251 lines
28 KiB
XML
251 lines
28 KiB
XML
<document id="503D2846F47E807CBBAF9BF82953709B" ID-CLB-Dataset="302547" ID-DOI="10.11646/phytotaxa.376.3.1" ID-GBIF-Dataset="10464e27-82f2-4df2-9169-da26984e2770" ID-ISSN="1179-3163" ID-Zenodo-Dep="13726654" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725676653091" checkinUser="felipe" docAuthor="Lu, Ngan Thi, Zhang, Liang, Zhou, Xin-Mao, Gao, Xin-Fen & Zhang, Li-Bing" docDate="2018" docId="E078202E28304719AE85577AFAA7FB08" docLanguage="en" docName="phytotaxa.376.3.1.pdf" docOrigin="Phytotaxa 376 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.376.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Arachniodes quangnamensis Li Bing Zhang, Ngan Thi Lu & Liang Zhang 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="131" masterDocId="1C4158562834471CAE0D5040FFB2FFE6" masterDocTitle="Three new species of the fern genus Arachniodes (Dryopteridaceae) from Vietnam" masterLastPageNumber="132" masterPageNumber="126" pageNumber="130" updateTime="1725689664931" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="5433AA18F7D3B38D71E424E1F2475980">
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<mods:title id="DDD64842AF44AEA032F8AAA93BC55BCC">Three new species of the fern genus Arachniodes (Dryopteridaceae) from Vietnam</mods:title>
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<mods:roleTerm id="3DF90302A6C11230238D9BD443D6C46B">Author</mods:roleTerm>
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<mods:namePart id="B2917937E3034CC1CFF2620A2CCFD977">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="7E8A67167CD980CCCE6B5B32CDCFAA7E">CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China & Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam & University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<mods:roleTerm id="6C7FC774347132F3F614A5CA83024104">Author</mods:roleTerm>
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<mods:namePart id="5ED96D2D04F376A77D7219EF988153F6">Zhang, Liang</mods:namePart>
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<mods:affiliation id="F49B4D1F8A2191DCC89D8778B4CAF8EA">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China.</mods:affiliation>
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<mods:namePart id="C49A04F4203D03276F10CC90D5002113">Zhou, Xin-Mao</mods:namePart>
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<mods:affiliation id="B1E6EC4D2D8028B7DA828A1565BD0720">School of Ecology and Environmental Science, Yunnan University, Kunming, 650091, Yunnan, China;</mods:affiliation>
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<mods:namePart id="CD7AE87B1B3709D39812E3BEEC502287">Gao, Xin-Fen</mods:namePart>
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<mods:affiliation id="D57F152EE6EF22A76B11EC770F58E822">CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China</mods:affiliation>
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<mods:namePart id="51643BD3F4E4E7C78B77F3B2C1132E58">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="3B2F46D04DDF9966AE18C69C7DDAC9CD">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A.</mods:affiliation>
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<mods:title id="7E33086BE068FEEF7AFACEFE6B06DF7D">Phytotaxa</mods:title>
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<mods:date id="FD873C3BFA8041364E3C3EE4F4015A49">2018</mods:date>
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<mods:number id="A1E01F014500C1F4F7CBAF043E5822B0">2018-11-19</mods:number>
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<subSubSection id="20CBC2B328304718AE85577AFA2FF8B3" box="[136,1437,1850,1877]" pageId="4" pageNumber="130" type="nomenclature">
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<paragraph id="686E913828304718AE85577AFA2FF8B3" blockId="4.[136,1437,1850,1877]" box="[136,1437,1850,1877]" pageId="4" pageNumber="130">
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<heading id="3326265428304718AE85577AFA2FF8B3" box="[136,1437,1850,1877]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="130" reason="3">
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<taxonomicName id="AFD1EABB28304718AE85577AFC6AF8B3" ID-CoL="G33F" authority="Li Bing Zhang, Ngan Thi Lu & Liang Zhang" authorityName="Li Bing Zhang, Ngan Thi Lu & Liang Zhang" authorityYear="2018" box="[136,984,1850,1877]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="130" phylum="Tracheophyta" rank="species" species="quangnamensis" status="sp. nov.">
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<emphasis id="5AA54D2A28304718AE85577AFE65F8B2" bold="true" box="[136,471,1850,1877]" italics="true" pageId="4" pageNumber="130">Arachniodes quangnamensis</emphasis>
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Li Bing Zhang, Ngan Thi Lu & Liang Zhang
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</taxonomicName>
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,
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<emphasis id="5AA54D2A28304718ADEE577CFB8AF8B3" box="[995,1080,1852,1877]" italics="true" pageId="4" pageNumber="130">
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<taxonomicNameLabel id="4196F05128304718ADEE577CFB8AF8B3" box="[995,1080,1852,1877]" pageId="4" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<figureCitation id="F0EA8DBD28304718AA33577CFB73F8B3" box="[1086,1217,1852,1877]" captionStart="FIGURE 1" captionStartId="1.[136,229,1623,1644]" captionTargetBox="[298,1290,315,1598]" captionTargetId="figure-73@1.[298,1290,314,1598]" captionTargetPageId="1" captionText="FIGURE 1. A–H. Arachniodes quangnamensis Li Bing Zhang, N.T. Lu & Liang Zhang,—A & B. Habit.—C. Sterile frond.—D. Lower portion of plant.—E. Fertile frond.—F. Portion of frond showing basal pinnae.—G. Portion of rachis and pinna.—H. Portion of pinnule showing sori and microscales. J–P. A. daklakensis Li Bing Zhang, N.T. Lu & X.M.Zhou.—J. Fronds.—K & M. Lowest pinnae.—L. Stipe base.—N. Lamina apex.—O. Portions of lateral pinnae.—P. Portion of pinnule showing sori." figureDoi="http://doi.org/10.5281/zenodo.13726656" httpUri="https://zenodo.org/record/13726656/files/figure.png" pageId="4" pageNumber="130">
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Figs. 1A
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<emphasis id="5AA54D2A28304718AA92577EFB1FF8B2" box="[1183,1197,1854,1876]" italics="true" pageId="4" pageNumber="130">‒</emphasis>
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H
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</figureCitation>
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&
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<figureCitation id="F0EA8DBD28304718AAE8577CFA2BF8B3" box="[1253,1433,1852,1877]" captionStart="FIGURE 3" captionStartId="3.[136,229,1862,1883]" captionTargetBox="[151,1436,190,1835]" captionTargetId="figure-18@3.[151,1436,190,1837]" captionTargetPageId="3" captionText="FIGURE 3. The morphology of scales, spores, and indusia. A–C, H, I & L. Arachniodes quangnamensis.—A. Rhizome scales.—B. Scales from stipe base.—C. Scales from distal portion of stipe.—H & I. SEM photographs of spores.—L. Ciliate indusia. D, E, J, K & N. A. longicaudata.—D. Scales from stipe base.—E. Scales from distal portion of stipe.—J & K. SEM photographs of spores.—N. Indusia with long and hairlike ciliate. F, G & M. A. daklakensis.—F. Scales from stipe base.—G. Scales from distal portion of stipe.—M. Indusia with long and hairlike ciliate." figureDoi="http://doi.org/10.5281/zenodo.13726662" httpUri="https://zenodo.org/record/13726662/files/figure.png" pageId="4" pageNumber="130">
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3A
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<emphasis id="5AA54D2A28304718AB0A577EFAA7F8B2" box="[1287,1301,1854,1876]" italics="true" pageId="4" pageNumber="130">‒</emphasis>
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C, H, I & L
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</figureCitation>
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.
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="20CBC2B328304719AE855722FB9FFC84" lastPageId="5" lastPageNumber="131" pageId="4" pageNumber="130" type="materials_examined">
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<paragraph id="686E913828304718AE855722FCA3F87D" blockId="4.[136,1452,1890,2091]" pageId="4" pageNumber="130">
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Type:—
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<collectingCountry id="10C6D1A828304718AEDB5722FEFAF891" box="[214,328,1890,1911]" name="Vietnam" pageId="4" pageNumber="130">VIETNAM</collectingCountry>
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. Quangnam Province: Songthanh Nature reserve, elev.
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<quantity id="AF293CDD28304718AD5B5722FC77F891" box="[854,965,1890,1911]" metricMagnitude="2" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="4" pageNumber="130" unit="m" value="430.0" valueMax="460.0" valueMin="400.0">400–460 m</quantity>
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,
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<date id="1C6FB7F828304718ADC35722FBC8F891" box="[974,1146,1890,1911]" pageId="4" pageNumber="130" value="2014-12-05">5 December 2014</date>
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,
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<emphasis id="5AA54D2A28304718AA8E5722FEDFF87D" italics="true" pageId="4" pageNumber="130">Li Bing Zhang, Liang Zhang & Ngan Thi Lu 7926</emphasis>
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(
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<typeStatus id="B76A2F9A28304718AF7457C6FE7DF87D" box="[377,463,1926,1947]" pageId="4" pageNumber="130" type="holotype">holotype</typeStatus>
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<collectionCode id="0EC009FD28304718AFD957C6FD97F87D" box="[468,549,1926,1947]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="4" pageNumber="130">VNMN</collectionCode>
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!,
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<typeStatus id="B76A2F9A28304718AC3F57C6FD33F87D" box="[562,641,1926,1947]" pageId="4" pageNumber="130" type="isotype">isotypes</typeStatus>
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<collectionCode id="0EC009FD28304718AC8B57C6FD74F87D" box="[646,710,1926,1947]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12796" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="4" pageNumber="130" type="Herbarium">CDBI</collectionCode>
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!,
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<collectionCode id="0EC009FD28304718ACD957C6FCB0F87D" box="[724,770,1926,1947]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="4" pageNumber="130" type="Herbarium">MO</collectionCode>
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!).
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</paragraph>
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<paragraph id="686E913828304719AE8557EAFE6BFF3F" blockId="4.[136,1452,1890,2091]" lastBlockId="5.[189,1452,160,217]" lastPageId="5" lastPageNumber="131" pageId="4" pageNumber="130">
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Diagnosis:—This species resembles
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<taxonomicName id="AFD1EABB28304718AFE157EAFB37F859" authority="(Hayata 1911: 428) Ching (1962: 257)" authorityName="Ching" authorityPageNumber="257" authorityYear="1962" baseAuthorityName="Hayata" baseAuthorityPageNumber="428" baseAuthorityYear="1911" box="[492,1157,1962,1983]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="130" phylum="Tracheophyta" rank="species" species="dimorphophylla">
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<emphasis id="5AA54D2A28304718AFE157EAFCB4F859" box="[492,774,1962,1983]" italics="true" pageId="4" pageNumber="130">Arachniodes dimorphophylla</emphasis>
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(
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<bibRefCitation id="0C40ECC928304718AD1957EAFC71F859" author="Hayata, B." box="[788,963,1962,1983]" pageId="4" pageNumber="130" pagination="1 - 471" refId="ref3830" refString="Hayata, B. (1911) Materials for a flora of Formosa. Journal of the College of Science, Imperial University of Tokyo 30 (1): 1 - 471." type="journal article" year="1911">Hayata 1911: 428</bibRefCitation>
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) Ching (1962: 257)
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</taxonomicName>
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in having fronds dimorphous, fertile fronds lanceolate, longer but narrower than sterile fronds, sterile fronds ovate-lanceolate, and sori small (
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||
<quantity id="AF293CDD28304718AAE1578EFAFDF805" box="[1260,1359,1998,2019]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="130" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
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in diam.) and closer to midribs than to segment margins, but the former bears rhizomes suberect or ascending (creeping rhizomes unseen), laminae more shallowly dissected, often 2-pinnate-pinnatifid, basal basiscopic segments of lowest pinnae less elongated, and indusia ciliate; in contrast, the latter has rhizomes often short-creeping, laminae 3-pinnate, basal basiscopic segments of lowest pinnae longelongated, and indusia entire.
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</paragraph>
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<paragraph id="686E913828314719AE855149FB9FFC84" blockId="5.[136,1452,265,1262]" pageId="5" pageNumber="131">
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Rhizome erect or ascending, 1.2–4.2 ×
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<quantity id="AF293CDD28314719AC455149FD78FEC4" box="[584,714,265,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="5" pageNumber="131" unit="cm" value="1.85" valueMax="2.5" valueMin="1.2">1.2–2.5 cm</quantity>
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, with remnant bases of old stipes, up to 18 fronds each rhizome; ca. 35 roots,
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<quantity id="AF293CDD28314719AF11516DFECDFEA0" box="[284,383,301,326]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="5" pageNumber="131" unit="cm" value="6.5" valueMax="10.0" valueMin="3.0">3–10 cm</quantity>
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×
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<quantity id="AF293CDD28314719AFAD516DFD98FEA0" box="[416,554,301,326]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="5" pageNumber="131" unit="mm" value="0.5" valueMax="0.7" valueMin="0.3">0.3–0.7 mm</quantity>
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, apex densely scaly; scales (brown to red-brown), linear-narrow lanceolate, tip tortuous, 4–10 × 0.3–1.0 mm, entire, membranous. Fronds dimorphic,
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||
<quantity id="AF293CDD28314719ADBC5111FB94FE8C" box="[945,1062,337,362]" metricMagnitude="-1" metricUnit="m" metricValue="4.8" metricValueMax="7.5" metricValueMin="2.1" pageId="5" pageNumber="131" unit="cm" value="48.0" valueMax="75.0" valueMin="21.0">21–75 cm</quantity>
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. Sterile lamina 14–32 ×
|
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<quantity id="AF293CDD28314719AB4C5111FA15FE8C" box="[1345,1447,337,362]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.8" metricValueMin="0.5" pageId="5" pageNumber="131" unit="cm" value="11.5" valueMax="18.0" valueMin="5.0">5–18 cm</quantity>
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, stipe
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||
<quantity id="AF293CDD28314719AECB5135FE9AFE68" box="[198,296,373,398]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" metricValueMax="2.9" metricValueMin="0.7" pageId="5" pageNumber="131" unit="cm" value="18.0" valueMax="29.0" valueMin="7.0">7–29 cm</quantity>
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×
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<quantity id="AF293CDD28314719AF475135FE7FFE68" box="[330,461,373,398]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="12.0" metricValueMin="3.2" pageId="5" pageNumber="131" unit="mm" value="7.6" valueMax="12.0" valueMin="3.2">3.2–12 mm</quantity>
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, base with scales similar to those on rhizome, scales sparser upward, blackish, thread– linear, 4–6 ×
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<quantity id="AF293CDD28314719AF2451D9FE76FE54" box="[297,452,409,434]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="6.0" metricValueMin="1.2" pageId="5" pageNumber="131" unit="mm" value="0.36" valueMax="0.6" valueMin="0.12">0.12–0.6 mm</quantity>
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, entire and thicker; lamina 2 pinnate-pinnatifid, green when dried, deltoid-lanceolate, papery, base acute, apex acuminate; rachis scaly as distal portion of stipe, sparse scales adaxially; pinnae 12–22 pairs, lowest pinnae 4–14 ×
|
||
<quantity id="AF293CDD28314719AFC051A1FD88FE1C" box="[461,570,481,506]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.5" metricValueMin="2.0" pageId="5" pageNumber="131" unit="cm" value="3.75" valueMax="5.5" valueMin="2.0">2–5.5 cm</quantity>
|
||
, stalks ca.
|
||
<quantity id="AF293CDD28314719ACB451A1FCB8FE1C" box="[697,778,481,506]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="131" unit="cm" value="0.7">0.7 cm</quantity>
|
||
, upper shorter even sessile, alternate, spreading, lanceolate or deltoid-lanceolate, base rounded or broadly cuneate, apex acute or obtuse; pinnules 5–14 pairs per pinna, shortly stalked, oblong, deltoid-lanceolate, sometimes elliptical-oval, 1.0–1.5 ×
|
||
<quantity id="AF293CDD28314719ADC05269FBE3FDA4" box="[973,1105,553,578]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="3.8" metricValueMin="1.5" pageId="5" pageNumber="131" unit="cm" value="2.65" valueMax="3.8" valueMin="1.5">1.5–3.8 cm</quantity>
|
||
, base acroscopically rounded, obtuse, cuneate basiscopically cuneate, apex obtuse, margin often aristate; costule, vein sparse minute brown filiform scales abaxially. Fertile fronds up to 75 ×
|
||
<quantity id="AF293CDD28314719AC615231FD61FD6C" box="[620,723,625,650]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="5" pageNumber="131" unit="cm" value="8.0" valueMax="12.0" valueMin="4.0">4–12 cm</quantity>
|
||
; lamina similar to that of sterile fronds in shape and dissection, often longer than sterile but narrower, papery to subleathery; pinnae 12–24 pairs, lowest pinnae 4–11.2 ×
|
||
<quantity id="AF293CDD28314719AB2552D5FA1AFD48" box="[1320,1448,661,686]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="5" pageNumber="131" unit="cm" value="2.5" valueMax="3.2" valueMin="1.8">1.8–3.2 cm</quantity>
|
||
, lanceolate, alternate, base acute, oblique-cordate, apex acute or acuminate; pinnule 8–13 pairs, lanceolate or oblong, 0.5–1.0 × 1.0–
|
||
<quantity id="AF293CDD28314719AF3E529DFE34FD10" box="[307,390,733,758]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="131" unit="cm" value="2.8">2.8 cm</quantity>
|
||
, base acroscopically truncate, obtuse, basiscopically cuneate, apex obtuse, margin lobed or pinnatifid, mucronate teeth. Sori small (ca. 2.0 mm), terminal on veinlets, approximate, 2–9 pairs per ultimate segment, closer to midvein than margin; spores elliptical
|
||
<typeStatus id="B76A2F9A28314719ACAA5365FD68FCD8" box="[679,730,805,830]" pageId="5" pageNumber="131">type</typeStatus>
|
||
, 31–38 × 35–46 um, inflated, echinate folder; indusia brown to dark brown, membranous, ciliate or irregularly denticulate on margin (ca.
|
||
<quantity id="AF293CDD28314719ADCA5309FB93FC84" box="[967,1057,841,866]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="131" unit="mm" value="2.4">2.4 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20CBC2B328314719AEB0532DFCA3FC4C" pageId="5" pageNumber="131" type="etymology">
|
||
<paragraph id="686E913828314719AEB0532DFCA3FC4C" blockId="5.[136,1452,265,1262]" pageId="5" pageNumber="131">
|
||
<emphasis id="5AA54D2A28314719AEB0532DFED6FC60" bold="true" box="[189,356,877,902]" pageId="5" pageNumber="131">Etymology:—</emphasis>
|
||
From
|
||
<emphasis id="5AA54D2A28314719AFA7532DFD92FC60" box="[426,544,877,902]" italics="true" pageId="5" pageNumber="131">quangnam</emphasis>
|
||
, the province name in central
|
||
<collectingCountry id="10C6D1A828314719AD88532DFC55FC60" box="[901,999,877,902]" name="Vietnam" pageId="5" pageNumber="131">Vietnam</collectingCountry>
|
||
, and the Latin ending
|
||
<emphasis id="5AA54D2A28314719AAE3532DFA82FC60" box="[1262,1328,877,902]" italics="true" pageId="5" pageNumber="131">-ensis</emphasis>
|
||
, of origin, referring to the
|
||
<typeStatus id="B76A2F9A28314719AF3A53D1FED5FC4C" box="[311,359,913,938]" pageId="5" pageNumber="131">type</typeStatus>
|
||
locality where we collected the plant.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20CBC2B328314719AEB053F5FAA7FB08" pageId="5" pageNumber="131" type="distribution">
|
||
<paragraph id="686E913828314719AEB053F5FF5DFC14" blockId="5.[136,1452,265,1262]" pageId="5" pageNumber="131">
|
||
<emphasis id="5AA54D2A28314719AEB053F5FEC7FC28" bold="true" box="[189,373,949,974]" pageId="5" pageNumber="131">Distribution:—</emphasis>
|
||
This species is currently only known from the
|
||
<typeStatus id="B76A2F9A28314719AD8553F5FC0AFC28" box="[904,952,949,974]" pageId="5" pageNumber="131">type</typeStatus>
|
||
locality and probably is endemic to Central
|
||
<collectingCountry id="10C6D1A828314719AE855399FF59FC14" box="[136,235,985,1010]" name="Vietnam" pageId="5" pageNumber="131">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="686E913828314719AEB053BDFDC2FB40" blockId="5.[136,1452,265,1262]" pageId="5" pageNumber="131">
|
||
<emphasis id="5AA54D2A28314719AEB053BDFE94FBF0" bold="true" box="[189,294,1021,1046]" pageId="5" pageNumber="131">Notes:—</emphasis>
|
||
In addition to
|
||
<taxonomicName id="AFD1EABB28314719AFC053BDFCA0FBF0" authorityName="Li Bing Zhang, Ngan Thi Lu & Liang Zhang" authorityYear="2018" box="[461,786,1021,1046]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="131" phylum="Tracheophyta" rank="species" species="quangnamensis">
|
||
<emphasis id="5AA54D2A28314719AFC053BDFCA0FBF0" box="[461,786,1021,1046]" italics="true" pageId="5" pageNumber="131">Arachniodes quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
, there are two more species in the genus with dimorphic fronds:
|
||
<taxonomicName id="AFD1EABB28314719AEE95461FCF3FBDC" authority="(Ching 1934: 44) Ching (1962: 258)" authorityName="Ching" authorityPageNumber="258" authorityYear="1962" baseAuthorityName="Ching" baseAuthorityPageNumber="44" baseAuthorityYear="1934" box="[228,833,1057,1082]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="131" phylum="Tracheophyta" rank="species" species="hainanenis">
|
||
<emphasis id="5AA54D2A28314719AEE95461FE31FBDC" box="[228,387,1057,1082]" italics="true" pageId="5" pageNumber="131">A. hainanenis</emphasis>
|
||
(Ching 1934: 44) Ching (1962: 258)
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="B76A2F9A28314719AD585461FC39FBDC" box="[853,907,1057,1082]" pageId="5" pageNumber="131">type</typeStatus>
|
||
:
|
||
<collectingRegion id="AA155FDA28314719AD975461FC5CFBDC" box="[922,1006,1057,1082]" country="China" name="Hainan" pageId="5" pageNumber="131">Hainan</collectingRegion>
|
||
,
|
||
<collectingCountry id="10C6D1A828314719ADF25461FBF6FBDC" box="[1023,1092,1057,1082]" name="China" pageId="5" pageNumber="131">China</collectingCountry>
|
||
) and
|
||
<taxonomicName id="AFD1EABB28314719AA815461FAD1FBDC" authorityName="Ching" authorityYear="1962" baseAuthorityName="Hayata" baseAuthorityYear="1911" box="[1164,1379,1057,1082]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="131" phylum="Tracheophyta" rank="species" species="dimorphophylla">
|
||
<emphasis id="5AA54D2A28314719AA815461FAD1FBDC" box="[1164,1379,1057,1082]" italics="true" pageId="5" pageNumber="131">A. dimorphophylla</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="B76A2F9A28314719AB7F5461FA1AFBDC" box="[1394,1448,1057,1082]" pageId="5" pageNumber="131">type</typeStatus>
|
||
:
|
||
<collectingRegion id="AA155FDA28314719AE855405FF42FBB8" box="[136,240,1093,1118]" country="Japan" name="Okinawa" pageId="5" pageNumber="131">Okinawa</collectingRegion>
|
||
,
|
||
<collectingCountry id="10C6D1A828314719AEF35405FEF1FBB8" box="[254,323,1093,1118]" name="Japan" pageId="5" pageNumber="131">Japan</collectingCountry>
|
||
). Phylogenetically,
|
||
<taxonomicName id="AFD1EABB28314719AC2F5405FD43FBB8" authorityName="Li Bing Zhang, Ngan Thi Lu & Liang Zhang" authorityYear="2018" box="[546,753,1093,1118]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="131" phylum="Tracheophyta" rank="species" species="quangnamensis">
|
||
<emphasis id="5AA54D2A28314719AC2F5405FD43FBB8" box="[546,753,1093,1118]" italics="true" pageId="5" pageNumber="131">A. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
was resolved as sister to
|
||
<taxonomicName id="AFD1EABB28314719AA105405FB76FBB8" authorityName="Ching" authorityYear="1962" baseAuthorityName="Ching" baseAuthorityYear="1934" box="[1053,1220,1093,1118]" class="Polypodiopsida" family="Dryopteridaceae" genus="Arachniodes" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="131" phylum="Tracheophyta" rank="species" species="hainanensis">
|
||
<emphasis id="5AA54D2A28314719AA105405FB76FBB8" box="[1053,1220,1093,1118]" italics="true" pageId="5" pageNumber="131">A. hainanensis</emphasis>
|
||
</taxonomicName>
|
||
, while the Japanese dimorphic species was nested in other clade (Lu
|
||
<emphasis id="5AA54D2A28314719ACA85429FD6FFB64" box="[677,733,1129,1154]" italics="true" pageId="5" pageNumber="131">et al.</emphasis>
|
||
, submitted).
|
||
<bibRefCitation id="0C40ECC928314719AD7D5429FBADFB64" author="He, H. & Wu, S. - G. & Xiang, J. - Y. & Barrington, D. S." box="[880,1055,1129,1154]" pageId="5" pageNumber="131" pagination="542 - 558" refId="ref3960" refString="He, H., Wu, S. - G., Xiang, J. - Y. & Barrington, D. S. (2013) Arachniodes Blume. In: Wu, Z. - Y., Raven, P. H. & Hong, D. - Y. (Eds.) Flora of China, Vols. 2 - 3. pp. 542 - 558." type="book chapter" year="2013">
|
||
He
|
||
<emphasis id="5AA54D2A28314719AD9A5429FC7CFB64" box="[919,974,1129,1154]" italics="true" pageId="5" pageNumber="131">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
noted a dimorphic species reported from
|
||
<collectingCountry id="10C6D1A828314719AEC854CDFEA8FB40" box="[197,282,1165,1190]" name="Taiwan" pageId="5" pageNumber="131">Taiwan</collectingCountry>
|
||
, which needs to be confirmed.
|
||
</paragraph>
|
||
<paragraph id="686E913828314719AEB054F1FAA7FB08" blockId="5.[136,1452,265,1262]" pageId="5" pageNumber="131">
|
||
<emphasis id="5AA54D2A28314719AEB054F1FDF8FB2C" bold="true" box="[189,586,1201,1226]" pageId="5" pageNumber="131">Additional material examined:—</emphasis>
|
||
<collectingCountry id="10C6D1A828314719AC4754F1FD7DFB2C" box="[586,719,1201,1226]" name="Vietnam" pageId="5" pageNumber="131">VIETNAM</collectingCountry>
|
||
. Quangnam Province: Songthanh Nature Reserve, elev.
|
||
<quantity id="AF293CDD28314719AB4454F1FF12FB08" metricMagnitude="2" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="5" pageNumber="131" unit="m" value="430.0" valueMax="460.0" valueMin="400.0">400–460 m</quantity>
|
||
,
|
||
<date id="1C6FB7F828314719AEA15495FEC4FB08" box="[172,374,1237,1262]" pageId="5" pageNumber="131" value="2014-12-05">5 December 2014</date>
|
||
,
|
||
<emphasis id="5AA54D2A28314719AF8C5495FC45FB08" box="[385,1015,1237,1262]" italics="true" pageId="5" pageNumber="131">Li Bing Zhang, Liang Zhang & Ngan Thi Lu 7915, 1918</emphasis>
|
||
(CDBI!, MO!, VNMN!).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |