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<document id="AC0490CEF3C640F2827E24F02E96496A" ID-CLB-Dataset="4791" ID-DOI="10.11646/zootaxa.3710.3.8" ID-GBIF-Dataset="05375634-2bf8-4058-b1c1-8c54f2dbb3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5978489" ID-ZooBank="43DBC4BA-684D-4196-BE71-1F4CFE0EF498" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629901958053" checkinUser="felipe" docAuthor="Frolov, Andrey V." docDate="2013" docId="49712475FFEBFF8752B3FA81FDA9F862" docLanguage="en" docName="zt03710p300.pdf" docOrigin="Zootaxa 3710 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orphnus planicollis Petrovitz 1971" docType="treatment" docVersion="8" lastPageNumber="297" masterDocId="B5485C0DFFEBFF875224FF9EFFEAFFE6" masterDocTitle="New and little known species of the genus Orphnus MacLeay (Coleoptera: Scarabaeidae: Orphninae) from the East African Rift" masterLastPageNumber="300" masterPageNumber="297" pageNumber="297" updateTime="1699144698176" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B30E126993FF07A3537BD95BFBEB918E">New and little known species of the genus Orphnus MacLeay (Coleoptera: Scarabaeidae: Orphninae) from the East African Rift</mods:title>
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<mods:namePart id="2CA9F94E21BCF1AD50687E4559854B4B">Frolov, Andrey V.</mods:namePart>
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<mods:date id="97C903B2CA3735E1ECA08B51AC97DF4F">2013</mods:date>
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<mods:number id="E22444B0AE0750E4ACA1B375E87B3D5F">3710</mods:number>
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<mods:identifier id="89C35ADA4E7DD50B39C3A6109B504963" type="DOI">10.11646/zootaxa.3710.3.8</mods:identifier>
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<treatment id="49712475FFEBFF8752B3FA81FDA9F862" ID-DOI="http://doi.org/10.5281/zenodo.6149848" ID-GBIF-Taxon="184961167" ID-Zenodo-Dep="6149848" LSID="urn:lsid:plazi:treatment:49712475FFEBFF8752B3FA81FDA9F862" httpUri="http://treatment.plazi.org/id/49712475FFEBFF8752B3FA81FDA9F862" lastPageNumber="297" pageId="0" pageNumber="297">
<subSubSection id="89C2C6E8FFEBFF8752B3FA81FDCBFAD1" box="[151,545,1311,1335]" pageId="0" pageNumber="297" type="nomenclature">
<paragraph id="C1679563FFEBFF8752B3FA81FDCBFAD1" blockId="0.[151,545,1311,1369]" box="[151,545,1311,1335]" pageId="0" pageNumber="297">
<heading id="9A2F220FFFEBFF8752B3FA81FDCBFAD1" bold="true" box="[151,545,1311,1335]" fontSize="10" level="2" pageId="0" pageNumber="297" reason="2">
<taxonomicName id="06D8EEE0FFEBFF8752B3FA81FDCBFAD1" authority="Petrovitz, 1971" authorityName="Petrovitz" authorityYear="1971" box="[151,545,1311,1335]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="species" species="planicollis">
<emphasis id="F3AC4971FFEBFF8752B3FA81FDCBFAD1" bold="true" box="[151,545,1311,1335]" pageId="0" pageNumber="297">
<emphasis id="F3AC4971FFEBFF8752B3FA81FE9BFAD1" bold="true" box="[151,369,1311,1335]" italics="true" pageId="0" pageNumber="297">Orphnus planicollis</emphasis>
<bibRefCitation id="A549E892FFEBFF87535CFABEFDCBFAD1" author="Petrovitz, R." box="[376,545,1312,1335]" pageId="0" pageNumber="297" pagination="1 - 46" refId="ref2000" refString="Petrovitz, R. (1971) Beitrag zur Kenntnis der Gattung Orphnus M'Leay (Orphninae, Scarabaeidae, Coleoptera). Revue de Zoologie et Botanique Africaine, 84, 1 - 46." type="journal article" year="1971">Petrovitz, 1971</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89C2C6E8FFEBFF8752B3FADCFE9BFABF" box="[151,369,1346,1369]" pageId="0" pageNumber="297" type="description">
<paragraph id="C1679563FFEBFF8752B3FADCFE9BFABF" blockId="0.[151,545,1311,1369]" box="[151,369,1346,1369]" pageId="0" pageNumber="297">
<figureCitation id="59E389E6FFEBFF8752B3FADCFF0AFABF" box="[151,224,1346,1369]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[174,1414,205,1635]" captionTargetId="figure-126@1.[164,1422,193,1652]" captionTargetPageId="1" captionText="FIGURES 1, 2. Orphnus, habitus of male. Fig. 1—O. planicollis. Fig. 2—O. demeyeri, holotype." figureDoi="http://doi.org/10.5281/zenodo.5978495" httpUri="https://zenodo.org/record/5978495/files/figure.png" pageId="0" pageNumber="297">Figs. 1</figureCitation>
,
<figureCitation id="59E389E6FFEBFF8752C9FADCFEA6FABF" box="[237,332,1346,1369]" captionStart="FIGURES 310" captionStartId="2.[151,264,1569,1592]" captionTargetBox="[151,1432,886,1541]" captionTargetId="figure-592@2.[151,1436,886,1548]" captionTargetPageId="2" captionText="FIGURES 310. Orphnus. Figs. 3, 4—base of pronotum; Figs. 58—internal sac sclerite (8—sclerite broken); Figs. 9, 10— aedeagus in lateral view and parameres in dorsal view. Figs. 3, 5, 6, 9—O. planicollis. Figs. 4, 7, 8, 10—O. demeyeri, holotype." figureDoi="http://doi.org/10.5281/zenodo.5978497" httpUri="https://zenodo.org/record/5978497/files/figure.png" pageId="0" pageNumber="297">3, 5, 6, 9</figureCitation>
,
<figureCitation id="59E389E6FFEBFF875373FADCFE9BFABF" box="[343,369,1346,1369]" captionStart="FIGURE 11" captionStartId="3.[151,249,1008,1030]" captionTargetBox="[181,1410,301,982]" captionTargetId="figure-232@3.[168,1419,298,983]" captionTargetPageId="3" captionText="FIGURE 11. Orphnus, map of known localities. •—O. planicollis; Δ—O. demeyeri." figureDoi="http://doi.org/10.5281/zenodo.5978501" httpUri="https://zenodo.org/record/5978501/files/figure.png" pageId="0" pageNumber="297">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="89C2C6E8FFEBFF8752B3FA16FEB5F9CF" pageId="0" pageNumber="297" type="materials_examined">
<paragraph id="C1679563FFEBFF8752B3FA16FC9FFA79" blockId="0.[151,1437,1416,1924]" box="[151,885,1416,1439]" pageId="0" pageNumber="297">
<emphasis id="F3AC4971FFEBFF8752B3FA16FED7FA79" bold="true" box="[151,317,1416,1439]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">Type material.</emphasis>
<materialsCitation id="71B09F3EFFEBFF875367FA16FC9FFA79" ID-GBIF-Occurrence="3338769302" box="[323,885,1416,1439]" collectionCode="MHNG" country="Malawi" location="Nyassaland" pageId="0" pageNumber="297" specimenCount="1" typeStatus="holotype">
<typeStatus id="1E632BC1FFEBFF875367FA16FE43FA79" box="[323,425,1416,1439]" pageId="0" pageNumber="297" type="holotype">Holotype</typeStatus>
,
<specimenCount id="D7DE5EEAFFEBFF875397FA16FE0DFA79" box="[435,487,1416,1439]" pageId="0" pageNumber="297" type="male">male</specimenCount>
with a label “
<collectingCountry id="B9CFD5F3FFEBFF87505FFA16FD10FA79" box="[635,762,1416,1439]" name="Malawi" pageId="0" pageNumber="297">Nyassaland</collectingCountry>
” (
<collectionCode id="A7C90DA6FFEBFF875135FA16FC83FA79" box="[785,873,1416,1439]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="297" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="C1679563FFEBFF8752E3FA34FEB5F9CF" blockId="0.[151,1437,1416,1924]" pageId="0" pageNumber="297">
<emphasis id="F3AC4971FFEBFF8752E3FA34FE44FA27" bold="true" box="[199,430,1450,1473]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">Additional material.</emphasis>
<materialsCitation id="71B09F3EFFEBFF875390FA34FC3BFA02" ID-GBIF-Occurrence="3338769307" collectingDate="1984-11-04" collectingDateMax="1984-11-09" collectingDateMin="1984-11-04" collectingMethod="pitfall traps" collectionCode="ZMUKK" collectorName="N. Scharff" country="Tanzania" elevation="1450" location="Chita village" municipality="Uzungwa Scarp Forest Reserve" pageId="0" pageNumber="297" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Iringa Region">
<collectingCountry id="B9CFD5F3FFEBFF875390FA34FDAAFA27" box="[436,576,1450,1473]" name="Tanzania" pageId="0" pageNumber="297">TANZANIA</collectingCountry>
:
<collectingRegion id="031C5B81FFEBFF87506EFA34FD0AFA27" box="[586,736,1450,1473]" country="Tanzania" name="Iringa" pageId="0" pageNumber="297">Iringa Region</collectingRegion>
,
<collectingMunicipality id="21030F19FFEBFF8750CFFA34FBD7FA27" box="[747,1085,1450,1473]" pageId="0" pageNumber="297">Uzungwa Scarp Forest Reserve</collectingMunicipality>
, above
<location id="C407C3B8FFEBFF8756A9FA34FAF2FA27" LSID="urn:lsid:plazi:treatment:49712475FFEBFF8752B3FA81FDA9F862:C407C3B8FFEBFF8756A9FA34FAF2FA27" box="[1165,1304,1450,1473]" country="Tanzania" municipality="Uzungwa Scarp Forest Reserve" name="Chita village" pageId="0" pageNumber="297" stateProvince="Iringa Region">Chita village</location>
,
<quantity id="06203886FFEBFF875707FA34FA9CFA27" box="[1315,1398,1450,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="0" pageNumber="297" unit="m" value="1450.0">
<elevation id="4AF57250FFEBFF875707FA34FA9CFA27" box="[1315,1398,1450,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="0" pageNumber="297" unit="m" value="1450.0">1450 m</elevation>
</quantity>
,
<date id="B566B3A3FFEBFF8757A5FA34FF14FA02" pageId="0" pageNumber="297" value="1984-11-04" valueMax="1984-11-09" valueMin="1984-11-04">
<collectingDate id="A5224A4BFFEBFF8757A5FA34FF14FA02" pageId="0" pageNumber="297" value="1984-11-04" valueMax="1984-11-09" valueMin="1984-11-04">4 9.xi.1984</collectingDate>
</date>
,
<collectingMethod id="1899ED74FFEBFF875329FA53FE64FA02" box="[269,398,1485,1508]" pageId="0" pageNumber="297">pitfall traps</collectingMethod>
,
<collectorName id="6C2DF0B5FFEBFF8753B8FA53FDE5FA02" box="[412,527,1485,1508]" pageId="0" pageNumber="297">N. Scharff</collectorName>
leg.,
<specimenCount id="D7DE5EEAFFEBFF875077FA53FD43FA02" box="[595,681,1485,1508]" pageId="0" pageNumber="297" type="male">7 males</specimenCount>
and
<specimenCount id="D7DE5EEAFFEBFF8750C1FA53FCBAFA02" box="[741,848,1485,1508]" pageId="0" pageNumber="297" type="female">5 females</specimenCount>
(
<collectionCode id="A7C90DA6FFEBFF875146FA53FC20FA02" box="[866,970,1485,1508]" pageId="0" pageNumber="297">ZMUKK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="71B09F3EFFEBFF8751C4FA53FD67F9E1" ID-GBIF-Occurrence="3338769304" collectingDate="1996-01-07" collectingDateMax="1996-01-19" collectingDateMin="1996-01-07" collectionCode="MCSNC" collectorName="G. Curletti" country="Tanzania" location="Mafinga" pageId="0" pageNumber="297" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Iringa Region">
<collectingRegion id="031C5B81FFEBFF8751C4FA53FB90FA02" box="[992,1146,1485,1508]" country="Tanzania" name="Iringa" pageId="0" pageNumber="297">Iringa Region</collectingRegion>
,
<location id="C407C3B8FFEBFF8756ADFA53FB0CFA02" LSID="urn:lsid:plazi:treatment:49712475FFEBFF8752B3FA81FDA9F862:C407C3B8FFEBFF8756ADFA53FB0CFA02" box="[1161,1254,1485,1508]" country="Tanzania" name="Mafinga" pageId="0" pageNumber="297" stateProvince="Iringa Region">Mafinga</location>
,
<date id="B566B3A3FFEBFF8756D1FA53FA9CFA02" box="[1269,1398,1485,1508]" pageId="0" pageNumber="297" value="1996-01-07" valueMax="1996-01-19" valueMin="1996-01-07">
<collectingDate id="A5224A4BFFEBFF8756D1FA53FA9CFA02" box="[1269,1398,1485,1508]" pageId="0" pageNumber="297" value="1996-01-07" valueMax="1996-01-19" valueMin="1996-01-07">719.i.1996</collectingDate>
</date>
,
<collectorName id="6C2DF0B5FFEBFF8757A1FA53FF03F9E1" pageId="0" pageNumber="297">G. Curletti</collectorName>
leg.,
<specimenCount id="D7DE5EEAFFEBFF875301FA6EFE9DF9E1" box="[293,375,1520,1543]" pageId="0" pageNumber="297" type="male">6 males</specimenCount>
and
<specimenCount id="D7DE5EEAFFEBFF875389FA6EFDFEF9E1" box="[429,532,1520,1543]" pageId="0" pageNumber="297" type="female">3 females</specimenCount>
(
<collectionCode id="A7C90DA6FFEBFF875007FA6EFD6CF9E1" box="[547,646,1520,1543]" country="Italy" httpUri="http://grbio.org/cool/wcy4-cagh" name="Museo Civico di Storia Naturale, Carmagnola" pageId="0" pageNumber="297">MCSNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="71B09F3EFFEBFF8750BDFA6EFB77F9E1" ID-GBIF-Occurrence="3338769301" box="[665,1181,1520,1543]" collectingDate="1996-01-07" collectingDateMax="1996-01-19" collectingDateMin="1996-01-07" collectionCode="ZMHUB" collectorName="G. Curletti" country="Tanzania" isTailEnumeration="true" location="Upangwa Dtsch." pageId="0" pageNumber="297" specimenCount="1" specimenCount-female="1" stateProvince="Iringa">
<location id="C407C3B8FFEBFF875087FA6EFCB2F9E1" LSID="urn:lsid:plazi:treatment:49712475FFEBFF8752B3FA81FDA9F862:C407C3B8FFEBFF875087FA6EFCB2F9E1" box="[675,856,1520,1543]" country="Tanzania" name="Upangwa Dtsch." pageId="0" pageNumber="297" stateProvince="Iringa">Upangwa Dtsch.</location>
O. Afr.”,
<specimenCount id="D7DE5EEAFFEBFF8751E0FA6EFBC8F9E1" box="[964,1058,1520,1543]" pageId="0" pageNumber="297" type="female">1 female</specimenCount>
(
<collectionCode id="A7C90DA6FFEBFF875614FA6EFB7CF9E1" box="[1072,1174,1520,1543]" pageId="0" pageNumber="297">ZMHUB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="71B09F3EFFEBFF87568DFA6EFEB1F9CF" ID-GBIF-Occurrence="3338769306" collectionCode="ZMHUB" country="Tanzania" location="Afrika" pageId="0" pageNumber="297" specimenCount="1" specimenCount-female="1">
“Deutsch-O.
<location id="C407C3B8FFEBFF87570BFA6EFA90F9E1" LSID="urn:lsid:plazi:treatment:49712475FFEBFF8752B3FA81FDA9F862:C407C3B8FFEBFF87570BFA6EFA90F9E1" box="[1327,1402,1520,1543]" country="Tanzania" name="Afrika" pageId="0" pageNumber="297">Afrika</location>
”,
<specimenCount id="D7DE5EEAFFEBFF8757AAFA6EFF35F9CF" pageId="0" pageNumber="297" type="female">1 female</specimenCount>
(
<collectionCode id="A7C90DA6FFEBFF8752CAF98CFEBEF9CF" box="[238,340,1554,1577]" pageId="0" pageNumber="297">ZMHUB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89C2C6E8FFEBFF8752E3F9ABFE36F931" pageId="0" pageNumber="297" type="description">
<paragraph id="C1679563FFEBFF8752E3F9ABFCCAF989" blockId="0.[151,1437,1416,1924]" pageId="0" pageNumber="297">
<emphasis id="F3AC4971FFEBFF8752E3F9ABFECBF9AA" bold="true" box="[199,289,1589,1612]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">Female.</emphasis>
Differs from male in having protibiae with elongate spur; clypeus without horn; pronotum narrower, feebly depressed on disc and without umbones and medial tubercle.
</paragraph>
<paragraph id="C1679563FFEBFF8752E3F9E4FE36F931" blockId="0.[151,1437,1416,1924]" pageId="0" pageNumber="297">
<emphasis id="F3AC4971FFEBFF8752E3F9E4FEACF977" bold="true" box="[199,326,1658,1681]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">Variability.</emphasis>
Body length of examined specimens varies from 6.59.0 mm. Clypeal horn of males varies from very short (wider than long) to 3 times longer than wide. Medial tubercle on pronotum and especially umbones are less developed in some specimens.
</paragraph>
</subSubSection>
<subSubSection id="89C2C6E8FFEBFF8752E3F97CFDA9F862" pageId="0" pageNumber="297" type="distribution">
<paragraph id="C1679563FFEBFF8752E3F97CFDA9F862" blockId="0.[151,1437,1416,1924]" pageId="0" pageNumber="297">
<emphasis id="F3AC4971FFEBFF8752E3F97CFEB3F91F" box="[199,345,1762,1785]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">
<emphasis id="F3AC4971FFEBFF8752E3F97CFEB8F91F" bold="true" box="[199,338,1762,1785]" pageId="0" pageNumber="297">Distribution</emphasis>
.
</emphasis>
The species was described from “Nyassa-Land” without more precise locality. The two exact localities known (Uzungwa Scarp Forest Reserve near Chita and Mafinga) are situated in Uzungwa—the southernmost and largest bloc of Eastern Arc Mountains (
<bibRefCitation id="A549E892FFEBFF8750A6F8B6FC8AF8D9" author="Burgess, N. D. &amp; Butynski, T. M. &amp; Cordeiro, N. J. &amp; Doggart, N. H. &amp; Fjeldsa, J. &amp; Howell, K. M. &amp; Kilahamaa, F. B. &amp; Loaderk, S. P. &amp; Lovettl, J. C. &amp; Mbilinyia, B. &amp; Menegonm, M. &amp; Moyern, D. C. &amp; Nashandaj, E. &amp; Perking, A. &amp; Roverom, F. &amp; Stanleyo, W. T. &amp; Stuartp, S. N." box="[642,864,1832,1855]" pageId="0" pageNumber="297" pagination="209 - 231" refId="ref1753" refString="Burgess, N. D., Butynski, T. M., Cordeiro, N. J., Doggart, N. H., Fjeldsa, J., Howell, K. M., Kilahamaa, F. B., Loaderk, S. P., Lovettl, J. C., Mbilinyia, B., Menegonm, M., Moyern, D. C., Nashandaj, E., Perking, A., Roverom, F., Stanleyo, W. T. &amp; Stuartp, S. N. (2007) The biological importance of the Eastern Arc Mountains of Tanzania and Kenya. Biological Conservation, 134, 209 - 231. http: // dx. doi. org / 10.1016 / j. biocon. 2006.08.015" type="journal article" year="2007">
Burgess
<emphasis id="F3AC4971FFEBFF8750C7F8B6FCFFF8D9" box="[739,789,1832,1855]" italics="true" pageId="0" pageNumber="297">et al</emphasis>
. 2007
</bibRefCitation>
). The other locality, Upangwa, refers to the part of Livingstone Mountains south of Uzungwa. Available data suggest that
<taxonomicName id="06D8EEE0FFEBFF8751B1F8D4FBCDF887" authorityName="Petrovitz" authorityYear="1971" box="[917,1063,1866,1889]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="species" species="planicollis">
<emphasis id="F3AC4971FFEBFF8751B1F8D4FBCDF887" box="[917,1063,1866,1889]" italics="true" pageId="0" pageNumber="297">O. planicollis</emphasis>
</taxonomicName>
inhabits mid-altitude afromontane forests and is apparently a litter dweller.
</paragraph>
</subSubSection>
</treatment>
</document>