347 lines
54 KiB
XML
347 lines
54 KiB
XML
<document id="6D6D299B56023E9258801A72DADE0718" ID-CLB-Dataset="27120" ID-DOI="10.11646/zootaxa.4582.1.1" ID-GBIF-Dataset="c79b6327-48fb-47bf-b17b-d20d2de295e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2643724" ID-ZooBank="9DC61794-4BD7-4F6D-BB8C-84D9855C8151" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555512297075" checkinUser="plazi" docAuthor="Mbo, Zingisile & Haddad, Charles R." docDate="2019" docId="B74F87AB9E47400953B3CAD766A99BBA" docLanguage="en" docName="zootaxa.4582.1.1.pdf.imf" docOrigin="Zootaxa 4582 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Drassodella amatola Mbo & Haddad 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="41" masterDocId="4B76FFD39E6140215324C93E62409D53" masterDocTitle="A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="39" updateTime="1698721989978" updateUser="ExternalLinkService">
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<mods:titleInfo id="BF56DBDBA702F4049011DEEAB47CCEEB">
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<mods:title id="03A7515BD4D52EFC9F85FAA654057F9C">A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae)</mods:title>
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<mods:namePart id="1CBA7F0622C35059AF965E00BA021454">Mbo, Zingisile</mods:namePart>
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<mods:roleTerm id="5223780A3E27E639F73FAA812C512B4D">Author</mods:roleTerm>
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<mods:namePart id="AC290C02035A6C6B013556163F37E374">Haddad, Charles R.</mods:namePart>
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<mods:typeOfResource id="1EEC81D4CEC8F691859E010177886691">text</mods:typeOfResource>
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<mods:title id="589F8318618116DA0FB4EBD87CD5FB7B">Zootaxa</mods:title>
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<mods:part id="ADA3FD0D1A07430121D544A7EC835FF8">
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<mods:date id="AB89535B0BF42BDE1F7900110EC08330">2019</mods:date>
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<mods:number id="0D485214EC639494E27736B9C28CE17A">2019-04-10</mods:number>
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<mods:number id="DCA30810305F32B27B717ABDE1A84561">4582</mods:number>
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<mods:classification id="4C7FB6473AA6033E8C74E32D7FB53262">journal article</mods:classification>
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<mods:identifier id="8C28E579AF7BB5B75FA34E4033711269" type="CLB-Dataset">27120</mods:identifier>
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<mods:identifier id="CC075AEC66B75CFA5A4C9861CF3253FA" type="DOI">10.11646/zootaxa.4582.1.1</mods:identifier>
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<treatment id="B74F87AB9E47400953B3CAD766A99BBA" ID-DOI="http://doi.org/10.5281/zenodo.5662628" ID-GBIF-Taxon="156208946" ID-Zenodo-Dep="5662628" LSID="urn:lsid:plazi:treatment:B74F87AB9E47400953B3CAD766A99BBA" httpUri="http://treatment.plazi.org/id/B74F87AB9E47400953B3CAD766A99BBA" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
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<subSubSection id="77FC65369E47400753B3CAD763B49950" box="[151,500,1001,1027]" pageId="38" pageNumber="39" type="nomenclature">
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<paragraph id="3F5936BD9E47400753B3CAD763B49950" blockId="38.[151,570,1001,1061]" box="[151,500,1001,1027]" pageId="38" pageNumber="39">
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<heading id="641181D19E47400753B3CAD763B49950" bold="true" box="[151,500,1001,1027]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
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<emphasis id="0D92EAAF9E47400753B3CAD763B49950" bold="true" box="[151,500,1001,1027]" pageId="38" pageNumber="39">
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<taxonomicName id="F8E64D3E9E47400753B3CAD763CF9950" ID-CoL="37LKM" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[151,399,1001,1027]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="amatola" status="sp. nov.">
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<emphasis id="0D92EAAF9E47400753B3CAD763CF9950" bold="true" box="[151,399,1001,1027]" italics="true" pageId="38" pageNumber="39">Drassodella amatola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="16A157D49E47400752B2CAD763B49950" box="[406,500,1001,1027]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="77FC65369E47400753B3CD32607A9976" box="[151,570,1036,1061]" pageId="38" pageNumber="39" type="description">
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<paragraph id="3F5936BD9E47400753B3CD32607A9976" blockId="38.[151,570,1001,1061]" box="[151,570,1036,1061]" pageId="38" pageNumber="39">
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<figureCitation id="A7DD2A389E47400753B3CD32629C9977" box="[151,220,1036,1061]" captionStart="FIGURES 1–4" captionStartId="5.[151,264,1326,1348]" captionTargetBox="[151,1435,406,1306]" captionTargetId="figure@5.[151,1435,406,1306]" captionTargetPageId="5" captionText="FIGURES 1–4. General habitus photographs of live Drassodella spiders: 1 D. amatola sp. nov. female from Hogsback, Eastern Cape; 2 D. quinquelabecula Tucker, 1923 female from Aardvark Nature Reserve, Western Cape; 3 D. septemmaculata (Strand, 1909) male from Fisherhaven, Western Cape; 4 D. vasivulva Tucker, 1923 male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb." httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="38" pageNumber="39">Figs 1</figureCitation>
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,
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<figureCitation id="A7DD2A389E47400753CECD3263489977" box="[234,264,1036,1060]" captionStart="FIGURES 5–17" captionStartId="6.[151,264,1691,1713]" captionTargetBox="[209,1363,193,1648]" captionTargetId="figure@6.[209,1363,193,1648]" captionTargetPageId="6" captionText="FIGURES 5–17. Digital microscope photographs of Drassodella somatic morphology: 5–8 dorsal view of carapace and chelicerae of D. flava sp. nov. female (5), D. guttata sp. nov. male (6), D. transversa sp. nov. male (7) and D. purcelli Tucker, 1923 female (8); 9 D. tenebrosa Lawrence, 1938 female, carapace, lateral view; 10 D. purcelli female, posterodorsal view of carapace, showing pedicel and sclerites; 11 same, ventral pedicel sclerite, ventral view; 12–14 dorsal view of abdomens of D. septemmaculata (Strand, 1909) male (12), D. aurostriata sp. nov. male (13) and D. amatola sp. nov. female (14); 15 D. tenebrosa female, abdomen, anterodorsal view; 16 D. septemmaculata male, abdomen, showing sclerites on venter; 17 D. tenebrosa female, spinnerets, posteroventral view. Abbreviations: AAS—anterior abdominal sclerite; CGS—cylindrical gland spigots; CON—condyle; DPS—dorsal pedicel sclerite; IS—inframamillary sclerite; PB—pleural bars; PES—post-epigastric sclerite; PPS—posterior pleural sclerite; PT—petiole; SC—dorsal scutum; SI—sigillum; VPS—ventral pedicel sclerite." httpUri="https://zenodo.org/record/2643730/files/figure.png" pageId="38" pageNumber="39">14</figureCitation>
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,
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<figureCitation id="A7DD2A389E4740075230CD3263729977" box="[276,306,1036,1060]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="38" pageNumber="39">70</figureCitation>
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,
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<figureCitation id="A7DD2A389E474007521ACD32631C9976" box="[318,348,1036,1061]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="38" pageNumber="39">86</figureCitation>
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,
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<figureCitation id="A7DD2A389E474007524CCD32638C9976" box="[360,460,1036,1061]" captionStart="FIGURES 157–171" captionStartId="39.[151,264,1272,1294]" captionTargetBox="[161,1426,193,1251]" captionTargetId="figure@39.[161,1426,193,1251]" captionTargetPageId="39" captionText="FIGURES 157–171. Dorsal habitus photographs of Drassodella species in the salisburyi species group: 157, 158 female and male of D. amatola sp. nov.; 159, 160 female and male of D. aurostriata sp. nov.; 161 male of D. baviaans sp. nov.; 162 female of D. purcelli Tucker, 1923; 163, 164 female and male of D. quinquelabecula Tucker, 1923; 165, 166 female and male of D. salisburyi Hewitt, 1916; 167, 168 female and male of D. septemmaculata (Strand, 1909); 169 female of D. trilineata sp. nov.; 170, 171 female and male of D. vasivulva Tucker, 1923. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643776/files/figure.png" pageId="38" pageNumber="39">157, 158</figureCitation>
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,
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<figureCitation id="A7DD2A389E47400752FCCD32607A9976" box="[472,570,1036,1061]" captionStart="FIGURES 172–175" captionStartId="40.[151,264,1395,1417]" captionTargetBox="[167,1425,622,1376]" captionTargetId="figure@40.[161,1426,611,1382]" captionTargetPageId="40" captionText="FIGURES 172–175. Genitalic morphology of Drassodella amatola sp. nov.: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="38" pageNumber="39">172–175</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="77FC65369E47400753B3CD6A631599C3" pageId="38" pageNumber="39" type="etymology">
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<paragraph id="3F5936BD9E47400753B3CD6A631599C3" blockId="38.[151,1437,1108,2033]" pageId="38" pageNumber="39">
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<emphasis id="0D92EAAF9E47400753B3CD6A635E993E" bold="true" box="[151,286,1108,1133]" pageId="38" pageNumber="39">Etymology.</emphasis>
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The species name is a noun in apposition of the mountain range (Amatola Mountains) where the
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<typeStatus id="E05D881F9E4740075648CD6B67DC993E" box="[1388,1436,1109,1133]" pageId="38" pageNumber="39">type</typeStatus>
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locality is found.
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</paragraph>
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</subSubSection>
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<subSubSection id="77FC65369E47400753E3CDA261D098DF" pageId="38" pageNumber="39" type="diagnosis">
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<paragraph id="3F5936BD9E47400753E3CDA261D098DF" blockId="38.[151,1437,1108,2033]" pageId="38" pageNumber="39">
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<emphasis id="0D92EAAF9E47400753E3CDA2630199E6" bold="true" box="[199,321,1180,1205]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
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<taxonomicName id="F8E64D3E9E474007526CCDA2607399E6" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[328,563,1180,1205]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="amatola" status="sp. nov.">
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<emphasis id="0D92EAAF9E474007526CCDA2607399E6" box="[328,563,1180,1205]" italics="true" pageId="38" pageNumber="39">Drassodella amatola</emphasis>
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</taxonomicName>
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<emphasis id="0D92EAAF9E474007511ECDA260D399E6" bold="true" box="[570,659,1180,1205]" pageId="38" pageNumber="39">
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<taxonomicNameLabel id="16A157D49E474007511ECDA260D399E6" box="[570,659,1180,1205]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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is distinct amongst species of this group by the absence of any dorsal markings on the abdomen, which is uniformly black. Furthermore, females can easily be distinguished by small, arc-shaped anterior hood, narrow median septum and large ST II that touch along their mesal margin (
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<figureCitation id="A7DD2A389E474007562DCDDA671F99AF" box="[1289,1375,1252,1277]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="38" pageNumber="39">Figs 70</figureCitation>
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,
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<figureCitation id="A7DD2A389E474007564FCDDA62849873" captionStart="FIGURES 172–175" captionStartId="40.[151,264,1395,1417]" captionTargetBox="[167,1425,622,1376]" captionTargetId="figure@40.[161,1426,611,1382]" captionTargetPageId="40" captionText="FIGURES 172–175. Genitalic morphology of Drassodella amatola sp. nov.: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="38" pageNumber="39">172, 173</figureCitation>
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), while clearly separated in the other species (
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<figureCitation id="A7DD2A389E47400751F1CC39611A9873" box="[725,858,1287,1312]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="38" pageNumber="39">Figs 71–77</figureCitation>
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). Males are easily distinguished from others in the genus by the slightly curved transverse embolus and the strongly prolaterally protruding basal half of the tegulum (
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<figureCitation id="A7DD2A389E47400753BBCC7162B7983B" box="[159,247,1359,1384]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="38" pageNumber="39">Figs 86</figureCitation>
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,
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<figureCitation id="A7DD2A389E4740075220CC6E632C983B" box="[260,364,1359,1384]" captionStart="FIGURES 172–175" captionStartId="40.[151,264,1395,1417]" captionTargetBox="[167,1425,622,1376]" captionTargetId="figure@40.[161,1426,611,1382]" captionTargetPageId="40" captionText="FIGURES 172–175. Genitalic morphology of Drassodella amatola sp. nov.: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="38" pageNumber="39">174, 175</figureCitation>
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). They are most similar to
|
||
<taxonomicName id="F8E64D3E9E4740075185CC6F616A983B" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[673,810,1359,1384]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="baviaans" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E4740075185CC6F616A983B" box="[673,810,1359,1384]" italics="true" pageId="38" pageNumber="39">D. baviaans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E4740075017CC7161CE983B" bold="true" box="[819,910,1359,1384]" pageId="38" pageNumber="39">
|
||
<taxonomicNameLabel id="16A157D49E4740075017CC7161CE983B" box="[819,910,1359,1384]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but can be recognized by the smaller median apophysis and the broader embolus tip (compare
|
||
<figureCitation id="A7DD2A389E474007519ACC4A616098DF" box="[702,800,1396,1421]" captionStart="FIGURES 172–175" captionStartId="40.[151,264,1395,1417]" captionTargetBox="[167,1425,622,1376]" captionTargetId="figure@40.[161,1426,611,1382]" captionTargetPageId="40" captionText="FIGURES 172–175. Genitalic morphology of Drassodella amatola sp. nov.: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="38" pageNumber="39">Figs 174</figureCitation>
|
||
and
|
||
<figureCitation id="A7DD2A389E4740075072CC4A61C498DF" box="[854,900,1396,1420]" captionStart="FIGURES 180–183" captionStartId="43.[151,264,1905,1927]" captionTargetBox="[167,1422,1059,1880]" captionTargetId="figure@43.[161,1426,1052,1884]" captionTargetPageId="43" captionText="FIGURES 180–183. Genitalic morphology of Drassodella baviaans sp. nov. (180, 181) and D. purcelli Tucker, 1923 (182, 183): 180 male palp, ventral view; 181 same, retrolateral view; 182 female epigyne, ventral view; 183 same, dorsal view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643786/files/figure.png" pageId="38" pageNumber="39">180</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="77FC65369E47400953E3CCA9604A9F6E" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="3F5936BD9E47400753E3CCA9677898AB" blockId="38.[151,1437,1108,2033]" pageId="38" pageNumber="39">
|
||
<emphasis id="0D92EAAF9E47400753E3CCA9613E98E3" bold="true" box="[199,894,1431,1456]" pageId="38" pageNumber="39">
|
||
Description. Female (
|
||
<typeStatus id="E05D881F9E474007529ACCA9606C98E3" box="[446,556,1431,1456]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
|
||
, Hogsback, NCA 2007/1188).
|
||
</emphasis>
|
||
Measurements: CL 2.78, CW 2.14, AL 3.60, AW 2.00, TL 6.52 (6.22–7.36), FL 0.22, SL 1.49, SW 1.32,
|
||
<collectingCountry id="47F1762D9E47400751D3CC82615D9887" box="[759,797,1468,1492]" name="Switzerland" pageId="38" pageNumber="39">CH</collectingCountry>
|
||
0.13, AME-AME 0.07, AME-ALE 0.02, ALE-ALE 0.25, PME-PME 0.19, PME-PLE 0.12, PLE-PLE 0.49, PERW 0.63, MOQAW 0.24, MOQPW 0.28, MOQL 0.17.
|
||
</paragraph>
|
||
<paragraph id="3F5936BD9E47400753E3CF3A662A9B13" blockId="38.[151,1437,1108,2033]" pageId="38" pageNumber="39">Length of leg segments: I 2.00 + 1.16 + 1.56 + 1.52 + 1.16 = 7.40, II 1.96 + 1.04 + 1.40 + 1.36 + 1.14 = 6.90, III 1.70 + 1.00 + 1.24 + 1.36 + 0.96 = 6.26, IV 2.44 + 1.28 + 1.90 + 2.56 + 1.20 = 9.38.</paragraph>
|
||
<paragraph id="3F5936BD9E47400753E3CF7267CC9AA3" blockId="38.[151,1437,1108,2033]" pageId="38" pageNumber="39">
|
||
General appearance as in
|
||
<figureCitation id="A7DD2A389E47400752DCCF7260029B37" box="[504,578,1612,1637]" captionStart="FIGURES 1–4" captionStartId="5.[151,264,1326,1348]" captionTargetBox="[151,1435,406,1306]" captionTargetId="figure@5.[151,1435,406,1306]" captionTargetPageId="5" captionText="FIGURES 1–4. General habitus photographs of live Drassodella spiders: 1 D. amatola sp. nov. female from Hogsback, Eastern Cape; 2 D. quinquelabecula Tucker, 1923 female from Aardvark Nature Reserve, Western Cape; 3 D. septemmaculata (Strand, 1909) male from Fisherhaven, Western Cape; 4 D. vasivulva Tucker, 1923 male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb." httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="38" pageNumber="39">Figs 1</figureCitation>
|
||
and
|
||
<figureCitation id="A7DD2A389E47400751A5CF7260EE9B36" box="[641,686,1612,1637]" captionStart="FIGURES 157–171" captionStartId="39.[151,264,1272,1294]" captionTargetBox="[161,1426,193,1251]" captionTargetId="figure@39.[161,1426,193,1251]" captionTargetPageId="39" captionText="FIGURES 157–171. Dorsal habitus photographs of Drassodella species in the salisburyi species group: 157, 158 female and male of D. amatola sp. nov.; 159, 160 female and male of D. aurostriata sp. nov.; 161 male of D. baviaans sp. nov.; 162 female of D. purcelli Tucker, 1923; 163, 164 female and male of D. quinquelabecula Tucker, 1923; 165, 166 female and male of D. salisburyi Hewitt, 1916; 167, 168 female and male of D. septemmaculata (Strand, 1909); 169 female of D. trilineata sp. nov.; 170, 171 female and male of D. vasivulva Tucker, 1923. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643776/files/figure.png" pageId="38" pageNumber="39">157</figureCitation>
|
||
. Carapace black; white feathery setae forming upsilon marking, running from PME towards posterior margin, diverging after fovea. Clypeus height equal to 1.86 AME diameter; AME smaller than ALE; AME separated by distance equal to their diameter, AME separated from ALE by distance equal to 0.29 AME diameter; PME smaller than PLE; PME separated by distance equal to 2.38 their diameter, PME separated from PLE by distance equal to 1.50 PME diameter. Chelicerae dark-brown, yellow at distal margin; promargin with three teeth, median tooth largest, proximal tooth smallest, median tooth closer to proximal tooth than distal tooth; retromargin with two teeth, distal tooth slightly larger. Endites dark brown, mottled on promarginal surface and medially. Labium dark brown, anterior margin pale yellow at maxillar hair tuft; slightly longer than wide. Sternum brown. Legs with proximal segments dark-brown, with black mottling, distal segments orange. Leg spination: femora: I do 1, II do 1, III do 5, IV do 2; patellae: spineless; tibiae: I & II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 2 plv 2 rlv 2 vt 2; metatarsi: I & II spineless, III pl 4 rl 4 plv 2 rlv 2, IV pl 4 do 2 rl 4 plv 1 rlv 2 vt 2; palpal spination: femora do 2 rl 1; patellae do 1 rl 1; tibiae do 1 rl 2; tarsi do 2 rl 2 plv 1.
|
||
</paragraph>
|
||
<caption id="6B9966359E46400653B3CDC6619298DA" httpUri="https://zenodo.org/record/2643776/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,264,1272,1294]" targetBox="[161,1426,193,1251]" targetPageId="39">
|
||
<paragraph id="3F5936BD9E46400653B3CDC6619298DA" blockId="39.[151,1436,1272,1417]" pageId="39" pageNumber="40">
|
||
<emphasis id="0D92EAAF9E46400653B3CDC6632E985D" bold="true" box="[151,366,1272,1294]" pageId="39" pageNumber="40">FIGURES 157–171.</emphasis>
|
||
Dorsal habitus photographs of
|
||
<taxonomicName id="F8E64D3E9E4640065195CDC6616D985D" authorityName="Hewitt" authorityYear="1916" box="[689,813,1272,1294]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0D92EAAF9E4640065195CDC6616D985D" box="[689,813,1272,1294]" italics="true" pageId="39" pageNumber="40">Drassodella</emphasis>
|
||
</taxonomicName>
|
||
species in the
|
||
<taxonomicName id="F8E64D3E9E46400650E1CDC66669985D" authorityName="Hewitt" authorityYear="1916" box="[965,1065,1272,1294]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="salisburyi">
|
||
<emphasis id="0D92EAAF9E46400650E1CDC66669985D" box="[965,1065,1272,1294]" italics="true" pageId="39" pageNumber="40">salisburyi</emphasis>
|
||
</taxonomicName>
|
||
species group: 157, 158 female and male of
|
||
<taxonomicName id="F8E64D3E9E46400653CBCC266323987F" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[239,355,1302,1325]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="amatola" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E46400653CBCC266323987F" box="[239,355,1302,1325]" italics="true" pageId="39" pageNumber="40">D. amatola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="16A157D49E4640065248CC2863FF987F" box="[364,447,1302,1324]" pageId="39" pageNumber="40" rank="species">
|
||
<emphasis id="0D92EAAF9E4640065248CC2863FF987F" bold="true" box="[364,447,1302,1324]" pageId="39" pageNumber="40">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 159, 160 female and male of
|
||
<taxonomicName id="F8E64D3E9E4640065023CC2661DA987E" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[775,922,1304,1325]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="aurostriata" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E4640065023CC2661DA987E" box="[775,922,1304,1325]" italics="true" pageId="39" pageNumber="40">D. aurostriata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E4640065086CC2861B5987F" bold="true" box="[930,1013,1302,1324]" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="16A157D49E4640065086CC2861B5987F" box="[930,1013,1302,1324]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 161 male of
|
||
<taxonomicName id="F8E64D3E9E46400657A9CC26674A987F" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[1165,1290,1302,1325]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="baviaans" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E46400657A9CC26674A987F" box="[1165,1290,1302,1325]" italics="true" pageId="39" pageNumber="40">D. baviaans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E4640065637CC286726987F" bold="true" box="[1299,1382,1302,1324]" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="16A157D49E4640065637CC286726987F" box="[1299,1382,1302,1324]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 162 female of
|
||
<taxonomicName id="F8E64D3E9E46400653DACC0863BB9818" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[254,507,1333,1355]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="purcelli">
|
||
<emphasis id="0D92EAAF9E46400653DACC08632C9818" box="[254,364,1333,1355]" italics="true" pageId="39" pageNumber="40">D. purcelli</emphasis>
|
||
Tucker, 1923
|
||
</taxonomicName>
|
||
; 163, 164 female and male of
|
||
<taxonomicName id="F8E64D3E9E4640065015CC0866C99818" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[817,1161,1333,1355]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="quinquelabecula">
|
||
<emphasis id="0D92EAAF9E4640065015CC0861BA9818" box="[817,1018,1333,1355]" italics="true" pageId="39" pageNumber="40">D. quinquelabecula</emphasis>
|
||
Tucker, 1923
|
||
</taxonomicName>
|
||
; 165, 166 female and male of
|
||
<taxonomicName id="F8E64D3E9E4640065397CC6B63859839" authority="Hewitt, 1916" authorityName="Hewitt" authorityYear="1916" box="[179,453,1364,1386]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="salisburyi">
|
||
<emphasis id="0D92EAAF9E4640065397CC6B63769839" box="[179,310,1364,1386]" italics="true" pageId="39" pageNumber="40">D. salisburyi</emphasis>
|
||
Hewitt, 1916
|
||
</taxonomicName>
|
||
; 167, 168 female and male of
|
||
<taxonomicName id="F8E64D3E9E46400651DFCC6B661B9839" authority="(Strand, 1909)" authorityName="Strand" authorityYear="1909" baseAuthorityName="Strand" baseAuthorityYear="1909" box="[763,1115,1364,1386]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="septemmaculata">
|
||
<emphasis id="0D92EAAF9E46400651DFCC6B61FE9839" box="[763,958,1364,1386]" italics="true" pageId="39" pageNumber="40">D. septemmaculata</emphasis>
|
||
(Strand, 1909)
|
||
</taxonomicName>
|
||
; 169 female
|
||
<emphasis id="0D92EAAF9E46400657FACC6A67379839" box="[1246,1399,1364,1386]" italics="true" pageId="39" pageNumber="40">
|
||
of
|
||
<taxonomicName id="F8E64D3E9E46400657DDCC6B67379839" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[1273,1399,1364,1386]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="trilineata" status="sp. nov.">D. trilineata</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="0D92EAAF9E4640065659CC6A628398DA" bold="true" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="16A157D49E4640065659CC6A628398DA" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 170, 171 female and male of
|
||
<taxonomicName id="F8E64D3E9E46400652DCCC4A614598DA" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[504,773,1395,1417]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vasivulva">
|
||
<emphasis id="0D92EAAF9E46400652DCCC4A603798DA" box="[504,631,1395,1417]" italics="true" pageId="39" pageNumber="40">D. vasivulva</emphasis>
|
||
Tucker, 1923
|
||
</taxonomicName>
|
||
. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3F5936BD9E46400653B3CCF066739BDD" blockId="39.[151,1437,1486,2036]" pageId="39" pageNumber="40">
|
||
Abdomen uniformly black (
|
||
<figureCitation id="A7DD2A389E46400652F2CCF0605F98B4" box="[470,543,1486,1511]" captionStart="FIGURES 1–4" captionStartId="5.[151,264,1326,1348]" captionTargetBox="[151,1435,406,1306]" captionTargetId="figure@5.[151,1435,406,1306]" captionTargetPageId="5" captionText="FIGURES 1–4. General habitus photographs of live Drassodella spiders: 1 D. amatola sp. nov. female from Hogsback, Eastern Cape; 2 D. quinquelabecula Tucker, 1923 female from Aardvark Nature Reserve, Western Cape; 3 D. septemmaculata (Strand, 1909) male from Fisherhaven, Western Cape; 4 D. vasivulva Tucker, 1923 male from De Hoop Nature Reserve, Western Cape. Photographs 1, 3, 4 by C. Haddad, 2 by P. Webb." httpUri="https://zenodo.org/record/2643726/files/figure.png" pageId="39" pageNumber="40">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E464006510BCCF0601C98B4" box="[559,604,1486,1511]" captionStart="FIGURES 157–171" captionStartId="39.[151,264,1272,1294]" captionTargetBox="[161,1426,193,1251]" captionTargetId="figure@39.[161,1426,193,1251]" captionTargetPageId="39" captionText="FIGURES 157–171. Dorsal habitus photographs of Drassodella species in the salisburyi species group: 157, 158 female and male of D. amatola sp. nov.; 159, 160 female and male of D. aurostriata sp. nov.; 161 male of D. baviaans sp. nov.; 162 female of D. purcelli Tucker, 1923; 163, 164 female and male of D. quinquelabecula Tucker, 1923; 165, 166 female and male of D. salisburyi Hewitt, 1916; 167, 168 female and male of D. septemmaculata (Strand, 1909); 169 female of D. trilineata sp. nov.; 170, 171 female and male of D. vasivulva Tucker, 1923. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643776/files/figure.png" pageId="39" pageNumber="40">157</figureCitation>
|
||
); lateral sigilla above booklungs sub-oval; venter pale with sparse hairs; two paired rows of tiny sclerites running from epigastric fold to spinnerets, inner rows with fewer sclerites. Spinnerets brown. Epigyne with small shallow anterior hood, anterior and posterior margins strongly recurved, lateral ends slightly rounded; lateral margins of median septum weakly sclerotised, converging mesally from anterior hood posteriorly to ST II; ST I, ST II and connecting ducts orange-brown; ST II bean-shaped, touching along their mesal margins, much larger than subspherical ST I (
|
||
<figureCitation id="A7DD2A389E4640065079CF4B61F49BDD" box="[861,948,1653,1678]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="39" pageNumber="40">Figs 70</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E46400650E4CF4866669BDD" box="[960,1062,1654,1678]" captionStart="FIGURES 172–175" captionStartId="40.[151,264,1395,1417]" captionTargetBox="[167,1425,622,1376]" captionTargetId="figure@40.[161,1426,611,1382]" captionTargetPageId="40" captionText="FIGURES 172–175. Genitalic morphology of Drassodella amatola sp. nov.: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="39" pageNumber="40">172, 173</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3F5936BD9E46400653E3CFA8668D9BA4" blockId="39.[151,1437,1486,2036]" pageId="39" pageNumber="40">
|
||
<emphasis id="0D92EAAF9E46400653E3CFA860909BFC" bold="true" box="[199,720,1686,1711]" pageId="39" pageNumber="40">
|
||
Male (
|
||
<typeStatus id="E05D881F9E4640065236CFA863C29BFC" box="[274,386,1686,1711]" pageId="39" pageNumber="40" type="paratype">Paratype</typeStatus>
|
||
, Hogsback,
|
||
<bibRefCitation id="5B774B4C9E4640065136CFA860D09BFC" author="Dippenaar-Schoeman, A. S. & Haddad, C. R." box="[530,656,1686,1711]" pageId="39" pageNumber="40" refId="ref44277" refString="Dippenaar-Schoeman, A. S. & Haddad, C. R. (2014) Spiders of the Grassland Biome. Agricultural Research Council, Pretoria, 122 pp." type="book" year="2014">NCA 2014</bibRefCitation>
|
||
/699).
|
||
</emphasis>
|
||
Measurements: CL 2.80, CW 2.08, AL 2.72, AW 1.40, TL 6.48 (6.32–6.68), FL 0.48, SL 1.42, SW 1.20,
|
||
<collectingCountry id="47F1762D9E4640065142CF8560CD9B80" box="[614,653,1723,1747]" name="Switzerland" pageId="39" pageNumber="40">CH</collectingCountry>
|
||
0.15, AME-AME 0.08, AME-ALE 0.02, ALE-ALE 0.25, PME-PME 0.10, PME-PLE 0.14, PLE-PLE 0.55, PERW 0.65, MOQAW 0.21, MOQPW 0.25, MOQL 0.32.
|
||
</paragraph>
|
||
<paragraph id="3F5936BD9E46400653E3CE3D662A9A6C" blockId="39.[151,1437,1486,2036]" pageId="39" pageNumber="40">Length of leg segments: I 2.08 + 1.04 + 1.60 + 1.44 + 1.14 = 7.30, II 1.80 + 0.96 + 1.36 + 1.36 + 1.06 = 6.54, III 1.64 + 0.84 + 1.06 + 1.46 + 0.88 = 5.88, IV 2.44 + 1.12 + 1.62 + 2.50 + 1.06 = 8.74.</paragraph>
|
||
<paragraph id="3F5936BD9E46400953E3CE75604A9F6E" blockId="39.[151,1437,1486,2036]" lastBlockId="40.[151,1437,151,573]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
|
||
General appearance as in
|
||
<figureCitation id="A7DD2A389E46400652C9CE75600E9A37" box="[493,590,1867,1892]" captionStart="FIGURES 157–171" captionStartId="39.[151,264,1272,1294]" captionTargetBox="[161,1426,193,1251]" captionTargetId="figure@39.[161,1426,193,1251]" captionTargetPageId="39" captionText="FIGURES 157–171. Dorsal habitus photographs of Drassodella species in the salisburyi species group: 157, 158 female and male of D. amatola sp. nov.; 159, 160 female and male of D. aurostriata sp. nov.; 161 male of D. baviaans sp. nov.; 162 female of D. purcelli Tucker, 1923; 163, 164 female and male of D. quinquelabecula Tucker, 1923; 165, 166 female and male of D. salisburyi Hewitt, 1916; 167, 168 female and male of D. septemmaculata (Strand, 1909); 169 female of D. trilineata sp. nov.; 170, 171 female and male of D. vasivulva Tucker, 1923. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643776/files/figure.png" pageId="39" pageNumber="40">Fig. 158</figureCitation>
|
||
. Carapace black, with faint radiating mottling; distinct upsilon marking of white feathery setae present, comprising median line from PME, diverging after fovea. Clypeus height equal to 2.50 AME diameter; AME smaller than ALE; AME separated by distance equal to 1.33 their diameter, AME separated from ALE by distance equal to 0.33 AME diameter; PME smaller than PLE; PME separated by distance equal to 1.43 their diameter, PME separated from PLE by distance twice PME diameter; Chelicerae dark brown anteriorly, brown to red-brown posteriorly; promargin with three teeth, proximal tooth smallest, median tooth largest, median tooth closer to proximal tooth than distal tooth; retromargin with two teeth, equal in size. Endites dark brown, paler brown at maxillar hair tuft. Labium dark brown. Sternum dark brown. All legs slightly mottled, proximal segments dark brown, distal segments yellow-brown. Leg spination: femora: I do 1, II do 1 rl 1, III do 4, IV do 2; patellae: spineless; tibiae: I & II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2, IV pl 2 do 2 plv 2 rlv 2; metatarsi: I & II spineless, III pl 4 do 1 rl 4 plv 2 rlv 2 vt 2, IV pl 4 do 1 rl 4 plv 1 rlv 2 vt 2; palpal spination: femora do 3 rl 1; patellae rl 1; tibiae pl 1 rl 3; tarsi rl 2. Abdomen uniformly black; lateral sigilla above booklungs sub-oval; venter pale, with sparse thin hairs. Spinnerets brown. Palp dark brown; RTA long, largely obscured in ventral view by retrolateral heel of cymbium; in retrolateral view triangular, with broad base and rounded tip; cymbium broad, 1.55 times longer than wide, distal end broadly rounded; tegulum protruding prominently on prolateral side basally; embolus originating from distal prolateral mound, transverse and directed retrolaterally; median apophysis stout, hook-shaped (
|
||
<figureCitation id="A7DD2A389E4940095211CB1A63CC9F6E" box="[309,396,548,573]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="40" pageNumber="41">Figs 86</figureCitation>
|
||
,
|
||
<figureCitation id="A7DD2A389E49400952B3CB1A63BD9F6E" box="[407,509,548,573]" captionStart="FIGURES 172–175" captionStartId="40.[151,264,1395,1417]" captionTargetBox="[167,1425,622,1376]" captionTargetId="figure@40.[161,1426,611,1382]" captionTargetPageId="40" captionText="FIGURES 172–175. Genitalic morphology of Drassodella amatola sp. nov.: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm." httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="40" pageNumber="41">174, 175</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6B9966359E49400953B3CC4D667C98FB" httpUri="https://zenodo.org/record/2643780/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,264,1395,1417]" targetBox="[167,1425,622,1376]" targetPageId="40">
|
||
<paragraph id="3F5936BD9E49400953B3CC4D667C98FB" blockId="40.[151,1436,1395,1448]" pageId="40" pageNumber="41">
|
||
<emphasis id="0D92EAAF9E49400953B3CC4D633098DA" bold="true" box="[151,368,1395,1417]" pageId="40" pageNumber="41">FIGURES 172–175.</emphasis>
|
||
Genitalic morphology of
|
||
<taxonomicName id="F8E64D3E9E49400951A5CC4D611998DA" authority="Mbo & Haddad, 2019" authorityName="Mbo & Haddad" authorityYear="2019" box="[641,857,1395,1417]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="amatola" status="sp. nov.">
|
||
<emphasis id="0D92EAAF9E49400951A5CC4D611998DA" box="[641,857,1395,1417]" italics="true" pageId="40" pageNumber="41">Drassodella amatola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0D92EAAF9E4940095046CC4D61F598DA" bold="true" box="[866,949,1395,1417]" pageId="40" pageNumber="41">
|
||
<taxonomicNameLabel id="16A157D49E4940095046CC4D61F598DA" box="[866,949,1395,1417]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 172 female epigyne, ventral view; 173 same, dorsal view; 174 male palp, ventral view; 175 same, retrolateral view. Scale bars = 0.25mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="77FC65369E49400953E3CCC663609B96" pageId="40" pageNumber="41" type="materials_examined">
|
||
<paragraph id="3F5936BD9E49400953E3CCC6635E9B0A" blockId="40.[151,1436,1528,1769]" pageId="40" pageNumber="41">
|
||
<emphasis id="0D92EAAF9E49400953E3CCC663399B42" bold="true" box="[199,377,1528,1553]" pageId="40" pageNumber="41">Type material.</emphasis>
|
||
<materialsCitation id="8F8E3CE09E49400952A0CCC6635E9B0A" ID-GBIF-Occurrence="2236308101" collectingDate="2007-03-27" collectionCode="NCA" collectorName="C. Haddad" country="South Africa" county="Amatola Mountains" latitude="-32.60475" location="Afromontane forest" longLatPrecision="1" longitude="26.943" municipality="Hogsback" pageId="40" pageNumber="41" specimenCode="NCA 2007" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape" typeStatus="holotype">
|
||
<typeStatus id="E05D881F9E49400952A0CCC663AC9B42" box="[388,492,1528,1553]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="29E0FD349E49400952D3CCC7604E9B43" box="[503,526,1529,1552]" pageId="40" pageNumber="41" type="female">♀</specimenCount>
|
||
:
|
||
<emphasis id="0D92EAAF9E4940095104CCC661439B42" bold="true" box="[544,771,1528,1553]" pageId="40" pageNumber="41">
|
||
<collectingCountry id="47F1762D9E4940095104CCC660BE9B42" box="[544,766,1528,1553]" name="South Africa" pageId="40" pageNumber="41">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="0D92EAAF9E494009502BCCC7665C9B43" box="[783,1052,1529,1552]" italics="true" pageId="40" pageNumber="41">
|
||
<collectingRegion id="FD22F85F9E494009502BCCC7665C9B43" box="[783,1052,1529,1552]" country="South Africa" name="Eastern Cape" pageId="40" pageNumber="41">Eastern Cape Province</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty id="D6384E319E494009570BCCC667569B42" box="[1071,1302,1528,1553]" pageId="40" pageNumber="41">Amatola Mountains</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="DF3DACC79E4940095601CCC667D89B42" box="[1317,1432,1528,1553]" pageId="40" pageNumber="41">Hogsback</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="5AD2507A9E49400953B3CF2263629B66" box="[151,290,1564,1589]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-32.60475">32°36.285'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5AD2507A9E4940095216CF2263809B66" box="[306,448,1564,1589]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="26.943">26°56.580'E</geoCoordinate>
|
||
,
|
||
<date id="4B58107D9E49400952EBCF2360119B66" box="[463,593,1565,1589]" pageId="40" pageNumber="41" value="2007-03-27">
|
||
<collectingDate id="5B1CE9959E49400952EBCF2360119B66" box="[463,593,1565,1589]" pageId="40" pageNumber="41" value="2007-03-27">27.III.2007</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="9213536B9E49400951B1CF2361529B66" box="[661,786,1564,1589]" pageId="40" pageNumber="41">C. Haddad</collectorName>
|
||
(active search, knee down,
|
||
<location id="3A3960669E4940095746CF2267059B66" LSID="urn:lsid:plazi:treatment:B74F87AB9E47400953B3CAD766A99BBA:3A3960669E4940095746CF2267059B66" box="[1122,1349,1564,1589]" country="South Africa" county="Amatola Mountains" latitude="-32.60475" longLatPrecision="1" longitude="26.943" municipality="Hogsback" name="Afromontane forest" pageId="40" pageNumber="41" stateProvince="Eastern Cape">Afromontane forest</location>
|
||
) (
|
||
<specimenCode id="6F409EC69E494009567BCF2362929B0A" collectionCode="NCA" pageId="40" pageNumber="41">NCA 2007</specimenCode>
|
||
/1188).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="3F5936BD9E49400953E3CF5B63609B96" blockId="40.[151,1436,1528,1769]" pageId="40" pageNumber="41">
|
||
<materialsCitation id="8F8E3CE09E49400953E3CF5B63329BF2" ID-GBIF-Occurrence="2236308086" collectionCode="III" country="South Africa" latitude="-32.60475" location="Hogsback" longLatPrecision="1" longitude="26.943" municipality="Amatola Mountains" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
|
||
<typeStatus id="E05D881F9E49400953E3CF5B63799B2E" box="[199,313,1637,1661]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<emphasis id="0D92EAAF9E494009526ACF5A60789B2E" bold="true" box="[334,568,1636,1661]" pageId="40" pageNumber="41">
|
||
<collectingCountry id="47F1762D9E494009526ACF5A60739B2E" box="[334,563,1636,1661]" name="South Africa" pageId="40" pageNumber="41">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="0D92EAAF9E494009516ECF5861259B2F" box="[586,869,1637,1661]" italics="true" pageId="40" pageNumber="41">
|
||
<collectingRegion id="FD22F85F9E494009516ECF5861259B2F" box="[586,869,1637,1661]" country="South Africa" name="Eastern Cape" pageId="40" pageNumber="41">Eastern Cape Province</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality id="DF3DACC79E494009505BCF5A662D9B2E" box="[895,1133,1636,1661]" pageId="40" pageNumber="41">Amatola Mountains</collectingMunicipality>
|
||
,
|
||
<location id="3A3960669E49400957A7CF5A66B69B2E" LSID="urn:lsid:plazi:treatment:B74F87AB9E47400953B3CAD766A99BBA:3A3960669E49400957A7CF5A66B69B2E" box="[1155,1270,1636,1661]" country="South Africa" latitude="-32.60475" longLatPrecision="1" longitude="26.943" municipality="Amatola Mountains" name="Hogsback" pageId="40" pageNumber="41" stateProvince="Eastern Cape">Hogsback</location>
|
||
,
|
||
<geoCoordinate id="5AD2507A9E4940095628CF5A67D79B2E" box="[1292,1431,1636,1661]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-32.60475">32°36.285'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5AD2507A9E49400953B3CFB663649BF2" box="[151,292,1672,1697]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="26.943">26°56.580'E</geoCoordinate>
|
||
,
|
||
<date id="4B58107D9E494009520BCFB663329BF2" box="[303,370,1672,1697]" pageId="40" pageNumber="41" value="2007-03-25">
|
||
25.
|
||
<collectionCode id="59F7AE789E4940095276CFB663329BF2" box="[338,370,1672,1697]" pageId="40" pageNumber="41">III</collectionCode>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="8F8E3CE09E4940095255CFB667629BF2" ID-GBIF-Occurrence="2236308088" box="[369,1314,1672,1697]" collectionCode="NCA" collectorName="C. Haddad" country="South Africa" location="Afromontane forest" pageId="40" pageNumber="41" specimenCode="NCA 2007" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape Province" typeStatus="paratype">
|
||
.2007, leg.
|
||
<collectorName id="9213536B9E49400952C8CFB760249BF2" box="[492,612,1672,1697]" pageId="40" pageNumber="41">C. Haddad</collectorName>
|
||
(sifting leaf litter,
|
||
<location id="3A3960669E4940095012CFB666569BF2" LSID="urn:lsid:plazi:treatment:B74F87AB9E47400953B3CAD766A99BBA:3A3960669E4940095012CFB666569BF2" box="[822,1046,1672,1697]" country="South Africa" name="Afromontane forest" pageId="40" pageNumber="41" stateProvince="Eastern Cape Province">Afromontane forest</location>
|
||
),
|
||
<specimenCount id="29E0FD349E494009570DCFB7660C9BF2" box="[1065,1100,1673,1697]" pageId="40" pageNumber="41" type="female">1♀</specimenCount>
|
||
(
|
||
<specimenCode id="6F409EC69E494009577FCFB766999BF2" box="[1115,1241,1672,1697]" collectionCode="NCA" pageId="40" pageNumber="41">NCA 2007</specimenCode>
|
||
/1159)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="8F8E3CE09E494009560ACFB7635C9B96" ID-GBIF-Occurrence="2236308035" collectionCode="NCA" collectorName="C. Haddad" country="South Africa" isTailEnumeration="true" location="Afromontane forest" pageId="40" pageNumber="41" specimenCode="NCA 2007" specimenCount="1" specimenCount-male="1" stateProvince="Eastern Cape Province" typeStatus="paratype">
|
||
<specimenCount id="29E0FD349E494009560ACFB767119BF2" box="[1326,1361,1673,1697]" pageId="40" pageNumber="41" type="male">1♂</specimenCount>
|
||
(
|
||
<specimenCode id="6F409EC69E494009567BCFB762929B96" collectionCode="NCA" pageId="40" pageNumber="41">NCA 2007</specimenCode>
|
||
/3841)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="77FC65369E49400953E3CFEE66A99BBA" box="[199,1257,1744,1769]" pageId="40" pageNumber="41" type="distribution">
|
||
<paragraph id="3F5936BD9E49400953E3CFEE66A99BBA" blockId="40.[151,1436,1528,1769]" box="[199,1257,1744,1769]" pageId="40" pageNumber="41">
|
||
<emphasis id="0D92EAAF9E49400953E3CFEE63209BBA" bold="true" box="[199,352,1744,1769]" pageId="40" pageNumber="41">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="E05D881F9E4940095144CFEF60D09BBA" box="[608,656,1745,1769]" pageId="40" pageNumber="41">type</typeStatus>
|
||
locality in the
|
||
<collectingRegion id="FD22F85F9E494009501CCFEF618F9BBA" box="[824,975,1745,1769]" country="South Africa" name="Eastern Cape" pageId="40" pageNumber="41">Eastern Cape</collectingRegion>
|
||
,
|
||
<collectingCountry id="47F1762D9E49400950FFCFEE662A9BBA" box="[987,1130,1744,1769]" name="South Africa" pageId="40" pageNumber="41">South Africa</collectingCountry>
|
||
(
|
||
<figureCitation id="A7DD2A389E494009575ECFEE669C9BBA" box="[1146,1244,1744,1769]" captionStart="FIGURE 188" captionStartId="46.[151,250,1794,1816]" captionTargetBox="[161,1426,807,1772]" captionTargetId="figure@46.[161,1426,807,1772]" captionTargetPageId="46" captionText="FIGURE 188. Distribution of Drassodella amatola sp. nov. (circles), D. aurostriata sp. nov. (stars), D. baviaans sp. nov. (square), D. purcelli Tucker, 1923 (triangles) and D. quinquelabecula Tucker, 1923 (inverted triangles) in southern Africa." httpUri="https://zenodo.org/record/2643790/files/figure.png" pageId="40" pageNumber="41">Fig. 188</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |