treatments-xml/data/9B/A4/BA/9BA4BAE1415D6D3B68490F4000B2FB5F.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.690.13751" ID-GBIF-Dataset="356a4c29-470b-4ca5-8503-76191b4fef6f" ID-PMC="PMC5672587" ID-Pensoft-Pub="1313-2970-690-1" ID-PubMed="29118595" ID-ZBK="C1B8D7C059E54C3A944F69F4FDE96B20" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-690-1" ModsDocOrigin="ZooKeys 690" ModsDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" checkinTime="1502920743207" checkinUser="pensoft" docAuthor="Shpeley, Danny, Hunting, Wesley &amp; Ball, George E." docDate="2017" docId="9BA4BAE1415D6D3B68490F4000B2FB5F" docLanguage="en" docName="ZooKeys 690: 1-195" docOrigin="ZooKeys 690" docSource="http://dx.doi.org/10.3897/zookeys.690.13751" docTitle="Selenophorus clypealis Ball &amp; Shpeley" docType="treatment" docVersion="5" lastPageNumber="42" masterDocId="FE026465FF86843AE44A7B6EFF8DC422" masterDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" masterLastPageNumber="195" masterPageNumber="1" pageNumber="41" updateTime="1668164690231" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shpeley, Danny</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hunting, Wesley</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ball, George E.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>690</mods:number>
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<mods:start>1</mods:start>
<mods:end>195</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.690.13751</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.690.13751</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-690-1</mods:identifier>
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<treatment ID-GBIF-Taxon="132785168" LSID="urn:lsid:plazi:treatment:9BA4BAE1415D6D3B68490F4000B2FB5F" httpUri="http://treatment.plazi.org/id/9BA4BAE1415D6D3B68490F4000B2FB5F" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<subSubSection pageId="40" pageNumber="41" type="nomenclature">
<paragraph pageId="40" pageNumber="41">
<taxonomicName authority="Ball &amp; Shpeley" class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus clypealis" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="clypealis">Selenophorus clypealis Ball &amp; Shpeley</taxonomicName>
Figs 25A, 27 
<normalizedToken originalValue="A–C">A-C</normalizedToken>
, 30
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="reference_group">
<paragraph pageId="40" pageNumber="41">
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus clypealis" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="clypealis">Selenophorus clypealis</taxonomicName>
Ball &amp; Shpeley, 1992: 101.- 
<bibRefCitation pageId="40" pageNumber="41">Ball 1992</bibRefCitation>
: 85.- 
<bibRefCitation author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="105" pageNumber="106" title="Systematic list of ground beetles of the World (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W." year="1998">Lorenz 1998</bibRefCitation>
: 355.- 
<bibRefCitation pageId="40" pageNumber="41">Lorenz 2005</bibRefCitation>
: 376.- 
<bibRefCitation author="Perez-Gelabert, DE" journalOrPublisher="Zootaxa" pageId="106" pageNumber="107" pagination="1 - 530" title="Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography." volume="1831" year="2008">Perez-Gelabert 2008</bibRefCitation>
: 79.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="type material">
<paragraph pageId="40" pageNumber="41">Type material.</paragraph>
<paragraph pageId="40" pageNumber="41">Complete label data for type material (holotype (MCZC) and allotype (WIBF)) are provided in the original description.</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="type locality">
<paragraph pageId="40" pageNumber="41">Type locality.</paragraph>
<paragraph pageId="40" pageNumber="41">Source of the Matelas (River), near Ennery, Artibonite Department, Haiti, Hispaniola.</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="diagnosis">
<paragraph pageId="40" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="40" pageNumber="41">This species is readily separated from the other four members of the hylacis species group on a combination of: clypeus with anterior margin markedly concave, small size and pronotum with hind angles rounded.</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="descriptive notes">
<paragraph pageId="40" pageNumber="41">Descriptive notes.</paragraph>
<paragraph pageId="40" pageNumber="41">
Data for SBL in Table 1. Habitus as in Fig. 25A. Labrum with anterior margin deeply notched; clypeus with anterior margin markedly concave. Antennae, mouthparts and legs testaceous. Dorsal and ventral surfaces rufo-brunneous to brunneo-piceous; lateral bead of pronotum paler. Head and pronotum shiny, microlines not visible at 100 
<normalizedToken originalValue="×">x</normalizedToken>
. Elytra shiny, with mesh pattern transverse, transverse microlines just visible at 100 
<normalizedToken originalValue="×">x</normalizedToken>
; iridescent, brighter than observed in 
<taxonomicName lsidName="S. parvus" pageId="40" pageNumber="41" rank="species" species="parvus">S. parvus</taxonomicName>
. Pronotum with posteriolateral impressions impunctate; posteriolateral angles rounded. Elytral striae impunctate, except the standard setigerous punctures in striae 2, 5 and 7. Males with adhesive vestiture on tarsomeres 1-4 of fore- and mid-tarsi; females without adhesive vestiture on tarsomeres 1-4 of fore- and mid-tarsi. Tarsomere 1 of fore- and mid-tarsus in females not expanded. Both males and females with four terminal setae near the posterior margin on sternum VII.
</paragraph>
<paragraph pageId="40" pageNumber="41">
Male genitalia. Fig. 27 
<normalizedToken originalValue="A–C">A-C</normalizedToken>
. Apical portion of phallic median lobe moderately long, trapezoidal, symmetrically broadly rounded in dorsal/ventral aspect, with narrow dorsal flange; endophallus medially with three patches of short, thin spines, one darkened microtrichial field near basal bulb in left lateral aspect; without lamina; ostium anopic. Ventral surface of shaft smooth.
</paragraph>
<paragraph pageId="40" pageNumber="41">
Ovipositor and female reproductive tract. Very similar to 
<taxonomicName lsidName="S. dessalinesi" pageId="40" pageNumber="41" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
, Fig. 29A. For details, see this topic for 
<taxonomicName lsidName="S. dessalinesi" pageId="40" pageNumber="41" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
, below.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="distribution">
<paragraph pageId="40" pageNumber="41">Geographical distribution.</paragraph>
<paragraph pageId="40" pageNumber="41">Fig. 30. This species is known only from the Greater Antillean island of Hispaniola and the island of Little St. James in the Virgin Islands.</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="chorological affinities and relationships">
<paragraph pageId="40" pageNumber="41">Chorological affinities and relationships.</paragraph>
<paragraph pageId="40" pageNumber="41">
Within the West Indian hylacis species group, the range of 
<taxonomicName lsidName="S. clypealis" pageId="40" pageNumber="41" rank="species" species="clypealis">S. clypealis</taxonomicName>
is overlapped by the ranges of 
<taxonomicName lsidName="S. subquadratus" pageId="40" pageNumber="41" rank="species" species="subquadratus">S. subquadratus</taxonomicName>
and 
<taxonomicName lsidName="S. dessalinesi" pageId="40" pageNumber="41" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
. Relationships of 
<taxonomicName lsidName="S. clypealis" pageId="40" pageNumber="41" rank="species" species="clypealis">S. clypealis</taxonomicName>
are not postulated beyond species group membership.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="materials_examined">
<paragraph pageId="41" pageNumber="42">
<pageBreakToken pageId="41" pageNumber="42" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="41" pageNumber="42">In addition to type material, we have seen a total of 6 specimens (1 male, 5 females). See Appendix for details.</paragraph>
<caption pageId="41" pageNumber="42">
<paragraph pageId="41" pageNumber="42">
Figure 25. Habitus digital images of 
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
species group, in part, dorsal aspect. A 
<taxonomicName lsidName="S. clypealis" pageId="41" pageNumber="42" rank="species" species="clypealis">S. clypealis</taxonomicName>
Ball &amp; Shpeley B 
<taxonomicName lsidName="S. dessalinesi" pageId="41" pageNumber="42" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
Ball &amp; Shpeley C 
<taxonomicName lsidName="S. dubius" pageId="41" pageNumber="42" rank="species" species="dubius">S. dubius</taxonomicName>
Putzeys Dparvus Darlington. Scale bars 5 mm.
</paragraph>
</caption>
<caption pageId="41" pageNumber="42">
<paragraph pageId="41" pageNumber="42">
Figure 26. Habitus digital image of 
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
species group, in part, dorsal aspect, 
<taxonomicName lsidName="S. subquadratus" pageId="41" pageNumber="42" rank="species" species="subquadratus">S. subquadratus</taxonomicName>
(Putzeys). Scale bar 5 mm.
</paragraph>
</caption>
<caption pageId="41" pageNumber="42">
<paragraph pageId="41" pageNumber="42">
Figure 27. Digital images of male genitalia of 
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
species group, in part. A, D right lateral aspect B, E dorsal aspect C, F left lateral aspect 
<normalizedToken originalValue="A–C">A-C</normalizedToken>
<taxonomicName lsidName="S. clypealis" pageId="41" pageNumber="42" rank="species" species="clypealis">S. clypealis</taxonomicName>
Ball &amp; Shpeley 
<normalizedToken originalValue="D–F">D-F</normalizedToken>
<taxonomicName lsidName="S. dessalinesi" pageId="41" pageNumber="42" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
Ball &amp; Shpeley. Scale bars 1 mm.
</paragraph>
</caption>
<caption pageId="41" pageNumber="42">
<paragraph pageId="41" pageNumber="42">
Figure 28. Digital images of male genitalia of 
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
species group, in part. A, D right lateral aspect B, E dorsal aspect C, F left lateral aspect 
<normalizedToken originalValue="A–C">A-C</normalizedToken>
<taxonomicName lsidName="S. parvus" pageId="41" pageNumber="42" rank="species" species="parvus">S. parvus</taxonomicName>
Darlington 
<normalizedToken originalValue="D–F">D-F</normalizedToken>
<taxonomicName lsidName="S. subquadratus" pageId="41" pageNumber="42" rank="species" species="subquadratus">S. subquadratus</taxonomicName>
(Putzeys). Scale bars 1 mm.
</paragraph>
</caption>
<caption pageId="41" pageNumber="42">
<paragraph pageId="41" pageNumber="42">
Figure 29. Line drawings of female reproductive tract of 
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
species group, in part, ventral aspect. A 
<taxonomicName lsidName="S. dessalinesi" pageId="41" pageNumber="42" rank="species" species="dessalinesi">S. dessalinesi</taxonomicName>
Ball &amp; Shpeley B 
<taxonomicName lsidName="S. parvus" pageId="41" pageNumber="42" rank="species" species="parvus">S. parvus</taxonomicName>
Darlington. Legend: bc bursa copulatrix co common oviduct gc1 gonocoxite 1 gc2 gonocoxite 2 sp spermatheca sp1 spermatheca 1 sp2 spermatheca 2 spg spermathecal gland spgd spermathecal gland duct; v valvifer. Scale bars 1 mm.
</paragraph>
</caption>
<caption pageId="41" pageNumber="42">
<paragraph pageId="41" pageNumber="42">
Figure 30. Map of West Indies showing known localities for species of 
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus hylacis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hylacis">Selenophorus hylacis</taxonomicName>
species group.
</paragraph>
</caption>
</subSubSection>
</treatment>
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