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†
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<taxonomicName id="4C374D25FFA5FFA418BB115FFD23FADF" authority="Vilhelmsen and Perrichot" authorityName="Vilhelmsen and Perrichot" box="[151,767,1283,1310]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="carstengroehni" status="gen.">
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Vilhelmsen and Perrichot, gen.
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(
|
||||
<figureCitation id="130C2A23FFA5FFA419B41111FE2EFAA6" box="[408,498,1357,1383]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[114,179,1713,1737]" captionStartId-1="4.[129,192,1671,1695]" captionStartId-2="5.[113,178,1815,1839]" captionStartId-3="6.[129,194,1682,1706]" captionStartId-4="7.[113,178,1669,1693]" captionTargetBox-0="[114,1458,144,1686]" captionTargetBox-1="[129,1473,144,1643]" captionTargetBox-2="[114,1458,144,1788]" captionTargetBox-3="[129,1473,144,1654]" captionTargetBox-4="[114,1454,149,1639]" captionTargetId-0="figure-88@3.[114,1458,144,1686]" captionTargetId-1="figure-89@4.[129,1473,144,1643]" captionTargetId-2="figure-7@5.[114,1458,144,1788]" captionTargetId-3="figure-64@6.[129,1473,144,1654]" captionTargetId-4="figure-101@7.[113,1460,144,1641]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" captionText-1="Figure 2. †Kryptovelona carstengroehni gen* et sp* nov*,female holotype (GPIH 5078, CCGG nº 3811)* A,C, photomicrographs* B, D, rendered from synchrotron scans* A, B, head, anterior view* C, head, lateroventral view* D, anterior view of tentorium* Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; greenarrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge*" captionText-2="Figure 3. A, B, †Cretorussus vilhelmseni Jouault et al.2021, female holotype IGR.BU-020. C, †Kryptovelona groehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †Kryptovelona in italics, with homologues in † Cretorussus in brackets); tb1 = fore tibia." captionText-3="Figure 4. †Kryptovelona carstengroehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum." captionText-4="Figure 5. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal overview of extent of ovipositor inside the body; anterior to the top* B, profurca in posterior view* C, profurca in anterolateral view* D, posterior part of abdomen, dorsal internal view* All views rendered from synchrotron scans* Red arrows = ovipositor; black arrows = medain apodemes on abdominal sternum 6 and 7; green arrows = median groove on profurcal bridge; dl1 = prodiscrimenal lamella; pOE = profurcal bridge*" pageId="2" pageNumber="3">Figs 1–5</figureCitation>
|
||||
<figureCitation id="130C2A23FFA5FFA419B41111FE2EFAA6" box="[408,498,1357,1383]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[114,179,1713,1737]" captionStartId-1="4.[129,192,1671,1695]" captionStartId-2="5.[113,178,1815,1839]" captionStartId-3="6.[129,194,1682,1706]" captionStartId-4="7.[113,178,1669,1693]" captionTargetBox-0="[114,1458,144,1686]" captionTargetBox-1="[129,1473,144,1643]" captionTargetBox-2="[114,1458,144,1788]" captionTargetBox-3="[129,1473,144,1654]" captionTargetBox-4="[114,1454,149,1639]" captionTargetId-0="figure-88@3.[114,1458,144,1686]" captionTargetId-1="figure-89@4.[129,1473,144,1643]" captionTargetId-2="figure-7@5.[114,1458,144,1788]" captionTargetId-3="figure-64@6.[129,1473,144,1654]" captionTargetId-4="figure-101@7.[113,1460,144,1641]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" captionText-1="Figure 2. †Kryptovelona carstengroehni gen* et sp* nov*,female holotype (GPIH 5078, CCGG nº 3811)* A,C, photomicrographs* B, D, rendered from synchrotron scans* A, B, head, anterior view* C, head, lateroventral view* D, anterior view of tentorium* Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; greenarrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge*" captionText-2="Figure 3. A, B, †Cretorussus vilhelmseni Jouault et al.2021, female holotype IGR.BU-020. C, †Kryptovelona groehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †Kryptovelona in italics, with homologues in † Cretorussus in brackets); tb1 = fore tibia." captionText-3="Figure 4. †Kryptovelona carstengroehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum." captionText-4="Figure 5. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal overview of extent of ovipositor inside the body; anterior to the top* B, profurca in posterior view* C, profurca in anterolateral view* D, posterior part of abdomen, dorsal internal view* All views rendered from synchrotron scans* Red arrows = ovipositor; black arrows = medain apodemes on abdominal sternum 6 and 7; green arrows = median groove on profurcal bridge; dl1 = prodiscrimenal lamella; pOE = profurcal bridge*" figureDoi-0="http://doi.org/10.5281/zenodo.14291327" figureDoi-1="http://doi.org/10.5281/zenodo.14291329" figureDoi-2="http://doi.org/10.5281/zenodo.14291331" figureDoi-3="http://doi.org/10.5281/zenodo.14291333" figureDoi-4="http://doi.org/10.5281/zenodo.14291335" httpUri-0="https://zenodo.org/record/14291327/files/figure.png" httpUri-1="https://zenodo.org/record/14291329/files/figure.png" httpUri-2="https://zenodo.org/record/14291331/files/figure.png" httpUri-3="https://zenodo.org/record/14291333/files/figure.png" httpUri-4="https://zenodo.org/record/14291335/files/figure.png" pageId="2" pageNumber="3">Figs 1–5</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</heading>
|
||||
|
|
@ -122,13 +121,11 @@ preserved in a piece of Baltic amber dated as
|
|||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32D652DFFA5FFA418AD12C1FC7AFF26" pageId="2" pageNumber="3" type="diagnosis">
|
||||
<paragraph id="8B8836A6FFA5FFA418AD12C1FCD5F86E" blockId="2.[129,778,1693,1905]" lastBlockId="2.[129,777,1912,1967]" pageId="2" pageNumber="3">
|
||||
<paragraph id="8B8836A6FFA5FFA418AD12C1FC7AFF26" blockId="2.[129,778,1693,1905]" lastBlockId="2.[825,1474,175,231]" pageId="2" pageNumber="3">
|
||||
<emphasis id="B943EAB4FFA5FFA418AD12C1FF3BF975" box="[129,231,1693,1716]" italics="true" pageId="2" pageNumber="3">Diagnosis:</emphasis>
|
||||
Ventral coronal teeth absent, fully developed ventral transverse longitudinal carina absent. Antennomeres 4 and 5 not conspicuously shortened, maxillary palp elongate, five-segmented. Median mesoscutal sulcus at most developed for a short distance anteriorly, notauli absent, mesoscutellum triangular, acute, raised compared to surrounding sclerites. Forewing vein 1r arises from approximately middle of pterostigma; vein 1r-Rs well developed, elongate, discal cell rectangular, proximal part not broader than distal part. Ovipositor
|
||||
Ventral coronal teeth absent, fully developed ventral transverse longitudinal carina absent. Antennomeres 4 and 5 not conspicuously shortened, maxillary palp elongate, five-segmented. Median mesoscutal sulcus at most developed for a short distance anteriorly, notauli absent, mesoscutellum triangular, acute, raised compared to surrounding sclerites. Forewing vein 1r arises from approximately middle of pterostigma; vein 1r-Rs well developed, elongate, discal cell rectangular, proximal part not broader than distal part. Ovipositor internalized, extending into thorax, profurca with median groove and abdominal sterna with median apodemes for handling ovipositor.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFA5FFA41B1514CCFC7AFF26" blockId="2.[825,1474,144,168]" lastBlockId="2.[825,1474,175,231]" pageId="2" pageNumber="3">internalized, extending into thorax, profurca with median groove and abdominal sterna with median apodemes for handling ovipositor.</paragraph>
|
||||
</subSubSection>
|
||||
</subSection>
|
||||
<subSubSection id="C32D652DFFA5FFAE1B151555FDEAFF06" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3" type="description">
|
||||
<paragraph id="8B8836A6FFA5FFA41B151555FB3EFE5C" blockId="2.[825,1474,264,413]" pageId="2" pageNumber="3">
|
||||
<emphasis id="B943EAB4FFA5FFA41B151555FC6CFEE1" box="[825,944,265,288]" italics="true" pageId="2" pageNumber="3">Description:</emphasis>
|
||||
|
|
@ -139,20 +136,22 @@ Female (Fig. 1). Body length 6.
|
|||
<paragraph id="8B8836A6FFA5FFA41B1515E3FC2EFA85" blockId="2.[825,1474,447,1348]" pageId="2" pageNumber="3">
|
||||
<emphasis id="B943EAB4FFA5FFA41B1515E3FCAAFE16" box="[825,886,447,471]" italics="true" pageId="2" pageNumber="3">Head:</emphasis>
|
||||
Ocellar corona narrow, distance between median ocellus and lateral coronal tooth subequal to ocellar width; ventral coronal teeth absent (
|
||||
<figureCitation id="130C2A23FFA5FFA41C2F15A1FB8FFDD4" box="[1027,1107,509,533]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" pageId="2" pageNumber="3">Figs 1A</figureCitation>
|
||||
<figureCitation id="130C2A23FFA5FFA41C2F15A1FB8FFDD4" box="[1027,1107,509,533]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" figureDoi="http://doi.org/10.5281/zenodo.14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="2" pageNumber="3">Figs 1A</figureCitation>
|
||||
,
|
||||
<figureCitation id="130C2A23FFA5FFA41C4F15A1FBA2FDD4" box="[1123,1150,509,533]" captionStart="Figure 2" captionStartId="4.[129,192,1671,1695]" captionTargetBox="[129,1473,144,1643]" captionTargetId="figure-89@4.[129,1473,144,1643]" captionTargetPageId="4" captionText="Figure 2. †Kryptovelona carstengroehni gen* et sp* nov*,female holotype (GPIH 5078, CCGG nº 3811)* A,C, photomicrographs* B, D, rendered from synchrotron scans* A, B, head, anterior view* C, head, lateroventral view* D, anterior view of tentorium* Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; greenarrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge*" pageId="2" pageNumber="3">2B</figureCitation>
|
||||
<figureCitation id="130C2A23FFA5FFA41C4F15A1FBA2FDD4" box="[1123,1150,509,533]" captionStart="Figure 2" captionStartId="4.[129,192,1671,1695]" captionTargetBox="[129,1473,144,1643]" captionTargetId="figure-89@4.[129,1473,144,1643]" captionTargetPageId="4" captionText="Figure 2. †Kryptovelona carstengroehni gen* et sp* nov*,female holotype (GPIH 5078, CCGG nº 3811)* A,C, photomicrographs* B, D, rendered from synchrotron scans* A, B, head, anterior view* C, head, lateroventral view* D, anterior view of tentorium* Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; greenarrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge*" figureDoi="http://doi.org/10.5281/zenodo.14291329" httpUri="https://zenodo.org/record/14291329/files/figure.png" pageId="2" pageNumber="3">2B</figureCitation>
|
||||
). Dorsal transverse and longitudinal frontal carinae absent. Ventral transverse frontal carina not continuous medially, extending across toruli and laterally continuous with lateral carina of subantennal groove (Fig. 2A, B). Frons and vertex areolate-punctate, genae with scattered punctures; frons, vertex, and genae covered with fine, slender hairs (Fig. 2A, C). Internally, median and lateral frontal septa well developed. No conspicuous hairs just posteriorly of eyes. Postocular and occipital carinae absent (Fig. 2C). Lateral carina of subantennal groove low and short, not extending beyond posterior margin of eye. Tentorium with broad median and lateral lamellae on anterior arm, lamellae terminating some distance from anterior tentorial pit; tentorial bridge narrow, arched, with short anterior process; dorsal tentorial arms arise posteriorly on anterior arms, short, apparently do not reach head capsule (Fig. 2D). Antenna with 10 antennomeres, covered with short hairs; scapus elongate, cylindrical, slightly curved; combined length of antennomeres 4 and 5 longer than antennomere 6; antennomere 9 swollen subapically, as long as combined length of antennomeres 7 and 8, without carina laterally; antennomere 10 short, cylindrical, inserted subapically on antennomere 9 (
|
||||
<figureCitation id="130C2A23FFA5FFA41B6810D3FC45FB66" box="[836,921,1167,1191]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" pageId="2" pageNumber="3">Figs 1C</figureCitation>
|
||||
<figureCitation id="130C2A23FFA5FFA41B6810D3FC45FB66" box="[836,921,1167,1191]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" figureDoi="http://doi.org/10.5281/zenodo.14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="2" pageNumber="3">Figs 1C</figureCitation>
|
||||
,
|
||||
<figureCitation id="130C2A23FFA5FFA41B8710D3FC16FB66" box="[939,970,1167,1191]" captionStart="Figure 3" captionStartId="5.[113,178,1815,1839]" captionTargetBox="[114,1458,144,1788]" captionTargetId="figure-7@5.[114,1458,144,1788]" captionTargetPageId="5" captionText="Figure 3. A, B, †Cretorussus vilhelmseni Jouault et al.2021, female holotype IGR.BU-020. C, †Kryptovelona groehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †Kryptovelona in italics, with homologues in † Cretorussus in brackets); tb1 = fore tibia." pageId="2" pageNumber="3">3C</figureCitation>
|
||||
<figureCitation id="130C2A23FFA5FFA41B8710D3FC16FB66" box="[939,970,1167,1191]" captionStart="Figure 3" captionStartId="5.[113,178,1815,1839]" captionTargetBox="[114,1458,144,1788]" captionTargetId="figure-7@5.[114,1458,144,1788]" captionTargetPageId="5" captionText="Figure 3. A, B, †Cretorussus vilhelmseni Jouault et al.2021, female holotype IGR.BU-020. C, †Kryptovelona groehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs. A, distal part of antenna. B, fore tibia and tarsus. C, distal part of antenna and fore tibia and tarsus. Red arrow = tarsal spur; blue arrow = tibial spur (slightly displaced); green arrow = groove in fore tibia; A[x] = antennomere [x]; t[x] = fore tarsomere [x] (putative composite structures and structures distal to them in †Kryptovelona in italics, with homologues in † Cretorussus in brackets); tb1 = fore tibia." figureDoi="http://doi.org/10.5281/zenodo.14291331" httpUri="https://zenodo.org/record/14291331/files/figure.png" pageId="2" pageNumber="3">3C</figureCitation>
|
||||
). Labrum visible between opened mandibles, small, globose, with distinct ventrally directed hairs (Fig. 2A). Mandibles chisel-like, no teeth developed, with conspicuous distally pointing hairs laterally (Fig. 2A, B). Maxillary palps elongate, five-segmented, labial palps short, distal palpomere swollen (Fig. 2C).
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFA5FFA41B151139FA1EF870" blockId="2.[825,1474,1381,1906]" lastBlockId="2.[825,1474,1914,1969]" pageId="2" pageNumber="3">
|
||||
<paragraph id="8B8836A6FFA5FFA21B151139FF1BF8BB" blockId="2.[825,1474,1381,1906]" lastBlockId="4.[129,777,1827,1914]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
|
||||
<emphasis id="B943EAB4FFA5FFA41B151139FC54FABD" box="[825,904,1381,1404]" italics="true" pageId="2" pageNumber="3">Thorax:</emphasis>
|
||||
Pronotum dorsally of equal length throughout, only shallowly incurved posteriorly; lateral sculpture obscured by pilosity/film (Fig. 4A, B). Prosternum mostly concealed by propleura, trapezoid, lateral margins converging anteriorly, discrimen and discrimenal lamella distinct; profurcal arms extend obliquely anteriorly from posterior corners of prosternum, profurcal bridge high and straight, with distinct median groove, not bent posteriorly (Fig. 5B, C). Fore coxa not expanded medially (Fig. 1C); fore femur without ventral carina; fore tibia swollen distally and subdivided by groove; fore tarsus with three tarsomeres, basitarsus elongate and terminating in spur overlapping tarsomere 2 (Fig. 3C). Mesoscutum finely punctate, mostly covered with short hairs, with scattered punctures laterally, median mesoscutal sulcus at most developed anteriorly, notauli absent, parapsides present laterally and transscutal articulation posteriorly (Fig. 4B, C); axillar flanges well developed; scutoscutellar sulcus well developed; mesoscutellum finely punctate, covered with hairs, acutely triangular posteriorly, raised and with carinate lateral margins; mesoscutellar arm raised,
|
||||
Pronotum dorsally of equal length throughout, only shallowly incurved posteriorly; lateral sculpture obscured by pilosity/film (Fig. 4A, B). Prosternum mostly concealed by propleura, trapezoid, lateral margins converging anteriorly, discrimen and discrimenal lamella distinct; profurcal arms extend obliquely anteriorly from posterior corners of prosternum, profurcal bridge high and straight, with distinct median groove, not bent posteriorly (Fig. 5B, C). Fore coxa not expanded medially (Fig. 1C); fore femur without ventral carina; fore tibia swollen distally and subdivided by groove; fore tarsus with three tarsomeres, basitarsus elongate and terminating in spur overlapping tarsomere 2 (Fig. 3C). Mesoscutum finely punctate, mostly covered with short hairs, with scattered punctures laterally, median mesoscutal sulcus at most developed anteriorly, notauli absent, parapsides present laterally and transscutal articulation posteriorly (Fig. 4B, C); axillar flanges well developed; scutoscutellar sulcus well developed; mesoscutellum finely punctate, covered with hairs, acutely triangular posteriorly, raised and with carinate lateral margins; mesoscutellar arm raised, without pit anteriorly; mesonotum continuous posterior to mesoscutellum (Fig. 4C). Mesopleuron areolate—punctate laterally, mesosubalar carina present, mesepisternal carina absent; with mid-coxa subdivided, with distinct lateral carina. Metanotum metascutellum not developed, median metanotal carina not observed, lateral metanotal carina present, reaching posterior margin of metanotum; metapleuron with few hairs, mostly glabrous (Fig. 4A); hind coxae not especially hairy laterally, with
|
||||
<pageTitle id="CBA8EEC1FFA4FFA51B0613DEFA6FF878" pageId="3" pageNumber="4">rounded medioventral margin and well-developed posterolateral carina; hind femur without ventral carina or denticles ventrally;</pageTitle>
|
||||
hind tibia with distinct pegs dorsally, longitudinal and ventral carinae not developed; apical hind tibial spurs short, of unequal length.
|
||||
</paragraph>
|
||||
<caption id="DF48662EFFA4FFA5185E12EDFA4EF8C3" pageId="3" pageNumber="4" startId="3.[114,179,1713,1737]" targetBox="[114,1458,144,1686]" targetPageId="3" targetType="figure">
|
||||
<caption id="DF48662EFFA4FFA5185E12EDFA4EF8C3" ID-DOI="http://doi.org/10.5281/zenodo.14291327" ID-Zenodo-Dep="14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="3" pageNumber="4" startId="3.[114,179,1713,1737]" targetBox="[114,1458,144,1686]" targetPageId="3" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA4FFA5185E12EDFA4EF8C3" blockId="3.[113,1438,1713,1794]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B943EAB4FFA4FFA5185E12EDFF16F908" bold="true" box="[114,202,1713,1737]" pageId="3" pageNumber="4">Figure 1.</emphasis>
|
||||
†
|
||||
|
|
@ -162,11 +161,7 @@ Pronotum dorsally of equal length throughout, only shallowly incurved posteriorl
|
|||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, dorsal habitus. B, lateral habitus. C, ventral habitus. White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8836A6FFA4FFA5185D1378FA6FF8BA" blockId="3.[113,763,1828,1915]" lastBlockId="3.[810,1459,1828,1915]" pageId="3" pageNumber="4">without pit anteriorly; mesonotum continuous posterior to mesoscutellum (Fig. 4C). Mesopleuron areolate—punctate laterally, mesosubalar carina present, mesepisternal carina absent; with mid-coxa subdivided, with distinct lateral carina. Metanotum metascutellum not developed, median metanotal carina not observed, lateral metanotal carina present, reaching posterior margin of metanotum; metapleuron with few hairs, mostly glabrous (Fig. 4A); hind coxae not especially hairy laterally, with</paragraph>
|
||||
<paragraph id="8B8836A6FFA4FFA51B0613DEFA6FF878" blockId="3.[810,1459,1922,1977]" pageId="3" pageNumber="4">
|
||||
<pageTitle id="CBA8EEC1FFA4FFA51B0613DEFA6FF878" pageId="3" pageNumber="4">rounded medioventral margin and well-developed posterolateral carina; hind femur without ventral carina or denticles ventrally;</pageTitle>
|
||||
</paragraph>
|
||||
<caption id="DF48662EFFA3FFA218AD12DBFA77F932" pageId="4" pageNumber="5" startId="4.[129,192,1671,1695]" targetBox="[129,1473,144,1643]" targetPageId="4" targetType="figure">
|
||||
<caption id="DF48662EFFA3FFA218AD12DBFA77F932" ID-DOI="http://doi.org/10.5281/zenodo.14291329" ID-Zenodo-Dep="14291329" httpUri="https://zenodo.org/record/14291329/files/figure.png" pageId="4" pageNumber="5" startId="4.[129,192,1671,1695]" targetBox="[129,1473,144,1643]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA3FFA218AD12DBFA77F932" blockId="4.[128,1474,1671,1779]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B943EAB4FFA3FFA218AD12DBFF0AF95E" bold="true" box="[129,214,1671,1695]" pageId="4" pageNumber="5">Figure 2.</emphasis>
|
||||
†
|
||||
|
|
@ -176,22 +171,31 @@ gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, dorsal habitus.
|
|||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, C, photomicrographs. B, D, rendered from synchrotron scans. A, B, head, anterior view. C, head, lateroventral view. D, anterior view of tentorium. Red arrows = ventral transverse frontal carinae; white arrow = labrum; black arrows = coronal teeth; green arrows = maxillary palps; blue arrows = labial palp; ata = anterior tentorial arm; dta = dorsal tentorial arm; md = mandible; ltl = lateral tentorial lamella; mtl = median tentorial lamella; sc = scapus; teb = tentorial bridge.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8836A6FFA3FFA218AD137FFF1BF8BB" blockId="4.[129,777,1827,1914]" pageId="4" pageNumber="5">hind tibia with distinct pegs dorsally, longitudinal and ventral carinae not developed; apical hind tibial spurs short, of unequal length.</paragraph>
|
||||
<paragraph id="8B8836A6FFA3FFA218AD13C0FCD5F813" blockId="4.[129,777,1947,2003]" pageId="4" pageNumber="5">
|
||||
<paragraph id="8B8836A6FFA3FFA218AD13C0FBB8F8BB" blockId="4.[129,777,1947,2003]" lastBlockId="4.[825,1474,1827,1914]" pageId="4" pageNumber="5">
|
||||
<pageTitle id="CBA8EEC1FFA3FFA218AD13C0FCD5F813" pageId="4" pageNumber="5">
|
||||
<emphasis id="B943EAB4FFA3FFA218AD13C0FF19F872" box="[129,197,1948,1971]" italics="true" pageId="4" pageNumber="5">Wings:</emphasis>
|
||||
Forewing vein 2r-rs arise from approximately middle of pterostigma; vein 1-Rs well developed, elongate, discal cell
|
||||
</pageTitle>
|
||||
rectangular, proximal part not broader than distal part; vein cu-a inserts on Cu1 close to middle of cell M (Fig. 1A). Hindwing venation cannot be observed.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFA3FFA21B15137FFBB8F8BB" blockId="4.[825,1474,1827,1914]" pageId="4" pageNumber="5">rectangular, proximal part not broader than distal part; vein cu-a inserts on Cu1 close to middle of cell M (Fig. 1A). Hindwing venation cannot be observed.</paragraph>
|
||||
<paragraph id="8B8836A6FFA3FFA21B1513C7FA1EF813" blockId="4.[825,1474,1947,2002]" pageId="4" pageNumber="5">
|
||||
<paragraph id="8B8836A6FFA3FFAE1B1513C7FDEAFF06" blockId="4.[825,1474,1947,2002]" lastBlockId="8.[129,566,175,199]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
|
||||
<pageTitle id="CBA8EEC1FFA3FFA21B1513C7FA1EF813" pageId="4" pageNumber="5">
|
||||
<emphasis id="B943EAB4FFA3FFA21B1513C7FC43F872" box="[825,927,1947,1971]" italics="true" pageId="4" pageNumber="5">Abdomen:</emphasis>
|
||||
Dorsally covered by wings, hence obscuring structural detail via light microscopy; laterally with dense pilosity
|
||||
</pageTitle>
|
||||
of short hairs. Tergum 1 and 2 coarsely areolate (Fig. 4C), postspiracular and subspiracular carinae on tergum 1 absent, median carina on tergum 2 absent; tergum 3–8 finely punctate. Median apodemes observed on several abdominal sterna (Fig. 5D); sternum 7 externally with narrow median part delimited laterally and projecting posteriorly. Tergum 9 covered with short hairs, without longitudinal carinae, smooth areas and depressions; internally, prominent anterior flange and chordate apodemes present. Ovipositor apparatus apparently not fully preserved (Fig. 5A); second valvifer with traces of ventral
|
||||
<collectionCode id="ED26AE63FFA0FFA118F713C3FF37F876" box="[219,235,1951,1975]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="8">T</collectionCode>
|
||||
9-second valvifer muscles [muscle
|
||||
<quantity id="4CCF9B43FFA0FFA11A7A13C2FDA2F876" box="[598,638,1950,1975]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="8" unit="in" value="6.0">6 in</quantity>
|
||||
Vilhelmsen
|
||||
<emphasis id="B943EAB4FFA0FFA11B06137EFC86F8F8" box="[810,858,1825,1849]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
(2001)] observable, possible base of ovipositor shaft with remains of processus medianus present; only parts of internalized ovipositor loop traceable, ventral/distal part extends
|
||||
<pageTitle id="CBA8EEC1FFA0FFA11B061323FA6FF876" pageId="7" pageNumber="8">anteriorly at least to mesofurca, manner of coiling in prothorax not observed; tip of ovipositor slightly protruding from tip of</pageTitle>
|
||||
abdomen, third valvulae not visible. Composite tergum 10/ cercus triangular, not continuous medially.
|
||||
</paragraph>
|
||||
<caption id="DF48662EFFA2FFA3185D134BFE26F861" pageId="5" pageNumber="6" startId="5.[113,178,1815,1839]" targetBox="[114,1458,144,1788]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA2FFA3185D134BFAA6F845" blockId="5.[113,1454,1815,1924]" pageId="5" pageNumber="6">
|
||||
</subSubSection>
|
||||
<caption id="DF48662EFFA2FFA3185D134BFE26F861" ID-DOI="http://doi.org/10.5281/zenodo.14291331" ID-Zenodo-Dep="14291331" httpUri="https://zenodo.org/record/14291331/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1815,1839]" targetBox="[114,1458,144,1788]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA2FFA3185D134BFE26F861" blockId="5.[113,1454,1815,1924]" lastBlockId="5.[125,506,1928,1952]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B943EAB4FFA2FFA3185D134BFF15F8EE" bold="true" box="[113,201,1815,1839]" pageId="5" pageNumber="6">Figure 3.</emphasis>
|
||||
A, B, †
|
||||
<taxonomicName id="4C374D25FFA2FFA319201344FD55F8F1" authority="Jouault et al. 2021" authorityName="Jouault" authorityYear="2021" box="[268,649,1816,1840]" class="Insecta" family="Burmorussidae" genus="Cretorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vilhelmseni">
|
||||
|
|
@ -210,32 +214,23 @@ gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). Photomicrographs.
|
|||
<taxonomicName id="4C374D25FFA2FFA31BCA1330FB83F845" box="[998,1119,1900,1924]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B943EAB4FFA2FFA31BCA1330FB83F845" box="[998,1119,1900,1924]" italics="true" pageId="5" pageNumber="6">Kryptovelona</emphasis>
|
||||
</taxonomicName>
|
||||
in italics, with homologues in
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFA2FFA3185D13D4FFA1F861" blockId="5.[113,125,1928,1952]" box="[113,125,1928,1952]" pageId="5" pageNumber="6">†</paragraph>
|
||||
<paragraph id="8B8836A6FFA2FFA3185113D4FE26F861" blockId="5.[125,506,1928,1952]" box="[125,506,1928,1952]" pageId="5" pageNumber="6">
|
||||
<heading id="D0C081CAFFA2FFA3185113D4FE26F861" box="[125,506,1928,1952]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="6" reason="2">
|
||||
in italics, with homologues in †
|
||||
<taxonomicName id="4C374D25FFA2FFA3185113D4FF39F85E" box="[125,229,1928,1951]" class="Insecta" family="Burmorussidae" genus="Cretorussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B943EAB4FFA2FFA3185113D4FF39F85E" box="[125,229,1928,1951]" italics="true" pageId="5" pageNumber="6">Cretorussus</emphasis>
|
||||
</taxonomicName>
|
||||
in brackets); tb1 = fore tibia.
|
||||
</heading>
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF48662EFFA1FFA018AD12CEFE57F8DB" pageId="6" pageNumber="7" startId="6.[129,194,1682,1706]" targetBox="[129,1473,144,1654]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA1FFA018AD12CEFA97F96B" blockId="6.[129,1453,1682,1818]" box="[129,1355,1682,1706]" pageId="6" pageNumber="7">
|
||||
<caption id="DF48662EFFA1FFA018AD12CEFE57F8DB" ID-DOI="http://doi.org/10.5281/zenodo.14291333" ID-Zenodo-Dep="14291333" httpUri="https://zenodo.org/record/14291333/files/figure.png" pageId="6" pageNumber="7" startId="6.[129,194,1682,1706]" targetBox="[129,1473,144,1654]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA1FFA018AD12CEFE57F8DB" blockId="6.[129,1453,1682,1818]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B943EAB4FFA1FFA018AD12CEFF35F96B" bold="true" box="[129,233,1682,1706]" pageId="6" pageNumber="7">Figure 4. †</emphasis>
|
||||
<taxonomicName id="4C374D25FFA1FFA018C512CEFE34F96B" box="[233,488,1682,1706]" class="Insecta" family="Orussidae" genus="Kryptovelona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carstengroehni">
|
||||
<emphasis id="B943EAB4FFA1FFA018C512CEFE34F96B" box="[233,488,1682,1706]" italics="true" pageId="6" pageNumber="7">Kryptovelona carstengroehni</emphasis>
|
||||
</taxonomicName>
|
||||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C,
|
||||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFA1FFA018AD12F2FA44F93F" blockId="6.[129,1453,1682,1818]" pageId="6" pageNumber="7">rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum;</paragraph>
|
||||
<paragraph id="8B8836A6FFA1FFA018AE135EFE57F8DB" blockId="6.[129,1453,1682,1818]" box="[130,395,1794,1818]" pageId="6" pageNumber="7">T1 = 1st abdominal tergum.</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8836A6FFA1FFA018AD133EFCD5F8BB" blockId="6.[129,777,1890,1915]" box="[129,777,1890,1915]" pageId="6" pageNumber="7">of short hairs. Tergum 1 and 2 coarsely areolate (Fig. 4C),</paragraph>
|
||||
<paragraph id="8B8836A6FFA1FFA018AC13DDFA1EF878" blockId="6.[128,778,1921,1977]" lastBlockId="6.[825,1474,1921,1977]" pageId="6" pageNumber="7">postspiracular and subspiracular carinae on tergum 1 absent, median carina on tergum 2 absent; tergum 3–8 finely punctate. Median apodemes observed on several abdominal sterna (Fig. 5D); sternum 7 externally with narrow median part delimited laterally and projecting posteriorly. Tergum 9</paragraph>
|
||||
<caption id="DF48662EFFA0FFA1185D12D9FB2CF930" pageId="7" pageNumber="8" startId="7.[113,178,1669,1693]" targetBox="[114,1454,149,1639]" targetPageId="7" targetType="figure">
|
||||
<caption id="DF48662EFFA0FFA1185D12D9FB2CF930" ID-DOI="http://doi.org/10.5281/zenodo.14291335" ID-Zenodo-Dep="14291335" httpUri="https://zenodo.org/record/14291335/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,178,1669,1693]" targetBox="[114,1454,149,1639]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8B8836A6FFA0FFA1185D12D9FB2CF930" blockId="7.[113,1421,1669,1778]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B943EAB4FFA0FFA1185D12D9FF15F95F" bold="true" box="[113,201,1669,1694]" pageId="7" pageNumber="8">Figure 5.</emphasis>
|
||||
†
|
||||
|
|
@ -245,20 +240,6 @@ gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrogr
|
|||
gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, dorsal overview of extent of ovipositor inside the body; anterior to the top. B, profurca in posterior view. C, profurca in anterolateral view. D, posterior part of abdomen, dorsal internal view. All views rendered from synchrotron scans. Red arrows = ovipositor; black arrows = medain apodemes on abdominal sternum 6 and 7; green arrows = median groove on profurcal bridge; dl1 = prodiscrimenal lamella; pfb = profurcal bridge.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8836A6FFA0FFA1185D137DFD26F8B6" blockId="7.[113,762,1825,1912]" pageId="7" pageNumber="8">covered with short hairs, without longitudinal carinae, smooth areas and depressions; internally, prominent anterior flange and chordate apodemes present. Ovipositor apparatus appar-</paragraph>
|
||||
<paragraph id="8B8836A6FFA0FFA1185D1323FA6FF876" blockId="7.[113,165,1919,1975]" lastBlockId="7.[810,1459,1919,1975]" pageId="7" pageNumber="8">
|
||||
ently not fully preserved (Fig. 5A); second valvifer with traces of ventral
|
||||
<collectionCode id="ED26AE63FFA0FFA118F713C3FF37F876" box="[219,235,1951,1975]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="8">T</collectionCode>
|
||||
9-second valvifer muscles [muscle
|
||||
<quantity id="4CCF9B43FFA0FFA11A7A13C2FDA2F876" box="[598,638,1950,1975]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="8" unit="in" value="6.0">6 in</quantity>
|
||||
Vilhelmsen
|
||||
<emphasis id="B943EAB4FFA0FFA11B06137EFC86F8F8" box="[810,858,1825,1849]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
(2001)] observable, possible base of ovipositor shaft with remains of processus medianus present; only parts of internalized ovipositor loop traceable, ventral/distal part extends
|
||||
<pageTitle id="CBA8EEC1FFA0FFA11B061323FA6FF876" pageId="7" pageNumber="8">anteriorly at least to mesofurca, manner of coiling in prothorax not observed; tip of ovipositor slightly protruding from tip of</pageTitle>
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAFFFAE18AD14CCFCD5FF69" blockId="8.[129,777,144,168]" box="[129,777,144,168]" pageId="8" pageNumber="9">abdomen, third valvulae not visible. Composite tergum 10/</paragraph>
|
||||
<paragraph id="8B8836A6FFAFFFAE18AD14ECFDEAFF06" blockId="8.[129,566,175,199]" box="[129,566,175,199]" pageId="8" pageNumber="9">cercus triangular, not continuous medially.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32D652DFFAFFFAE18AD14B5FD72FEBF" pageId="8" pageNumber="9" type="etymology">
|
||||
<paragraph id="8B8836A6FFAFFFAE18AD14B5FD72FEBF" blockId="8.[129,778,233,382]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B943EAB4FFAFFFAE18AD14B5FF2CFEC0" box="[129,240,233,257]" italics="true" pageId="8" pageNumber="9">Etymology:</emphasis>
|
||||
|
|
@ -289,9 +270,9 @@ occur in Baltic amber, there are sufficient differences to justify recognizing t
|
|||
<emphasis id="B943EAB4FFAFFFAE1A4115A1FD2CFDD4" box="[621,752,509,533]" italics="true" pageId="8" pageNumber="9">Kryptovelona</emphasis>
|
||||
</taxonomicName>
|
||||
is placed closer to the clade comprising the majority of the extant taxa. The RoguePlots (Supporting Information,
|
||||
<figureCitation id="130C2A23FFAFFFAE1A441660FD6DFD95" box="[616,689,572,596]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" pageId="8" pageNumber="9">Figs S1</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1A441660FD6DFD95" box="[616,689,572,596]" captionStart="Figure 1" captionStartId="3.[114,179,1713,1737]" captionTargetBox="[114,1458,144,1686]" captionTargetId="figure-88@3.[114,1458,144,1686]" captionTargetPageId="3" captionText="Figure 1. †Kryptovelona carstengroehni gen* et sp* nov*, female holotype (GPIH 5078, CCGG nº 3811)* A, dorsal habitus* B, lateral habitus* C, ventral habitus* White arrow = vein 1-Rs; red arrow = vein cu-a; black arrows = coronal teeth; cx3 = hind coxa; dc = discal cell; fe3 = hind femur; n1 = pronotum; n2 = mesoscutum; pl1 = propleuron; pl2 = mesopleuron; pt = pterostigma; sc2 = mesoscutellum; tb1 = fore tibia*" figureDoi="http://doi.org/10.5281/zenodo.14291327" httpUri="https://zenodo.org/record/14291327/files/figure.png" pageId="8" pageNumber="9">Figs S1</figureCitation>
|
||||
, S
|
||||
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<figureCitation id="130C2A23FFAFFFAE1AE41660FD09FD95" box="[712,725,572,596]" captionStart="Figure 4" captionStartId="6.[129,194,1682,1706]" captionTargetBox="[129,1473,144,1654]" captionTargetId="figure-64@6.[129,1473,144,1654]" captionTargetPageId="6" captionText="Figure 4. †Kryptovelona carstengroehni gen. et sp. nov., female holotype (GPIH 5078, CCGG nº 3811). A, B, photomicrographs. C, rendered from synchrotron scans. A, thorax, lateral view. B, thorax, lateral view. C, thorax and anterior abdomen, dorsolateral view. White arrow = metapleuron; red arrow = spiracle on 1st abdominal tergum; green arrows = transscutal articulation; blue arrow = parapside; yellow arrow = scutellar arm; n1 = pronotum; n2 = mesonotum; n3 = metanotum; pl1 = propleuron; pl2 = mesopleuron; sc2 = mesoscutellum; T1 = 1st abdominal tergum." figureDoi="http://doi.org/10.5281/zenodo.14291333" httpUri="https://zenodo.org/record/14291333/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
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) flag a possible †
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<figureCitation id="130C2A23FFAFFFAE19B410E1FE2EFB16" box="[408,498,1213,1239]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStart-3="Figure 9" captionStartId-0="9.[113,178,1265,1289]" captionStartId-1="10.[129,194,1523,1547]" captionStartId-2="11.[113,178,1631,1655]" captionStartId-3="12.[129,194,1517,1541]" captionTargetBox-0="[114,1458,144,1237]" captionTargetBox-1="[129,1473,144,1495]" captionTargetBox-2="[114,1438,144,1603]" captionTargetBox-3="[132,1466,152,1489]" captionTargetId-0="figure-289@9.[114,1458,144,1237]" captionTargetId-1="figure-211@10.[129,1473,144,1495]" captionTargetId-2="figure-112@11.[114,1458,144,1603]" captionTargetId-3="figure-149@12.[129,1473,144,1489]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="Figure 6. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). Dorsal habitus (A). Photomicrograph. B, rendered from synchrotron scans (head and wings removed in reconstruction). White arrow = vein 1-Rs; red arrow = vein cu-a; green arrow = median metanotal carina; blue arrow = lateral metanotal carina; yellow arrow = mesoscutellar arm; dc = discal cell; n1 = pronotum; n2 = mesoscutum; pt = pterostigma; sc2 = mesoscutellum; t[n] = abdominal tergum [n]." captionText-1="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." captionText-2="Figure 8. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* A, photomicrograph* B–F, rendered from synchrotron scans* A, head, anterior* B, head, posterior* C, head, lateral, showing position of tentorium* D, tentorium in lateral view* E, head, dorsal, showing position of tentorium* F, tentorium in dorsal view* Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge*" captionText-3="Figure 9. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* Ovipositor; Rendered from synchrotron scans* A, habitus, lateral with position of views indicated* B, profurca, posterior view* C, profurca and ovipositor loop, posterolateral view* D, complete ovipositor loop, lateral view* Black arrow = median groove on profurcal bridge; green arrows = dorsal branch of ovipositor loop; red arrows = ventral branch of ovipositor loop; blue arrows = profurca; fua = profurcal arm; fub = profurcal bridge; S1 = prosternum; 3vv = 3rd valvula*" pageId="8" pageNumber="9">Figs 6–9</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE19B410E1FE2EFB16" box="[408,498,1213,1239]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStart-3="Figure 9" captionStartId-0="9.[113,178,1265,1289]" captionStartId-1="10.[129,194,1523,1547]" captionStartId-2="11.[113,178,1631,1655]" captionStartId-3="12.[129,194,1517,1541]" captionTargetBox-0="[114,1458,144,1237]" captionTargetBox-1="[129,1473,144,1495]" captionTargetBox-2="[114,1438,144,1603]" captionTargetBox-3="[132,1466,152,1489]" captionTargetId-0="figure-289@9.[114,1458,144,1237]" captionTargetId-1="figure-211@10.[129,1473,144,1495]" captionTargetId-2="figure-112@11.[114,1458,144,1603]" captionTargetId-3="figure-149@12.[129,1473,144,1489]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="Figure 6. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). Dorsal habitus (A). Photomicrograph. B, rendered from synchrotron scans (head and wings removed in reconstruction). White arrow = vein 1-Rs; red arrow = vein cu-a; green arrow = median metanotal carina; blue arrow = lateral metanotal carina; yellow arrow = mesoscutellar arm; dc = discal cell; n1 = pronotum; n2 = mesoscutum; pt = pterostigma; sc2 = mesoscutellum; t[n] = abdominal tergum [n]." captionText-1="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." captionText-2="Figure 8. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* A, photomicrograph* B–F, rendered from synchrotron scans* A, head, anterior* B, head, posterior* C, head, lateral, showing position of tentorium* D, tentorium in lateral view* E, head, dorsal, showing position of tentorium* F, tentorium in dorsal view* Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge*" captionText-3="Figure 9. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* Ovipositor; Rendered from synchrotron scans* A, habitus, lateral with position of views indicated* B, profurca, posterior view* C, profurca and ovipositor loop, posterolateral view* D, complete ovipositor loop, lateral view* Black arrow = median groove on profurcal bridge; green arrows = dorsal branch of ovipositor loop; red arrows = ventral branch of ovipositor loop; blue arrows = profurca; fua = profurcal arm; fub = profurcal bridge; S1 = prosternum; 3vv = 3rd valvula*" figureDoi-0="http://doi.org/10.5281/zenodo.14291337" figureDoi-1="http://doi.org/10.5281/zenodo.14291339" figureDoi-2="http://doi.org/10.5281/zenodo.14291341" figureDoi-3="http://doi.org/10.5281/zenodo.14291343" httpUri-0="https://zenodo.org/record/14291337/files/figure.png" httpUri-1="https://zenodo.org/record/14291339/files/figure.png" httpUri-2="https://zenodo.org/record/14291341/files/figure.png" httpUri-3="https://zenodo.org/record/14291343/files/figure.png" pageId="8" pageNumber="9">Figs 6–9</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</heading>
|
||||
|
|
@ -118,38 +119,36 @@ preserved in a piece of Baltic amber dated as
|
|||
<paragraph id="8B8836A6FFAFFFAE18AD1193FD75F8E1" blockId="8.[129,778,1487,1824]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B943EAB4FFAFFFAE18AD1193FF3BFA27" box="[129,231,1487,1510]" italics="true" pageId="8" pageNumber="9">Diagnosis:</emphasis>
|
||||
Ventral coronal teeth present, connected by faint U-shaped carina (Fig. 8A). Ventral transverse longitudinal carina fully developed, with median notch (
|
||||
<figureCitation id="130C2A23FFAFFFAE1A5D1251FD1AF9E7" box="[625,710,1549,1574]" captionStart="Figure 7" captionStartId="10.[129,194,1523,1547]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-211@10.[129,1473,144,1495]" captionTargetPageId="10" captionText="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." pageId="8" pageNumber="9">Figs 7A</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1A5D1251FD1AF9E7" box="[625,710,1549,1574]" captionStart="Figure 7" captionStartId="10.[129,194,1523,1547]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-211@10.[129,1473,144,1495]" captionTargetPageId="10" captionText="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." figureDoi="http://doi.org/10.5281/zenodo.14291339" httpUri="https://zenodo.org/record/14291339/files/figure.png" pageId="8" pageNumber="9">Figs 7A</figureCitation>
|
||||
,
|
||||
<figureCitation id="130C2A23FFAFFFAE1AF71251FD25F9E4" box="[731,761,1549,1573]" captionStart="Figure 8" captionStartId="11.[113,178,1631,1655]" captionTargetBox="[114,1438,144,1603]" captionTargetId="figure-112@11.[114,1458,144,1603]" captionTargetPageId="11" captionText="Figure 8. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* A, photomicrograph* B–F, rendered from synchrotron scans* A, head, anterior* B, head, posterior* C, head, lateral, showing position of tentorium* D, tentorium in lateral view* E, head, dorsal, showing position of tentorium* F, tentorium in dorsal view* Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge*" pageId="8" pageNumber="9">8A</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1AF71251FD25F9E4" box="[731,761,1549,1573]" captionStart="Figure 8" captionStartId="11.[113,178,1631,1655]" captionTargetBox="[114,1438,144,1603]" captionTargetId="figure-112@11.[114,1458,144,1603]" captionTargetPageId="11" captionText="Figure 8. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* A, photomicrograph* B–F, rendered from synchrotron scans* A, head, anterior* B, head, posterior* C, head, lateral, showing position of tentorium* D, tentorium in lateral view* E, head, dorsal, showing position of tentorium* F, tentorium in dorsal view* Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge*" figureDoi="http://doi.org/10.5281/zenodo.14291341" httpUri="https://zenodo.org/record/14291341/files/figure.png" pageId="8" pageNumber="9">8A</figureCitation>
|
||||
). Antennomeres 4 and 5 combined approximately the same length as antennomere 6, maxillary palp elongate, five-segmented (Fig. 8B). Fore coxa expanded medially. Median mesoscutal sulcus and notauli at most weakly developed, mesoscutellum triangular, acute, raised compared to surrounding sclerites (Fig. 6A, B). Forewing vein 1r arises from approximately middle of pterostigma; vein 1r-Rs well developed, elongate, discal cell trapezoid, proximal part broader than distal part (Fig. 6A).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32D652DFFAFFFAF18AD131EFC5AF8FD" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
|
||||
<paragraph id="8B8836A6FFAFFFAE18AD131EFCD5F876" blockId="8.[129,778,1857,1913]" lastBlockId="8.[129,777,1920,1975]" pageId="8" pageNumber="9">
|
||||
<paragraph id="8B8836A6FFAFFFAE18AD131EFB57FF06" blockId="8.[129,778,1857,1913]" lastBlockId="8.[825,1163,175,199]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B943EAB4FFAFFFAE18AD131EFF2BF898" box="[129,247,1858,1881]" italics="true" pageId="8" pageNumber="9">Description:</emphasis>
|
||||
Female (
|
||||
<figureCitation id="130C2A23FFAFFFAE1971131DFE40F898" box="[349,412,1857,1881]" captionStart="Figure 6" captionStartId="9.[113,178,1265,1289]" captionTargetBox="[114,1458,144,1237]" captionTargetId="figure-289@9.[114,1458,144,1237]" captionTargetPageId="9" captionText="Figure 6. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). Dorsal habitus (A). Photomicrograph. B, rendered from synchrotron scans (head and wings removed in reconstruction). White arrow = vein 1-Rs; red arrow = vein cu-a; green arrow = median metanotal carina; blue arrow = lateral metanotal carina; yellow arrow = mesoscutellar arm; dc = discal cell; n1 = pronotum; n2 = mesoscutum; pt = pterostigma; sc2 = mesoscutellum; t[n] = abdominal tergum [n]." pageId="8" pageNumber="9">Figs 6</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1971131DFE40F898" box="[349,412,1857,1881]" captionStart="Figure 6" captionStartId="9.[113,178,1265,1289]" captionTargetBox="[114,1458,144,1237]" captionTargetId="figure-289@9.[114,1458,144,1237]" captionTargetPageId="9" captionText="Figure 6. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). Dorsal habitus (A). Photomicrograph. B, rendered from synchrotron scans (head and wings removed in reconstruction). White arrow = vein 1-Rs; red arrow = vein cu-a; green arrow = median metanotal carina; blue arrow = lateral metanotal carina; yellow arrow = mesoscutellar arm; dc = discal cell; n1 = pronotum; n2 = mesoscutum; pt = pterostigma; sc2 = mesoscutellum; t[n] = abdominal tergum [n]." figureDoi="http://doi.org/10.5281/zenodo.14291337" httpUri="https://zenodo.org/record/14291337/files/figure.png" pageId="8" pageNumber="9">Figs 6</figureCitation>
|
||||
,
|
||||
<figureCitation id="130C2A23FFAFFFAE1986131EFE6BF898" box="[426,439,1858,1881]" captionStart="Figure 7" captionStartId="10.[129,194,1523,1547]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-211@10.[129,1473,144,1495]" captionTargetPageId="10" captionText="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." pageId="8" pageNumber="9">7</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1986131EFE6BF898" box="[426,439,1858,1881]" captionStart="Figure 7" captionStartId="10.[129,194,1523,1547]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-211@10.[129,1473,144,1495]" captionTargetPageId="10" captionText="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." figureDoi="http://doi.org/10.5281/zenodo.14291339" httpUri="https://zenodo.org/record/14291339/files/figure.png" pageId="8" pageNumber="9">7</figureCitation>
|
||||
). Body length 3.
|
||||
<quantity id="4CCF9B43FFAFFFAE1A41131DFD7AF898" box="[621,678,1857,1881]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="mm" value="8.0">8 mm</quantity>
|
||||
. Original body coloration not observable. Specimen very well preserved and well observable from most angles except for distal margin of right forewing, which is obscured by narrow cracks extending
|
||||
. Original body coloration not observable. Specimen very well preserved and well observable from most angles except for distal margin of right forewing, which is obscured by narrow cracks extending from the wings. Forewings folded above abdomen, obscuring it and the hindwings in dorsal view.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAFFFAE1B1514CCFB57FF06" blockId="8.[825,1474,144,168]" lastBlockId="8.[825,1163,175,199]" pageId="8" pageNumber="9">from the wings. Forewings folded above abdomen, obscuring it and the hindwings in dorsal view.</paragraph>
|
||||
<paragraph id="8B8836A6FFAFFFAE1B1514B5FA58FB4C" blockId="8.[825,1474,233,1165]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B943EAB4FFAFFFAE1B1514B5FCAAFEC0" box="[825,886,233,257]" italics="true" pageId="8" pageNumber="9">Head:</emphasis>
|
||||
Ocellar corona narrow, distance between median ocellus and lateral coronal tooth less than ocellar width; ventral coronal teeth present, situated slightly medial to median coronal teeth; faint median frontal carinae extend ventrally from ventral coronal teeth and converge medially, forming U-shaped loop (Fig. 8A). Dorsal transverse and longitudinal frontal carinae absent. Ventral transverse frontal carina continuous medially and with slight incision, covering toruli in anterior view (Fig. 8A); carina laterally continuous with lateral carina of subantennal groove (Fig. 7). Frons and vertex areolate (Fig. 8A), genae with scattered punctures; frons, vertex, and genae without conspicuous pilosity. Postocular carina well developed for most of height of eye, occipital carinae present (Fig. 8B), continuous with distinct lateral carina of subantennal groove ventrally. Tentorium with median lamellae on anterior arm broadening posteriorly (Fig. 8E, F), lateral lamellae hardly developed; tentorial bridge narrow, arched, with distinct anterior process (Fig. 8F); dorsal tentorial arms arise posteriorly on anterior arms, short, apparently do not reach head capsule (Fig. 8C, D). Antennae with 10 antennomeres, covered with short hairs; scapus short, subcylindrical (Fig. 8A); combined length of antennomeres 4 and 5 approximately equal to antennomere 6; antennomere 9 longer than antennomeres 7 and 8 combined, swollen subapically, without carina laterally; antennomere 10 short, cylindrical, inserted subapically on antennomere 9 (Fig. 7). Labrum visible between mandibles (
|
||||
<figureCitation id="130C2A23FFAFFFAE1DB41784FC8BFBD1" captionStart="Figure 7" captionStartId="10.[129,194,1523,1547]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-211@10.[129,1473,144,1495]" captionTargetPageId="10" captionText="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." pageId="8" pageNumber="9">Figs 7A</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1DB41784FC8BFBD1" captionStart="Figure 7" captionStartId="10.[129,194,1523,1547]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-211@10.[129,1473,144,1495]" captionTargetPageId="10" captionText="Figure 7. †Orussus juttagroehnae sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia." figureDoi="http://doi.org/10.5281/zenodo.14291339" httpUri="https://zenodo.org/record/14291339/files/figure.png" pageId="8" pageNumber="9">Figs 7A</figureCitation>
|
||||
,
|
||||
<figureCitation id="130C2A23FFAFFFAE1B4B17A4FC58FBD1" box="[871,900,1016,1040]" captionStart="Figure 8" captionStartId="11.[113,178,1631,1655]" captionTargetBox="[114,1438,144,1603]" captionTargetId="figure-112@11.[114,1458,144,1603]" captionTargetPageId="11" captionText="Figure 8. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* A, photomicrograph* B–F, rendered from synchrotron scans* A, head, anterior* B, head, posterior* C, head, lateral, showing position of tentorium* D, tentorium in lateral view* E, head, dorsal, showing position of tentorium* F, tentorium in dorsal view* Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge*" pageId="8" pageNumber="9">8A</figureCitation>
|
||||
<figureCitation id="130C2A23FFAFFFAE1B4B17A4FC58FBD1" box="[871,900,1016,1040]" captionStart="Figure 8" captionStartId="11.[113,178,1631,1655]" captionTargetBox="[114,1438,144,1603]" captionTargetId="figure-112@11.[114,1458,144,1603]" captionTargetPageId="11" captionText="Figure 8. Orussus jutagroehnae sp* nov*, female holotype (GPIH 5081, CCGG nº 1288)* A, photomicrograph* B–F, rendered from synchrotron scans* A, head, anterior* B, head, posterior* C, head, lateral, showing position of tentorium* D, tentorium in lateral view* E, head, dorsal, showing position of tentorium* F, tentorium in dorsal view* Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge*" figureDoi="http://doi.org/10.5281/zenodo.14291341" httpUri="https://zenodo.org/record/14291341/files/figure.png" pageId="8" pageNumber="9">8A</figureCitation>
|
||||
), small, globose, with distinct ventrally directed hairs. Mandibles chisel-like, no teeth developed, with conspicuous distally pointing hairs laterally. Maxillary palps elongate, five-segmented, labial palps short, three-segmented (Fig. 8B); glossa of labium folded longitudinally, concave in posterior view.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAFFFAE1B1510F3FA1EF8B6" blockId="8.[825,1474,1198,1911]" pageId="8" pageNumber="9">
|
||||
<paragraph id="8B8836A6FFAFFFAF1B1510F3FF64F9A0" blockId="8.[825,1474,1198,1911]" lastBlockId="9.[113,762,1421,1633]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||||
<emphasis id="B943EAB4FFAFFFAE1B1510F3FC54FB07" box="[825,904,1199,1222]" italics="true" pageId="8" pageNumber="9">Thorax:</emphasis>
|
||||
Pronotum dorsally short, of equal length throughout, deeply incurved posteriorly (Fig. 6B), laterally evenly punctured. Prosternum mostly concealed by propleura, diamondshaped, lateral margins diverging anteriorly, discrimen and discrimenal lamella distinct; profurcal arms extend obliquely anteriorly from posterior corners of prosternum, profurcal bridge low and straight, with distinct median groove accommodating lower branch of curved ovipositor (Fig. 9B), bridge not bent posteriorly. Fore coxa expanded medially; ventral carina on fore femur not observed; fore tibia swollen distally and subdivided by groove (Fig. 7); fore tarsus with three tarsomeres, basitarsus elongate and terminating in spur overlapping tarsomere 2. Mesoscutum finely punctate, not densely covered with hairs, median mesoscutal sulcus absent, notauli only developed as shallow grooves not reaching transscutal articulation posteriorly, parapsides present laterally and transscutal articulation posteriorly (Fig. 6); axillar flanges well developed; scutoscutellar sulcus well developed; mesoscutellum areolate-punctate, rounded triangular posteriorly (Fig. 6), raised and with smooth carinate lateral margins; mesoscutellar arms curved posteriorly, area anterior to arms raised to same level as arms; mesonotum continuous posterior to mesoscutellum. Mesopleuron areolate–punctate lateroventrally, mesosubalar carina present,
|
||||
Pronotum dorsally short, of equal length throughout, deeply incurved posteriorly (Fig. 6B), laterally evenly punctured. Prosternum mostly concealed by propleura, diamondshaped, lateral margins diverging anteriorly, discrimen and discrimenal lamella distinct; profurcal arms extend obliquely anteriorly from posterior corners of prosternum, profurcal bridge low and straight, with distinct median groove accommodating lower branch of curved ovipositor (Fig. 9B), bridge not bent posteriorly. Fore coxa expanded medially; ventral carina on fore femur not observed; fore tibia swollen distally and subdivided by groove (Fig. 7); fore tarsus with three tarsomeres, basitarsus elongate and terminating in spur overlapping tarsomere 2. Mesoscutum finely punctate, not densely covered with hairs, median mesoscutal sulcus absent, notauli only developed as shallow grooves not reaching transscutal articulation posteriorly, parapsides present laterally and transscutal articulation posteriorly (Fig. 6); axillar flanges well developed; scutoscutellar sulcus well developed; mesoscutellum areolate-punctate, rounded triangular posteriorly (Fig. 6), raised and with smooth carinate lateral margins; mesoscutellar arms curved posteriorly, area anterior to arms raised to same level as arms; mesonotum continuous posterior to mesoscutellum. Mesopleuron areolate–punctate lateroventrally, mesosubalar carina present, mesepisternal carina absent; mid-coxa subdivided, with distinct lateral carina. Metanotum with median and lateral metanotal carinae well developed (Fig. 6B); metapleuron areolate across most of its surface; hind coxae not especially hairy laterally, with angled medioventral margin and well-developed posterolateral carina; ventral part of hind femur concealed; hind tibia with distinct pegs dorsally, longitudinal and ventral carinae not observed; apical hind tibial spurs short, of approximately equal length.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAFFFAE1B151323FA1EF877" blockId="8.[825,1474,1919,1974]" pageId="8" pageNumber="9">mesepisternal carina absent; mid-coxa subdivided, with distinct lateral carina. Metanotum with median and lateral metanotal</paragraph>
|
||||
<caption id="DF48662EFFAEFFAF185D10ADFC63FA9C" pageId="9" pageNumber="10" startId="9.[113,178,1265,1289]" targetBox="[114,1458,144,1237]" targetPageId="9" targetType="figure">
|
||||
<caption id="DF48662EFFAEFFAF185D10ADFC63FA9C" ID-DOI="http://doi.org/10.5281/zenodo.14291337" ID-Zenodo-Dep="14291337" httpUri="https://zenodo.org/record/14291337/files/figure.png" pageId="9" pageNumber="10" startId="9.[113,178,1265,1289]" targetBox="[114,1458,144,1237]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8B8836A6FFAEFFAF185D10ADFC63FA9C" blockId="9.[113,1440,1265,1373]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B943EAB4FFAEFFAF185D10ADFF15FAC8" bold="true" box="[113,201,1265,1289]" pageId="9" pageNumber="10">Figure 6.</emphasis>
|
||||
†
|
||||
|
|
@ -159,25 +158,22 @@ Pronotum dorsally short, of equal length throughout, deeply incurved posteriorly
|
|||
sp. nov., female holotype (GPIH 5081, CCGG nº 1288). Dorsal habitus (A). Photomicrograph. B, rendered from synchrotron scans (head and wings removed in reconstruction). White arrow = vein 1-Rs; red arrow = vein cu-a; green arrow = median metanotal carina; blue arrow = lateral metanotal carina; yellow arrow = mesoscutellar arm; dc = discal cell; n1 = pronotum; n2 = mesoscutum; pt = pterostigma; sc2 = mesoscutellum; t[n] = abdominal tergum [n].
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8836A6FFAEFFAF185D11D1FF64F9A0" blockId="9.[113,762,1421,1633]" pageId="9" pageNumber="10">carinae well developed (Fig. 6B); metapleuron areolate across most of its surface; hind coxae not especially hairy laterally, with angled medioventral margin and well-developed posterolateral carina; ventral part of hind femur concealed; hind tibia with distinct pegs dorsally, longitudinal and ventral carinae not observed; apical hind tibial spurs short, of approximately equal length.</paragraph>
|
||||
<paragraph id="8B8836A6FFAEFFAF185D12D5FEA4F8FC" blockId="9.[113,763,1672,1853]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B943EAB4FFAEFFAF185D12D5FF6AF961" box="[113,182,1673,1696]" italics="true" pageId="9" pageNumber="10">Wings:</emphasis>
|
||||
Forewing apparently without marked bands of infuscation; vein 2r-rs arise from approximately middle of pterostigma; vein 1-Rs well developed, elongate, discal cell trapezoid, proximal part higher than distal part (Fig. 6A); vein cu-a inserts on Cu1 only slightly distally to vein M. Hindwing venation cannot be observed.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAEFFAF185D1302FD26F854" blockId="9.[113,762,1886,1910]" lastBlockId="9.[113,762,1917,1973]" pageId="9" pageNumber="10">
|
||||
<paragraph id="8B8836A6FFAEFFAF185D1302FC5AF8FD" blockId="9.[113,762,1886,1910]" lastBlockId="9.[810,1459,1421,1852]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B943EAB4FFAEFFAF185D1302FF0AF8B7" box="[113,214,1886,1910]" italics="true" pageId="9" pageNumber="10">Abdomen:</emphasis>
|
||||
Dorsally covered by wings, hence obscuring structural detail; laterally not particularly hairy, sculpture finely im-
|
||||
Dorsally covered by wings, hence obscuring structural detail; laterally not particularly hairy, sculpture finely imbricate. Tergum 1 with weakly developed postspiracular carina, subspiracular carinae absent. Tergum 2 without median carina (Fig. 6B), with rectangular smooth area laterally abutting antecosta. Median apodemes observed on several abdominal sterna; sternum 7 externally with narrow median part delimited laterally and projecting posteriorly (Fig. 7A). Tergum 9 punctate, with low, longitudinal carinae delimiting concave depression medially, smooth areas and depressions absent. Tip of ovipositor slightly protruding from tip of abdomen, third valvulae exposed ventrally (Fig. 7A). Internally, complete ovipositor loop can be traced (Fig. 9D), loop simple, hairpin-like, anterior part of ventral branch runs through dorsal longitudinal groove of profurcal bridge (Fig. 9C); traces of ovipositor sack containing loop observed. Composite tergum 10/cercus triangular, not continuous medially.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAEFFAF185D13C1FC5AF8FD" blockId="9.[113,762,1917,1973]" lastBlockId="9.[810,1459,1421,1852]" pageId="9" pageNumber="10">bricate. Tergum 1 with weakly developed postspiracular carina, subspiracular carinae absent. Tergum 2 without median carina (Fig. 6B), with rectangular smooth area laterally abutting antecosta. Median apodemes observed on several abdominal sterna; sternum 7 externally with narrow median part delimited laterally and projecting posteriorly (Fig. 7A). Tergum 9 punctate, with low, longitudinal carinae delimiting concave depression medially, smooth areas and depressions absent. Tip of ovipositor slightly protruding from tip of abdomen, third valvulae exposed ventrally (Fig. 7A). Internally, complete ovipositor loop can be traced (Fig. 9D), loop simple, hairpin-like, anterior part of ventral branch runs through dorsal longitudinal groove of profurcal bridge (Fig. 9C); traces of ovipositor sack containing loop observed. Composite tergum 10/cercus triangular, not continuous medially.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32D652DFFAEFFAF1B061301FCA9F875" pageId="9" pageNumber="10" type="etymology">
|
||||
<paragraph id="8B8836A6FFAEFFAF1B061301FA6FF8B4" blockId="9.[810,1459,1885,1909]" box="[810,1459,1885,1909]" pageId="9" pageNumber="10">
|
||||
<paragraph id="8B8836A6FFAEFFAF1B061301FCA9F875" blockId="9.[810,1459,1885,1909]" lastBlockId="9.[810,1459,1916,1972]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B943EAB4FFAEFFAF1B061301FC45F8B4" box="[810,921,1885,1909]" italics="true" pageId="9" pageNumber="10">Etymology:</emphasis>
|
||||
The species epithet honours Mrs Jutta Gröhn, wife of
|
||||
The species epithet honours Mrs Jutta Gröhn, wife of Carsten Gröhn and longterm supporter of his passion for amber insects.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFAEFFAF1B061321FCA9F875" blockId="9.[810,1459,1916,1972]" pageId="9" pageNumber="10">Carsten Gröhn and longterm supporter of his passion for amber insects.</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF48662EFFADFFAC18AD11AFFE79F99E" pageId="10" pageNumber="11" startId="10.[129,194,1523,1547]" targetBox="[129,1473,144,1495]" targetPageId="10" targetType="figure">
|
||||
<caption id="DF48662EFFADFFAC18AD11AFFE79F99E" ID-DOI="http://doi.org/10.5281/zenodo.14291339" ID-Zenodo-Dep="14291339" httpUri="https://zenodo.org/record/14291339/files/figure.png" pageId="10" pageNumber="11" startId="10.[129,194,1523,1547]" targetBox="[129,1473,144,1495]" targetPageId="10" targetType="figure">
|
||||
<paragraph id="8B8836A6FFADFFAC18AD11AFFE79F99E" blockId="10.[129,1476,1523,1632]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B943EAB4FFADFFAC18AD11AFFF04F9CA" bold="true" box="[129,216,1523,1547]" pageId="10" pageNumber="11">Figure 7.</emphasis>
|
||||
†
|
||||
|
|
@ -187,8 +183,8 @@ The species epithet honours Mrs Jutta Gröhn, wife of
|
|||
sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, ventral habitus. B, lateral habitus. Green arrows = 3rd valvulae; white arrow = labrum; red arrows = ventral frontal carina; blue arrows = lateral carina of subantennal groove; black arrows = coronal teeth; cx3 = hind coxa; fe3 = hind femur; n1 = pronotum; pl1 = propleuron; pl2 = mesopleuron; s7 = abdominal sternum 7; t9 = tergum 9; tb1 = fore tibia.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C32D652DFFADFFAA18AD12CCFCD5F8BD" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="discussion">
|
||||
<paragraph id="8B8836A6FFADFFAC18AD12CCFCD5F800" blockId="10.[129,778,1679,1923]" lastBlockId="10.[129,778,1930,1985]" pageId="10" pageNumber="11">
|
||||
<subSubSection id="C32D652DFFADFFAA18AD12CCFD8AF87A" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="discussion">
|
||||
<paragraph id="8B8836A6FFADFFAC18AD12CCFADFF800" blockId="10.[129,778,1679,1923]" lastBlockId="10.[825,1474,1930,1986]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B943EAB4FFADFFAC18AD12CCFF2DF966" box="[129,241,1680,1703]" italics="true" pageId="10" pageNumber="11">Comments:</emphasis>
|
||||
†
|
||||
<taxonomicName id="4C374D25FFADFFAC192612CCFE35F966" box="[266,489,1679,1703]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
|
|
@ -226,8 +222,6 @@ could not be confirmed in †
|
|||
<emphasis id="B943EAB4FFADFFAC198413D6FD97F863" box="[424,587,1930,1954]" italics="true" pageId="10" pageNumber="11">O. juttagroehnae</emphasis>
|
||||
is the presence of white spots on at least some legs (Vilhelmsen 2003) and also
|
||||
</pageTitle>
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFADFFAC1B1512CCFA1EF843" blockId="10.[825,1475,1679,1923]" pageId="10" pageNumber="11">
|
||||
on other body parts (e.g. face, antennae, pronotum, abdominal tip) in some species; given that the natural colour of the fossil specimen is not observable, white spots might well be present. †
|
||||
<taxonomicName id="4C374D25FFADFFAC1B6A12B2FBFCF8C4" box="[838,1056,1773,1797]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
<emphasis id="B943EAB4FFADFFAC1B6A12B2FBFCF8C4" box="[838,1056,1773,1797]" italics="true" pageId="10" pageNumber="11">Orussus juttagroehnae</emphasis>
|
||||
|
|
@ -241,16 +235,14 @@ spp
|
|||
<taxonomicName id="4C374D25FFADFFAC1BDC1317FB15F8A2" box="[1008,1225,1867,1891]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
<emphasis id="B943EAB4FFADFFAC1BDC1317FB15F8A2" box="[1008,1225,1867,1891]" italics="true" pageId="10" pageNumber="11">Orussus juttagroehnae</emphasis>
|
||||
</taxonomicName>
|
||||
is the first fossil species described from the genus, and the oldest member of an extant
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFADFFAC1B1513D6FADFF800" blockId="10.[825,1474,1930,1986]" pageId="10" pageNumber="11">
|
||||
is the first fossil species described from the genus, and the oldest member of an extant
|
||||
<pageTitle id="CBA8EEC1FFADFFAC1B1513D6FADFF800" pageId="10" pageNumber="11">
|
||||
orussid genus recorded so far; †
|
||||
<emphasis id="B943EAB4FFADFFAC1CA813D6FA9AF860" box="[1156,1350,1930,1954]" italics="true" pageId="10" pageNumber="11">Ophrynopus peritus</emphasis>
|
||||
Engel 2008 is from the much younger Dominican amber.
|
||||
</pageTitle>
|
||||
</paragraph>
|
||||
<caption id="DF48662EFFACFFAD185D1203FAF2F8C2" pageId="11" pageNumber="12" startId="11.[113,178,1631,1655]" targetBox="[114,1438,144,1603]" targetPageId="11" targetType="figure">
|
||||
<caption id="DF48662EFFACFFAD185D1203FAF2F8C2" ID-DOI="http://doi.org/10.5281/zenodo.14291341" ID-Zenodo-Dep="14291341" httpUri="https://zenodo.org/record/14291341/files/figure.png" pageId="11" pageNumber="12" startId="11.[113,178,1631,1655]" targetBox="[114,1438,144,1603]" targetPageId="11" targetType="figure">
|
||||
<paragraph id="8B8836A6FFACFFAD185D1203FAF2F8C2" blockId="11.[113,1456,1631,1795]" pageId="11" pageNumber="12">
|
||||
<emphasis id="B943EAB4FFACFFAD185D1203FF15F9B6" bold="true" box="[113,201,1631,1655]" pageId="11" pageNumber="12">Figure 8.</emphasis>
|
||||
<taxonomicName id="4C374D25FFACFFAD18E2123CFE49F9B6" box="[206,405,1631,1655]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
|
|
@ -259,8 +251,7 @@ Engel 2008 is from the much younger Dominican amber.
|
|||
sp. nov., female holotype (GPIH 5081, CCGG nº 1288). A, photomicrograph. B–F, rendered from synchrotron scans. A, head, anterior. B, head, posterior. C, head, lateral, showing position of tentorium. D, tentorium in lateral view. E, head, dorsal, showing position of tentorium. F, tentorium in dorsal view. Black arrows = coronal teeth (ventral tooth with white head); white arrows = median longitudinal frontal carina; blue arrow = lateral carina of subantennal groove; red arrows = ventral frontal carina; yellow arrow = labrum; green arrows = occipital carina; violet arrow = glossa of labium; ata = anterior tentorial bridge; dta = dorsal tentorial bridge; lp = labial palp; md = mandible; mp = maxillary palp; mtl = median tentorial lamella; of = occipital foramen; sc = scapus; teb = tentorial bridge.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSection id="E2B82F4DFFACFFAD18A11368FA6FF808" pageId="11" pageNumber="12" type="multiple">
|
||||
<paragraph id="8B8836A6FFACFFAD18A11368FA6FF8AA" blockId="11.[113,764,1843,1899]" lastBlockId="11.[810,1460,1843,1899]" pageId="11" pageNumber="12">
|
||||
<paragraph id="8B8836A6FFACFFAA18A11368FD8AF87A" blockId="11.[113,764,1843,1899]" lastBlockId="12.[129,777,1923,1979]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||||
The
|
||||
<typeStatus id="548C8804FFACFFAD1896136FFECAF88A" box="[186,278,1843,1867]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||||
of †
|
||||
|
|
@ -281,38 +272,18 @@ Middlekauff 1983
|
|||
</taxonomicName>
|
||||
at the lower end of the size range for this genus; the upper limit observed by Vilhelmsen (2004: table 2) being 16.0 mm (e.g.
|
||||
<taxonomicName id="4C374D25FFACFFAD1A8013EDFBD2F88A" authority="Scopoli 1763" authorityName="Scopoli" authorityYear="1763" box="[684,1038,1843,1992]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="abietinus">
|
||||
<emphasis id="B943EAB4FFACFFAD1A8013EDFC5DF88A" box="[684,897,1843,1992]" italics="true" pageId="11" pageNumber="12">
|
||||
Orussus
|
||||
<heading id="D0C081CAFFACFFAD1B06136FFC5DF88A" box="[810,897,1843,1867]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="12" reason="2">abietinus</heading>
|
||||
</emphasis>
|
||||
<emphasis id="B943EAB4FFACFFAD1A8013EDFC5DF88A" box="[684,897,1843,1992]" italics="true" pageId="11" pageNumber="12">Orussus abietinus</emphasis>
|
||||
Scopoli 1763
|
||||
</taxonomicName>
|
||||
). This is the more remarkable as the †
|
||||
<taxonomicName id="4C374D25FFACFFAD1DB01368FC72F8AA" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
<emphasis id="B943EAB4FFACFFAD1DB01368FC72F8AA" italics="true" pageId="11" pageNumber="12">O. juttagroehnae</emphasis>
|
||||
</taxonomicName>
|
||||
specimen is a female and females tend to be sig-
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFACFFAD1B06132EFA6FF808" blockId="11.[810,1459,1906,1993]" pageId="11" pageNumber="12">
|
||||
nificantly larger than males; the smallest specimen of
|
||||
specimen is a female and females tend to be significantly larger than males; the smallest specimen of
|
||||
<taxonomicName id="4C374D25FFACFFAD1D49132EFCA5F868" authorityName="Middlekauff" authorityYear="1983" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="minutus">
|
||||
<emphasis id="B943EAB4FFACFFAD1D49132EFCA5F868" italics="true" pageId="11" pageNumber="12">Orussus minutus</emphasis>
|
||||
</taxonomicName>
|
||||
measured by Vilhelmsen (2004) was a male. There is, of course, preservational bias favouring smaller insect fossils
|
||||
</paragraph>
|
||||
</subSection>
|
||||
<caption id="DF48662EFFABFFAA18AD11B1FF14F9B4" pageId="12" pageNumber="13" startId="12.[129,194,1517,1541]" targetBox="[132,1466,152,1489]" targetPageId="12" targetType="figure">
|
||||
<paragraph id="8B8836A6FFABFFAA18AD11B1FAA7F9C4" blockId="12.[129,1460,1517,1653]" box="[129,1403,1517,1541]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B943EAB4FFABFFAA18AD11B1FF04F9C4" bold="true" box="[129,216,1517,1541]" pageId="12" pageNumber="13">Figure 9.</emphasis>
|
||||
<taxonomicName id="4C374D25FFABFFAA18F111B1FE78F9C4" box="[221,420,1517,1541]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
<emphasis id="B943EAB4FFABFFAA18F111B1FE78F9C4" box="[221,420,1517,1541]" italics="true" pageId="12" pageNumber="13">Orussus juttagroehnae</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., female holotype (GPIH 5081, CCGG nº 1288). Ovipositor; Rendered from synchrotron scans.
|
||||
</paragraph>
|
||||
<paragraph id="8B8836A6FFABFFAA18AD1255FF14F9B4" blockId="12.[129,1460,1517,1653]" pageId="12" pageNumber="13">A, habitus, lateral with position of views indicated. B, profurca, posterior view. C, profurca and ovipositor loop, posterolateral view. D, complete ovipositor loop, lateral view. Black arrow = median groove on profurcal bridge; green arrows = dorsal branch of ovipositor loop; red arrows = ventral branch of ovipositor loop; blue arrows = profurca; fua = profurcal arm; fub = profurcal bridge; S1 = prosternum; 3vv = 3rd valvula.</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B8836A6FFABFFAA18AD129BFCD5F8BD" blockId="12.[128,777,1735,1916]" pageId="12" pageNumber="13">
|
||||
in amber, but †
|
||||
measured by Vilhelmsen (2004) was a male. There is, of course, preservational bias favouring smaller insect fossils in amber, but †
|
||||
<taxonomicName id="4C374D25FFABFFAA19081294FE14F91E" box="[292,456,1735,1759]" class="Insecta" family="Orussidae" genus="Orussus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jutagroehnae">
|
||||
<emphasis id="B943EAB4FFABFFAA19081294FE14F91E" box="[292,456,1735,1759]" italics="true" pageId="12" pageNumber="13">O. juttagroehnae</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
@ -339,12 +310,8 @@ Basibuyuk
|
|||
<emphasis id="B943EAB4FFABFFAA18C51339FE11F8BD" box="[233,461,1893,1916]" italics="true" pageId="12" pageNumber="13">Mesorussus taimyrensis</emphasis>
|
||||
<bibRefCitation id="EFA64B57FFABFFAA19FB1338FDA5F8BD" author="Rasnitsyn" box="[471,633,1892,1916]" firstAuthor="Rasnitsyn" pageId="12" pageNumber="13" pagination="349 - 357" refId="ref11217" refString="Rasnitsyn AP * New Hymenoptera from the Jurassic and Cretaceous of Asia * Paleontological Journal 1977; 11: 349 - 357 [English translation from Russian of Paleontologiceskij Zurnal 3: 98 - 108] *" type="journal article" year="1977">Rasnitsyn 1977</bibRefCitation>
|
||||
</taxonomicName>
|
||||
[female, pos-
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32D652DFFABFFAA18AD13DFFD8AF87A" pageId="12" pageNumber="13" type="multiple">
|
||||
<paragraph id="8B8836A6FFABFFAA18AD13DFFD8AF87A" blockId="12.[129,777,1923,1979]" pageId="12" pageNumber="13">
|
||||
<pageTitle id="CBA8EEC1FFABFFAA18AD13DFFD8AF87A" pageId="12" pageNumber="13">sibly 1.75 mm—body length of incomplete fossil estimated by Vilhelmsen (2004)] are the smallest recorded.</pageTitle>
|
||||
[female,
|
||||
<pageTitle id="CBA8EEC1FFABFFAA18AD13DFFD8AF87A" pageId="12" pageNumber="13">possibly 1.75 mm—body length of incomplete fossil estimated by Vilhelmsen (2004)] are the smallest recorded.</pageTitle>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
|
|
|
|||
Loading…
Reference in a new issue