treatments-xml/data/19/6C/87/196C87F3FF8DFFD27B8AACF9FC04B231.xml
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<document id="FC34C7058274A6514DAD91053A8C0ED3" ID-DOI="10.1649/828.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="10105234" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173236196" checkinUser="tatiana" docAuthor="Morón, Miguel-Angel &amp; Riley, Edward G." docDate="2005" docId="196C87F3FF8DFFD27B8AACF9FC04B231" docLanguage="en" docName="828.1.pdf.imf" docOrigin="The Coleopterists Bulletin 59 (3)" docSource="http://dx.doi.org/10.1649/828.1" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Phyllophaga (Phyllophaga) nandalumia Moron and Riley, new species" docType="treatment" docVersion="3" lastPageNumber="397" masterDocId="E555FF8BFF89FFD47B6FA94AFFD0B057" masterDocTitle="Four New Species of Phyllophaga from Mexico and Central America (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="399" masterPageNumber="391" pageNumber="395" updateTime="1699632931453" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="70455DB739D25F75DEDEF370B6F887A2">Four New Species of Phyllophaga from Mexico and Central America (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:namePart id="57E6FEE3D205ADA39F13320E68E0B266">Morón, Miguel-Angel</mods:namePart>
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<mods:namePart id="34CCF8AD6F99226E9E81166A2B0505B6">Riley, Edward G.</mods:namePart>
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<mods:title id="4812A80AE8387669F1B9712553FBDF02">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="90E0724EF5A9B859100AE95EAC4F5857">
<mods:date id="33F9769BE6BB84B61A673DEF971B7ADB">2005</mods:date>
<mods:detail id="2DB34A6C4DFAC13F300474B6227D8FBF" type="pubDate">
<mods:number id="4164C0E39E4A29DB58999AA3F5FA6C22">2005-09-30</mods:number>
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<mods:classification id="92F3F4A84EC8A83C8E6A7E9C544AD406">journal article</mods:classification>
<mods:identifier id="FB97EEF80501EFF55AC026E004940FE3" type="DOI">10.1649/828.1</mods:identifier>
<mods:identifier id="21E31B81C46F9CE328BE67C3686C5552" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="32B7534581CFFE2641C8E18CB27241AE" type="Zenodo-Dep">10105234</mods:identifier>
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<treatment id="196C87F3FF8DFFD27B8AACF9FC04B231" LSID="urn:lsid:plazi:treatment:196C87F3FF8DFFD27B8AACF9FC04B231" httpUri="http://treatment.plazi.org/id/196C87F3FF8DFFD27B8AACF9FC04B231" lastPageId="6" lastPageNumber="397" pageId="4" pageNumber="395">
<subSubSection id="D9DF656EFF8DFFD07B8AACF9FC66B59F" box="[229,950,1459,1480]" pageId="4" pageNumber="395" type="nomenclature">
<paragraph id="917A36E5FF8DFFD07B8AACF9FC66B59F" blockId="4.[229,950,1459,1507]" box="[229,950,1459,1480]" pageId="4" pageNumber="395">
<heading id="CA328189FF8DFFD07B8AACF9FC66B59F" box="[229,950,1459,1480]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="395" reason="2">
<emphasis id="A3B1EAF7FF8DFFD07B8AACF9FC66B59F" bold="true" box="[229,950,1459,1480]" pageId="4" pageNumber="395">
<taxonomicName id="56C54D66FF8DFFD07B8AACF9FCE2B59F" authority="Moron and Riley" authorityName="Moron and Riley" box="[229,818,1459,1480]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="nandalumia" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="A3B1EAF7FF8DFFD07B8AACF9FEB2B59F" bold="true" box="[229,354,1459,1480]" italics="true" pageId="4" pageNumber="395">Phyllophaga</emphasis>
(
<emphasis id="A3B1EAF7FF8DFFD07A1DACF9FE3FB59F" bold="true" box="[370,495,1459,1480]" italics="true" pageId="4" pageNumber="395">Phyllophaga</emphasis>
)
<emphasis id="A3B1EAF7FF8DFFD07A90ACF9FDA7B59F" bold="true" box="[511,631,1459,1480]" italics="true" pageId="4" pageNumber="395">nandalumia</emphasis>
Morón and Riley
</taxonomicName>
,
<taxonomicNameLabel id="B882578CFF8DFFD07850ACF9FC66B59F" box="[831,950,1459,1480]" pageId="4" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9DF656EFF8DFFD27962AC84FE59B080" lastPageId="6" lastPageNumber="397" pageId="4" pageNumber="395" type="description">
<paragraph id="917A36E5FF8DFFD07962AC84FD5FB5B4" blockId="4.[229,950,1459,1507]" box="[525,655,1486,1507]" pageId="4" pageNumber="395">
(
<figureCitation id="09FE2A60FF8DFFD0797BAC84FD58B5B4" box="[532,648,1486,1507]" captionStart="Figs" captionStartId="5.[169,208,1179,1198]" captionTargetBox="[149,977,198,1157]" captionTargetId="figure-148@5.[148,978,194,1158]" captionTargetPageId="5" captionText="Figs. 1418. Phyllophaga nandalumia. 14) Genital capsule, lateral view; 15) parameres and aedeagus, lateral view; 16) phallobase, parameres and aedeagus, dorsal view; 17) parameres and aedeagus, ventral view; 18) parameres, distal view. Scale line ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105240" httpUri="https://zenodo.org/record/10105240/files/figure.png" pageId="4" pageNumber="395">Figs. 1418</figureCitation>
)
</paragraph>
<paragraph id="917A36E5FF8DFFD07BAAACBDFC3DB675" blockId="4.[175,1005,1527,1570]" pageId="4" pageNumber="395">
<emphasis id="A3B1EAF7FF8DFFD07BAAACBDFE46B65D" bold="true" box="[197,406,1527,1546]" pageId="4" pageNumber="395">
Description.
<typeStatus id="4E7E8847FF8DFFD07A52ACBDFE41B65D" box="[317,401,1527,1546]" pageId="4" pageNumber="395" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Body shiny brownish black. Antennal club as long as the preceding five segments. Anterior half of lateral border of pronotum crenulate. Elytra densely and
</paragraph>
<caption id="C5BA666DFF8CFFD17BC6ADD1FD2AB489" ID-DOI="http://doi.org/10.5281/zenodo.10105240" ID-Zenodo-Dep="10105240" httpUri="https://zenodo.org/record/10105240/files/figure.png" pageId="5" pageNumber="396" startId="5.[169,208,1179,1198]" targetBox="[149,977,198,1157]" targetPageId="5">
<paragraph id="917A36E5FF8CFFD17BC6ADD1FD2AB489" blockId="5.[148,977,1179,1247]" pageId="5" pageNumber="396">
<emphasis id="A3B1EAF7FF8CFFD17BC6ADD1FEC6B4F9" bold="true" box="[169,278,1179,1198]" pageId="5" pageNumber="396">Figs. 1418.</emphasis>
<taxonomicName id="56C54D66FF8CFFD17A44ADD1FDD2B4F9" box="[299,514,1179,1198]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="396" phylum="Arthropoda" rank="species" species="nandalumia">
<emphasis id="A3B1EAF7FF8CFFD17A44ADD1FDD2B4F9" box="[299,514,1179,1198]" italics="true" pageId="5" pageNumber="396">Phyllophaga nandalumia</emphasis>
</taxonomicName>
.
<emphasis id="A3B1EAF7FF8CFFD17961ADD1FDFAB4F9" bold="true" box="[526,554,1179,1198]" pageId="5" pageNumber="396">14)</emphasis>
Genital capsule, lateral view;
<emphasis id="A3B1EAF7FF8CFFD1785DADD1FC9EB4F9" bold="true" box="[818,846,1179,1198]" pageId="5" pageNumber="396">15)</emphasis>
parameres and aedeagus, lateral view;
<emphasis id="A3B1EAF7FF8CFFD17A32ADF9FEA9B491" bold="true" box="[349,377,1203,1222]" pageId="5" pageNumber="396">16)</emphasis>
phallobase, parameres and aedeagus, dorsal view;
<emphasis id="A3B1EAF7FF8CFFD1785DADF9FC9EB491" bold="true" box="[818,846,1203,1222]" pageId="5" pageNumber="396">17)</emphasis>
parameres and aedeagus, ventral view;
<emphasis id="A3B1EAF7FF8CFFD17A0BAD81FE50B489" bold="true" box="[356,384,1227,1246]" pageId="5" pageNumber="396">18)</emphasis>
parameres, distal view. Scale line
<emphasis id="A3B1EAF7FF8CFFD179C3AD80FD6CB488" bold="true" box="[684,700,1226,1247]" italics="true" pageId="5" pageNumber="396">¼</emphasis>
1 mm.
</paragraph>
</caption>
<paragraph id="917A36E5FF8CFFD17BFBAC42FEDCB52C" blockId="5.[148,978,1288,1570]" pageId="5" pageNumber="396">
deeply punctate, without setae. Abdominal sternite with erect setae at sides. Sternites 3 to 5 convex, smooth, near midline, with scattered short setae at sides. Pygidium moderately convex, densely punctate, without setae. Tarsal claws with short, acute ventral tooth located toward basal dilatation. Other external characters as in the species group
<emphasis id="A3B1EAF7FF8CFFD179DBAC1AFCC6B534" box="[692,790,1360,1379]" italics="true" pageId="5" pageNumber="396">schizorhina</emphasis>
, subgroup 7 (
<emphasis id="A3B1EAF7FF8CFFD178CDAC1AFC01B534" box="[930,977,1360,1379]" italics="true" pageId="5" pageNumber="396">sensu</emphasis>
Morón 2003).
</paragraph>
<paragraph id="917A36E5FF8CFFD17BC3ACCAFE73B675" blockId="5.[148,978,1288,1570]" pageId="5" pageNumber="396">
Phallobase enlarged, distal border slightly asymmetrical, deeply notched; basal border rounded (
<figureCitation id="09FE2A60FF8CFFD17BF5ACD2FED8B5FC" box="[154,264,1432,1451]" captionStart="Figs" captionStartId="5.[169,208,1179,1198]" captionTargetBox="[149,977,198,1157]" captionTargetId="figure-148@5.[148,978,194,1158]" captionTargetPageId="5" captionText="Figs. 1418. Phyllophaga nandalumia. 14) Genital capsule, lateral view; 15) parameres and aedeagus, lateral view; 16) phallobase, parameres and aedeagus, dorsal view; 17) parameres and aedeagus, ventral view; 18) parameres, distal view. Scale line ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105240" httpUri="https://zenodo.org/record/10105240/files/figure.png" pageId="5" pageNumber="396">Figs. 14, 16</figureCitation>
). Parameres nearly as long as width of phallobase, with external sides widened, blade-like, prominent, distal half briefly curved toward mid-line and apex rounded (
<figureCitation id="09FE2A60FF8CFFD17830ACE5FC17B595" box="[863,967,1455,1474]" captionStart="Figs" captionStartId="5.[169,208,1179,1198]" captionTargetBox="[149,977,198,1157]" captionTargetId="figure-148@5.[148,978,194,1158]" captionTargetPageId="5" captionText="Figs. 1418. Phyllophaga nandalumia. 14) Genital capsule, lateral view; 15) parameres and aedeagus, lateral view; 16) phallobase, parameres and aedeagus, dorsal view; 17) parameres and aedeagus, ventral view; 18) parameres, distal view. Scale line ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105240" httpUri="https://zenodo.org/record/10105240/files/figure.png" pageId="5" pageNumber="396">Figs. 1418</figureCitation>
). Aedeagus mainly symmetrical, with wide tube-like, strongly sclerotized support, compressed laterally and strongly expanded toward apex, with one subapical spur (
<figureCitation id="09FE2A60FF8CFFD1786FAC95FCBAB5A5" box="[768,874,1503,1522]" captionStart="Figs" captionStartId="5.[169,208,1179,1198]" captionTargetBox="[149,977,198,1157]" captionTargetId="figure-148@5.[148,978,194,1158]" captionTargetPageId="5" captionText="Figs. 1418. Phyllophaga nandalumia. 14) Genital capsule, lateral view; 15) parameres and aedeagus, lateral view; 16) phallobase, parameres and aedeagus, dorsal view; 17) parameres and aedeagus, ventral view; 18) parameres, distal view. Scale line ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105240" httpUri="https://zenodo.org/record/10105240/files/figure.png" pageId="5" pageNumber="396">Figs. 1417</figureCitation>
). Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="563D9B00FF8CFFD179B4ACBDFCF2B65D" box="[731,802,1527,1546]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="396" unit="mm" value="5.3">5.3 mm</quantity>
. Total body length:
<quantity id="563D9B00FF8CFFD17BFBAF45FF03B675" box="[148,211,1551,1570]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="396" unit="mm" value="20.0">20 mm</quantity>
. Humeral width:
<quantity id="563D9B00FF8CFFD17A05AF45FE4EB675" box="[362,414,1551,1570]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="396" unit="mm" value="8.0">8 mm</quantity>
.
</paragraph>
<paragraph id="917A36E5FF8FFFD27BA8A988FE59B080" blockId="6.[175,1005,194,614]" box="[199,393,194,215]" pageId="6" pageNumber="397">
<emphasis id="A3B1EAF7FF8FFFD27BA8A988FEC8B080" bold="true" box="[199,280,194,215]" pageId="6" pageNumber="397">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D9DF656EFF8FFFD27BA8A997FECCB1C6" pageId="6" pageNumber="397" type="materials_examined">
<paragraph id="917A36E5FF8FFFD27BA8A997FC71B15A" blockId="6.[175,1005,194,614]" pageId="6" pageNumber="397">
<emphasis id="A3B1EAF7FF8FFFD27BA8A997FE97B0A5" bold="true" box="[199,327,221,242]" pageId="6" pageNumber="397">Type Series.</emphasis>
(
<materialsCitation id="21AD3CB8FF8FFFD27A36A997FC71B15A" collectingDate="1990-09-12" collectionCode="IEXA" collectorName="R. W. Jones" country="Mexico" elevation="1000" location="Berriozabal" pageId="6" pageNumber="397" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="holotype">
Described from
<specimenCount id="87C3FD6CFF8FFFD27A95A997FD86B0A5" box="[506,598,221,242]" pageId="6" pageNumber="397" type="male">one male</specimenCount>
).
<typeStatus id="4E7E8847FF8FFFD27906A997FD11B0A5" box="[617,705,221,242]" pageId="6" pageNumber="397" type="holotype">Holotype</typeStatus>
male;
<collectingCountry id="E9D27675FF8FFFD27868A997FCB8B0A5" box="[775,872,221,242]" name="Mexico" pageId="6" pageNumber="397">MEXICO</collectingCountry>
:
<collectingRegion id="5301F807FF8FFFD2781CA997FC12B0A5" box="[883,962,221,242]" country="Mexico" name="Chiapas" pageId="6" pageNumber="397">Chiapas</collectingRegion>
,
<quantity id="563D9B00FF8FFFD278A1A997FF1DB15A" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="397" unit="km" value="7.5">7.5 km</quantity>
N
<location id="941A603EFF8FFFD27B80A9B2FE8FB15A" LSID="urn:lsid:plazi:treatment:196C87F3FF8DFFD27B8AACF9FC04B231:941A603EFF8FFFD27B80A9B2FE8FB15A" box="[239,351,248,269]" country="Mexico" name="Berriozabal" pageId="6" pageNumber="397" stateProvince="Chiapas">Berriozabal</location>
,
<date id="E57B1025FF8FFFD27A04A9B2FE0DB15A" box="[363,477,248,269]" pageId="6" pageNumber="397" value="1990-09-12">
<collectingDate id="F53FE9CDFF8FFFD27A04A9B2FE0DB15A" box="[363,477,248,269]" pageId="6" pageNumber="397" value="1990-09-12">12-IX-1990</collectingDate>
</date>
, taken at light,
<quantity id="563D9B00FF8FFFD27919A9B2FD18B15A" box="[630,712,248,269]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="397" unit="m" value="1000.0">
<elevation id="1AE8D1D6FF8FFFD27919A9B2FD18B15A" box="[630,712,248,269]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="397" unit="m" value="1000.0">1,000 m</elevation>
</quantity>
,
<collectorName id="3C305333FF8FFFD279BBA9B2FC9BB15A" box="[724,843,248,269]" pageId="6" pageNumber="397">R. W. Jones</collectorName>
(
<collectionCode id="F7D4AE20FF8FFFD27836A9B2FC47B15A" box="[857,919,248,269]" pageId="6" pageNumber="397">IEXA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="917A36E5FF8FFFD27BA8A858FDA6B115" blockId="6.[175,1005,194,614]" pageId="6" pageNumber="397">
<materialsCitation id="21AD3CB8FF8FFFD27BA8A858FDA1B115" country="Mexico" latitude="8.869166" location="Mesa de Ocozocuautla" longLatPrecision="21" longitude="-8.469166" municipality="Meseta Central" pageId="6" pageNumber="397" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="A3B1EAF7FF8FFFD27BA8A858FEB4B170" bold="true" box="[199,356,274,295]" pageId="6" pageNumber="397">
<typeStatus id="4E7E8847FF8FFFD27BA8A858FF2BB170" box="[199,251,274,295]" pageId="6" pageNumber="397">Type</typeStatus>
Locality.
</emphasis>
<location id="941A603EFF8FFFD27A1FA858FD8DB170" LSID="urn:lsid:plazi:treatment:196C87F3FF8DFFD27B8AACF9FC04B231:941A603EFF8FFFD27A1FA858FD8DB170" box="[368,605,274,295]" country="Mexico" latitude="8.869166" longLatPrecision="21" longitude="-8.469166" municipality="Meseta Central" name="Mesa de Ocozocuautla" pageId="6" pageNumber="397" stateProvince="Chiapas">Mesa de Ocozocuautla</location>
,
<collectingMunicipality id="711EAC9FFF8FFFD27900A858FCD8B170" box="[623,776,274,295]" pageId="6" pageNumber="397">Meseta Central</collectingMunicipality>
de
<collectingRegion id="5301F807FF8FFFD27852A858FC5CB170" box="[829,908,274,295]" country="Mexico" name="Chiapas" pageId="6" pageNumber="397">Chiapas</collectingRegion>
, state of
<collectingRegion id="5301F807FF8FFFD27BC0A867FF2EB115" box="[175,254,301,322]" country="Mexico" name="Chiapas" pageId="6" pageNumber="397">Chiapas</collectingRegion>
,
<collectingCountry id="E9D27675FF8FFFD27A65A867FE82B115" box="[266,338,301,322]" name="Mexico" pageId="6" pageNumber="397">México</collectingCountry>
(approx. 16
<geoCoordinate id="F4F15022FF8FFFD27AA6A866FDD5B115" box="[457,517,300,322]" degrees="8" direction="north" minutes="52" orientation="latitude" pageId="6" pageNumber="397" precision="15" seconds="9" value="8.869166">
<emphasis id="A3B1EAF7FF8FFFD27AA6A866FE01B115" bold="true" box="[457,465,300,322]" pageId="6" pageNumber="397">8</emphasis>
52
<emphasis id="A3B1EAF7FF8FFFD27A86A866FE21B115" bold="true" box="[489,497,300,322]" pageId="6" pageNumber="397">9</emphasis>
N
</geoCoordinate>
; 93
<geoCoordinate id="F4F15022FF8FFFD27946A866FDBBB115" box="[553,619,300,322]" degrees="8" direction="west" minutes="28" orientation="longitude" pageId="6" pageNumber="397" precision="15" seconds="9" value="-8.469166">
<emphasis id="A3B1EAF7FF8FFFD27946A866FDE1B115" bold="true" box="[553,561,300,322]" pageId="6" pageNumber="397">8</emphasis>
28
<emphasis id="A3B1EAF7FF8FFFD27926A866FD81B115" bold="true" box="[585,593,300,322]" pageId="6" pageNumber="397">9</emphasis>
W
</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="917A36E5FF8FFFD27BA8A80DFECCB1C6" blockId="6.[175,1005,194,614]" pageId="6" pageNumber="397">
<emphasis id="A3B1EAF7FF8FFFD27BA8A80DFEBEB10B" bold="true" box="[199,366,327,348]" pageId="6" pageNumber="397">Biological Data.</emphasis>
This species inhabits the remnants of tropical deciduous forest located near
<quantity id="563D9B00FF8FFFD27A5AA828FE5AB120" box="[309,394,354,375]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="397" unit="m" value="1000.0">1,000 m</quantity>
altitude. The single known specimen was collected during September.
</paragraph>
</subSubSection>
<subSubSection id="D9DF656EFF8FFFD27BA8A8DDFE93B266" pageId="6" pageNumber="397" type="discussion">
<paragraph id="917A36E5FF8FFFD27BA8A8DDFE93B266" blockId="6.[175,1005,194,614]" pageId="6" pageNumber="397">
<emphasis id="A3B1EAF7FF8FFFD27BA8A8DDFEFAB1FB" bold="true" box="[199,298,407,428]" pageId="6" pageNumber="397">Remarks.</emphasis>
<taxonomicName id="56C54D66FF8FFFD27A59A8DDFDFCB1FB" box="[310,556,407,428]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="397" phylum="Arthropoda" rank="species" species="nandalumia">
<emphasis id="A3B1EAF7FF8FFFD27A59A8DDFDFCB1FB" box="[310,556,407,428]" italics="true" pageId="6" pageNumber="397">Phyllophaga nandalumia</emphasis>
</taxonomicName>
belongs to the species group
<emphasis id="A3B1EAF7FF8FFFD27805A8DDFC09B1FB" box="[874,985,407,428]" italics="true" pageId="6" pageNumber="397">schizorhina</emphasis>
, subgroup 7 (
<emphasis id="A3B1EAF7FF8FFFD27A48A8F8FE8CB190" box="[295,348,434,455]" italics="true" pageId="6" pageNumber="397">sensu</emphasis>
Morón 2003). Externally the new species is similar to
<taxonomicName id="56C54D66FF8FFFD2780FA8F8FF20B1B6" authority="Moron" authorityName="Moron" authorityYear="2003" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="397" phylum="Arthropoda" rank="species" species="ocozocuana">
<emphasis id="A3B1EAF7FF8FFFD2780FA8F8FC3CB190" box="[864,1004,434,455]" italics="true" pageId="6" pageNumber="397">P. ocozocuana</emphasis>
Morón
</taxonomicName>
from Ocozocuautla and El Bosque,
<collectingRegion id="5301F807FF8FFFD27907A886FD67B1B6" box="[616,695,460,481]" country="Mexico" name="Chiapas" pageId="6" pageNumber="397">Chiapas</collectingRegion>
,
<collectingCountry id="E9D27675FF8FFFD279A9A886FCC1B1B6" box="[710,785,460,481]" name="Mexico" pageId="6" pageNumber="397">Mexico</collectingCountry>
, but the body size is smaller and the male genital capsule is much different, having the phallobase not expanded dorso-basally and the sclerotized support of the aedeagus not twisted as in
<taxonomicName id="56C54D66FF8FFFD27BC0AB56FEEEB266" authorityName="Moron" authorityYear="2003" box="[175,318,540,561]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="397" phylum="Arthropoda" rank="species" species="ocozocuana">
<emphasis id="A3B1EAF7FF8FFFD27BC0AB56FEEEB266" box="[175,318,540,561]" italics="true" pageId="6" pageNumber="397">P. ocozocuana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D9DF656EFF8FFFD27BA8AB7CFC04B231" pageId="6" pageNumber="397" type="etymology">
<paragraph id="917A36E5FF8FFFD27BA8AB7CFC04B231" blockId="6.[175,1005,194,614]" pageId="6" pageNumber="397">
<emphasis id="A3B1EAF7FF8FFFD27BA8AB7CFEEBB21C" bold="true" box="[199,315,566,587]" pageId="6" pageNumber="397">Etymology.</emphasis>
Derived from part of the name of the ancient Chiapa indigenous settlement
<emphasis id="A3B1EAF7FF8FFFD27A49AB1BFE08B231" box="[294,472,593,614]" italics="true" pageId="6" pageNumber="397">Socton Nandalumi</emphasis>
, located near Chiapa de Corzo (Vega
<emphasis id="A3B1EAF7FF8FFFD27832AB1BFC5FB231" box="[861,911,593,614]" italics="true" pageId="6" pageNumber="397">et al.</emphasis>
1984).
</paragraph>
</subSubSection>
</treatment>
</document>