267 lines
29 KiB
XML
267 lines
29 KiB
XML
<document id="2B97A2BF776EC8DA5C2B5373285C5E95" ID-CLB-Dataset="35978" ID-DOI="http://dx.doi.org/10.3897/zookeys.637.9384" ID-GBIF-Dataset="7cc290d2-0f7b-4a0c-9e87-8023da6b5418" ID-PMC="PMC5240120" ID-Pensoft-Pub="1313-2970-637-33" ID-PubMed="28138273" ID-ZooBank="401C312B74D04FC8AD063191A6E29979" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-637-33" ModsDocOrigin="ZooKeys 637" ModsDocTitle="Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula" checkinTime="1480398151478" checkinUser="pensoft" docAuthor="Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki & Park, Sun-Jae" docDate="2016" docId="C8FCCFC2D8533E2F4BC6BFE9A606810F" docLanguage="en" docName="ZooKeys 637: 33-45" docOrigin="ZooKeys 637" docSource="http://dx.doi.org/10.3897/zookeys.637.9384" docTitle="Ptomaphagus hayashii Wang, Ruzicka, Perreau, Nishikawa & Park, 2016, sp. n." docType="treatment" docUuid="C468947A-D211-4411-B09B-775DFF6A7045" docUuidSource="ZooBank" docVersion="10" lastPageNumber="38" masterDocId="FF89FF87FFCBFF82FFC0FFDEFF8FA166" masterDocTitle="Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula" masterLastPageNumber="45" masterPageNumber="33" pageNumber="34" updateTime="1731874387632" updateUser="admin">
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<mods:title id="85657BA533E8992ED4B1FEBBAB591893">Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula</mods:title>
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<mods:namePart id="34E5A7B229509E59E800B7918E89962C">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="F7B4B51F1806BD78B3E385273C73D828">Ruzicka, Jan</mods:namePart>
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<mods:namePart id="5EA84DDDB7C8E14704CBF720ECCB05B0">Perreau, Michel</mods:namePart>
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<mods:namePart id="CE684A3E42C02686EB4BD0B882B60CF8">Nishikawa, Masaaki</mods:namePart>
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<mods:namePart id="CF948ED1FAC0B1ABA4E0A2CCEB9FE213">Park, Sun-Jae</mods:namePart>
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<mods:title id="BBF28348D37D4595B1B88ED8F3158EC7">ZooKeys</mods:title>
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<mods:date id="8879645218FBB9FAAB3D586BF4B6A091">2016</mods:date>
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<mods:number id="4894A25AC1C51B3E9780B2FB0BA3935C">637</mods:number>
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<mods:identifier id="1F97720590941FB188BE0B74AC9D78E7" type="DOI">http://dx.doi.org/10.3897/zookeys.637.9384</mods:identifier>
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<treatment id="C8FCCFC2D8533E2F4BC6BFE9A606810F" ID-GBIF-Taxon="127857844" LSID="urn:lsid:zoobank.org:act:C468947A-D211-4411-B09B-775DFF6A7045" httpUri="http://treatment.plazi.org/id/C8FCCFC2D8533E2F4BC6BFE9A606810F" lastPageId="5" lastPageNumber="38" pageId="1" pageNumber="34">
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<subSubSection id="017C7157212CD556CDE9C82FF654D849" pageId="1" pageNumber="34" type="nomenclature">
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<paragraph id="5112F63838660F7A0F722B7FA3B712D8" pageId="1" pageNumber="34">
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<taxonomicName id="C3C91ABA3E15E430EB21CB27852F8F89" ID-CoL="4Q9F6" LSID="http://zoobank.org/C468947A-D211-4411-B09B-775DFF6A7045" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus hayashii" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
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<taxonomicNameLabel id="4DEE1BC1F5934F492B29C1438866449B" pageId="1" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
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Figs 1B, C; 2; 3
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</paragraph>
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</subSubSection>
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<subSubSection id="C7D28C53E8A6464F58DF6E41453E0DC1" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="type material">
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<paragraph id="E92DF7FE043599689B3EA36AF8193B4B" pageId="1" pageNumber="34">Type material.</paragraph>
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<paragraph id="BD92BE3DA036159E57D3C1E412690A55" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
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Holotype: 1♂, USSR: Sadgorod [ca.
