577 lines
141 KiB
XML
577 lines
141 KiB
XML
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<mods:title id="7AFB27883B7DBD2513F0EAEC5BF420D2">A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae)</mods:title>
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<mods:title id="58915D4AFE4D453947AA2A2E2BF684D6">Zootaxa</mods:title>
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Genus
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<taxonomicName id="F8E64D3E9E65402553C9CFFB60639B8C" ID-CoL="486N" ID-ENA="2880603" authority="Hewitt, 1916" authorityName="Hewitt" authorityYear="1916" box="[237,547,1733,1759]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="0D92EAAF9E65402553C9CFFB633C9B8C" bold="true" box="[237,380,1733,1759]" italics="true" pageId="4" pageNumber="5">Drassodella</emphasis>
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<bibRefCitation id="5B774B4C9E65402552A0CFFB60639B8C" author="Hewitt, J." box="[388,547,1733,1759]" pageId="4" pageNumber="5" pagination="180 - 213" refId="ref44666" refString="Hewitt, J. (1916) Descriptions of new South African spiders. Annals of the Transvaal Museum, 5, 180 - 213." type="journal article" year="1916">Hewitt, 1916</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="77FC65369E65402553B3CE3266039A13" pageId="4" pageNumber="5" type="reference_group">
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<paragraph id="3F5936BD9E65402553B3CE3266039A13" blockId="4.[151,1436,1804,1856]" pageId="4" pageNumber="5">
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<taxonomicName id="F8E64D3E9E65402553B3CE3263A19A71" ID-CoL="486N" authority="Hewitt, 1916: 209" authorityName="Hewitt" authorityPageNumber="209" authorityYear="1916" box="[151,481,1804,1826]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="0D92EAAF9E65402553B3CE3263539A71" box="[151,275,1804,1826]" italics="true" pageId="4" pageNumber="5">Drassodella</emphasis>
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<treatmentCitation id="BE4710AC9E6540255239CE3263A19A71" author="Hewitt, J." box="[285,481,1804,1826]" page="209" pageId="4" pageNumber="5" year="1916">
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<bibRefCitation id="5B774B4C9E6540255239CE3263E99A71" author="Hewitt, J." box="[285,425,1804,1826]" pageId="4" pageNumber="5" pagination="180 - 213" refId="ref44666" refString="Hewitt, J. (1916) Descriptions of new South African spiders. Annals of the Transvaal Museum, 5, 180 - 213." type="journal article" year="1916">Hewitt, 1916</bibRefCitation>
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: 209
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="BE4710AC9E65402552D4CE3260F49A71" author="Tucker, R. W. E." box="[496,692,1804,1826]" page="309" pageId="4" pageNumber="5" year="1923">
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<bibRefCitation id="5B774B4C9E65402552D4CE32603D9A71" author="Tucker, R. W. E." box="[496,637,1804,1826]" pageId="4" pageNumber="5" pagination="251 - 437" refId="ref45524" refString="Tucker, R. W. E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251 - 437." type="journal article" year="1923">Tucker, 1923</bibRefCitation>
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: 309
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||
</treatmentCitation>
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;
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<treatmentCitation id="BE4710AC9E65402551E7CE3361CF9A71" author="Roewer, C. F." box="[707,911,1804,1826]" page="384" pageId="4" pageNumber="5" year="1954">
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<bibRefCitation id="5B774B4C9E65402551E7CE3361199A71" author="Roewer, C. F." box="[707,857,1804,1826]" pageId="4" pageNumber="5" refId="ref45416" refString="Roewer, C. F. (1954) Katalog der Araneae von 1758 bis 1940, bzw. 1954. Uol. 2. Institut royal des sciences naturelles de Belgique, Bruxelles, 1751 pp." type="book" year="1954">Roewer, 1954</bibRefCitation>
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||
: 384
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||
</treatmentCitation>
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, 1467;
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<treatmentCitation id="BE4710AC9E65402550C6CE3366F79A71" author="Bonnet, P." box="[994,1207,1804,1826]" page="1556" pageId="4" pageNumber="5" year="1956">
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<bibRefCitation id="5B774B4C9E65402550C6CE3366339A71" author="Bonnet, P." box="[994,1139,1804,1826]" pageId="4" pageNumber="5" pagination="919 - 1926" refId="ref44102" refString="Bonnet, P. (1956) Bibliographia araneorum. 2 (2). Freres Douladoure, Toulouse, 1008 pp. [pp. 919 - 1926]" type="book chapter" year="1956">Bonnet, 1956</bibRefCitation>
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: 1556
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</treatmentCitation>
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;
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<treatmentCitation id="BE4710AC9E65402557E2CE3267D89A71" author="Platnick, N. I." box="[1222,1432,1804,1826]" page="457" pageId="4" pageNumber="5" year="1989">
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<bibRefCitation id="5B774B4C9E65402557E2CE3267209A71" author="Platnick, N. I." box="[1222,1376,1804,1826]" pageId="4" pageNumber="5" refId="ref45106" refString="Platnick, N. I. (1989) Advances in Spider Taxonomy 1981 - 1987: a supplement to Brignoli's A Catalog of the Araneae described between 1940 and 1981 (edited by P. Merrett). Manchester University Press, Manschester, 673 pp." type="book" year="1989">Platnick, 1989</bibRefCitation>
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: 457
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="BE4710AC9E65402553E3CE14603E9A13" author="Dippenaar-Schoeman, A. S. & Jocque, R." box="[199,638,1834,1856]" page="166" pageId="4" pageNumber="5" year="1997">
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<bibRefCitation id="5B774B4C9E65402553E3CE14600B9A13" author="Dippenaar-Schoeman, A. S. & Jocque, R." box="[199,587,1834,1856]" pageId="4" pageNumber="5" refId="ref44412" refString="Dippenaar-Schoeman, A. S. & Jocque, R. (1997) African spiders: an identification manual. Plant Protection Research Institute handbook 9. Biosystematics Division, ARC-Plant Research Institute, Pretoria, 392 pp." type="book" year="1997">Dippenaar-Schoeman & Jocqué, 1997</bibRefCitation>
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||
: 166
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||
</treatmentCitation>
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;
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||
<treatmentCitation id="BE4710AC9E65402551ACCE14667F9A13" author="Jocque, R. & Dippenaar-Schoeman, A. S." box="[648,1087,1834,1856]" page="126" pageId="4" pageNumber="5" year="2006">
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<bibRefCitation id="5B774B4C9E65402551ACCE14664C9A13" author="Jocque, R. & Dippenaar-Schoeman, A. S." box="[648,1036,1834,1856]" pageId="4" pageNumber="5" refId="ref44761" refString="Jocque, R. & Dippenaar-Schoeman, A. S. (2006) Spider families of the World. Royal Museum for Central Africa, Tervuren, 334 pp. Available from: https: // www. africamuseum. be / sites / default / files / media / docs / research / publications / rmca / online / zoology-documentation / spider-families _ of _ the _ world. pdf (Accessed 8 Apr. 2019)" type="book" year="2006">Jocqué & Dippenaar-Schoeman, 2006</bibRefCitation>
|
||
: 126
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||
</treatmentCitation>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="77FC65369E65402553B3CE5661D99A2D" box="[151,921,1896,1918]" pageId="4" pageNumber="5" type="materials_examined">
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<paragraph id="3F5936BD9E65402553B3CE5661D99A2D" blockId="4.[151,921,1896,1918]" box="[151,921,1896,1918]" pageId="4" pageNumber="5">
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<emphasis id="0D92EAAF9E65402553B3CE5663669A2D" bold="true" box="[151,294,1896,1918]" pageId="4" pageNumber="5">
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<typeStatus id="E05D881F9E65402553B3CE56628C9A2D" box="[151,204,1896,1918]" pageId="4" pageNumber="5">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="F8E64D3E9E654025520FCE5660DE9A2D" authority="Hewitt, 1916" authorityName="Hewitt" authorityYear="1916" box="[299,670,1896,1918]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="salisburyi">
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<emphasis id="0D92EAAF9E654025520FCE5660519A2D" box="[299,529,1896,1918]" italics="true" pageId="4" pageNumber="5">Drassodella salisburyi</emphasis>
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<bibRefCitation id="5B774B4C9E654025513CCE5660DE9A2D" author="Hewitt, J." box="[536,670,1896,1918]" pageId="4" pageNumber="5" pagination="180 - 213" refId="ref44666" refString="Hewitt, J. (1916) Descriptions of new South African spiders. Annals of the Transvaal Museum, 5, 180 - 213." type="journal article" year="1916">Hewitt, 1916</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<subSubSection id="77FC65369E65402453B3CE94614E9C36" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="3F5936BD9E65402453B3CE94614E9C36" blockId="4.[151,1437,1962,2023]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<emphasis id="0D92EAAF9E65402553B3CE9463539A90" bold="true" box="[151,275,1962,1987]" pageId="4" pageNumber="5">Diagnosis:</emphasis>
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<taxonomicName id="F8E64D3E9E654025523ECE9463E39A90" authorityName="Hewitt" authorityYear="1916" box="[282,419,1962,1987]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="0D92EAAF9E654025523ECE9463E39A90" box="[282,419,1962,1987]" italics="true" pageId="4" pageNumber="5">Drassodella</emphasis>
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</taxonomicName>
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can be recognised from
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<taxonomicName id="F8E64D3E9E654025519CCE94610A9A90" authorityName="Lawrence" authorityYear="1938" box="[696,842,1962,1987]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E654025519CCE94610A9A90" box="[696,842,1962,1987]" italics="true" pageId="4" pageNumber="5">Austrachelas</emphasis>
|
||
</taxonomicName>
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by the following: 1) carapace almost always with an median upsilon marking of white feathery setae and the lateral margins with a narrow fringe of white feathery setae, which are absent in
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<taxonomicName id="F8E64D3E9E644024529AC9A960109DE3" authorityName="Lawrence" authorityYear="1938" box="[446,592,151,176]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="0D92EAAF9E644024529AC9A960109DE3" box="[446,592,151,176]" italics="true" pageId="5" pageNumber="6">Austrachelas</emphasis>
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||
</taxonomicName>
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||
; 2) usually with striped or spotted dorsal markings on the abdomen, which are rarely absent (e.g.
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<figureCitation id="A7DD2A389E64402452AAC98263B19D87" box="[398,497,188,213]" captionStart="FIGURES 1–4" captionStartId="5.[151,264,1326,1348]" captionTargetBox="[151,1435,406,1306]" captionTargetId="figure@5.[151,1435,406,1306]" captionTargetPageId="5" captionText="FIGURES 1–4. General habitus photographs of live Drassodella spiders: 1 D. amatola sp. nov. female from Hogsback, Eastern Cape; 2 D. quinquelabecula Tucker, 1923 female from Aardvark Nature Reserve, Western Cape; 3 D. septemmaculata (Strand, 1909) male from Fisherhaven, Western Cape; 4 D. vasivulva Tucker, 1923 male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb." httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="5" pageNumber="6">Figs 1–4</figureCitation>
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||
), while
|
||
<taxonomicName id="F8E64D3E9E644024516FC982609D9D86" authorityName="Lawrence" authorityYear="1938" box="[587,733,188,213]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E644024516FC982609D9D86" box="[587,733,188,213]" italics="true" pageId="5" pageNumber="6">Austrachelas</emphasis>
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||
</taxonomicName>
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||
have chevron markings (see
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||
<bibRefCitation id="5B774B4C9E6440245706C98266BB9D87" author="Haddad, C. R. & Lyle, R. & Bosselaers, J. & Ramirez, M. J." box="[1058,1275,188,213]" pageId="5" pageNumber="6" pagination="1 - 38" refId="ref44545" refString="Haddad, C. R., Lyle, R., Bosselaers, J. & Ramirez, M. J. (2009) A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae). Zootaxa, 2296, 1 - 38." type="journal article" year="2009">
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||
Haddad
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||
<emphasis id="0D92EAAF9E64402457A4C98366F89D86" box="[1152,1208,188,213]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
: figs 1–15); 3) shield-shaped sternum (
|
||
<figureCitation id="A7DD2A389E6440245286C9E163B59DAB" box="[418,501,223,248]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">Fig. 26</figureCitation>
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||
), while oval in
|
||
<taxonomicName id="F8E64D3E9E6440245180C9E161769DAB" authorityName="Lawrence" authorityYear="1938" box="[676,822,223,248]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E6440245180C9E161769DAB" box="[676,822,223,248]" italics="true" pageId="5" pageNumber="6">Austrachelas</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="5B774B4C9E6440245056C9E1660C9DAB" author="Haddad, C. R. & Lyle, R. & Bosselaers, J. & Ramirez, M. J." box="[882,1100,223,248]" pageId="5" pageNumber="6" pagination="1 - 38" refId="ref44545" refString="Haddad, C. R., Lyle, R., Bosselaers, J. & Ramirez, M. J. (2009) A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae). Zootaxa, 2296, 1 - 38." type="journal article" year="2009">
|
||
Haddad
|
||
<emphasis id="0D92EAAF9E64402450F4C9DF66499DAB" box="[976,1033,223,248]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
: fig. 17); 4) anterior legs with weakly scopulate metatarsi and tarsi (
|
||
<figureCitation id="A7DD2A389E6440245164C83A60D89C4F" box="[576,664,260,285]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="5" pageNumber="6">Figs 34</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6440245180C83A60AE9C4F" box="[676,750,260,284]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="5" pageNumber="6">39–42</figureCitation>
|
||
), while densely scopulate in
|
||
<taxonomicName id="F8E64D3E9E6440245711C83A66879C4E" authorityName="Lawrence" authorityYear="1938" box="[1077,1223,260,285]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E6440245711C83A66879C4E" box="[1077,1223,260,285]" italics="true" pageId="5" pageNumber="6">Austrachelas</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="5B774B4C9E6440245620C83A62949C13" author="Haddad, C. R. & Lyle, R. & Bosselaers, J. & Ramirez, M. J." pageId="5" pageNumber="6" pagination="1 - 38" refId="ref44545" refString="Haddad, C. R., Lyle, R., Bosselaers, J. & Ramirez, M. J. (2009) A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae). Zootaxa, 2296, 1 - 38." type="journal article" year="2009">
|
||
Haddad
|
||
<emphasis id="0D92EAAF9E6440245647C83B67DC9C4E" box="[1379,1436,260,285]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
: figs 24–26, 33); 5) posterior metatarsi with the pro- and retrolateral dorsal spines occurring singly (i.e. pl 3 rl 3), while usually paired in
|
||
<taxonomicName id="F8E64D3E9E64402452E7C87260159C36" authorityName="Lawrence" authorityYear="1938" box="[451,597,332,357]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E64402452E7C87260159C36" box="[451,597,332,357]" italics="true" pageId="5" pageNumber="6">Austrachelas</emphasis>
|
||
</taxonomicName>
|
||
(i.e. pl 6 rl 4-6).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6B9966359E64402453B3CC10615F98F3" httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,264,1326,1348]" targetBox="[151,1435,406,1306]" targetPageId="5">
|
||
<paragraph id="3F5936BD9E64402453B3CC10615F98F3" blockId="5.[151,1436,1326,1440]" pageId="5" pageNumber="6">
|
||
<emphasis id="0D92EAAF9E64402453B3CC10637E9817" bold="true" box="[151,318,1326,1348]" pageId="5" pageNumber="6">FIGURES 1–4.</emphasis>
|
||
General habitus photographs of live
|
||
<taxonomicName id="F8E64D3E9E64402451EACC10610A9817" authorityName="Hewitt" authorityYear="1916" box="[718,842,1326,1348]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E64402451EACC10610A9817" box="[718,842,1326,1348]" italics="true" pageId="5" pageNumber="6">Drassodella</emphasis>
|
||
</taxonomicName>
|
||
spiders: 1
|
||
<taxonomicName id="F8E64D3E9E64402450E1CC11667B9817" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[965,1083,1326,1348]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="amatola" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E64402450E1CC11667B9817" box="[965,1083,1326,1348]" italics="true" pageId="5" pageNumber="6">D. amatola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6440245762CC1066DA9817" bold="true" box="[1094,1178,1326,1348]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="16A157D49E6440245762CC1066DA9817" box="[1094,1178,1326,1348]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female from Hogsback, Eastern Cape; 2
|
||
<taxonomicName id="F8E64D3E9E644024521ACC7060D29830" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[318,658,1357,1379]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quinquelabecula">
|
||
<emphasis id="0D92EAAF9E644024521ACC7060469830" box="[318,518,1357,1379]" italics="true" pageId="5" pageNumber="6">D. quinquelabecula</emphasis>
|
||
Tucker, 1923
|
||
</taxonomicName>
|
||
female from Aardvark Nature Reserve, Western Cape; 3
|
||
<taxonomicName id="F8E64D3E9E64402457FCCC70636C98D1" authority="(Strand, 1909)" authorityName="Strand" authorityYear="1909" baseAuthorityName="Strand" baseAuthorityYear="1909" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="septemmaculata">
|
||
<emphasis id="0D92EAAF9E64402457FCCC7067DC9830" box="[1240,1436,1357,1379]" italics="true" pageId="5" pageNumber="6">D. septemmaculata</emphasis>
|
||
(Strand, 1909)
|
||
</taxonomicName>
|
||
male from Fisherhaven, Western Cape; 4
|
||
<taxonomicName id="F8E64D3E9E64402451D7CC53664998D1" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[755,1033,1388,1410]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vasivulva">
|
||
<emphasis id="0D92EAAF9E64402451D7CC53613698D1" box="[755,886,1388,1410]" italics="true" pageId="5" pageNumber="6">D. vasivulva</emphasis>
|
||
Tucker, 1923
|
||
</taxonomicName>
|
||
male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="77FC65369E64402A53E3CCF3672D9F33" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6" type="description">
|
||
<paragraph id="3F5936BD9E64402453E3CCF3611A9AC6" blockId="5.[151,1436,1485,1942]" pageId="5" pageNumber="6">
|
||
<emphasis id="0D92EAAF9E64402453E3CCF3631B98B5" bold="true" box="[199,347,1485,1510]" pageId="5" pageNumber="6">Description:</emphasis>
|
||
Small to medium spiders, body length
|
||
<quantity id="F81E9B589E644024500ACCF3619798B6" box="[814,983,1485,1510]" metricMagnitude="-3" metricUnit="m" metricValue="6.205" metricValueMax="9.85" metricValueMin="2.56" pageId="5" pageNumber="6" unit="mm" value="6.205" valueMax="9.85" valueMin="2.56">2.56–9.85 mm</quantity>
|
||
. Carapace broad, oval, cephalic region slightly narrowed, broadest between coxae II and III (
|
||
<figureCitation id="A7DD2A389E6440245028CCCF61369B59" box="[780,886,1521,1546]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="5" pageNumber="6">Figs 5–8</figureCitation>
|
||
); in lateral profile raised evenly along midline, sloping gently posteriorly (
|
||
<figureCitation id="A7DD2A389E64402452EECF2B60509B7D" box="[458,528,1557,1582]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
|
||
); posterior margin rebordered, moderately concave above pedicel (
|
||
<figureCitation id="A7DD2A389E644024562ECF2B67229B7D" box="[1290,1378,1557,1582]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="5" pageNumber="6">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E644024564BCF2867CE9B7D" box="[1391,1422,1558,1582]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">18</figureCitation>
|
||
); eight eyes in two rows (4:4); lateral eyes set on low tubercles; all eyes surrounded by black rings; AER slightly recurved when viewed dorsally (
|
||
<figureCitation id="A7DD2A389E6440245120CF63600D9B25" box="[516,589,1629,1654]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="5" pageNumber="6">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E644024517CCF6060369B25" box="[600,630,1630,1654]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">19</figureCitation>
|
||
), straight or slightly recurved when viewed from front; AME smaller or equal to ALE; AME separated by more than half their diameter, closer to ALE than to each other; PER slightly recurved, broader than AER (
|
||
<figureCitation id="A7DD2A389E64402452CDCF9B607D9BED" box="[489,573,1701,1726]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
|
||
); PME flattened, transversely oval, more or less the same distance between each other and PLE; PME smaller than PLE. Clypeus slightly larger than AME diameter, with few long erect setae; surface smooth, sparsely covered in short straight and feathery setae; usually black, rarely yellow-orange or brown with reticulate black mottling, usually with white feathery setae forming an inverted Y-shaped (upsilon) dorsal marking along midline, with split posterior to fovea (
|
||
<figureCitation id="A7DD2A389E6440245027CE0B611A9A1D" box="[771,858,1845,1870]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">Fig. 18</figureCitation>
|
||
); lateral margins with narrow fringe composed of white feathery setae (
|
||
<figureCitation id="A7DD2A389E64402452A2CE6760479A22" box="[390,519,1881,1906]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">Figs 20, 21</figureCitation>
|
||
), rarely without dorsal markings; several long erect setae scattered between fovea and posterior margin (
|
||
<figureCitation id="A7DD2A389E64402452ABCE4363A29AC5" box="[399,482,1917,1942]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="5" pageNumber="6">Fig. 18</figureCitation>
|
||
); fovea short to elongate, narrow.
|
||
</paragraph>
|
||
<caption id="6B9966359E67402753B3CFA567129AF4" httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,264,1691,1713]" targetBox="[209,1363,193,1648]" targetPageId="6">
|
||
<paragraph id="3F5936BD9E67402753B3CFA567129AF4" blockId="6.[151,1437,1691,1959]" pageId="6" pageNumber="7">
|
||
<emphasis id="0D92EAAF9E67402753B3CFA5630B9BE2" bold="true" box="[151,331,1691,1713]" pageId="6" pageNumber="7">FIGURES 5–17.</emphasis>
|
||
Digital microscope photographs of
|
||
<taxonomicName id="F8E64D3E9E67402751EBCFA5610B9BE2" authorityName="Hewitt" authorityYear="1916" box="[719,843,1691,1713]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E67402751EBCFA5610B9BE2" box="[719,843,1691,1713]" italics="true" pageId="6" pageNumber="7">Drassodella</emphasis>
|
||
</taxonomicName>
|
||
somatic morphology: 5–8 dorsal view of carapace and chelicerae of
|
||
<taxonomicName id="F8E64D3E9E6740275204CF8563329B83" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[288,370,1722,1744]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6740275204CF8563329B83" box="[288,370,1722,1744]" italics="true" pageId="6" pageNumber="7">D. flava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E674027525CCF8463899B83" bold="true" box="[376,457,1722,1744]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="16A157D49E674027525CCF8463899B83" box="[376,457,1722,1744]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female (5),
|
||
<taxonomicName id="F8E64D3E9E6740275161CF8560ED9B83" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[581,685,1723,1744]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guttata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6740275161CF8560ED9B83" box="[581,685,1723,1744]" italics="true" pageId="6" pageNumber="7">D. guttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6740275197CF8461449B83" bold="true" box="[691,772,1722,1744]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="16A157D49E6740275197CF8461449B83" box="[691,772,1722,1744]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male (6),
|
||
<taxonomicName id="F8E64D3E9E6740275049CF8561B79B83" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[877,1015,1723,1744]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="transversa" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6740275049CF8561B79B83" box="[877,1015,1723,1744]" italics="true" pageId="6" pageNumber="7">D. transversa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E67402750D9CF84660E9B83" bold="true" box="[1021,1102,1722,1744]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="16A157D49E67402750D9CF84660E9B83" box="[1021,1102,1722,1744]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male (7) and
|
||
<taxonomicName id="F8E64D3E9E67402757F8CF85628A9BBC" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="purcelli">
|
||
<emphasis id="0D92EAAF9E67402757F8CF8567099B83" box="[1244,1353,1722,1744]" italics="true" pageId="6" pageNumber="7">D. purcelli</emphasis>
|
||
Tucker, 1923
|
||
</taxonomicName>
|
||
female (8); 9
|
||
<taxonomicName id="F8E64D3E9E674027527ACFE460CD9BBC" authority="Lawrence, 1938" authorityName="Lawrence" authorityYear="1938" box="[350,653,1753,1775]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="0D92EAAF9E674027527ACFE463A19BBC" box="[350,481,1753,1775]" italics="true" pageId="6" pageNumber="7">D. tenebrosa</emphasis>
|
||
Lawrence, 1938
|
||
</taxonomicName>
|
||
female, carapace, lateral view; 10
|
||
<taxonomicName id="F8E64D3E9E67402750D1CFE466249BBC" authorityName="Tucker" authorityYear="1923" box="[1013,1124,1753,1775]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="purcelli">
|
||
<emphasis id="0D92EAAF9E67402750D1CFE466249BBC" box="[1013,1124,1753,1775]" italics="true" pageId="6" pageNumber="7">D. purcelli</emphasis>
|
||
</taxonomicName>
|
||
female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of
|
||
<taxonomicName id="F8E64D3E9E67402756A7CFC7639B9A7F" authority="(Strand, 1909)" authorityName="Strand" authorityYear="1909" baseAuthorityName="Strand" baseAuthorityYear="1909" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="septemmaculata">
|
||
<emphasis id="0D92EAAF9E67402756A7CFC7637C9A7F" italics="true" pageId="6" pageNumber="7">D. septemmaculata</emphasis>
|
||
(Strand, 1909)
|
||
</taxonomicName>
|
||
male (12),
|
||
<taxonomicName id="F8E64D3E9E674027517FCE2960AF9A7F" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[603,751,1815,1836]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aurostriata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E674027517FCE2960AF9A7F" box="[603,751,1815,1836]" italics="true" pageId="6" pageNumber="7">D. aurostriata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E67402751DDCE28610E9A7F" bold="true" box="[761,846,1814,1836]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="16A157D49E67402751DDCE28610E9A7F" box="[761,846,1814,1836]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male (13) and
|
||
<taxonomicName id="F8E64D3E9E67402750D2CE29662C9A7F" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[1014,1132,1814,1836]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="amatola" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E67402750D2CE29662C9A7F" box="[1014,1132,1814,1836]" italics="true" pageId="6" pageNumber="7">D. amatola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6740275752CE28668B9A7F" bold="true" box="[1142,1227,1814,1836]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="16A157D49E6740275752CE28668B9A7F" box="[1142,1227,1814,1836]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female (14); 15
|
||
<taxonomicName id="F8E64D3E9E67402756A7CE2962BA9A18" authorityName="Lawrence" authorityYear="1938" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="0D92EAAF9E67402756A7CE2962BA9A18" italics="true" pageId="6" pageNumber="7">D. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
female, abdomen, anterodorsal view; 16
|
||
<taxonomicName id="F8E64D3E9E6740275196CE0861389A18" baseAuthorityName="Strand" baseAuthorityYear="1909" box="[690,888,1845,1867]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="septemmaculata">
|
||
<emphasis id="0D92EAAF9E6740275196CE0861389A18" box="[690,888,1845,1867]" italics="true" pageId="6" pageNumber="7">D. septemmaculata</emphasis>
|
||
</taxonomicName>
|
||
male, abdomen, showing sclerites on venter; 17
|
||
<taxonomicName id="F8E64D3E9E67402756A7CE0862BA9A3A" authorityName="Lawrence" authorityYear="1938" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="0D92EAAF9E67402756A7CE0862BA9A3A" italics="true" pageId="6" pageNumber="7">D. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F5936BD9E66402653E3C9A9618B9C83" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="8">
|
||
Chelicerae usually moderately protruding in males, slightly angled and/or vertically orientated in females, slightly tapering distally (
|
||
<figureCitation id="A7DD2A389E6640265299C98260659D86" box="[445,549,188,213]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="7" pageNumber="8">Figs 5–8</figureCitation>
|
||
); surface finely granulate, with many moderately long erect setae on anterior surface of paturon; cheliceral escort seta (
|
||
<emphasis id="0D92EAAF9E6640265159C9DF60FB9DAB" box="[637,699,225,248]" italics="true" pageId="7" pageNumber="8">sensu</emphasis>
|
||
<bibRefCitation id="5B774B4C9E66402651E1C9DE612A9DAB" author="Ramirez, M. J." box="[709,874,224,248]" pageId="7" pageNumber="8" pagination="1 - 374" refId="ref45369" refString="Ramirez, M. J. (2014) The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. https: // doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014</bibRefCitation>
|
||
: fig. 15B) present, promarginally near fang base (
|
||
<figureCitation id="A7DD2A389E66402653BBC83A62B49C4F" box="[159,244,260,285]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
|
||
); condyle (boss) small to large (
|
||
<figureCitation id="A7DD2A389E664026517EC83A60DF9C4F" box="[602,671,260,285]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
|
||
); fangs moderately elongate (
|
||
<figureCitation id="A7DD2A389E66402650C8C83A667F9C4F" box="[1004,1087,260,285]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
|
||
), with fine ridges near opening of venom ducts; promargin with three teeth, retromargin with two teeth (
|
||
<figureCitation id="A7DD2A389E664026509AC81966519C13" box="[958,1041,295,320]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="7" pageNumber="8">Fig. 23</figureCitation>
|
||
). Chilum single, distinct, triangular in shape; endites obliquely depressed, very broad, slightly sinuous, widest medially, without median groove, with distinct dense maxillar hair tuft and serrula (
|
||
<figureCitation id="A7DD2A389E66402651E4C85161139CDB" box="[704,851,367,392]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="7" pageNumber="8">Figs 24, 25</figureCitation>
|
||
); labium slightly longer than broad or as long, posterolateral margins of labium with distinct sclerotized groove; anterior portion slightly inclined, posterior portion short; anterior margin with erect setae, slightly concave medially.
|
||
</paragraph>
|
||
<caption id="6B9966359E66402653B3CF31639B9BED" httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,264,1551,1573]" targetBox="[162,1424,517,1529]" targetPageId="7">
|
||
<paragraph id="3F5936BD9E66402653B3CF31639B9BED" blockId="7.[151,1436,1551,1726]" pageId="7" pageNumber="8">
|
||
<emphasis id="0D92EAAF9E66402653B3CF3163139B76" bold="true" box="[151,339,1551,1573]" pageId="7" pageNumber="8">FIGURES 18–26.</emphasis>
|
||
Scanning electron microscope photographs of
|
||
<taxonomicName id="F8E64D3E9E6640265009CF3161B79B76" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[813,1015,1551,1573]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guttata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6640265009CF3161B79B76" box="[813,1015,1551,1573]" italics="true" pageId="7" pageNumber="8">Drassodella guttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E66402650D9CF31660D9B76" bold="true" box="[1021,1101,1551,1573]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="16A157D49E66402650D9CF31660D9B76" box="[1021,1101,1551,1573]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female (18–21, 24–26) and male (23), and
|
||
<taxonomicName id="F8E64D3E9E66402653DDCF1163249B10" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[249,356,1581,1604]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E66402653DDCF1163249B10" box="[249,356,1581,1604]" italics="true" pageId="7" pageNumber="8">D. tolkieni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E664026524FCF1363FB9B10" bold="true" box="[363,443,1581,1603]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="16A157D49E664026524FCF1363FB9B10" box="[363,443,1581,1603]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F5936BD9E66402653E3CE3161B79AC0" blockId="7.[151,1436,1807,1940]" pageId="7" pageNumber="8">
|
||
Sternum shield-shaped, longer than broad, with blunt apex posteriorly (
|
||
<figureCitation id="A7DD2A389E66402650D4CE3166079A74" box="[1008,1095,1807,1832]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="7" pageNumber="8">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6640265771CE3166359A7B" box="[1109,1141,1807,1832]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="7" pageNumber="8">26</figureCitation>
|
||
); anterior margin concave or straight, widest at coxae II. Intercoxal sclerites present between coxae I & II, II & III, III & IV; precoxal triangles present; pleural bars weakly to moderately sclerotised, isolated or fused (
|
||
<figureCitation id="A7DD2A389E664026576ACE6966D59A3C" box="[1102,1173,1879,1904]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
|
||
). Pedicel composed of broad dorsal sclerite and inverted Y- or V-shaped ventral sclerite (
|
||
<figureCitation id="A7DD2A389E6640265053CE4561AB9AC0" box="[887,1003,1915,1940]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="7" pageNumber="8">Figs 9–11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="6B9966359E69402953B3CF0B639F9A71" httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,264,1589,1611]" targetBox="[158,1429,193,1568]" targetPageId="8">
|
||
<paragraph id="3F5936BD9E69402953B3CF0B639F9A71" blockId="8.[151,1437,1589,1826]" pageId="8" pageNumber="9">
|
||
<emphasis id="0D92EAAF9E69402953B3CF0B63169B18" bold="true" box="[151,342,1589,1611]" pageId="8" pageNumber="9">FIGURES 27–38.</emphasis>
|
||
Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34)
|
||
<taxonomicName id="F8E64D3E9E6940295604CF0B629F9B39" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="guttata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6940295604CF0B629F9B39" italics="true" pageId="8" pageNumber="9">Drassodella guttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E69402953C0CF6A63749B39" bold="true" box="[228,308,1620,1642]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="16A157D49E69402953C0CF6A63749B39" box="[228,308,1620,1642]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and female
|
||
<taxonomicName id="F8E64D3E9E694029528BCF6B605A9B39" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[431,538,1620,1642]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E694029528BCF6B605A9B39" box="[431,538,1620,1642]" italics="true" pageId="8" pageNumber="9">D. tolkieni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E694029513BCF6A602F9B39" bold="true" box="[543,623,1620,1642]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="16A157D49E694029513BCF6A602F9B39" box="[543,623,1620,1642]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F5936BD9E69402853E3CE5866FA9C36" blockId="8.[151,1437,1893,2026]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
Legs long and moderately thin (
|
||
<figureCitation id="A7DD2A389E694029511CCE5B60DF9A2D" box="[568,671,1893,1918]" captionStart="FIGURES 1–4" captionStartId="5.[151,264,1326,1348]" captionTargetBox="[151,1435,406,1306]" captionTargetId="figure@5.[151,1435,406,1306]" captionTargetPageId="5" captionText="FIGURES 1–4. General habitus photographs of live Drassodella spiders: 1 D. amatola sp. nov. female from Hogsback, Eastern Cape; 2 D. quinquelabecula Tucker, 1923 female from Aardvark Nature Reserve, Western Cape; 3 D. septemmaculata (Strand, 1909) male from Fisherhaven, Western Cape; 4 D. vasivulva Tucker, 1923 male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb." httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="8" pageNumber="9">Figs 1–4</figureCitation>
|
||
), leg formula 4123; anterior pairs of legs weakly spined, posterior pairs of legs moderately to strongly spined; trochanters notched; femora with curved elongate ventral setae (
|
||
<figureCitation id="A7DD2A389E694029564ECEB762A49A95" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="8" pageNumber="9">Figs 27, 28</figureCitation>
|
||
); patellar indentation narrow, with large lyriform organ at proximal end (
|
||
<figureCitation id="A7DD2A389E694029571ECE9366839A95" box="[1082,1219,1965,1990]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="8" pageNumber="9">Figs 29, 30</figureCitation>
|
||
); tibiae with distal lyriform organ retrolaterally (
|
||
<figureCitation id="A7DD2A389E69402952D9CEEF601A9AB9" box="[509,602,2001,2026]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="8" pageNumber="9">Fig. 31</figureCitation>
|
||
); tibiae, metatarsi and tarsi with several trichobothria, feathery and chemosensory setae (
|
||
<figureCitation id="A7DD2A389E68402852AEC9A960259DE3" box="[394,613,151,176]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="9" pageNumber="10">Figs 31–33, 35–37</figureCitation>
|
||
); posterior legs with single or paired ventral metatarsal spines; feathery setae present on all leg segments (e.g.
|
||
<figureCitation id="A7DD2A389E6840285166C98260AA9D87" box="[578,746,188,213]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="9" pageNumber="10">Figs 29, 33, 34</figureCitation>
|
||
), usually dorsally and laterally situated on metatarsi and tarsi; metatarsi and tarsi weakly scopulate (
|
||
<figureCitation id="A7DD2A389E684028511BC9E160D69DAB" box="[575,662,223,248]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="9" pageNumber="10">Figs 34</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6840285187C9DE60AD9DAB" box="[675,749,224,248]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="9" pageNumber="10">39–42</figureCitation>
|
||
), anterior metatarsi usually scopulate in distal half only, tarsi scopulate for entire length; tarsal organ oval and slightly elevated (
|
||
<figureCitation id="A7DD2A389E68402850A5C83A61959C4F" box="[897,981,260,285]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="9" pageNumber="10">Fig. 38</figureCitation>
|
||
); tarsi with paired dentate claws, usually with 3-5 teeth, situated dorsally to claw tufts comprising tenent setae (
|
||
<figureCitation id="A7DD2A389E684028509FC81966539C13" box="[955,1043,295,320]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="9" pageNumber="10">Figs 35</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6840285705C81666019C13" box="[1057,1089,296,320]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="9" pageNumber="10">43</figureCitation>
|
||
); claw tufts denser on anterior (
|
||
<figureCitation id="A7DD2A389E68402853BBC87262B19C37" box="[159,241,332,357]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="9" pageNumber="10">Fig. 40</figureCitation>
|
||
) than posterior tarsi (
|
||
<figureCitation id="A7DD2A389E68402852C5C87260759C37" box="[481,565,332,357]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="9" pageNumber="10">Fig. 43</figureCitation>
|
||
); females with ventral row of erect setae on palpal femur.
|
||
</paragraph>
|
||
<caption id="6B9966359E68402853B3CFD6612E9A87" httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1768,1790]" targetBox="[161,1426,399,1747]" targetPageId="9">
|
||
<paragraph id="3F5936BD9E68402853B3CFD6612E9A87" blockId="9.[151,1436,1768,2004]" pageId="9" pageNumber="10">
|
||
<emphasis id="0D92EAAF9E68402853B3CFD663169BAD" bold="true" box="[151,342,1768,1790]" pageId="9" pageNumber="10">FIGURES 39–50.</emphasis>
|
||
Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43)
|
||
<taxonomicName id="F8E64D3E9E6840285604CFD6629F9A4E" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="guttata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6840285604CFD6629F9A4E" italics="true" pageId="9" pageNumber="10">Drassodella guttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E68402853C2CE3863779A4F" bold="true" box="[230,311,1798,1820]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="16A157D49E68402853C2CE3863779A4F" box="[230,311,1798,1820]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and female
|
||
<taxonomicName id="F8E64D3E9E6840285299CE3660699A4F" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[445,553,1798,1821]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6840285299CE3660699A4F" box="[445,553,1798,1821]" italics="true" pageId="9" pageNumber="10">D. tolkieni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6840285114CE3860C19A4F" bold="true" box="[560,641,1798,1820]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="16A157D49E6840285114CE3860C19A4F" box="[560,641,1798,1820]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F5936BD9E6B402B53E3C9A967209CFF" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="11">
|
||
Abdomen oval-elongate, with yellow, white or cream dorsal stripes or spots, or without markings, on black or grey background (
|
||
<figureCitation id="A7DD2A389E6B402B5240C98263899D87" box="[356,457,188,213]" captionStart="FIGURES 1–4" captionStartId="5.[151,264,1326,1348]" captionTargetBox="[151,1435,406,1306]" captionTargetId="figure@5.[151,1435,406,1306]" captionTargetPageId="5" captionText="FIGURES 1–4. General habitus photographs of live Drassodella spiders: 1 D. amatola sp. nov. female from Hogsback, Eastern Cape; 2 D. quinquelabecula Tucker, 1923 female from Aardvark Nature Reserve, Western Cape; 3 D. septemmaculata (Strand, 1909) male from Fisherhaven, Western Cape; 4 D. vasivulva Tucker, 1923 male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb." httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="10" pageNumber="11">Figs 1–4</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6B402B52F2C98260609D87" box="[470,544,188,212]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">12–14</figureCitation>
|
||
); dorsum covered in short erect and feathery setae, markings comprising dense feathery setae (
|
||
<figureCitation id="A7DD2A389E6B402B5265C9E163D69DAB" box="[321,406,223,248]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="10" pageNumber="11">Fig. 44</figureCitation>
|
||
); dorsum with two pairs of small sigilla, posterior pair more widely separated (
|
||
<figureCitation id="A7DD2A389E6B402B5629C9E167CE9DAB" box="[1293,1422,223,248]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Figs 12, 14</figureCitation>
|
||
), often with additional pair of sclerites on anterior margin above pedicel (
|
||
<figureCitation id="A7DD2A389E6B402B50F3C83A666A9C4E" box="[983,1066,260,285]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Fig. 15</figureCitation>
|
||
) and laterally above booklungs; males with short dorsal abdominal scutum anteriorly, absent in females; several fine straight erect setae on anterior margin above pedicel; pale ventrally (
|
||
<figureCitation id="A7DD2A389E6B402B516CC87260DD9C36" box="[584,669,332,357]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Fig. 16</figureCitation>
|
||
), with one or two lines of tiny sclerites usually present, sometimes absent, with few scattered feathery setae between short straight setae (
|
||
<figureCitation id="A7DD2A389E6B402B50DAC851661B9CDB" box="[1022,1115,367,392]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="10" pageNumber="11">Fig. 45</figureCitation>
|
||
); epigastric region weakly sclerotised, and post-epigastric sclerites weakly to strongly sclerotized (
|
||
<figureCitation id="A7DD2A389E6B402B509FC8AA664D9CFE" box="[955,1037,404,429]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Fig. 16</figureCitation>
|
||
), between short straight setae.
|
||
</paragraph>
|
||
<paragraph id="3F5936BD9E6B402B53E3C88961219FD7" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="11">
|
||
Spinnerets (analysed with SEM observations only in females of
|
||
<taxonomicName id="F8E64D3E9E6B402B5085C88766549C83" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[929,1044,441,464]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="guttata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6B402B5085C88766549C83" box="[929,1044,441,464]" italics="true" pageId="10" pageNumber="11">D. guttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6B402B5738C88966379C83" bold="true" box="[1052,1143,439,464]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="16A157D49E6B402B5738C88966379C83" box="[1052,1143,439,464]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="F8E64D3E9E6B402B5794C88767689C83" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[1200,1320,439,464]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6B402B5794C88767689C83" box="[1200,1320,439,464]" italics="true" pageId="10" pageNumber="11">D. tolkieni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6B402B5614C88967CB9C83" bold="true" box="[1328,1419,439,464]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="16A157D49E6B402B5614C88967CB9C83" box="[1328,1419,439,464]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
): ALS conical, nearly cylindrical, not widely separated, with sclerotized apical segment (
|
||
<figureCitation id="A7DD2A389E6B402B5752C8E2668E9CA7" box="[1142,1230,476,501]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6B402B57FFC8E266BB9CA7" box="[1243,1275,476,500]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="10" pageNumber="11">47</figureCitation>
|
||
); single major ampullate gland spigot and numerous piriform gland spigots present (
|
||
<figureCitation id="A7DD2A389E6B402B509CC8C1664E9F4B" box="[952,1038,511,536]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="10" pageNumber="11">Fig. 48</figureCitation>
|
||
); PMS with two (
|
||
<figureCitation id="A7DD2A389E6B402B57FAC8C167719F4B" box="[1246,1329,511,536]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Fig. 17</figureCitation>
|
||
) or three rows of cylindrical gland spigots, single minor ampullate gland spigot and many aciniform gland spigots (
|
||
<figureCitation id="A7DD2A389E6B402B561FCB1A67CF9F6F" box="[1339,1423,548,573]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="10" pageNumber="11">Fig. 49</figureCitation>
|
||
); PLS with aciniform gland spigots, cylindrical gland spigots, single minor ampullate gland spigot and nubbins (
|
||
<figureCitation id="A7DD2A389E6B402B564BCB7962F79FD6" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="10" pageNumber="11">Fig. 50</figureCitation>
|
||
); PLS longer and more widely separated than ALS (
|
||
<figureCitation id="A7DD2A389E6B402B5024CB5261139FD7" box="[768,851,620,645]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="10" pageNumber="11">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="6B9966359E6B402B53B3CFF2616C9AC9" httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1740,1762]" targetBox="[157,1429,700,1719]" targetPageId="10">
|
||
<paragraph id="3F5936BD9E6B402B53B3CFF2616C9AC9" blockId="10.[151,1436,1740,1946]" pageId="10" pageNumber="11">
|
||
<emphasis id="0D92EAAF9E6B402B53B3CFF263169BB1" bold="true" box="[151,342,1740,1762]" pageId="10" pageNumber="11">FIGURES 51–59.</emphasis>
|
||
Scanning electron microscope photographs of female (51, 52) and male (55–59)
|
||
<taxonomicName id="F8E64D3E9E6B402B578DCFF267359BB1" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[1193,1397,1740,1762]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="guttata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6B402B578DCFF267359BB1" box="[1193,1397,1740,1762]" italics="true" pageId="10" pageNumber="11">Drassodella guttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6B402B5659CFF262839A52" bold="true" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="16A157D49E6B402B5659CFF262839A52" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and female
|
||
<taxonomicName id="F8E64D3E9E6B402B526DCFD263F59A52" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[329,437,1771,1793]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E6B402B526DCFD263F59A52" box="[329,437,1771,1793]" italics="true" pageId="10" pageNumber="11">D. tolkieni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E6B402B5298CFD5604D9A52" bold="true" box="[444,525,1771,1793]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="16A157D49E6B402B5298CFD5604D9A52" box="[444,525,1771,1793]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F5936BD9E6A402A53E3C9A966EC9CDB" blockId="11.[151,1437,151,608]" pageId="11" pageNumber="12">
|
||
Female epigynes with distinct anterior hoods, lateral hoods only present in
|
||
<taxonomicName id="F8E64D3E9E6A402A5730C9A766CA9DE3" authorityName="Tucker" authorityYear="1923" box="[1044,1162,151,176]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melana">
|
||
<emphasis id="0D92EAAF9E6A402A5730C9A766CA9DE3" box="[1044,1162,151,176]" italics="true" pageId="11" pageNumber="12">D. melana</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A561EC9A967CE9DE3" box="[1338,1422,151,176]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Fig. 51</figureCitation>
|
||
); anterior hood transverse, median septum broad and subrectangular in
|
||
<taxonomicName id="F8E64D3E9E6A402A50F4C983660A9D86" authorityName="Tucker" authorityYear="1923" box="[976,1098,188,213]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melana">
|
||
<emphasis id="0D92EAAF9E6A402A50F4C983660A9D86" box="[976,1098,188,213]" italics="true" pageId="11" pageNumber="12">D. melana</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A562CC98267D09D86" box="[1288,1424,188,213]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="11" pageNumber="12">Figs 60–69</figureCitation>
|
||
), variable but generally more longitudinal in
|
||
<taxonomicName id="F8E64D3E9E6A402A51A7C9DF61549DAB" authorityName="Hewitt" authorityYear="1916" box="[643,788,223,248]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="salisburyi">
|
||
<emphasis id="0D92EAAF9E6A402A51A7C9DF61549DAB" box="[643,788,223,248]" italics="true" pageId="11" pageNumber="12">D. salisburyi</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A50E2C9E1660A9DAB" box="[966,1098,223,248]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="11" pageNumber="12">Figs 70–77</figureCitation>
|
||
); epigyne muscle sigilla (
|
||
<figureCitation id="A7DD2A389E6A402A564EC9E162A79C4E" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Figs 51, 52</figureCitation>
|
||
) broad or elongate, sometimes absent; longitudinal ridge of median septum sometimes broadened posteriorly; copulatory openings forming narrow slit laterally on median septum (
|
||
<figureCitation id="A7DD2A389E6A402A571EC81966CF9C13" box="[1082,1167,295,320]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Fig. 51</figureCitation>
|
||
); copulatory ducts very short, entering ST II along mesal margin (
|
||
<figureCitation id="A7DD2A389E6A402A514DC87260FC9C36" box="[617,700,332,357]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Fig. 53</figureCitation>
|
||
); ST II anterior to ST I, with broad, medially-curving duct linking them (
|
||
<figureCitation id="A7DD2A389E6A402A53FAC85163219CDB" box="[222,353,367,392]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="11" pageNumber="12">Figs 60–69</figureCitation>
|
||
); ST II with widely spaced glandular pores on spermathecal head (
|
||
<figureCitation id="A7DD2A389E6A402A576FC85166E09CDB" box="[1099,1184,367,392]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Fig. 54</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3F5936BD9E6A402A53E3C8AA672D9F33" blockId="11.[151,1437,151,608]" pageId="11" pageNumber="12">
|
||
Male palp with broad oval cymbium, tapering distally, with sparse to dense cymbial scopula present dorsally (
|
||
<figureCitation id="A7DD2A389E6A402A53BBC88963629C83" box="[159,290,439,464]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Figs 55–57</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6A402A520BC886633A9C83" box="[303,378,440,464]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="11" pageNumber="12">78–92</figureCitation>
|
||
); embolus in
|
||
<taxonomicName id="F8E64D3E9E6A402A5131C88760CC9C83" authorityName="Tucker" authorityYear="1923" box="[533,652,439,464]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melana">
|
||
<emphasis id="0D92EAAF9E6A402A5131C88760CC9C83" box="[533,652,439,464]" italics="true" pageId="11" pageNumber="12">D. melana</emphasis>
|
||
</taxonomicName>
|
||
species group originating prolaterally, forming broad arch, with fine retrolateral tip (
|
||
<figureCitation id="A7DD2A389E6A402A5262C8E263899CA6" box="[326,457,476,501]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Figs 57–59</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E6A402A52F1C8E2605E9CA6" box="[469,542,476,501]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="11" pageNumber="12">78–85</figureCitation>
|
||
), subtriangular and variably orientated in
|
||
<taxonomicName id="F8E64D3E9E6A402A50D2C8E366C79CA6" authorityName="Hewitt" authorityYear="1916" box="[1014,1159,476,501]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="salisburyi">
|
||
<emphasis id="0D92EAAF9E6A402A50D2C8E366C79CA6" box="[1014,1159,476,501]" italics="true" pageId="11" pageNumber="12">D. salisburyi</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A561CC8E262F79F4B" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="11" pageNumber="12">Figs 86– 92</figureCitation>
|
||
); embolus with one or two basal prongs prolaterally in
|
||
<taxonomicName id="F8E64D3E9E6A402A5066CB3F61FA9F4B" authorityName="Tucker" authorityYear="1923" box="[834,954,511,536]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melana">
|
||
<emphasis id="0D92EAAF9E6A402A5066CB3F61FA9F4B" box="[834,954,511,536]" italics="true" pageId="11" pageNumber="12">D. melana</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A5756C8C166B79F4B" box="[1138,1271,511,536]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Figs 57–59</figureCitation>
|
||
), absent in
|
||
<taxonomicName id="F8E64D3E9E6A402A56A5CB3F63459F6E" authorityName="Hewitt" authorityYear="1916" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="salisburyi">
|
||
<emphasis id="0D92EAAF9E6A402A56A5CB3F63459F6E" italics="true" pageId="11" pageNumber="12">D. salisburyi</emphasis>
|
||
</taxonomicName>
|
||
species group; median apophysis hook-shaped in
|
||
<taxonomicName id="F8E64D3E9E6A402A506FCB1B61829F6E" authorityName="Tucker" authorityYear="1923" box="[843,962,548,573]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melana">
|
||
<emphasis id="0D92EAAF9E6A402A506FCB1B61829F6E" box="[843,962,548,573]" italics="true" pageId="11" pageNumber="12">D. melana</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A575ECB1A66909F6E" box="[1146,1232,548,573]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="11" pageNumber="12">Fig. 59</figureCitation>
|
||
), subtriangular or forming stout hook in
|
||
<taxonomicName id="F8E64D3E9E6A402A52B5CB7760629F33" authorityName="Hewitt" authorityYear="1916" box="[401,546,583,608]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="salisburyi">
|
||
<emphasis id="0D92EAAF9E6A402A52B5CB7760629F33" box="[401,546,583,608]" italics="true" pageId="11" pageNumber="12">D. salisburyi</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<figureCitation id="A7DD2A389E6A402A51F5CB7961149F33" box="[721,852,583,608]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="11" pageNumber="12">Figs 86–92</figureCitation>
|
||
); retrolateral tibial apophysis simple, triangular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |