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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.418.7629" ID-PMC="PMC4104466" ID-Pensoft-Pub="1313-2970-418-1" ID-Pensoft-UUID="F034FFA3FFAFFFBEB578C42DD65AFFA2" ID-PubMed="25061358" ID-Zenodo-Dep="578347" ID-ZooBank="D71C8A3DA6CA40A5B3A034A1FD1C16A0" ModsDocID="1313-2970-418-1" checkinTime="1451245725588" checkinUser="pensoft" docAuthor="Skvarla, Michael J., Fisher, J. Ray &amp; Dowling, Ashley P. G." docDate="2014" docId="5EE57B0B89DB4510277EB48DAC3F1E9E" docLanguage="en" docName="ZooKeys 418: 1-103" docOrigin="ZooKeys 418" docPubDate="2014-06-20" docSource="http://dx.doi.org/10.3897/zookeys.418.7629" docTitle="Scutopalus Den Heyer 1979" docType="treatment" docVersion="5" id="F034FFA3FFAFFFBEB578C42DD65AFFA2" lastPageNumber="40" masterDocId="F034FFA3FFAFFFBEB578C42DD65AFFA2" masterDocTitle="A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records" masterLastPageNumber="103" masterPageNumber="1" pageNumber="37" updateTime="1668158708043" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skvarla, Michael J.</mods:namePart>
<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, J. Ray</mods:namePart>
<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>418</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>103</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.418.7629</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.418.7629</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-418-1</mods:identifier>
<mods:identifier type="ZooBank">D71C8A3DA6CA40A5B3A034A1FD1C16A0</mods:identifier>
<mods:identifier type="Pensoft-UUID">F034FFA3FFAFFFBEB578C42DD65AFFA2</mods:identifier>
<mods:identifier type="Zenodo-Dep">578347</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152053721" LSID="urn:lsid:plazi:treatment:5EE57B0B89DB4510277EB48DAC3F1E9E" httpUri="http://treatment.plazi.org/id/5EE57B0B89DB4510277EB48DAC3F1E9E" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37">
<subSubSection pageId="36" pageNumber="37" type="nomenclature">
<paragraph pageId="36" pageNumber="37">
<taxonomicName LSID="5EE57B0B-89DB-4510-277E-B48DAC3F1E9E" authority="Den Heyer, 1979" authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scutopalus" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Scutopalus Den Heyer, 1979</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="historical review">
<paragraph pageId="36" pageNumber="37">Historical review.</paragraph>
<paragraph pageId="36" pageNumber="37">
<bibRefCitation author="Den Heyer, J" journalOrPublisher="Acarologia" pageId="86" pageNumber="87" pagination="187 - 193" refId="B81" refString="Den Heyer, J, 1979c. Scutopalus, a new cunaxid genus from the Ethiopian Region (Prostigmata: Acari). Acarologia 21: 187 - 193" title="Scutopalus, a new cunaxid genus from the Ethiopian Region (Prostigmata: Acari)." volume="21" year="1979 c">Den Heyer (1979c)</bibRefCitation>
erected
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus</emphasis>
</taxonomicName>
for
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus arboreus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="latisetosus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus latisetosus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Shiba, M" journalOrPublisher="Nature and Life in South East Asia" pageId="93" pageNumber="94" pagination="83 - 229" refId="B232" refString="Shiba, M, 1978. Taxonomic investigation on free-living Prostigmata from the Malay Peninsula. Nature and Life in South East Asia 7: 83 - 229" title="Taxonomic investigation on free-living Prostigmata from the Malay Peninsula." volume="7" year="1978">Shiba (1978)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="clavatus">
<emphasis italics="true" pageId="36" pageNumber="37">Cunaxoides clavatus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Kuznetzov, NN" journalOrPublisher="Trudy Gosudarstvennogo Nikitskogo Botanicheskogo Sada" pageId="89" pageNumber="90" pagination="51 - 105" refId="B163" refString="Kuznetzov, NN, Livshitz, IZ, 1979. Predatory mites of the Nikita Botanical Gardens (Acariformes: Bdellidae, Cunaxidae Camarobiidae). Trudy Gosudarstvennogo Nikitskogo Botanicheskogo Sada 79: 51 - 105" title="Predatory mites of the Nikita Botanical Gardens (Acariformes: Bdellidae, Cunaxidae Camarobiidae)." volume="79" year="1979">Kuznetzov and Livshitz (1979)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="trepidus">
<emphasis italics="true" pageId="36" pageNumber="37">Cunaxoides trepidus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Tseng, YH" journalOrPublisher="Quarterly Journal of the Taiwan Museum" pageId="94" pageNumber="95" pagination="253 - 277" refId="B250" refString="Tseng, YH, 1980. Taxonomical study of the mite family Cunaxidae from Taiwan (Acarina: Tromoidiformes). Quarterly Journal of the Taiwan Museum 33 (3-4): 253 - 277" title="Taxonomical study of the mite family Cunaxidae from Taiwan (Acarina: Tromoidiformes)." volume="33" year="1980">Tseng (1980)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="osseus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides osseus</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="unguianalis">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides unguianalis</emphasis>
</taxonomicName>
.
<bibRefCitation author="Gupta, SK" journalOrPublisher="Records of the Zoological Survey of India" pageId="88" pageNumber="89" pagination="189 - 213" refId="B138" refString="Gupta, SK, Ghosh, SK, 1980. Some prostigmatid mites (Acarina) from Andaman and Nicobar Islands. Records of the Zoological Survey of India 77: 189 - 213" title="Some prostigmatid mites (Acarina) from Andaman and Nicobar Islands." volume="77" year="1980">Gupta and Ghosh (1980)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pradhani">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides pradhani</emphasis>
</taxonomicName>
.
<bibRefCitation author="Smiley, RL" journalOrPublisher="Indira Publishing House, West Bloomington, Michigan" pageId="93" pageNumber="94" refId="B239" refString="Smiley, RL, 1992. The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomington, Michigan" title="The predatory mite family Cunaxidae (Acari) of the world with a new classification." year="1992">Smiley (1992)</bibRefCitation>
synonymized
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus</emphasis>
</taxonomicName>
with
<taxonomicName authorityName="Smiley" authorityYear="1975" class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides</emphasis>
</taxonomicName>
and transferred
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="trepidus">
<emphasis italics="true" pageId="36" pageNumber="37">Cunaxoides trepidus</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Smiley" authorityYear="1975" class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides</emphasis>
</taxonomicName>
.
<bibRefCitation author="Corpuz-Raros, LA" journalOrPublisher="Asia Life Sciences" pageId="85" pageNumber="86" pagination="125 - 140" refId="B60" refString="Corpuz-Raros, LA, 1996c. Philippine predatory mites of the family Cunaxidae (Acari). 6. Genus Neocunaxoides Smiley with a new species record from Central Kalimantan, Borneo, Indonesia. Asia Life Sciences 5 (2): 125 - 140" title="Philippine predatory mites of the family Cunaxidae (Acari). 6. Genus Neocunaxoides Smiley with a new species record from Central Kalimantan, Borneo, Indonesia." volume="5" year="1996 c">Corpuz-Raros (1996c)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="makapalus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides makapalus</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides philippinensis</emphasis>
</taxonomicName>
, and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides rugosus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Lin, LZ" journalOrPublisher="Fujian Sciences and Technology Press, Fuzhou" pageId="90" pageNumber="91" pagination="109 - 121" refId="B176" refString="Lin, LZ, Zhang, YZ, 2000. Fauna of Insects in Fuijan Province of China. Fujian Sciences and Technology Press, Fuzhou: 109 - 121" title="Fauna of Insects in Fuijan Province of China" year="2000">Lin and Zhang (2000)</bibRefCitation>
recorded
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="clavatus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides clavatus</emphasis>
</taxonomicName>
from tea in China.
<bibRefCitation author="Sionti, PG" journalOrPublisher="International Journal of Acarology" pageId="93" pageNumber="94" pagination="315 - 325" publicationUrl="10.1080/01647950308684347" refId="B234" refString="Sionti, PG, Papadoulis, GT, 2003. Cunaxid mites of Greece (Acari: Cunaxidae). International Journal of Acarology 29 (4): 315 - 325, DOI: 10.1080/01647950308684347" title="Cunaxid mites of Greece (Acari: Cunaxidae)." url="10.1080/01647950308684347" volume="29" year="2003">Sionti and Papadoulis (2003)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="abiesae">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides abiesae</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="smolikensis">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides smolikensis</emphasis>
</taxonomicName>
.
<bibRefCitation author="Bashir, MH" journalOrPublisher="Pakistan Entomologist" pageId="83" pageNumber="84" pagination="29 - 32" refId="B18" refString="Bashir, MH, Afzal, M, 2004b. A new species of genus Neocunaxoides (Cunaxidae) from Punjab, Pakistan. Pakistan Entomologist 26 (2): 29 - 32" title="A new species of genus Neocunaxoides (Cunaxidae) from Punjab, Pakistan." volume="26" year="2004 b">Bashir and Afzal (2004b)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gilbertoi">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides gilbertoi</emphasis>
</taxonomicName>
. Castro and
<bibRefCitation author="Den Heyer, J" journalOrPublisher="Arthropod fauna of the UAE" pageId="86" pageNumber="87" pagination="17 - 25" refId="B92" refString="Den Heyer, J, 2009. . Arthropod fauna of the UAE Vol. 2: 17 - 25" volume="Vol. 2" year="2009">Den Heyer (2009)</bibRefCitation>
transferred
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="trepidus">
<emphasis italics="true" pageId="36" pageNumber="37">Pulaeus trepidus</emphasis>
</taxonomicName>
(=
<taxonomicName genus="Neounaxoides" lsidName="" pageId="36" pageNumber="37" rank="species" species="trepidus">
<emphasis italics="true" pageId="36" pageNumber="37">Neounaxoides trepidus</emphasis>
</taxonomicName>
) to
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Rocha, MDS" journalOrPublisher="Zootaxa" pageId="92" pageNumber="93" pagination="38 - 44" publicationUrl="10.11646/zootaxa.3734.1.4" refId="B221" refString="Rocha, MDS, Skvarla, MJ, Ferla, NJ, 2013. A new species of Scutopalus (Acari: Cunaxidae: Cunaxoidinae) from Rio Grande do Sul State, Brazil with a key to world species. Zootaxa 3734 (1): 38 - 44, DOI: 10.11646/zootaxa.3734.1.4" title="A new species of Scutopalus (Acari: Cunaxidae: Cunaxoidinae) from Rio Grande do Sul State, Brazil with a key to world species." url="10.11646/zootaxa.3734.1.4" volume="3734" year="2013">Rocha et al. (2013)</bibRefCitation>
described
<taxonomicName authorityName="Rocha, Skvarla &amp; Ferla" authorityYear="2013" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="tomentosus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus tomentosus</emphasis>
</taxonomicName>
and transferred
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="makapalus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides makapalus</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides philippinensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides rugosus</emphasis>
</taxonomicName>
, and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="unguianalis">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides unguianalis</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="diagnosis">
<paragraph pageId="36" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="36" pageNumber="37">
<emphasis italics="true" pageId="36" pageNumber="37">Gnathosoma</emphasis>
.
<emphasis bold="true" pageId="36" pageNumber="37">Pedipalps</emphasis>
3-segmented. Femurogenu complimented with 5 sts. Tibiotarsi at least twice as long as wide and complemented with 5 sts, 1 asl. Subterminal pointed process on pedipalp tibiotarsal claw absent; small teeth on pedipalp tibiotarsal claw absent.
<emphasis bold="true" pageId="36" pageNumber="37">Subcapitulum</emphasis>
with 6 pairs of setae (
<emphasis italics="true" pageId="36" pageNumber="37">hg1</emphasis>
-
<emphasis italics="true" pageId="36" pageNumber="37">4</emphasis>
and 2 pairs of adoral setae).
<emphasis bold="true" pageId="36" pageNumber="37">Chelicera</emphasis>
without seta.
</paragraph>
<paragraph pageId="36" pageNumber="37">
<emphasis italics="true" pageId="36" pageNumber="37">Idiosoma, dorsal</emphasis>
. Proterosoma with a well-defined shield present, complemented with 2 pairs of setae (
<emphasis italics="true" pageId="36" pageNumber="37">lps</emphasis>
and
<emphasis italics="true" pageId="36" pageNumber="37">mps</emphasis>
) and 2 pairs of setose sensillae (
<emphasis italics="true" pageId="36" pageNumber="37">at</emphasis>
and
<emphasis italics="true" pageId="36" pageNumber="37">pt</emphasis>
). Dorsal hysterosoma with a well-defined plate fused to the proterosomal plate. Small platelets may be present laterad and posterior to the dorsal shield. Setae
<emphasis italics="true" pageId="36" pageNumber="37">c1</emphasis>
-
<emphasis italics="true" pageId="36" pageNumber="37">h1</emphasis>
,
<emphasis italics="true" pageId="36" pageNumber="37">c2</emphasis>
, and
<emphasis italics="true" pageId="36" pageNumber="37">h2</emphasis>
present. Setae
<emphasis italics="true" pageId="36" pageNumber="37">f2</emphasis>
absent. Integument not covered in shields or plates striated.
</paragraph>
<paragraph pageId="36" pageNumber="37">
<emphasis italics="true" pageId="36" pageNumber="37">Idiosoma, ventral</emphasis>
.
<emphasis bold="true" pageId="36" pageNumber="37">Coxae</emphasis>
well-sclerotized. Coxae I-II fused medially. Coxae III-IV fused. Genital plates each bear 4 setae (
<emphasis italics="true" pageId="36" pageNumber="37">g1</emphasis>
-
<emphasis italics="true" pageId="36" pageNumber="37">4</emphasis>
); 2 pairs of genital papillae visible underneath the plates. A small platelet may be present laterad the genital plate. Integument not covered in shields or plates striated.
<emphasis bold="true" pageId="36" pageNumber="37">Legs.</emphasis>
Basifemora I-IV setal formula 3-4-2-0 sts. Telofemora I-IV setal formula 5-5-4-3. Tibiae III with 1 bsl, 5 sts. Tibiae IV with 5 sts (4 short, 1 long).
</paragraph>
</subSubSection>
<subSubSection lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37" type="key to female scutopalus">
<paragraph pageId="36" pageNumber="37">
Key to female
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="36" pageNumber="37">
(modified from
<bibRefCitation author="Rocha, MDS" journalOrPublisher="Zootaxa" pageId="92" pageNumber="93" pagination="38 - 44" publicationUrl="10.11646/zootaxa.3734.1.4" refId="B221" refString="Rocha, MDS, Skvarla, MJ, Ferla, NJ, 2013. A new species of Scutopalus (Acari: Cunaxidae: Cunaxoidinae) from Rio Grande do Sul State, Brazil with a key to world species. Zootaxa 3734 (1): 38 - 44, DOI: 10.11646/zootaxa.3734.1.4" title="A new species of Scutopalus (Acari: Cunaxidae: Cunaxoidinae) from Rio Grande do Sul State, Brazil with a key to world species." url="10.11646/zootaxa.3734.1.4" volume="3734" year="2013">Rocha et al. 2013</bibRefCitation>
).
</paragraph>
<paragraph pageId="36" pageNumber="37">
As suggested by
<bibRefCitation author="Den Heyer, J" editor="Roskov, Y" journalOrPublisher="Reading, UK" pageId="86" pageNumber="87" publicationUrl="10.1080/01647954.2010.495953" refId="B94" refString="Den Heyer, J, 2011b. BdelloideaBase: Bdellid &amp; Cunaxid Databases (version Sep 2011). In: Roskov, Y, Kunze, T, Paglinawan, L, Orrell, T, Nicolson, D, Culham, A, Bailly, N, Kirk, P, Bourgoin, T, Baillargeon, G, Hernandez, F, De Wever, A, Eds., Species 2000 &amp; ITIS Catalogue of Life, 2013 Annual Checklist. Reading, UK, DOI: 10.1080/01647954.2010.495953" title="BdelloideaBase: Bdellid &amp; Cunaxid Databases (version Sep 2011)." url="10.1080/01647954.2010.495953" volumeTitle="Species 2000 &amp; ITIS Catalogue of Life, 2013 Annual Checklist." year="2011 b">Den Heyer (2011b)</bibRefCitation>
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pradhani">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides pradhani</emphasis>
</taxonomicName>
(
<bibRefCitation author="Gupta, SK" journalOrPublisher="Records of the Zoological Survey of India" pageId="88" pageNumber="89" pagination="189 - 213" refId="B138" refString="Gupta, SK, Ghosh, SK, 1980. Some prostigmatid mites (Acarina) from Andaman and Nicobar Islands. Records of the Zoological Survey of India 77: 189 - 213" title="Some prostigmatid mites (Acarina) from Andaman and Nicobar Islands." volume="77" year="1980">Gupta and Ghosh 1980</bibRefCitation>
) and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gilbertoi">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides gilbertoi</emphasis>
</taxonomicName>
(Bashir and Afzal 2004) are transferred to
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Scutopalus</emphasis>
</taxonomicName>
as they posses 5 setae on the femurogenu instead of 6 as in
<taxonomicName authorityName="Smiley" authorityYear="1975" class="Arachnida" family="Cunaxidae" genus="Neocunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="36" pageNumber="37">Neocunaxoides</emphasis>
</taxonomicName>
and have well-demarcated plates.
</paragraph>
<paragraph lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37">
<table inLine="true" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37">
<tr pageId="36" pageNumber="37">
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">1</td>
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">
Coxae I-II faintly or totally divided (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="36" pageNumber="37">Fig. 40a, b</figureCitation>
)
</td>
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">2</td>
</tr>
<tr pageId="36" pageNumber="37">
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">-</td>
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">
Coxae I-II fused medially (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="36" pageNumber="37">Fig. 40c</figureCitation>
)
</td>
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">7</td>
</tr>
<tr pageId="36" pageNumber="37">
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">2 (1)</td>
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">
Coxae I-II faintly divided (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="36" pageNumber="37">Fig. 40a</figureCitation>
)
</td>
<td colspan="1" pageId="36" pageNumber="37" rowspan="1">3</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<pageBreakToken pageId="37" pageNumber="38" start="start">-</pageBreakToken>
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Coxae I-II totally divided (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 40b</figureCitation>
)
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">4</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">3 (2)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Sternal shield bearing 6 pairs of setae; setae
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
and
<emphasis italics="true" pageId="37" pageNumber="38">mps</emphasis>
simple; coxae II with 2 setae; basifemora I-IV setal formula 3-3-2-0; Greece
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Sionti &amp; Papadoulis" baseAuthorityYear="2003" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="abiesae">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus abiesae</emphasis>
</taxonomicName>
Sionti &amp; Papadoulis, 2003
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Sternal shield bearing 5 pairs of setae; setae
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
and
<emphasis italics="true" pageId="37" pageNumber="38">mps</emphasis>
setose; coxae II with 1 setae; basifemora I-IV setal formula 2-2-2-1; South Africa
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus arboreus</emphasis>
</taxonomicName>
Den Heyer, 1979
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">4 (2)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
At least 2 pairs of thick rod-like setae on the dorsum (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 41</figureCitation>
); India
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Gupta &amp; Ghosh" baseAuthorityYear="1980" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pradhani">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus pradhani</emphasis>
</taxonomicName>
(Gupta &amp; Ghosh, 1980)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">Rod-like setae on dorsal shield absent</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">5</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">5 (4)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">Coxae II with 2 sts</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">6</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">Coxae II with 3 sts; Pakistan</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Bashir &amp; Afzal" baseAuthorityYear="2004" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="gilbertoi">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus gilbertoi</emphasis>
</taxonomicName>
(Bashir &amp; Afzal, 2004)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">6 (4)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Setae
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
and
<emphasis italics="true" pageId="37" pageNumber="38">h1</emphasis>
on small platelets; ratio
<emphasis italics="true" pageId="37" pageNumber="38">c1</emphasis>
:
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
2:1; genua I with 4 asl, 5 sts; genua II with 2 asl, 5 sts; South Africa
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="latisetosus">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus latisetosus</emphasis>
</taxonomicName>
Den Heyer, 1979
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Setae
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
and
<emphasis italics="true" pageId="37" pageNumber="38">h1</emphasis>
on integument; ratio
<emphasis italics="true" pageId="37" pageNumber="38">c1</emphasis>
:
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
1:1; genua I with 3 asl, 5 sts; genua II with 1 asl, 5 sts; Greece
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Sionti &amp; Papadoulis" baseAuthorityYear="2003" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="smolikensis">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus smolikensis</emphasis>
</taxonomicName>
Sionti &amp; Papadoulis, 2003
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">7 (1)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Dorsal shield smooth and/or punctate (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 42a</figureCitation>
)
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">8</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Dorsal shield sparse granulate, rugose, or reticulate (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 42b-d</figureCitation>
)
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">12</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">8 (7)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">Coxae II and IV with 2 setae</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">9</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">Coxae II and IV with 3 setae</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">11</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">9 (8)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Setae
<emphasis italics="true" pageId="37" pageNumber="38">mps</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">d1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">e1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
clavate (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 43</figureCitation>
); a small subscutum situated posterior to the dorsal shield present; Malaysia
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Shiba" baseAuthorityYear="1978" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="clavatus">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus clavatus</emphasis>
</taxonomicName>
(Shiba, 1978)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Setae
<emphasis italics="true" pageId="37" pageNumber="38">mps</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">d1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">e1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
setiform; a small subscutum situated posterior to the dorsal shield absent
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">10</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">10 (9)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Setae
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
on dorsal shield; setae
<emphasis italics="true" pageId="37" pageNumber="38">lps</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">mps</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">d1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">e1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
set on tubercles (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 44</figureCitation>
); area between
<emphasis italics="true" pageId="37" pageNumber="38">pt</emphasis>
more heavily sclerotized, forming ridges; Taiwan
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Tseng" baseAuthorityYear="1980" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="osseus">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus osseus</emphasis>
</taxonomicName>
(Tseng, 1980)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
Setae
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
on integument; setae
<emphasis italics="true" pageId="37" pageNumber="38">lps</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">mps</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">c2</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">d1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">e1</emphasis>
,
<emphasis italics="true" pageId="37" pageNumber="38">f1</emphasis>
set normally; area between
<emphasis italics="true" pageId="37" pageNumber="38">pt</emphasis>
normally sclerotized, not forming ridges; Ukraine
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Kuznetzov &amp; Livshitz" baseAuthorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="trepidus">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus trepidus</emphasis>
</taxonomicName>
(Kuznetzov &amp; Livshitz, 1979)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">11 (8)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">4 pairs of hysterosomal setae around genital shield; long slender platelet laterad genital shield present; with a narrow transverse sclertie behind main shield; Philippines</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus philippinensis</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">3 pairs of hystersomal setae around genital shield; long slender platelet laterad genital shield absent; dorsal sclerites absent; Philippines</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="makapalus">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus makapalus</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">12 (7)</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
1 or more dorsal sclerites present (behind or laterad dorsal shield); dorsal shield rugose or reticulate (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="37" pageNumber="38">Fig. 42b, c</figureCitation>
); basifemora IV with 1 seta; pedipalpal tibiotarsus with 6 setae present and apophysis absent
</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">13</td>
</tr>
<tr pageId="37" pageNumber="38">
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">-</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">Dorsal sclerites absent; dorsal shield sparsely granulate; basifemora IV with 2 setae; pedipalpal tibiotarsus with 5 setae and a rod-shaped dorsal apophysis present; Taiwan</td>
<td colspan="1" pageId="37" pageNumber="38" rowspan="1">
<taxonomicName baseAuthorityName="Tseng" baseAuthorityYear="1980" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="unguianalis">
<emphasis italics="true" pageId="37" pageNumber="38">Scutopalus unguianalis</emphasis>
</taxonomicName>
(Tseng, 1980)
</td>
</tr>
<tr pageId="38" pageNumber="39">
<td colspan="1" pageId="38" pageNumber="39" rowspan="1">
<pageBreakToken pageId="38" pageNumber="39" start="start">13</pageBreakToken>
(12)
</td>
<td colspan="1" pageId="38" pageNumber="39" rowspan="1">
Dorsal shield rugose (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="38" pageNumber="39">Fig. 42b</figureCitation>
); setae
<emphasis italics="true" pageId="38" pageNumber="39">f1</emphasis>
and
<emphasis italics="true" pageId="38" pageNumber="39">h1</emphasis>
on integument; dorsal setae (except
<emphasis italics="true" pageId="38" pageNumber="39">c2</emphasis>
and
<emphasis italics="true" pageId="38" pageNumber="39">h2</emphasis>
) distally rod-like (slightly clavate), with minute barbs; narrow transverse shield behind main dorsal shield present; Philippines
</td>
<td colspan="1" pageId="38" pageNumber="39" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis italics="true" pageId="38" pageNumber="39">Scutopalus rugosus</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="38" pageNumber="39">
<td colspan="1" pageId="38" pageNumber="39" rowspan="1">-</td>
<td colspan="1" pageId="38" pageNumber="39" rowspan="1">
Dorsal shield reticulate (
<figureCitation captionStart="Figures 40–44" captionStartId="F13" captionText="Figures 40 - 44. Scutopalus key illustrations. 40 a Coxae I-II faintly divided 40 b Coxae I-II totally divided 41 Coxae I-II fused medially 42 Dorsal shield with thick, rod-like setae present 43 Dorsal shield smooth or punctate 44 a Dorsal shield rugose 44 b Dorsal shield reticulate 44 c Dorsal shield sparsely granulate 45 a Setae mps, c 1, c 2, d 1, e 1, f 1 clavate 45 b Setae mps, c 1, c 2, d 1, e 1, f 1 setiform 46 Setae lps, mps, c 1, c 2, d 1, e 1, f 1 set on tubercles." httpUri="https://binary.pensoft.net/fig/29887" pageId="38" pageNumber="39">Fig. 42c</figureCitation>
); setae
<emphasis italics="true" pageId="38" pageNumber="39">f1</emphasis>
and
<emphasis italics="true" pageId="38" pageNumber="39">h1</emphasis>
on small platelets; dorsal setae (except
<emphasis italics="true" pageId="38" pageNumber="39">c2</emphasis>
and
<emphasis italics="true" pageId="38" pageNumber="39">h2</emphasis>
) broad and serrate; sclerites laterad and behind dorsal shield present; Brazil
</td>
<td colspan="1" pageId="38" pageNumber="39" rowspan="1">
<taxonomicName authorityName="Rocha, Skvarla &amp; Ferla" authorityYear="2013" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tomentosus">
<emphasis italics="true" pageId="38" pageNumber="39">Scutopalus tomentosus</emphasis>
</taxonomicName>
Rocha, Skvarla &amp; Ferla, 2013
</td>
</tr>
</table>
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/29887" pageId="39" pageNumber="40" start="Figures 40–44" startId="F13">
<paragraph pageId="39" pageNumber="40">
<emphasis bold="true" pageId="39" pageNumber="40">
<pageBreakToken pageId="39" pageNumber="40" start="start">Figures</pageBreakToken>
40-44.
</emphasis>
<taxonomicName authorityName="Den Heyer" authorityYear="1979" class="Arachnida" family="Cunaxidae" genus="Scutopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="39" pageNumber="40">Scutopalus</emphasis>
</taxonomicName>
key illustrations.
<emphasis bold="true" pageId="39" pageNumber="40">40a</emphasis>
Coxae I-II faintly divided
<emphasis bold="true" pageId="39" pageNumber="40">40b</emphasis>
Coxae I-II totally divided
<emphasis bold="true" pageId="39" pageNumber="40">41</emphasis>
Coxae I-II fused medially
<emphasis bold="true" pageId="39" pageNumber="40">42</emphasis>
Dorsal shield with thick, rod-like setae present
<emphasis bold="true" pageId="39" pageNumber="40">43</emphasis>
Dorsal shield smooth or punctate
<emphasis bold="true" pageId="39" pageNumber="40">44a</emphasis>
Dorsal shield rugose
<emphasis bold="true" pageId="39" pageNumber="40">44b</emphasis>
Dorsal shield reticulate
<emphasis bold="true" pageId="39" pageNumber="40">44c</emphasis>
Dorsal shield sparsely granulate
<emphasis bold="true" pageId="39" pageNumber="40">45a</emphasis>
Setae
<emphasis italics="true" pageId="39" pageNumber="40">mps</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">c1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">c2</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">d1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">e1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">f1</emphasis>
clavate
<emphasis bold="true" pageId="39" pageNumber="40">45b</emphasis>
Setae
<emphasis italics="true" pageId="39" pageNumber="40">mps</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">c1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">c2</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">d1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">e1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">f1</emphasis>
setiform
<emphasis bold="true" pageId="39" pageNumber="40">46</emphasis>
Setae
<emphasis italics="true" pageId="39" pageNumber="40">lps</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">mps</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">c1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">c2</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">d1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">e1</emphasis>
,
<emphasis italics="true" pageId="39" pageNumber="40">f1</emphasis>
set on tubercles.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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