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<document id="F54CC31E2E50413BFADD2C4A0C345A06" ID-DOI="10.11646/zootaxa.5523.4.5" ID-GBIF-Dataset="c8fd3aa9-6203-4d1b-ae7b-367bf1d498b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13949669" ID-ZooBank="999559E0-4E12-45DC-AED9-88374AA3E3D1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729236181491" checkinUser="plazi" docAuthor="Kuwahara, Gregory K., Marshall, Stephen A. &amp; Yau, Tiffany" docDate="2024" docId="03A1976C12663A7973CA6803FAD0F459" docLanguage="en" docName="Zootaxa.5523.4.5.pdf" docOrigin="Zootaxa 5523 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helicosina valga Kuwahara &amp; Marshall &amp; Yau 2024, sp. nov." docType="treatment" docVersion="8" lastPageNumber="470" masterDocId="FF98EF14126C3A72735D6E59FFA1F664" masterDocTitle="Helicosina, a new genus of Neotropical Limosininae (Diptera: Sphaeroceridae)" masterLastPageNumber="471" masterPageNumber="459" pageNumber="469" updateTime="1729684655948" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8DD61946C4ABEDC802A6E596662126B7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7CCFDAC258DB053F6A12E20F93014BC4">
<mods:title id="E3A2E3C45F0DCB56AC2B964173D3C197">Helicosina, a new genus of Neotropical Limosininae (Diptera: Sphaeroceridae)</mods:title>
</mods:titleInfo>
<mods:name id="56BCB8303683BDC140F1325F8C21ACA7" type="personal">
<mods:role id="8D0E80B44E757E86BE9F8572897D9D2C">
<mods:roleTerm id="48D5B0152F61AA3F784430401AC9CBEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1582EF477FDBFEC0CA2FCEB763F64DCB">Kuwahara, Gregory K.</mods:namePart>
<mods:nameIdentifier id="49FB6142410492DE0BF514D293F617E0" type="email">gkuwahar@uoguelph.ca</mods:nameIdentifier>
</mods:name>
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<mods:role id="A150A3B610426C3728F07645510206FD">
<mods:roleTerm id="AAD5C0E9EB5E21D884463BB9CF3E0683">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="504C9993410EC9AAE1425AEA9594B3C2">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="8991132E1C6DFA45F5E52DE299F3239C" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="6CC622F8C172C7315B66796B05D9644E" type="personal">
<mods:role id="58645D748E84D8C652B0BBA57901AD3A">
<mods:roleTerm id="4FF035B4F48D53BAB1C737266A1976CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6FB96988590537437B28A7F0F6AE5E81">Yau, Tiffany</mods:namePart>
<mods:nameIdentifier id="BF4CD9F0EAB10507F77335B2BC9E6B89" type="email">tyau@alumni.uoguelph.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="358CD9850F0AFA4A28D4652588A5C04C">text</mods:typeOfResource>
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<mods:title id="A97409171781B0DB80E5AB97935FBD3E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BE67016D52BB3F6F30048B3A0A7E33C8">
<mods:date id="8C9ABA04C4CE0503DD4B74A017EEF5CC">2024</mods:date>
<mods:detail id="E9C0AD6BA6E5A4098A380816F3BCB532" type="pubDate">
<mods:number id="C43453EC3E26AB883817B6D3FF83BC76">2024-10-17</mods:number>
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<mods:detail id="32F99A142BA5B43FAF4AEA36E5A8FB41" type="volume">
<mods:number id="63C9D918D715694E5A64C33EE673FC7C">5523</mods:number>
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<mods:detail id="42B0F93B169BC6501D4A3A56E2FB7CDA" type="issue">
<mods:number id="F9D688944B9A6113477C6E1ADF626945">4</mods:number>
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<mods:classification id="79F709FA5E6326EA2676AB256AB392AA">journal article</mods:classification>
<mods:identifier id="B9CD359AC4EFBEDA1116C7AD5525B066" type="DOI">10.11646/zootaxa.5523.4.5</mods:identifier>
<mods:identifier id="55E03A080B558C52175460283D0E5397" type="GBIF-Dataset">c8fd3aa9-6203-4d1b-ae7b-367bf1d498b6</mods:identifier>
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<treatment id="03A1976C12663A7973CA6803FAD0F459" ID-DOI="http://doi.org/10.5281/zenodo.13949733" ID-GBIF-Taxon="244134453" ID-Zenodo-Dep="13949733" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459" httpUri="http://treatment.plazi.org/id/03A1976C12663A7973CA6803FAD0F459" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469">
<subSubSection id="C31275F112663A7873CA6803FDE1F011" box="[151,576,1626,1653]" pageId="10" pageNumber="469" type="nomenclature">
<paragraph id="8BB7267A12663A7873CA6803FDE1F011" blockId="10.[151,576,1626,1689]" box="[151,576,1626,1653]" pageId="10" pageNumber="469">
<heading id="D0FF911612663A7873CA6803FDE1F011" bold="true" box="[151,576,1626,1653]" fontSize="11" level="1" pageId="10" pageNumber="469" reason="1">
<emphasis id="B97CFA6812663A7873CA6803FDE1F011" bold="true" box="[151,576,1626,1653]" pageId="10" pageNumber="469">
<taxonomicName id="4C085DF912663A7873CA6803FE7AF011" authority="Marshall" authorityName="Kuwahara &amp; Marshall &amp; Yau" authorityYear="2024" box="[151,475,1626,1653]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="469" phylum="Arthropoda" rank="species" species="valga" status="sp. nov.">
<emphasis id="B97CFA6812663A7873CA6803FEC3F011" bold="true" box="[151,354,1626,1653]" italics="true" pageId="10" pageNumber="469">Helicosina valga</emphasis>
Marshall
</taxonomicName>
<taxonomicNameLabel id="A24F471312663A7872BE6802FDE1F011" box="[483,576,1627,1653]" pageId="10" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31275F112663A7973CA6826FA9DF69D" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469" type="description">
<paragraph id="8BB7267A12663A7873CA6826FE28F0FD" blockId="10.[151,576,1626,1689]" box="[151,393,1663,1689]" pageId="10" pageNumber="469">
(
<figureCitation id="13333AFF12663A7873C26826FEBDF0FD" box="[159,284,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="10" pageNumber="469">Figs. 2E–F</figureCitation>
,
<figureCitation id="13333AFF12663A78727A6826FEEBF0FD" box="[295,330,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982427" httpUri="https://zenodo.org/record/13982427/files/figure.png" pageId="10" pageNumber="469">3C</figureCitation>
,
<figureCitation id="13333AFF12663A78720B6826FEC7F0FD" box="[342,358,1663,1689]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="10" pageNumber="469">6</figureCitation>
,
<figureCitation id="13333AFF12663A78722F6826FE21F0FD" box="[370,384,1663,1689]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13982439" httpUri="https://zenodo.org/record/13982439/files/figure.png" pageId="10" pageNumber="469">8</figureCitation>
)
</paragraph>
<paragraph id="8BB7267A12663A7873CA689FFC30F085" blockId="10.[151,1436,1734,2013]" box="[151,913,1734,1761]" pageId="10" pageNumber="469">
<emphasis id="B97CFA6812663A7873CA689FFE88F084" bold="true" box="[151,297,1734,1760]" pageId="10" pageNumber="469">Description.</emphasis>
<emphasis id="B97CFA6812663A78726D689EFE17F085" box="[304,438,1735,1761]" italics="true" pageId="10" pageNumber="469">Body length</emphasis>
: Males
<quantity id="4CF08B9F12663A78714D689EFD38F085" box="[528,665,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="10" pageNumber="469" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
, females 2.0–
<quantity id="4CF08B9F12663A787069689EFC2CF085" box="[820,909,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="469" unit="mm" value="2.2">2.2 mm</quantity>
.
</paragraph>
<paragraph id="8BB7267A12663A78739A68B2FE91F14D" blockId="10.[151,1436,1734,2013]" pageId="10" pageNumber="469">
<emphasis id="B97CFA6812663A78739A68B2FE82F161" box="[199,291,1771,1797]" italics="true" pageId="10" pageNumber="469">General</emphasis>
(
<figureCitation id="13333AFF12663A78726A68B2FE32F161" box="[311,403,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="10" pageNumber="469">Fig. 2E</figureCitation>
): Male fore and hind femora strongly thickened (length 2.9–3.0 × and 3.5–3.7 × width, respectively).
</paragraph>
<paragraph id="8BB7267A12663A79739A696AFA9DF69D" blockId="10.[151,1436,1734,2013]" lastBlockId="11.[151,1436,151,573]" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469">
<emphasis id="B97CFA6812663A78739A696AFED0F129" box="[199,369,1843,1869]" italics="true" pageId="10" pageNumber="469">Male abdomen</emphasis>
(
<figureCitation id="13333AFF12663A7872DE696AFE6AF129" box="[387,459,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="10" pageNumber="469">Fig. 6</figureCitation>
): S5 broad, gently curved, broadly desclerotized posteromedially, and densely setose laterally. Posterior opening of epandrium oval (medial height 1.3–1.4 × medial width); cercus separated medially, forming a weakly-developed subanal plate. Surstylus strongly sinuate, basal half broad and subtriangular with the apex curved posteriorly, apical half strongly bent downwards and constricted with several anterior setae. Postgonite small, broad, and apically tapered with a small subapical patch of microtomentum. Phallapodeme elongate and well sclerotized. Basiphallus stout, slightly curved, with a stout, apically pointed epiphallus; distiphallus stout, largely membranous but including a trough-shaped basal sclerite with upper surface bent inwards to form a “roof”, a shieldshaped upper sclerite, and paired elongate lower-lateral sclerites; lateral membrane densely microtrichose.
</paragraph>
</subSubSection>
<subSubSection id="C31275F112673A79739A6F5BFBD2F47D" pageId="11" pageNumber="470" type="materials_examined">
<paragraph id="8BB7267A12673A79739A6F5BFC7CF725" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="470">
<emphasis id="B97CFA6812673A79739A6F5BFE11F778" bold="true" box="[199,432,258,284]" pageId="11" pageNumber="470">Material examined.</emphasis>
<materialsCitation id="3B602C2712673A7972EB6F5AFC7CF725" ID-GBIF-Occurrence="4958089310" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" county="Rio San Lorencito" elevation="900" location="Curso" municipality="San Ramon Forest Reserve" pageId="11" pageNumber="470" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
<typeStatus id="54B398D812673A7972EB6F5AFD82F779" box="[438,547,259,285]" pageId="11" pageNumber="470" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F31F66EA12673A7971716F5AFD70F779" box="[556,721,259,285]" name="Costa Rica" pageId="11" pageNumber="470">COSTA RICA</collectingCountry>
:
<emphasis id="B97CFA6812673A7971866F5BFCEBF778" bold="true" box="[731,842,258,284]" pageId="11" pageNumber="470">
<collectingRegion id="49CCE89812673A7971866F5BFCE4F778" box="[731,837,258,284]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="470">Alajuela</collectingRegion>
:
</emphasis>
<collectingCounty id="62D65EF612673A79700D6F5AFBBEF779" box="[848,1055,259,285]" pageId="11" pageNumber="470">Rio San Lorencito</collectingCounty>
,
<collectingMunicipality id="6BD3BC0012673A7977746F5AFAFBF779" box="[1065,1370,259,285]" pageId="11" pageNumber="470">San Ramón Forest Reserve</collectingMunicipality>
,
<locationDeviation id="9A053F5812673A7976396F5BFE2AF725" location="Colonia Palmarena" pageId="11" pageNumber="470">
<quantity id="4CF08B9F12673A7976396F5BFA3DF779" box="[1380,1436,258,285]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="470" unit="km" value="5.0">5 km</quantity>
N Colonia Palmareña
</locationDeviation>
,
<elevation id="0025C14912673A7972CB6F7EFDBBF725" box="[406,538,295,321]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="470" unit="m" value="900.0">
<quantity id="4CF08B9F12673A7972CB6F7EFE7CF725" box="[406,477,295,321]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="470" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB600BA12673A79717B6F7EFDDEF725" box="[550,639,295,321]" pageId="11" pageNumber="470" value="1990-03">
<collectingDate id="EFF2F95212673A79717B6F7EFDDEF725" box="[550,639,295,321]" pageId="11" pageNumber="470" value="1990-03">iii.1990</collectingDate>
</date>
,
<location id="8ED770A112673A7971D66F7EFD6FF725" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A7971D66F7EFD6FF725" box="[651,718,295,321]" country="Costa Rica" county="Rio San Lorencito" municipality="San Ramon Forest Reserve" name="Curso" pageId="11" pageNumber="470" stateProvince="Alajuela">Curso</location>
<taxonomicName id="4C085DF912673A7971896F7EFCE7F725" authorityName="Latreille" authorityYear="1802" box="[724,838,295,321]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="470" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
(
<specimenCount id="9D0EEDF312673A79700B6F7EFCCCF725" box="[854,877,295,321]" count="1" pageId="11" pageNumber="470" type="male">♂</specimenCount>
,
<collectionCode id="ED19BEBF12673A7970246F7EFC70F725" box="[889,977,295,321]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="470">MNCR</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB7267A12673A79739A6F12FCDCF791" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="470">
<materialsCitation id="3B602C2712673A79739A6F12FBB8F701" ID-GBIF-Occurrence="4958089312" box="[199,1049,330,357]" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" county="Rio San Lorencito" elevation="900" location="Curso" municipality="San Ramon Forest Reserve" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
<typeStatus id="54B398D812673A79739A6F12FE99F701" box="[199,312,331,357]" pageId="11" pageNumber="470" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F31F66EA12673A7972186F12FE4DF701" box="[325,492,331,357]" name="Costa Rica" pageId="11" pageNumber="470">COSTA RICA</collectingCountry>
:
<emphasis id="B97CFA6812673A7972A46F13FDC6F700" bold="true" box="[505,615,330,356]" pageId="11" pageNumber="470">
<collectingRegion id="49CCE89812673A7972A46F13FDC2F700" box="[505,611,330,356]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="470">Alajuela</collectingRegion>
:
</emphasis>
same data as holotype (
<specimenCount id="9D0EEDF312673A7970226F12FC08F701" box="[895,937,331,357]" count="1" pageId="11" pageNumber="470" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED19BEBF12673A7970E46F12FBB0F701" box="[953,1041,331,357]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="470">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B602C2712673A7977786F12FCE8F7ED" ID-GBIF-Occurrence="4958089319" collectingDate="1990-05" collectionCode="MNCR" collectorName="II Curso Parataxon" country="Costa Rica" elevation="600" location="Finca San Gabriel" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
<location id="8ED770A112673A7977786F12FB57F701" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A7977786F12FB57F701" box="[1061,1270,331,357]" country="Costa Rica" name="Finca San Gabriel" pageId="11" pageNumber="470" stateProvince="Alajuela">Finca San Gabriel</location>
,
<locationDeviation id="9A053F5812673A79765F6F12FF6CF7ED" location="Dos Rios" pageId="11" pageNumber="470">
<quantity id="4CF08B9F12673A79765F6F12FA9CF701" box="[1282,1341,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="470" unit="km" value="2.0">2 km</quantity>
SE Dos Rios
</locationDeviation>
,
<elevation id="0025C14912673A7973876F36FEFEF7ED" box="[218,351,367,393]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="470" unit="m" value="600.0">
<quantity id="4CF08B9F12673A7973876F36FE83F7ED" box="[218,290,367,393]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="470" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB600BA12673A7972306F36FE1AF7ED" box="[365,443,367,393]" pageId="11" pageNumber="470" value="1990-05">
<collectingDate id="EFF2F95212673A7972306F36FE1AF7ED" box="[365,443,367,393]" pageId="11" pageNumber="470" value="1990-05">v.1990</collectingDate>
</date>
,
<collectorName id="26FD43AC12673A79729A6F36FD3CF7ED" box="[455,669,367,393]" pageId="11" pageNumber="470">II Curso Parataxon</collectorName>
(
<specimenCount id="9D0EEDF312673A7971ED6F36FD7BF7ED" box="[688,730,367,393]" count="1" pageId="11" pageNumber="470" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED19BEBF12673A7971B46F36FCE0F7ED" box="[745,833,367,393]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="470">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B602C2712673A7970086F36FB8BF7C9" ID-GBIF-Occurrence="4958089313" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="DEBU, MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="11" pageNumber="470" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B97CFA6812673A7970086F36FC46F7ED" bold="true" box="[853,999,367,393]" pageId="11" pageNumber="470">
<collectingRegion id="49CCE89812673A7970086F36FC43F7ED" box="[853,994,367,393]" country="Costa Rica" name="Puntarenas" pageId="11" pageNumber="470">Puntarenas</collectingRegion>
:
</emphasis>
<location id="8ED770A112673A7970B26F36FAF9F7ED" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A7970B26F36FAF9F7ED" box="[1007,1368,367,393]" country="Costa Rica" name="Monteverde Biological Reserve" pageId="11" pageNumber="470" stateProvince="Puntarenas">Monteverde Biological Reserve</location>
,
<elevation id="0025C14912673A7976396F37FF4DF7C9" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="470" unit="m" value="1500.0">
<quantity id="4CF08B9F12673A7976396F37FF0CF7C9" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="470" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
, cloud forest,
<date id="FFB600BA12673A7972CF6FCAFD97F7C9" box="[402,566,403,429]" pageId="11" pageNumber="470" value="2000-06-11" valueMax="2000-06-13" valueMin="2000-06-11">
<collectingDate id="EFF2F95212673A7972CF6FCAFD97F7C9" box="[402,566,403,429]" pageId="11" pageNumber="470" value="2000-06-11" valueMax="2000-06-13" valueMin="2000-06-11">11–13.vi.2000</collectingDate>
</date>
,
<collectorName id="26FD43AC12673A79711E6FCAFD43F7C9" box="[579,738,403,429]" pageId="11" pageNumber="470">S.A. Marshall</collectorName>
(
<specimenCount id="9D0EEDF312673A7971A96FCAFC82F7C9" box="[756,803,403,429]" count="2" pageId="11" pageNumber="470" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D0EEDF312673A79706C6FCAFCFFF7C9" box="[817,862,403,429]" count="1" pageId="11" pageNumber="470" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED19BEBF12673A7970336FCAFC1CF7C9" box="[878,957,403,429]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="11" pageNumber="470">DEBU</collectionCode>
,
<collectionCode id="ED19BEBF12673A7970976FCAFB83F7C9" box="[970,1058,403,429]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="470">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B602C2712673A79776A6FCAFBB5F7B5" ID-GBIF-Occurrence="4958089305" collectingDate="2000-06-13" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Station" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<location id="8ED770A112673A79776A6FCAFA39F7C9" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A79776A6FCAFA39F7C9" box="[1079,1432,403,429]" country="Costa Rica" name="Monteverde Biological Station" pageId="11" pageNumber="470" stateProvince="Puntarenas">Monteverde Biological Station</location>
,
<elevation id="0025C14912673A7973CA6FEFFE86F7B5" box="[151,295,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="470" unit="m" value="1500.0">
<quantity id="4CF08B9F12673A7973CA6FEFFF4AF7B5" box="[151,235,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="470" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
, white pans in kitchen compost,
<date id="FFB600BA12673A7971CD6FEEFCABF7B5" box="[656,778,439,465]" pageId="11" pageNumber="470" value="2000-06-13">
<collectingDate id="EFF2F95212673A7971CD6FEEFCABF7B5" box="[656,778,439,465]" pageId="11" pageNumber="470" value="2000-06-13">13.vi.2000</collectingDate>
</date>
,
<collectorName id="26FD43AC12673A7970486FEEFCD7F7B5" box="[789,886,439,465]" pageId="11" pageNumber="470">M. Buck</collectorName>
(
<specimenCount id="9D0EEDF312673A7970DA6FEEFC0FF7B5" box="[903,942,439,465]" count="1" pageId="11" pageNumber="470" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED19BEBF12673A7970E66FEEFBADF7B5" box="[955,1036,439,465]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="11" pageNumber="470">DEBU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B602C2712673A7977436FEEFCD9F791" ID-GBIF-Occurrence="4958089302" collectingDate="1974-07-05" collectionCode="CAS" collectorName="C. W. &amp; L. O'Brien &amp; Marshall" country="Panama" location="Cerro Campana" municipality="Oeste" pageId="11" pageNumber="470" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
<collectingCountry id="F31F66EA12673A7977436FEEFB38F7B5" box="[1054,1177,439,465]" name="Panama" pageId="11" pageNumber="470">PANAMA</collectingCountry>
:
<emphasis id="B97CFA6812673A7977FF6FEFFAF7F7B4" bold="true" box="[1186,1366,438,465]" pageId="11" pageNumber="470">
<collectingRegion id="49CCE89812673A7977FF6FEFFAA5F7B5" box="[1186,1284,438,465]" country="Panama" name="Panama" pageId="11" pageNumber="470">Panamá</collectingRegion>
<collectingMunicipality id="6BD3BC0012673A7976576FEFFAF0F7B4" box="[1290,1361,438,464]" pageId="11" pageNumber="470">Oeste</collectingMunicipality>
:
</emphasis>
<location id="8ED770A112673A7976016FEEFEA5F791" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A7976016FEEFEA5F791" country="Panama" municipality="Oeste" name="Cerro Campana" pageId="11" pageNumber="470" stateProvince="Panama">Cerro Campana</location>
,
<date id="FFB600BA12673A7972526F83FE22F791" box="[271,387,474,501]" pageId="11" pageNumber="470" value="1974-07-05">
<collectingDate id="EFF2F95212673A7972526F83FE22F791" box="[271,387,474,501]" pageId="11" pageNumber="470" value="1974-07-05">5.vii.1974</collectingDate>
</date>
,
<collectorName id="26FD43AC12673A7972D26F82FDC7F791" box="[399,614,475,501]" pageId="11" pageNumber="470">C.W. &amp; L. O’Brien</collectorName>
&amp;
<collectorName id="26FD43AC12673A7971D46F82FD4CF791" box="[649,749,475,501]" pageId="11" pageNumber="470">Marshall</collectorName>
(
<specimenCount id="9D0EEDF312673A7971A26F82FC88F791" box="[767,809,475,501]" count="1" pageId="11" pageNumber="470" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED19BEBF12673A7970686F82FCD1F791" box="[821,880,475,501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="470" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB7267A12673A79739A6FA6FBD2F47D" blockId="11.[151,1436,151,573]" box="[199,1139,510,537]" pageId="11" pageNumber="470">
<emphasis id="B97CFA6812673A79739A6FA6FEC7F47C" bold="true" box="[199,358,510,537]" pageId="11" pageNumber="470">Type locality.</emphasis>
<collectingCountry id="F31F66EA12673A7972316FA6FE48F47D" box="[364,489,511,537]" name="Costa Rica" pageId="11" pageNumber="470">Costa Rica</collectingCountry>
,
<collectingRegion id="49CCE89812673A7972AE6FA6FDF4F47D" box="[499,597,511,537]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="470">Alajuela</collectingRegion>
, Rio San Lorencito, San Ramón Forest Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C31275F112673A79739A6C7AFAD0F459" box="[199,1393,547,573]" pageId="11" pageNumber="470" type="etymology">
<paragraph id="8BB7267A12673A79739A6C7AFAD0F459" blockId="11.[151,1436,151,573]" box="[199,1393,547,573]" pageId="11" pageNumber="470">
<emphasis id="B97CFA6812673A79739A6C7AFEEFF459" bold="true" box="[199,334,547,573]" pageId="11" pageNumber="470">Etymology.</emphasis>
The specific name (from the Latin ‘valgus’: bent out) refers to the distinctively bent surstylus.
</paragraph>
</subSubSection>
</treatment>
</document>
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