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Figs 1A,
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; 4; 5
@ -68,37 +68,38 @@ Figs 1A,
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(s. str.) sibiricus
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<taxonomicName id="761BB91B1B4A4A62D866BF454BC3C5AA" authority="Jeannel 1934: 165" authorityName="Jeannel" authorityPageNumber="165" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus sibiricus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="sibiricus">
<pageBreakToken id="ADD589A2EC095FD67EC06AB4133A146E" pageId="6" pageNumber="39" start="start">Ptomaphagus</pageBreakToken>
(s. str.) sibiricus
<bibRefCitation id="28CF1FD325F6EA26365F622634E1AF3D" author="Jeannel, R" journalOrPublisher="Revue Francaise d'Entomologie" pageId="11" pageNumber="44" pagination="161 - 170" title="Les Ptomaphagus palearctiques (Col.)." volume="1" year="1934">Jeannel 1934</bibRefCitation>
: 165 (
<taxonomicName id="CDF021FC29D0942416BC968CA83A8B18" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
: 165
</taxonomicName>
(
<taxonomicName id="CDF021FC29D0942416BC968CA83A8B18" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); type locality: [RUSSIA, Far East] Wladiwostok; SDEI);
<bibRefCitation id="FBAD41962F0703F5818F7C3B9757BED7" author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle (N. S.)" pageId="11" pageNumber="44" pagination="1 - 433" title="Monographie des Catopidae (Insectes Coleopteres)." volume="1" year="1936">Jeannel 1936</bibRefCitation>
: 72, 84 (
<taxonomicName id="0856BB9AD61EB5E2331532DB119A4E48" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="0856BB9AD61EB5E2331532DB119A4E48" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); in key; distribution);
<bibRefCitation id="3088C41BCF29B54151A6FC4DED97ADD2" author="Nishikawa, M" journalOrPublisher="Coleopterists' Association of Japan, Tokyo" pageId="11" pageNumber="44" pagination="1 - 6" title="Family Catopidae." volumeTitle="Check-list of Coleoptera of Japan" year="1983">Nishikawa 1983</bibRefCitation>
: 1 (
<taxonomicName id="49D7154683F1D3965EB2624B5EC26D32" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="49D7154683F1D3965EB2624B5EC26D32" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in check-list);
<bibRefCitation id="0796C375E98534DD8B602DC03563064B" author="Perreau, M" journalOrPublisher="Memoires de la Societe entomologique de France" pageId="11" pageNumber="44" pagination="1 - 460" title="Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae." volume="4" year="2000">Perreau 2000</bibRefCitation>
: 364 (
<taxonomicName id="EFFF213047CCDC69C5D713DF573ADE14" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(s. str.); in catalogue);
<bibRefCitation id="64D2B1A87DFFBCFF20D4DC44B3FAEB64" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="11" pageNumber="44" pagination="133 - 203" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2: Hydrophiloidea - Histeroidea - Staphylinoidea" year="2004">Perreau 2004</bibRefCitation>
: 178 (
<taxonomicName id="934D1DDAD1D0CDABB81DD8D98D0FF537" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="934D1DDAD1D0CDABB81DD8D98D0FF537" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue);
<bibRefCitation id="CFFD9153AC5B8BD80A45C743466024F9" author="Zinchenko, VK" journalOrPublisher="Evraziatskii Entomologicheskii Zhurnal" pageId="11" pageNumber="44" pagination="339 - 340" title="Notes on the occurrence of round fungus beetles (Leiodidae, Cholevinae) in the Russian Far East." volume="7" year="2008">Zinchenko and Lyubechanskii 2008</bibRefCitation>
: 340 (
<taxonomicName id="8A44F60B8B55BDB23BF691BD6DF2AA2C" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="8A44F60B8B55BDB23BF691BD6DF2AA2C" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; distribution);
<bibRefCitation id="784D74271C8BD79F87B0F673176ADD24" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Brill, Leiden" pageId="11" pageNumber="44" pagination="180 - 290" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Hydrophiloidea - Staphylinoidea" year="2015">Perreau 2015</bibRefCitation>
: 249 (
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<taxonomicName id="E537A85176734399F67BC83D409E7A92" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue).
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@ -109,8 +110,8 @@ Type material.Holotype: ♀, [RUSSIA, Far East] Wladiwostok [ca.
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<geoCoordinate id="C282F3A35454F52DB88CA6DDAC384AAA" direction="east" orientation="longitude" precision="925" value="132.0">132°00'E</geoCoordinate>
] // Reitter // Coll. Koltze // Pt. variicornis / Rosenh. //
<taxonomicName id="E309D491C1CFAEB22DB639A11C93D89F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus sibiricus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus sibiricus</taxonomicName>
Jeann. / type / R. Jeannel det. // DEI
<taxonomicName id="BA2E9FCD4874C05CD104C90DB403FB65" authority="Jeann." authorityName="Jeann." class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus sibiricus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus sibiricus Jeann.</taxonomicName>
/ type / R. Jeannel det. // DEI
<normalizedToken id="49DAF505DF3EA7E037CC67885A40870D" originalValue="Müncheberg">Muencheberg</normalizedToken>
/ Col - 07069 (SDEI).
