronotum and mesopleuron, lateral view; H: fore wing." pageId="25" pageNumber="60">Fig. 17 E</figureCitation>
). Antenna with 43–45 (HT: 45) flagellomeres. FL I 4.15–4.8 (HT: 4.3) × as long as maximum depth in lateral view and 1.6–1.8 (HT: 1.6) × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E91B4A56FF6CF894FDB3FE02" blockId="25.[151,1437,1295,2042]" lastBlockId="26.[151,1437,151,393]" lastPageId="26" lastPageNumber="61" pageId="25" pageNumber="60">
Mesosoma coarsely and densely punctate, 1.4–1.45 (HT: 1.4) × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate ventrally, sparsely punctate dorsally; speculum with a large smooth area, its ventral part with a rather deep concavity (
<figureCitation id="67B114D7E91B4A55FE3DF878FDDBF85D" box="[406,549,1980,2006]" captionStart="FIGURE 17" captionStartId="26.[151,250,1938,1962]" captionTargetBox="[151,1436,402,1922]" captionTargetId="figure-131@26.[151,1436,402,1922]" captionTargetPageId="26" captionText="FIGURE 17. Syzeuctus rufiapicalis sp. nov., females, holotype (A-C, E, F, H) and paratype (D, G)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F, G: pronotum and mesopleuron, lateral view; H: fore wing." pageId="25" pageNumber="60">Figs 17 F, G</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without or with angulation (
<figureCitation id="67B114D7E91B4A55FC2CF824FC20F871" box="[903,990,2016,2042]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="25" pageNumber="60">Fig. 1 I</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum largely absent medially, present laterally. Fore wing length 7.5–9.75 (HT: 7.6) mm. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole longer than areolet (
<figureCitation id="67B114D7E9184A56FE67FF1BFDC2FF72" box="[460,572,223,249]" captionStart="FIGURE 17" captionStartId="26.[151,250,1938,1962]" captionTargetBox="[151,1436,402,1922]" captionTargetId="figure-131@26.[151,1436,402,1922]" captionTargetPageId="26" captionText="FIGURE 17. Syzeuctus rufiapicalis sp. nov., females, holotype (A-C, E, F, H) and paratype (D, G)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F, G: pronotum and mesopleuron, lateral view; H: fore wing." pageId="26" pageNumber="61">Fig. 17 H</figureCitation>
). Fore wing vein 1cu-a slightly inclivous to subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 6.0–6.5 (HT: 6.5) × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.0–4.1 (HT: 4.0): 2.0: 1.3–1.45 (HT: 1.4): 0.6–0.75 (HT: 0.7): 0.8–0.95 (HT: 0.9). Hind tarsal claws simple except for minute subbasal teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9184A56FE74FEABFDBFFE02" box="[479,577,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="26" pageNumber="61">Fig. 2 K</figureCitation>
).
</paragraph>
<caption id="ABF558DAE9184A56FF3CF856FEA0F879" pageId="26" pageNumber="61" startId="26.[151,250,1938,1962]" targetBox="[151,1436,402,1922]" targetPageId="26" targetType="figure">
<paragraph id="FF350852E9184A56FF3CF856FEA0F879" blockId="26.[151,1436,1938,2034]" pageId="26" pageNumber="61">
<emphasis id="CDFED440E9184A56FF3CF856FEE1F821" bold="true" box="[151,287,1938,1962]" pageId="26" pageNumber="61">FIGURE 17.</emphasis>
<taxonomicName id="388A73D1E9184A56FE8FF856FDFFF821" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[292,513,1938,1962]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="61" phylum="Arthropoda" rank="species" species="rufiapicalis" status="sp. nov.">
<emphasis id="CDFED440E9184A56FE8FF856FDFFF821" box="[292,513,1938,1962]" italics="true" pageId="26" pageNumber="61">Syzeuctus rufiapicalis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9184A56FDACF856FDA8F821" bold="true" box="[519,598,1938,1962]" pageId="26" pageNumber="61">
<taxonomicNameLabel id="D6CD693BE9184A56FDACF856FDA8F821" box="[519,598,1938,1962]" pageId="26" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, females, holotype (A-C, E, F, H) and paratype (D, G)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F, G: pronotum and mesopleuron, lateral view; H: fore wing.
</paragraph>
</caption>
<caption id="ABF558DAE9194A57FF3CFD2EFA62FC89" box="[151,1436,746,770]" pageId="27" pageNumber="62" startId="27.[151,246,746,770]" targetBox="[151,1436,181,722]" targetPageId="27" targetType="figure">
<paragraph id="FF350852E9194A57FF3CFD2EFA62FC89" blockId="27.[151,1436,746,770]" box="[151,1436,746,770]" pageId="27" pageNumber="62">
<emphasis id="CDFED440E9194A57FF3CFD2EFEE4FC89" bold="true" box="[151,282,746,770]" pageId="27" pageNumber="62">FIGURE 18.</emphasis>
<taxonomicName id="388A73D1E9194A57FE8BFD2EFE0BFC89" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[288,501,746,770]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="rufiapicalis" status="sp. nov.">
<emphasis id="CDFED440E9194A57FE8BFD2EFE0BFC89" box="[288,501,746,770]" italics="true" pageId="27" pageNumber="62">Syzeuctus rufiapicalis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9194A57FE50FD2EFDB7FC89" bold="true" box="[507,585,746,770]" pageId="27" pageNumber="62">
<taxonomicNameLabel id="D6CD693BE9194A57FE50FD2EFDB7FC89" box="[507,585,746,770]" pageId="27" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, paratype—A: lateral habitus; B: head, frontal view; C: T I and T II, lateral view.
</paragraph>
</caption>
<paragraph id="FF350852E9194A57FF6CFCF5FF1BFC51" blockId="27.[151,1437,816,2031]" pageId="27" pageNumber="62">Metasoma. T I 2.55–2.95 (HT: 2.85) × as long as maximum width, finely punctate except for smooth areas on median longitudinal area, area along posterior margin and lateral sides. T II 1.85–2.1 (HT: 1.85) × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 1.95–2.05 (HT: 1.95) × as long as hind tibia, ca. 0.7–0.8 (HT: 0.75) × as long as fore wing, and shorter than body length.</paragraph>
<paragraph id="FF350852E9194A57FF6CFC20FE54FA58" blockId="27.[151,1437,816,2031]" pageId="27" pageNumber="62">
Colouration (
<figureCitation id="67B114D7E9194A57FEF6FC20FE0BFC75" box="[349,501,996,1022]" captionStart="FIGURE 17" captionStartId="26.[151,250,1938,1962]" captionTargetBox="[151,1436,402,1922]" captionTargetId="figure-131@26.[151,1436,402,1922]" captionTargetPageId="26" captionText="FIGURE 17. Syzeuctus rufiapicalis sp. nov., females, holotype (A-C, E, F, H) and paratype (D, G)—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C, D: head, frontal view; E: head, dorsal view; F, G: pronotum and mesopleuron, lateral view; H: fore wing." pageId="27" pageNumber="62">Figs 17 A–H</figureCitation>
). Body (excluding wings and legs) black. Clypeus, mandible except for teeth, face except for three longitudinal stripes and area around anterior tentorial pits, malar space, inner orbit on frons and vertex, and outer orbit yellow. Antenna blackish-brown to brown. Pronotum with a pair of transverse yellow stripes. Mesoscutum with a pair of antero-lateral yellow markings and a median yellow marking (the latter sometimes divided into a pair of markings). Subtegular ridge, antero-dorsal spot of mesopleuron, tegula, scutellum except a median small black spot, postscutellum, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of metapleuron and propodeum yellow. Mesopleuron sometimes (including
<typeStatus id="2031B6F0E9194A57FC4BFB78FBBAFB5D" box="[992,1092,1212,1238]" pageId="27" pageNumber="62" type="holotype">holotype</typeStatus>
) with an antero-median yellow marking. Upper division of metapleuron sometimes (including
<typeStatus id="2031B6F0E9194A57FCCBFB24FC3AFB71" box="[864,964,1248,1274]" pageId="27" pageNumber="62" type="holotype">holotype</typeStatus>
) yellow. Base and posterior margin of T I yellow. Posterior margins of T II and T III yellow. T IV to T VII largely reddish-yellow to red. Membranous parts of metasomal sternites largely yellow. Subgenital plate reddish-yellow. Ovipositor reddish-brown. Wings hyaline. Apical clouded area of fore wing slightly present. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow except for black area of mid coxa and black mid fifth tarsomere. Hind leg blackish-brown to black except for a dorsal small yellow spot of coxa, trochantellus, apex of femur, bases of tibia and first tarsomere.
</paragraph>
<paragraph id="FF350852E9194A57FF6CFA18FDB5F9E8" blockId="27.[151,1437,816,2031]" pageId="27" pageNumber="62">
<emphasis id="CDFED440E9194A57FF6CFA18FEFAFA7D" bold="true" box="[199,260,1500,1526]" pageId="27" pageNumber="62">Male</emphasis>
(n = 1). Similar to female (
<figureCitation id="67B114D7E9194A57FD92FA18FD33FA7D" box="[569,717,1500,1526]" captionStart="FIGURE 18" captionStartId="27.[151,246,746,770]" captionTargetBox="[151,1436,181,722]" captionTargetId="figure-19@27.[151,1436,181,722]" captionTargetPageId="27" captionText="FIGURE 18. Syzeuctus rufiapicalis sp. nov., male, paratype—A: lateral habitus; B: head, frontal view; C: T I and T II, lateral view." pageId="27" pageNumber="62">Figs 18 A–C</figureCitation>
). Malar space 0.9 × as long as basal mandibular width. Antenna with 46 flagellomeres. Tergal spiracles enlarged and with circular felt-like area, its length more than 0.2 × as long as T I (
<figureCitation id="67B114D7E9194A57FF43F9E0FE95F9B5" box="[232,363,1572,1599]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="27" pageNumber="62">Figs 2 P, Q</figureCitation>
,
<figureCitation id="67B114D7E9194A57FEDCF9E0FE2DF9B5" box="[375,467,1572,1598]" captionStart="FIGURE 18" captionStartId="27.[151,246,746,770]" captionTargetBox="[151,1436,181,722]" captionTargetId="figure-19@27.[151,1436,181,722]" captionTargetPageId="27" captionText="FIGURE 18. Syzeuctus rufiapicalis sp. nov., male, paratype—A: lateral habitus; B: head, frontal view; C: T I and T II, lateral view." pageId="27" pageNumber="62">18 A, C</figureCitation>
). Basal part of T I almost smooth (
<figureCitation id="67B114D7E9194A57FCC9F9E0FC38F9B5" box="[866,966,1572,1598]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="27" pageNumber="62">Fig. 2 Q</figureCitation>
). Yellow area of body larger than female. Mesosternum with a large yellow area.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9194A57FF6CF9A8FD4DF90D" box="[199,691,1644,1671]" pageId="27" pageNumber="62" type="distribution">
<paragraph id="FF350852E9194A57FF6CF9A8FD4DF90D" blockId="27.[151,1437,816,2031]" box="[199,691,1644,1671]" pageId="27" pageNumber="62">
<emphasis id="CDFED440E9194A57FF6CF9A8FEA7F90D" bold="true" box="[199,345,1644,1670]" pageId="27" pageNumber="62">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9194A57FECCF9A9FE59F90C" box="[359,423,1645,1671]" name="Japan" pageId="27" pageNumber="62">Japan</collectingCountry>
(Honshu and Shikoku).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9194A57FF6CF954FE36F945" pageId="27" pageNumber="62" type="etymology">
<paragraph id="FF350852E9194A57FF6CF954FBDEF921" blockId="27.[151,1437,816,2031]" box="[199,1056,1680,1707]" pageId="27" pageNumber="62">
<emphasis id="CDFED440E9194A57FF6CF954FEB6F921" bold="true" box="[199,328,1680,1706]" pageId="27" pageNumber="62">Etymology</emphasis>
. The specific name is from red apical part of metasomal tergites.
</paragraph>
<paragraph id="FF350852E9194A57FF6CF970FE36F945" blockId="27.[151,1437,816,2031]" box="[199,456,1716,1743]" pageId="27" pageNumber="62">
<emphasis id="CDFED440E9194A57FF6CF970FEBDF945" bold="true" box="[199,323,1716,1742]" pageId="27" pageNumber="62">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9194A57FF6CF91CFAFEF865" pageId="27" pageNumber="62" type="discussion">
<paragraph id="FF350852E9194A57FF6CF91CFAFEF865" blockId="27.[151,1437,816,2031]" pageId="27" pageNumber="62">
<emphasis id="CDFED440E9194A57FF6CF91CFECAF979" bold="true" box="[199,308,1752,1778]" pageId="27" pageNumber="62">Remarks</emphasis>
. In the key provided by
<bibRefCitation id="9B1B75A3E9194A57FDE0F91CFD17F979" author="Riedel, M." box="[587,745,1752,1778]" pageId="27" pageNumber="62" pagination="1013 - 1098" refId="ref23284" refString="Riedel, M. (2021) The genus Syzeuctus Forster in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species. Linzer biologische Beitrage, 52 (2), 1013 - 1098." type="journal article" year="2021">Riedel (2021)</bibRefCitation>
, the female runs to couplet 44 (
<taxonomicName id="388A73D1E9194A57FBFBF91CFA6EF979" authority="Riedel, 2021" authorityName="Riedel" authorityYear="2021" box="[1104,1424,1752,1778]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="CDFED440E9194A57FBFBF91CFB0AF979" box="[1104,1268,1752,1778]" italics="true" pageId="27" pageNumber="62">S. maroccanus</emphasis>
<bibRefCitation id="9B1B75A3E9194A57FB57F91CFA6EF979" author="Riedel, M." box="[1276,1424,1752,1778]" pageId="27" pageNumber="62" pagination="1013 - 1098" refId="ref23284" refString="Riedel, M. (2021) The genus Syzeuctus Forster in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species. Linzer biologische Beitrage, 52 (2), 1013 - 1098." type="journal article" year="2021">Riedel, 2021</bibRefCitation>
</taxonomicName>
), but differs by the T I and (at least) basal part of T II black (red in
<taxonomicName id="388A73D1E9194A57FC2CF938FBD2F89D" authorityName="Riedel" authorityYear="2021" box="[903,1068,1788,1814]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="CDFED440E9194A57FC2CF938FBD2F89D" box="[903,1068,1788,1814]" italics="true" pageId="27" pageNumber="62">S. maroccanus</emphasis>
</taxonomicName>
), the fore wing without a strong apical clouded area (with a strong brown apical spot in
<taxonomicName id="388A73D1E9194A57FCA2F8E4FC53F8B1" authorityName="Riedel" authorityYear="2021" box="[777,941,1824,1850]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="CDFED440E9194A57FCA2F8E4FC53F8B1" box="[777,941,1824,1850]" italics="true" pageId="27" pageNumber="62">S. maroccanus</emphasis>
</taxonomicName>
), and the malar space 0.75–0.8 × as long as basal mandibular width (0.5 × in
<taxonomicName id="388A73D1E9194A57FDA4F880FD4AF8D5" authorityName="Riedel" authorityYear="2021" box="[527,692,1860,1886]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="CDFED440E9194A57FDA4F880FD4AF8D5" box="[527,692,1860,1886]" italics="true" pageId="27" pageNumber="62">S. maroccanus</emphasis>
</taxonomicName>
). The male runs to couplet 7 (
<taxonomicName id="388A73D1E9194A57FBA1F880FAE8F8D5" authority="Seyrig, 1928" authorityName="Seyrig" authorityYear="1928" box="[1034,1302,1860,1886]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9194A57FBA1F880FB80F8D5" box="[1034,1150,1860,1886]" italics="true" pageId="27" pageNumber="62">S. dusmeti</emphasis>
Seyrig, 1928
</taxonomicName>
) of the key, but differs by the posterior transversal carina of propodeum more or less complete (absent medially in
<taxonomicName id="388A73D1E9194A57FAB4F8ACFA6DF809" authorityName="Seyrig" authorityYear="1928" box="[1311,1427,1896,1922]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9194A57FAB4F8ACFA6DF809" box="[1311,1427,1896,1922]" italics="true" pageId="27" pageNumber="62">S. dusmeti</emphasis>
</taxonomicName>
) and T II and T III without a conspicuous yellow posterior band (with yellow bands in
<taxonomicName id="388A73D1E9194A57FBE1F848FB42F82D" authorityName="Seyrig" authorityYear="1928" box="[1098,1212,1932,1958]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9194A57FBE1F848FB42F82D" box="[1098,1212,1932,1958]" italics="true" pageId="27" pageNumber="62">S. dusmeti</emphasis>
</taxonomicName>
). The female of this species also resembles
<taxonomicName id="388A73D1E9194A57FE34F874FD56F841" authority="Sheng, 2009" authorityName="Sheng" authorityYear="2009" box="[415,680,1968,1994]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="sparsus">
<emphasis id="CDFED440E9194A57FE34F874FDECF841" box="[415,530,1968,1994]" italics="true" pageId="27" pageNumber="62">S. sparsus</emphasis>
<bibRefCitation id="9B1B75A3E9194A57FDB1F874FD56F841" author="Sheng, M-L. &amp; Sun, S." box="[538,680,1968,1994]" pageId="27" pageNumber="62" refId="ref23401" refString="Sheng, M-L. &amp; Sun, S. (2009) Insect fauna of Henan, Hymenoptera: Ichneumonidae. Science Press, Beijing, 340 pp. [In Chinese with English summary]" type="book" year="2009">Sheng, 2009</bibRefCitation>
</taxonomicName>
but can be distinguished by the antenna with 43–45 flagellomeres (51–52 flagellomeres in
<taxonomicName id="388A73D1E9194A57FE01F810FDE2F865" authorityName="Sheng" authorityYear="2009" box="[426,540,2004,2030]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="sparsus">
<emphasis id="CDFED440E9194A57FE01F810FDE2F865" box="[426,540,2004,2030]" italics="true" pageId="27" pageNumber="62">S. sparsus</emphasis>
</taxonomicName>
) and a longer malar space (0.75–0.8 × versus 0.5 × in
<taxonomicName id="388A73D1E9194A57FBD5F810FB0EF865" authorityName="Sheng" authorityYear="2009" box="[1150,1264,2004,2030]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="62" phylum="Arthropoda" rank="species" species="sparsus">
<emphasis id="CDFED440E9194A57FBD5F810FB0EF865" box="[1150,1264,2004,2030]" italics="true" pageId="27" pageNumber="62">S. sparsus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="FF350852E91C4A52FF3CFB43FDB7FB2B" blockId="30.[151,585,1158,1221]" box="[151,585,1158,1185]" pageId="30" pageNumber="65">
<heading id="A47DBF3EE91C4A52FF3CFB43FDB7FB2B" bold="true" box="[151,585,1158,1185]" fontSize="11" level="1" pageId="30" pageNumber="65" reason="1">
<taxonomicName id="388A73D1E91C4A52FF3CFB43FDB7FB2B" ID-CoL="545JL" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[151,585,1158,1185]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="65" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E91C4A52FF3CFB43FDB7FB2B" bold="true" box="[151,585,1158,1185]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF3CFB43FE60FB2A" bold="true" box="[151,414,1158,1185]" italics="true" pageId="30" pageNumber="65">Syzeuctus takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E91C4A52FE0DFB42FDB7FB2B" author="Uchida, T." box="[422,585,1158,1184]" pageId="30" pageNumber="65" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</emphasis>
</taxonomicName>
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</subSubSection>
<subSubSection id="B7905BD9E91C4A52FF3CFB6FFE56FB4E" box="[151,424,1195,1221]" pageId="30" pageNumber="65" type="description">
<paragraph id="FF350852E91C4A52FF3CFB6FFE56FB4E" blockId="30.[151,585,1158,1221]" box="[151,424,1195,1221]" pageId="30" pageNumber="65">
(
<figureCitation id="67B114D7E91C4A52FF34FB6FFEFAFB4E" box="[159,260,1195,1221]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="30" pageNumber="65">Figs 1 K</figureCitation>
,
<figureCitation id="67B114D7E91C4A52FEA4FB6FFEC1FB4E" box="[271,319,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="30" pageNumber="65">2 M</figureCitation>
,
<figureCitation id="67B114D7E91C4A52FEE0FB6FFE5EFB4E" box="[331,416,1195,1221]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." pageId="30" pageNumber="65">21 A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91C4A52FF3CFB2BFDD0FA8C" box="[151,558,1263,1287]" pageId="30" pageNumber="65" type="reference_group">
<paragraph id="FF350852E91C4A52FF3CFB2BFDD0FA8C" blockId="30.[151,558,1263,1287]" box="[151,558,1263,1287]" pageId="30" pageNumber="65">
<treatmentCitationGroup id="DF9A2F7CE91C4A52FF3CFB2BFDD0FA8C" box="[151,558,1263,1287]" pageId="30" pageNumber="65">
<treatmentCitation id="7E2B2E43E91C4A52FF3CFB2BFDD4FA8C" author="Uchida, T." box="[151,554,1263,1287]" page="95" pageId="30" pageNumber="65" year="1928">
<taxonomicName id="388A73D1E91C4A52FF3CFB2BFDD4FA8C" ID-CoL="545JL" authority="Uchida, 1928: 95" authorityName="Uchida" authorityPageNumber="95" authorityYear="1928" box="[151,554,1263,1287]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="65" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E91C4A52FF3CFB2BFE8CFA8C" box="[151,370,1263,1287]" italics="true" pageId="30" pageNumber="65">Syzeuctus takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E91C4A52FED3FB2BFDD4FA8C" author="Uchida, T." box="[376,554,1263,1287]" pageId="30" pageNumber="65" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928: 95</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91C4A52FF3CFAF1FC28F988" pageId="30" pageNumber="65" type="materials_examined">
<paragraph id="FF350852E91C4A52FF3CFAF1FC28F988" blockId="30.[151,1437,1333,2008]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF3CFAF1FE79FAC4" bold="true" box="[151,391,1333,1359]" pageId="30" pageNumber="65">Materials examined</emphasis>
.
<materialsCitation id="4FE2020FE91C4A52FE3CFAF1FA7EFAC4" box="[407,1408,1333,1360]" collectingDate="1966-07-15" collectingDateMax="1966-07-18" collectingDateMin="1966-07-15" collectionCode="SEHU" collectorName="K. Kusigemati" country="Japan" location="Mt. Daisetsu" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hokkaido">
<emphasis id="CDFED440E91C4A52FE3CFAF1FE0FFAC4" bold="true" box="[407,497,1333,1359]" pageId="30" pageNumber="65">
<collectingCountry id="879D48C2E91C4A52FE3CFAF1FE0FFAC4" box="[407,497,1333,1359]" name="Japan" pageId="30" pageNumber="65">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E91C4A52FDA0FAF1FD7FFAC4" bold="true" box="[523,641,1333,1359]" pageId="30" pageNumber="65">
<collectingRegion id="3D4EC6B0E91C4A52FDA0FAF1FD7FFAC4" box="[523,641,1333,1359]" country="Japan" name="Hokkaido" pageId="30" pageNumber="65">Hokkaido</collectingRegion>
</emphasis>
] 1 F,
<location id="FA555E89E91C4A52FD62FAF2FCA3FAC4" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FD62FAF2FCA3FAC4" box="[713,861,1333,1360]" country="Japan" name="Mt. Daisetsu" pageId="30" pageNumber="65" stateProvince="Hokkaido">Mt. Daisetsu</location>
,
<date id="8B342E92E91C4A52FCC0FAF1FBCDFAC4" box="[875,1075,1333,1360]" pageId="30" pageNumber="65" value="1966-07-15" valueMax="1966-07-18" valueMin="1966-07-15">
<collectingDate id="9B70D77AE91C4A52FCC0FAF1FBCDFAC4" box="[875,1075,1333,1360]" pageId="30" pageNumber="65" value="1966-07-15" valueMax="1966-07-18" valueMin="1966-07-15">15–18. VII. 1966</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FBEBFAF2FB1BFAC4" box="[1088,1253,1333,1360]" pageId="30" pageNumber="65">K. Kusigemati</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FA82FAF1FA86FAC4" box="[1321,1400,1333,1359]" pageId="30" pageNumber="65">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FA25FAF1FC23FAF8" collectingDate="2012-07-16" collectionCode="KPMNH" collectorName="M. Ito" country="Japan" county="Horokanai Town" location="Uryu" municipality="Moshiri" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hokkaido">
1 F,
<collectingCounty id="165470DEE91C4A52FF1FFA9DFE8BFAFF" box="[180,373,1369,1396]" pageId="30" pageNumber="65">Horokanai Town</collectingCounty>
,
<collectingMunicipality id="1F519228E91C4A52FE29FA9DFE20FAF8" box="[386,478,1369,1395]" pageId="30" pageNumber="65">Moshiri</collectingMunicipality>
,
<location id="FA555E89E91C4A52FE40FA9EFDD9FAFF" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FE40FA9EFDD9FAFF" box="[491,551,1370,1396]" country="Japan" county="Horokanai Town" municipality="Moshiri" name="Uryu" pageId="30" pageNumber="65" stateProvince="Hokkaido">Uryu</location>
,
<date id="8B342E92E91C4A52FD9FFA9DFD2FFAF8" box="[564,721,1369,1396]" pageId="30" pageNumber="65" value="2012-07-16">
<collectingDate id="9B70D77AE91C4A52FD9FFA9DFD2FFAF8" box="[564,721,1369,1396]" pageId="30" pageNumber="65" value="2012-07-16">16. VII. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FD75FA9EFCD8FAFF" box="[734,806,1370,1396]" pageId="30" pageNumber="65">M. Ito</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FCC2FA9DFC2BFAF8" box="[873,981,1369,1395]" pageId="30" pageNumber="65">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FC42FA9DFDFBFA1C" collectingDate="2007-06-22" collectingDateMax="2007-07-10" collectingDateMin="2007-06-22" collectionCode="ZSM" collectorName="A. Ueda" country="Japan" location="Kuriyama Town" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hokkaido">
1 F,
<location id="FA555E89E91C4A52FBB5FA9DFB26FAFF" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FBB5FA9DFB26FAFF" box="[1054,1240,1369,1396]" country="Japan" name="Kuriyama Town" pageId="30" pageNumber="65" stateProvince="Hokkaido">Kuriyama Town</location>
,
<date id="8B342E92E91C4A52FB4EFA9DFF2CFA1C" pageId="30" pageNumber="65" value="2007-06-22" valueMax="2007-07-10" valueMin="2007-06-22">
<collectingDate id="9B70D77AE91C4A52FB4EFA9DFF2CFA1C" pageId="30" pageNumber="65" value="2007-06-22" valueMax="2007-07-10" valueMin="2007-06-22">22. VI.–10. VII. 2007</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FF77FABAFEC4FA1C" box="[220,314,1405,1432]" pageId="30" pageNumber="65">A. Ueda</collectorName>
leg., MsT (
<collectionCode id="999B9097E91C4A52FE16FAB9FE02FA1C" box="[445,508,1405,1431]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="30" pageNumber="65" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FDBAFABAFF08FA30" collectingDate="1926-07-30" collectionCode="SEHU" collectorName="S. Hirayama" country="Japan" location="Mt. Takaosan" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Tokyo" typeStatus="lectotype">
[
<emphasis id="CDFED440E91C4A52FDB1FAB9FD89FA1C" bold="true" box="[538,631,1405,1431]" pageId="30" pageNumber="65">Honshu</emphasis>
] 1 F (
<typeStatus id="2031B6F0E91C4A52FD16FAB9FCD4FA1C" box="[701,810,1405,1431]" pageId="30" pageNumber="65" type="lectotype">lectotype</typeStatus>
),
<collectingRegion id="3D4EC6B0E91C4A52FC96FAB9FC79FA1C" box="[829,903,1405,1431]" country="Japan" name="Tokyo" pageId="30" pageNumber="65">Tokyo</collectingRegion>
,
<location id="FA555E89E91C4A52FC39FABAFBD3FA1C" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FC39FABAFBD3FA1C" box="[914,1069,1405,1432]" country="Japan" name="Mt. Takaosan" pageId="30" pageNumber="65" stateProvince="Tokyo">Mt. Takaosan</location>
,
<date id="8B342E92E91C4A52FB92FAB9FB2DFA1C" box="[1081,1235,1405,1432]" pageId="30" pageNumber="65" value="1926-07-30">
<collectingDate id="9B70D77AE91C4A52FB92FAB9FB2DFA1C" box="[1081,1235,1405,1432]" pageId="30" pageNumber="65" value="1926-07-30">30. VII. 1926</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FB74FAB9FA95FA1C" box="[1247,1387,1405,1431]" pageId="30" pageNumber="65">S. Hirayama</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FF34FA65FF10FA30" box="[159,238,1441,1467]" pageId="30" pageNumber="65">SEHU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FEA9FA65FB30FA30" box="[258,1230,1441,1468]" collectingDate="2004-05-26" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" location="Enkaizan" municipality="Yokohama City" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Kanagawa" typeStatus="lectotype">
1 F,
<collectingRegion id="3D4EC6B0E91C4A52FE9FFA66FE15FA30" box="[308,491,1441,1468]" country="Japan" name="Kanagawa" pageId="30" pageNumber="65">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91C4A52FE5EFA65FD58FA30" box="[501,678,1441,1467]" pageId="30" pageNumber="65">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E91C4A52FD19FA65FCE3FA30" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FD19FA65FCE3FA30" box="[690,797,1441,1467]" country="Japan" municipality="Yokohama City" name="Enkaizan" pageId="30" pageNumber="65" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E91C4A52FC82FA65FC53FA30" box="[809,941,1441,1468]" pageId="30" pageNumber="65" value="2004-05-26">
<collectingDate id="9B70D77AE91C4A52FC82FA65FC53FA30" box="[809,941,1441,1468]" pageId="30" pageNumber="65" value="2004-05-26">26. V. 2004</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FC12FA66FBE4FA30" box="[953,1050,1441,1468]" pageId="30" pageNumber="65">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FBF1FA65FB38FA30" box="[1114,1222,1441,1467]" pageId="30" pageNumber="65">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FB71FA65FC7BFA54" collectingDate="1951-05-22" collectionCode="MNHAH" collectorName="A. Nagatomi" country="Japan" location="Sakazukiyama" municipality="Tamba" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Hyogo" typeStatus="lectotype">
1 F,
<collectingRegion id="3D4EC6B0E91C4A52FAA7FA66FA66FA30" box="[1292,1432,1441,1468]" country="Japan" name="Hyogo" pageId="30" pageNumber="65">Hyogo Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91C4A52FF3CFA01FF18FA54" box="[151,230,1477,1503]" pageId="30" pageNumber="65">Tamba</collectingMunicipality>
,
<location id="FA555E89E91C4A52FF58FA01FE67FA54" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FF58FA01FE67FA54" box="[243,409,1477,1503]" country="Japan" municipality="Tamba" name="Sakazukiyama" pageId="30" pageNumber="65" stateProvince="Hyogo">Sakazukiyama</location>
,
<date id="8B342E92E91C4A52FE0DFA01FDD5FA6B" box="[422,555,1477,1504]" pageId="30" pageNumber="65" value="1951-05-22">
<collectingDate id="9B70D77AE91C4A52FE0DFA01FDD5FA6B" box="[422,555,1477,1504]" pageId="30" pageNumber="65" value="1951-05-22">22. V. 1951</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FD9CFA02FD37FA54" box="[567,713,1477,1504]" pageId="30" pageNumber="65">A. Nagatomi</collectorName>
leg. (
<collectionCode id="999B9097E91C4A52FCA7FA01FC83FA54" box="[780,893,1477,1503]" pageId="30" pageNumber="65">MNHAH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91C4A52FC3AFA02FC2CF988" collectingDate="2014-05-21" collectingDateMax="2014-05-25" collectingDateMin="2014-05-21" collectionCode="EUM" collectorName="E. Yamamoto" country="Japan" location="Matsuyamasougou-park" municipality="Matsuyama City" pageId="30" pageNumber="65" specimenCount="1" stateProvince="Ehime" typeStatus="lectotype">
[
<emphasis id="CDFED440E91C4A52FC30FA01FBF6FA54" bold="true" box="[923,1032,1477,1503]" pageId="30" pageNumber="65">Shikoku]</emphasis>
1 F,
<collectingRegion id="3D4EC6B0E91C4A52FBEFFA01FB31FA54" box="[1092,1231,1477,1503]" country="Japan" name="Ehime" pageId="30" pageNumber="65">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E91C4A52FB70FA02FA66FA54" box="[1243,1432,1477,1504]" pageId="30" pageNumber="65">Matsuyama City</collectingMunicipality>
,
<location id="FA555E89E91C4A52FF3CFA2DFE57F988" LSID="urn:lsid:plazi:treatment:7723B944E91C4A6CFF3CFB43FD1BFDEA:FA555E89E91C4A52FF3CFA2DFE57F988" box="[151,425,1513,1539]" country="Japan" municipality="Matsuyama City" name="Matsuyamasougou-park" pageId="30" pageNumber="65" stateProvince="Ehime">Matsuyamasougou-park</location>
,
<date id="8B342E92E91C4A52FE1FFA2DFD9FF988" box="[436,609,1513,1540]" pageId="30" pageNumber="65" value="2014-05-21" valueMax="2014-05-25" valueMin="2014-05-21">
<collectingDate id="9B70D77AE91C4A52FE1FFA2DFD9FF988" box="[436,609,1513,1540]" pageId="30" pageNumber="65" value="2014-05-21" valueMax="2014-05-25" valueMin="2014-05-21">21–25. V. 2014</collectingDate>
</date>
,
<collectorName id="527F6D84E91C4A52FDC7FA2EFCFAF98F" box="[620,772,1514,1540]" pageId="30" pageNumber="65">E. Yamamoto</collectorName>
leg., MsT (
<collectionCode id="999B9097E91C4A52FC2EFA2DFC37F988" box="[901,969,1513,1539]" pageId="30" pageNumber="65">EUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91C4A6CFF6CF9C9FDD9FE26" lastPageId="32" lastPageNumber="67" pageId="30" pageNumber="65" type="description">
<paragraph id="FF350852E91C4A52FF6CF9C9FCABF9C7" blockId="30.[151,1437,1333,2008]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF6CF9C9FEAFF9AC" bold="true" box="[199,337,1549,1575]" pageId="30" pageNumber="65">Description</emphasis>
.
<emphasis id="CDFED440E91C4A52FEF5F9C9FE4BF9AC" bold="true" box="[350,437,1549,1575]" pageId="30" pageNumber="65">Female</emphasis>
(n = 6). Body polished and covered with punctures and silvery setae; length 9.0–
<quantity id="3872A5B7E91C4A52FA9FF9C9FA67F9A3" box="[1332,1433,1549,1576]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="30" pageNumber="65" unit="mm" value="10.2">10.2 mm</quantity>
. Setae on head and mesosoma not exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E91C4A52FF6CF991FDF3F8E0" blockId="30.[151,1437,1333,2008]" pageId="30" pageNumber="65">
Head 0.6 × as long as wide. Combined length of clypeus and face 1.05–1.1 × facial width. Clypeus smooth except for dorsal margin punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.4 × as long as basal mandibular width. Genal carina low ventrally, reaching mandibular base. POL 1.65–1.9 × as long as OD. OOL 1.4–1.5 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E91C4A52FC22F8CDFC05F8A8" box="[905,1019,1801,1827]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." pageId="30" pageNumber="65">Fig. 21 D</figureCitation>
). Antenna with 38–42 flagellomeres. FL I 3.8–4.4 × as long as maximum depth in lateral view and 1.55–1.75 × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E91C4A53FF6CF8B2FE45FE02" blockId="30.[151,1437,1333,2008]" lastBlockId="31.[151,1437,151,393]" lastPageId="31" lastPageNumber="66" pageId="30" pageNumber="65">
Mesosoma coarsely and densely punctate, 1.4 × as long as maximum height. Pronotum with rugulose area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate; speculum with a large smooth area (
<figureCitation id="67B114D7E91C4A52FD9AF879FD61F85C" box="[561,671,1981,2007]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." pageId="30" pageNumber="65">Fig. 21 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end remote from anterior margin of mesopleuron, without angulation (
<figureCitation id="67B114D7E91D4A53FC60FF53FBCDFF3A" box="[971,1075,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="31" pageNumber="66">Fig. 1 K</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum present and complete. Fore wing length
<quantity id="3872A5B7E91D4A53FAE3FF7FFF38FF72" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="8.9" metricValueMin="6.9" pageId="31" pageNumber="66" unit="mm" value="7.9" valueMax="8.9" valueMin="6.9">6.9–8.9 mm</quantity>
. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole shorter than areolet (
<figureCitation id="67B114D7E91D4A53FF34FEC7FEF3FE96" box="[159,269,259,285]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." pageId="31" pageNumber="66">Fig. 21 F</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.1–5.9 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 3.9–4.1: 2.0: 1.4: 0.7: 0.8. Hind tarsal claws pectinate by short teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E91D4A53FEE2FEABFE51FE02" box="[329,431,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="31" pageNumber="66">Fig. 2 M</figureCitation>
).
</paragraph>
<caption id="ABF558DAE91D4A53FF3CF933FAFEF8B8" pageId="31" pageNumber="66" startId="31.[151,250,1783,1807]" targetBox="[151,1436,417,1759]" targetPageId="31" targetType="figure">
<paragraph id="FF350852E91D4A53FF3CF933FAFEF8B8" blockId="31.[151,1437,1783,1843]" pageId="31" pageNumber="66">
<emphasis id="CDFED440E91D4A53FF3CF933FEDEF884" bold="true" box="[151,288,1783,1807]" pageId="31" pageNumber="66">FIGURE 21.</emphasis>
<taxonomicName id="388A73D1E91D4A53FE8CF933FD6DF884" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[295,659,1783,1807]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="66" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E91D4A53FE8CF933FDFCF884" box="[295,514,1783,1807]" italics="true" pageId="31" pageNumber="66">Syzeuctus takaozanus</emphasis>
<bibRefCitation id="9B1B75A3E91D4A53FDA3F933FD6DF884" author="Uchida, T." box="[520,659,1783,1807]" pageId="31" pageNumber="66" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings.
</paragraph>
</caption>
<paragraph id="FF350852E91D4A53FF6CF8A6FBE1F86C" blockId="31.[151,1436,1889,2024]" pageId="31" pageNumber="66">Metasoma. T I 1.9–1.95 × as long as maximum width, finely punctate except for smooth areas on median longitudinal area, area along posterior margin and lateral sides. T II 1.25–1.4 × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine punctures. Ovipositor sheath 3.35–3.6 × as long as hind tibia, ca. 1.4 × as long as fore wing, and as long as or slightly longer than body length.</paragraph>
<paragraph id="FF350852E9224A6CFF6CFF53FCABFE02" blockId="32.[151,1437,151,609]" pageId="32" pageNumber="67">
Colouration (
<figureCitation id="67B114D7E9224A6CFEF4FF53FE08FF3A" box="[351,502,151,177]" captionStart="FIGURE 21" captionStartId="31.[151,250,1783,1807]" captionTargetBox="[151,1436,417,1759]" captionTargetId="figure-129@31.[151,1436,417,1759]" captionTargetPageId="31" captionText="FIGURE 21. Syzeuctus takaozanus Uchida, 1928, female from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: wings." pageId="32" pageNumber="67">Figs 21 A–F</figureCitation>
). Body (excluding wings and legs) black. Clypeus yellow except for dorsal part, mandible except for base and teeth, ventral part of pedicel, and tegula yellow. Ventral part of scape and basal segments of flagellum tinged with brown to reddish-brown. Fore and mid legs yellow except for largely black coxae and partly darkened mid trochanter. Hind leg black to blackish-brown except for trochantellus, base and apex of femur, and basal part of tibia yellow to yellowish-brown. Membranous parts of metasomal sternites largely dark yellowish-brown. Ovipositor reddish-brown. Wings hyaline, without distinct apical clouded area of fore wing. Veins and pterostigma brown except for yellowish-brown wing base.
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFE57FDD9FE26" blockId="32.[151,1437,151,609]" box="[199,551,403,429]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE57FEFAFE26" bold="true" box="[199,260,403,429]" pageId="32" pageNumber="67">Male</emphasis>
. Unknown (see remarks).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF6CFE72FE36FE7E" pageId="32" pageNumber="67" type="distribution">
<paragraph id="FF350852E9224A6CFF6CFE72FA95FE5A" blockId="32.[151,1437,151,609]" box="[199,1387,438,465]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE72FEA7FE5B" bold="true" box="[199,345,438,464]" pageId="32" pageNumber="67">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9224A6CFECCFE73FE59FE5A" box="[359,423,439,465]" name="Japan" pageId="32" pageNumber="67">Japan</collectingCountry>
(
<collectingRegion id="3D4EC6B0E9224A6CFE1DFE73FDD7FE5A" box="[438,553,439,465]" country="Japan" name="Hokkaido" pageId="32" pageNumber="67">Hokkaido</collectingRegion>
, Honshu, and Shikoku),
<collectingCountry id="879D48C2E9224A6CFC96FE73FC7EFE5A" box="[829,896,439,465]" name="China" pageId="32" pageNumber="67">China</collectingCountry>
and
<collectingCountry id="879D48C2E9224A6CFC1DFE73FC00FE5A" box="[950,1022,439,465]" name="South Korea" pageId="32" pageNumber="67">Korea</collectingCountry>
. New to Hokkaido and Shikoku.
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFE1EFE36FE7E" blockId="32.[151,1437,151,609]" box="[199,456,474,501]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE1EFEBDFE7F" bold="true" box="[199,323,474,500]" pageId="32" pageNumber="67">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF6CFE3BFD1BFDEA" pageId="32" pageNumber="67" type="discussion">
<paragraph id="FF350852E9224A6CFF6CFE3BFD1BFDEA" blockId="32.[151,1437,151,609]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFE3BFECAFD92" bold="true" box="[199,308,511,537]" pageId="32" pageNumber="67">Remarks</emphasis>
.
<bibRefCitation id="9B1B75A3E9224A6CFEEEFE3BFE2FFD92" author="Iwata, K." box="[325,465,510,537]" pageId="32" pageNumber="67" pagination="63 - 74" refId="ref23060" refString="Iwata, K. (1958) Ovarian eggs of 233 species of the Japanese Ichneumonidae (Hymenoptera). Acta Hymenopterologica, 1 (1), 63 - 74." type="journal article" year="1958">Iwata (1958</bibRefCitation>
,
<bibRefCitation id="9B1B75A3E9224A6CFE74FE3BFDE6FD92" author="Iwata, K." box="[479,536,511,537]" pageId="32" pageNumber="67" pagination="115 - 169" refId="ref23092" refString="Iwata, K. (1960) The comparative anatomy of the ovary in Hymenoptera. Part V. Ichneumonidae. Acta Hymenopterologica, 1 (2), 115 - 169." type="journal article" year="1960">1960</bibRefCitation>
) described the ovarian eggs and ovary.
<bibRefCitation id="9B1B75A3E9224A6CFC4DFE3BFB45FD92" author="Kang, G-W. &amp; Kolarov, J. &amp; Lee, J. - W." box="[998,1211,511,537]" pageId="32" pageNumber="67" pagination="17 - 30" refId="ref23125" refString="Kang, G-W., Kolarov, J. &amp; Lee, J. - W. (2020) Review of the genus Syzeuctus (Hymenoptera, Ichneumonidae, Banchinae) from South Korea. Journal of Hymenoptera Research, 80, 17 - 30. https: // doi. org / 10.3897 / jhr. 80.57667" type="journal article" year="2020">
Kang
<emphasis id="CDFED440E9224A6CFB87FE3BFB99FD92" box="[1068,1127,511,537]" italics="true" pageId="32" pageNumber="67">et al.</emphasis>
(2020)
</bibRefCitation>
described the male of this species, but it may be misidentified because of the clouded apex of fore wing in his male in contrast to the entirely hyaline wing in the female of
<taxonomicName id="388A73D1E9224A6CFDEFFD83FD20FDEA" authorityName="Uchida" authorityYear="1928" box="[580,734,583,609]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="67" phylum="Arthropoda" rank="species" species="takaozanus">
<emphasis id="CDFED440E9224A6CFDEFFD83FD20FDEA" box="[580,734,583,609]" italics="true" pageId="32" pageNumber="67">S. takaozanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="FF350852E91E4A50FF3CF933FDCCF89A" blockId="28.[151,562,1783,1846]" box="[151,562,1783,1810]" pageId="28" pageNumber="63">
<heading id="A47DBF3EE91E4A50FF3CF933FDCCF89A" bold="true" box="[151,562,1783,1810]" fontSize="11" level="1" pageId="28" pageNumber="63" reason="1">
<taxonomicName id="388A73D1E91E4A50FF3CF933FDCCF89A" ID-CoL="545JC" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[151,562,1783,1810]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="63" phylum="Arthropoda" rank="species" species="sambonis">
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<emphasis id="CDFED440E91E4A50FF3CF933FE79F899" bold="true" box="[151,391,1783,1810]" italics="true" pageId="28" pageNumber="63">Syzeuctus sambonis</emphasis>
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<paragraph id="FF350852E91E4A50FF3CF8DFFE02F8BE" blockId="28.[151,562,1783,1846]" box="[151,508,1819,1846]" pageId="28" pageNumber="63">
(
<figureCitation id="67B114D7E91E4A50FF34F8DFFF04F8BD" box="[159,250,1819,1846]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="28" pageNumber="63">Figs 1 J</figureCitation>
,
<figureCitation id="67B114D7E91E4A50FEADF8DFFED0F8BD" box="[262,302,1819,1846]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="28" pageNumber="63">2 L</figureCitation>
,
<figureCitation id="67B114D7E91E4A50FE91F8DFFE6FF8BE" box="[314,401,1819,1845]" captionStart="FIGURE 19" captionStartId="28.[151,250,1641,1665]" captionTargetBox="[151,1436,181,1617]" captionTargetId="figure-19@28.[151,1436,181,1617]" captionTargetPageId="28" captionText="FIGURE 19. Syzeuctus sambonis Uchida, 1928, females from Japan—A: lateral habitus; B: head, mesosoma, wings, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="28" pageNumber="63">19 A–E</figureCitation>
,
<figureCitation id="67B114D7E91E4A50FE36F8DFFE0AF8BE" box="[413,500,1819,1845]" captionStart="FIGURE 20" captionStartId="29.[151,250,1767,1791]" captionTargetBox="[151,1436,777,1742]" captionTargetId="figure-310@29.[151,1436,777,1743]" captionTargetPageId="29" captionText="FIGURE 20. Syzeuctus sambonis Uchida, 1928, males from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view." pageId="28" pageNumber="63">20 A–C</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="DF9A2F7CE91E4A50FF3CF8A4FDE2F8F3" box="[151,540,1888,1912]" pageId="28" pageNumber="63">
<treatmentCitation id="7E2B2E43E91E4A50FF3CF8A4FDE6F8F3" author="Uchida, T." box="[151,536,1888,1912]" page="94" pageId="28" pageNumber="63" year="1928">
<taxonomicName id="388A73D1E91E4A50FF3CF8A4FDE6F8F3" ID-CoL="545JC" authority="Uchida, 1928: 94" authorityName="Uchida" authorityPageNumber="94" authorityYear="1928" box="[151,536,1888,1912]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="63" phylum="Arthropoda" rank="species" species="sambonis">
<emphasis id="CDFED440E91E4A50FF3CF8A4FE9EF8F3" box="[151,352,1888,1912]" italics="true" pageId="28" pageNumber="63">Syzeuctus sambonis</emphasis>
<bibRefCitation id="9B1B75A3E91E4A50FECDF8A4FDE6F8F3" author="Uchida, T." box="[358,536,1888,1912]" pageId="28" pageNumber="63" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928: 94</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91E4A51FF3CF861FD04FE02" lastPageId="29" lastPageNumber="64" pageId="28" pageNumber="63" type="materials_examined">
<paragraph id="FF350852E91E4A50FF3CF861FA62F86F" blockId="28.[151,1436,1957,2021]" pageId="28" pageNumber="63">
<emphasis id="CDFED440E91E4A50FF3CF861FE7CF834" bold="true" box="[151,386,1957,1983]" pageId="28" pageNumber="63">
<collectingRegion id="3D4EC6B0E91E4A50FF3CF861FEF4F834" box="[151,266,1957,1983]" pageId="28" pageNumber="63">Materials</collectingRegion>
examined
</emphasis>
.
<materialsCitation id="4FE2020FE91E4A50FE26F861FA66F86F" collectingDate="1962-07-25" collectingDateMax="1981-07-26" collectingDateMin="1962-07-25" collectionCode="KPMNH" collectorName="H. Nagase &amp; H. Suda" country="Japan" location="Kamakura" pageId="28" pageNumber="63" specimenCount="1" stateProvince="Kanagawa">
<emphasis id="CDFED440E91E4A50FE26F861FE19F834" bold="true" box="[397,487,1957,1983]" pageId="28" pageNumber="63">
<collectingCountry id="879D48C2E91E4A50FE26F861FE19F834" box="[397,487,1957,1983]" name="Japan" pageId="28" pageNumber="63">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E91E4A50FE56F862FDA4F84B" bold="true" box="[509,602,1958,1984]" pageId="28" pageNumber="63">Honshu</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E91E4A50FD34F862FCADF84B" box="[671,851,1958,1984]" country="Japan" name="Kanagawa" pageId="28" pageNumber="63">Kanagawa Pref.</collectingRegion>
,
<location id="FA555E89E91E4A50FCF0F862FC2DF84B" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91E4A50FCF0F862FC2DF84B" box="[859,979,1958,1984]" country="Japan" name="Kamakura" pageId="28" pageNumber="63" stateProvince="Kanagawa">Kamakura</location>
,
<date id="8B342E92E91E4A50FC70F862FB91F84B" box="[987,1135,1958,1984]" pageId="28" pageNumber="63" value="1981-07-26">
<collectingDate id="9B70D77AE91E4A50FC70F862FB91F84B" box="[987,1135,1958,1984]" pageId="28" pageNumber="63" value="1981-07-26">26. VII. 1981</collectingDate>
</date>
,
<collectorName id="527F6D84E91E4A50FBD3F862FB14F84B" box="[1144,1258,1958,1984]" pageId="28" pageNumber="63">H. Nagase</collectorName>
leg. (
<collectionCode id="999B9097E91E4A50FA8FF862FA6EF84B" box="[1316,1424,1958,1984]" pageId="28" pageNumber="63">KPMNH</collectionCode>
); 1 F,
<collectingRegion id="3D4EC6B0E91E4A50FF61F80EFEB0F86F" box="[202,334,1994,2020]" country="Japan" name="Chiba" pageId="28" pageNumber="63">Chiba Pref.</collectingRegion>
, Kimitsu City, Minekami Vil., Toyooka,
<date id="8B342E92E91E4A50FCB6F80EFC47F86F" box="[797,953,1994,2021]" pageId="28" pageNumber="63" value="1962-07-25">
<collectingDate id="9B70D77AE91E4A50FCB6F80EFC47F86F" box="[797,953,1994,2021]" pageId="28" pageNumber="63" value="1962-07-25">25. VII. 1962</collectingDate>
</date>
,
<collectorName id="527F6D84E91E4A50FC6EF80EFBDEF86F" box="[965,1056,1994,2020]" pageId="28" pageNumber="63">H. Suda</collectorName>
leg. (
<collectionCode id="999B9097E91E4A50FBC9F80EFB30F86F" box="[1122,1230,1994,2020]" pageId="28" pageNumber="63">KPMNH</collectionCode>
); 1 F,
<collectingRegion id="3D4EC6B0E91E4A50FABFF80EFA66F86F" box="[1300,1432,1994,2020]" country="Japan" name="Chiba" pageId="28" pageNumber="63">Chiba Pref.</collectingRegion>
</materialsCitation>
,
</paragraph>
<paragraph id="FF350852E91F4A51FF3CFF53FD04FE02" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="64">
<materialsCitation id="4FE2020FE91F4A51FF3CFF53FC61FF3A" box="[151,927,151,177]" collectingDate="2002-07-04" collectionCode="KPMNH" collectorName="H. Suda" location="Ichikawa City" municipality="Kitakokubun" pageId="29" pageNumber="64" specimenCount="1">
<location id="FA555E89E91F4A51FF3CFF53FEC7FF3A" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FF3CFF53FEC7FF3A" box="[151,313,151,177]" municipality="Kitakokubun" name="Ichikawa City" pageId="29" pageNumber="64">Ichikawa City</location>
,
<collectingMunicipality id="1F519228E91F4A51FEECFF53FE22FF3A" box="[327,476,151,177]" pageId="29" pageNumber="64">Kitakokubun</collectingMunicipality>
,
<date id="8B342E92E91F4A51FE41FF53FD85FF3A" box="[490,635,151,177]" pageId="29" pageNumber="64" value="2002-07-04">
<collectingDate id="9B70D77AE91F4A51FE41FF53FD85FF3A" box="[490,635,151,177]" pageId="29" pageNumber="64" value="2002-07-04">4. VII. 2002</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FD22FF53FD18FF3A" box="[649,742,151,177]" pageId="29" pageNumber="64">H. Suda</collectorName>
leg. (
<collectionCode id="999B9097E91F4A51FC80FF53FC69FF3A" box="[811,919,151,177]" pageId="29" pageNumber="64">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FC07FF53FE26FF5E" collectingDate="2011-06-30" collectionCode="ZSM" collectorName="M. Ito" county="Nakamachi" location="Nara Pref." municipality="Nara City" pageId="29" pageNumber="64" specimenCount="1">
1 F,
<location id="FA555E89E91F4A51FC49FF53FBA2FF3A" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FC49FF53FBA2FF3A" box="[994,1116,151,177]" county="Nakamachi" municipality="Nara City" name="Nara Pref." pageId="29" pageNumber="64">Nara Pref.</location>
,
<collectingMunicipality id="1F519228E91F4A51FBC2FF53FB24FF3A" box="[1129,1242,151,177]" pageId="29" pageNumber="64">Nara City</collectingMunicipality>
,
<collectingCounty id="165470DEE91F4A51FB42FF53FA95FF3A" box="[1257,1387,151,177]" pageId="29" pageNumber="64">Nakamachi</collectingCounty>
,
<date id="8B342E92E91F4A51FAD2FF53FF03FF5E" pageId="29" pageNumber="64" value="2011-06-30">
<collectingDate id="9B70D77AE91F4A51FAD2FF53FF03FF5E" pageId="29" pageNumber="64" value="2011-06-30">30. VI. 2011</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FEA2FF7FFEAEFF5E" box="[265,336,187,213]" pageId="29" pageNumber="64">M. Ito</collectorName>
leg. (
<collectionCode id="999B9097E91F4A51FE3BFF7FFE31FF5E" box="[400,463,187,213]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="29" pageNumber="64" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FE4FFF7FFEFDFF72" collectingDate="2003-07-06" collectionCode="OMNH" collectorName="O. Tominaga" county="Tonomine-kogen" location="Hyogo Pref." municipality="Okawachi Town" pageId="29" pageNumber="64" specimenCount="1">
1 F,
<location id="FA555E89E91F4A51FDBEFF7FFD5CFF5E" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FDBEFF7FFD5CFF5E" box="[533,674,187,213]" county="Tonomine-kogen" municipality="Okawachi Town" name="Hyogo Pref." pageId="29" pageNumber="64">Hyogo Pref.</location>
,
<collectingMunicipality id="1F519228E91F4A51FD05FF7FFC96FF5E" box="[686,872,187,213]" pageId="29" pageNumber="64">Okawachi Town</collectingMunicipality>
,
<collectingCounty id="165470DEE91F4A51FCDFFF7FFBC8FF5E" box="[884,1078,187,213]" pageId="29" pageNumber="64">Tonomine-kogen</collectingCounty>
,
<date id="8B342E92E91F4A51FBEAFF7FFB30FF5E" box="[1089,1230,187,213]" pageId="29" pageNumber="64" value="2003-07-06">
<collectingDate id="9B70D77AE91F4A51FBEAFF7FFB30FF5E" box="[1089,1230,187,213]" pageId="29" pageNumber="64" value="2003-07-06">6. VII. 2003</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FB72FF7FFA92FF5E" box="[1241,1388,187,213]" pageId="29" pageNumber="64">O. Tominaga</collectorName>
leg. (
<collectionCode id="999B9097E91F4A51FF34FF1BFF05FF72" box="[159,251,223,249]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="29" pageNumber="64" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FEBBFF1BFBCDFF72" box="[272,1075,223,249]" collectingDate="1968-08-07" collectionCode="EUM" collectorName="K. Hatta" location="Hyogo Pref." municipality="Mt. Hyonosen" pageId="29" pageNumber="64" specimenCount="1">
1 M,
<location id="FA555E89E91F4A51FEFBFF1BFE20FF72" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FEFBFF1BFE20FF72" box="[336,478,223,249]" municipality="Mt. Hyonosen" name="Hyogo Pref." pageId="29" pageNumber="64">Hyogo Pref.</location>
,
<collectingMunicipality id="1F519228E91F4A51FE40FF1BFD6EFF72" box="[491,656,223,249]" pageId="29" pageNumber="64">Mt. Hyonosen</collectingMunicipality>
,
<date id="8B342E92E91F4A51FD36FF1BFCC8FF72" box="[669,822,223,249]" pageId="29" pageNumber="64" value="1968-08-07">
<collectingDate id="9B70D77AE91F4A51FD36FF1BFCC8FF72" box="[669,822,223,249]" pageId="29" pageNumber="64" value="1968-08-07">7. VIII. 1968</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FCE8FF1BFC5AFF72" box="[835,932,223,249]" pageId="29" pageNumber="64">K. Hatta</collectorName>
leg. (
<collectionCode id="999B9097E91F4A51FC4CFF1BFBD5FF72" box="[999,1067,223,249]" pageId="29" pageNumber="64">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FBEBFF1BFD9DFE96" collectingDate="1953-07-17" collectionCode="MNHAH" collectorName="Taniguchi" location="Hyogo Pref." municipality="Sekinomiya" pageId="29" pageNumber="64" specimenCount="1">
1 F,
<location id="FA555E89E91F4A51FBDEFF1BFAFDFF72" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FBDEFF1BFAFDFF72" box="[1141,1283,223,249]" municipality="Sekinomiya" name="Hyogo Pref." pageId="29" pageNumber="64">Hyogo Pref.</location>
,
<collectingMunicipality id="1F519228E91F4A51FAA4FF1BFA66FF72" box="[1295,1432,223,249]" pageId="29" pageNumber="64">Sekinomiya</collectingMunicipality>
,
<date id="8B342E92E91F4A51FF3CFEC7FECFFE96" box="[151,305,258,285]" pageId="29" pageNumber="64" value="1953-07-17">
<collectingDate id="9B70D77AE91F4A51FF3CFEC7FECFFE96" box="[151,305,258,285]" pageId="29" pageNumber="64" value="1953-07-17">17. VII. 1953</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FE90FEC7FE55FE96" box="[315,427,259,285]" pageId="29" pageNumber="64">Taniguchi</collectorName>
leg. (
<collectionCode id="999B9097E91F4A51FE41FEC7FDA5FE96" box="[490,603,259,285]" pageId="29" pageNumber="64">MNHAH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FDC5FEC7FE03FECA" collectingDate="2008-07-30" collectionCode="KPMNH" collectorName="Miyakejima Is. &amp; H. Makihara" location="Tokyo" municipality="Tsubota" pageId="29" pageNumber="64" specimenCount="1">
[
<emphasis id="CDFED440E91F4A51FDDCFEC7FD2BFE96" bold="true" box="[631,725,259,285]" pageId="29" pageNumber="64">Izu Isls.</emphasis>
] 1 F &amp; 1 M,
<location id="FA555E89E91F4A51FCC2FEC7FC4DFE96" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FCC2FEC7FC4DFE96" box="[873,947,259,285]" municipality="Tsubota" name="Tokyo" pageId="29" pageNumber="64">Tokyo</location>
,
<collectorName id="527F6D84E91F4A51FC14FEC7FB94FE96" box="[959,1130,259,285]" pageId="29" pageNumber="64">Miyakejima Is.</collectorName>
,
<collectingMunicipality id="1F519228E91F4A51FBDDFEC7FB2FFE96" box="[1142,1233,259,285]" pageId="29" pageNumber="64">Tsubota</collectingMunicipality>
,
<date id="8B342E92E91F4A51FB77FEC7FA88FE96" box="[1244,1398,259,285]" pageId="29" pageNumber="64" value="2008-07-30">
<collectingDate id="9B70D77AE91F4A51FB77FEC7FA88FE96" box="[1244,1398,259,285]" pageId="29" pageNumber="64" value="2008-07-30">30. VII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FA2AFEC7FEFCFECA" pageId="29" pageNumber="64">H. Makihara</collectorName>
leg., MsT, (
<collectionCode id="999B9097E91F4A51FE22FEE3FE0BFECA" box="[393,501,295,321]" pageId="29" pageNumber="64">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FDA3FEE3FF15FEEE" collectingDate="2008-07-07" collectingDateMax="2008-07-30" collectingDateMin="2008-07-07" collectionCode="EUM" collectorName="Miyakejima Is. &amp; H. Makihara" location="Tokyo" municipality="Ako" pageId="29" pageNumber="64" specimenCount="1">
1 F &amp; 1 M,
<location id="FA555E89E91F4A51FD27FEE3FD28FECA" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FD27FEE3FD28FECA" box="[652,726,295,321]" municipality="Ako" name="Tokyo" pageId="29" pageNumber="64">Tokyo</location>
,
<collectorName id="527F6D84E91F4A51FD49FEE3FC73FECA" box="[738,909,295,321]" pageId="29" pageNumber="64">Miyakejima Is.</collectorName>
,
<collectingMunicipality id="1F519228E91F4A51FC33FEE3FC35FECA" box="[920,971,295,321]" pageId="29" pageNumber="64">Ako</collectingMunicipality>
,
<date id="8B342E92E91F4A51FC7DFEE3FB75FECA" box="[982,1163,295,321]" pageId="29" pageNumber="64" value="2008-07-07" valueMax="2008-07-30" valueMin="2008-07-07">
<collectingDate id="9B70D77AE91F4A51FC7DFEE3FB75FECA" box="[982,1163,295,321]" pageId="29" pageNumber="64" value="2008-07-07" valueMax="2008-07-30" valueMin="2008-07-07">7–30. VII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FB3DFEE3FADAFECA" box="[1174,1316,295,321]" pageId="29" pageNumber="64">H. Makihara</collectorName>
leg., MsT, (
<collectionCode id="999B9097E91F4A51FF34FE8FFF1DFEEE" box="[159,227,331,357]" pageId="29" pageNumber="64">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE91F4A51FF5DFE8FFB0FFEEE" box="[246,1265,330,357]" collectingDate="2005-08-15" collectionCode="EUM" collectorName="E. Yamamoto" county="Yurano" location="Ehime Pref." municipality="Kumakogen" pageId="29" pageNumber="64" specimenCount="1">
[
<emphasis id="CDFED440E91F4A51FF54FE8EFE9DFEEF" bold="true" box="[255,355,330,356]" pageId="29" pageNumber="64">Shikoku</emphasis>
] 1 F,
<location id="FA555E89E91F4A51FE0AFE8FFDD7FEEE" LSID="urn:lsid:plazi:treatment:7723B944E91E4A52FF3CF933FC3BFBBE:FA555E89E91F4A51FE0AFE8FFDD7FEEE" box="[417,553,331,357]" county="Yurano" municipality="Kumakogen" name="Ehime Pref." pageId="29" pageNumber="64">Ehime Pref.</location>
,
<collectingMunicipality id="1F519228E91F4A51FD98FE8FFD41FEEE" box="[563,703,331,357]" pageId="29" pageNumber="64">Kumakogen</collectingMunicipality>
,
<collectingCounty id="165470DEE91F4A51FD63FE8FFCE2FEEE" box="[712,796,331,357]" pageId="29" pageNumber="64">Yurano</collectingCounty>
,
<date id="8B342E92E91F4A51FC8DFE8EFC38FEEE" box="[806,966,330,357]" pageId="29" pageNumber="64" value="2005-08-15">
<collectingDate id="9B70D77AE91F4A51FC8DFE8EFC38FEEE" box="[806,966,330,357]" pageId="29" pageNumber="64" value="2005-08-15">15. VIII. 2005</collectingDate>
</date>
,
<collectorName id="527F6D84E91F4A51FC7AFE8FFB96FEEE" box="[977,1128,331,357]" pageId="29" pageNumber="64">E. Yamamoto</collectorName>
leg. (
<collectionCode id="999B9097E91F4A51FB0EFE8FFB17FEEE" box="[1189,1257,331,357]" pageId="29" pageNumber="64">EUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE91F4A51FB50FE8EFD08FE02" collectionCode="SEHU" country="South Korea" location="KOREA" pageId="29" pageNumber="64" specimenCount="1" typeStatus="holotype">
<emphasis id="CDFED440E91F4A51FB50FE8EFA9CFEEF" bold="true" box="[1275,1378,330,356]" pageId="29" pageNumber="64">
<collectingCountry id="879D48C2E91F4A51FB50FE8EFA9CFEEF" box="[1275,1378,330,356]" name="South Korea" pageId="29" pageNumber="64">KOREA</collectingCountry>
</emphasis>
: 1 M (
<typeStatus id="2031B6F0E91F4A51FF34FEABFEF9FE02" box="[159,263,367,393]" pageId="29" pageNumber="64" type="holotype">holotype</typeStatus>
), “Chosen Takano Uchida”, “27/7” (
<collectionCode id="999B9097E91F4A51FD34FEABFD10FE02" box="[671,750,367,393]" pageId="29" pageNumber="64">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91F4A52FF6CFE56FDF2FC42" lastPageId="30" lastPageNumber="65" pageId="29" pageNumber="64" type="description">
<paragraph id="FF350852E91F4A51FF6CFE56FC9AFE5A" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="64">
<emphasis id="CDFED440E91F4A51FF6CFE56FEAFFE27" bold="true" box="[199,337,402,428]" pageId="29" pageNumber="64">Description</emphasis>
.
<emphasis id="CDFED440E91F4A51FECAFE57FE46FE26" bold="true" box="[353,440,403,429]" pageId="29" pageNumber="64">Female</emphasis>
(n = 8). Body polished and covered with punctures and silvery setae; length
<quantity id="3872A5B7E91F4A51FA86FE56FF38FE5A" metricMagnitude="-2" metricUnit="m" metricValue="1.285" metricValueMax="1.42" metricValueMin="1.15" pageId="29" pageNumber="64" unit="mm" value="12.85" valueMax="14.2" valueMin="11.5">11.5–14.2 mm</quantity>
. Setae on head and mesosoma exceptionally long and erect.
</paragraph>
<paragraph id="FF350852E91F4A51FF6CFE1FFD5CFD7A" blockId="29.[151,1437,151,753]" pageId="29" pageNumber="64">
Head 0.55 × as long as wide. Combined length of clypeus and face 1.05–1.1 × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.3–0.4 × as long as basal mandibular width. Genal carina lamella-like ventrally, reaching mandibular base. POL 1.5–1.7 × as long as OD. OOL 1.3–1.6 × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E91F4A51FB80FD4BFB61FD22" box="[1067,1183,655,681]" captionStart="FIGURE 19" captionStartId="28.[151,250,1641,1665]" captionTargetBox="[151,1436,181,1617]" captionTargetId="figure-19@28.[151,1436,181,1617]" captionTargetPageId="28" captionText="FIGURE 19. Syzeuctus sambonis Uchida, 1928, females from Japan—A: lateral habitus; B: head, mesosoma, wings, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="29" pageNumber="64">Fig. 19 D</figureCitation>
). Antenna with 49–53 flagellomeres. FL I 5.0–5.2 × as long as maximum depth in lateral view and 1.9–2.0 × as long as FL II. Preapical flagellomere 1.2 × as long as maximum width.
</paragraph>
<caption id="ABF558DAE91F4A51FF3CF923FE08F8A8" pageId="29" pageNumber="64" startId="29.[151,250,1767,1791]" targetBox="[151,1436,777,1742]" targetPageId="29" targetType="figure">
<paragraph id="FF350852E91F4A51FF3CF923FE08F8A8" blockId="29.[151,1437,1767,1827]" pageId="29" pageNumber="64">
<emphasis id="CDFED440E91F4A51FF3CF923FEDCF974" bold="true" box="[151,290,1767,1791]" pageId="29" pageNumber="64">FIGURE 20.</emphasis>
<taxonomicName id="388A73D1E91F4A51FE81F923FD74F974" authority="Uchida, 1928" authorityName="Uchida" authorityYear="1928" box="[298,650,1767,1791]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="sambonis">
<emphasis id="CDFED440E91F4A51FE81F923FE0BF974" box="[298,501,1767,1791]" italics="true" pageId="29" pageNumber="64">Syzeuctus sambonis</emphasis>
<bibRefCitation id="9B1B75A3E91F4A51FE56F923FD74F974" author="Uchida, T." box="[509,650,1767,1791]" pageId="29" pageNumber="64" pagination="1 - 115" refId="ref23517" refString="Uchida, T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University, 25, 1 - 115." type="journal article" year="1928">Uchida, 1928</bibRefCitation>
</taxonomicName>
, males from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view.
</paragraph>
</caption>
<paragraph id="FF350852E91F4A52FF6CF896FC3CFECA" blockId="29.[151,1437,1873,2044]" lastBlockId="30.[151,1437,151,1077]" lastPageId="30" lastPageNumber="65" pageId="29" pageNumber="64">
Mesosoma coarsely and densely punctate, 1.4 × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate; speculum with a large smooth area (
<figureCitation id="67B114D7E91F4A51FD9AF85DFD61F838" box="[561,671,1945,1971]" captionStart="FIGURE 19" captionStartId="28.[151,250,1641,1665]" captionTargetBox="[151,1436,181,1617]" captionTargetId="figure-19@28.[151,1436,181,1617]" captionTargetPageId="28" captionText="FIGURE 19. Syzeuctus sambonis Uchida, 1928, females from Japan—A: lateral habitus; B: head, mesosoma, wings, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="29" pageNumber="64">Fig. 19 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally, its dorsal end close to anterior margin of mesopleuron, with angulation (
<figureCitation id="67B114D7E91F4A51FCDFF879FC2FF85C" box="[884,977,1981,2007]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="29" pageNumber="64">Fig. 1 J</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum present and complete. Fore wing length
<quantity id="3872A5B7E91F4A51FAABF825FA67F877" box="[1280,1433,2017,2044]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" metricValueMax="10.5" metricValueMin="9.3" pageId="29" pageNumber="64" unit="mm" value="9.9" valueMax="10.5" valueMin="9.3">9.3–10.5 mm</quantity>
. Areolet receiving vein 2m-cu basal to the outer angle, petiolated anteriorly; petiole slightly shorter than areolet (
<figureCitation id="67B114D7E91C4A52FF34FF7FFEF0FF5E" box="[159,270,187,213]" captionStart="FIGURE 19" captionStartId="28.[151,250,1641,1665]" captionTargetBox="[151,1436,181,1617]" captionTargetId="figure-19@28.[151,1436,181,1617]" captionTargetPageId="28" captionText="FIGURE 19. Syzeuctus sambonis Uchida, 1928, females from Japan—A: lateral habitus; B: head, mesosoma, wings, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="30" pageNumber="65">Fig. 19 B</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 6.0–6.6 × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 3.8–4.0: 2.0: 1.4: 0.5–0.6: 0.65–0.7. Hind tarsal claws simple except for minute subbasal teeth, sometimes with a small tooth, strongly bent apically (ca. 90°) (
<figureCitation id="67B114D7E91C4A52FCFCFEE3FC4BFECA" box="[855,949,295,321]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="30" pageNumber="65">Fig. 2 L</figureCitation>
).
</paragraph>
<paragraph id="FF350852E91C4A52FF6CFE8FFBDAFE26" blockId="30.[151,1437,151,1077]" pageId="30" pageNumber="65">Metasoma. T I 2.0–2.05 × as long as maximum width, punctate except for median longitudinal smooth area. T II 1.45–1.5 × as long as maximum width. T II to T V coriaceous with fine punctures. Ovipositor sheath 1.25–1.4 × as long as hind tibia, ca. 0.5 × as long as fore wing, and shorter than body length.</paragraph>
<paragraph id="FF350852E91C4A52FF6CFE73FD8CFCD6" blockId="30.[151,1437,151,1077]" pageId="30" pageNumber="65">
Colouration (
<figureCitation id="67B114D7E91C4A52FEF7FE73FE10FE5A" box="[348,494,439,465]" captionStart="FIGURE 19" captionStartId="28.[151,250,1641,1665]" captionTargetBox="[151,1436,181,1617]" captionTargetId="figure-19@28.[151,1436,181,1617]" captionTargetPageId="28" captionText="FIGURE 19. Syzeuctus sambonis Uchida, 1928, females from Japan—A: lateral habitus; B: head, mesosoma, wings, and metasoma, dorsal view; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view." pageId="30" pageNumber="65">Figs 19 A–E</figureCitation>
). Body (excluding wings and legs) black. Clypeus, mandible except for teeth, malar space, face except for area around anterior tentorial pits and a median longitudinal spot, inner orbit on frons and vertex, small stripe of outer orbit, and palpi yellow. Antenna blackish-brown to reddish-brown except for small yellow markings of scape and pedicel. Dorsal longitudinal stripe of pronotum, subtegular ridge, scutellum except for a median black spot, postscutellum, and posterior transverse marking of propodeum yellow. Mesoscutum usually with a median small yellow spot. Posterior margins of each metasomal tergites and bases of T III and T IV tinged with reddish-brown to red. Reddish-brown area of metasomal tergites sometimes enlarged. Metasomal sternites blackish-brown except for median parts of membranous parts and posterior margin of subgenital plate tinged with yellowish-brown. Ovipositor reddish-brown. Wings hyaline. Fore wing without conspicuous apical clouded area of fore wing. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow except for black mid coxa; fore coxa with small black spots. Hind leg black to blackish-brown. Apex of femur and base of tibia tinged with yellowish-brown.
</paragraph>
<paragraph id="FF350852E91C4A52FF6CFCA3FDF2FC42" blockId="30.[151,1437,151,1077]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF6CFCA3FEFAFC0A" bold="true" box="[199,260,871,897]" pageId="30" pageNumber="65">Male</emphasis>
(n = 4). Similar to female (
<figureCitation id="67B114D7E91C4A52FDF8FCA3FD11FC0A" box="[595,751,871,897]" captionStart="FIGURE 20" captionStartId="29.[151,250,1767,1791]" captionTargetBox="[151,1436,777,1742]" captionTargetId="figure-310@29.[151,1436,777,1743]" captionTargetPageId="29" captionText="FIGURE 20. Syzeuctus sambonis Uchida, 1928, males from Japan—A: lateral habitus; B: head, mesosoma, and metasoma, dorsal view; C: head, frontal view." pageId="30" pageNumber="65">Figs 20 A–C</figureCitation>
). Tergal spiracles not enlarged. T II 1.38–1.5 × as long as maximum width. Face entirely yellow. Hind tibia paler than female. Antero-lateral part of mesoscutum sometimes with a pair of small yellow spots.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91C4A52FF6CFC16FE36FB9A" pageId="30" pageNumber="65" type="distribution">
<paragraph id="FF350852E91C4A52FF6CFC16FA89FC66" blockId="30.[151,1437,151,1077]" box="[199,1399,978,1005]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF6CFC16FEA7FC67" bold="true" box="[199,345,978,1004]" pageId="30" pageNumber="65">Distribution</emphasis>
.
<collectingCountry id="879D48C2E91C4A52FECCFC17FE59FC66" box="[359,423,979,1005]" name="Japan" pageId="30" pageNumber="65">Japan</collectingCountry>
(Honshu, Miyakejima Is., and Shikoku),
<collectingCountry id="879D48C2E91C4A52FCD3FC17FC45FC66" box="[888,955,979,1005]" name="China" pageId="30" pageNumber="65">China</collectingCountry>
and
<collectingCountry id="879D48C2E91C4A52FC59FC17FBC4FC66" box="[1010,1082,979,1005]" name="South Korea" pageId="30" pageNumber="65">Korea</collectingCountry>
. New to Miyakejima Island.
</paragraph>
<paragraph id="FF350852E91C4A52FF6CFC32FE36FB9A" blockId="30.[151,1437,151,1077]" box="[199,456,1014,1041]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF6CFC32FEBDFB9B" bold="true" box="[199,323,1014,1040]" pageId="30" pageNumber="65">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E91C4A52FF6CFBDFFC3BFBBE" box="[199,965,1050,1077]" pageId="30" pageNumber="65" type="discussion">
<paragraph id="FF350852E91C4A52FF6CFBDFFC3BFBBE" blockId="30.[151,1437,151,1077]" box="[199,965,1050,1077]" pageId="30" pageNumber="65">
<emphasis id="CDFED440E91C4A52FF6CFBDFFECAFBBE" bold="true" box="[199,308,1051,1077]" pageId="30" pageNumber="65">Remarks</emphasis>
.
<bibRefCitation id="9B1B75A3E91C4A52FEE9FBDFFE32FBBE" author="Iwata, K." box="[322,460,1050,1077]" pageId="30" pageNumber="65" pagination="63 - 74" refId="ref23060" refString="Iwata, K. (1958) Ovarian eggs of 233 species of the Japanese Ichneumonidae (Hymenoptera). Acta Hymenopterologica, 1 (1), 63 - 74." type="journal article" year="1958">Iwata (1958</bibRefCitation>
,
<bibRefCitation id="9B1B75A3E91C4A52FE7CFBDFFDEEFBBE" author="Iwata, K." box="[471,528,1051,1077]" pageId="30" pageNumber="65" pagination="115 - 169" refId="ref23092" refString="Iwata, K. (1960) The comparative anatomy of the ovary in Hymenoptera. Part V. Ichneumonidae. Acta Hymenopterologica, 1 (2), 115 - 169." type="journal article" year="1960">1960</bibRefCitation>
) described the ovarian eggs and ovary.
</paragraph>
</subSubSection>
</treatment>
</document>

458
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<paragraph id="FF350852E9224A6CFF3CFD77FE08FD46" blockId="32.[151,578,691,753]" box="[151,502,691,717]" pageId="32" pageNumber="67">
<heading id="A47DBF3EE9224A6CFF3CFD77FE08FD46" bold="true" box="[151,502,691,717]" fontSize="11" level="1" pageId="32" pageNumber="67" reason="1">
<emphasis id="CDFED440E9224A6CFF3CFD77FE08FD46" bold="true" box="[151,502,691,717]" pageId="32" pageNumber="67">
<taxonomicName id="388A73D1E9224A6CFF3CFD77FE6FFD46" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[151,401,691,717]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="67" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9224A6CFF3CFD77FE6FFD46" bold="true" box="[151,401,691,717]" italics="true" pageId="32" pageNumber="67">Syzeuctus yamatonis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6CD693BE9224A6CFE32FD77FE08FD46" box="[409,502,691,717]" pageId="32" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="67B114D7E9224A6CFF34FD13FEFEFD7A" box="[159,256,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="32" pageNumber="67">Figs 1 L</figureCitation>
,
<figureCitation id="67B114D7E9224A6CFEA7FD13FE8BFD7A" box="[268,373,727,753]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="32" pageNumber="67">2 N, R, S</figureCitation>
,
<figureCitation id="67B114D7E9224A6CFE2AFD13FE28FD7A" box="[385,470,727,753]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." pageId="32" pageNumber="67">22 A–F</figureCitation>
,
<figureCitation id="67B114D7E9224A6CFE4BFD13FDC4FD7A" box="[480,570,727,753]" captionStart="FIGURE 23" captionStartId="34.[151,250,1958,1982]" captionTargetBox="[151,1436,417,1935]" captionTargetId="figure-127@34.[151,1436,417,1935]" captionTargetPageId="34" captionText="FIGURE 23. Syzeuctus yamatonis sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view." pageId="32" pageNumber="67">23 A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6CFF3CFCDBFB7AFBF6" pageId="32" pageNumber="67" type="materials_examined">
<paragraph id="FF350852E9224A6CFF3CFCDBFB7AFBF6" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF3CFCDBFEE1FCB3" bold="true" box="[151,287,798,825]" pageId="32" pageNumber="67">Type series</emphasis>
.
<materialsCitation id="4FE2020FE9224A6CFE84FCDBFEBAFCD6" collectingDate="2009-05-27" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" county="Honshu" location="Enkaizan" municipality="Yokohama City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Kanagawa" typeStatus="holotype">
<emphasis id="CDFED440E9224A6CFE84FCDBFE62FCB2" bold="true" box="[303,412,799,825]" pageId="32" pageNumber="67">
<typeStatus id="2031B6F0E9224A6CFE84FCDBFE62FCB2" box="[303,412,799,825]" pageId="32" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
: F,
<emphasis id="CDFED440E9224A6CFE60FCDAFDDBFCB3" bold="true" box="[459,549,798,824]" pageId="32" pageNumber="67">
<collectingCountry id="879D48C2E9224A6CFE60FCDAFDDBFCB3" box="[459,549,798,824]" name="Japan" pageId="32" pageNumber="67">JAPAN</collectingCountry>
</emphasis>
,
<emphasis id="CDFED440E9224A6CFD9DFCDBFD6DFCB2" bold="true" box="[566,659,799,825]" pageId="32" pageNumber="67">
<collectingCounty id="165470DEE9224A6CFD9DFCDBFD6DFCB2" box="[566,659,799,825]" pageId="32" pageNumber="67">Honshu</collectingCounty>
</emphasis>
,
<collectingRegion id="3D4EC6B0E9224A6CFD0FFCDBFCA3FCB2" box="[676,861,799,825]" country="Japan" name="Kanagawa" pageId="32" pageNumber="67">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFCC2FCDBFBE2FCB2" box="[873,1052,799,825]" pageId="32" pageNumber="67">Yokohama City</collectingMunicipality>
,
<location id="FA555E89E9224A6CFB81FCDBFB6BFCB2" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFB81FCDBFB6BFCB2" box="[1066,1173,799,825]" country="Japan" county="Honshu" municipality="Yokohama City" name="Enkaizan" pageId="32" pageNumber="67" stateProvince="Kanagawa">Enkaizan</location>
,
<date id="8B342E92E9224A6CFB08FCDBFAD5FCB2" box="[1187,1323,799,825]" pageId="32" pageNumber="67" value="2009-05-27">
<collectingDate id="9B70D77AE9224A6CFB08FCDBFAD5FCB2" box="[1187,1323,799,825]" pageId="32" pageNumber="67" value="2009-05-27">27. V. 2009</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFA92FCDBFA62FCB2" box="[1337,1436,799,825]" pageId="32" pageNumber="67">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFF7BFC87FEC2FCD6" box="[208,316,835,861]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4FE2020FE9224A6CFEE4FC87FE70FC0A" collectingDate="2008-08-18" collectingDateMax="2008-08-25" collectingDateMin="2008-08-18" collectionCode="KPMNH" collectorName="K. Konishi" country="Japan" location="Hitsujigaoka" municipality="Sapporo City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<emphasis id="CDFED440E9224A6CFEE4FC87FE39FCD6" bold="true" box="[335,455,835,861]" pageId="32" pageNumber="67">
<typeStatus id="2031B6F0E9224A6CFEE4FC87FE39FCD6" box="[335,455,835,861]" pageId="32" pageNumber="67" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="CDFED440E9224A6CFE7CFC86FDCFFCD7" bold="true" box="[471,561,834,860]" pageId="32" pageNumber="67">
<collectingCountry id="879D48C2E9224A6CFE7CFC86FDCFFCD7" box="[471,561,834,860]" name="Japan" pageId="32" pageNumber="67">JAPAN</collectingCountry>
</emphasis>
: [
<emphasis id="CDFED440E9224A6CFDE1FC86FD3EFCD7" bold="true" box="[586,704,834,860]" pageId="32" pageNumber="67">
<collectingRegion id="3D4EC6B0E9224A6CFDE1FC86FD3EFCD7" box="[586,704,834,860]" country="Japan" name="Hokkaido" pageId="32" pageNumber="67">Hokkaido</collectingRegion>
</emphasis>
] 1 F,
<collectingMunicipality id="1F519228E9224A6CFCAFFC87FC65FCD6" box="[772,923,835,861]" pageId="32" pageNumber="67">Sapporo City</collectingMunicipality>
,
<location id="FA555E89E9224A6CFC0CFC87FBC7FCD6" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFC0CFC87FBC7FCD6" box="[935,1081,835,861]" country="Japan" municipality="Sapporo City" name="Hitsujigaoka" pageId="32" pageNumber="67" stateProvince="Hokkaido">Hitsujigaoka</location>
,
<date id="8B342E92E9224A6CFBEEFC86FAEAFCD6" box="[1093,1300,834,861]" pageId="32" pageNumber="67" value="2008-08-18" valueMax="2008-08-25" valueMin="2008-08-18">
<collectingDate id="9B70D77AE9224A6CFBEEFC86FAEAFCD6" box="[1093,1300,834,861]" pageId="32" pageNumber="67" value="2008-08-18" valueMax="2008-08-25" valueMin="2008-08-18">18–25. VIII. 2008</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFA8BFC87FA62FCD6" box="[1312,1436,835,861]" pageId="32" pageNumber="67">K. Konishi</collectorName>
leg., MsT (
<collectionCode id="999B9097E9224A6CFEB1FCA3FE78FC0A" box="[282,390,871,897]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFE36FCA3FB95FC0A" box="[413,1131,870,897]" collectingDate="2010-07-23" collectingDateMax="2010-07-25" collectingDateMin="2010-07-23" collectionCode="EUM" collectorName="K. Takashino" country="Japan" location="Hitsujigaoka" municipality="Sapporo City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
1 F, ditto,
<date id="8B342E92E9224A6CFDB7FCA2FD16FC0A" box="[540,744,870,897]" pageId="32" pageNumber="67" value="2010-07-23" valueMax="2010-07-25" valueMin="2010-07-23">
<collectingDate id="9B70D77AE9224A6CFDB7FCA2FD16FC0A" box="[540,744,870,897]" pageId="32" pageNumber="67" value="2010-07-23" valueMax="2010-07-25" valueMin="2010-07-23">23–25. VII. 2010</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFD5CFCA3FC6EFC0A" box="[759,912,871,897]" pageId="32" pageNumber="67">K. Takashino</collectorName>
leg., MsT (
<collectionCode id="999B9097E9224A6CFBB5FCA3FB9CFC0A" box="[1054,1122,871,897]" pageId="32" pageNumber="67">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFBD1FCA3FB5EFC2E" collectingDate="2012-08-21" collectionCode="KPMNH" collectorName="Y. Matsubara &amp; K. Fukuda" country="Japan" location="Mamurogawa Town" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Yamagata" typeStatus="paratype">
[
<emphasis id="CDFED440E9224A6CFB2FFCA3FB1FFC0A" bold="true" box="[1156,1249,871,897]" pageId="32" pageNumber="67">Honshu</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9224A6CFA85FCA3FF2EFC2E" country="Japan" name="Yamagata" pageId="32" pageNumber="67">Yamagata Pref.</collectingRegion>
,
<location id="FA555E89E9224A6CFF70FC4FFE3EFC2E" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFF70FC4FFE3EFC2E" box="[219,448,907,933]" country="Japan" name="Mamurogawa Town" pageId="32" pageNumber="67" stateProvince="Yamagata">Mamurogawa Town</location>
,
<date id="8B342E92E9224A6CFE60FC4FFD91FC2E" box="[459,623,907,933]" pageId="32" pageNumber="67" value="2012-08-21">
<collectingDate id="9B70D77AE9224A6CFE60FC4FFD91FC2E" box="[459,623,907,933]" pageId="32" pageNumber="67" value="2012-08-21">21. VIII. 2012</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFDD2FC4FFCF0FC2E" box="[633,782,907,933]" pageId="32" pageNumber="67">Y. Matsubara</collectorName>
&amp;
<collectorName id="527F6D84E9224A6CFC98FC4FFC54FC2E" box="[819,938,907,933]" pageId="32" pageNumber="67">K. Fukuda</collectorName>
leg., MsT (
<collectionCode id="999B9097E9224A6CFB87FC4FFB66FC2E" box="[1068,1176,907,933]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFB07FC4FFD40FC42" collectingDate="2009-06-19" collectionCode="KPMNH" collectorName="K. Kubo" country="Japan" location="Mamurogawa Town" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
1 M, same locality of
<typeStatus id="2031B6F0E9224A6CFF3CFC6BFF02FC42" box="[151,252,943,969]" pageId="32" pageNumber="67" type="holotype">holotype</typeStatus>
,
<date id="8B342E92E9224A6CFEA3FC6BFE65FC42" box="[264,411,943,969]" pageId="32" pageNumber="67" value="2009-06-19">
<collectingDate id="9B70D77AE9224A6CFEA3FC6BFE65FC42" box="[264,411,943,969]" pageId="32" pageNumber="67" value="2009-06-19">19. VI. 2009</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFE0CFC6BFDF7FC42" box="[423,521,943,969]" pageId="32" pageNumber="67">K. Kubo</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFDE1FC6BFD48FC42" box="[586,694,943,969]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFD60FC6BFD8CFC66" collectingDate="2013-07-16" collectionCode="KPMNH" collectorName="S. Shimizu" country="Japan" county="Myokou" location="Suginosawa" municipality="City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Niigata" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9224A6CFD56FC6BFC6FFC42" box="[765,913,943,969]" country="Japan" name="Niigata" pageId="32" pageNumber="67">Niigata Pref.</collectingRegion>
,
<collectingCounty id="165470DEE9224A6CFC36FC6BFC01FC42" box="[925,1023,943,969]" pageId="32" pageNumber="67">Myokou</collectingCounty>
,
<collectingMunicipality id="1F519228E9224A6CFBA0FC6BFBC2FC42" box="[1035,1084,943,969]" pageId="32" pageNumber="67">City</collectingMunicipality>
,
<location id="FA555E89E9224A6CFBE2FC6BFB2DFC42" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFBE2FC6BFB2DFC42" box="[1097,1235,943,969]" country="Japan" county="Myokou" municipality="City" name="Suginosawa" pageId="32" pageNumber="67" stateProvince="Niigata">Suginosawa</location>
,
<location id="FA555E89E9224A6CFB74FC6BFA66FC42" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFB74FC6BFA66FC42" box="[1247,1432,943,969]" country="Japan" county="Myokou" municipality="City" name="Mt. Sasagamine" pageId="32" pageNumber="67" stateProvince="Niigata">Mt. Sasagamine</location>
,
<date id="8B342E92E9224A6CFF3CFC17FECDFC66" box="[151,307,979,1005]" pageId="32" pageNumber="67" value="2013-07-16">
<collectingDate id="9B70D77AE9224A6CFF3CFC17FECDFC66" box="[151,307,979,1005]" pageId="32" pageNumber="67" value="2013-07-16">16. VII. 2013</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFEEBFC17FE42FC66" box="[320,444,979,1005]" pageId="32" pageNumber="67">S. Shimizu</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFE55FC17FD94FC66" box="[510,618,979,1005]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFDD5FC17FC1FFC66" box="[638,993,979,1005]" collectingDate="2013-08-07" collectionCode="ZSM" collectorName="S. Shimizu" country="Japan" county="Myokou" location="Suginosawa" municipality="City" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
1 F, ditto,
<date id="8B342E92E9224A6CFD5FFC17FC77FC66" box="[756,905,979,1005]" pageId="32" pageNumber="67" value="2013-08-07">
<collectingDate id="9B70D77AE9224A6CFD5FFC17FC77FC66" box="[756,905,979,1005]" pageId="32" pageNumber="67" value="2013-08-07">7. VIII. 2013</collectingDate>
</date>
(
<collectionCode id="999B9097E9224A6CFC32FC17FC26FC66" box="[921,984,979,1005]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="32" pageNumber="67" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFC45FC17FD01FB9A" collectingDate="2017-05-31" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Kouyoudai" municipality="Narusawa Vil." pageId="32" pageNumber="67" specimenCount="1" stateProvince="Yamanashi" typeStatus="paratype">
1 M,
<collectingRegion id="3D4EC6B0E9224A6CFB87FC17FB16FC66" box="[1068,1256,979,1005]" country="Japan" name="Yamanashi" pageId="32" pageNumber="67">Yamanashi Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFB5FFC17FA66FC66" box="[1268,1432,979,1005]" pageId="32" pageNumber="67">Narusawa Vil.</collectingMunicipality>
,
<location id="FA555E89E9224A6CFF3CFC33FEE8FB9A" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFF3CFC33FEE8FB9A" box="[151,278,1015,1041]" country="Japan" municipality="Narusawa Vil." name="Kouyoudai" pageId="32" pageNumber="67" stateProvince="Yamanashi">Kouyoudai</location>
,
<date id="8B342E92E9224A6CFE88FC33FE54FB9A" box="[291,426,1015,1041]" pageId="32" pageNumber="67" value="2017-05-31">
<collectingDate id="9B70D77AE9224A6CFE88FC33FE54FB9A" box="[291,426,1015,1041]" pageId="32" pageNumber="67" value="2017-05-31">31. V. 2017</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFE1CFC33FDB7FB9A" box="[439,585,1015,1041]" pageId="32" pageNumber="67">K. Watanabe</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFD20FC33FD09FB9A" box="[651,759,1015,1041]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFCA0FC33FDFCFBBE" collectingDate="1982-07-29" collectionCode="KPMNH" collectorName="T. Murota" country="Japan" county="Oono City" location="Ropponnoki" municipality="Koike" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Fukui" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9224A6CFC94FC33FC3DFB9A" box="[831,963,1015,1041]" country="Japan" name="Fukui" pageId="32" pageNumber="67">Fukui Pref.</collectingRegion>
,
<collectingCounty id="165470DEE9224A6CFC64FC33FBB9FB9A" box="[975,1095,1015,1041]" pageId="32" pageNumber="67">Oono City</collectingCounty>
,
<collectingMunicipality id="1F519228E9224A6CFBFFFC33FB65FB9A" box="[1108,1179,1015,1041]" pageId="32" pageNumber="67">Koike</collectingMunicipality>
,
<location id="FA555E89E9224A6CFB02FC33FAC8FB9A" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFB02FC33FAC8FB9A" box="[1193,1334,1015,1041]" country="Japan" county="Oono City" municipality="Koike" name="Ropponnoki" pageId="32" pageNumber="67" stateProvince="Fukui">Ropponnoki</location>
,
<date id="8B342E92E9224A6CFAE8FC33FF2CFBBE" pageId="32" pageNumber="67" value="1982-07-29">
<collectingDate id="9B70D77AE9224A6CFAE8FC33FF2CFBBE" pageId="32" pageNumber="67" value="1982-07-29">29. VII. 1982</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFF76FBDFFEB3FBBE" box="[221,333,1051,1077]" pageId="32" pageNumber="67">T. Murota</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFE25FBDFFE04FBBE" box="[398,506,1051,1077]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFDA5FBDFFE72FBD2" collectingDate="2011-08-06" collectionCode="KPMNH" collectorName="K. Watanabe" country="Japan" location="Mt. Hyonosen" municipality="Wakasa Town" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Tottori" typeStatus="paratype">
1 F,
<collectingRegion id="3D4EC6B0E9224A6CFD94FBDFFD32FBBE" box="[575,716,1051,1077]" country="Japan" name="Tottori" pageId="32" pageNumber="67">Tottori Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFD7CFBDFFC89FBBE" box="[727,887,1051,1077]" pageId="32" pageNumber="67">Wakasa Town</collectingMunicipality>
,
<location id="FA555E89E9224A6CFC28FBDFFBD8FBBE" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFC28FBDFFBD8FBBE" box="[899,1062,1051,1077]" country="Japan" municipality="Wakasa Town" name="Mt. Hyonosen" pageId="32" pageNumber="67" stateProvince="Tottori">Mt. Hyonosen</location>
,
<date id="8B342E92E9224A6CFB98FBDFFB36FBBE" box="[1075,1224,1051,1077]" pageId="32" pageNumber="67" value="2011-08-06">
<collectingDate id="9B70D77AE9224A6CFB98FBDFFB36FBBE" box="[1075,1224,1051,1077]" pageId="32" pageNumber="67" value="2011-08-06">6. VIII. 2011</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFB7FFBDFFA9AFBBE" box="[1236,1380,1051,1077]" pageId="32" pageNumber="67">K. Watanabe</collectorName>
leg.,
<collectingMethod id="26CB7045E9224A6CFF3CFBFBFEF6FBD2" box="[151,264,1087,1113]" pageId="32" pageNumber="67">Light trap</collectingMethod>
(
<collectionCode id="999B9097E9224A6CFEB3FBFBFE7AFBD2" box="[280,388,1087,1113]" pageId="32" pageNumber="67">KPMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFE33FBFBFA66FBD2" box="[408,1432,1086,1113]" collectingDate="1955-07-29" collectionCode="EUM" collectorName="S. Ueda" country="Japan" location="Jyojyusha" municipality="Mt. Ishizuchi" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Ehime" typeStatus="paratype">
[
<emphasis id="CDFED440E9224A6CFE09FBFAFDF8FBD3" bold="true" box="[418,518,1086,1112]" pageId="32" pageNumber="67">Shikoku</emphasis>
] 1 F,
<collectingRegion id="3D4EC6B0E9224A6CFDE1FBFBFD2BFBD2" box="[586,725,1087,1113]" country="Japan" name="Ehime" pageId="32" pageNumber="67">Ehime Pref.</collectingRegion>
,
<collectingMunicipality id="1F519228E9224A6CFD4AFBFBFC85FBD2" box="[737,891,1087,1113]" pageId="32" pageNumber="67">Mt. Ishizuchi</collectingMunicipality>
,
<location id="FA555E89E9224A6CFC2CFBFBFC04FBD2" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFC2CFBFBFC04FBD2" box="[903,1018,1087,1113]" country="Japan" municipality="Mt. Ishizuchi" name="Jyojyusha" pageId="32" pageNumber="67" stateProvince="Ehime">Jyojyusha</location>
,
<date id="8B342E92E9224A6CFBACFBFBFB5DFBD2" box="[1031,1187,1086,1113]" pageId="32" pageNumber="67" value="1955-07-29">
<collectingDate id="9B70D77AE9224A6CFBACFBFBFB5DFBD2" box="[1031,1187,1086,1113]" pageId="32" pageNumber="67" value="1955-07-29">29. VII. 1955</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFB04FBFBFAF7FBD2" box="[1199,1289,1087,1113]" pageId="32" pageNumber="67">S. Ueda</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFAE7FBFBFA6EFBD2" box="[1356,1424,1087,1113]" pageId="32" pageNumber="67">EUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4FE2020FE9224A6CFF3CFBA7FB7EFBF6" box="[151,1152,1123,1149]" collectingDate="1961-07-11" collectionCode="MNHAH" collectorName="A. Nakanishi" country="Japan" location="Cape Sata" pageId="32" pageNumber="67" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
[
<emphasis id="CDFED440E9224A6CFF0AFBA7FF00FBF6" bold="true" box="[161,254,1123,1149]" pageId="32" pageNumber="67">Kyushu</emphasis>
] 1 M,
<collectingRegion id="3D4EC6B0E9224A6CFEE1FBA7FDF4FBF6" box="[330,522,1123,1149]" country="Japan" name="Kagoshima" pageId="32" pageNumber="67">Kagoshima Pref.</collectingRegion>
,
<location id="FA555E89E9224A6CFDBFFBA7FD76FBF6" LSID="urn:lsid:plazi:treatment:7723B944E9224A6FFF3CFD77FC69FC9E:FA555E89E9224A6CFDBFFBA7FD76FBF6" box="[532,648,1123,1149]" country="Japan" name="Cape Sata" pageId="32" pageNumber="67" stateProvince="Kagoshima">Cape Sata</location>
,
<date id="8B342E92E9224A6CFD38FBA7FCD4FBF6" box="[659,810,1123,1149]" pageId="32" pageNumber="67" value="1961-07-11">
<collectingDate id="9B70D77AE9224A6CFD38FBA7FCD4FBF6" box="[659,810,1123,1149]" pageId="32" pageNumber="67" value="1961-07-11">11. VII. 1961</collectingDate>
</date>
,
<collectorName id="527F6D84E9224A6CFC9EFBA7FC37FBF6" box="[821,969,1123,1149]" pageId="32" pageNumber="67">A. Nakanishi</collectorName>
leg. (
<collectionCode id="999B9097E9224A6CFBA3FBA7FB87FBF6" box="[1032,1145,1123,1149]" pageId="32" pageNumber="67">MNHAH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9224A6FFF6CFB42FA75FEEE" lastPageId="35" lastPageNumber="70" pageId="32" pageNumber="67" type="description">
<paragraph id="FF350852E9224A6CFF6CFB42FC36FB4E" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
<emphasis id="CDFED440E9224A6CFF6CFB42FEAFFB2B" bold="true" box="[199,337,1158,1184]" pageId="32" pageNumber="67">Description</emphasis>
.
<emphasis id="CDFED440E9224A6CFEF5FB43FE4BFB2A" bold="true" box="[350,437,1159,1185]" pageId="32" pageNumber="67">Female</emphasis>
(n = 8). Body polished and covered with punctures and silvery setae; length 9.0–10.8 (HT: 10.8) mm. Setae on head and mesosoma not exceptionally long and erect
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFB0BFA7BFA6E" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
Head 0.6 × as long as wide. Combined length of clypeus and face 0.95–1.1 (HT: 1.0) × facial width. Clypeus smooth except for dorsal margin sparsely punctate, slightly convex in lateral view, without central transverse ridge. Face densely punctate, slightly convex medially in lateral view, without a pair of convexities just above of each anterior tentorial pit. Frons slightly concave, without horns or convexities. Malar space 0.5–0.6 (HT: 0.5) × as long as basal mandibular width. Genal carina lamella-like ventrally, reaching mandibular base. POL 1.6–1.8 (HT: 1.7) × as long as OD. OOL 1.2–1.4 (HT: 1.2) × as long as OD. Temple weakly roundly narrowed behind eye in dorsal view (
<figureCitation id="67B114D7E9224A6CFF34FA63FEECFA4A" box="[159,274,1447,1473]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." pageId="32" pageNumber="67">Fig. 22 D</figureCitation>
). Antenna with 42–44 (HT: 44) flagellomeres. FL I 4.1–4.7 (HT: 4.7) × as long as maximum depth in lateral view and 1.5–1.85 (HT: 1.75) × as long as FL II. Preapical flagellomere 1.0 × as long as maximum width.
</paragraph>
<paragraph id="FF350852E9224A6CFF6CFA2BFAD9F8C6" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">
Mesosoma coarsely and densely punctate, 1.4–1.45 (HT: 1.4) × as long as maximum height. Pronotum with small smooth area just behind of epomia. Epomia present, its dorsal end not reaching dorsal margin. Mesopleuron densely punctate ventrally, sparsely punctate dorsally; speculum with a large smooth area (
<figureCitation id="67B114D7E9224A6CFB31F9F3FAF6F9DA" box="[1178,1288,1591,1617]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." pageId="32" pageNumber="67">Fig. 22 E</figureCitation>
). Epicnemial carina present latero-ventrally and ventrally (
<figureCitation id="67B114D7E9224A6CFD09F99FFCF9F9FE" box="[674,775,1627,1653]" captionStart="FIGURE 1" captionStartId="4.[151,250,984,1008]" captionTargetBox="[159,1432,313,958]" captionTargetId="figure-85@4.[151,1436,309,961]" captionTargetPageId="4" captionText="FIGURE 1. Epicnemial carina of Japanese species of Syzeuctus spp., females—A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D: S. flavitarsis sp. nov. (paratype); E: S. flavofacialis Kang &amp; Lee, 2020; F: S. laoticus Riedel, 2022; G: S. maculatus Sheng, 2009; H: S. nigrus sp. nov. (holotype); I: S. rufiapicalis sp. nov. (holotype); J: S. sambonis Uchida, 1928; K: S. takaozanus Uchida, 1928; L: S. yamatonis sp. nov. (holotype)." pageId="32" pageNumber="67">Fig. 1 L</figureCitation>
). Dorsal surface of propodeum without transverse rugae. Posterior transverse carina of propodeum largely absent medially, obsolete laterally. Fore wing length 7.6–8.2 (HT: 8.2) mm. Areolet receiving vein 2m-cu slightly basal to the outer angle, petiolated anteriorly; petiole as long as areolet (
<figureCitation id="67B114D7E9224A6CFEBDF903FE79F96A" box="[278,391,1735,1761]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." pageId="32" pageNumber="67">Fig. 22 B</figureCitation>
). Fore wing vein 1cu-a subvertical, slightly postfurcal to vein M&amp;RS. Nervellus reclivous, intercepted posterior to the middle. Hind femur 5.2–5.7 (HT: 5.25) × as long as maximum depth in lateral view. Ratio of length of first to fifth hind tarsomeres 4.0–4.1 (HT: 4.1): 2.0: 1.3: 0.6–0.7 (HT: 0.7): 0.7–0.8 (HT: 0.8). Hind tarsal claws simple except for minute subbasal teeth, moderately bent apically (less than 90°) (
<figureCitation id="67B114D7E9224A6CFB12F8F7FAE4F8C6" box="[1209,1306,1843,1869]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="32" pageNumber="67">Fig. 2 N</figureCitation>
).
</paragraph>
<paragraph id="FF350852E9224A6CFF6CF893FDDBF856" blockId="32.[151,1437,798,2013]" pageId="32" pageNumber="67">Metasoma. T I 1.9–2.4 (HT: 2.2) × as long as maximum width, finely punctate laterally, smooth medially. T II 1.35–1.5 (HT: 1.5) × as long as maximum width, finely and densely punctate. T III to T V coriaceous with fine and dense punctures. Ovipositor sheath 2.1–2.5 (HT: 2.3) × as long as hind tibia, ca. 0.8–0.85 (HT: 0.8) × as long as fore wing, and shorter than body length.</paragraph>
<caption id="ABF558DAE9234A6DFF3CF9A8FA6AF923" pageId="33" pageNumber="68" startId="33.[151,248,1644,1668]" targetBox="[151,1436,181,1619]" targetPageId="33" targetType="figure">
<paragraph id="FF350852E9234A6DFF3CF9A8FA6AF923" blockId="33.[151,1437,1644,1704]" pageId="33" pageNumber="68">
<emphasis id="CDFED440E9234A6DFF3CF9A8FEE0F90F" bold="true" box="[151,286,1644,1668]" pageId="33" pageNumber="68">FIGURE 22.</emphasis>
<taxonomicName id="388A73D1E9234A6DFE8EF9A9FE0DF90F" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[293,499,1645,1668]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="68" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9234A6DFE8EF9A9FE0DF90F" box="[293,499,1645,1668]" italics="true" pageId="33" pageNumber="68">Syzeuctus yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9234A6DFE51F9A8FDB4F90F" bold="true" box="[506,586,1644,1668]" pageId="33" pageNumber="68">
<taxonomicNameLabel id="D6CD693BE9234A6DFE51F9A8FDB4F90F" box="[506,586,1644,1668]" pageId="33" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus.
</paragraph>
</caption>
<paragraph id="FF350852E9234A6EFF6CF912FA9DFE02" blockId="33.[151,1437,1750,2029]" lastBlockId="34.[151,1437,151,393]" lastPageId="34" lastPageNumber="69" pageId="33" pageNumber="68">
Colouration (
<figureCitation id="67B114D7E9234A6DFEF7F912FE10F97A" box="[348,494,1750,1777]" captionStart="FIGURE 22" captionStartId="33.[151,248,1644,1668]" captionTargetBox="[151,1436,181,1619]" captionTargetId="figure-19@33.[151,1436,181,1620]" captionTargetPageId="33" captionText="FIGURE 22. Syzeuctus yamatonis sp. nov., females, paratype (A) and holotype (B-F)—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: head, dorsal view; E: pronotum and mesopleuron, lateral view; F: apical part of hind tibia and hind tarsus." pageId="33" pageNumber="68">Figs 22 A–F</figureCitation>
). Body (excluding wings and legs) black. Clypeus, mandible except for teeth, malar space, a pair of small yellow spots below antennal sockets, orbits on face, frons, vertex and gena, and palpi yellow. Antenna blackish-brown to reddish-brown. Scape and pedicel usually partly tinged with yellow ventrally. Two spots of collar, a L-shape marking of dorsal margin of pronotum, a pair of antero-lateral markings and a median marking of mesoscutum, subtegular ridge, antero-dorsal and postero-ventral spots of mesopleuron, scutellum except for a median black spot, postscutellum, posterior spot of metapleuron, a pair of antero-lateral spots of propodeum, and a posterior triangle marking of propodeum yellow. Scutellum sometimes entirely yellow. Metasomal tergites black except for a pair of basal spots of T I and posterior margins of T I to T III yellow to ivory. Posterior margins of T IV sometimes tinged with yellow. Membranous parts of metasomal sternites largely yellow. Subgenital plate blackish-brown, usually partly tinged with yellowish-brown. Ovipositor reddish-brown. Wings hyaline.Apical clouded area of fore wing slightly to weakly present. Veins and pterostigma blackish-brown except for yellowish-brown wing base. Fore and mid legs yellow; mid coxa partly tinged with blackish-brown basally. Hind coxa black except for a dorsal spot and apex yellow to yellowish-brown. Hind trochanter black to blackish-brown. Hind trochantellus reddish-yellow. Hind femur reddish-brown except for apex tinged with yellow. Hind tibia and tibial spurs yellowish-brown to whitish-brown except for apical part of tibia blackish-brown. Hind tarsus brown, partly tinged with yellow.
</paragraph>
<caption id="ABF558DAE9204A6EFF3CF862FE73F868" pageId="34" pageNumber="69" startId="34.[151,250,1958,1982]" targetBox="[151,1436,417,1935]" targetPageId="34" targetType="figure">
<paragraph id="FF350852E9204A6EFF3CF862FE73F868" blockId="34.[151,1437,1958,2019]" pageId="34" pageNumber="69">
<emphasis id="CDFED440E9204A6EFF3CF862FEDFF835" bold="true" box="[151,289,1958,1982]" pageId="34" pageNumber="69">FIGURE 23.</emphasis>
<taxonomicName id="388A73D1E9204A6EFE83F863FE07F835" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[296,505,1959,1982]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="69" phylum="Arthropoda" rank="species" species="yamatonis" status="sp. nov.">
<emphasis id="CDFED440E9204A6EFE83F863FE07F835" box="[296,505,1959,1982]" italics="true" pageId="34" pageNumber="69">Syzeuctus yamatonis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9204A6EFDAAF863FDACF834" bold="true" box="[513,594,1959,1983]" pageId="34" pageNumber="69">
<taxonomicNameLabel id="D6CD693BE9204A6EFDAAF863FDACF834" box="[513,594,1959,1983]" pageId="34" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view.
</paragraph>
</caption>
<paragraph id="FF350852E9214A6FFF6CFF53FA75FEEE" blockId="35.[151,1437,150,789]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFF53FEFAFF3A" bold="true" box="[199,260,151,177]" pageId="35" pageNumber="70">Male</emphasis>
(n = 3). Similar to female (
<figureCitation id="67B114D7E9214A6FFD9CFF53FD32FF3A" box="[567,716,151,177]" captionStart="FIGURE 23" captionStartId="34.[151,250,1958,1982]" captionTargetBox="[151,1436,417,1935]" captionTargetId="figure-127@34.[151,1436,417,1935]" captionTargetPageId="34" captionText="FIGURE 23. Syzeuctus yamatonis sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view." pageId="35" pageNumber="70">Figs 23 A–D</figureCitation>
). A male with body length
<quantity id="3872A5B7E9214A6FFC51FF52FBADFF3A" box="[1018,1107,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="35" pageNumber="70" unit="mm" value="8.5">8.5 mm</quantity>
. Malar space 0.9 × as long as basal mandibular width. Tergal spiracles enlarged and with circular felt-like area, its length less than 0.2× as long as T I (
<figureCitation id="67B114D7E9214A6FFF4FFF1BFE9DFF72" box="[228,355,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="35" pageNumber="70">Figs 2 R, S</figureCitation>
,
<figureCitation id="67B114D7E9214A6FFEDBFF1BFE55FF72" box="[368,427,223,249]" captionStart="FIGURE 23" captionStartId="34.[151,250,1958,1982]" captionTargetBox="[151,1436,417,1935]" captionTargetId="figure-127@34.[151,1436,417,1935]" captionTargetPageId="34" captionText="FIGURE 23. Syzeuctus yamatonis sp. nov., males, paratypes—A: lateral habitus; B: dorsal habitus; C: head, frontal view; D: T I to T III, lateral view." pageId="35" pageNumber="70">23 D</figureCitation>
). Basal part of T I punctate laterally (
<figureCitation id="67B114D7E9214A6FFCE7FF1BFC59FF72" box="[844,935,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1698,1722]" captionTargetBox="[163,1423,399,1663]" captionTargetId="figure-86@5.[151,1435,381,1675]" captionTargetPageId="5" captionText="FIGURE 2. Hind tarsal claws and metasomal tergites of Japanese species of Syzeuctus spp., females (A-N) and males (O–S)— A: S. apicifer (Walker, 1874); B: S. brevicaudus sp. nov. (holotype); C: S. coreanus Uchida, 1928; D, O: S. flavitarsis sp. nov. (holotype); E–G: S. flavofacialis Kang &amp; Lee, 2020; H: S. laoticus Riedel, 2022; I: S. maculatus Sheng, 2009; J: S. nigrus sp. nov. (holotype); K, P, Q: S. rufiapicalis sp. nov. (K: holotype; P, Q: paratype); L: S. sambonis Uchida, 1928; M: S. takaozanus Uchida, 1928; N, R, S: S. yamatonis sp. nov. (N: holotype; R, S: paratype)—A–E, G–N: outer claw; F: inner claw; O, P, R: T I and T II, lateral view; Q, S: T I and T II, dorsal view." pageId="35" pageNumber="70">Fig. 2 S</figureCitation>
). Hind femur 5.3–6.3 × as long as maximum depth in lateral view. T I 2.4–2.8 × as long as maximum width. T II 1.7–2.05 × as long as maximum width. Yellow area of body larger than female. Face yellow except for a small median longitudinal spot. Two yellow spots of mesopleuron united into a single C-shaped marking. Posterior segments of metasoma tinged with reddish-brown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9214A6FFF6CFEAAFC62FE02" box="[199,924,366,393]" pageId="35" pageNumber="70" type="distribution">
<paragraph id="FF350852E9214A6FFF6CFEAAFC62FE02" blockId="35.[151,1437,150,789]" box="[199,924,366,393]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFEAAFEA7FE03" bold="true" box="[199,345,366,392]" pageId="35" pageNumber="70">Distribution</emphasis>
.
<collectingCountry id="879D48C2E9214A6FFECCFEABFE59FE02" box="[359,423,367,393]" name="Japan" pageId="35" pageNumber="70">Japan</collectingCountry>
(
<collectingRegion id="3D4EC6B0E9214A6FFE1DFEABFDD7FE02" box="[438,553,367,393]" country="Japan" name="Hokkaido" pageId="35" pageNumber="70">Hokkaido</collectingRegion>
, Honshu, Shikoku, and Kyushu).
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9214A6FFF6CFE57FE36FE5A" pageId="35" pageNumber="70" type="etymology">
<paragraph id="FF350852E9214A6FFF6CFE57FC35FE26" blockId="35.[151,1437,150,789]" box="[199,971,403,429]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFE57FEB6FE26" bold="true" box="[199,328,403,429]" pageId="35" pageNumber="70">Etymology</emphasis>
. The specific name is from old name of
<collectingCountry id="879D48C2E9214A6FFCA5FE57FCB0FE26" box="[782,846,403,429]" name="Japan" pageId="35" pageNumber="70">Japan</collectingCountry>
“Yamato”.
</paragraph>
<paragraph id="FF350852E9214A6FFF6CFE72FE36FE5A" blockId="35.[151,1437,150,789]" box="[199,456,438,465]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFE72FEBDFE5B" bold="true" box="[199,323,438,464]" pageId="35" pageNumber="70">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B7905BD9E9214A6FFF6CFE1FFC69FC9E" pageId="35" pageNumber="70" type="discussion">
<paragraph id="FF350852E9214A6FFF6CFE1FFC69FC9E" blockId="35.[151,1437,150,789]" pageId="35" pageNumber="70">
<emphasis id="CDFED440E9214A6FFF6CFE1FFECAFE7E" bold="true" box="[199,308,475,501]" pageId="35" pageNumber="70">Remarks</emphasis>
. This species is closely related to
<taxonomicName id="388A73D1E9214A6FFD69FE1FFCAEFE7E" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[706,848,475,501]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="flavitarsis" status="sp. nov.">
<emphasis id="CDFED440E9214A6FFD69FE1FFCAEFE7E" box="[706,848,475,501]" italics="true" pageId="35" pageNumber="70">S. flavitarsis</emphasis>
</taxonomicName>
<emphasis id="CDFED440E9214A6FFCF2FE1FFC4BFE7E" bold="true" box="[857,949,475,501]" pageId="35" pageNumber="70">
<taxonomicNameLabel id="D6CD693BE9214A6FFCF2FE1FFC4BFE7E" box="[857,949,475,501]" pageId="35" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Both species were collected same locality (
<collectingRegion id="3D4EC6B0E9214A6FFF35FE3BFEB1FD92" box="[158,335,511,537]" country="Japan" name="Yamagata" pageId="35" pageNumber="70">Yamagata Pref.</collectingRegion>
, Mamurogawa Vil.) on same day. This species can be distinguished by the punctures on upper part of mesopleuron which are irregular and sparser than lower part (same density in
<taxonomicName id="388A73D1E9214A6FFBBDFDE7FB5FFDB6" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[1046,1185,547,573]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="flavitarsis">
<emphasis id="CDFED440E9214A6FFBBDFDE7FB5FFDB6" box="[1046,1185,547,573]" italics="true" pageId="35" pageNumber="70">S. flavitarsis</emphasis>
</taxonomicName>
) and the first to fourth hind tarsomeres usually more or less darkened (ivory in
<taxonomicName id="388A73D1E9214A6FFCB7FD83FC57FDEA" authorityName="Watanabe &amp; Riedel" authorityYear="2024" box="[796,937,583,609]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="flavitarsis">
<emphasis id="CDFED440E9214A6FFCB7FD83FC57FDEA" box="[796,937,583,609]" italics="true" pageId="35" pageNumber="70">S. flavitarsis</emphasis>
</taxonomicName>
). In the key provided by
<bibRefCitation id="9B1B75A3E9214A6FFB65FD83FA93FDEA" author="Riedel, M." box="[1230,1389,583,609]" pageId="35" pageNumber="70" pagination="1013 - 1098" refId="ref23284" refString="Riedel, M. (2021) The genus Syzeuctus Forster in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species. Linzer biologische Beitrage, 52 (2), 1013 - 1098." type="journal article" year="2021">Riedel (2021)</bibRefCitation>
, the females and males runs to couplet 47 (female of
<taxonomicName id="388A73D1E9214A6FFD14FDAFFCCCFD0E" authorityName="Seyrig" authorityYear="1928" box="[703,818,619,645]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFD14FDAFFCCCFD0E" box="[703,818,619,645]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
) and 7 (male of
<taxonomicName id="388A73D1E9214A6FFC5BFDAFFB9AFD0E" authorityName="Seyrig" authorityYear="1928" box="[1008,1124,619,645]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFC5BFDAFFB9AFD0E" box="[1008,1124,619,645]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), but differs by the antenna with 42–44 flagellomeres (38–40 flagellomeres in female of
<taxonomicName id="388A73D1E9214A6FFC95FD4BFC4FFD22" authorityName="Seyrig" authorityYear="1928" box="[830,945,655,681]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFC95FD4BFC4FFD22" box="[830,945,655,681]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), the malar space 0.5–0.6 × as long as basal mandibular width (
<quantity id="3872A5B7E9214A6FFEC5FD77FE4FFD46" box="[366,433,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.7779999999999998" pageId="35" pageNumber="70" unit="in" value="0.7">0.7 in</quantity>
female of
<taxonomicName id="388A73D1E9214A6FFD87FD77FD5EFD46" authorityName="Seyrig" authorityYear="1928" box="[556,672,691,717]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFD87FD77FD5EFD46" box="[556,672,691,717]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), the median part of face largely black (largely yellow in female of
<taxonomicName id="388A73D1E9214A6FFF3CFD13FEF4FD7A" authorityName="Seyrig" authorityYear="1928" box="[151,266,727,753]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFF3CFD13FEF4FD7A" box="[151,266,727,753]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
), the yellow markings of frons and vertex narrower than
<taxonomicName id="388A73D1E9214A6FFC23FD13FC05FD7A" authorityName="Seyrig" authorityYear="1928" box="[904,1019,727,753]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFC23FD13FC05FD7A" box="[904,1019,727,753]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
, and the hind coxa black with yellow apical marking(s) (yellow with black spot in the male of
<taxonomicName id="388A73D1E9214A6FFCB8FD3FFC78FC9E" authorityName="Seyrig" authorityYear="1928" box="[787,902,763,789]" class="Insecta" family="Ichneumonidae" genus="Syzeuctus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="70" phylum="Arthropoda" rank="species" species="dusmeti">
<emphasis id="CDFED440E9214A6FFCB8FD3FFC78FC9E" box="[787,902,763,789]" italics="true" pageId="35" pageNumber="70">S. dusmeti</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>