325 lines
46 KiB
XML
325 lines
46 KiB
XML
<document id="DB4B10DD725E8987B69360D7B388722C" ID-DOI="10.1093/zoolinnean/zlae021" ID-ISSN="0024-4082" ID-Zenodo-Dep="14291325" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1732847136500" checkinUser="plazi" docAuthor="Vilhelmsen, Lars, Boudinot, Brendon, Jenkins Shaw, Josh, Hammel, Jörg U & Perrichot, Vincent" docDate="2024" docId="039E87B0FFA5FFAE18A7115FFF34FBA8" docLanguage="en" docName="zlae021.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae021" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Kryptovelona carstengroehni Vilhelmsen and Perrichot" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="FFA7FFC8FFA7FFA6182C145CFFDCFFC1" masterDocTitle="Echoes from the Cretaceous: new fossils shed light on the evolution of host detection and concealed ovipositor apparatus in the parasitoid wasp superfamily Orussoidea (Hymenoptera)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="3" updateTime="1733518441416" updateUser="ExternalLinkService">
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<mods:title id="C23E4EEA248323EF4792FBBA513B5B63">Echoes from the Cretaceous: new fossils shed light on the evolution of host detection and concealed ovipositor apparatus in the parasitoid wasp superfamily Orussoidea (Hymenoptera)</mods:title>
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<mods:roleTerm id="9770C320D4A0F3779E112C7CEB2EF46F">Author</mods:roleTerm>
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<mods:namePart id="BC775730EC6381707F3E877B091076A8">Vilhelmsen, Lars</mods:namePart>
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<mods:roleTerm id="E046AE3EB5DA21867793953A2BABE137">Author</mods:roleTerm>
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<mods:namePart id="9CF1E751F0549DB44D8E2C84C360EE8C">Boudinot, Brendon</mods:namePart>
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<mods:roleTerm id="8C6578B5F5E22E44D22E3555AB2FAB36">Author</mods:roleTerm>
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<mods:namePart id="4B89461710A079173B8C90052E01C727">Jenkins Shaw, Josh</mods:namePart>
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<mods:name id="269B1FB801B12295965CDB4E91D358DD" type="personal">
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<mods:roleTerm id="5A9FEB1CD0779C1012E9469BCD77788A">Author</mods:roleTerm>
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<mods:namePart id="DB3EFC2AB7367C1F6898496378F9F4F8">Hammel, Jörg U</mods:namePart>
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<mods:name id="F4F3B6DF382921BACE7EA50F6D856824" type="personal">
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<mods:roleTerm id="AC6A7E370951709B75198E6C5FF217B8">Author</mods:roleTerm>
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<mods:namePart id="307F1D9BF00EB1D40B5201DB5A79AA5C">Perrichot, Vincent</mods:namePart>
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<mods:typeOfResource id="00D487A733B2823F114F9FEA8EFE2F52">text</mods:typeOfResource>
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<mods:title id="946C7C60B67B9309AF840D2697D245F9">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="134707E54D34EBE2EEA4C06BC6C4BDF1">
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<mods:date id="D60FA09B9C945D0ECB911D4CF3C3D85E">2024</mods:date>
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<mods:title id="B52B43DBF786E2B9B10AFB1493726235">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="AA5BFAE218EC7C8EE73A933DD1F7E9B3">2024-02-28</mods:number>
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<mods:number id="93287D57C94D1B4C73C1A2385FE085A5">202</mods:number>
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<mods:number id="156D3DA8CDBE6845F1A86E1B901020C3">3</mods:number>
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<mods:start id="DB66FD8E5757ECF7D86B237FD7B1F482">1</mods:start>
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<mods:url id="56EFB3E9435072BF3F1BD7AEB5EEFFC5">https://doi.org/10.1093/zoolinnean/zlae021</mods:url>
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<mods:classification id="2EC63B2803F34D026E9C66DD3A872318">journal article</mods:classification>
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<mods:identifier id="0C94E62E401778B2442B6C46E0DC5ED9" type="DOI">10.1093/zoolinnean/zlae021</mods:identifier>
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<mods:identifier id="812966805DE31A831DEECE817A0AC952" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="830033A5982191EB1E7843205E52B3B5" type="Zenodo-Dep">14291325</mods:identifier>
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<treatment id="039E87B0FFA5FFAE18A7115FFF34FBA8" LSID="urn:lsid:plazi:treatment:039E87B0FFA5FFAE18A7115FFF34FBA8" httpUri="http://treatment.plazi.org/id/039E87B0FFA5FFAE18A7115FFF34FBA8" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3">
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<subSubSection id="C32D652DFFA5FFA418A7115FFE2BFAFC" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B8836A6FFA5FFA418A7115FFE2BFAFC" blockId="2.[139,767,1283,1341]" pageId="2" pageNumber="3">
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<emphasis id="B943EAB4FFA5FFA418A7115FFD23FADF" bold="true" box="[139,767,1283,1310]" pageId="2" pageNumber="3">
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†
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<taxonomicName id="4C374D25FFA5FFA418BB115FFD23FADF" authority="Vilhelmsen and Perrichot" authorityName="Vilhelmsen and Perrichot" box="[151,767,1283,1310]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="carstengroehni" status="gen.">
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<emphasis id="B943EAB4FFA5FFA418BB115FFE69FADF" bold="true" box="[151,437,1283,1310]" italics="true" pageId="2" pageNumber="3">Kryptovelona carstengroehni</emphasis>
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Vilhelmsen and Perrichot, gen.
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</taxonomicName>
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</emphasis>
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<emphasis id="B943EAB4FFA5FFA419BE117FFE2BFAFC" bold="true" box="[402,503,1315,1341]" pageId="2" pageNumber="3">et sp. nov.</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32D652DFFA5FFA419A11110FE29F9C2" pageId="2" pageNumber="3" type="description">
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<paragraph id="8B8836A6FFA5FFA419A11110FE21FAA7" blockId="2.[397,509,1356,1383]" box="[397,509,1356,1383]" pageId="2" pageNumber="3">
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<heading id="D0C081CAFFA5FFA419A11110FE21FAA7" box="[397,509,1356,1383]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="3" reason="3">
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<emphasis id="B943EAB4FFA5FFA419A11110FE21FAA7" bold="true" box="[397,509,1356,1383]" pageId="2" pageNumber="3">
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(
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<figureCitation id="130C2A23FFA5FFA419B41111FE2EFAA6" box="[408,498,1357,1383]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[114,179,1713,1737]" captionStartId-1="4.[129,192,1671,1695]" captionStartId-2="5.[113,178,1815,1839]" captionStartId-3="6.[129,194,1682,1706]" captionStartId-4="7.[113,178,1669,1693]" captionTargetBox-0="[114,1458,144,1686]" captionTargetBox-1="[129,1473,144,1643]" captionTargetBox-2="[114,1458,144,1788]" captionTargetBox-3="[129,1473,144,1654]" captionTargetBox-4="[114,1454,149,1639]" captionTargetId-0="figure-88@3.[114,1458,144,1686]" captionTargetId-1="figure-89@4.[129,1473,144,1643]" captionTargetId-2="figure-7@5.[114,1458,144,1788]" captionTargetId-3="figure-64@6.[129,1473,144,1654]" captionTargetId-4="figure-101@7.[113,1460,144,1641]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" captionText-1="Figure 2. †Kryptovelona carstengroehni gen* et sp* nov*,female holotype (GPIH 5078, CCGG nº 3811)* A,C, photomicrographs* B, D, rendered from synchrotron scans* A, B, head, anterior view* C, head, lateroventral view* D, anterior view of tentorium* Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; greenarrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge*" captionText-2="Figure 3. A, B, †Cretorussus vilhelmseni Jouault et al.2021, female holotype IGR.BU-020. C, †Kryptovelona groehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †Kryptovelona in italics, with homologues in † Cretorussus in brackets); tb1 = fore tibia." captionText-3="Figure 4. †Kryptovelona carstengroehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum." captionText-4="Figure 5. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal overview of extent of ovipositor inside the body; anterior to the top* B, profurca in posterior view* C, profurca in anterolateral view* D, posterior part of abdomen, dorsal internal view* All views rendered from synchrotron scans* Red arrows = ovipositor; black arrows = medain apodemes on abdominal sternum 6 and 7; green arrows = median groove on profurcal bridge; dl1 = prodiscrimenal lamella; pOE = profurcal bridge*" figureDoi-0="http://doi.org/10.5281/zenodo.14291327" figureDoi-1="http://doi.org/10.5281/zenodo.14291329" figureDoi-2="http://doi.org/10.5281/zenodo.14291331" figureDoi-3="http://doi.org/10.5281/zenodo.14291333" figureDoi-4="http://doi.org/10.5281/zenodo.14291335" httpUri-0="https://zenodo.org/record/14291327/files/figure.png" httpUri-1="https://zenodo.org/record/14291329/files/figure.png" httpUri-2="https://zenodo.org/record/14291331/files/figure.png" httpUri-3="https://zenodo.org/record/14291333/files/figure.png" httpUri-4="https://zenodo.org/record/14291335/files/figure.png" pageId="2" pageNumber="3">Figs 1–5</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8836A6FFA5FFA418AD11D1FE29F9C2" blockId="2.[129,777,1421,1539]" pageId="2" pageNumber="3">
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<emphasis id="B943EAB4FFA5FFA418AD11D1FE8AFA64" box="[129,342,1421,1445]" italics="true" pageId="2" pageNumber="3">ZooBank registration</emphasis>
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||
: urn:lsid:zoobank.org:act:
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<uuid id="FF910C73FFA5FFA41A5711D1FE36FA04" pageId="2" pageNumber="3">FCDEC20D-16ED-46AF-9866-4E792B0E7B1F</uuid>
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(for
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<taxonomicName id="4C374D25FFA5FFA41A5811F1FD2BFA04" box="[628,759,1453,1477]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B943EAB4FFA5FFA41A5811F1FD2BFA04" box="[628,759,1453,1477]" italics="true" pageId="2" pageNumber="3">Kryptovelona</emphasis>
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</taxonomicName>
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||
); urn:lsid:zoobank.org:act:
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<uuid id="FF910C73FFA5FFA419B41190FEF4F9C2" pageId="2" pageNumber="3">DCE94EBD-773F-438A-A441- A059C79149F8</uuid>
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||
(for
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<taxonomicName id="4C374D25FFA5FFA4197511B7FE38F9C2" box="[345,484,1515,1539]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="carstengroehni">
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<emphasis id="B943EAB4FFA5FFA4197511B7FE38F9C2" box="[345,484,1515,1539]" italics="true" pageId="2" pageNumber="3">carstengroehni</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D652DFFA5FFA418AD1279FE52F9BA" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8B8836A6FFA5FFA418AD1279FE52F9BA" blockId="2.[129,778,1572,1659]" pageId="2" pageNumber="3">
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<emphasis id="B943EAB4FFA5FFA418AD1279FE9DF9FC" box="[129,321,1573,1597]" italics="true" pageId="2" pageNumber="3">Material examined:</emphasis>
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<materialsCitation id="3B5F3CFBFFA5FFA419641279FE52F9BA" collectionCode="GPIH" pageId="2" pageNumber="3" specimenCode="GPIH 5078" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="548C8804FFA5FFA419641279FE77F9FC" box="[328,427,1573,1597]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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<specimenCode id="DB919EDDFFA5FFA4199D1279FDF7F9FD" box="[433,555,1572,1597]" pageId="2" pageNumber="3">GPIH 5078</specimenCode>
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(CCGG no. 3811), a complete
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<specimenCount id="9D31FD2FFFA5FFA418C41218FEF0F99D" box="[232,300,1604,1628]" pageId="2" pageNumber="3" type="female">female</specimenCount>
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preserved in a piece of Baltic amber dated as
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<geologicalTimeScale id="F917B60DFFA5FFA418AD1238FF23F9BA" box="[129,255,1636,1659]" pageId="2" pageNumber="3">Late Eocene</geologicalTimeScale>
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(Priabonian).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32D652DFFA5FFA418AD12C1FC7AFF26" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="8B8836A6FFA5FFA418AD12C1FC7AFF26" blockId="2.[129,778,1693,1905]" lastBlockId="2.[825,1474,175,231]" pageId="2" pageNumber="3">
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<emphasis id="B943EAB4FFA5FFA418AD12C1FF3BF975" box="[129,231,1693,1716]" italics="true" pageId="2" pageNumber="3">Diagnosis:</emphasis>
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Ventral coronal teeth absent, fully developed ventral transverse longitudinal carina absent. Antennomeres 4 and 5 not conspicuously shortened, maxillary palp elongate, five-segmented. Median mesoscutal sulcus at most developed for a short distance anteriorly, notauli absent, mesoscutellum triangular, acute, raised compared to surrounding sclerites. Forewing vein 1r arises from approximately middle of pterostigma; vein 1r-Rs well developed, elongate, discal cell rectangular, proximal part not broader than distal part. Ovipositor internalized, extending into thorax, profurca with median groove and abdominal sterna with median apodemes for handling ovipositor.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D652DFFA5FFAE1B151555FDEAFF06" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3" type="description">
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||
<paragraph id="8B8836A6FFA5FFA41B151555FB3EFE5C" blockId="2.[825,1474,264,413]" pageId="2" pageNumber="3">
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||
<emphasis id="B943EAB4FFA5FFA41B151555FC6CFEE1" box="[825,944,265,288]" italics="true" pageId="2" pageNumber="3">Description:</emphasis>
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Female (Fig. 1). Body length 6.
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<quantity id="4CCF9B43FFA5FFA41D381554FA91FEDE" box="[1300,1357,264,288]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="3" unit="mm" value="8.0">8 mm</quantity>
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. Specimen generally well preserved; body with opaque film somewhat obscuring sculpture and pilosity, as well as original coloration; cracks in matrix around specimen also obscures certain features. Right hind tarsus cut off during trimming.
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</paragraph>
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<paragraph id="8B8836A6FFA5FFA41B1515E3FC2EFA85" blockId="2.[825,1474,447,1348]" pageId="2" pageNumber="3">
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<emphasis id="B943EAB4FFA5FFA41B1515E3FCAAFE16" box="[825,886,447,471]" italics="true" pageId="2" pageNumber="3">Head:</emphasis>
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Ocellar corona narrow, distance between median ocellus and lateral coronal tooth subequal to ocellar width; ventral coronal teeth absent (
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<figureCitation id="130C2A23FFA5FFA41C2F15A1FB8FFDD4" box="[1027,1107,509,533]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" figureDoi="http://doi.org/10.5281/zenodo.14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="2" pageNumber="3">Figs 1A</figureCitation>
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,
|
||
<figureCitation id="130C2A23FFA5FFA41C4F15A1FBA2FDD4" box="[1123,1150,509,533]" captionStart="Figure 2" captionStartId="4.[129,192,1671,1695]" captionTargetBox="[129,1473,144,1643]" captionTargetId="figure-89@4.[129,1473,144,1643]" captionTargetPageId="4" captionText="Figure 2. †Kryptovelona carstengroehni gen* et sp* nov*,female holotype (GPIH 5078, CCGG nº 3811)* A,C, photomicrographs* B, D, rendered from synchrotron scans* A, B, head, anterior view* C, head, lateroventral view* D, anterior view of tentorium* Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; greenarrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge*" figureDoi="http://doi.org/10.5281/zenodo.14291329" httpUri="https://zenodo.org/record/14291329/files/figure.png" pageId="2" pageNumber="3">2B</figureCitation>
|
||
). Dorsal transverse and longitudinal frontal carinae absent. Ventral transverse frontal carina not continuous medially, extending across toruli and laterally continuous with lateral carina of subantennal groove (Fig. 2A, B). Frons and vertex areolate-punctate, genae with scattered punctures; frons, vertex, and genae covered with fine, slender hairs (Fig. 2A, C). Internally, median and lateral frontal septa well developed. No conspicuous hairs just posteriorly of eyes. Postocular and occipital carinae absent (Fig. 2C). Lateral carina of subantennal groove low and short, not extending beyond posterior margin of eye. Tentorium with broad median and lateral lamellae on anterior arm, lamellae terminating some distance from anterior tentorial pit; tentorial bridge narrow, arched, with short anterior process; dorsal tentorial arms arise posteriorly on anterior arms, short, apparently do not reach head capsule (Fig. 2D). Antenna with 10 antennomeres, covered with short hairs; scapus elongate, cylindrical, slightly curved; combined length of antennomeres 4 and 5 longer than antennomere 6; antennomere 9 swollen subapically, as long as combined length of antennomeres 7 and 8, without carina laterally; antennomere 10 short, cylindrical, inserted subapically on antennomere 9 (
|
||
<figureCitation id="130C2A23FFA5FFA41B6810D3FC45FB66" box="[836,921,1167,1191]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" figureDoi="http://doi.org/10.5281/zenodo.14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="2" pageNumber="3">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation id="130C2A23FFA5FFA41B8710D3FC16FB66" box="[939,970,1167,1191]" captionStart="Figure 3" captionStartId="5.[113,178,1815,1839]" captionTargetBox="[114,1458,144,1788]" captionTargetId="figure-7@5.[114,1458,144,1788]" captionTargetPageId="5" captionText="Figure 3. A, B, †Cretorussus vilhelmseni Jouault et al.2021, female holotype IGR.BU-020. C, †Kryptovelona groehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †Kryptovelona in italics, with homologues in † Cretorussus in brackets); tb1 = fore tibia." figureDoi="http://doi.org/10.5281/zenodo.14291331" httpUri="https://zenodo.org/record/14291331/files/figure.png" pageId="2" pageNumber="3">3C</figureCitation>
|
||
). Labrum visible between opened mandibles, small, globose, with distinct ventrally directed hairs (Fig. 2A). Mandibles chisel-like, no teeth developed, with conspicuous distally pointing hairs laterally (Fig. 2A, B). Maxillary palps elongate, five-segmented, labial palps short, distal palpomere swollen (Fig. 2C).
|
||
</paragraph>
|
||
<paragraph id="8B8836A6FFA5FFA21B151139FF1BF8BB" blockId="2.[825,1474,1381,1906]" lastBlockId="4.[129,777,1827,1914]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
|
||
<emphasis id="B943EAB4FFA5FFA41B151139FC54FABD" box="[825,904,1381,1404]" italics="true" pageId="2" pageNumber="3">Thorax:</emphasis>
|
||
Pronotum dorsally of equal length throughout, only shallowly incurved posteriorly; lateral sculpture obscured by pilosity/film (Fig. 4A, B). Prosternum mostly concealed by propleura, trapezoid, lateral margins converging anteriorly, discrimen and discrimenal lamella distinct; profurcal arms extend obliquely anteriorly from posterior corners of prosternum, profurcal bridge high and straight, with distinct median groove, not bent posteriorly (Fig. 5B, C). Fore coxa not expanded medially (Fig. 1C); fore femur without ventral carina; fore tibia swollen distally and subdivided by groove; fore tarsus with three tarsomeres, basitarsus elongate and terminating in spur overlapping tarsomere 2 (Fig. 3C). Mesoscutum finely punctate, mostly covered with short hairs, with scattered punctures laterally, median mesoscutal sulcus at most developed anteriorly, notauli absent, parapsides present laterally and transscutal articulation posteriorly (Fig. 4B, C); axillar flanges well developed; scutoscutellar sulcus well developed; mesoscutellum finely punctate, covered with hairs, acutely triangular posteriorly, raised and with carinate lateral margins; mesoscutellar arm raised, without pit anteriorly; mesonotum continuous posterior to mesoscutellum (Fig. 4C). Mesopleuron areolate—punctate laterally, mesosubalar carina present, mesepisternal carina absent; with mid-coxa subdivided, with distinct lateral carina. Metanotum metascutellum not developed, median metanotal carina not observed, lateral metanotal carina present, reaching posterior margin of metanotum; metapleuron with few hairs, mostly glabrous (Fig. 4A); hind coxae not especially hairy laterally, with
|
||
<pageTitle id="CBA8EEC1FFA4FFA51B0613DEFA6FF878" pageId="3" pageNumber="4">rounded medioventral margin and well-developed posterolateral carina; hind femur without ventral carina or denticles ventrally;</pageTitle>
|
||
hind tibia with distinct pegs dorsally, longitudinal and ventral carinae not developed; apical hind tibial spurs short, of unequal length.
|
||
</paragraph>
|
||
<caption id="DF48662EFFA4FFA5185E12EDFA4EF8C3" ID-DOI="http://doi.org/10.5281/zenodo.14291327" ID-Zenodo-Dep="14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="3" pageNumber="4" startId="3.[114,179,1713,1737]" targetBox="[114,1458,144,1686]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B8836A6FFA4FFA5185E12EDFA4EF8C3" blockId="3.[113,1438,1713,1794]" pageId="3" pageNumber="4">
|
||
<emphasis id="B943EAB4FFA4FFA5185E12EDFF16F908" bold="true" box="[114,202,1713,1737]" pageId="3" pageNumber="4">Figure 1.</emphasis>
|
||
†
|
||
<taxonomicName id="4C374D25FFA4FFA518F612EEFE06F90B" box="[218,474,1714,1738]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="carstengroehni">
|
||
<emphasis id="B943EAB4FFA4FFA518F612EEFE06F90B" box="[218,474,1714,1738]" italics="true" pageId="3" pageNumber="4">Kryptovelona carstengroehni</emphasis>
|
||
</taxonomicName>
|
||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, dorsal habitus. B, lateral habitus. C, ventral habitus. White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF48662EFFA3FFA218AD12DBFA77F932" ID-DOI="http://doi.org/10.5281/zenodo.14291329" ID-Zenodo-Dep="14291329" httpUri="https://zenodo.org/record/14291329/files/figure.png" pageId="4" pageNumber="5" startId="4.[129,192,1671,1695]" targetBox="[129,1473,144,1643]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8836A6FFA3FFA218AD12DBFA77F932" blockId="4.[128,1474,1671,1779]" pageId="4" pageNumber="5">
|
||
<emphasis id="B943EAB4FFA3FFA218AD12DBFF0AF95E" bold="true" box="[129,214,1671,1695]" pageId="4" pageNumber="5">Figure 2.</emphasis>
|
||
†
|
||
<taxonomicName id="4C374D25FFA3FFA218CA12DBFE03F95E" box="[230,479,1671,1695]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="carstengroehni">
|
||
<emphasis id="B943EAB4FFA3FFA218CA12DBFE03F95E" box="[230,479,1671,1695]" italics="true" pageId="4" pageNumber="5">Kryptovelona carstengroehni</emphasis>
|
||
</taxonomicName>
|
||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, C, photomicrographs. B, D, rendered from synchrotron scans. A, B, head, anterior view. C, head, lateroventral view. D, anterior view of tentorium. Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; green arrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8836A6FFA3FFA218AD13C0FBB8F8BB" blockId="4.[129,777,1947,2003]" lastBlockId="4.[825,1474,1827,1914]" pageId="4" pageNumber="5">
|
||
<pageTitle id="CBA8EEC1FFA3FFA218AD13C0FCD5F813" pageId="4" pageNumber="5">
|
||
<emphasis id="B943EAB4FFA3FFA218AD13C0FF19F872" box="[129,197,1948,1971]" italics="true" pageId="4" pageNumber="5">Wings:</emphasis>
|
||
Forewing vein 2r-rs arise from approximately middle of pterostigma; vein 1-Rs well developed, elongate, discal cell
|
||
</pageTitle>
|
||
rectangular, proximal part not broader than distal part; vein cu-a inserts on Cu1 close to middle of cell M (Fig. 1A). Hindwing venation cannot be observed.
|
||
</paragraph>
|
||
<paragraph id="8B8836A6FFA3FFAE1B1513C7FDEAFF06" blockId="4.[825,1474,1947,2002]" lastBlockId="8.[129,566,175,199]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
|
||
<pageTitle id="CBA8EEC1FFA3FFA21B1513C7FA1EF813" pageId="4" pageNumber="5">
|
||
<emphasis id="B943EAB4FFA3FFA21B1513C7FC43F872" box="[825,927,1947,1971]" italics="true" pageId="4" pageNumber="5">Abdomen:</emphasis>
|
||
Dorsally covered by wings, hence obscuring structural detail via light microscopy; laterally with dense pilosity
|
||
</pageTitle>
|
||
of short hairs. Tergum 1 and 2 coarsely areolate (Fig. 4C), postspiracular and subspiracular carinae on tergum 1 absent, median carina on tergum 2 absent; tergum 3–8 finely punctate. Median apodemes observed on several abdominal sterna (Fig. 5D); sternum 7 externally with narrow median part delimited laterally and projecting posteriorly. Tergum 9 covered with short hairs, without longitudinal carinae, smooth areas and depressions; internally, prominent anterior flange and chordate apodemes present. Ovipositor apparatus apparently not fully preserved (Fig. 5A); second valvifer with traces of ventral
|
||
<collectionCode id="ED26AE63FFA0FFA118F713C3FF37F876" box="[219,235,1951,1975]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="8">T</collectionCode>
|
||
9-second valvifer muscles [muscle
|
||
<quantity id="4CCF9B43FFA0FFA11A7A13C2FDA2F876" box="[598,638,1950,1975]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="8" unit="in" value="6.0">6 in</quantity>
|
||
Vilhelmsen
|
||
<emphasis id="B943EAB4FFA0FFA11B06137EFC86F8F8" box="[810,858,1825,1849]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
(2001)] observable, possible base of ovipositor shaft with remains of processus medianus present; only parts of internalized ovipositor loop traceable, ventral/distal part extends
|
||
<pageTitle id="CBA8EEC1FFA0FFA11B061323FA6FF876" pageId="7" pageNumber="8">anteriorly at least to mesofurca, manner of coiling in prothorax not observed; tip of ovipositor slightly protruding from tip of</pageTitle>
|
||
abdomen, third valvulae not visible. Composite tergum 10/ cercus triangular, not continuous medially.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF48662EFFA2FFA3185D134BFE26F861" ID-DOI="http://doi.org/10.5281/zenodo.14291331" ID-Zenodo-Dep="14291331" httpUri="https://zenodo.org/record/14291331/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1815,1839]" targetBox="[114,1458,144,1788]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8836A6FFA2FFA3185D134BFE26F861" blockId="5.[113,1454,1815,1924]" lastBlockId="5.[125,506,1928,1952]" pageId="5" pageNumber="6">
|
||
<emphasis id="B943EAB4FFA2FFA3185D134BFF15F8EE" bold="true" box="[113,201,1815,1839]" pageId="5" pageNumber="6">Figure 3.</emphasis>
|
||
A, B, †
|
||
<taxonomicName id="4C374D25FFA2FFA319201344FD55F8F1" authority="Jouault et al. 2021" authorityName="Jouault" authorityYear="2021" box="[268,649,1816,1840]" class="Insecta" family="Burmorussidae" genus="Cretorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vilhelmseni">
|
||
<emphasis id="B943EAB4FFA2FFA319201344FE00F8F1" box="[268,476,1816,1840]" italics="true" pageId="5" pageNumber="6">Cretorussus vilhelmseni</emphasis>
|
||
<bibRefCitation id="EFA64B57FFA2FFA319CE1344FD55F8F1" author="Jouault" box="[482,649,1816,1840]" etAl="et al." firstAuthor="Jouault" pageId="5" pageNumber="6" pagination="324 - 31" refId="ref10723" refString="Jouault C, Perrichot V, Nel A * Taxonomic addition to early diverged parasitic wasps (Hymenoptea: Orussoidea) from mid-Cretaceous Burmese amber * Proceedings of the Geologists' Association 2021; 132: 324 - 31 * hups: // doi * org / 10 * 1016 / j * pgeola * 2021 * 02 * 002" type="journal article" year="2021">
|
||
Jouault
|
||
<emphasis id="B943EAB4FFA2FFA31A061344FD8AF8F1" box="[554,598,1816,1840]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, female holotype IGR.BU-020. C, †
|
||
<taxonomicName id="4C374D25FFA2FFA31BF21344FB42F8F1" box="[990,1182,1816,1840]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="groehni">
|
||
<emphasis id="B943EAB4FFA2FFA31BF21344FB42F8F1" box="[990,1182,1816,1840]" italics="true" pageId="5" pageNumber="6">Kryptovelona groehni</emphasis>
|
||
</taxonomicName>
|
||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †
|
||
<taxonomicName id="4C374D25FFA2FFA31BCA1330FB83F845" box="[998,1119,1900,1924]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFA2FFA31BCA1330FB83F845" box="[998,1119,1900,1924]" italics="true" pageId="5" pageNumber="6">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
in italics, with homologues in †
|
||
<taxonomicName id="4C374D25FFA2FFA3185113D4FF39F85E" box="[125,229,1928,1951]" class="Insecta" family="Burmorussidae" genus="Cretorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFA2FFA3185113D4FF39F85E" box="[125,229,1928,1951]" italics="true" pageId="5" pageNumber="6">Cretorussus</emphasis>
|
||
</taxonomicName>
|
||
in brackets); tb1 = fore tibia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF48662EFFA1FFA018AD12CEFE57F8DB" ID-DOI="http://doi.org/10.5281/zenodo.14291333" ID-Zenodo-Dep="14291333" httpUri="https://zenodo.org/record/14291333/files/figure.png" pageId="6" pageNumber="7" startId="6.[129,194,1682,1706]" targetBox="[129,1473,144,1654]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B8836A6FFA1FFA018AD12CEFE57F8DB" blockId="6.[129,1453,1682,1818]" pageId="6" pageNumber="7">
|
||
<emphasis id="B943EAB4FFA1FFA018AD12CEFF35F96B" bold="true" box="[129,233,1682,1706]" pageId="6" pageNumber="7">Figure 4. †</emphasis>
|
||
<taxonomicName id="4C374D25FFA1FFA018C512CEFE34F96B" box="[233,488,1682,1706]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carstengroehni">
|
||
<emphasis id="B943EAB4FFA1FFA018C512CEFE34F96B" box="[233,488,1682,1706]" italics="true" pageId="6" pageNumber="7">Kryptovelona carstengroehni</emphasis>
|
||
</taxonomicName>
|
||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF48662EFFA0FFA1185D12D9FB2CF930" ID-DOI="http://doi.org/10.5281/zenodo.14291335" ID-Zenodo-Dep="14291335" httpUri="https://zenodo.org/record/14291335/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,178,1669,1693]" targetBox="[114,1454,149,1639]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B8836A6FFA0FFA1185D12D9FB2CF930" blockId="7.[113,1421,1669,1778]" pageId="7" pageNumber="8">
|
||
<emphasis id="B943EAB4FFA0FFA1185D12D9FF15F95F" bold="true" box="[113,201,1669,1694]" pageId="7" pageNumber="8">Figure 5.</emphasis>
|
||
†
|
||
<taxonomicName id="4C374D25FFA0FFA118F612D9FE05F95C" box="[218,473,1669,1693]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="carstengroehni">
|
||
<emphasis id="B943EAB4FFA0FFA118F612D9FE05F95C" box="[218,473,1669,1693]" italics="true" pageId="7" pageNumber="8">Kryptovelona carstengroehni</emphasis>
|
||
</taxonomicName>
|
||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, dorsal overview of extent of ovipositor inside the body; anterior to the top. B, profurca in posterior view. C, profurca in anterolateral view. D, posterior part of abdomen, dorsal internal view. All views rendered from synchrotron scans. Red arrows = ovipositor; black arrows = medain apodemes on abdominal sternum 6 and 7; green arrows = median groove on profurcal bridge; dl1 = prodiscrimenal lamella; pfb = profurcal bridge.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32D652DFFAFFFAE18AD14B5FD72FEBF" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="8B8836A6FFAFFFAE18AD14B5FD72FEBF" blockId="8.[129,778,233,382]" pageId="8" pageNumber="9">
|
||
<emphasis id="B943EAB4FFAFFFAE18AD14B5FF2CFEC0" box="[129,240,233,257]" italics="true" pageId="8" pageNumber="9">Etymology:</emphasis>
|
||
The genus name is a combination of
|
||
<emphasis id="B943EAB4FFAFFFAE1AB414B5FD04FEC0" box="[664,728,233,257]" italics="true" pageId="8" pageNumber="9">krypto</emphasis>
|
||
and
|
||
<emphasis id="B943EAB4FFAFFFAE18AD1554FF63FEE1" box="[129,191,264,288]" italics="true" pageId="8" pageNumber="9">velona</emphasis>
|
||
, the Greek words for hidden and needle, respectively, alluding to the concealed ovipositor of the family. The species’ epithet honours Mr Carsten Gröhn (Glinde,
|
||
<collectingCountry id="F3207636FFAFFFAE1A44151BFD17FE9E" box="[616,715,327,351]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
|
||
) who generously made the
|
||
<typeStatus id="548C8804FFAFFFAE1974153AFE5AFEBF" box="[344,390,358,382]" pageId="8" pageNumber="9">type</typeStatus>
|
||
specimen available for study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D652DFFAFFFAE18AD15FCFF34FBA8" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="8B8836A6FFAFFFAE18AD15FCFDC4FD10" blockId="8.[129,778,415,1129]" pageId="8" pageNumber="9">
|
||
<emphasis id="B943EAB4FFAFFFAE18AD15FCFF2DFE76" box="[129,241,416,439]" italics="true" pageId="8" pageNumber="9">Comments:</emphasis>
|
||
Although both †
|
||
<taxonomicName id="4C374D25FFAFFFAE19B015C3FDC3FE76" box="[412,543,415,439]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE19B015C3FDC3FE76" box="[412,543,415,439]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C374D25FFAFFFAE1A7015C3FD17FE76" authorityName="Schedl" authorityYear="2011" box="[604,715,415,439]" class="Insecta" family="Orussidae" genus="Baltorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE1A7015C3FD17FE76" box="[604,715,415,439]" italics="true" pageId="8" pageNumber="9">Baltorussus</emphasis>
|
||
</taxonomicName>
|
||
occur in Baltic amber, there are sufficient differences to justify recognizing them as separate genera. Besides, they do not group together in the phylogenetic analyses, where †
|
||
<taxonomicName id="4C374D25FFAFFFAE1A4115A1FD2CFDD4" box="[621,752,509,533]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE1A4115A1FD2CFDD4" box="[621,752,509,533]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
is placed closer to the clade comprising the majority of the extant taxa. The RoguePlots (Supporting Information,
|
||
<figureCitation id="130C2A23FFAFFFAE1A441660FD6DFD95" box="[616,689,572,596]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" figureDoi="http://doi.org/10.5281/zenodo.14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="8" pageNumber="9">Figs S1</figureCitation>
|
||
, S
|
||
<figureCitation id="130C2A23FFAFFFAE1AE41660FD09FD95" box="[712,725,572,596]" captionStart="Figure 4" captionStartId="6.[129,194,1682,1706]" captionTargetBox="[129,1473,144,1654]" captionTargetId="figure-64@6.[129,1473,144,1654]" captionTargetPageId="6" captionText="Figure 4. †Kryptovelona carstengroehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum." figureDoi="http://doi.org/10.5281/zenodo.14291333" httpUri="https://zenodo.org/record/14291333/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
|
||
) flag a possible †
|
||
<taxonomicName id="4C374D25FFAFFFAE19291607FE54FDB2" box="[261,392,603,627]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE19291607FE54FDB2" box="[261,392,603,627]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
–†
|
||
<taxonomicName id="4C374D25FFAFFFAE198F1607FDCEFDB2" authorityName="Schedl" authorityYear="2011" box="[419,530,603,627]" class="Insecta" family="Orussidae" genus="Baltorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE198F1607FDCEFDB2" box="[419,530,603,627]" italics="true" pageId="8" pageNumber="9">Baltorussus</emphasis>
|
||
</taxonomicName>
|
||
sister-relationship, but with lower probability than in the preferred Bayesian tree; other alternative placements of †
|
||
<taxonomicName id="4C374D25FFAFFFAE19BE16C6FDC9FD73" box="[402,533,666,690]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE19BE16C6FDC9FD73" box="[402,533,666,690]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
closer to the base of extant
|
||
<taxonomicName id="4C374D25FFAFFFAE188216E5FECAFD10" authorityName="Newman" authorityYear="1834" box="[174,278,697,721]" class="Insecta" family="Orussidae" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Orussidae</taxonomicName>
|
||
have even lower support.
|
||
</paragraph>
|
||
<paragraph id="8B8836A6FFAFFFAE18B01685FF34FBA8" blockId="8.[129,778,415,1129]" pageId="8" pageNumber="9">
|
||
The sculpture of the frons is highly diagnostic in †
|
||
<taxonomicName id="4C374D25FFAFFFAE1AB61684FCD5FD31" authorityName="Schedl" authorityYear="2011" box="[666,777,728,752]" class="Insecta" family="Orussidae" genus="Baltorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE1AB61684FCD5FD31" box="[666,777,728,752]" italics="true" pageId="8" pageNumber="9">Baltorussus</emphasis>
|
||
</taxonomicName>
|
||
(Vilhelmsen and Zimmermann 2014: fig. 3A, B); this is not observed in †
|
||
<taxonomicName id="4C374D25FFAFFFAE1922174BFE4DFCEE" authority="Ne" authorityName="Ne" box="[270,401,791,815]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE1922174BFE4DFCEE" box="[270,401,791,815]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
. The latter has much denser pilosity over large areas of the body, in particular the dorsal part of the thorax. The median mesoscutellar sulcus is well developed in †
|
||
<taxonomicName id="4C374D25FFAFFFAE18A21729FF21FC4C" authorityName="Schedl" authorityYear="2011" box="[142,253,885,909]" class="Insecta" family="Orussidae" genus="Baltorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE18A21729FF21FC4C" box="[142,253,885,909]" italics="true" pageId="8" pageNumber="9">Baltorussus</emphasis>
|
||
</taxonomicName>
|
||
, extending all the way to the transscutal articulation (Vilhelmsen and Zimmermann 2014: fig. 7), whereas it is absent in †
|
||
<taxonomicName id="4C374D25FFAFFFAE18DC17E8FEAFFC0D" box="[240,371,948,972]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE18DC17E8FEAFFC0D" box="[240,371,948,972]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
; the notauli are also more developed in the former, extending approximately halfway to the articulation, whereas they are not developed in †
|
||
<taxonomicName id="4C374D25FFAFFFAE1A2C17AEFD5FFBCB" authority="In" authorityName="In" box="[512,643,1010,1034]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE1A2C17AEFD5FFBCB" box="[512,643,1010,1034]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
. In the forewing of †
|
||
<taxonomicName id="4C374D25FFAFFFAE18CE104EFEB9FBEB" box="[226,357,1042,1066]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE18CE104EFEB9FBEB" box="[226,357,1042,1066]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||
</taxonomicName>
|
||
, vein cu-a arises from approximately the middle of the discal cell; in †
|
||
<taxonomicName id="4C374D25FFAFFFAE198A106DFDC9FB88" authorityName="Schedl" authorityYear="2011" box="[422,533,1073,1097]" class="Insecta" family="Orussidae" genus="Baltorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B943EAB4FFAFFFAE198A106DFDC9FB88" box="[422,533,1073,1097]" italics="true" pageId="8" pageNumber="9">Baltorussus</emphasis>
|
||
</taxonomicName>
|
||
it is placed much closer to vein M.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |