Add updates up until 2024-10-23 11:57:51
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<mods:namePart id="F0F44C9C3280CD8603F84886A4641443">Marshall, Stephen A.</mods:namePart>
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<mods:namePart id="CD3008F3C5F6DC8CE995AFCEBB3A7E3C">Yau, Tiffany</mods:namePart>
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@ -75,15 +73,15 @@ Marshall
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<figureCitation id="13333AFF12663A7873C26D1AFF4CF539" box="[159,237,835,861]" captionStart="FIGURE 1" captionStartId="2.[152,255,1795,1819]" captionTargetBox="[151,1435,306,1769]" captionTargetId="figure-76@2.[151,1435,306,1769]" captionTargetPageId="2" captionText="FIGURE 1. Helicosina sica gen. et sp. nov., living male, Tiputini Biological Station, Ecuador." pageId="10" pageNumber="469">Figs. 1</figureCitation>
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<figureCitation id="13333AFF12663A7873C26D1AFF4CF539" box="[159,237,835,861]" captionStart="FIGURE 1" captionStartId="2.[152,255,1795,1819]" captionTargetBox="[151,1435,306,1769]" captionTargetId="figure-76@2.[151,1435,306,1769]" captionTargetPageId="2" captionText="FIGURE 1. Helicosina sica gen. et sp. nov., living male, Tiputini Biological Station, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.13982423" httpUri="https://zenodo.org/record/13982423/files/figure.png" pageId="10" pageNumber="469">Figs. 1</figureCitation>
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,
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<figureCitation id="13333AFF12663A7873A66D1AFEE1F539" box="[251,320,835,861]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="10" pageNumber="469">2C–D</figureCitation>
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<figureCitation id="13333AFF12663A7873A66D1AFEE1F539" box="[251,320,835,861]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="10" pageNumber="469">2C–D</figureCitation>
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,
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<figureCitation id="13333AFF12663A7872116D1AFECEF539" box="[332,367,835,861]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." pageId="10" pageNumber="469">3B</figureCitation>
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<figureCitation id="13333AFF12663A7872116D1AFECEF539" box="[332,367,835,861]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982427" httpUri="https://zenodo.org/record/13982427/files/figure.png" pageId="10" pageNumber="469">3B</figureCitation>
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,
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<figureCitation id="13333AFF12663A7872276D1BFE2BF539" box="[378,394,834,861]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="10" pageNumber="469">5</figureCitation>
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<figureCitation id="13333AFF12663A7872276D1BFE2BF539" box="[378,394,834,861]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="10" pageNumber="469">5</figureCitation>
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,
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<figureCitation id="13333AFF12663A7872CB6D1AFE05F539" box="[406,420,835,861]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." pageId="10" pageNumber="469">8</figureCitation>
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<figureCitation id="13333AFF12663A7872CB6D1AFE05F539" box="[406,420,835,861]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13982439" httpUri="https://zenodo.org/record/13982439/files/figure.png" pageId="10" pageNumber="469">8</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12663A7873CA6DD3FCF4F5C1" blockId="10.[151,1437,906,1545]" box="[151,853,906,933]" pageId="10" pageNumber="469">
|
||||
|
|
@ -98,20 +96,20 @@ Marshall
|
|||
<paragraph id="8BB7267A12663A78739A6DF6FE91F589" blockId="10.[151,1437,906,1545]" pageId="10" pageNumber="469">
|
||||
<emphasis id="B97CFA6812663A78739A6DF6FE82F5AD" box="[199,291,943,969]" italics="true" pageId="10" pageNumber="469">General</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF12663A78726B6DF6FE35F5AD" box="[310,404,943,969]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="10" pageNumber="469">Fig. 2C</figureCitation>
|
||||
<figureCitation id="13333AFF12663A78726B6DF6FE35F5AD" box="[310,404,943,969]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="10" pageNumber="469">Fig. 2C</figureCitation>
|
||||
): Male fore and hind femora strongly thickened (length 3.0–3.2 × and 3.1–3.5 × width, respectively).
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12663A78739A6DAEFD9AF369" blockId="10.[151,1437,906,1545]" pageId="10" pageNumber="469">
|
||||
<emphasis id="B97CFA6812663A78739A6DAEFECFF275" box="[199,366,1015,1041]" italics="true" pageId="10" pageNumber="469">Male abdomen</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF12663A7872266DAEFE1EF275" box="[379,447,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="10" pageNumber="469">Fig. 5</figureCitation>
|
||||
<figureCitation id="13333AFF12663A7872266DAEFE1EF275" box="[379,447,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="10" pageNumber="469">Fig. 5</figureCitation>
|
||||
): S5 broad, desclerotized posteromedially, and densely setose laterally. Posterior opening of epandrium circular (medial height subequal to medial width); subanal plate with a dark, sclerotized I-shaped medial pattern. Surstylus strongly sinuate, basal half bulbous and setulose externally, apical half strongly constricted with several setae anterobasally. Postgonite small, truncate apically. Phallapodeme relatively short and weakly developed. Basiphallus stout, slightly curved, with an elongate, apically pointed epiphallus; distiphallus stout, largely membranous and supported by several sclerites: a V-shaped posterior sclerite, a triangular upper sclerite, and paired V-shaped upper-lateral sclerites; lower surface grooved with scattered darkened spots; lateral membrane with an elongate row of stout setulae.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31275F112663A78739A6B4FFCAEF3A5" pageId="10" pageNumber="469" type="materials_examined">
|
||||
<paragraph id="8BB7267A12663A78739A6B4FFB80F331" blockId="10.[151,1437,906,1545]" pageId="10" pageNumber="469">
|
||||
<emphasis id="B97CFA6812663A78739A6B4FFE11F354" bold="true" box="[199,432,1302,1328]" pageId="10" pageNumber="469">Material examined.</emphasis>
|
||||
<materialsCitation id="3B602C2712663A7872EB6B4EFB80F331" collectingDate="2002-04-30" collectingDateMax="2002-05-07" collectingDateMin="2002-04-30" collectionCode="QCAZ" country="Ecuador" county="Jatun Sacha Reserve" elevation="450" location="Buck" municipality="Malaise" pageId="10" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
|
||||
<materialsCitation id="3B602C2712663A7872EB6B4EFB80F331" ID-GBIF-Occurrence="4958089307" collectingDate="2002-04-30" collectingDateMax="2002-05-07" collectingDateMin="2002-04-30" collectionCode="QCAZ" country="Ecuador" county="Jatun Sacha Reserve" elevation="450" location="Buck" municipality="Malaise" pageId="10" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
|
||||
<typeStatus id="54B398D812663A7872EB6B4EFD82F355" box="[438,547,1303,1329]" pageId="10" pageNumber="469" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<collectingCountry id="F31F66EA12663A7871706B4EFD18F355" box="[557,697,1303,1329]" name="Ecuador" pageId="10" pageNumber="469">ECUADOR</collectingCountry>
|
||||
|
|
@ -148,7 +146,7 @@ a.s.l.
|
|||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12663A78739A6B06FBCFF3F9" blockId="10.[151,1437,906,1545]" pageId="10" pageNumber="469">
|
||||
<materialsCitation id="3B602C2712663A78739A6B06FBC6F3F9" collectingDate="2011-05" collectionCode="DEBU, QCAZ" collectorName="S. A. Marshall" country="Ecuador" location="Tiputini Biodiversity Station" pageId="10" pageNumber="469" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Napo" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712663A78739A6B06FBC6F3F9" ID-GBIF-Occurrence="4958089304" collectingDate="2011-05" collectionCode="DEBU, QCAZ" collectorName="S. A. Marshall" country="Ecuador" location="Tiputini Biodiversity Station" pageId="10" pageNumber="469" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Napo" typeStatus="paratype">
|
||||
<typeStatus id="54B398D812663A78739A6B06FE99F31D" box="[199,312,1375,1401]" pageId="10" pageNumber="469" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<collectingCountry id="F31F66EA12663A78721F6B06FE6FF31D" box="[322,462,1375,1401]" name="Ecuador" pageId="10" pageNumber="469">ECUADOR</collectingCountry>
|
||||
|
|
@ -176,10 +174,8 @@ furl,
|
|||
<collectionCode id="ED19BEBF12663A7872E96BDAFDA2F3F9" box="[436,515,1411,1437]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="10" pageNumber="469">DEBU</collectionCode>
|
||||
,
|
||||
<collectionCode id="ED19BEBF12663A7871536BDAFDFEF3F9" box="[526,607,1411,1437]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="469" type="Museum">QCAZ</collectionCode>
|
||||
). One of the male
|
||||
<typeStatus id="54B398D812663A7870716BDAFC36F3F9" box="[812,919,1411,1437]" pageId="10" pageNumber="469" type="paratype">paratypes</typeStatus>
|
||||
is shown in
|
||||
<figureCitation id="13333AFF12663A7877786BDAFBC6F3F9" box="[1061,1127,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[152,255,1795,1819]" captionTargetBox="[151,1435,306,1769]" captionTargetId="figure-76@2.[151,1435,306,1769]" captionTargetPageId="2" captionText="FIGURE 1. Helicosina sica gen. et sp. nov., living male, Tiputini Biological Station, Ecuador." pageId="10" pageNumber="469">Fig. 1</figureCitation>
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||||
). One of the male paratypes is shown in
|
||||
<figureCitation id="13333AFF12663A7877786BDAFBC6F3F9" box="[1061,1127,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[152,255,1795,1819]" captionTargetBox="[151,1435,306,1769]" captionTargetId="figure-76@2.[151,1435,306,1769]" captionTargetPageId="2" captionText="FIGURE 1. Helicosina sica gen. et sp. nov., living male, Tiputini Biological Station, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.13982423" httpUri="https://zenodo.org/record/13982423/files/figure.png" pageId="10" pageNumber="469">Fig. 1</figureCitation>
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||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,64 +1,62 @@
|
|||
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<mods:extent id="D359BAD8A9586970A0317DC4C6DEE4A4" unit="page">
|
||||
<mods:start id="9A71790A3FCA25E311FB6246357894E4">459</mods:start>
|
||||
<mods:end id="543AE232B1F017E2AB01D753ECA2DF49">471</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="F8E6DA136861DC9A9CC279C55A637BD2">
|
||||
<mods:url id="B052384DE5C79A8523C02494957CED96">http://dx.doi.org/10.11646/zootaxa.5523.4.5</mods:url>
|
||||
<mods:location id="0BD8E07F348B9F1B7E268BE460E3EC5B">
|
||||
<mods:url id="78D7C0728F1BE71970927FE680554403">http://dx.doi.org/10.11646/zootaxa.5523.4.5</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="565B6BF008BEE34B3D5FD003259E435F">journal article</mods:classification>
|
||||
<mods:identifier id="4A27858C6BB80E9B62AD03427E8BABA9" type="DOI">10.11646/zootaxa.5523.4.5</mods:identifier>
|
||||
<mods:identifier id="7F40F40F5C42786EAC9EAADFF09493A4" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="2C93F4CC17001893FF07E2397D404254" type="Zenodo-Dep">13949669</mods:identifier>
|
||||
<mods:identifier id="2C7E5567C1C49C090749A6CF3B57DAEC" type="ZooBank">999559E0-4E12-45DC-AED9-88374AA3E3D1</mods:identifier>
|
||||
<mods:classification id="79F709FA5E6326EA2676AB256AB392AA">journal article</mods:classification>
|
||||
<mods:identifier id="B9CD359AC4EFBEDA1116C7AD5525B066" type="DOI">10.11646/zootaxa.5523.4.5</mods:identifier>
|
||||
<mods:identifier id="55E03A080B558C52175460283D0E5397" type="GBIF-Dataset">c8fd3aa9-6203-4d1b-ae7b-367bf1d498b6</mods:identifier>
|
||||
<mods:identifier id="0C2B58D96F29E065715710F95E98F06F" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="4639032784DEE1B5170E410E9CAC2749" type="Zenodo-Dep">13949669</mods:identifier>
|
||||
<mods:identifier id="5435294C1E12C69F909C75BE299ACBDC" type="ZooBank">999559E0-4E12-45DC-AED9-88374AA3E3D1</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03A1976C12663A7973CA6803FAD0F459" ID-Zenodo-Dep="13949733" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459" httpUri="http://treatment.plazi.org/id/03A1976C12663A7973CA6803FAD0F459" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469">
|
||||
<treatment id="03A1976C12663A7973CA6803FAD0F459" ID-DOI="http://doi.org/10.5281/zenodo.13949733" ID-GBIF-Taxon="244134453" ID-Zenodo-Dep="13949733" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459" httpUri="http://treatment.plazi.org/id/03A1976C12663A7973CA6803FAD0F459" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469">
|
||||
<subSubSection id="C31275F112663A7873CA6803FDE1F011" box="[151,576,1626,1653]" pageId="10" pageNumber="469" type="nomenclature">
|
||||
<paragraph id="8BB7267A12663A7873CA6803FDE1F011" blockId="10.[151,576,1626,1689]" box="[151,576,1626,1653]" pageId="10" pageNumber="469">
|
||||
<heading id="D0FF911612663A7873CA6803FDE1F011" bold="true" box="[151,576,1626,1653]" fontSize="11" level="1" pageId="10" pageNumber="469" reason="1">
|
||||
|
|
@ -75,13 +73,13 @@ Marshall
|
|||
<subSubSection id="C31275F112663A7973CA6826FA9DF69D" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469" type="description">
|
||||
<paragraph id="8BB7267A12663A7873CA6826FE28F0FD" blockId="10.[151,576,1626,1689]" box="[151,393,1663,1689]" pageId="10" pageNumber="469">
|
||||
(
|
||||
<figureCitation id="13333AFF12663A7873C26826FEBDF0FD" box="[159,284,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="10" pageNumber="469">Figs. 2E–F</figureCitation>
|
||||
<figureCitation id="13333AFF12663A7873C26826FEBDF0FD" box="[159,284,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="10" pageNumber="469">Figs. 2E–F</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12663A78727A6826FEEBF0FD" box="[295,330,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." pageId="10" pageNumber="469">3C</figureCitation>
|
||||
<figureCitation id="13333AFF12663A78727A6826FEEBF0FD" box="[295,330,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982427" httpUri="https://zenodo.org/record/13982427/files/figure.png" pageId="10" pageNumber="469">3C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12663A78720B6826FEC7F0FD" box="[342,358,1663,1689]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="10" pageNumber="469">6</figureCitation>
|
||||
<figureCitation id="13333AFF12663A78720B6826FEC7F0FD" box="[342,358,1663,1689]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="10" pageNumber="469">6</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12663A78722F6826FE21F0FD" box="[370,384,1663,1689]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." pageId="10" pageNumber="469">8</figureCitation>
|
||||
<figureCitation id="13333AFF12663A78722F6826FE21F0FD" box="[370,384,1663,1689]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13982439" httpUri="https://zenodo.org/record/13982439/files/figure.png" pageId="10" pageNumber="469">8</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12663A7873CA689FFC30F085" blockId="10.[151,1436,1734,2013]" box="[151,913,1734,1761]" pageId="10" pageNumber="469">
|
||||
|
|
@ -96,20 +94,20 @@ Marshall
|
|||
<paragraph id="8BB7267A12663A78739A68B2FE91F14D" blockId="10.[151,1436,1734,2013]" pageId="10" pageNumber="469">
|
||||
<emphasis id="B97CFA6812663A78739A68B2FE82F161" box="[199,291,1771,1797]" italics="true" pageId="10" pageNumber="469">General</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF12663A78726A68B2FE32F161" box="[311,403,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="10" pageNumber="469">Fig. 2E</figureCitation>
|
||||
<figureCitation id="13333AFF12663A78726A68B2FE32F161" box="[311,403,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="10" pageNumber="469">Fig. 2E</figureCitation>
|
||||
): Male fore and hind femora strongly thickened (length 2.9–3.0 × and 3.5–3.7 × width, respectively).
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12663A79739A696AFA9DF69D" blockId="10.[151,1436,1734,2013]" lastBlockId="11.[151,1436,151,573]" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469">
|
||||
<emphasis id="B97CFA6812663A78739A696AFED0F129" box="[199,369,1843,1869]" italics="true" pageId="10" pageNumber="469">Male abdomen</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF12663A7872DE696AFE6AF129" box="[387,459,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="10" pageNumber="469">Fig. 6</figureCitation>
|
||||
<figureCitation id="13333AFF12663A7872DE696AFE6AF129" box="[387,459,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="10" pageNumber="469">Fig. 6</figureCitation>
|
||||
): S5 broad, gently curved, broadly desclerotized posteromedially, and densely setose laterally. Posterior opening of epandrium oval (medial height 1.3–1.4 × medial width); cercus separated medially, forming a weakly-developed subanal plate. Surstylus strongly sinuate, basal half broad and subtriangular with the apex curved posteriorly, apical half strongly bent downwards and constricted with several anterior setae. Postgonite small, broad, and apically tapered with a small subapical patch of microtomentum. Phallapodeme elongate and well sclerotized. Basiphallus stout, slightly curved, with a stout, apically pointed epiphallus; distiphallus stout, largely membranous but including a trough-shaped basal sclerite with upper surface bent inwards to form a “roof”, a shieldshaped upper sclerite, and paired elongate lower-lateral sclerites; lateral membrane densely microtrichose.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31275F112673A79739A6F5BFBD2F47D" pageId="11" pageNumber="470" type="materials_examined">
|
||||
<paragraph id="8BB7267A12673A79739A6F5BFC7CF725" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="470">
|
||||
<emphasis id="B97CFA6812673A79739A6F5BFE11F778" bold="true" box="[199,432,258,284]" pageId="11" pageNumber="470">Material examined.</emphasis>
|
||||
<materialsCitation id="3B602C2712673A7972EB6F5AFC7CF725" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" county="Rio San Lorencito" elevation="900" location="Curso" municipality="San Ramon Forest Reserve" pageId="11" pageNumber="470" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
|
||||
<materialsCitation id="3B602C2712673A7972EB6F5AFC7CF725" ID-GBIF-Occurrence="4958089310" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" county="Rio San Lorencito" elevation="900" location="Curso" municipality="San Ramon Forest Reserve" pageId="11" pageNumber="470" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
|
||||
<typeStatus id="54B398D812673A7972EB6F5AFD82F779" box="[438,547,259,285]" pageId="11" pageNumber="470" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<collectingCountry id="F31F66EA12673A7971716F5AFD70F779" box="[556,721,259,285]" name="Costa Rica" pageId="11" pageNumber="470">COSTA RICA</collectingCountry>
|
||||
|
|
@ -146,7 +144,7 @@ a.s.l.
|
|||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12673A79739A6F12FCDCF791" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="470">
|
||||
<materialsCitation id="3B602C2712673A79739A6F12FBB8F701" box="[199,1049,330,357]" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" county="Rio San Lorencito" elevation="900" location="Curso" municipality="San Ramon Forest Reserve" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712673A79739A6F12FBB8F701" ID-GBIF-Occurrence="4958089312" box="[199,1049,330,357]" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" county="Rio San Lorencito" elevation="900" location="Curso" municipality="San Ramon Forest Reserve" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<typeStatus id="54B398D812673A79739A6F12FE99F701" box="[199,312,331,357]" pageId="11" pageNumber="470" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<collectingCountry id="F31F66EA12673A7972186F12FE4DF701" box="[325,492,331,357]" name="Costa Rica" pageId="11" pageNumber="470">COSTA RICA</collectingCountry>
|
||||
|
|
@ -162,7 +160,7 @@ same data as holotype (
|
|||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B602C2712673A7977786F12FCE8F7ED" collectingDate="1990-05" collectionCode="MNCR" collectorName="II Curso Parataxon" country="Costa Rica" elevation="600" location="Finca San Gabriel" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712673A7977786F12FCE8F7ED" ID-GBIF-Occurrence="4958089319" collectingDate="1990-05" collectionCode="MNCR" collectorName="II Curso Parataxon" country="Costa Rica" elevation="600" location="Finca San Gabriel" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<location id="8ED770A112673A7977786F12FB57F701" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A7977786F12FB57F701" box="[1061,1270,331,357]" country="Costa Rica" name="Finca San Gabriel" pageId="11" pageNumber="470" stateProvince="Alajuela">Finca San Gabriel</location>
|
||||
,
|
||||
<locationDeviation id="9A053F5812673A79765F6F12FF6CF7ED" location="Dos Rios" pageId="11" pageNumber="470">
|
||||
|
|
@ -187,7 +185,7 @@ a.s.l.
|
|||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B602C2712673A7970086F36FB8BF7C9" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="DEBU, MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="11" pageNumber="470" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712673A7970086F36FB8BF7C9" ID-GBIF-Occurrence="4958089313" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="DEBU, MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="11" pageNumber="470" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<emphasis id="B97CFA6812673A7970086F36FC46F7ED" bold="true" box="[853,999,367,393]" pageId="11" pageNumber="470">
|
||||
<collectingRegion id="49CCE89812673A7970086F36FC43F7ED" box="[853,994,367,393]" country="Costa Rica" name="Puntarenas" pageId="11" pageNumber="470">Puntarenas</collectingRegion>
|
||||
:
|
||||
|
|
@ -215,7 +213,7 @@ a.s.l.
|
|||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B602C2712673A79776A6FCAFBB5F7B5" collectingDate="2000-06-13" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Station" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712673A79776A6FCAFBB5F7B5" ID-GBIF-Occurrence="4958089305" collectingDate="2000-06-13" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Station" pageId="11" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||||
<location id="8ED770A112673A79776A6FCAFA39F7C9" LSID="urn:lsid:plazi:treatment:03A1976C12663A7973CA6803FAD0F459:8ED770A112673A79776A6FCAFA39F7C9" box="[1079,1432,403,429]" country="Costa Rica" name="Monteverde Biological Station" pageId="11" pageNumber="470" stateProvince="Puntarenas">Monteverde Biological Station</location>
|
||||
,
|
||||
<elevation id="0025C14912673A7973CA6FEFFE86F7B5" box="[151,295,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="470" unit="m" value="1500.0">
|
||||
|
|
@ -235,7 +233,7 @@ a.s.l.
|
|||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B602C2712673A7977436FEEFCD9F791" collectingDate="1974-07-05" collectionCode="CAS" collectorName="C. W. & L. O'Brien & Marshall" country="Panama" location="Cerro Campana" municipality="Oeste" pageId="11" pageNumber="470" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
|
||||
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@ -79,11 +77,11 @@ Key to the species of
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1 Body length ≤
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. Male fore and hind femora slightly thickened (length 1.9–2.0 × and 4.4–5.6 × width, respectively) (
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<figureCitation id="13333AFF12693A7776286D19FEA2F516" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="5" pageNumber="464">Fig. 2A</figureCitation>
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<figureCitation id="13333AFF12693A7776286D19FEA2F516" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="5" pageNumber="464">Fig. 2A</figureCitation>
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). Surstylus straight along anterior margin (
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<figureCitation id="13333AFF12693A7771FA6D04FD54F516" box="[679,757,861,883]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 4C</figureCitation>
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<figureCitation id="13333AFF12693A7771FA6D04FD54F516" box="[679,757,861,883]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="5" pageNumber="464">Fig. 4C</figureCitation>
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). Posterior opening of epandrium roughly triangular, widest ventrally (
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<figureCitation id="13333AFF12693A7773B46D23FE95F5EB" box="[233,308,890,912]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 4D</figureCitation>
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<figureCitation id="13333AFF12693A7773B46D23FE95F5EB" box="[233,308,890,912]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="5" pageNumber="464">Fig. 4D</figureCitation>
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)..........................................................................
|
||||
<taxonomicName id="4C085DF912693A7777E66D22FAAEF5F4" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[1211,1295,891,912]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="species" species="crena" status="sp. nov.">
|
||||
<emphasis id="B97CFA6812693A7777E66D22FAAEF5F4" box="[1211,1295,891,912]" italics="true" pageId="5" pageNumber="464">H. crena</emphasis>
|
||||
|
|
@ -97,15 +95,15 @@ Key to the species of
|
|||
- Body length ≥
|
||||
<quantity id="4CF08B9F12693A7772286DC1FE63F5C9" box="[373,450,920,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="464" unit="mm" value="1.9">1.9 mm</quantity>
|
||||
. Male fore and hind femora distinctly thickened (length 2.9–3.2 × and 3.1–3.7 × width, respectively) (
|
||||
<figureCitation id="13333AFF12693A7773B46DECFEFFF5AE" box="[233,350,949,971]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="5" pageNumber="464">Figs 2C, 2E</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7773B46DECFEFFF5AE" box="[233,350,949,971]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="5" pageNumber="464">Figs 2C, 2E</figureCitation>
|
||||
). Surstylus sinuate, curved along anterior margin (
|
||||
<figureCitation id="13333AFF12693A7770606DECFC2AF5AF" box="[829,907,949,971]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Figs 5C</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7770606DECFC2AF5AF" box="[829,907,949,971]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Figs 5C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12693A7770C86DECFC15F5AF" box="[917,948,949,971]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">6C</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7770C86DECFC15F5AF" box="[917,948,949,971]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">6C</figureCitation>
|
||||
). Posterior opening of epandrium ovoid or circular, widest around the middle (
|
||||
<figureCitation id="13333AFF12693A7772BF6D8BFD90F58C" box="[482,561,978,1000]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Figs 5D</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7772BF6D8BFD90F58C" box="[482,561,978,1000]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Figs 5D</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12693A7771616D8BFDFBF58C" box="[572,602,978,1000]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">6D</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7771616D8BFDFBF58C" box="[572,602,978,1000]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">6D</figureCitation>
|
||||
)................................................................... 2
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
|
|
@ -114,15 +112,15 @@ Key to the species of
|
|||
<paragraph id="8BB7267A12693A7773CA6DA9FA3CF239" blockId="5.[151,1437,831,1235]" pageId="5" pageNumber="464">
|
||||
<keyLead id="30F9854F12693A7773CA6DA9FA3CF239" pageId="5" pageNumber="464">
|
||||
2 Eye height 3.1–3.2 × genal height; face only slightly protruding. Male S5 longer, width 2.4–2.7 × length (
|
||||
<figureCitation id="13333AFF12693A7777BB6DA9FA92F261" box="[1254,1331,1007,1030]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 5A</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7777BB6DA9FA92F261" box="[1254,1331,1007,1030]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5A</figureCitation>
|
||||
). Posterior opening of epandrium almost circular (height 0.9–1.0 × width) (
|
||||
<figureCitation id="13333AFF12693A7770106A54FC3BF247" box="[845,922,1037,1059]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 5D</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7770106A54FC3BF247" box="[845,922,1037,1059]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5D</figureCitation>
|
||||
). Surstylus bulging basally, apical half gently sinuate (
|
||||
<figureCitation id="13333AFF12693A7773B46A73FE92F224" box="[233,307,1066,1088]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 5C</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7773B46A73FE92F224" box="[233,307,1066,1088]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5C</figureCitation>
|
||||
). Postgonite parallel-sided basally, truncate apically (
|
||||
<figureCitation id="13333AFF12693A7770706A73FCD9F224" box="[813,888,1066,1088]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 5B</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7770706A73FCD9F224" box="[813,888,1066,1088]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5B</figureCitation>
|
||||
). Distiphallus with a lateral row of thickened setulae (
|
||||
<figureCitation id="13333AFF12693A7776286A73FF5EF239" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 5B</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7776286A73FF5EF239" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5B</figureCitation>
|
||||
)...............................................................................
|
||||
<taxonomicName id="4C085DF912693A77779F6A11FAA6F239" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[1218,1287,1096,1117]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="species" species="sica" status="sp. nov.">
|
||||
<emphasis id="B97CFA6812693A77779F6A11FAA6F239" box="[1218,1287,1096,1117]" italics="true" pageId="5" pageNumber="464">H. sica</emphasis>
|
||||
|
|
@ -134,15 +132,15 @@ Key to the species of
|
|||
<paragraph id="8BB7267A12693A7773CA6A3CFA3DF2B6" blockId="5.[151,1437,831,1235]" pageId="5" pageNumber="464">
|
||||
<keyLead id="30F9854F12693A7773CA6A3CFA3DF2B6" pageId="5" pageNumber="464">
|
||||
- Eye height 3.6–3.7 × genal height; face distinctly protruding. Male S5 shorter, width 4.2–4.5 × length (
|
||||
<figureCitation id="13333AFF12693A7777BE6A3CFA92F21F" box="[1251,1331,1125,1147]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 6A</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7777BE6A3CFA92F21F" box="[1251,1331,1125,1147]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6A</figureCitation>
|
||||
). Posterior opening of epandrium roughly ovoid (height 1.3–1.4 × width) (
|
||||
<figureCitation id="13333AFF12693A7770166ADBFC38F2FC" box="[843,921,1154,1176]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 6D</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7770166ADBFC38F2FC" box="[843,921,1154,1176]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6D</figureCitation>
|
||||
). Surstylus roughly parallel-sided basally, apical half strongly bent at about a right angle (
|
||||
<figureCitation id="13333AFF12693A7771636AF9FD2BF2D2" box="[574,650,1184,1206]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 6C</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7771636AF9FD2BF2D2" box="[574,650,1184,1206]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6C</figureCitation>
|
||||
). Postgonite broadened basally, triangular apically (
|
||||
<figureCitation id="13333AFF12693A7777216AF9FB69F2D2" box="[1148,1224,1184,1206]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 6B</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7777216AF9FB69F2D2" box="[1148,1224,1184,1206]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6B</figureCitation>
|
||||
). Distiphallus densely microtomentose dorsally but without longer setulae (
|
||||
<figureCitation id="13333AFF12693A7771806AE4FC87F2B7" box="[733,806,1213,1235]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 6B</figureCitation>
|
||||
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)............................
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<taxonomicName id="4C085DF912693A7777DF6AE7FB75F2B6" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[1154,1236,1213,1235]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="species" species="valga" status="sp. nov.">
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,
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<figureCitation id="13333AFF12693A77723D6BD2FED1F3C1" box="[352,368,1419,1445]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">4</figureCitation>
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<figureCitation id="13333AFF12693A77723D6BD2FED1F3C1" box="[352,368,1419,1445]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="5" pageNumber="464">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12693A7772216BD2FE2DF3C1" box="[380,396,1419,1445]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[184,1404,186,1915]" captionTargetId="figure-19@9.[179,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Helicosina crena sp. nov. female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D. single spermatheca; E, paired spermathecae. Abbreviations: cer—cercus, hyp—hypoproct. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">7</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7772216BD2FE2DF3C1" box="[380,396,1419,1445]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[184,1404,186,1915]" captionTargetId="figure-19@9.[179,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Helicosina crena sp. nov. female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D. single spermatheca; E, paired spermathecae. Abbreviations: cer—cercus, hyp—hypoproct. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982437" httpUri="https://zenodo.org/record/13982437/files/figure.png" pageId="5" pageNumber="464">7</figureCitation>
|
||||
,
|
||||
<figureCitation id="13333AFF12693A7772C56BD2FE07F3C1" box="[408,422,1419,1445]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." pageId="5" pageNumber="464">8</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7772C56BD2FE07F3C1" box="[408,422,1419,1445]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13982439" httpUri="https://zenodo.org/record/13982439/files/figure.png" pageId="5" pageNumber="464">8</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12693A7773CA6B8BFC30F389" blockId="5.[151,1436,1490,1841]" box="[151,913,1490,1517]" pageId="5" pageNumber="464">
|
||||
|
|
@ -98,20 +96,20 @@ Marshall
|
|||
<paragraph id="8BB7267A12693A77739A6BAEFE91F051" blockId="5.[151,1436,1490,1841]" pageId="5" pageNumber="464">
|
||||
<emphasis id="B97CFA6812693A77739A6BAEFE82F075" box="[199,291,1527,1553]" italics="true" pageId="5" pageNumber="464">General</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF12693A77726A6BAEFE36F075" box="[311,407,1527,1553]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="5" pageNumber="464">Fig. 2A</figureCitation>
|
||||
<figureCitation id="13333AFF12693A77726A6BAEFE36F075" box="[311,407,1527,1553]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="5" pageNumber="464">Fig. 2A</figureCitation>
|
||||
): Male fore and hind femora slightly thickened (length 1.9–2.0 × and 4.4–5.6 × width, respectively).
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A12693A77739A6866FC5DF08D" blockId="5.[151,1436,1490,1841]" pageId="5" pageNumber="464">
|
||||
<emphasis id="B97CFA6812693A77739A6866FECFF03D" box="[199,366,1599,1625]" italics="true" pageId="5" pageNumber="464">Male abdomen</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF12693A7772206866FE63F03D" box="[381,450,1599,1625]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." pageId="5" pageNumber="464">Fig. 4</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7772206866FE63F03D" box="[381,450,1599,1625]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="5" pageNumber="464">Fig. 4</figureCitation>
|
||||
): S5 broad, desclerotized posteromedially, and uniformly long-setose. Posterior opening of epandrium triangular (medial height subequal to ventral width); subanal plate long and very narrow. Surstylus folded into inner and outer lobes; outer lobe rounded and setose, inner lobe flat with a few marginal setae and a large, transparent, posteroventral scale. Cercus broad, lobate, narrowly fused dorsally. Distiphallus short-cylindrical with a very broad, bilobed opening subtended by broad, flap-like distal lobes.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C31275F112693A78739A68ABFC48F4E1" lastPageId="10" lastPageNumber="469" pageId="5" pageNumber="464" type="materials_examined">
|
||||
<paragraph id="8BB7267A12693A77739A68ABFD66F155" blockId="5.[151,1436,1490,1841]" pageId="5" pageNumber="464">
|
||||
<emphasis id="B97CFA6812693A77739A68ABFE13F168" bold="true" box="[199,434,1778,1804]" pageId="5" pageNumber="464">Material examined.</emphasis>
|
||||
<materialsCitation id="3B602C2712693A7772E668AAFD66F155" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="5" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||||
<materialsCitation id="3B602C2712693A7772E668AAFD66F155" ID-GBIF-Occurrence="4958089314" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="5" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||||
<typeStatus id="54B398D812693A7772E668AAFD89F169" box="[443,552,1779,1805]" pageId="5" pageNumber="464" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<collectingCountry id="F31F66EA12693A77716E68AAFD7AF169" box="[563,731,1779,1805]" name="Costa Rica" pageId="5" pageNumber="464">COSTA RICA</collectingCountry>
|
||||
|
|
@ -138,7 +136,7 @@ Marshall
|
|||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<caption id="DF7776F2126A3A7473C5692CFC5CF19C" pageId="6" pageNumber="465" startId="6.[152,255,1909,1933]" targetBox="[186,1401,186,1880]" targetPageId="6" targetType="figure">
|
||||
<caption id="DF7776F2126A3A7473C5692CFC5CF19C" ID-DOI="http://doi.org/10.5281/zenodo.13982429" ID-Zenodo-Dep="13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="6" pageNumber="465" startId="6.[152,255,1909,1933]" targetBox="[186,1401,186,1880]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8BB7267A126A3A7473C5692CFC5CF19C" blockId="6.[151,1437,1909,2041]" pageId="6" pageNumber="465">
|
||||
<emphasis id="B97CFA68126A3A7473C5692CFEB6F1E9" bold="true" box="[152,279,1909,1933]" pageId="6" pageNumber="465">FIGURE 4.</emphasis>
|
||||
<taxonomicName id="4C085DF9126A3A747240692CFE68F1E9" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[285,457,1909,1933]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="crena" status="sp. nov.">
|
||||
|
|
@ -148,7 +146,7 @@ Marshall
|
|||
male paratype terminalia. A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF7776F2126B3A7573C569CEFA29F1B7" pageId="7" pageNumber="466" startId="7.[152,255,1943,1967]" targetBox="[179,1412,186,1915]" targetPageId="7" targetType="figure">
|
||||
<caption id="DF7776F2126B3A7573C569CEFA29F1B7" ID-DOI="http://doi.org/10.5281/zenodo.13982433" ID-Zenodo-Dep="13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="7" pageNumber="466" startId="7.[152,255,1943,1967]" targetBox="[179,1412,186,1915]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BB7267A126B3A7573C569CEFA29F1B7" blockId="7.[151,1436,1943,2004]" pageId="7" pageNumber="466">
|
||||
<emphasis id="B97CFA68126B3A7573C569CEFEB9F1D4" bold="true" box="[152,280,1943,1968]" pageId="7" pageNumber="466">FIGURE 5.</emphasis>
|
||||
<taxonomicName id="4C085DF9126B3A75724269C1FE1DF1CB" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[287,444,1944,1968]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="sica" status="sp. nov.">
|
||||
|
|
@ -158,7 +156,7 @@ male paratype terminalia. A, S5–8 and postabdomen, ventral. B, aedeagus and as
|
|||
male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF7776F212643A7A73C569CEFA29F1B7" pageId="8" pageNumber="467" startId="8.[152,255,1943,1967]" targetBox="[181,1406,186,1915]" targetPageId="8" targetType="figure">
|
||||
<caption id="DF7776F212643A7A73C569CEFA29F1B7" ID-DOI="http://doi.org/10.5281/zenodo.13982435" ID-Zenodo-Dep="13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="8" pageNumber="467" startId="8.[152,255,1943,1967]" targetBox="[181,1406,186,1915]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BB7267A12643A7A73C569CEFA29F1B7" blockId="8.[151,1436,1943,2004]" pageId="8" pageNumber="467">
|
||||
<emphasis id="B97CFA6812643A7A73C569CEFEB6F1CB" bold="true" box="[152,279,1943,1967]" pageId="8" pageNumber="467">FIGURE 6.</emphasis>
|
||||
<taxonomicName id="4C085DF912643A7A724069C1FE69F1D4" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[285,456,1944,1968]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="467" phylum="Arthropoda" rank="species" species="valga" status="sp. nov.">
|
||||
|
|
@ -168,7 +166,7 @@ male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and as
|
|||
male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF7776F212653A7B73C569CEFAB4F1B7" pageId="9" pageNumber="468" startId="9.[152,255,1943,1967]" targetBox="[184,1404,186,1915]" targetPageId="9" targetType="figure">
|
||||
<caption id="DF7776F212653A7B73C569CEFAB4F1B7" ID-DOI="http://doi.org/10.5281/zenodo.13982437" ID-Zenodo-Dep="13982437" httpUri="https://zenodo.org/record/13982437/files/figure.png" pageId="9" pageNumber="468" startId="9.[152,255,1943,1967]" targetBox="[184,1404,186,1915]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8BB7267A12653A7B73C569CEFAB4F1B7" blockId="9.[151,1437,1943,2004]" pageId="9" pageNumber="468">
|
||||
<emphasis id="B97CFA6812653A7B73C569CEFEB8F1D4" bold="true" box="[152,281,1943,1968]" pageId="9" pageNumber="468">FIGURE 7.</emphasis>
|
||||
<taxonomicName id="4C085DF912653A7B727C69C1FE6EF1CB" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[289,463,1944,1968]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="468" phylum="Arthropoda" rank="species" species="crena" status="sp. nov.">
|
||||
|
|
@ -179,7 +177,7 @@ female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D.
|
|||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB7267A12663A78739A6ECEFD96F405" blockId="10.[151,1437,150,717]" pageId="10" pageNumber="469">
|
||||
<materialsCitation id="3B602C2712663A78739A6ECEFAC2F6D5" box="[199,1379,150,177]" collectingDate="1986-08-18" collectionCode="DEBU" collectorName="L. Masner" country="Costa Rica" elevation="700" location="Rio Penas Blancas" pageId="10" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712663A78739A6ECEFAC2F6D5" ID-GBIF-Occurrence="4958089303" box="[199,1379,150,177]" collectingDate="1986-08-18" collectionCode="DEBU" collectorName="L. Masner" country="Costa Rica" elevation="700" location="Rio Penas Blancas" pageId="10" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="paratype">
|
||||
<typeStatus id="54B398D812663A78739A6ECEFE99F6D5" box="[199,312,151,177]" pageId="10" pageNumber="469" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<collectingCountry id="F31F66EA12663A7872186ECEFE4CF6D5" box="[325,493,151,177]" name="Costa Rica" pageId="10" pageNumber="469">COSTA RICA</collectingCountry>
|
||||
|
|
@ -206,7 +204,7 @@ female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D.
|
|||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B602C2712663A7876326ECFFEBAF69D" collectingDate="2012-09-05" collectingDateMax="2012-09-08" collectingDateMin="2012-09-05" collectionCode="MNCR" collectorName="W. Porras" country="Costa Rica" elevation="1600" location="Zurqui de Moravia" municipality="Moravia" pageId="10" pageNumber="469" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712663A7876326ECFFEBAF69D" ID-GBIF-Occurrence="4958089316" collectingDate="2012-09-05" collectingDateMax="2012-09-08" collectingDateMin="2012-09-05" collectionCode="MNCR" collectorName="W. Porras" country="Costa Rica" elevation="1600" location="Zurqui de Moravia" municipality="Moravia" pageId="10" pageNumber="469" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
|
||||
<emphasis id="B97CFA6812663A7876326ECFFF75F6B1" bold="true" pageId="10" pageNumber="469">
|
||||
<collectingRegion id="49CCE89812663A7876326ECFFF6EF6B1" country="Costa Rica" name="San Jose" pageId="10" pageNumber="469">San José</collectingRegion>
|
||||
:
|
||||
|
|
@ -233,7 +231,7 @@ female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D.
|
|||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B602C2712663A78727A6E86FE0CF779" collectingDate="2013-08-05" collectingDateMax="2013-08-06" collectingDateMin="2013-08-05" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1600" location="Zurqui de Moravia" municipality="Moravia" pageId="10" pageNumber="469" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712663A78727A6E86FE0CF779" ID-GBIF-Occurrence="4958089318" collectingDate="2013-08-05" collectingDateMax="2013-08-06" collectingDateMin="2013-08-05" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1600" location="Zurqui de Moravia" municipality="Moravia" pageId="10" pageNumber="469" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
|
||||
<collectingMunicipality id="6BD3BC0012663A78727A6E86FE28F69D" box="[295,393,223,249]" pageId="10" pageNumber="469">Moravia</collectingMunicipality>
|
||||
,
|
||||
<location id="8ED770A112663A7872C86E86FDCFF69D" LSID="urn:lsid:plazi:treatment:03A1976C12693A7873CA6B3FFDEBF4A9:8ED770A112663A7872C86E86FDCFF69D" box="[405,622,223,249]" country="Costa Rica" municipality="Moravia" name="Zurqui de Moravia" pageId="10" pageNumber="469" stateProvince="San Jose">Zurquí de Moravia</location>
|
||||
|
|
@ -254,7 +252,7 @@ female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D.
|
|||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B602C2712663A7872E16F5AFCB8F725" collectingDate="2013-05-24" collectingDateMax="2013-05-31" collectingDateMin="2013-05-24" collectionCode="MNCR" collectorName="Proyecto" country="Costa Rica" county="Moravia" elevation="1600" location="Tower" municipality="Zurqui de Moravia" pageId="10" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
|
||||
<materialsCitation id="3B602C2712663A7872E16F5AFCB8F725" ID-GBIF-Occurrence="4958089308" collectingDate="2013-05-24" collectingDateMax="2013-05-31" collectingDateMin="2013-05-24" collectionCode="MNCR" collectorName="Proyecto" country="Costa Rica" county="Moravia" elevation="1600" location="Tower" municipality="Zurqui de Moravia" pageId="10" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
|
||||
<collectingCounty id="62D65EF612663A7872E16F5AFDBFF779" box="[444,542,259,285]" pageId="10" pageNumber="469">Moravia</collectingCounty>
|
||||
,
|
||||
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@ -279,7 +277,7 @@ ZADBI (
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@ -304,7 +302,7 @@ ZADBI (
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@ -329,7 +327,7 @@ ZADBI (
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@ -379,7 +377,7 @@ ZADBI (
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@ -406,7 +404,7 @@ ZADBI (
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<mods:identifier id="084FDEB366442CB2D28CC1EE5D5B8ED0" type="DOI">10.11646/zootaxa.5523.4.5</mods:identifier>
|
||||
<mods:identifier id="161CC4118342E8F4D284AFFC4A72491D" type="GBIF-Dataset">c8fd3aa9-6203-4d1b-ae7b-367bf1d498b6</mods:identifier>
|
||||
<mods:identifier id="CF6FC1A23B886D09B25C9F7891E9590E" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="1E6A7C7B2367D2718F04639588E309FD" type="Zenodo-Dep">13949669</mods:identifier>
|
||||
<mods:identifier id="1AFC37D84A6FF6670B685FA7CA41CDF6" type="ZooBank">999559E0-4E12-45DC-AED9-88374AA3E3D1</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03A1976C126D3A7773CA6FCBFC7BF405" ID-Zenodo-Dep="13949727" LSID="urn:lsid:plazi:treatment:03A1976C126D3A7773CA6FCBFC7BF405" httpUri="http://treatment.plazi.org/id/03A1976C126D3A7773CA6FCBFC7BF405" lastPageId="5" lastPageNumber="464" pageId="1" pageNumber="460">
|
||||
<treatment id="03A1976C126D3A7773CA6FCBFC7BF405" ID-DOI="http://doi.org/10.5281/zenodo.13949727" ID-GBIF-Taxon="244134449" ID-Zenodo-Dep="13949727" LSID="urn:lsid:plazi:treatment:03A1976C126D3A7773CA6FCBFC7BF405" httpUri="http://treatment.plazi.org/id/03A1976C126D3A7773CA6FCBFC7BF405" lastPageId="5" lastPageNumber="464" pageId="1" pageNumber="460">
|
||||
<subSubSection id="C31275F1126D3A7373CA6FCBFDE5F7C9" box="[151,580,402,429]" pageId="1" pageNumber="460" type="nomenclature">
|
||||
<paragraph id="8BB7267A126D3A7373CA6FCBFDE5F7C9" blockId="1.[151,580,402,429]" box="[151,580,402,429]" pageId="1" pageNumber="460">
|
||||
<heading id="D0FF9116126D3A7373CA6FCBFDE5F7C9" bold="true" box="[151,580,402,429]" fontSize="11" level="1" pageId="1" pageNumber="460" reason="1">
|
||||
|
|
@ -102,7 +100,7 @@ is easily recognized by the combination of a single cruciate pair of interfronta
|
|||
, females
|
||||
<quantity id="4CF08B9F126D3A7370F66CEAFB99F4A9" box="[939,1080,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="1" pageNumber="460" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6–2.2 mm</quantity>
|
||||
. General colour brown to dark brown, legs pale brown to yellow (
|
||||
<figureCitation id="13333AFF126D3A7371406C8EFD28F495" box="[541,649,727,753]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,1795,1819]" captionStartId-1="3.[152,255,1868,1892]" captionTargetBox-0="[151,1435,306,1769]" captionTargetBox-1="[151,1435,309,1845]" captionTargetId-0="figure-76@2.[151,1435,306,1769]" captionTargetId-1="figure-73@3.[151,1435,309,1845]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Helicosina sica gen. et sp. nov., living male, Tiputini Biological Station, Ecuador." captionText-1="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." pageId="1" pageNumber="460">Figs. 1–2</figureCitation>
|
||||
<figureCitation id="13333AFF126D3A7371406C8EFD28F495" box="[541,649,727,753]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,1795,1819]" captionStartId-1="3.[152,255,1868,1892]" captionTargetBox-0="[151,1435,306,1769]" captionTargetBox-1="[151,1435,309,1845]" captionTargetId-0="figure-76@2.[151,1435,306,1769]" captionTargetId-1="figure-73@3.[151,1435,309,1845]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Helicosina sica gen. et sp. nov., living male, Tiputini Biological Station, Ecuador." captionText-1="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13982423" figureDoi-1="http://doi.org/10.5281/zenodo.13982425" httpUri-0="https://zenodo.org/record/13982423/files/figure.png" httpUri-1="https://zenodo.org/record/13982425/files/figure.png" pageId="1" pageNumber="460">Figs. 1–2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB7267A126D3A73739A6CA2FE26F219" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="460">
|
||||
|
|
@ -120,7 +118,7 @@ is easily recognized by the combination of a single cruciate pair of interfronta
|
|||
<paragraph id="8BB7267A126D3A73739A686EFAA0F085" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="460">
|
||||
<emphasis id="B97CFA68126D3A73739A686EFEA0F034" box="[199,257,1591,1616]" italics="true" pageId="1" pageNumber="460">Wing</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF126D3A73724F686EFEF9F035" box="[274,344,1591,1617]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." pageId="1" pageNumber="460">Fig. 3</figureCitation>
|
||||
<figureCitation id="13333AFF126D3A73724F686EFEF9F035" box="[274,344,1591,1617]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[174,1414,181,1919]" captionTargetId="figure-29@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Wing photos of Helicosina species.A, Helicosina crena sp. nov. paratype; B, Helicosina sica sp. nov. paratype; C, Helicosina valga sp. nov. paratype. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982427" httpUri="https://zenodo.org/record/13982427/files/figure.png" pageId="1" pageNumber="460">Fig. 3</figureCitation>
|
||||
): Slightly infuscate, relatively elongate (length 2.2–2.4 × width). CS2 1.1–1.2 × CS3. Vein R
|
||||
<subScript id="178C243F126D3A7376DE6810FA3DF033" attach="left" box="[1411,1436,1609,1623]" fontSize="6" pageId="1" pageNumber="460">2+3</subScript>
|
||||
sinuate, distal curve much stronger than basal curve; R
|
||||
|
|
@ -137,7 +135,7 @@ extending beyond dm-m by 1.2–1.3 × length of dm-m. CuA+CuP long and sinuate a
|
|||
<emphasis id="B97CFA68126D3A73739A68B2FECEF161" box="[199,367,1771,1797]" italics="true" pageId="1" pageNumber="460">Male abdomen</emphasis>
|
||||
: T2–5 and S2–4 brown to dark brown, broad, slightly desclerotized around margins, uniformly long setose in posterior half. S5 broad, curved, relatively simple, and long setose at least laterally. Synsternite 6+7 well-developed, S6 portion extending dextrally beyond middle of S5. Epandrium relatively large, saddle-shaped, uniformly long-setose; cercus usually fused medially, forming a well-developed subanal plate; subepandrial sclerite thickened medially. Hypandrium well-developed, Y-shaped, anterior apodeme elongate, slightly sinuate, strongly fused to lateral arms, lateral arms fused to anteroventral corners of epandrium. Surstylus large (subequal in length to epandrial height), elongate (height 2.8–3.4 × length) and tapered, largely bare externally with longer setae along inner-anterior edge. Postgonite small, stout. Phallapodeme large, curved. Basiphallus stout, straight to slightly elbowed, sometimes with a posterior patch of dense setulae, usually with a conical epiphallus; distiphallus large and broad, bulbous, largely membranous and supported by several flattened sclerites, laterally and sometimes dorsally setulose or microtomentose.
|
||||
</paragraph>
|
||||
<caption id="DF7776F2126E3A7073C5695AFBFAF17F" box="[152,1115,1795,1820]" pageId="2" pageNumber="461" startId="2.[152,255,1795,1819]" targetBox="[151,1435,306,1769]" targetPageId="2" targetType="figure">
|
||||
<caption id="DF7776F2126E3A7073C5695AFBFAF17F" ID-DOI="http://doi.org/10.5281/zenodo.13982423" ID-Zenodo-Dep="13982423" box="[152,1115,1795,1820]" httpUri="https://zenodo.org/record/13982423/files/figure.png" pageId="2" pageNumber="461" startId="2.[152,255,1795,1819]" targetBox="[151,1435,306,1769]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="8BB7267A126E3A7073C5695AFBFAF17F" blockId="2.[152,1115,1795,1820]" box="[152,1115,1795,1820]" pageId="2" pageNumber="461">
|
||||
<emphasis id="B97CFA68126E3A7073C5695AFEB8F17F" bold="true" box="[152,281,1795,1819]" pageId="2" pageNumber="461">FIGURE 1.</emphasis>
|
||||
<taxonomicName id="4C085DF9126E3A707242695AFE1DF17F" authorityName="Kuwahara & Marshall & Yau" authorityYear="2024" box="[287,444,1795,1819]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="sica" status="gen. et sp. nov.">
|
||||
|
|
@ -150,11 +148,11 @@ extending beyond dm-m by 1.2–1.3 × length of dm-m. CuA+CuP long and sinuate a
|
|||
<paragraph id="8BB7267A126E3A71739A6934FCA6F779" blockId="2.[151,1437,1901,2036]" lastBlockId="3.[151,1437,150,285]" lastPageId="3" lastPageNumber="462" pageId="2" pageNumber="461">
|
||||
<emphasis id="B97CFA68126E3A70739A6934FE28F1E3" box="[199,393,1901,1927]" italics="true" pageId="2" pageNumber="461">Female abdomen</emphasis>
|
||||
(
|
||||
<figureCitation id="13333AFF126E3A7072C56934FE7FF1E3" box="[408,478,1901,1927]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[184,1404,186,1915]" captionTargetId="figure-19@9.[179,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Helicosina crena sp. nov. female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D. single spermatheca; E, paired spermathecae. Abbreviations: cer—cercus, hyp—hypoproct. Scale bars indicate 0.10 mm." pageId="2" pageNumber="461">Fig. 7</figureCitation>
|
||||
<figureCitation id="13333AFF126E3A7072C56934FE7FF1E3" box="[408,478,1901,1927]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[184,1404,186,1915]" captionTargetId="figure-19@9.[179,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Helicosina crena sp. nov. female paratype terminalia. A, dorsal view; B, ventral view; C, lateral view; D. single spermatheca; E, paired spermathecae. Abbreviations: cer—cercus, hyp—hypoproct. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982437" httpUri="https://zenodo.org/record/13982437/files/figure.png" pageId="2" pageNumber="461">Fig. 7</figureCitation>
|
||||
): T2–T6 and S2–S6 brown, broad, well sclerotized medially but desclerotized around the margins, uniformly long setose along entire surface. T7 broad, well sclerotized, and uniformly long setose; T8 broad, desclerotized anteromedially and protruding posteromedially (though this may be the epiproct fused to T8) with six long posterior setae. Epiproct absent or fused to posterior margin of T8. Cercus stout, subconical with long dorsal, apical, and apicolateral setae. S7 large (1.5 × length of T7), subquadrate, and relatively densely setose; S8 strongly reduced or absent. Hypoproct broad, U-shaped, densely microtomentose with 8–10 posterior setae. Spermathecae (2+1) stout, bulb spherical/cylindrical with a large and deep apical invagination, stem elongate but thickened below bulb; ducts elongate and membranous.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF7776F2126F3A7173C56915FDE9F1B4" pageId="3" pageNumber="462" startId="3.[152,255,1868,1892]" targetBox="[151,1435,309,1845]" targetPageId="3" targetType="figure">
|
||||
<caption id="DF7776F2126F3A7173C56915FDE9F1B4" ID-DOI="http://doi.org/10.5281/zenodo.13982425" ID-Zenodo-Dep="13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="3" pageNumber="462" startId="3.[152,255,1868,1892]" targetBox="[151,1435,309,1845]" targetPageId="3" targetType="figure">
|
||||
<paragraph id="8BB7267A126F3A7173C56915FDE9F1B4" blockId="3.[151,1437,1868,2000]" pageId="3" pageNumber="462">
|
||||
<emphasis id="B97CFA68126F3A7173C56915FEB6F100" bold="true" box="[152,279,1868,1892]" pageId="3" pageNumber="462">FIGURE 2.</emphasis>
|
||||
Habitus and head photos of
|
||||
|
|
@ -195,7 +193,7 @@ paratype habitus, lateral; F,
|
|||
paratype head, anterolateral. Scale bars indicate 0.50 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF7776F212683A7673C569CEFC89F1B7" pageId="4" pageNumber="463" startId="4.[152,255,1943,1967]" targetBox="[174,1414,181,1919]" targetPageId="4" targetType="figure">
|
||||
<caption id="DF7776F212683A7673C569CEFC89F1B7" ID-DOI="http://doi.org/10.5281/zenodo.13982427" ID-Zenodo-Dep="13982427" httpUri="https://zenodo.org/record/13982427/files/figure.png" pageId="4" pageNumber="463" startId="4.[152,255,1943,1967]" targetBox="[174,1414,181,1919]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8BB7267A12683A7673C569CEFC89F1B7" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="463">
|
||||
<emphasis id="B97CFA6812683A7673C569CEFEB9F1CB" bold="true" box="[152,280,1943,1967]" pageId="4" pageNumber="463">FIGURE 3.</emphasis>
|
||||
Wing photos of
|
||||
|
|
@ -292,7 +290,7 @@ Known from
|
|||
, and
|
||||
<collectingCountry id="F31F66EA12693A77707D6C1EFCD8F405" box="[800,889,583,609]" name="Panama" pageId="5" pageNumber="464">Panama</collectingCountry>
|
||||
(
|
||||
<figureCitation id="13333AFF12693A7770D56C1EFC6DF405" box="[904,972,583,609]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." pageId="5" pageNumber="464">Fig. 8</figureCitation>
|
||||
<figureCitation id="13333AFF12693A7770D56C1EFC6DF405" box="[904,972,583,609]" captionStart="FIGURE 8" captionStartId="11.[152,255,1563,1587]" captionTargetBox="[190,1398,597,1539]" captionTargetId="figure-197@11.[190,1398,597,1539]" captionTargetPageId="11" captionText="FIGURE 8. Distribution map for Helicosina. Blue circles indicate H. crena sp. nov., red circles indicate H. sica sp. nov., and green circles indicate H. valga sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13982439" httpUri="https://zenodo.org/record/13982439/files/figure.png" pageId="5" pageNumber="464">Fig. 8</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
Loading…
Reference in a new issue