</paragraph>
@ -124,13 +125,11 @@ Additional material.RUSSIA, Far East: 1♀, FE.Russia, SW Khabarovsky / kray reg
<geoCoordinate id="9B340C777A683FC51F0386BDA1C18C26" direction="north" orientation="latitude" precision="925" value="43.166668">43°10'N</geoCoordinate>
<geoCoordinate id="3BF673772B63050862F87675D8B51FCE" direction="east" orientation="longitude" precision="925" value="132.0">132°00'E</geoCoordinate>
] / Christov [leg.] IX.[18]76 //
<taxonomicName id="4ED2C64D9593F3A0B5C75DD88A95BFDD" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus sibiricus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus sibiricus</taxonomicName>
<taxonomicName id="4ED2C64D9593F3A0B5C75DD88A95BFDD" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus sibiricus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus sibiricus</taxonomicName>
/ Jeannel det. //
<taxonomicName id="2319C74519F8F235E6C362D664C8C92B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/ (
<taxonomicName id="BCEDEFD15FD000AB985660FFB2648A2F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
)
<taxonomicName id="358A7B5AB652B37EBCE1D0E0088233AB" class="Magnoliopsida" family="Malvaceae" genus="Staphylinoidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Staphylinoidea sibiricus" order="Malvales" pageId="6" pageNumber="39" phylum="Tracheophyta" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="2319C74519F8F235E6C362D664C8C92B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/
<taxonomicName id="9E94A4EB345B594AE1E87E6696B0CA0A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="sibiricus" subGenus="Ptomaphagus">(Ptomaphagus) sibiricus</taxonomicName>
/ Jeannel, 1934 / Ex. M. Nishikawa, 2008 / # MNHN 103377Ch1S ♀ (MNHN); 10♂♂10♀♀, RUSSIA: Primorsky / 30 km NE Vladivostok / Tajvaza / 29.
<normalizedToken id="CCE3967EDB380651E4899FB566FBD75A" originalValue="VII">VII-</normalizedToken>
5.VIII.1992 / B. D. Gill [leg.] (CJRZ); 1♀, RUSSIA: Primoryi / Nakhodka [ca.
@ -144,19 +143,19 @@ Additional material.RUSSIA, Far East: 1♀, FE.Russia, SW Khabarovsky / kray reg
,
<geoCoordinate id="8BBC898154742C105DF05329598F0979" direction="east" orientation="longitude" precision="555" value="132.34">132.34°E</geoCoordinate>
, poch.l. [= pitfall trap], V. Zinchenko &amp; A. Korshunov [leg.], 9.-19.viii.2011 //
<taxonomicName id="A016910FA6B1BA64836EE5F8F0D5AF83" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="A016910FA6B1BA64836EE5F8F0D5AF83" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/
<taxonomicName id="C0C999F29E49A07CFDCCBBDA4A095FA0" class="Magnoliopsida" family="Malvaceae" genus="Staphylinoidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Staphylinoidea sibiricus" order="Malvales" pageId="6" pageNumber="39" phylum="Tracheophyta" rank="species" species="sibiricus">sibiricus</taxonomicName>
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/ V. Zinchenko det. 2002 (CCBW). South Korea: 1♂, KOREA: Gangwon Prov., / Pyeongchang-gun, Jinbu-myeon, / Dongsan-ri, Mt. Odaesan, Sangwonsa [ca.
<geoCoordinate id="6857A895C2F525CD2AA0FB6EAD4E0294" direction="north" orientation="latitude" precision="925" value="37.783333">37°47'N</geoCoordinate>
<geoCoordinate id="71A340E5369C2916DB0ECAD6714DB723" direction="east" orientation="longitude" precision="925" value="128.55">128°33'E</geoCoordinate>
], / 4.VI.-22.VI.2001, K.-J. Ahn, S.-J. / Park, M.-S. Kim, M.-J. Jeon [leg.], ex FIT //
<taxonomicName id="F53E6ACBFA8850160E8532F3334B00F6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="F53E6ACBFA8850160E8532F3334B00F6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/ baekamsanensis / n. sp. / det. S.-J. Park 2005 // 1 (CNUIC); 1♂, KOREA: Gangwon Prov., / Hongcheon-gun [ca.
<geoCoordinate id="26A40F0C1C10FD7C693C45D20B32329B" direction="north" orientation="latitude" precision="925" value="37.683334">37°41'N</geoCoordinate>
<geoCoordinate id="D0DAF6653F87FE086C9E1CDC5E324986" direction="east" orientation="longitude" precision="925" value="127.833336">127°50'E</geoCoordinate>
], Mt. Baekamsan, / 25.V.-20.VI.2002, K J Ahn, / C W Shin, J S Park, ex FIT //
<taxonomicName id="129EAA846D85BAC63EBE1E838683F546" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="129EAA846D85BAC63EBE1E838683F546" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
koreanus / new species / det. Sun-Jae Park (CNUIC); 1♀, KOREA, Chungbuk Prov., / Danyanggun, Youngchunmyeon,
<pageBreakToken id="ABBFFF72BD16AE7993D35981ADB7232A" pageId="7" pageNumber="40" start="start">/</pageBreakToken>
Namcheonri, Namcheon valley, / Mt. Sobaeksan [ca.
@ -169,7 +168,7 @@ Namcheonri, Namcheon valley, / Mt. Sobaeksan [ca.
<geoCoordinate id="602C6A5ADD3FFAB649266BB471B05282" direction="north" orientation="latitude" precision="925" value="35.433334">35°26'N</geoCoordinate>
<geoCoordinate id="C8F784F01BA20DA566F407DDD712EC2D" direction="east" orientation="longitude" precision="925" value="126.86667">126°52'E</geoCoordinate>
] // vi.16.1999 / D. S. Kim coll. / ex bait trap //
<taxonomicName id="DC8CBA035B1D9BD091E510E34FE3EBCE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="DC8CBA035B1D9BD091E510E34FE3EBCE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/ sp. / Det. M. Nishikawa // [5, 6, 13, 14 &amp; 15, respectively] (CNUIC); 1♂2♀♀, KOREA: Chungbuk Prov., / Boeun-gun, Mt. Sokrisan [ca.
<geoCoordinate id="D3D267361AB0E440EA938D73E0F3BB6B" direction="north" orientation="latitude" precision="925" value="36.533333">36°32'N</geoCoordinate>
<geoCoordinate id="332D1302B964E418CDA693B1495FE086" direction="east" orientation="longitude" precision="925" value="127.9">127°54'E</geoCoordinate>
@ -229,10 +228,9 @@ quite transverse, HW/HL = 1.7. Clypeofrontal suture absent. Clypeus with anterio
<paragraph id="C60EE09A958236A199EAD9E0FD469509" pageId="10" pageNumber="43">Diagnosis.</paragraph>
<paragraph id="8B73E4DEC086BD54EF38AAB1A5D5C42A" pageId="10" pageNumber="43">
See under
<taxonomicName id="959D67DCB6ED09E8C64867100B2D202A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName id="DAEBACDC6785FDDCB8C19FFB352175FC" class="Magnoliopsida" family="Malvaceae" genus="Staphylinoidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Staphylinoidea hayashii" order="Malvales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. above. Other remarks on this species see
<taxonomicName id="28DA8DE263AD38476C8A3FF5250243B8" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="0A3A0D740820C2BD2C58B4A74C5AD0BD" rank="species">sp. n.</taxonomicNameLabel>
above. Other remarks on this species see
<bibRefCitation id="F4AE7E84B4E9D2669BA8EEEF16157514" author="Wang, C-B" journalOrPublisher="ZooKeys" pageId="11" pageNumber="44" pagination="119 - 144" title="Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan." url="10.3897/zookeys.607.9074" volume="607" year="2016 a">Wang et al. (2016a)</bibRefCitation>
.
</paragraph>

View file

@ -1,287 +1,264 @@
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<taxonomicName LSID="http://zoobank.org/C468947A-D211-4411-B09B-775DFF6A7045" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus hayashii" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="hayashii">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="34">sp. n.</taxonomicNameLabel>
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<subSubSection id="017C7157212CD556CDE9C82FF654D849" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="5112F63838660F7A0F722B7FA3B712D8" pageId="1" pageNumber="34">
<taxonomicName id="C3C91ABA3E15E430EB21CB27852F8F89" ID-CoL="4Q9F6" LSID="http://zoobank.org/C468947A-D211-4411-B09B-775DFF6A7045" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus hayashii" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="4DEE1BC1F5934F492B29C1438866449B" pageId="1" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
Figs 1B, C; 2; 3
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="type material">
<paragraph pageId="1" pageNumber="34">Type material.</paragraph>
<paragraph lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
<subSubSection id="C7D28C53E8A6464F58DF6E41453E0DC1" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="type material">
<paragraph id="E92DF7FE043599689B3EA36AF8193B4B" pageId="1" pageNumber="34">Type material.</paragraph>
<paragraph id="BD92BE3DA036159E57D3C1E412690A55" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
Holotype: 1♂, USSR: Sadgorod [ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="43.25">43°15'N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="925" value="132.05">132°03'E</geoCoordinate>
<geoCoordinate id="C42FB1B7D6CD74F4C5ABD5288E544BD5" direction="north" orientation="latitude" precision="925" value="43.25">43°15'N</geoCoordinate>
<geoCoordinate id="06A49A5A8F4FF0190AD360EC8C1C631D" direction="east" orientation="longitude" precision="925" value="132.05">132°03'E</geoCoordinate>
] / (in forest; trap with / bait), Vladivostok // Primorskyi Kray / 16.VI.1978 / E. Berlov leg. (NSMT). Paratypes: 1♂1♀, FE.Russia, SW Khabarovsky / kray reg.,
<normalizedToken originalValue="Strelnikova">Strel'nikova</normalizedToken>
<normalizedToken id="C40A636E1527609D1275C07833110C69" originalValue="Strelnikova">Strel'nikova</normalizedToken>
/ range Mts.,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.716667">46°43'N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="925" value="134.13333">134°08'E</geoCoordinate>
<geoCoordinate id="2558F20685435FCBBA793ACA88B46BC8" direction="north" orientation="latitude" precision="925" value="46.716667">46°43'N</geoCoordinate>
<geoCoordinate id="768BA55CCA14C55E739D1B29BE61EA7D" direction="east" orientation="longitude" precision="925" value="134.13333">134°08'E</geoCoordinate>
/ Samur Riv., 130-550 m alt. / 25.VI.-28.VII.2014 / A. Plutenko leg. (CMNE); 1♀, same data as holotype (CMNE); 1♀, USSR Ussuri reg. / NOVOVARVAROVKA [ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="43.966667">43°58'N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="925" value="132.98334">132°59'E</geoCoordinate>
<geoCoordinate id="12CD9867246DE669417C7E0FCB03ECFF" direction="north" orientation="latitude" precision="925" value="43.966667">43°58'N</geoCoordinate>
<geoCoordinate id="FDC87CC874B70B9330EE087470BEE0D2" direction="east" orientation="longitude" precision="925" value="132.98334">132°59'E</geoCoordinate>
] / 6-10.VII.1989 / R.
<normalizedToken originalValue="Červenka">Cervenka</normalizedToken>
<normalizedToken id="CDC49BFF4EB547F95D5408D10A02945B" originalValue="Červenka">Cervenka</normalizedToken>
lgt. //
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="5053BA93DFB5EF0887C194CDE44FAE3A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) /
<taxonomicName lsidName="sibiricus" pageId="1" pageNumber="34" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="15B54690EC12DAFA2D2C655848897F05" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="sibiricus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="sibiricus">sibiricus</taxonomicName>
/ Jeannel, 1934 / Jan
<normalizedToken originalValue="Růžička">Ruzicka</normalizedToken>
<normalizedToken id="6681632577B2C2263AC2890E073D92CD" originalValue="Růžička">Ruzicka</normalizedToken>
det. 2001 // ♀ (CJRZ); 1♂1♀, USSR Ussuri reg. / JASNOE [= Yasnoye, ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="43.45">43°27'N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="925" value="132.15">132°09'E</geoCoordinate>
<geoCoordinate id="6189C1705F3A5E4152CD25A31EE28F28" direction="north" orientation="latitude" precision="925" value="43.45">43°27'N</geoCoordinate>
<geoCoordinate id="613528F110FC4212504009B9B7FE3B2C" direction="east" orientation="longitude" precision="925" value="132.15">132°09'E</geoCoordinate>
] VII.1989 / St.
<normalizedToken originalValue="Bečvář">Becvar</normalizedToken>
<normalizedToken id="C731FA6526C4F1D8907DB1BA124F76DB" originalValue="Bečvář">Becvar</normalizedToken>
lgt. //
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="2CA465B2A616531CF467424D793B9E86" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) /
<taxonomicName lsidName="sibiricus" pageId="1" pageNumber="34" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="4942F90BF108F50A0537076A40629374" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="sibiricus" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="sibiricus">sibiricus</taxonomicName>
/ Jeannel, 1934 / Jan
<normalizedToken originalValue="Růžička">Ruzicka</normalizedToken>
<normalizedToken id="DC9B15691EB737C077DF9F99DE867BD0" originalValue="Růžička">Ruzicka</normalizedToken>
det. 2001 (CJRZ); 1♂1♀, USSR Ussuri reg. / JASNOE [= Yasnoye] / VII.1989 12-16 / St.
<normalizedToken originalValue="Bečvář">Becvar</normalizedToken>
<pageBreakToken pageId="2" pageNumber="35" start="start">lgt</pageBreakToken>
<normalizedToken id="5F68A245F65810BFEBED239CE474E30F" originalValue="Bečvář">Becvar</normalizedToken>
<pageBreakToken id="6693BCE089EB0B38406CD5AFE6001593" pageId="2" pageNumber="35" start="start">lgt</pageBreakToken>
. // MOUNT. OBLATCHNAJA [= Oblachnaya] / 400-800 m (CJRZ); 1♀, RUSSIA, Far East: / S Primorye region, / LAZO env. [ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="43.366665">43°22'N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="925" value="133.9">133°54'E</geoCoordinate>
<geoCoordinate id="4833D3623454472FF603862F4A291F9B" direction="north" orientation="latitude" precision="925" value="43.366665">43°22'N</geoCoordinate>
<geoCoordinate id="1AD0C8D46A2892705BA15D7FAE86C8B2" direction="east" orientation="longitude" precision="925" value="133.9">133°54'E</geoCoordinate>
], 2.VII. / 2002 //
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="2F7B5CF33B909C4E2905F821114C64CB" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) /
<taxonomicName lsidName="sibiricus" pageId="2" pageNumber="35" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="4C7804E472431DE39E0E6F9CCE113ED4" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="sibiricus" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="sibiricus">sibiricus</taxonomicName>
/ Jeannel, 1934 / Jan
<normalizedToken originalValue="Růžička">Ruzicka</normalizedToken>
<normalizedToken id="D41151AFE6A3A2E4937DEA7E45847C77" originalValue="Růžička">Ruzicka</normalizedToken>
det 2009 / ♀ (CJRZ); 1♂, RUSSIA: Primorsky / 30 km NE Vladivostok / Tajvaza / 29.
<normalizedToken originalValue="VII">VII-</normalizedToken>
<normalizedToken id="3A2B6AE03F4CDBBDB3BADC4CBABCD356" originalValue="VII">VII-</normalizedToken>
5.VIII.1992 / B. D. Gill [leg.] (CJRZ); 1♂, Russian Far East, Primorskii krai / Lazovski Zapovednik, c. 170 km E / Vladivostok, Korpad, 1.-14.v.2001 / 175 m;
<geoCoordinate direction="north" orientation="latitude" precision="15" value="43.263054">43°15'47&quot;N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="15" value="134.12889">134°07'44&quot;E</geoCoordinate>
<geoCoordinate id="CD4B5161DCEBD8558B9F02D844F00772" direction="north" orientation="latitude" precision="15" value="43.263054">43°15'47&quot;N</geoCoordinate>
<geoCoordinate id="A662A7CBC24847D3D19EF2E640E37B38" direction="east" orientation="longitude" precision="15" value="134.12889">134°07'44&quot;E</geoCoordinate>
/ floodplain of Priamushka / Malaise trap 440; / M. Quest coll. BMNH (E) 2009-59 (BMNH); 1♀, same data as previous except: 13.-31.vii.2001 /
<geoCoordinate direction="north" orientation="latitude" precision="15" value="43.264446">43°15'52&quot;N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="15" value="134.12917">134°07'45&quot;E</geoCoordinate>
<geoCoordinate id="924CD3AFE08001F3137312B53C97EEA5" direction="north" orientation="latitude" precision="15" value="43.264446">43°15'52&quot;N</geoCoordinate>
<geoCoordinate id="6B29B84D6C81FB4ADA2BC19C074D04DF" direction="east" orientation="longitude" precision="15" value="134.12917">134°07'45&quot;E</geoCoordinate>
; 174 m; / Mountain top, Malaise trap 481 (BMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="36" type="description">
<paragraph pageId="3" pageNumber="36">
<pageBreakToken pageId="3" pageNumber="36" start="start">Description</pageBreakToken>
<subSubSection id="299D6994EC14DCB32A57CD1A224A6FDA" pageId="3" pageNumber="36" type="description">
<paragraph id="9F50B4379A7E077CF66DD0163CCFEC44" pageId="3" pageNumber="36">
<pageBreakToken id="EDB71575A3BC06F6B02321C5760B89B7" pageId="3" pageNumber="36" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="3" pageNumber="36">
<paragraph id="7ED33EE04356F255E9088C7E9E13E66A" pageId="3" pageNumber="36">
Male. EBL: 3.8-4.3 mm (4.2 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.7: 1.1: 1.1: 2.3 mm; width: HW: EW: PW: ELW = 1.1: 0.1: 1.6: 1.8 mm. Proportion of antennomeres from base to tip in
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="52B632BD433C0D388924097E1DFEA3CA" originalValue="μm">μm</normalizedToken>
(length
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="4293E593789F514A04E01FFC9926A510" originalValue="×">x</normalizedToken>
width): 149
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9D8CDE8F9BF675D7E493633AE4ADBF74" originalValue="×">x</normalizedToken>
75, 109
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="55F41DD75297B2611D85FD5CA5094363" originalValue="×">x</normalizedToken>
67, 95
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="3C149A4FFB263B009FE759C381DE5D87" originalValue="×">x</normalizedToken>
70, 64
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="661095573224C5CCFD4B29EBF8BEF5FA" originalValue="×">x</normalizedToken>
79, 58
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="8E348C10C01D730DE017A6619C1AF8C8" originalValue="×">x</normalizedToken>
92, 41
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="ECBE5344C69C755178E038D1336B04C2" originalValue="×">x</normalizedToken>
100, 91
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="521D2AD240C8164746C1FC1E7187BDAD" originalValue="×">x</normalizedToken>
128, 39
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="869AD5BB2CBAF573DFA8BF9836363EBE" originalValue="×">x</normalizedToken>
128, 87
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="347F03BCF2829DBE9D8E7A418FEE6A70" originalValue="×">x</normalizedToken>
151, 90
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="E1AD997DDDE7CEC955AB5B26F2CE14A7" originalValue="×">x</normalizedToken>
156, 161
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="BDABA457BFD617267E9CE3CB59C5442E" originalValue="×">x</normalizedToken>
143.
</paragraph>
<paragraph pageId="3" pageNumber="36">Habitus (Fig. 1B) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, antennae (apical half of ultimate antennomere yellowish) and tarsi reddish brown. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
<caption pageId="3" pageNumber="36">
<paragraph pageId="3" pageNumber="36">
<paragraph id="F01CC3EBFD19EB7BF84B5FBB65F56659" pageId="3" pageNumber="36">Habitus (Fig. 1B) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, antennae (apical half of ultimate antennomere yellowish) and tarsi reddish brown. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
<caption id="1211AE70008E9E77FB697C4D7BECBB02" pageId="3" pageNumber="36">
<paragraph id="333F8DD59B26D0CD9CCC19A658C62183" pageId="3" pageNumber="36">
Figure 1. Habitus of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) spp. (dorsal view). A,
<normalizedToken originalValue="DF">D-F</normalizedToken>
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="3" pageNumber="36" rank="species" species="sibiricus">sibiricus</taxonomicName>
Jeannel, 1934 A ♀ (holotype; Vladivostok) D ♂ (Vladivostok) E ♂ (Pyeongchang-gun) F ♀ (Boeun-gun) B, C
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="3" pageNumber="36" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. B ♂ (holotype; Vladivostok) C ♀ (paratype; Ussuri region). Scale: 1 mm.
<taxonomicName id="9527CF76CF4A8A8D1C2493DB8DDF2255" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">Ptomaphagus (s. str.) spp.</taxonomicName>
(dorsal view). A,
<normalizedToken id="491D06A90C869C586A7CB9B346F30883" originalValue="DF">D-F</normalizedToken>
<taxonomicName id="4E5602ABC3EC76AD7729847E33377195" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus Jeannel, 1934</taxonomicName>
A ♀ (holotype; Vladivostok) D ♂ (Vladivostok) E ♂ (Pyeongchang-gun) F ♀ (Boeun-gun) B, C
<taxonomicName id="4B0718F1D15DDE40A93EA7330377E248" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="44AAC9658886C472E7628EB7AB1DC65B" rank="species">sp. n.</taxonomicNameLabel>
B ♂ (holotype; Vladivostok) C ♀ (paratype; Ussuri region). Scale: 1 mm.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="36">Head transverse, HW/HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.11. Antennae (Fig. 2A) slender, AL/HW = 1.0; antennomere III shorter than II; VI with length/width = 0.4; XI peach-shape.</paragraph>
<caption pageId="3" pageNumber="36">
<paragraph pageId="3" pageNumber="36">
<paragraph id="D52D8A59C6ABBC9364A3970F9545B194" pageId="3" pageNumber="36">Head transverse, HW/HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.11. Antennae (Fig. 2A) slender, AL/HW = 1.0; antennomere III shorter than II; VI with length/width = 0.4; XI peach-shape.</paragraph>
<caption id="4B630E847518681C26469F378E441095" pageId="3" pageNumber="36">
<paragraph id="302DD9545BDADCA2FDC6F08E6D60504D" pageId="3" pageNumber="36">
Figure 2.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="3" pageNumber="36" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. (♂: holotype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scales: 0.1 mm.
<taxonomicName id="1AF488DCD5AC97D5F1BC29AE3DE7692F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="C0A00D3C945D4519852B0648A6E377FF" rank="species">sp. n.</taxonomicNameLabel>
(♂: holotype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="36">Pronotum (Fig. 2B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, simply narrowing from posterior to anterior; hind angles slightly projected backwards and acute. Posterior margin widely protruded in middle part, emarginate near hind angles.</paragraph>
<paragraph pageId="3" pageNumber="36">Elytra oval, widest at about basal 1/4, ELL/EW = 1.3. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 2G) narrowly rounded. Sutural striae present. Metathoracic wings fully developed.</paragraph>
<paragraph pageId="3" pageNumber="36">Prolegs robust, with basal three protarsomeres (Fig. 2C) strongly expanded: TW/BTW = 1.2. Protibiae (Fig. 2E) strongly expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph pageId="3" pageNumber="36">Abdominal ventrite VIII (Fig. 2I) round at posterior edge and with an inconspicuous median notch. Spiculum gastrale (Fig. 2J) of genital segment with about 1/5 of length protruding beyond anterior edge of epipleurite IX.</paragraph>
<paragraph pageId="3" pageNumber="36">Aedeagus (Fig. 3A) long and slender, with median lobe gradually narrowing towards a lanceolate apex and terminated to a widely rounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 3C) inserted with 5 ventrally oriented setae on both sides; parameres narrow, reaching about apical 1/5 of median lobe, each apex (Fig. 3D) with 2 lateral setae and 1 apical seta distinctly shorter. In lateral view (Fig. 3B), median lobe regularly bent ventrad and gradually tapering to a flat apex. Endophallus with stylus quite slender, a subelliptical nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.</paragraph>
<caption pageId="3" pageNumber="36">
<paragraph pageId="3" pageNumber="36">
<paragraph id="89F1006B2AD06F9738EC010B1B268AE1" pageId="3" pageNumber="36">Pronotum (Fig. 2B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, simply narrowing from posterior to anterior; hind angles slightly projected backwards and acute. Posterior margin widely protruded in middle part, emarginate near hind angles.</paragraph>
<paragraph id="35A633AAF3BD552387C6224B5DA14478" pageId="3" pageNumber="36">Elytra oval, widest at about basal 1/4, ELL/EW = 1.3. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 2G) narrowly rounded. Sutural striae present. Metathoracic wings fully developed.</paragraph>
<paragraph id="D5AA733EF8DB7BE596303425B4C6796C" pageId="3" pageNumber="36">Prolegs robust, with basal three protarsomeres (Fig. 2C) strongly expanded: TW/BTW = 1.2. Protibiae (Fig. 2E) strongly expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph id="C8E3006562F9480A92AF46395B803264" pageId="3" pageNumber="36">Abdominal ventrite VIII (Fig. 2I) round at posterior edge and with an inconspicuous median notch. Spiculum gastrale (Fig. 2J) of genital segment with about 1/5 of length protruding beyond anterior edge of epipleurite IX.</paragraph>
<paragraph id="F88A5AEB587FF454F4B9E3E3A679529E" pageId="3" pageNumber="36">Aedeagus (Fig. 3A) long and slender, with median lobe gradually narrowing towards a lanceolate apex and terminated to a widely rounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 3C) inserted with 5 ventrally oriented setae on both sides; parameres narrow, reaching about apical 1/5 of median lobe, each apex (Fig. 3D) with 2 lateral setae and 1 apical seta distinctly shorter. In lateral view (Fig. 3B), median lobe regularly bent ventrad and gradually tapering to a flat apex. Endophallus with stylus quite slender, a subelliptical nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.</paragraph>
<caption id="4E9F578E65CCD173F03FFE51E131FB92" pageId="3" pageNumber="36">
<paragraph id="EFD4E902EC3F4D2F2733C7ADD4435EE7" pageId="3" pageNumber="36">
Figure 3.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="3" pageNumber="36" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. (♂: holotype). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scales: 0.1 mm.
<taxonomicName id="C2201254412A72A0E513E08918E48384" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="7B45EC56087B9D889BED1654D3F05FEE" rank="species">sp. n.</taxonomicNameLabel>
(♂: holotype). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="36">Female. Similar to male in general appearance (Fig. 1C), including elytral apices (Fig. 2H), but distinguished by the following characteristics: protarsi (Fig. 2D) simply linear; protibiae (Fig. 2F) narrower; abdominal ventrite VIII (Fig. 3E) narrowly rounded at posterior edge; genital segment and ovipositor as shown in Fig. 3F; spermatheca (Fig. 3F) curved in distal part and coiled in proximal part.</paragraph>
<paragraph id="9377C57EADA110CC8E6AFA5778762CBF" pageId="3" pageNumber="36">Female. Similar to male in general appearance (Fig. 1C), including elytral apices (Fig. 2H), but distinguished by the following characteristics: protarsi (Fig. 2D) simply linear; protibiae (Fig. 2F) narrower; abdominal ventrite VIII (Fig. 3E) narrowly rounded at posterior edge; genital segment and ovipositor as shown in Fig. 3F; spermatheca (Fig. 3F) curved in distal part and coiled in proximal part.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37" type="diagnosis">
<paragraph pageId="4" pageNumber="37">
<pageBreakToken pageId="4" pageNumber="37" start="start">Diagnosis</pageBreakToken>
<subSubSection id="BC1477D7472EB63EA20296505BDE13D0" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37" type="diagnosis">
<paragraph id="F6CA9015A541464C3CF43E232475D200" pageId="4" pageNumber="37">
<pageBreakToken id="72A7ABF0255CA73D23B2012087EE7A8E" pageId="4" pageNumber="37" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="37">
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="4" pageNumber="37" rank="species" species="sibiricus">sibiricus</taxonomicName>
<paragraph id="76EE9D151812EE4C4316059258B0C858" pageId="4" pageNumber="37">
<taxonomicName id="FF52FF89AD07B4849637C03C6B36AEFA" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
is sympatric with
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="4" pageNumber="37" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. in some locations of the Russian Far East, and they are very similar to each other (including spermatheca (Figs 3F; 5F), which is only more roundly curved in the stem part in
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="4" pageNumber="37" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n.). For external characters, the body size of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="4" pageNumber="37" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="FD9549191F147BA279C3BE921D6F5496" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="B94F53DC733BAD602E6CE05E93B70B2F" rank="species">sp. n.</taxonomicNameLabel>
in some locations of the Russian Far East, and they are very similar to each other (including spermatheca (Figs 3F; 5F), which is only more roundly curved in the stem part in
<taxonomicName id="737ACF468E69EFE00194708F3F8A0F6F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="5FB1E5940CA94AD1F2A85FD06142C777" rank="species">sp. n.</taxonomicNameLabel>
). For external characters, the body size of
<taxonomicName id="956CEE881609848C5DE6BF3FCD064275" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
(Figs 1A,
<normalizedToken originalValue="DF">D-F</normalizedToken>
<normalizedToken id="1BC129C70B4F2DAF79ED1BEFD83BA0FE" originalValue="DF">D-F</normalizedToken>
) is a little smaller than
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="4" pageNumber="37" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. (Figs 1B, C). However, their aedeagi provide critical characters to distinguish the two species: in
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="4" pageNumber="37" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n., the aedeagus is much larger and more slender (Fig. 3A), the right apicoventral piece of the median lobe is slenderly lanceolate (Fig. 3C), the apical half of median lobe much flatter in lateral view (Fig. 3B); while in
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="4" pageNumber="37" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="25B3B00989469819C7ED11A6A5B4711B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="510BC363946A3A41DF9200A20AEE88AD" rank="species">sp. n.</taxonomicNameLabel>
(Figs 1B, C). However, their aedeagi provide critical characters to distinguish the two species: in
<taxonomicName id="1685E37C7BADD2A3AD4EE41BEBF0C2C1" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="1CF8BEE34E7DF6C79984D7E3D6839333" rank="species">sp. n.</taxonomicNameLabel>
, the aedeagus is much larger and more slender (Fig. 3A), the right apicoventral piece of the median lobe is slenderly lanceolate (Fig. 3C), the apical half of median lobe much flatter in lateral view (Fig. 3B); while in
<taxonomicName id="69C64A728579EDC044494BB6C8D9DE06" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
, the aedeagus is stouter (Fig. 5A), the right apicoventral piece of median lobe is much wider and subpentagonal (Fig. 5C), the apical half of the median lobe thicker in lateral view (Fig. 5B).
</paragraph>
<caption pageId="4" pageNumber="37">
<paragraph pageId="4" pageNumber="37">
<caption id="D291266382DEBC481CF8051B2444CB0E" pageId="4" pageNumber="37">
<paragraph id="1DB64A54880380C8700BF02C6FF6B1D1" pageId="4" pageNumber="37">
Figure 4.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="4" pageNumber="37" rank="species" species="sibiricus">sibiricus</taxonomicName>
Jeannel, 1934 (Vladivostok). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scales: 0.1 mm.
<taxonomicName id="6E7897AAAB07B24071AA4A91AEF92661" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus Jeannel, 1934</taxonomicName>
(Vladivostok). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scales: 0.1 mm.
</paragraph>
</caption>
<caption pageId="4" pageNumber="37">
<paragraph pageId="4" pageNumber="37">
<caption id="292CC59BC9ED570F77B1C370DD2173F6" pageId="4" pageNumber="37">
<paragraph id="44CA9B1613ACBDAC87002C7E67C80915" pageId="4" pageNumber="37">
Figure 5.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="4" pageNumber="37" rank="species" species="sibiricus">sibiricus</taxonomicName>
Jeannel, 1934 (Vladivostok). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scales: 0.1 mm.
<taxonomicName id="214261599E61F564EBC557F8A45BB86E" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus Jeannel, 1934</taxonomicName>
(Vladivostok). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="38">
<pageBreakToken pageId="5" pageNumber="38" start="start">Furthermore</pageBreakToken>
<paragraph id="87A205672AF252BE5F6976B9AB55488F" pageId="5" pageNumber="38">
<pageBreakToken id="AC7D1FFE0C5C5BD35EE39754377E22DA" pageId="5" pageNumber="38" start="start">Furthermore</pageBreakToken>
, based on specimens examined,
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="sibiricus" pageId="5" pageNumber="38" rank="species" species="sibiricus">sibiricus</taxonomicName>
<taxonomicName id="EB99BD009659B21E0A7823B0A1BD86C7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="sibiricus">Ptomaphagus (s. str.) sibiricus</taxonomicName>
seems to be much more widely distributed, extending southward to South Korea; while
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.)
<taxonomicName lsidName="hayashii" pageId="5" pageNumber="38" rank="species" species="hayashii">hayashii</taxonomicName>
sp. n. is presently known only in the Russian Far East.
<taxonomicName id="297E91393140618E90F3CA4C5B18ACE2" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="hayashii" status="sp. nov.">Ptomaphagus (s. str.) hayashii</taxonomicName>
<taxonomicNameLabel id="9717E2B37784069712F812E3F48E32E3" rank="species">sp. n.</taxonomicNameLabel>
is presently known only in the Russian Far East.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="38" type="etymology">
<paragraph pageId="5" pageNumber="38">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="38">
<subSubSection id="B9C5D362C60B45F3D6CD2AA454278884" pageId="5" pageNumber="38" type="etymology">
<paragraph id="44EECF5F0391AD6DA34F64F7549179C5" pageId="5" pageNumber="38">Etymology.</paragraph>
<paragraph id="4B560A3669CB5E5FFF58BE0400771DAF" pageId="5" pageNumber="38">
The specific epithet is dedicated to Mr. Yasuhiko Hayashi (Kawanishi, Japan), a famous independent taxonomist on
<taxonomicName lsidName="Staphylinoidea" pageId="5" pageNumber="38" rank="superFamily" superFamily="Staphylinoidea">Staphylinoidea</taxonomicName>
<taxonomicName id="F7D8AA7EAF411AB04AF2796832E1F97B" class="Insecta" kingdom="Animalia" lsidName="Staphylinoidea" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="superFamily" superFamily="Staphylinoidea">Staphylinoidea</taxonomicName>
, for his continual generous help to our study.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="38" type="distribution">
<paragraph pageId="5" pageNumber="38">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="38">Russia (Far East) (Fig. 6).</paragraph>
<caption pageId="5" pageNumber="38">
<paragraph pageId="5" pageNumber="38">
<subSubSection id="9670C6098D94077270F8D26A4FA4361C" pageId="5" pageNumber="38" type="distribution">
<paragraph id="3827E36AAC1E8F90F95980FDB3126CD8" pageId="5" pageNumber="38">Distribution.</paragraph>
<paragraph id="580AF0B2E9515491243182F47F30FFF4" pageId="5" pageNumber="38">Russia (Far East) (Fig. 6).</paragraph>
<caption id="720B2B8CE1A6E67266FE3A4B2A07763E" pageId="5" pageNumber="38">
<paragraph id="FA7225B46FF3D1481CE2DEEE28397FA7" pageId="5" pageNumber="38">
Figure 6. Distribution map of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="127713778AB5096751DF16547082D331" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
species from the Russian Far East and the Korean Peninsula.
</paragraph>
</caption>