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<geoCoordinate id="C42FB1B7D6CD74F4C5ABD5288E544BD5" direction="north" orientation="latitude" precision="925" value="43.25">43°15'N</geoCoordinate>
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<geoCoordinate id="06A49A5A8F4FF0190AD360EC8C1C631D" direction="east" orientation="longitude" precision="925" value="132.05">132°03'E</geoCoordinate>
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] / (in forest; trap with / bait), Vladivostok // Primorskyi Kray / 16.VI.1978 / E. Berlov leg. (NSMT). Paratypes: 1♂1♀, FE.Russia, SW Khabarovsky / kray reg.,
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<normalizedToken id="C40A636E1527609D1275C07833110C69" originalValue="Strel’nikova">Strel'nikova</normalizedToken>
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/ range Mts.,
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<geoCoordinate id="2558F20685435FCBBA793ACA88B46BC8" direction="north" orientation="latitude" precision="925" value="46.716667">46°43'N</geoCoordinate>
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<geoCoordinate id="768BA55CCA14C55E739D1B29BE61EA7D" direction="east" orientation="longitude" precision="925" value="134.13333">134°08'E</geoCoordinate>
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/ Samur Riv., 130-550 m alt. / 25.VI.-28.VII.2014 / A. Plutenko leg. (CMNE); 1♀, same data as holotype (CMNE); 1♀, USSR Ussuri reg. / NOVOVARVAROVKA [ca.
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<geoCoordinate id="12CD9867246DE669417C7E0FCB03ECFF" direction="north" orientation="latitude" precision="925" value="43.966667">43°58'N</geoCoordinate>
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<geoCoordinate id="FDC87CC874B70B9330EE087470BEE0D2" direction="east" orientation="longitude" precision="925" value="132.98334">132°59'E</geoCoordinate>
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] / 6-10.VII.1989 / R.
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<normalizedToken id="CDC49BFF4EB547F95D5408D10A02945B" originalValue="Červenka">Cervenka</normalizedToken>
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lgt. //
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<taxonomicName id="5053BA93DFB5EF0887C194CDE44FAE3A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(s. str.) /
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<taxonomicName id="15B54690EC12DAFA2D2C655848897F05" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="sibiricus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="sibiricus">sibiricus</taxonomicName>
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/ Jeannel, 1934 / Jan
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<normalizedToken id="6681632577B2C2263AC2890E073D92CD" originalValue="Růžička">Ruzicka</normalizedToken>
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det. 2001 // ♀ (CJRZ); 1♂1♀, USSR Ussuri reg. / JASNOE [= Yasnoye, ca.
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<geoCoordinate id="6189C1705F3A5E4152CD25A31EE28F28" direction="north" orientation="latitude" precision="925" value="43.45">43°27'N</geoCoordinate>
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<geoCoordinate id="613528F110FC4212504009B9B7FE3B2C" direction="east" orientation="longitude" precision="925" value="132.15">132°09'E</geoCoordinate>
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] VII.1989 / St.
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<normalizedToken id="C731FA6526C4F1D8907DB1BA124F76DB" originalValue="Bečvář">Becvar</normalizedToken>
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lgt. //
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<taxonomicName id="2CA465B2A616531CF467424D793B9E86" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(s. str.) /
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<taxonomicName id="4942F90BF108F50A0537076A40629374" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="sibiricus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="sibiricus">sibiricus</taxonomicName>
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/ Jeannel, 1934 / Jan
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<normalizedToken id="DC9B15691EB737C077DF9F99DE867BD0" originalValue="Růžička">Ruzicka</normalizedToken>
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det. 2001 (CJRZ); 1♂1♀, USSR Ussuri reg. / JASNOE [= Yasnoye] / VII.1989 12-16 / St.
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<pageBreakToken id="6693BCE089EB0B38406CD5AFE6001593" pageId="2" pageNumber="35" start="start">lgt</pageBreakToken>
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. // MOUNT. OBLATCHNAJA [= Oblachnaya] / 400-800 m (CJRZ); 1♀, RUSSIA, Far East: / S Primorye region, / LAZO env. [ca.
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<geoCoordinate id="4833D3623454472FF603862F4A291F9B" direction="north" orientation="latitude" precision="925" value="43.366665">43°22'N</geoCoordinate>
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<geoCoordinate id="1AD0C8D46A2892705BA15D7FAE86C8B2" direction="east" orientation="longitude" precision="925" value="133.9">133°54'E</geoCoordinate>
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], 2.VII. / 2002 //
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<taxonomicName id="2F7B5CF33B909C4E2905F821114C64CB" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(s. str.) /
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<taxonomicName id="4C7804E472431DE39E0E6F9CCE113ED4" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="sibiricus" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="sibiricus">sibiricus</taxonomicName>
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/ Jeannel, 1934 / Jan
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<normalizedToken id="D41151AFE6A3A2E4937DEA7E45847C77" originalValue="Růžička">Ruzicka</normalizedToken>
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det 2009 / ♀ (CJRZ); 1♂, RUSSIA: Primorsky / 30 km NE Vladivostok / Tajvaza / 29.
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<normalizedToken id="3A2B6AE03F4CDBBDB3BADC4CBABCD356" originalValue="VII–">VII-</normalizedToken>
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5.VIII.1992 / B. D. Gill [leg.] (CJRZ); 1♂, Russian Far East, Primorskii krai / Lazovski Zapovednik, c. 170 km E / Vladivostok, Korpad, 1.-14.v.2001 / 175 m;
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<geoCoordinate id="CD4B5161DCEBD8558B9F02D844F00772" direction="north" orientation="latitude" precision="15" value="43.263054">43°15'47"N</geoCoordinate>
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<geoCoordinate id="A662A7CBC24847D3D19EF2E640E37B38" direction="east" orientation="longitude" precision="15" value="134.12889">134°07'44"E</geoCoordinate>
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/ floodplain of Priamushka / Malaise trap 440; / M. Quest coll. BMNH (E) 2009-59 (BMNH); 1♀, same data as previous except: 13.-31.vii.2001 /
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<geoCoordinate id="924CD3AFE08001F3137312B53C97EEA5" direction="north" orientation="latitude" precision="15" value="43.264446">43°15'52"N</geoCoordinate>
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<geoCoordinate id="6B29B84D6C81FB4ADA2BC19C074D04DF" direction="east" orientation="longitude" precision="15" value="134.12917">134°07'45"E</geoCoordinate>
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; 174 m; / Mountain top, Malaise trap 481 (BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="299D6994EC14DCB32A57CD1A224A6FDA" pageId="3" pageNumber="36" type="description">
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<paragraph id="9F50B4379A7E077CF66DD0163CCFEC44" pageId="3" pageNumber="36">
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<pageBreakToken id="EDB71575A3BC06F6B02321C5760B89B7" pageId="3" pageNumber="36" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph id="7ED33EE04356F255E9088C7E9E13E66A" pageId="3" pageNumber="36">
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Male. EBL: 3.8-4.3 mm (4.2 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.7: 1.1: 1.1: 2.3 mm; width: HW: EW: PW: ELW = 1.1: 0.1: 1.6: 1.8 mm. Proportion of antennomeres from base to tip in
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<normalizedToken id="52B632BD433C0D388924097E1DFEA3CA" originalValue="μm">μm</normalizedToken>
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(length
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<normalizedToken id="4293E593789F514A04E01FFC9926A510" originalValue="×">x</normalizedToken>
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width): 149
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75, 109
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67, 95
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70, 64
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79, 58
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<normalizedToken id="8E348C10C01D730DE017A6619C1AF8C8" originalValue="×">x</normalizedToken>
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92, 41
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<normalizedToken id="ECBE5344C69C755178E038D1336B04C2" originalValue="×">x</normalizedToken>
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100, 91
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<normalizedToken id="521D2AD240C8164746C1FC1E7187BDAD" originalValue="×">x</normalizedToken>
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128, 39
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<normalizedToken id="869AD5BB2CBAF573DFA8BF9836363EBE" originalValue="×">x</normalizedToken>
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128, 87
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<normalizedToken id="347F03BCF2829DBE9D8E7A418FEE6A70" originalValue="×">x</normalizedToken>
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151, 90
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<normalizedToken id="E1AD997DDDE7CEC955AB5B26F2CE14A7" originalValue="×">x</normalizedToken>
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156, 161
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<normalizedToken id="BDABA457BFD617267E9CE3CB59C5442E" originalValue="×">x</normalizedToken>
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143.
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</paragraph>
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<paragraph id="F01CC3EBFD19EB7BF84B5FBB65F56659" pageId="3" pageNumber="36">Habitus (Fig. 1B) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, antennae (apical half of ultimate antennomere yellowish) and tarsi reddish brown. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
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<caption id="1211AE70008E9E77FB697C4D7BECBB02" pageId="3" pageNumber="36">
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<paragraph id="333F8DD59B26D0CD9CCC19A658C62183" pageId="3" pageNumber="36">
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Figure 1. Habitus of
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<taxonomicName id="9527CF76CF4A8A8D1C2493DB8DDF2255" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">Ptomaphagus (s. str.) spp.</taxonomicName>
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(dorsal view). A,
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<normalizedToken id="491D06A90C869C586A7CB9B346F30883" originalValue="D–F">D-F</normalizedToken>
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<taxonomicName id="4E5602ABC3EC76AD7729847E33377195" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus Jeannel, 1934</taxonomicName>
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A ♀ (holotype; Vladivostok) D ♂ (Vladivostok) E ♂ (Pyeongchang-gun) F ♀ (Boeun-gun) B, C
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<taxonomicName id="4B0718F1D15DDE40A93EA7330377E248" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
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<taxonomicNameLabel id="44AAC9658886C472E7628EB7AB1DC65B" rank="species">sp. n.</taxonomicNameLabel>
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B ♂ (holotype; Vladivostok) C ♀ (paratype; Ussuri region). Scale: 1 mm.
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</paragraph>
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</caption>
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<paragraph id="D52D8A59C6ABBC9364A3970F9545B194" pageId="3" pageNumber="36">Head transverse, HW/HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.11. Antennae (Fig. 2A) slender, AL/HW = 1.0; antennomere III shorter than II; VI with length/width = 0.4; XI peach-shape.</paragraph>
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<caption id="4B630E847518681C26469F378E441095" pageId="3" pageNumber="36">
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<paragraph id="302DD9545BDADCA2FDC6F08E6D60504D" pageId="3" pageNumber="36">
|
||
Figure 2.
|
||
<taxonomicName id="1AF488DCD5AC97D5F1BC29AE3DE7692F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="C0A00D3C945D4519852B0648A6E377FF" rank="species">sp. n.</taxonomicNameLabel>
|
||
(♂: holotype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scales: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="89F1006B2AD06F9738EC010B1B268AE1" pageId="3" pageNumber="36">Pronotum (Fig. 2B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, simply narrowing from posterior to anterior; hind angles slightly projected backwards and acute. Posterior margin widely protruded in middle part, emarginate near hind angles.</paragraph>
|
||
<paragraph id="35A633AAF3BD552387C6224B5DA14478" pageId="3" pageNumber="36">Elytra oval, widest at about basal 1/4, ELL/EW = 1.3. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 2G) narrowly rounded. Sutural striae present. Metathoracic wings fully developed.</paragraph>
|
||
<paragraph id="D5AA733EF8DB7BE596303425B4C6796C" pageId="3" pageNumber="36">Prolegs robust, with basal three protarsomeres (Fig. 2C) strongly expanded: TW/BTW = 1.2. Protibiae (Fig. 2E) strongly expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
|
||
<paragraph id="C8E3006562F9480A92AF46395B803264" pageId="3" pageNumber="36">Abdominal ventrite VIII (Fig. 2I) round at posterior edge and with an inconspicuous median notch. Spiculum gastrale (Fig. 2J) of genital segment with about 1/5 of length protruding beyond anterior edge of epipleurite IX.</paragraph>
|
||
<paragraph id="F88A5AEB587FF454F4B9E3E3A679529E" pageId="3" pageNumber="36">Aedeagus (Fig. 3A) long and slender, with median lobe gradually narrowing towards a lanceolate apex and terminated to a widely rounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 3C) inserted with 5 ventrally oriented setae on both sides; parameres narrow, reaching about apical 1/5 of median lobe, each apex (Fig. 3D) with 2 lateral setae and 1 apical seta distinctly shorter. In lateral view (Fig. 3B), median lobe regularly bent ventrad and gradually tapering to a flat apex. Endophallus with stylus quite slender, a subelliptical nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.</paragraph>
|
||
<caption id="4E9F578E65CCD173F03FFE51E131FB92" pageId="3" pageNumber="36">
|
||
<paragraph id="EFD4E902EC3F4D2F2733C7ADD4435EE7" pageId="3" pageNumber="36">
|
||
Figure 3.
|
||
<taxonomicName id="C2201254412A72A0E513E08918E48384" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="7B45EC56087B9D889BED1654D3F05FEE" rank="species">sp. n.</taxonomicNameLabel>
|
||
(♂: holotype). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scales: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9377C57EADA110CC8E6AFA5778762CBF" pageId="3" pageNumber="36">Female. Similar to male in general appearance (Fig. 1C), including elytral apices (Fig. 2H), but distinguished by the following characteristics: protarsi (Fig. 2D) simply linear; protibiae (Fig. 2F) narrower; abdominal ventrite VIII (Fig. 3E) narrowly rounded at posterior edge; genital segment and ovipositor as shown in Fig. 3F; spermatheca (Fig. 3F) curved in distal part and coiled in proximal part.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BC1477D7472EB63EA20296505BDE13D0" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37" type="diagnosis">
|
||
<paragraph id="F6CA9015A541464C3CF43E232475D200" pageId="4" pageNumber="37">
|
||
<pageBreakToken id="72A7ABF0255CA73D23B2012087EE7A8E" pageId="4" pageNumber="37" start="start">Diagnosis</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="76EE9D151812EE4C4316059258B0C858" pageId="4" pageNumber="37">
|
||
<taxonomicName id="FF52FF89AD07B4849637C03C6B36AEFA" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
|
||
is sympatric with
|
||
<taxonomicName id="FD9549191F147BA279C3BE921D6F5496" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="B94F53DC733BAD602E6CE05E93B70B2F" rank="species">sp. n.</taxonomicNameLabel>
|
||
in some locations of the Russian Far East, and they are very similar to each other (including spermatheca (Figs 3F; 5F), which is only more roundly curved in the stem part in
|
||
<taxonomicName id="737ACF468E69EFE00194708F3F8A0F6F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="5FB1E5940CA94AD1F2A85FD06142C777" rank="species">sp. n.</taxonomicNameLabel>
|
||
). For external characters, the body size of
|
||
<taxonomicName id="956CEE881609848C5DE6BF3FCD064275" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
|
||
(Figs 1A,
|
||
<normalizedToken id="1BC129C70B4F2DAF79ED1BEFD83BA0FE" originalValue="D–F">D-F</normalizedToken>
|
||
) is a little smaller than
|
||
<taxonomicName id="25B3B00989469819C7ED11A6A5B4711B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="510BC363946A3A41DF9200A20AEE88AD" rank="species">sp. n.</taxonomicNameLabel>
|
||
(Figs 1B, C). However, their aedeagi provide critical characters to distinguish the two species: in
|
||
<taxonomicName id="1685E37C7BADD2A3AD4EE41BEBF0C2C1" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="1CF8BEE34E7DF6C79984D7E3D6839333" rank="species">sp. n.</taxonomicNameLabel>
|
||
, the aedeagus is much larger and more slender (Fig. 3A), the right apicoventral piece of the median lobe is slenderly lanceolate (Fig. 3C), the apical half of median lobe much flatter in lateral view (Fig. 3B); while in
|
||
<taxonomicName id="69C64A728579EDC044494BB6C8D9DE06" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
|
||
, the aedeagus is stouter (Fig. 5A), the right apicoventral piece of median lobe is much wider and subpentagonal (Fig. 5C), the apical half of the median lobe thicker in lateral view (Fig. 5B).
|
||
</paragraph>
|
||
<caption id="D291266382DEBC481CF8051B2444CB0E" pageId="4" pageNumber="37">
|
||
<paragraph id="1DB64A54880380C8700BF02C6FF6B1D1" pageId="4" pageNumber="37">
|
||
Figure 4.
|
||
<taxonomicName id="6E7897AAAB07B24071AA4A91AEF92661" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus Jeannel, 1934</taxonomicName>
|
||
(Vladivostok). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scales: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="292CC59BC9ED570F77B1C370DD2173F6" pageId="4" pageNumber="37">
|
||
<paragraph id="44CA9B1613ACBDAC87002C7E67C80915" pageId="4" pageNumber="37">
|
||
Figure 5.
|
||
<taxonomicName id="214261599E61F564EBC557F8A45BB86E" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus Jeannel, 1934</taxonomicName>
|
||
(Vladivostok). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scales: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="87A205672AF252BE5F6976B9AB55488F" pageId="5" pageNumber="38">
|
||
<pageBreakToken id="AC7D1FFE0C5C5BD35EE39754377E22DA" pageId="5" pageNumber="38" start="start">Furthermore</pageBreakToken>
|
||
, based on specimens examined,
|
||
<taxonomicName id="EB99BD009659B21E0A7823B0A1BD86C7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
|
||
seems to be much more widely distributed, extending southward to South Korea; while
|
||
<taxonomicName id="297E91393140618E90F3CA4C5B18ACE2" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
|
||
<taxonomicNameLabel id="9717E2B37784069712F812E3F48E32E3" rank="species">sp. n.</taxonomicNameLabel>
|
||
is presently known only in the Russian Far East.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B9C5D362C60B45F3D6CD2AA454278884" pageId="5" pageNumber="38" type="etymology">
|
||
<paragraph id="44EECF5F0391AD6DA34F64F7549179C5" pageId="5" pageNumber="38">Etymology.</paragraph>
|
||
<paragraph id="4B560A3669CB5E5FFF58BE0400771DAF" pageId="5" pageNumber="38">
|
||
The specific epithet is dedicated to Mr. Yasuhiko Hayashi (Kawanishi, Japan), a famous independent taxonomist on
|
||
<taxonomicName id="F7D8AA7EAF411AB04AF2796832E1F97B" class="Insecta" kingdom="Animalia" lsidName="Staphylinoidea" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="superFamily" superFamily="Staphylinoidea">Staphylinoidea</taxonomicName>
|
||
, for his continual generous help to our study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9670C6098D94077270F8D26A4FA4361C" pageId="5" pageNumber="38" type="distribution">
|
||
<paragraph id="3827E36AAC1E8F90F95980FDB3126CD8" pageId="5" pageNumber="38">Distribution.</paragraph>
|
||
<paragraph id="580AF0B2E9515491243182F47F30FFF4" pageId="5" pageNumber="38">Russia (Far East) (Fig. 6).</paragraph>
|
||
<caption id="720B2B8CE1A6E67266FE3A4B2A07763E" pageId="5" pageNumber="38">
|
||
<paragraph id="FA7225B46FF3D1481CE2DEEE28397FA7" pageId="5" pageNumber="38">
|
||
Figure 6. Distribution map of
|
||
<taxonomicName id="127713778AB5096751DF16547082D331" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
|
||
species from the Russian Far East and the Korean Peninsula.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |