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<document ID-DOI="http://doi.org/10.5281/zenodo.4760725" ID-GBIF-Dataset="1abcb62d-e2d8-4daf-9d33-7acb0cbd9580" ID-ISSN="1854-0392" ID-Zenodo-Dep="4760725" IM.materialsCitations_approvedBy="diego" IM.tables_requiresApprovalFor="existingObjects" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1620963958573" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Kondratieff, Boris C." docDate="2012" docId="C86087E19326F457403CFD3BFADFFC6E" docLanguage="en" docName="Illiesia.8.5.45-77.pdf" docOrigin="Illiesia 8 (5)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Allocapnia RICKERI" docType="treatment" docVersion="6" lastPageNumber="66" masterDocId="3459FF999333F442411EFFB0FFBEFFFA" masterDocTitle="Epiproct And Dorsal Process Structure In The Allocapnia Forbesi Frison, A. Pygmaea (Burmeister), And A. Rickeri Frison Species Groups (Plecoptera: Capniidae), And Inclusion Of A. Minima (Newport) In A New Species Group" masterLastPageNumber="77" masterPageNumber="45" pageNumber="66" updateTime="1680606490160" updateUser="diego" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Epiproct And Dorsal Process Structure In The Allocapnia Forbesi Frison, A. Pygmaea (Burmeister), And A. Rickeri Frison Species Groups (Plecoptera: Capniidae), And Inclusion Of A. Minima (Newport) In A New Species Group</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>and Boris C. Kondratieff &amp; Department of Biology, Box 4045, Mississippi College, Clinton, Mississippi 39058, U. S. A. E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kondratieff, Boris C.</mods:namePart>
<mods:affiliation>Department of Bioagricultural Sciences and Pest Management,</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>77</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760725</mods:identifier>
<mods:identifier type="GBIF-Dataset">1abcb62d-e2d8-4daf-9d33-7acb0cbd9580</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4760725</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4763006" ID-GBIF-Taxon="182073852" ID-Zenodo-Dep="4763006" LSID="urn:lsid:plazi:treatment:C86087E19326F457403CFD3BFADFFC6E" httpUri="http://treatment.plazi.org/id/C86087E19326F457403CFD3BFADFFC6E" lastPageNumber="66" pageId="21" pageNumber="66">
<subSubSection box="[290,696,651,675]" pageId="21" pageNumber="66" type="nomenclature">
<paragraph blockId="21.[290,696,651,675]" box="[290,696,651,675]" pageId="21" pageNumber="66">
<heading allCaps="true" bold="true" box="[290,696,651,675]" centered="true" fontSize="10" level="1" pageId="21" pageNumber="66" reason="1">
<emphasis bold="true" box="[290,696,651,675]" pageId="21" pageNumber="66">
<taxonomicName ID-CoL="8MPYS" ID-ENA="143702" authority="RICKERI" authorityName="RICKERI" box="[290,589,651,675]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[290,589,651,675]" italics="true" pageId="21" pageNumber="66">ALLOCAPNIA RICKERI</emphasis>
</taxonomicName>
GROUP
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="66" type="discussion">
<paragraph blockId="21.[189,798,730,1927]" pageId="21" pageNumber="66">
<bibRefCitation author="Ross, H. H. &amp; W. E. Ricker" box="[227,500,730,755]" pageId="21" pageNumber="66" refId="ref13207" refString="Ross, H. H. &amp; W. E. Ricker. 1971. The classification, evolution, and dispersal of the winter stonefly genus Allocapnia. Illinois Biological Monographs 45: University of Illinois Press, Urbana, Illinois. 166 pp." type="book" year="1971">Ross &amp; Ricker (1971)</bibRefCitation>
placed six species (
<taxonomicName authority="Ross &amp; Ricker 1971" authorityName="Ross &amp; Ricker" authorityYear="1971" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="cunninghami">
<emphasis italics="true" pageId="21" pageNumber="66">A. cunninghami</emphasis>
<bibRefCitation author="Ross, H. H. &amp; W. E. Ricker" box="[341,571,766,790]" pageId="21" pageNumber="66" refId="ref13207" refString="Ross, H. H. &amp; W. E. Ricker. 1971. The classification, evolution, and dispersal of the winter stonefly genus Allocapnia. Illinois Biological Monographs 45: University of Illinois Press, Urbana, Illinois. 166 pp." type="book" year="1971">Ross &amp; Ricker 1971</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Ross &amp; Ricker 1971" authorityName="Ross &amp; Ricker" authorityYear="1971" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis box="[583,707,766,790]" italics="true" pageId="21" pageNumber="66">A. perplexa</emphasis>
<bibRefCitation author="Ross, H. H. &amp; W. E. Ricker" pageId="21" pageNumber="66" refId="ref13207" refString="Ross, H. H. &amp; W. E. Ricker. 1971. The classification, evolution, and dispersal of the winter stonefly genus Allocapnia. Illinois Biological Monographs 45: University of Illinois Press, Urbana, Illinois. 166 pp." type="book" year="1971">Ross &amp; Ricker 1971</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Frison 1942" authorityName="Frison" authorityYear="1942" box="[355,623,801,826]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">
<emphasis box="[355,463,801,825]" italics="true" pageId="21" pageNumber="66">A. rickeri</emphasis>
<bibRefCitation author="Frison, T. H." box="[478,623,801,826]" pageId="21" pageNumber="66" pagination="235 - 355" refId="ref12315" refString="Frison, T. H. 1942. Studies of North American Plecoptera with special reference to the fauna of Illinois. Bulletin of the Illinois Natural History Survey, 22: 235 - 355." type="journal article" year="1942">Frison 1942</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Ricker 1952" authorityName="Ricker" authorityYear="1952" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="sandersoni">
<emphasis box="[642,797,801,825]" italics="true" pageId="21" pageNumber="66">A. sandersoni</emphasis>
<bibRefCitation author="Ricker, W. E." box="[189,331,837,861]" pageId="21" pageNumber="66" refId="ref13098" refString="Ricker, W. E. 1952. Systematic studies in Plecoptera. Indiana University Publications, Science Series No. 18. Indiana University Press, Bloomington, Indiana. 200 pp." type="book" year="1952">Ricker 1952</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Ross 1964" authorityName="Ross" authorityYear="1964" box="[346,618,837,861]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="stannardi">
<emphasis box="[346,486,837,861]" italics="true" pageId="21" pageNumber="66">A. stannardi</emphasis>
Ross 1964
</taxonomicName>
;
<taxonomicName authority="Ricker 1952" authorityName="Ricker" authorityYear="1952" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="zola">
<emphasis box="[632,712,837,861]" italics="true" pageId="21" pageNumber="66">A. zola</emphasis>
<bibRefCitation author="Ricker, W. E." pageId="21" pageNumber="66" refId="ref13098" refString="Ricker, W. E. 1952. Systematic studies in Plecoptera. Indiana University Publications, Science Series No. 18. Indiana University Press, Bloomington, Indiana. 200 pp." type="book" year="1952">Ricker 1952</bibRefCitation>
</taxonomicName>
) in this group and suggested, primarily on the basis of female characters, these form two “well- marked complexes of three species each”. The line of fusion between abdominal sterna 7-8 includes a sclerotized ridge in females of
<taxonomicName authorityName="Ross &amp; Ricker" authorityYear="1971" box="[573,754,1015,1039]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="cunninghami">
<emphasis box="[573,754,1015,1039]" italics="true" pageId="21" pageNumber="66">A. cunninghami</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Ross &amp; Ricker" authorityYear="1971" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis italics="true" pageId="21" pageNumber="66">A. perplexa</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Ricker" authorityYear="1952" box="[338,413,1050,1074]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="zola">
<emphasis box="[338,413,1050,1074]" italics="true" pageId="21" pageNumber="66">A. zola</emphasis>
</taxonomicName>
, whereas females of
<emphasis italics="true" pageId="21" pageNumber="66">
<taxonomicName box="[658,761,1050,1074]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">A. rickeri</taxonomicName>
,
<taxonomicName authorityName="Ricker" authorityYear="1952" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="sandersoni">A. sandersoni</taxonomicName>
</emphasis>
and most
<taxonomicName authorityName="Ross" authorityYear="1964" box="[422,556,1086,1110]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="stannardi">
<emphasis box="[422,556,1086,1110]" italics="true" pageId="21" pageNumber="66">A. stannardi</emphasis>
</taxonomicName>
lack such a structure. Subsequently,
<taxonomicName authority="Kirchner 1980" authorityName="Kirchner" authorityYear="1980" box="[360,637,1121,1145]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="harperi">
<emphasis box="[360,468,1121,1145]" italics="true" pageId="21" pageNumber="66">A. harperi</emphasis>
<bibRefCitation author="Kirchner, R. F." box="[473,637,1121,1145]" pageId="21" pageNumber="66" pagination="19 - 21" refId="ref12638" refString="Kirchner, R. F. 1980. A new Allocapnia from Virginia (Plecoptera: Capniidae). Entomological News, 91: 19 - 21." type="journal article" year="1980">Kirchner 1980</bibRefCitation>
</taxonomicName>
, was added to the second complex.
</paragraph>
<paragraph blockId="21.[189,798,730,1927]" pageId="21" pageNumber="66">
The epiproct upper limb for species in this group, is typified by a long basal segment and a relatively short, somewhat bulbous apical segment (
<figureCitation box="[668,755,1263,1287]" captionStart="Figs" captionStartId="22.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1695]" captionTargetId="figure-64@22.[249,1383,315,1695]" captionTargetPageId="22" captionText="Figs. 79-84. Allocapnia cunninghami, tributary Little Trammel Creek, Sumner Co., Tennessee. 79. Male terminalia, dorsal. 80. Apical segment upper limb of epiproct tip, dorsal. 81. Surface detail apical segment upper limb of epiproct, dorsal. 82. 8th tergal process and epiproct tip, lateral. 83. 8th tergal process and epiproct tip, anterodorsal. 84. 7th tergal process, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760754" httpUri="https://zenodo.org/record/4760754/files/figure.png" pageId="21" pageNumber="66">Figs. 79</figureCitation>
,
<figureCitation box="[764,792,1263,1287]" captionStart="Figs" captionStartId="23.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1698]" captionTargetId="figure-64@23.[249,1383,315,1698]" captionTargetPageId="23" captionText="Figs. 85-90. Allocapnia harperi, Station Spring Creek, Tazewell Co., Virginia. 85. Male terminalia, dorsal. 86. Apical segment upper limb of epiproct, dorsal. 87. Apical segment upper limb of epiproct tip, dorsal. 88. Surface detail apical segment upper limb of epiproct, dorsal. 89. 8th tergal process and epiproct tip, lateral. 90. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760758" httpUri="https://zenodo.org/record/4760758/files/figure.png" pageId="21" pageNumber="66">85</figureCitation>
,
<figureCitation box="[189,218,1299,1323]" captionStart="Figs" captionStartId="25.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1699]" captionTargetId="figure-64@25.[249,1383,315,1699]" captionTargetPageId="25" captionText="Figs. 91-96. Allocapnia perplexa, East Fork Bledsoe Creek, Sumner Co., Tennessee. 91. Male terminalia, dorsal. 92. Apical segment upper limb of epiproct tip, dorsal. 93. Surface detail apical segment upper limb of epiproct, dorsal. 94. 8th tergal process and epiproct tip, lateral. 95. 8th tergal process and epiproct tip, dorsal. 96. 8th tergal process, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760760" httpUri="https://zenodo.org/record/4760760/files/figure.png" pageId="21" pageNumber="66">91</figureCitation>
,
<figureCitation box="[235,265,1299,1323]" captionStart="Figs" captionStartId="26.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1696]" captionTargetId="figure-64@26.[249,1383,315,1696]" captionTargetPageId="26" captionText="Figs. 97-102. Allocapnia rickeri, Turkey Creek, Scioto Co., Ohio. 97. Male terminalia, dorsal. 98. Apical segment upper limb of epiproct tip, dorsal. 99. Male terminalia, lateral. 100. 8th tergal process and epiproct tip, lateral. 101. 8th tergal process and epiproct tip, dorsal. 102. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760762" httpUri="https://zenodo.org/record/4760762/files/figure.png" pageId="21" pageNumber="66">97</figureCitation>
), a feature shared with members of the
<taxonomicName authorityName=": Ross &amp; Ricker" authorityYear="1971" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="forbesi">
<emphasis italics="true" pageId="21" pageNumber="66">A. forbesi</emphasis>
</taxonomicName>
group. The apical segment bears a distal patch of densely packed, wave-like spikes (
<figureCitation box="[617,704,1370,1394]" captionStart="Figs" captionStartId="22.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1695]" captionTargetId="figure-64@22.[249,1383,315,1695]" captionTargetPageId="22" captionText="Figs. 79-84. Allocapnia cunninghami, tributary Little Trammel Creek, Sumner Co., Tennessee. 79. Male terminalia, dorsal. 80. Apical segment upper limb of epiproct tip, dorsal. 81. Surface detail apical segment upper limb of epiproct, dorsal. 82. 8th tergal process and epiproct tip, lateral. 83. 8th tergal process and epiproct tip, anterodorsal. 84. 7th tergal process, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760754" httpUri="https://zenodo.org/record/4760754/files/figure.png" pageId="21" pageNumber="66">Figs. 81</figureCitation>
,
<figureCitation box="[714,741,1370,1394]" captionStart="Figs" captionStartId="23.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1698]" captionTargetId="figure-64@23.[249,1383,315,1698]" captionTargetPageId="23" captionText="Figs. 85-90. Allocapnia harperi, Station Spring Creek, Tazewell Co., Virginia. 85. Male terminalia, dorsal. 86. Apical segment upper limb of epiproct, dorsal. 87. Apical segment upper limb of epiproct tip, dorsal. 88. Surface detail apical segment upper limb of epiproct, dorsal. 89. 8th tergal process and epiproct tip, lateral. 90. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760758" httpUri="https://zenodo.org/record/4760758/files/figure.png" pageId="21" pageNumber="66">87</figureCitation>
) and the basal segment has a relatively broad, shallow longitudinal groove. The lower limb bears a distinct notch offsetting the apical spoon (
<figureCitation box="[672,785,1476,1501]" captionStart="Figs" captionStartId="26.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1696]" captionTargetId="figure-64@26.[249,1383,315,1696]" captionTargetPageId="26" captionText="Figs. 97-102. Allocapnia rickeri, Turkey Creek, Scioto Co., Ohio. 97. Male terminalia, dorsal. 98. Apical segment upper limb of epiproct tip, dorsal. 99. Male terminalia, lateral. 100. 8th tergal process and epiproct tip, lateral. 101. 8th tergal process and epiproct tip, dorsal. 102. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760762" httpUri="https://zenodo.org/record/4760762/files/figure.png" pageId="21" pageNumber="66">Fig. 100</figureCitation>
). Abdominal tergum 8, or 7 and 8, bear dorsal processes (
<figureCitation box="[319,412,1547,1572]" captionStart="Figs" captionStartId="23.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1698]" captionTargetId="figure-64@23.[249,1383,315,1698]" captionTargetPageId="23" captionText="Figs. 85-90. Allocapnia harperi, Station Spring Creek, Tazewell Co., Virginia. 85. Male terminalia, dorsal. 86. Apical segment upper limb of epiproct, dorsal. 87. Apical segment upper limb of epiproct tip, dorsal. 88. Surface detail apical segment upper limb of epiproct, dorsal. 89. 8th tergal process and epiproct tip, lateral. 90. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760758" httpUri="https://zenodo.org/record/4760758/files/figure.png" pageId="21" pageNumber="66">Figs. 90</figureCitation>
,
<figureCitation box="[426,455,1547,1571]" captionStart="Figs" captionStartId="25.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1699]" captionTargetId="figure-64@25.[249,1383,315,1699]" captionTargetPageId="25" captionText="Figs. 91-96. Allocapnia perplexa, East Fork Bledsoe Creek, Sumner Co., Tennessee. 91. Male terminalia, dorsal. 92. Apical segment upper limb of epiproct tip, dorsal. 93. Surface detail apical segment upper limb of epiproct, dorsal. 94. 8th tergal process and epiproct tip, lateral. 95. 8th tergal process and epiproct tip, dorsal. 96. 8th tergal process, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760760" httpUri="https://zenodo.org/record/4760760/files/figure.png" pageId="21" pageNumber="66">95</figureCitation>
,
<figureCitation box="[468,511,1548,1572]" captionStart="Figs" captionStartId="29.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1701]" captionTargetId="figure-64@29.[249,1383,315,1701]" captionTargetPageId="29" captionText="Figs. 109-114. Allocapnia stannardi, 1 mile W Newfound Gap, Sevier Co., Tennessee. 109. Male terminalia, dorsal. 110. Apical segment upper limb of epiproct tip, dorsal. 111. Surface detail apical segment upper limb of epiporct, dorsal. 112. Male terminalia, lateral. 113. Abdominal tergal processes, dorsal. 114. Abdominal tergal processes, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760766" httpUri="https://zenodo.org/record/4760766/files/figure.png" pageId="21" pageNumber="66">112</figureCitation>
); those on tergum 8 are typically bilobed but in one species,
<emphasis box="[654,742,1583,1607]" italics="true" pageId="21" pageNumber="66">
<taxonomicName authorityName="Ricker" authorityYear="1952" box="[654,738,1583,1607]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="zola">A. zola</taxonomicName>
,
</emphasis>
this structure is trilobed and a second process, also present on segment 8, is displaced toward the anterior segmental margin (
<figureCitation box="[539,642,1689,1714]" captionStart="Figs" captionStartId="30.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1701]" captionTargetId="figure-64@30.[249,1383,315,1701]" captionTargetPageId="30" captionText="Figs. 115-120. Allocapnia zola, Stoney Fork Reed Creek, Wythe Co., Virginia. 115. Male terminalia, dorsal. 116. Apical segment upper limb of epiproct tip, dorsal. 117. Surface detail apical segment upper limb of epiproct, dorsal. 118. Abdominal tergal processes, dorsal. 119. Male terminalia, lateral. 120. Abdominal tergal processes, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760770" httpUri="https://zenodo.org/record/4760770/files/figure.png" pageId="21" pageNumber="66">Fig. 115</figureCitation>
). In Ross &amp; Ricker’s analysis,
<taxonomicName authorityName="Ross" authorityYear="1964" box="[410,551,1725,1749]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="stannardi">
<emphasis box="[410,551,1725,1749]" italics="true" pageId="21" pageNumber="66">A. stannardi</emphasis>
</taxonomicName>
and
<taxonomicName box="[621,727,1725,1749]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">
<emphasis box="[621,727,1725,1749]" italics="true" pageId="21" pageNumber="66">A. rickeri</emphasis>
</taxonomicName>
were considered sister species with
<taxonomicName authorityName="Ricker" authorityYear="1952" box="[570,723,1761,1785]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="sandersoni">
<emphasis box="[570,723,1761,1785]" italics="true" pageId="21" pageNumber="66">A. sandersoni</emphasis>
</taxonomicName>
more distantly related, and the other complex was presented as an unresolved trichotomy. The
<taxonomicName box="[701,797,1832,1856]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">
<emphasis box="[701,797,1832,1856]" italics="true" pageId="21" pageNumber="66">A. rickeri</emphasis>
</taxonomicName>
group shares “ancestor number 15” with
<taxonomicName authority="Ross 1964" authorityName="Ross" authorityYear="1964" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="jeanae">
<emphasis box="[696,797,1868,1891]" italics="true" pageId="21" pageNumber="66">A. jeanae</emphasis>
Ross 1964
</taxonomicName>
, in this analysis (
<bibRefCitation author="Ross, H. H. &amp; W. E. Ricker" box="[507,736,1903,1927]" pageId="21" pageNumber="66" refId="ref13207" refString="Ross, H. H. &amp; W. E. Ricker. 1971. The classification, evolution, and dispersal of the winter stonefly genus Allocapnia. Illinois Biological Monographs 45: University of Illinois Press, Urbana, Illinois. 166 pp." type="book" year="1971">Ross &amp; Ricker 1971</bibRefCitation>
).
</paragraph>
<paragraph blockId="21.[835,1444,253,916]" pageId="21" pageNumber="66">
We are unable to provide a definitive hypothesis for relationships within the
<taxonomicName box="[1191,1298,288,312]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">
<emphasis box="[1191,1298,288,312]" italics="true" pageId="21" pageNumber="66">A. rickeri</emphasis>
</taxonomicName>
group, but based primarily on tergal lobe variations, we could propose one subgroup in which the 8
<superScript attach="left" box="[1292,1307,359,373]" fontSize="6" pageId="21" pageNumber="66">th</superScript>
tergal lobe notch is at least 75 μm wide (
<emphasis italics="true" pageId="21" pageNumber="66">
<taxonomicName authorityName="Kirchner" authorityYear="1980" box="[1167,1278,395,419]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="harperi">A. harperi</taxonomicName>
,
<taxonomicName authorityName="Ross &amp; Ricker" authorityYear="1971" box="[1287,1409,395,419]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="perplexa">A. perplexa</taxonomicName>
,
<taxonomicName class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">A. rickeri</taxonomicName>
,
<taxonomicName authorityName="Ross" authorityYear="1964" box="[915,1049,430,454]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="stannardi">A. stannardi</taxonomicName>
</emphasis>
;
<figureCitation box="[1061,1151,430,455]" captionStart="Figs" captionStartId="23.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1698]" captionTargetId="figure-64@23.[249,1383,315,1698]" captionTargetPageId="23" captionText="Figs. 85-90. Allocapnia harperi, Station Spring Creek, Tazewell Co., Virginia. 85. Male terminalia, dorsal. 86. Apical segment upper limb of epiproct, dorsal. 87. Apical segment upper limb of epiproct tip, dorsal. 88. Surface detail apical segment upper limb of epiproct, dorsal. 89. 8th tergal process and epiproct tip, lateral. 90. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760758" httpUri="https://zenodo.org/record/4760758/files/figure.png" pageId="21" pageNumber="66">Figs. 90</figureCitation>
,
<figureCitation box="[1160,1189,430,454]" captionStart="Figs" captionStartId="25.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1699]" captionTargetId="figure-64@25.[249,1383,315,1699]" captionTargetPageId="25" captionText="Figs. 91-96. Allocapnia perplexa, East Fork Bledsoe Creek, Sumner Co., Tennessee. 91. Male terminalia, dorsal. 92. Apical segment upper limb of epiproct tip, dorsal. 93. Surface detail apical segment upper limb of epiproct, dorsal. 94. 8th tergal process and epiproct tip, lateral. 95. 8th tergal process and epiproct tip, dorsal. 96. 8th tergal process, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760760" httpUri="https://zenodo.org/record/4760760/files/figure.png" pageId="21" pageNumber="66">96</figureCitation>
,
<figureCitation box="[1199,1241,431,455]" captionStart="Figs" captionStartId="26.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1696]" captionTargetId="figure-64@26.[249,1383,315,1696]" captionTargetPageId="26" captionText="Figs. 97-102. Allocapnia rickeri, Turkey Creek, Scioto Co., Ohio. 97. Male terminalia, dorsal. 98. Apical segment upper limb of epiproct tip, dorsal. 99. Male terminalia, lateral. 100. 8th tergal process and epiproct tip, lateral. 101. 8th tergal process and epiproct tip, dorsal. 102. 8th tergal process and epiproct tip, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760762" httpUri="https://zenodo.org/record/4760762/files/figure.png" pageId="21" pageNumber="66">101</figureCitation>
,
<figureCitation box="[1251,1292,431,455]" captionStart="Figs" captionStartId="29.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1701]" captionTargetId="figure-64@29.[249,1383,315,1701]" captionTargetPageId="29" captionText="Figs. 109-114. Allocapnia stannardi, 1 mile W Newfound Gap, Sevier Co., Tennessee. 109. Male terminalia, dorsal. 110. Apical segment upper limb of epiproct tip, dorsal. 111. Surface detail apical segment upper limb of epiporct, dorsal. 112. Male terminalia, lateral. 113. Abdominal tergal processes, dorsal. 114. Abdominal tergal processes, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760766" httpUri="https://zenodo.org/record/4760766/files/figure.png" pageId="21" pageNumber="66">114</figureCitation>
) and another in which the 8
<superScript attach="left" box="[1001,1016,465,479]" fontSize="6" pageId="21" pageNumber="66">th</superScript>
tergal lobe notch is no more than 45 μm wide (
<emphasis box="[955,1382,502,526]" italics="true" pageId="21" pageNumber="66">
<taxonomicName authorityName="Ross &amp; Ricker" authorityYear="1971" box="[955,1133,502,526]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="cunninghami">A. cunninghami</taxonomicName>
,
<taxonomicName authorityName="Ricker" authorityYear="1952" box="[1143,1292,502,526]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="sandersoni">A. sandersoni</taxonomicName>
,
<taxonomicName authorityName="Ricker" authorityYear="1952" box="[1301,1376,502,526]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="zola">A. zola</taxonomicName>
;
</emphasis>
<figureCitation captionStart="Figs" captionStartId="22.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1695]" captionTargetId="figure-64@22.[249,1383,315,1695]" captionTargetPageId="22" captionText="Figs. 79-84. Allocapnia cunninghami, tributary Little Trammel Creek, Sumner Co., Tennessee. 79. Male terminalia, dorsal. 80. Apical segment upper limb of epiproct tip, dorsal. 81. Surface detail apical segment upper limb of epiproct, dorsal. 82. 8th tergal process and epiproct tip, lateral. 83. 8th tergal process and epiproct tip, anterodorsal. 84. 7th tergal process, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760754" httpUri="https://zenodo.org/record/4760754/files/figure.png" pageId="21" pageNumber="66">Figs. 83</figureCitation>
,
<figureCitation box="[879,922,537,561]" captionStart="Figs" captionStartId="28.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1696]" captionTargetId="figure-64@28.[249,1383,315,1696]" captionTargetPageId="28" captionText="Figs. 103-108. Allocapnia sandersoni, Little Red River, Searcy Co., Arkansas. 103. Male terminalia, dorsal. 104. Apical segment upper limb of epiproct tip, dorsal. 105. Male terminalia, lateral. 106. 8th tergal process and epiproct tip, lateral. 107. 8th tergal process and epiproct tip, dorsal. 108. 8th tergal process, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760764" httpUri="https://zenodo.org/record/4760764/files/figure.png" pageId="21" pageNumber="66">108</figureCitation>
,
<figureCitation box="[936,980,537,561]" captionStart="Figs" captionStartId="30.[189,241,1780,1804]" captionTargetBox="[249,1383,315,1701]" captionTargetId="figure-64@30.[249,1383,315,1701]" captionTargetPageId="30" captionText="Figs. 115-120. Allocapnia zola, Stoney Fork Reed Creek, Wythe Co., Virginia. 115. Male terminalia, dorsal. 116. Apical segment upper limb of epiproct tip, dorsal. 117. Surface detail apical segment upper limb of epiproct, dorsal. 118. Abdominal tergal processes, dorsal. 119. Male terminalia, lateral. 120. Abdominal tergal processes, anterodorsal." figureDoi="http://doi.org/10.5281/zenodo.4760770" httpUri="https://zenodo.org/record/4760770/files/figure.png" pageId="21" pageNumber="66">120</figureCitation>
). Within the first group,
<emphasis italics="true" pageId="21" pageNumber="66">
<taxonomicName authorityName="Kirchner" authorityYear="1980" box="[1288,1404,537,561]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="harperi">A. harperi</taxonomicName>
,
<taxonomicName authorityName="Ross" authorityYear="1964" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="stannardi">A. stannardi</taxonomicName>
,
</emphasis>
and
<taxonomicName box="[1001,1101,573,597]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="rickeri">
<emphasis box="[1001,1101,573,597]" italics="true" pageId="21" pageNumber="66">A. rickeri</emphasis>
</taxonomicName>
share ear-like lobes on the 8
<superScript attach="right" box="[1429,1444,572,586]" fontSize="6" pageId="21" pageNumber="66">th</superScript>
tergal process, whereas those of
<taxonomicName authorityName="Ross &amp; Ricker" authorityYear="1971" box="[1226,1350,608,632]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis box="[1226,1350,608,632]" italics="true" pageId="21" pageNumber="66">A. perplexa</emphasis>
</taxonomicName>
are not ear shaped, however we offer no resolution for the former trichotomy. In the
<emphasis italics="true" pageId="21" pageNumber="66">
<taxonomicName authorityName="Ross &amp; Ricker" authorityYear="1971" box="[1140,1318,679,703]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="cunninghami">A. cunninghami</taxonomicName>
,
<taxonomicName authorityName="Ricker" authorityYear="1952" box="[1329,1407,679,703]" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="zola">A. zola</taxonomicName>
,
<taxonomicName authorityName="Ricker" authorityYear="1952" class="Insecta" family="Capniidae" genus="Allocapnia" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="66" phylum="Arthropoda" rank="species" species="sandersoni">A. sandersoni</taxonomicName>
</emphasis>
subgroup, the former two species each share a small lobe on tergum 7 and also a small median projection on the 8
<superScript attach="left" box="[1167,1182,785,799]" fontSize="6" pageId="21" pageNumber="66">th</superScript>
tergal process which supports a sister group relationship between these two species. Unfortunately, we have no probable apomorphy which unites this latter subgroup.
</paragraph>
</subSubSection>
</treatment>
</document>
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