One new genus and four new species of Liocranidae Simon, 1897 (Arachnida, Araneae) from China and Vietnam Author Chu, Chang https://orcid.org/0000-0003-3520-5463 College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China Author Li, Shuqiang https://orcid.org/0000-0002-3290-5416 Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China lisq@ioz.ac.cn Author Yao, Yanbin https://orcid.org/0000-0002-2560-9299 Jinshan College of Fujian Agriculture and Forestry University, Fuzhou 350002, Fujian, China Author Yao, Zhiyuan https://orcid.org/0000-0002-1631-0949 College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China yaozy@synu.edu.cn text ZooKeys 2023 2023-10-06 1181 219 240 http://dx.doi.org/10.3897/zookeys.1181.108822 journal article http://dx.doi.org/10.3897/zookeys.1181.108822 1313-2970-1181-219 1DF5630C74594525892B647A84C2098F 6BA38450891853158C21659EC537DFD4 Koppe ninger Chu & Li sp. nov. Figs 1 , 2 Type material. Holotype : 1♀ (IZCAS-Ar44617), China , Yunnan, Pu'er , Ning'er County, Jinpaoshan Park, 23°3.658′N , 101°3.466′E , hand catch in leaf litter, 26 July 2022, F. Gao leg. Etymology. The specific name refers to the type locality and is a noun in apposition. Diagnosis. The new species resembles K. princeps Deeleman-Reinhold, 2001 (cf. Figs 1 , 2 and Deeleman-Reinhold 2001 : 283, figs 391-396) by the similar rectangular epigynal window (Fig. 1A ), posteriorly located copulatory openings (Fig. 1A ), thin copulatory ducts (Fig. 1B ) and globular secondary spermathecae (Fig. 1B ). Females can be distinguished by the epigyne with a pair of long, oblique sclerotized area laterally (Fig. 1A ; vs. epigyne with a pair of short, similar point-shaped sclerotized area laterally), by the intertubular ducts globular (Fig. 1B ; vs. intertubular ducts tubular), by the primary spermathecae elliptical, separated by about their diameter (Fig. 1B ; vs. primary spermathecae kidney-shaped, separated by more than twice their diameter), by the secondary spermathecae separated by less than half of their diameter (Fig. 1B ; vs. secondary spermathecae separated by more than their diameter), and by the fertilization ducts pointing antero-laterally (Fig. 1B ; vs. fertilization ducts pointing postero-laterally). This species also resembles K. fusca Sankaran, 2022 (cf. Figs 1 , 2 and Sankaran 2022 : 438, figs 1, 2) by the similar rectangular epigynal window (Fig. 1A ), posteriorly located copulatory openings (Fig. 1A ), thin copulatory ducts (Fig. 1B ), globular secondary spermathecae (Fig. 1B ) and antero-laterally pointed fertilization ducts (Fig. 1B ). Females can be distinguished by the epigyne with a pair of long, oblique sclerotized area laterally (Fig. 1A ; absent), by the epigyne without distinct median flap (Fig. 1A ; present), by the intertubular ducts globular (Fig. 1B ; vs. intertubular ducts tubular, with anterior twist), by the primary spermathecae elliptical, separated by about their diameter (Fig. 1B ; vs. primary spermathecae kidney-shaped, separated by more than their diameter), and by the secondary spermathecae separated by less than half of their diameter (Fig. 1B ; vs. secondary spermathecae connected to each other). Male unknown. Description. Female (holotype; Fig. 2A-C ). Total body length 5.35, carapace 2.22 long, 1.60 wide; opisthosoma 3.13 long, 2.11 wide. Eye sizes and interdistances: AME 0.09, ALE 0.08, PME 0.09, PLE 0.08; AME-AME 0.11, AME-ALE 0.10, PME-PME 0.18, PME-PLE 0.17, AME-PME 0.12, ALE-PLE 0.09. Carapace reddish-brown, smooth, with distinct radial grooves; fovea longitudinal, slit-like. Chelicerae reddish-brown, massive, with several setae on anterior surface, with three promarginal and seven retromarginal teeth. Endites and labium reddish-brown; endites narrower in middle, subapically with large, semicircular membranous area and dense scopula; labium longer than wide, with subbasal constriction and sparse scopula apically. Sternum reddish-brown, shield-shaped, with intercoxal sclerites between coxae; posterior margin extending between coxae IV; intercoxal sclerites distinctly enlarged, especially between coxae I and II, II and III. Legs yellowish-brown. Leg spination: femur I pl 1; tibiae I plv 9 rlv 8, II plv 7 rlv 6; metatarsi I plv 6 rlv 6, II plv 5 rlv 4. Palp and leg measurements: palp 2.97 (0.92, 0.43, 0.62, -, 1.00), I 8.76 (2.10, 0.69, 2.36, 2.23, 1.38), II 6.78 (1.84, 0.62, 1.61, 1.64, 1.07), III 5.89 (1.56, 0.60, 1.16, 1.59, 0.98), IV 8.29 (2.10, 0.70, 1.87, 2.37, 1.25). Leg formula: 1423. Dorsal opisthosoma brown with grey patterns, oval, with scutum covering half of dorsum surface. Lateral opisthosoma with pale stripes. Ventral opisthosoma yellowish with brown patterns posteriorly, epigastric scutum reddish-brown. Spinnerets yellowish. Epigyne (Fig. 1A, B ). Epigynal field nearly fan-shaped, with a pair of long, oblique sclerotized area laterally; posterior part medially with weakly sclerotized epigynal window. Copulatory openings hidden under epigynal plate. Copulatory ducts thin. Intertubular ducts globular. Primary spermathecae elliptical, separated by about their diameter; secondary spermathecae globular, separated by less than half of their diameter. Fertilization ducts pointing antero-laterally. Distribution. China (Yunnan, type locality; Fig. 12 ).