190 lines
30 KiB
XML
190 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1181.108822" ID-Pensoft-Pub="1313-2970-1181-219" ID-Pensoft-UUID="6BA38450891853158C21659EC537DFD4" ID-ZooBank="1DF5630C74594525892B647A84C2098F" ModsDocID="1313-2970-1181-219" checkinTime="1696648901044" checkinUser="pensoft" docAuthor="Chu, Chang, Li, Shuqiang, Yao, Yanbin & Yao, Zhiyuan" docDate="2023" docId="1692FDEC1E8D592DA2D61A6828B54BD1" docLanguage="en" docName="ZooKeys 1181: 219-240" docOrigin="ZooKeys 1181" docPubDate="2023-10-06" docSource="http://dx.doi.org/10.3897/zookeys.1181.108822" docTitle="Koppe ninger Chu & Li 2023, sp. nov." docType="treatment" docUuid="3D1D9CAA-0F7B-4BEC-845E-BE9F5C67CB9D" docUuidSource="ZooBank" docVersion="1" id="6BA38450891853158C21659EC537DFD4" lastPageNumber="219" masterDocId="6BA38450891853158C21659EC537DFD4" masterDocTitle="One new genus and four new species of Liocranidae Simon, 1897 (Arachnida, Araneae) from China and Vietnam" masterLastPageNumber="240" masterPageNumber="219" pageNumber="219" updateTime="1696648901044" updateUser="pensoft">
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<mods:titleInfo id="2E23954D9D32FB1D047E003AFDD84CE9">
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<mods:title id="9B5878BA7BBB2C11475A61D3F5571EFC">One new genus and four new species of Liocranidae Simon, 1897 (Arachnida, Araneae) from China and Vietnam</mods:title>
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<mods:name id="7EC9DC73EE2E6DEC6DBD354D0B888E76" type="personal">
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<mods:roleTerm id="0CC72535FFDD61F014FA06B8E473B1CC">Author</mods:roleTerm>
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<mods:namePart id="1E6DE7F43930481B4CDE91BBA76D58F1">Chu, Chang</mods:namePart>
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<mods:nameIdentifier id="BEDD35B743C9B5E045CBE7D94F8E78C0" type="ORCID">https://orcid.org/0000-0003-3520-5463</mods:nameIdentifier>
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<mods:affiliation id="24125DEB735EEC6E246D603F08CAA6B5">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:roleTerm id="F9E9EFBFB7E8A172ADE6E7ADF93FF090">Author</mods:roleTerm>
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<mods:namePart id="C6B410EF497BD358F76F4FACD83D2784">Li, Shuqiang</mods:namePart>
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<mods:nameIdentifier id="9EB86B55DBE9F365CC15A5EA8917D062" type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
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<mods:affiliation id="1FDD8442BCA9A60D371B229EA00E25D0">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier id="03EDADF35D0519037238838BFE6F509C" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:name id="39C0856E91E2B068C6451F8128911E94" type="personal">
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<mods:roleTerm id="069FD89A5461FF73CC12531269024D55">Author</mods:roleTerm>
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<mods:namePart id="116B79176FBDA03B7CFD0C34C28338C1">Yao, Yanbin</mods:namePart>
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<mods:nameIdentifier id="0F40DC56073403833E6646519D453DBA" type="ORCID">https://orcid.org/0000-0002-2560-9299</mods:nameIdentifier>
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<mods:affiliation id="50CE9BB6BD9AE9B943F3966A105410AD">Jinshan College of Fujian Agriculture and Forestry University, Fuzhou 350002, Fujian, China</mods:affiliation>
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<mods:namePart id="5C0F753D7EEC0AA3F8716C9BF1522BEE">Yao, Zhiyuan</mods:namePart>
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<mods:nameIdentifier id="03E56924D2D4C5A168C85EF4C615EAA7" type="ORCID">https://orcid.org/0000-0002-1631-0949</mods:nameIdentifier>
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<mods:affiliation id="D8083A6AC717AEF23F729210207C0D93">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:nameIdentifier id="29CF4C4F7E24647897F565B34AFF326D" type="email">yaozy@synu.edu.cn</mods:nameIdentifier>
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<mods:title id="1E8E3DE2D9472E978256FECBF359563F">ZooKeys</mods:title>
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<mods:part id="8F1C56CFE38AB522E10F2CAA683DF293">
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<mods:date id="1B87F21A40FFEB318B00A76EF518870F">2023</mods:date>
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<mods:number id="6DD37F87BE34AE8B17DA92CF91CCB635">2023-10-06</mods:number>
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<mods:number id="B5C80878B1B9845022D0F21BC3E271D5">1181</mods:number>
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<mods:url id="41A80E8ACC3446E22878EB8E3C1BDCBA">http://dx.doi.org/10.3897/zookeys.1181.108822</mods:url>
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<mods:classification id="A4483B9DC51875164AADF059594BD31B">journal article</mods:classification>
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<mods:identifier id="3FD090FF95049166298420F44FD7F15C" type="DOI">http://dx.doi.org/10.3897/zookeys.1181.108822</mods:identifier>
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<treatment id="1692FDEC1E8D592DA2D61A6828B54BD1" LSID="urn:lsid:zoobank.org:act:3D1D9CAA-0F7B-4BEC-845E-BE9F5C67CB9D" httpUri="http://treatment.plazi.org/id/1692FDEC1E8D592DA2D61A6828B54BD1" lastPageNumber="219" pageId="0" pageNumber="219">
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<subSubSection id="452FF27F61A7275DE1B8E0393E1610A2" pageId="0" pageNumber="219" type="nomenclature">
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<paragraph id="B3D3AA49EB54868A35840DD9C867F2ED" pageId="0" pageNumber="219">
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<taxonomicName id="6E566676F9913C52D4BC5DC5B0F67CAE" LSID="https://zoobank.org/3D1D9CAA-0F7B-4BEC-845E-BE9F5C67CB9D" authority="Chu & Li" authorityName="Chu & Li" authorityYear="2023" class="Arachnida" family="Liocranidae" genus="Koppe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Koppe ninger" order="Araneae" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ninger" status="sp. nov.">Koppe ninger Chu & Li</taxonomicName>
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<taxonomicNameLabel id="24F373C9B837F6BADEB4510BC2376737" pageId="0" pageNumber="219">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="7DA97488C7D955DEF3872213EC5A3587" pageId="0" pageNumber="219" type="description">
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<paragraph id="79F6A71BBBCF35FA0E6D07D60199060D" pageId="0" pageNumber="219">
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<figureCitation id="A1D2D923473053C25BB8897D020A2814" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">Figs 1</figureCitation>
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<figureCitation id="3816E0FD107D5288862583C16868327E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Koppe ninger sp. nov., holotype female (A-C) habitus A dorsal view B lateral view C ventral view. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1181.108822.figure2" httpUri="https://binary.pensoft.net/fig/917721" pageId="0" pageNumber="219">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="37EA09919AE46DFA7B3FA867241070EF" pageId="0" pageNumber="219" type="type material">
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<paragraph id="77E8AA061924A21C94806288E775C33B" pageId="0" pageNumber="219">Type material.</paragraph>
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<paragraph id="B795E53B41781C3094394CFAC7E4CF36" pageId="0" pageNumber="219">
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<emphasis id="B3EB0B1061B01E7B01205C85FD83CD35" bold="true" italics="true" pageId="0" pageNumber="219">Holotype</emphasis>
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: 1♀ (IZCAS-Ar44617),
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<emphasis id="2FDE3DFEBDDE6B76581D1547E8F33B39" bold="true" pageId="0" pageNumber="219">China</emphasis>
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, Yunnan,
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<normalizedToken id="B1C36F636964C2C4078569186F8F3E70" originalValue="Pu’er">Pu'er</normalizedToken>
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,
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<normalizedToken id="A5DCF3DEDA03D43FB002E5587193B49B" originalValue="Ning’er">Ning'er</normalizedToken>
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County, Jinpaoshan Park,
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<geoCoordinate id="72872F9783C92A4A9C2CC3A4DB352347" degrees="23" direction="north" minutes="3.658" orientation="latitude" precision="1" value="23.060966">23°3.658′N</geoCoordinate>
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,
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<geoCoordinate id="6ACE93743A0919312EFE8A5FC6C64B1B" degrees="101" direction="east" minutes="3.466" orientation="longitude" precision="1" value="101.05777">101°3.466′E</geoCoordinate>
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, hand catch in leaf litter, 26 July 2022, F. Gao leg.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0593CE2B5B10170506FBFD869B83199" pageId="0" pageNumber="219" type="etymology">
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<paragraph id="9002243A27BAD03DF369E11176009EAA" pageId="0" pageNumber="219">Etymology.</paragraph>
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<paragraph id="9694B7DD0A595C316F548A240145F5D3" pageId="0" pageNumber="219">The specific name refers to the type locality and is a noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection id="56946456A808EEAC77D9ACF4118EF1EA" pageId="0" pageNumber="219" type="diagnosis">
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<paragraph id="D51F6BFE889FC2E69EE2E6AD54474EB2" pageId="0" pageNumber="219">Diagnosis.</paragraph>
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<paragraph id="EF0DF3F4A12F7BD12FB5A85534D82BD2" pageId="0" pageNumber="219">
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The new species resembles
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<taxonomicName id="586889E50BA4C856CC1B3E052CF877D6" lsidName="K. princeps" pageId="0" pageNumber="219" rank="species" species="princeps">
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<emphasis id="EE69484A91DA5E64C8B9F197A71E61D6" italics="true" pageId="0" pageNumber="219">K. princeps</emphasis>
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</taxonomicName>
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Deeleman-Reinhold, 2001 (cf. Figs
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<figureCitation id="0F3557A46C88D00434C018D92A862F35" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1</figureCitation>
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,
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<figureCitation id="5D4AE9DDD7C534FD38A50EB8EAE30959" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Koppe ninger sp. nov., holotype female (A-C) habitus A dorsal view B lateral view C ventral view. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1181.108822.figure2" httpUri="https://binary.pensoft.net/fig/917721" pageId="0" pageNumber="219">2</figureCitation>
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and
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<bibRefCitation id="BCBA3873A71B52E5063B4D9B0D5DD950" DOI="https://doi.org/10.1163/9789004475588" author="Deeleman-Reinhold, CL" journalOrPublisher="Kavkazskij Entomologiceskij Bjulleten [Caucasian Entomological Bulletin]" pageId="0" pageNumber="219" refId="B8" refString="Deeleman-Reinhold, CL, 2001. Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill Leiden, 591 pp. https://doi.org/10.1163/9789004475588" title="Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill Leiden, 591 pp." url="https://doi.org/10.1163/9789004475588" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
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: 283, figs 391-396) by the similar rectangular epigynal window (Fig.
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<figureCitation id="BECFD627EBA659D87EA26C024E2DE539" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
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), posteriorly located copulatory openings (Fig.
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<figureCitation id="636AD9775E6083EBD2FC3AF918E816B9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
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), thin copulatory ducts (Fig.
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<figureCitation id="2DBC3727798306D593A3702412A39BF7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
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) and globular secondary spermathecae (Fig.
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<figureCitation id="EFFDC9060A9FF0A449DB235D3A01E78C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
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). Females can be distinguished by the epigyne with a pair of long, oblique sclerotized area laterally (Fig.
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<figureCitation id="04A74FE479E21802DABC2475EEACF679" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
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; vs. epigyne with a pair of short, similar point-shaped sclerotized area laterally), by the intertubular ducts globular (Fig.
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<figureCitation id="1E36E4FC5F280FE1486566FB1228BEA0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
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; vs. intertubular ducts tubular), by the primary spermathecae elliptical, separated by about their diameter (Fig.
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<figureCitation id="5071DC3FA1DD7D8B449EE3241BF5CB5F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
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||
; vs. primary spermathecae kidney-shaped, separated by more than twice their diameter), by the secondary spermathecae separated by less than half of their diameter (Fig.
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<figureCitation id="78B60E494CE147626E9154246B5D593E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
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; vs. secondary spermathecae separated by more than their diameter), and by the fertilization ducts pointing antero-laterally (Fig.
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<figureCitation id="B4E0DDEA73CBD1E303372D9C598EB4D4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
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; vs. fertilization ducts pointing postero-laterally). This species also resembles
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<taxonomicName id="FBAACDEA1AF5F891A9FA1CB0E547265A" lsidName="K. fusca" pageId="0" pageNumber="219" rank="species" species="fusca">
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<emphasis id="2AA9128C035BE7AB3E94FAF18EB53C52" italics="true" pageId="0" pageNumber="219">K. fusca</emphasis>
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</taxonomicName>
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Sankaran, 2022 (cf. Figs
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<figureCitation id="53D01E224623C510AC0FB4131697B304" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1</figureCitation>
|
||
,
|
||
<figureCitation id="5F9742F11E3A9666677D3F32FB1438B2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Koppe ninger sp. nov., holotype female (A-C) habitus A dorsal view B lateral view C ventral view. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1181.108822.figure2" httpUri="https://binary.pensoft.net/fig/917721" pageId="0" pageNumber="219">2</figureCitation>
|
||
and
|
||
<bibRefCitation id="F1E7CFC753B36E999587D31E2EEACDAC" DOI="https://doi.org/10.11646/zootaxa.5104.3.7" author="Sankaran, PM" journalOrPublisher="Zootaxa" pageId="0" pageNumber="219" pagination="436 - 440" refId="B22" refString="Sankaran, PM, 2022. Remarks on the spider genus Koppe Deeleman-Reinhold, 2001 (Araneae: Liocranidae), including the first records from India. Zootaxa 5104 (3): 436 - 440, DOI: https://doi.org/10.11646/zootaxa.5104.3.7" title="Remarks on the spider genus Koppe Deeleman-Reinhold, 2001 (Araneae: Liocranidae), including the first records from India." url="https://doi.org/10.11646/zootaxa.5104.3.7" volume="5104" year="2022">Sankaran 2022</bibRefCitation>
|
||
: 438, figs 1, 2) by the similar rectangular epigynal window (Fig.
|
||
<figureCitation id="7A6B10936530B55E3424F8E17877A6A1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
|
||
), posteriorly located copulatory openings (Fig.
|
||
<figureCitation id="25DB4ECE3447DCE6657ED3DA69848F66" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
|
||
), thin copulatory ducts (Fig.
|
||
<figureCitation id="C9BFD0DCAB9C40E31DCBC8E6F02DD21C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
|
||
), globular secondary spermathecae (Fig.
|
||
<figureCitation id="FF496B4262CC5F3F35AE98C934CB35EF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
|
||
) and antero-laterally pointed fertilization ducts (Fig.
|
||
<figureCitation id="07E006B4009BC6FC4401464DDA777079" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
|
||
). Females can be distinguished by the epigyne with a pair of long, oblique sclerotized area laterally (Fig.
|
||
<figureCitation id="08412332F81A611977FADA1917F2E047" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
|
||
; absent), by the epigyne without distinct median flap (Fig.
|
||
<figureCitation id="215B6DAB61C1A6840AEFB73247120A23" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A</figureCitation>
|
||
; present), by the intertubular ducts globular (Fig.
|
||
<figureCitation id="F03AF2E11D98ED54C73FBC9F78EC1E0C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
|
||
; vs. intertubular ducts tubular, with anterior twist), by the primary spermathecae elliptical, separated by about their diameter (Fig.
|
||
<figureCitation id="08BE578F1FF436EC901DE1BD5572BC0B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
|
||
; vs. primary spermathecae kidney-shaped, separated by more than their diameter), and by the secondary spermathecae separated by less than half of their diameter (Fig.
|
||
<figureCitation id="10DFC2FEC675CFBACC45541B0D2A4C49" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1B</figureCitation>
|
||
; vs. secondary spermathecae connected to each other). Male unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3CBCD6915734A0AE13D75EFD6C3E8113" pageId="0" pageNumber="219" type="description">
|
||
<paragraph id="0DC821F7585849AB87DC8DDD2E1504B9" pageId="0" pageNumber="219">Description.</paragraph>
|
||
<paragraph id="61B70104427CFB78C8398C8B8912E86A" pageId="0" pageNumber="219">
|
||
<emphasis id="1018B5642CA08F03931240B78A552480" bold="true" pageId="0" pageNumber="219">Female</emphasis>
|
||
(holotype; Fig.
|
||
<figureCitation id="8FADDBAC74968702B6DA7A0D417F5DE0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Koppe ninger sp. nov., holotype female (A-C) habitus A dorsal view B lateral view C ventral view. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1181.108822.figure2" httpUri="https://binary.pensoft.net/fig/917721" pageId="0" pageNumber="219">2A-C</figureCitation>
|
||
). Total body length 5.35, carapace 2.22 long, 1.60 wide; opisthosoma 3.13 long, 2.11 wide. Eye sizes and interdistances: AME 0.09, ALE 0.08, PME 0.09, PLE 0.08; AME-AME 0.11, AME-ALE 0.10, PME-PME 0.18, PME-PLE 0.17, AME-PME 0.12, ALE-PLE 0.09. Carapace reddish-brown, smooth, with distinct radial grooves; fovea longitudinal, slit-like. Chelicerae reddish-brown, massive, with several setae on anterior surface, with three promarginal and seven retromarginal teeth. Endites and labium reddish-brown; endites narrower in middle, subapically with large, semicircular membranous area and dense scopula; labium longer than wide, with subbasal constriction and sparse scopula apically. Sternum reddish-brown, shield-shaped, with intercoxal sclerites between coxae; posterior margin extending between coxae IV; intercoxal sclerites distinctly enlarged, especially between coxae I and II, II and III. Legs yellowish-brown. Leg spination: femur I pl 1; tibiae I plv 9 rlv 8, II plv 7 rlv 6; metatarsi I plv 6 rlv 6, II plv 5 rlv 4. Palp and leg measurements: palp 2.97 (0.92, 0.43, 0.62, -, 1.00), I 8.76 (2.10, 0.69, 2.36, 2.23, 1.38), II 6.78 (1.84, 0.62, 1.61, 1.64, 1.07), III 5.89 (1.56, 0.60, 1.16, 1.59, 0.98), IV 8.29 (2.10, 0.70, 1.87, 2.37, 1.25). Leg formula: 1423. Dorsal opisthosoma brown with grey patterns, oval, with scutum covering half of dorsum surface. Lateral opisthosoma with pale stripes. Ventral opisthosoma yellowish with brown patterns posteriorly, epigastric scutum reddish-brown. Spinnerets yellowish.
|
||
</paragraph>
|
||
<paragraph id="5EF81079131A802B26985F3FECD72F57" pageId="0" pageNumber="219">
|
||
<emphasis id="3588B79F3F0313A5AE206F804E5B6927" bold="true" italics="true" pageId="0" pageNumber="219">Epigyne</emphasis>
|
||
(Fig.
|
||
<figureCitation id="B2D310C2CFD5EF9C6736E962BA448912" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Koppe ninger sp. nov., holotype female (A, B) A epigyne, ventral view B vulva, dorsal view. Abbreviations: CD = copulatory duct, CO = copulatory opening, EW = epigynal window, FD = fertilization duct, ITD = intertubular duct, SP I = spermatheca I, SP II = spermatheca II. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1181.108822.figure1" httpUri="https://binary.pensoft.net/fig/917720" pageId="0" pageNumber="219">1A, B</figureCitation>
|
||
). Epigynal field nearly fan-shaped, with a pair of long, oblique sclerotized area laterally; posterior part medially with weakly sclerotized epigynal window. Copulatory openings hidden under epigynal plate. Copulatory ducts thin. Intertubular ducts globular. Primary spermathecae elliptical, separated by about their diameter; secondary spermathecae globular, separated by less than half of their diameter. Fertilization ducts pointing antero-laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FA6B80F9492C37534DB67A92C826981D" pageId="0" pageNumber="219" type="distribution">
|
||
<paragraph id="08D739CECF9EEDFD55A627A70C7B8FBD" pageId="0" pageNumber="219">Distribution.</paragraph>
|
||
<paragraph id="B068B64D2A7CE495AB2E1087C811ED3B" pageId="0" pageNumber="219">
|
||
China (Yunnan, type locality; Fig.
|
||
<figureCitation id="441162F888585C66D197AC8908EFA4B0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. New distribution records of liocranid spiders from China and Vietnam 1 Koppe ninger sp. nov. 2 Sinocranum menghai sp. nov. 3 Xantharia baizilongi sp. nov. 4 X. cucphuong sp. nov." figureDoi="10.3897/zookeys.1181.108822.figure12" httpUri="https://binary.pensoft.net/fig/917731" pageId="0" pageNumber="219">12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |