200 lines
15 KiB
XML
200 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.690.13751" ID-GBIF-Dataset="356a4c29-470b-4ca5-8503-76191b4fef6f" ID-PMC="PMC5672587" ID-Pensoft-Pub="1313-2970-690-1" ID-PubMed="29118595" ID-ZBK="C1B8D7C059E54C3A944F69F4FDE96B20" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-690-1" ModsDocOrigin="ZooKeys 690" ModsDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" checkinTime="1502920743207" checkinUser="pensoft" docAuthor="Shpeley, Danny, Hunting, Wesley & Ball, George E." docDate="2017" docId="9BA4BAE1415D6D3B68490F4000B2FB5F" docLanguage="en" docName="ZooKeys 690: 1-195" docOrigin="ZooKeys 690" docSource="http://dx.doi.org/10.3897/zookeys.690.13751" docTitle="Selenophorus clypealis Ball & Shpeley" docType="treatment" docVersion="5" lastPageNumber="42" masterDocId="FE026465FF86843AE44A7B6EFF8DC422" masterDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" masterLastPageNumber="195" masterPageNumber="1" pageNumber="41" updateTime="1668164690231" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shpeley, Danny</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunting, Wesley</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ball, George E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>690</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>195</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.690.13751</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.690.13751</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-690-1</mods:identifier>
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<mods:identifier type="ZBK">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
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<mods:identifier type="ZooBank">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="132785168" LSID="urn:lsid:plazi:treatment:9BA4BAE1415D6D3B68490F4000B2FB5F" httpUri="http://treatment.plazi.org/id/9BA4BAE1415D6D3B68490F4000B2FB5F" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<subSubSection pageId="40" pageNumber="41" type="nomenclature">
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<paragraph pageId="40" pageNumber="41">
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<taxonomicName authority="Ball & Shpeley" class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus clypealis" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="clypealis">Selenophorus clypealis Ball & Shpeley</taxonomicName>
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Figs 25A, 27
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 30
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="reference_group">
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<paragraph pageId="40" pageNumber="41">
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus clypealis" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="clypealis">Selenophorus clypealis</taxonomicName>
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Ball & Shpeley, 1992: 101.-
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<bibRefCitation pageId="40" pageNumber="41">Ball 1992</bibRefCitation>
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: 85.-
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="105" pageNumber="106" title="Systematic list of ground beetles of the World (InsectaColeoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W." year="1998">Lorenz 1998</bibRefCitation>
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: 355.-
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<bibRefCitation pageId="40" pageNumber="41">Lorenz 2005</bibRefCitation>
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: 376.-
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<bibRefCitation author="Perez-Gelabert, DE" journalOrPublisher="Zootaxa" pageId="106" pageNumber="107" pagination="1 - 530" title="Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography." volume="1831" year="2008">Perez-Gelabert 2008</bibRefCitation>
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: 79.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="type material">
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<paragraph pageId="40" pageNumber="41">Type material.</paragraph>
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<paragraph pageId="40" pageNumber="41">Complete label data for type material (holotype (MCZC) and allotype (WIBF)) are provided in the original description.</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="type locality">
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<paragraph pageId="40" pageNumber="41">Type locality.</paragraph>
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<paragraph pageId="40" pageNumber="41">Source of the Matelas (River), near Ennery, Artibonite Department, Haiti, Hispaniola.</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="diagnosis">
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<paragraph pageId="40" pageNumber="41">Diagnosis.</paragraph>
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<paragraph pageId="40" pageNumber="41">This species is readily separated from the other four members of the hylacis species group on a combination of: clypeus with anterior margin markedly concave, small size and pronotum with hind angles rounded.</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="descriptive notes">
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<paragraph pageId="40" pageNumber="41">Descriptive notes.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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Data for SBL in Table 1. Habitus as in Fig. 25A. Labrum with anterior margin deeply notched; clypeus with anterior margin markedly concave. Antennae, mouthparts and legs testaceous. Dorsal and ventral surfaces rufo-brunneous to brunneo-piceous; lateral bead of pronotum paler. Head and pronotum shiny, microlines not visible at 100
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<normalizedToken originalValue="×">x</normalizedToken>
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. Elytra shiny, with mesh pattern transverse, transverse microlines just visible at 100
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<normalizedToken originalValue="×">x</normalizedToken>
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; iridescent, brighter than observed in
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<taxonomicName lsidName="S. parvus" pageId="40" pageNumber="41" rank="species" species="parvus">S. parvus</taxonomicName>
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. Pronotum with posteriolateral impressions impunctate; posteriolateral angles rounded. Elytral striae impunctate, except the standard setigerous punctures in striae 2, 5 and 7. Males with adhesive vestiture on tarsomeres 1-4 of fore- and mid-tarsi; females without adhesive vestiture on tarsomeres 1-4 of fore- and mid-tarsi. Tarsomere 1 of fore- and mid-tarsus in females not expanded. Both males and females with four terminal setae near the posterior margin on sternum VII.
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</paragraph>
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<paragraph pageId="40" pageNumber="41">
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Male genitalia. Fig. 27
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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. Apical portion of phallic median lobe moderately long, trapezoidal, symmetrically broadly rounded in dorsal/ventral aspect, with narrow dorsal flange; endophallus medially with three patches of short, thin spines, one darkened microtrichial field near basal bulb in left lateral aspect; without lamina; ostium anopic. Ventral surface of shaft smooth.
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</paragraph>
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<paragraph pageId="40" pageNumber="41">
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Ovipositor and female reproductive tract. Very similar to
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<taxonomicName lsidName="S. dessalinesi" pageId="40" pageNumber="41" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
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, Fig. 29A. For details, see this topic for
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<taxonomicName lsidName="S. dessalinesi" pageId="40" pageNumber="41" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
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, below.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="distribution">
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<paragraph pageId="40" pageNumber="41">Geographical distribution.</paragraph>
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<paragraph pageId="40" pageNumber="41">Fig. 30. This species is known only from the Greater Antillean island of Hispaniola and the island of Little St. James in the Virgin Islands.</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="chorological affinities and relationships">
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<paragraph pageId="40" pageNumber="41">Chorological affinities and relationships.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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Within the West Indian hylacis species group, the range of
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<taxonomicName lsidName="S. clypealis" pageId="40" pageNumber="41" rank="species" species="clypealis">S. clypealis</taxonomicName>
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is overlapped by the ranges of
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<taxonomicName lsidName="S. subquadratus" pageId="40" pageNumber="41" rank="species" species="subquadratus">S. subquadratus</taxonomicName>
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and
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<taxonomicName lsidName="S. dessalinesi" pageId="40" pageNumber="41" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
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. Relationships of
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<taxonomicName lsidName="S. clypealis" pageId="40" pageNumber="41" rank="species" species="clypealis">S. clypealis</taxonomicName>
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are not postulated beyond species group membership.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="materials_examined">
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<paragraph pageId="41" pageNumber="42">
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<pageBreakToken pageId="41" pageNumber="42" start="start">Material</pageBreakToken>
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examined.
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</paragraph>
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<paragraph pageId="41" pageNumber="42">In addition to type material, we have seen a total of 6 specimens (1 male, 5 females). See Appendix for details.</paragraph>
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<caption pageId="41" pageNumber="42">
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<paragraph pageId="41" pageNumber="42">
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Figure 25. Habitus digital images of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
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species group, in part, dorsal aspect. A
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<taxonomicName lsidName="S. clypealis" pageId="41" pageNumber="42" rank="species" species="clypealis">S. clypealis</taxonomicName>
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Ball & Shpeley B
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<taxonomicName lsidName="S. dessalinesi" pageId="41" pageNumber="42" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
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Ball & Shpeley C
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<taxonomicName lsidName="S. dubius" pageId="41" pageNumber="42" rank="species" species="dubius">S. dubius</taxonomicName>
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Putzeys Dparvus Darlington. Scale bars 5 mm.
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</paragraph>
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</caption>
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<caption pageId="41" pageNumber="42">
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<paragraph pageId="41" pageNumber="42">
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Figure 26. Habitus digital image of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
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species group, in part, dorsal aspect,
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<taxonomicName lsidName="S. subquadratus" pageId="41" pageNumber="42" rank="species" species="subquadratus">S. subquadratus</taxonomicName>
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(Putzeys). Scale bar 5 mm.
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</paragraph>
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</caption>
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<caption pageId="41" pageNumber="42">
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<paragraph pageId="41" pageNumber="42">
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Figure 27. Digital images of male genitalia of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
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species group, in part. A, D right lateral aspect B, E dorsal aspect C, F left lateral aspect
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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<taxonomicName lsidName="S. clypealis" pageId="41" pageNumber="42" rank="species" species="clypealis">S. clypealis</taxonomicName>
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Ball & Shpeley
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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<taxonomicName lsidName="S. dessalinesi" pageId="41" pageNumber="42" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
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Ball & Shpeley. Scale bars 1 mm.
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</paragraph>
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</caption>
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<caption pageId="41" pageNumber="42">
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<paragraph pageId="41" pageNumber="42">
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Figure 28. Digital images of male genitalia of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
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species group, in part. A, D right lateral aspect B, E dorsal aspect C, F left lateral aspect
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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<taxonomicName lsidName="S. parvus" pageId="41" pageNumber="42" rank="species" species="parvus">S. parvus</taxonomicName>
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Darlington
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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<taxonomicName lsidName="S. subquadratus" pageId="41" pageNumber="42" rank="species" species="subquadratus">S. subquadratus</taxonomicName>
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(Putzeys). Scale bars 1 mm.
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</paragraph>
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</caption>
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<caption pageId="41" pageNumber="42">
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<paragraph pageId="41" pageNumber="42">
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Figure 29. Line drawings of female reproductive tract of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
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species group, in part, ventral aspect. A
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<taxonomicName lsidName="S. dessalinesi" pageId="41" pageNumber="42" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
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Ball & Shpeley B
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<taxonomicName lsidName="S. parvus" pageId="41" pageNumber="42" rank="species" species="parvus">S. parvus</taxonomicName>
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Darlington. Legend: bc bursa copulatrix co common oviduct gc1 gonocoxite 1 gc2 gonocoxite 2 sp spermatheca sp1 spermatheca 1 sp2 spermatheca 2 spg spermathecal gland spgd spermathecal gland duct; v valvifer. Scale bars 1 mm.
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</paragraph>
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</caption>
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<caption pageId="41" pageNumber="42">
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<paragraph pageId="41" pageNumber="42">
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Figure 30. Map of West Indies showing known localities for species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
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species group.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |