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<document id="009DFEB19A3D5665370468136A860A5A" ID-DOI="10.3853/j.2201-4349.69.2017.1654" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237682" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629534585749" checkinUser="felipe" docAuthor="Irwin, Nancy" docDate="2017" docId="706DD361D44BFFE9FF29A91798F9CBE6" docLanguage="en" docName="RecAustMus.69.2.73-100.pdf" docOrigin="Records of the Australian Museum 69 (2)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1654" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Nyctimene certans , Andersen 1912" docType="treatment" docVersion="3" lastPageNumber="84" masterDocId="8C54AB19D443FFE2FFD6A84F9963CD48" masterDocTitle="A new Tube-nosed Fruit Bat from New Guinea, Nyctimene wrightae sp. nov., A Re-diagnosis of N. certans and N. cyclotis (Pteropodidae: Chiroptera), and a Review of their Conservation Status" masterLastPageNumber="100" masterPageNumber="73" pageNumber="81" updateTime="1699076801224" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="FBA7DE0BDC4BE201B8330DA826FD33A0">A new Tube-nosed Fruit Bat from New Guinea, Nyctimene wrightae sp. nov., A Re-diagnosis of N. certans and N. cyclotis (Pteropodidae: Chiroptera), and a Review of their Conservation Status</mods:title>
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<mods:namePart id="A1F66C68A2219E461ACFBC1D395FD59D">Irwin, Nancy</mods:namePart>
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<mods:date id="6CB5A0743ABAF7F5A25E2BB3EEB047AD">2017</mods:date>
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<paragraph id="F87B6277D44BFFEAFF29A9179BA2CC3A" blockId="8.[255,705,344,371]" box="[255,705,344,371]" pageId="8" pageNumber="81">
<heading id="A333D51BD44BFFEAFF29A9179BA2CC3A" box="[255,705,344,371]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="81" reason="2">
<taxonomicName id="3FC419F4D44BFFEAFF29A9179BA2CC3A" ID-CoL="485LD" authority="Andersen, 1912 b" authorityName=", Andersen" authorityYear="1912" box="[255,705,344,371]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFF29A9179BA2CC3A" bold="true" box="[255,705,344,371]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF29A9179887CC3B" bold="true" box="[255,484,344,371]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
<bibRefCitation id="9C551F86D44BFFEAFE3CA9179BA2CC3A" author="Andersen, K" box="[490,705,344,370]" pageId="8" pageNumber="81" pagination="95 - 96" refId="ref20612" refString="Andersen, K. 1912 b. A new Nyctimene from New Guinea. Annals and Magazine of Natural History (series 9) (8): 95 - 96. https: // doi. org / 10.1080 / 00222931208693111" type="journal article" year="1912">Andersen, 1912b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B0DE31FCD44BFFEAFE50A9C19B59CCE0" box="[390,570,398,425]" pageId="8" pageNumber="81" type="description">
<paragraph id="F87B6277D44BFFEAFE50A9C19B59CCE0" blockId="8.[390,570,398,425]" box="[390,570,398,425]" pageId="8" pageNumber="81">
<figureCitation id="60FF7EF2D44BFFEAFE50A9C19892CCE1" box="[390,497,398,425]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStartId-0="7.[518,581,1894,1916]" captionStartId-1="8.[835,898,932,954]" captionStartId-2="9.[121,184,1872,1894]" captionStartId-3="10.[166,229,1782,1804]" captionTargetBox-0="[122,1395,156,2011]" captionTargetBox-1="[835,1464,155,917]" captionTargetBox-2="[121,1421,151,1857]" captionTargetBox-3="[167,1467,151,1766]" captionTargetId-0="figure-10@7.[121,1398,151,2017]" captionTargetId-1="figure-620@8.[829,1468,151,920]" captionTargetId-2="figure-9@9.[121,1421,151,1857]" captionTargetId-3="figure-129@10.[167,1467,151,1766]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Figure 2. Holotype skull and mandible of Nyctimene cyclotis NHMUK 1910.7.16.9; (a) mandible, right side is broken and left M is missing; (b) ventral view of cranium, right P4 2" captionText-1="Figure 3. Plot of 5-Metacarpal divided by the 1st phalanx of the 5 digit, verses 2nd phalanx of the 2nd digit, indicating separation of Nyctimene certans and N. cyclotis using 3 simple wing measurements for 255 specimens. Nyctimene certans (squares, n = 35), N. a. papuanus (circles, n = 128), N. wrightae sp. nov. (triangles, n = 90) and N. cyclotis (diamonds, n = 2). Holotype specimens are indicated for each species by an asterisk. Full or closed symbols indicate individuals identified genetically (n = 77), open symbols are those identified morphologically (n = 179)." captionText-2="Figure 4. Dorsal and ventral views of skulls and mandibles of Nyctimene certans (young adult) (AM M.17888), N.certans (old, showing extensive tooth wear) (AM M.30648), N. wrightae sp. nov. (holotype) (AM M.16423) and N. a. papuanus (AM M.16425)." captionText-3="Figure 5. Dorsal and ventral photographs of the holotypes of Nyctimene certans and N. cyclotis. (a, b) Holotype of N. certans NHMUK 1911.11.29.1; photographs courtesyof HarryTaylor (NHMUK).(c, d) Holotype ofN.cyclotis NHMUK 1910.7.16.9; photographs courtesy of Jeff Streicher (NHMUK). The ears are short and rounded in N. certans and N. cyclotis. Both FAs of N. cyclotis are clearly broken. The dorsal stripe is difficult to distinguish in the base of the dorsal in N. certans and N. cyclotis. Overall there is a grizzled grey-brown appearance in the fur in the dorsal of N. certans and N. cyclotis, compared to a more even brown colour in N. wrightae sp. nov. and N. a. papuanus (see Fig. 6)." figureDoi-0="http://doi.org/10.5281/zenodo.5237686" figureDoi-1="http://doi.org/10.5281/zenodo.5237689" figureDoi-2="http://doi.org/10.5281/zenodo.5237691" figureDoi-3="http://doi.org/10.5281/zenodo.5237693" httpUri-0="https://zenodo.org/record/5237686/files/figure.png" httpUri-1="https://zenodo.org/record/5237689/files/figure.png" httpUri-2="https://zenodo.org/record/5237691/files/figure.png" httpUri-3="https://zenodo.org/record/5237693/files/figure.png" pageId="8" pageNumber="81">Figs 25</figureCitation>
,
<figureCitation id="60FF7EF2D44BFFEAFE28A9C19B59CCE0" box="[510,570,398,424]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStart-4="Figure 11" captionStart-5="Figure 12" captionStartId-0="14.[166,229,1886,1908]" captionStartId-1="15.[121,183,894,916]" captionStartId-2="15.[794,857,894,916]" captionStartId-3="16.[170,233,983,1005]" captionStartId-4="16.[845,908,982,1004]" captionStartId-5="17.[121,184,1902,1924]" captionTargetBox-0="[166,1466,151,1873]" captionTargetBox-1="[112,1424,149,872]" captionTargetBox-3="[157,1469,153,963]" captionTargetBox-5="[121,1421,151,1878]" captionTargetId-0="figure-7@14.[166,1466,151,1873]" captionTargetId-5="figure-48@17.[121,1421,151,1883]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-3="16" captionTargetPageId-5="17" captionText-0="Figure 7. Photographs of Nyctimene wrightae sp. nov. and N. certans. (a) Female with young pup attached, showing characteristic long ears with thickened edge and short, brown hair (photograph courtesy of Debra Wright); (b) N. certans, showing darker fur than N. wrightae sp. nov. and round, broad ears (photograph courtesy of Debra Wright); (c) N. wrightae sp. nov. yellow colour morph with large brown spots caught in O-Pio, Chimbu Prov. (field number NRI 721), the dorsal stripe is clear and becomes thinner in upper part of back." captionText-1="Figure 8. First two axes of a LDFA based on 14 factors of combined skull, dental and external body measurements (listed in Table 4), showing separation between N. a. papuanus (circles, n = 44), N. certans (squares, n = 15) and N. wrightae sp. nov. (triangles, n = 11). Holotype of N. wrightae sp. nov. is indicated by star symbol." captionText-2="Figure 9. Separation of three species on the first two axes of a LDFA based on 14 skull and dental measurements (listed in Table 5). Nyctimene a. papuanus (circles, n = 39), N. certans (squares, n = 19), N. wrightae sp. nov. (triangles, n = 10). Star indicates holotype of N. wrightae sp. nov." captionText-3="Figure 10. Plot of breadth against M1 breadth for Nyctimene a. papuanus (circles,n =58) N.certans (squares,n = 27), N.wrightae sp. nov. (triangles, n = 11) and N. cyclotis (diamond, n = 2). Filled symbols are genetically identified. Holotypes of each species indicated by star symbols. Smaller outliers of N. certans are old individuals with smaller teeth due to extensive tooth wear.Outliers of N. wrightae sp. nov. with larger teeth than their conspecifics are young individuals with little tooth wear." captionText-4="Figure 11. First two axes of a LDFA of three Nyctimene species based on 7 external wing measurements (listed in Table 6), n = 242. Separation is achieved between individuals of N. certans (squares, n = 32) from N. wrightae sp. nov. (triangles, n = 88) and N. a. papuanus (circles, n = 122) but there remains some overlap between N. wrightae sp. nov. and N. a. papuanus. Holotypes are indicated by star symbol. The holotype of N. cyclotis was plotted a posteriori (diamond)." captionText-5="Figure 12. Distribution of specimens assigned in this study to N. cyclotis (stars, n = 2), N. certans (squares, n = 26), N. wrightae sp. nov. (triangles n = 22) and N. a. papuanus (circles n = 48)." figureDoi-0="http://doi.org/10.5281/zenodo.5237697" figureDoi-1="http://doi.org/10.5281/zenodo.5237699" figureDoi-3="http://doi.org/10.5281/zenodo.5237701" figureDoi-5="http://doi.org/10.5281/zenodo.5237703" httpUri-0="https://zenodo.org/record/5237697/files/figure.png" httpUri-1="https://zenodo.org/record/5237699/files/figure.png" httpUri-3="https://zenodo.org/record/5237701/files/figure.png" httpUri-5="https://zenodo.org/record/5237703/files/figure.png" pageId="8" pageNumber="81">712</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B0DE31FCD44BFFEAFF70A9F29BCBCF67" pageId="8" pageNumber="81" type="materials_examined">
<paragraph id="F87B6277D44BFFEAFF70A9F29A6ACC9D" blockId="8.[166,777,445,469]" box="[166,777,445,469]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF70A9F2985BCC9D" bold="true" box="[166,312,445,469]" pageId="8" pageNumber="81">Type locality</emphasis>
. Mount Goliath, Prov. of
<collectingRegion id="3A00AC95D44BFFEAFD9DA9F29BECCC9C" box="[587,655,445,468]" country="Indonesia" name="Papua Barat" pageId="8" pageNumber="81">Papua</collectingRegion>
,
<collectingCountry id="80D322E7D44BFFEAFD4FA9F29A67CC9D" box="[665,772,445,469]" name="Indonesia" pageId="8" pageNumber="81">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="F87B6277D44BFFEAFF70A9A59A74CF4A" blockId="8.[166,791,490,514]" box="[166,791,490,514]" pageId="8" pageNumber="81">
<materialsCitation id="48AC682AD44BFFEAFF70A9A59A74CF4A" box="[166,791,490,514]" collectionCode="NHMUK" pageId="8" pageNumber="81" specimenCode="NHMUK 1911.11" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="CAB0BE65D44BFFEAFF70A9A5986DCF4A" bold="true" box="[166,270,490,514]" pageId="8" pageNumber="81">
<typeStatus id="277FDCD5D44BFFEAFF70A9A5986DCF4A" box="[166,270,490,514]" pageId="8" pageNumber="81" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="A862CA0CD44BFFEAFECDA9A49882CF4A" box="[283,481,490,514]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="81" type="Museum">NHMUK 1911.11</specimenCode>
.29.1.
<specimenCount id="EEC2A9FED44BFFEAFDC8A9A49B51CF4A" box="[542,562,491,514]" pageId="8" pageNumber="81" type="male"></specimenCount>
adult, skin and skull.
</materialsCitation>
</paragraph>
<paragraph id="F87B6277D44BFFEAFF70AA589BCBCF67" blockId="8.[166,680,535,560]" box="[166,680,535,560]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF70AA589837CF78" bold="true" box="[166,340,535,560]" pageId="8" pageNumber="81">Common name</emphasis>
. Mountain tube-nosed fruit bat.
</paragraph>
</subSubSection>
<subSubSection id="B0DE31FCD44BFFEAFF70AA0A98D7C927" pageId="8" pageNumber="81" type="distribution">
<paragraph id="F87B6277D44BFFEAFF70AA0A98C3CFDF" blockId="8.[166,794,581,663]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF70AA0A9852CF15" bold="true" box="[166,305,581,605]" pageId="8" pageNumber="81">Distribution</emphasis>
. Upper montane forest of New
<collectingCountry id="80D322E7D44BFFEAFD56AA0A9BB3CF15" box="[640,720,581,605]" name="Guinea-Bissau" pageId="8" pageNumber="81">Guinea</collectingCountry>
, above
<quantity id="3F3CCF92D44BFFEAFF70AA2D998ECF32" box="[166,237,610,634]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="81" unit="m" value="700.0">700 m</quantity>
up to
<quantity id="3F3CCF92D44BFFEAFEECAA2D98ECCF32" box="[314,399,610,634]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="81" unit="m" value="3000.0">3000 m</quantity>
asl but typically most localities lie between
<quantity id="3F3CCF92D44BFFEAFED0AA3098FFCFDF" box="[262,412,639,663]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="8" pageNumber="81" unit="m" value="2050.0" valueMax="2500.0" valueMin="1600.0">16002500 m</quantity>
.
</paragraph>
<paragraph id="F87B6277D44BFFEAFF70AAE29849CFAA" blockId="8.[166,793,685,738]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF70AAE298C0CF8D" bold="true" box="[166,419,685,709]" pageId="8" pageNumber="81">Specimens examined</emphasis>
. 39 including the
<typeStatus id="277FDCD5D44BFFEAFD56AAE29B85CF8D" box="[640,742,685,709]" pageId="8" pageNumber="81" type="holotype">holotype</typeStatus>
, see Appendix 1.
</paragraph>
<paragraph id="F87B6277D44BFFEAFF70AAB898D7C927" blockId="8.[164,795,759,1135]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF70AAB89817CE47" bold="true" box="[166,372,759,783]" pageId="8" pageNumber="81">Revised diagnosis.</emphasis>
<taxonomicName id="3FC419F4D44BFFEAFEAFAAB79B5FCE47" authorityName=", Andersen" authorityYear="1912" box="[377,572,760,783]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFEAFAAB79B5FCE47" box="[377,572,760,783]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
differs from all other
<taxonomicName id="3FC419F4D44BFFEAFF70AB59980ACE65" box="[166,361,789,813]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="CAB0BE65D44BFFEAFF70AB599874CE65" box="[166,279,790,813]" italics="true" pageId="8" pageNumber="81">Nyctimene</emphasis>
species
</taxonomicName>
in having the shortest upper (&lt;32%) and lower (&lt;36%) tooth rows relative to skull length (
<tableCitation id="B54657CCD44BFFEAFD17AB7D9A70CE02" box="[705,787,818,842]" captionStart="Table 2" captionStartId="6.[166,229,171,195]" captionText="Table 2. Summary statistics of selected external, cranial and dental variables used in this study for Nyctimene wrightae sp. nov., N. cyclotis, N. certans and N. a. papuanus (mm). H = holotype of N. wrightae sp. nov., AM.M16423." pageId="8" pageNumber="81">Table 2</tableCitation>
,
<figureCitation id="60FF7EF2D44BFFEAFF71AB00998ACE20" box="[167,233,847,872]" captionStart="Figure 2" captionStartId="7.[518,581,1894,1916]" captionTargetBox="[122,1395,156,2011]" captionTargetId="figure-10@7.[121,1398,151,2017]" captionTargetPageId="7" captionText="Figure 2. Holotype skull and mandible of Nyctimene cyclotis NHMUK 1910.7.16.9; (a) mandible, right side is broken and left M is missing; (b) ventral view of cranium, right P4 2" figureDoi="http://doi.org/10.5281/zenodo.5237686" httpUri="https://zenodo.org/record/5237686/files/figure.png" pageId="8" pageNumber="81">Figs 2</figureCitation>
,
<figureCitation id="60FF7EF2D44BFFEAFF24AB1F9861CE20" box="[242,258,848,872]" captionStart="Figure 4" captionStartId="9.[121,184,1872,1894]" captionTargetBox="[121,1421,151,1857]" captionTargetId="figure-9@9.[121,1421,151,1857]" captionTargetPageId="9" captionText="Figure 4. Dorsal and ventral views of skulls and mandibles of Nyctimene certans (young adult) (AM M.17888), N.certans (old, showing extensive tooth wear) (AM M.30648), N. wrightae sp. nov. (holotype) (AM M.16423) and N. a. papuanus (AM M.16425)." figureDoi="http://doi.org/10.5281/zenodo.5237691" httpUri="https://zenodo.org/record/5237691/files/figure.png" pageId="8" pageNumber="81">4</figureCitation>
).
<taxonomicName id="3FC419F4D44BFFEAFEC4AB1F98B4CE2F" authorityName=", Andersen" authorityYear="1912" box="[274,471,848,871]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFEC4AB1F98B4CE2F" box="[274,471,848,871]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="3FC419F4D44BFFEAFD7DAB1F9A7ACE2F" box="[683,793,847,871]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D44BFFEAFD7DAB1F9A7ACE2F" box="[683,793,847,871]" italics="true" pageId="8" pageNumber="81">N. cyclotis</emphasis>
</taxonomicName>
in having a relatively broader palate, with a P
<superScript id="0FB1CF3FD44BFFEAFD51AB249BECCE31" attach="left" box="[647,655,875,889]" fontSize="6" pageId="8" pageNumber="81">3</superScript>
-P
<superScript id="0FB1CF3FD44BFFEAFD71AB249BCCCE31" attach="left" box="[679,687,875,889]" fontSize="6" pageId="8" pageNumber="81">3</superScript>
markedly longer than the upper post-canine tooth row (P
<superScript id="0FB1CF3FD44BFFEAFD66ABC69BDBCEDF" attach="left" box="[688,696,905,919]" fontSize="6" pageId="8" pageNumber="81">1</superScript>
-M
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), vs. slightly longer than P
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-M
<quantity id="3F3CCF92D44BFFEAFE66ABE998B1CEF7" box="[432,466,934,959]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="81" unit="in" value="1.0">
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in
</quantity>
<emphasis id="CAB0BE65D44BFFEAFE00ABE79BA2CEF7" box="[470,705,935,959]" italics="true" pageId="8" pageNumber="81">
<taxonomicName id="3FC419F4D44BFFEAFE00ABE79B24CEF7" box="[470,583,935,959]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">N. cyclotis</taxonomicName>
;
<taxonomicName id="3FC419F4D44BFFEAFD83ABE79BA2CEF7" authorityName=", Andersen" authorityYear="1912" box="[597,705,936,959]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">N. certans</taxonomicName>
</emphasis>
also has a longer second digit, especially the 2M-
<specimenCount id="EEC2A9FED44BFFEAFD89AB8A9B1CCE95" box="[607,639,965,989]" pageId="8" pageNumber="81" type="generic">2P.</specimenCount>
On average it is larger and darker dorsally than any other medium-sized
<taxonomicName id="3FC419F4D44BFFEAFF7EAC4F9808C95F" box="[168,363,1023,1047]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="CAB0BE65D44BFFEAFF7EAC4F987AC95F" box="[168,281,1024,1047]" italics="true" pageId="8" pageNumber="81">Nyctimene</emphasis>
species
</taxonomicName>
from mainland New
<collectingCountry id="80D322E7D44BFFEAFD84ABB09BC0C95F" box="[594,675,1023,1047]" name="Guinea-Bissau" pageId="8" pageNumber="81">Guinea</collectingCountry>
. It also has the broadest cheek-teeth relative to skull size (esp. P
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), which are almost as broad as they are long, compared to any other species of
<taxonomicName id="3FC419F4D44BFFEAFEC1AC1898D3C927" authorityName="Miller" authorityYear="1907" box="[279,432,1111,1135]" pageId="8" pageNumber="93" rank="subFamily" subFamily="Nyctimeninae">Nyctimeninae</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="ACBB32FFD44BFFEAFC95ABEB9CDBC9D8" ID-DOI="http://doi.org/10.5281/zenodo.5237689" ID-Zenodo-Dep="5237689" httpUri="https://zenodo.org/record/5237689/files/figure.png" pageId="8" pageNumber="81" startId="8.[835,898,932,954]" targetBox="[835,1464,155,917]" targetPageId="8">
<paragraph id="F87B6277D44BFFEAFC95ABEB9CDBC9D8" blockId="8.[835,1467,932,1168]" pageId="8" pageNumber="81">
Figure 3. Plot of 5-Metacarpal divided by the 1st phalanx of the 5 digit, verses 2nd phalanx of the 2nd digit, indicating separation of
<taxonomicName id="3FC419F4D44BFFEAFCB2AB959D44CEA7" authorityName=", Andersen" authorityYear="1912" box="[868,1063,986,1007]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFCB2AB959D44CEA7" box="[868,1063,986,1007]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
and
<taxonomicName id="3FC419F4D44BFFEAFBB7AB959DB0CEA7" box="[1121,1235,986,1007]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D44BFFEAFBB7AB959DB0CEA7" box="[1121,1235,986,1007]" italics="true" pageId="8" pageNumber="81">N. cyclotis</emphasis>
</taxonomicName>
using 3 simple wing measurements for 255 specimens.
<taxonomicName id="3FC419F4D44BFFEAFB76ABBA9C3BC942" authorityName=", Andersen" authorityYear="1912" box="[1184,1368,1013,1034]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFB76ABBA9C3BC942" box="[1184,1368,1013,1034]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
(squares,
<emphasis id="CAB0BE65D44BFFEAFC92AC5F9A0BC96D" box="[836,872,1040,1061]" italics="true" pageId="8" pageNumber="81">n =</emphasis>
35),
<emphasis id="CAB0BE65D44BFFEAFC4BAC5F9D52C96D" box="[925,1073,1040,1061]" italics="true" pageId="8" pageNumber="81">
N. a.
<taxonomicName id="3FC419F4D44BFFEAFC02AC5F9D52C96D" authorityName="Andersen" authorityYear="1910" box="[980,1073,1040,1061]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="papuanus">papuanus</taxonomicName>
</emphasis>
(circles,
<emphasis id="CAB0BE65D44BFFEAFB59AC5F9DD1C96D" box="[1167,1202,1040,1061]" italics="true" pageId="8" pageNumber="81">n =</emphasis>
128),
<taxonomicName id="3FC419F4D44BFFEAFB24AC5F9C06C96C" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[1266,1381,1039,1061]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="wrightae" status="sp. nov.">
<emphasis id="CAB0BE65D44BFFEAFB24AC5F9C06C96C" box="[1266,1381,1039,1061]" italics="true" pageId="8" pageNumber="81">N. wrightae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D183031ED44BFFEAFAB8AC409CD8C96C" box="[1390,1467,1039,1060]" pageId="8" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
(triangles,
<emphasis id="CAB0BE65D44BFFEAFC7BAC659AB2C977" box="[941,977,1066,1087]" italics="true" pageId="8" pageNumber="81">n =</emphasis>
90) and
<taxonomicName id="3FC419F4D44BFFEAFBFBAC659DF7C977" box="[1069,1172,1066,1087]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D44BFFEAFBFBAC659DF7C977" box="[1069,1172,1066,1087]" italics="true" pageId="8" pageNumber="81">N. cyclotis</emphasis>
</taxonomicName>
(diamonds,
<emphasis id="CAB0BE65D44BFFEAFAC6AC659C57C977" box="[1296,1332,1066,1087]" italics="true" pageId="8" pageNumber="81">n =</emphasis>
2). Holotype specimens are indicated for each species by an asterisk. Full or closed symbols indicate individuals identified genetically (
<emphasis id="CAB0BE65D44BFFEAFA43AC2F9CC2C93D" box="[1429,1441,1120,1141]" italics="true" pageId="8" pageNumber="81">n</emphasis>
= 77), open symbols are those identified morphologically (
<emphasis id="CAB0BE65D44BFFEAFA89AC359CE3C9C7" box="[1375,1408,1146,1167]" italics="true" pageId="8" pageNumber="81">n =</emphasis>
179).
</paragraph>
</caption>
<subSubSection id="B0DE31FCD44BFFEAFF70ACCA9A95CA5A" pageId="8" pageNumber="81" type="description">
<paragraph id="F87B6277D44BFFEAFF70ACCA98C4C85A" blockId="8.[164,796,1157,2002]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFF70ACCA984FC9D5" bold="true" box="[166,300,1157,1181]" pageId="8" pageNumber="81">Description</emphasis>
. Essentially Andersens original description stands (
<bibRefCitation id="9C551F86D44BFFEAFF21ACED98D0C9F2" author="Andersen, K" box="[247,435,1186,1210]" pageId="8" pageNumber="81" pagination="95 - 96" refId="ref20612" refString="Andersen, K. 1912 b. A new Nyctimene from New Guinea. Annals and Magazine of Natural History (series 9) (8): 95 - 96. https: // doi. org / 10.1080 / 00222931208693111" type="journal article" year="1912">Andersen, 1912b</bibRefCitation>
), validated by a larger series than the
<specimenCount id="EEC2A9FED44BFFEAFF1AAC8F9805C99F" box="[204,358,1215,1239]" pageId="8" pageNumber="81" type="generic">two specimens</specimenCount>
available to Andersen.
<taxonomicName id="3FC419F4D44BFFEAFD8EAC8F9A79C99F" authorityName=", Andersen" authorityYear="1912" box="[600,794,1216,1239]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFD8EAC8F9A79C99F" box="[600,794,1216,1239]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
is a medium-sized species for the subfamily with a forearm range of
<quantity id="3F3CCF92D44BFFEAFED3ACB598C0C85A" box="[261,419,1274,1298]" metricMagnitude="-2" metricUnit="m" metricValue="6.11" metricValueMax="6.74" metricValueMin="5.4799999999999995" pageId="8" pageNumber="81" unit="mm" value="61.1" valueMax="67.4" valueMin="54.8">54.867.4 mm</quantity>
.
</paragraph>
<paragraph id="F87B6277D44BFFEAFF13AD589807C8ED" blockId="8.[164,796,1157,2002]" pageId="8" pageNumber="81">
The fur colour is dark grey and grizzled with wood-brown on the back; it has for the subfamily, the largest proportion of the base of the hair shaft being dark brown (3040%) and likewise the tips of the hairs dark brown (1030% of the hair shaft) (
<figureCitation id="60FF7EF2D44BFFEAFF26ADC29858C8ED" box="[240,315,1421,1445]" captionStart="Figure 5" captionStartId="10.[166,229,1782,1804]" captionTargetBox="[167,1467,151,1766]" captionTargetId="figure-129@10.[167,1467,151,1766]" captionTargetPageId="10" captionText="Figure 5. Dorsal and ventral photographs of the holotypes of Nyctimene certans and N. cyclotis. (a, b) Holotype of N. certans NHMUK 1911.11.29.1; photographs courtesyof HarryTaylor (NHMUK).(c, d) Holotype ofN.cyclotis NHMUK 1910.7.16.9; photographs courtesy of Jeff Streicher (NHMUK). The ears are short and rounded in N. certans and N. cyclotis. Both FAs of N. cyclotis are clearly broken. The dorsal stripe is difficult to distinguish in the base of the dorsal in N. certans and N. cyclotis. Overall there is a grizzled grey-brown appearance in the fur in the dorsal of N. certans and N. cyclotis, compared to a more even brown colour in N. wrightae sp. nov. and N. a. papuanus (see Fig. 6)." figureDoi="http://doi.org/10.5281/zenodo.5237693" httpUri="https://zenodo.org/record/5237693/files/figure.png" pageId="8" pageNumber="81">Figs. 5</figureCitation>
,
<figureCitation id="60FF7EF2D44BFFEAFE91ADC29834C8ED" box="[327,343,1421,1445]" captionStart="Figure 7" captionStartId="14.[166,229,1886,1908]" captionTargetBox="[166,1466,151,1873]" captionTargetId="figure-7@14.[166,1466,151,1873]" captionTargetPageId="14" captionText="Figure 7. Photographs of Nyctimene wrightae sp. nov. and N. certans. (a) Female with young pup attached, showing characteristic long ears with thickened edge and short, brown hair (photograph courtesy of Debra Wright); (b) N. certans, showing darker fur than N. wrightae sp. nov. and round, broad ears (photograph courtesy of Debra Wright); (c) N. wrightae sp. nov. yellow colour morph with large brown spots caught in O-Pio, Chimbu Prov. (field number NRI 721), the dorsal stripe is clear and becomes thinner in upper part of back." figureDoi="http://doi.org/10.5281/zenodo.5237697" httpUri="https://zenodo.org/record/5237697/files/figure.png" pageId="8" pageNumber="81">7</figureCitation>
).
</paragraph>
<paragraph id="F87B6277D44BFFEAFF13ADE4981BCB1D" blockId="8.[164,796,1157,2002]" pageId="8" pageNumber="81">
<taxonomicName id="3FC419F4D44BFFEAFF13ADE498F9C88A" authorityName=", Andersen" authorityYear="1912" box="[197,410,1451,1474]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFF13ADE498F9C88A" box="[197,410,1451,1474]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
has a significantly longer skull, with CBL, C
<subScript id="64406032D44BFFEAFEE5AD9B9858C8AA" attach="left" box="[307,315,1492,1506]" fontSize="6" pageId="8" pageNumber="81">1</subScript>
-M
<subScript id="64406032D44BFFEAFE8AAD9B9807C8AA" attach="right" box="[348,356,1492,1506]" fontSize="6" pageId="8" pageNumber="81">2</subScript>
, Zygo-L larger than the other sympatric medium-sized species (
<tableCitation id="B54657CCD44BFFEAFE7FADAA9B6BC8B5" box="[425,520,1509,1533]" captionStart="Table 1" captionStartId="1.[121,184,1197,1221]" captionText="Table 1. A summary of different taxonomic arrangements of Nyctimene taxa by selected authors subsequent to Andersen." pageId="8" pageNumber="81">Tables 1</tableCitation>
and
<tableCitation id="B54657CCD44BFFEAFD92ADAA9B37C8B5" box="[580,596,1509,1533]" captionStart="Table 2" captionStartId="6.[166,229,171,195]" captionText="Table 2. Summary statistics of selected external, cranial and dental variables used in this study for Nyctimene wrightae sp. nov., N. cyclotis, N. certans and N. a. papuanus (mm). H = holotype of N. wrightae sp. nov., AM.M16423." pageId="8" pageNumber="81">2</tableCitation>
). Its brain case is longer but not as broad as the similar
<taxonomicName id="3FC419F4D44BFFEAFD97AE4C9BA3CB52" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[577,704,1538,1562]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="wrightae" status="sp. nov.">
<emphasis id="CAB0BE65D44BFFEAFD97AE4C9BA3CB52" box="[577,704,1538,1562]" italics="true" pageId="8" pageNumber="81">N. wrightae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D183031ED44BFFEAFD1EAE4C9A7FCB52" box="[712,796,1539,1562]" pageId="8" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="60FF7EF2D44BFFEAFF78AE509990CB70" box="[174,243,1567,1592]" captionStart="Figure 4" captionStartId="9.[121,184,1872,1894]" captionTargetBox="[121,1421,151,1857]" captionTargetId="figure-9@9.[121,1421,151,1857]" captionTargetPageId="9" captionText="Figure 4. Dorsal and ventral views of skulls and mandibles of Nyctimene certans (young adult) (AM M.17888), N.certans (old, showing extensive tooth wear) (AM M.30648), N. wrightae sp. nov. (holotype) (AM M.16423) and N. a. papuanus (AM M.16425)." figureDoi="http://doi.org/10.5281/zenodo.5237691" httpUri="https://zenodo.org/record/5237691/files/figure.png" pageId="8" pageNumber="81">Fig. 4</figureCitation>
). Further specimens may also show this to be the case for
<emphasis id="CAB0BE65D44BFFEAFED7AE71981BCB1D" box="[257,376,1597,1621]" italics="true" pageId="8" pageNumber="81">
<taxonomicName id="3FC419F4D44BFFEAFED7AE719816CB1D" box="[257,373,1597,1621]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">N. cyclotis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="F87B6277D44BFFEAFF13AE1598CBCA08" blockId="8.[164,796,1157,2002]" pageId="8" pageNumber="81">
The skull of the
<typeStatus id="277FDCD5D44BFFEAFEABAE1598B9CB3A" box="[381,474,1626,1650]" pageId="8" pageNumber="81" type="holotype">holotype</typeStatus>
specimen is badly broken as noted previously (
<bibRefCitation id="9C551F86D44BFFEAFEBEAE379B6FCBD8" author="Peterson, R" box="[360,524,1656,1680]" pageId="8" pageNumber="81" pagination="26 - 41" refId="ref23034" refString="Peterson, R. L. 1991. Systematic variation in the Megachiropteran Tube-nosed Bats Nyctimene cyclotis and N. certans. Bulletin of the American Museum of Natural History 206: 26 - 41." type="journal article" year="1991">Peterson, 1991</bibRefCitation>
;
<bibRefCitation id="9C551F86D44BFFEAFDC0AE389BB1CBC7" author="Andersen, K" box="[534,722,1655,1679]" pageId="8" pageNumber="81" pagination="95 - 96" refId="ref20612" refString="Andersen, K. 1912 b. A new Nyctimene from New Guinea. Annals and Magazine of Natural History (series 9) (8): 95 - 96. https: // doi. org / 10.1080 / 00222931208693111" type="journal article" year="1912">Andersen, 1912b</bibRefCitation>
). Only the anterior half of the skull remains, consisting of the palate, rostrum and half of the left zygomatic arch (
<figureCitation id="60FF7EF2D44BFFEAFD5DAEFD9BADCB82" box="[651,718,1714,1738]" captionStart="Figure 2" captionStartId="7.[518,581,1894,1916]" captionTargetBox="[122,1395,156,2011]" captionTargetId="figure-10@7.[121,1398,151,2017]" captionTargetPageId="7" captionText="Figure 2. Holotype skull and mandible of Nyctimene cyclotis NHMUK 1910.7.16.9; (a) mandible, right side is broken and left M is missing; (b) ventral view of cranium, right P4 2" figureDoi="http://doi.org/10.5281/zenodo.5237686" httpUri="https://zenodo.org/record/5237686/files/figure.png" pageId="8" pageNumber="81">Fig. 2</figureCitation>
), from which the characteristic rounded dental arcade, and large, broad, cuspidate teeth are apparent. The lower mandible is in better condition with only the right angular process broken off and missing (
<figureCitation id="60FF7EF2D44BFFEAFE8FAF6898F8CA08" box="[345,411,1831,1856]" captionStart="Figure 2" captionStartId="7.[518,581,1894,1916]" captionTargetBox="[122,1395,156,2011]" captionTargetId="figure-10@7.[121,1398,151,2017]" captionTargetPageId="7" captionText="Figure 2. Holotype skull and mandible of Nyctimene cyclotis NHMUK 1910.7.16.9; (a) mandible, right side is broken and left M is missing; (b) ventral view of cranium, right P4 2" figureDoi="http://doi.org/10.5281/zenodo.5237686" httpUri="https://zenodo.org/record/5237686/files/figure.png" pageId="8" pageNumber="81">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="F87B6277D44BFFEAFF13AF0A9ABEC887" blockId="8.[164,796,1157,2002]" lastBlockId="8.[837,1468,1258,1810]" pageId="8" pageNumber="81">
<bibRefCitation id="9C551F86D44BFFEAFF13AF0A9809CA15" author="Peterson, R" box="[197,362,1861,1885]" pageId="8" pageNumber="81" pagination="26 - 41" refId="ref23034" refString="Peterson, R. L. 1991. Systematic variation in the Megachiropteran Tube-nosed Bats Nyctimene cyclotis and N. certans. Bulletin of the American Museum of Natural History 206: 26 - 41." type="journal article" year="1991">Peterson (1991)</bibRefCitation>
discussed the possibility that the
<typeStatus id="277FDCD5D44BFFEAFD68AF0A9A79CA15" box="[702,794,1861,1885]" pageId="8" pageNumber="81" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3FC419F4D44BFFEAFF13AF2C9856CA32" authorityName=", Andersen" authorityYear="1912" box="[197,309,1891,1914]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFF13AF2C9856CA32" box="[197,309,1891,1914]" italics="true" pageId="8" pageNumber="81">N. certans</emphasis>
</taxonomicName>
is a sub-adult due to the metacarpal lengths being relatively short, together with the highly cuspidate teeth. Indeed the type specimen has one of the shortest FA and metacarpals of the specimens assigned to
<taxonomicName id="3FC419F4D44BFFEAFDAAAFF49B86CA9A" authorityName=", Andersen" authorityYear="1912" box="[636,741,1979,2002]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFDAAAFF49B86CA9A" box="[636,741,1979,2002]" italics="true" pageId="8" pageNumber="81">N. certans</emphasis>
</taxonomicName>
here and the 5M-1P and 5M-2P are particularly short. The teeth are large and compacted; additionally the cusps are well defined similar to a sub-adult. However the teeth do show wear, as indicated by exposed dentine on the molars, suggestive of some use, more than that anticipated in a juvenile. Therefore overall, although the individual is clearly not very old, I concur with
<bibRefCitation id="9C551F86D44BFFEAFC10ADD59DE7C8FA" author="Andersen, K" box="[966,1156,1434,1458]" pageId="8" pageNumber="81" pagination="95 - 96" refId="ref20612" refString="Andersen, K. 1912 b. A new Nyctimene from New Guinea. Annals and Magazine of Natural History (series 9) (8): 95 - 96. https: // doi. org / 10.1080 / 00222931208693111" type="journal article" year="1912">Andersen (1912b)</bibRefCitation>
who considered the specimen to be an adult.
</paragraph>
<paragraph id="F87B6277D44BFFEAFCB3AD9B9A95CA5A" blockId="8.[837,1468,1258,1810]" pageId="8" pageNumber="81">
The wings of
<taxonomicName id="3FC419F4D44BFFEAFBD4AD9A9D10C8A4" authorityName=", Andersen" authorityYear="1912" box="[1026,1139,1493,1516]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFBD4AD9A9D10C8A4" box="[1026,1139,1493,1516]" italics="true" pageId="8" pageNumber="81">N. certans</emphasis>
</taxonomicName>
are deeper than its sympatric congeneric species; primarily caused by a significantly longer 5th digit, especially the 1st phalanx (
<tableCitation id="B54657CCD44BFFEAFB13AE409C43CB6F" box="[1221,1312,1551,1575]" captionStart="Table 2" captionStartId="6.[166,229,171,195]" captionText="Table 2. Summary statistics of selected external, cranial and dental variables used in this study for Nyctimene wrightae sp. nov., N. cyclotis, N. certans and N. a. papuanus (mm). H = holotype of N. wrightae sp. nov., AM.M16423." pageId="8" pageNumber="81">Tables 2</tableCitation>
,
<tableCitation id="B54657CCD44BFFEAFAFCAE409C54CB6F" box="[1322,1335,1551,1575]" captionStart="Table 3" captionStartId="11.[121,184,167,191]" captionText="Table 3. Welch Two Sample ttests for each pair of species, on 24 variables used in the LDFA." pageId="8" pageNumber="81">3</tableCitation>
). The wings also, on average, have significantly longer 2M-2P than their sympatric congenerics (
<tableCitation id="B54657CCD44BFFEAFB98AE059DCCCB2A" box="[1102,1199,1610,1634]" captionStart="Table 2" captionStartId="6.[166,229,171,195]" captionText="Table 2. Summary statistics of selected external, cranial and dental variables used in this study for Nyctimene wrightae sp. nov., N. cyclotis, N. certans and N. a. papuanus (mm). H = holotype of N. wrightae sp. nov., AM.M16423." pageId="8" pageNumber="81">Tables 2</tableCitation>
,
<tableCitation id="B54657CCD44BFFEAFB69AE059DADCB2A" box="[1215,1230,1610,1634]" captionStart="Table 3" captionStartId="11.[121,184,167,191]" captionText="Table 3. Welch Two Sample ttests for each pair of species, on 24 variables used in the LDFA." pageId="8" pageNumber="81">3</tableCitation>
). Bats with a 2M-2P above 8.0 mm and a ratio 5-MET / 5M-1P below 2.0 are therefore likely to be
<taxonomicName id="3FC419F4D44BFFEAFBF8AECA9DFECBD4" authorityName=", Andersen" authorityYear="1912" box="[1070,1181,1669,1692]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFBF8AECA9DFECBD4" box="[1070,1181,1669,1692]" italics="true" pageId="8" pageNumber="81">N. certans</emphasis>
</taxonomicName>
(
<figureCitation id="60FF7EF2D44BFFEAFB7DAECB9D8FCBD5" box="[1195,1260,1668,1693]" captionStart="Figure 3" captionStartId="8.[835,898,932,954]" captionTargetBox="[835,1464,155,917]" captionTargetId="figure-620@8.[829,1468,151,920]" captionTargetPageId="8" captionText="Figure 3. Plot of 5-Metacarpal divided by the 1st phalanx of the 5 digit, verses 2nd phalanx of the 2nd digit, indicating separation of Nyctimene certans and N. cyclotis using 3 simple wing measurements for 255 specimens. Nyctimene certans (squares, n = 35), N. a. papuanus (circles, n = 128), N. wrightae sp. nov. (triangles, n = 90) and N. cyclotis (diamonds, n = 2). Holotype specimens are indicated for each species by an asterisk. Full or closed symbols indicate individuals identified genetically (n = 77), open symbols are those identified morphologically (n = 179)." figureDoi="http://doi.org/10.5281/zenodo.5237689" httpUri="https://zenodo.org/record/5237689/files/figure.png" pageId="8" pageNumber="81">Fig. 3</figureCitation>
). A combination of these three external wing measurements therefore typically separates
<taxonomicName id="3FC419F4D44BFFEAFC79AE8F9D7FCB9F" authorityName=", Andersen" authorityYear="1912" box="[943,1052,1728,1751]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFC79AE8F9D7FCB9F" box="[943,1052,1728,1751]" italics="true" pageId="8" pageNumber="81">N. certans</emphasis>
</taxonomicName>
from the other sympatric
<taxonomicName id="3FC419F4D44BFFEAFAFBAE8F9CFDCB9F" authorityName="Borkhausen" authorityYear="1797" box="[1325,1438,1728,1751]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis id="CAB0BE65D44BFFEAFAFBAE8F9CFDCB9F" box="[1325,1438,1728,1751]" italics="true" pageId="8" pageNumber="81">Nyctimene</emphasis>
</taxonomicName>
of similar body-size including the externally very similar
<taxonomicName id="3FC419F4D44BFFEAFA74AE929AFBCA5A" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D44BFFEAFA74AE929AFBCA5A" italics="true" pageId="8" pageNumber="81">N. cyclotis</emphasis>
</taxonomicName>
(
<figureCitation id="60FF7EF2D44BFFEAFC71AEB59A8BCA5A" box="[935,1000,1786,1810]" captionStart="Figure 3" captionStartId="8.[835,898,932,954]" captionTargetBox="[835,1464,155,917]" captionTargetId="figure-620@8.[829,1468,151,920]" captionTargetPageId="8" captionText="Figure 3. Plot of 5-Metacarpal divided by the 1st phalanx of the 5 digit, verses 2nd phalanx of the 2nd digit, indicating separation of Nyctimene certans and N. cyclotis using 3 simple wing measurements for 255 specimens. Nyctimene certans (squares, n = 35), N. a. papuanus (circles, n = 128), N. wrightae sp. nov. (triangles, n = 90) and N. cyclotis (diamonds, n = 2). Holotype specimens are indicated for each species by an asterisk. Full or closed symbols indicate individuals identified genetically (n = 77), open symbols are those identified morphologically (n = 179)." figureDoi="http://doi.org/10.5281/zenodo.5237689" httpUri="https://zenodo.org/record/5237689/files/figure.png" pageId="8" pageNumber="81">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B0DE31FCD44BFFE9FC91AF6898F9CBE6" lastPageId="11" lastPageNumber="84" pageId="8" pageNumber="81" type="discussion">
<paragraph id="F87B6277D44BFFEAFC91AF689DBFCADF" blockId="8.[837,1468,1831,2002]" pageId="8" pageNumber="81">
<emphasis id="CAB0BE65D44BFFEAFC91AF689AD5CA77" bold="true" box="[839,950,1831,1855]" pageId="8" pageNumber="81">Remarks.</emphasis>
Smith &amp; Hood (1983) based their concept of
<taxonomicName id="3FC419F4D44BFFEAFA74AF679AFBCA14" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D44BFFEAFA74AF679AFBCA14" italics="true" pageId="8" pageNumber="81">N. cyclotis</emphasis>
</taxonomicName>
from specimen BPBM 28398 (photographed in fig. 1c of their paper) but I concur with
<bibRefCitation id="9C551F86D44BFFEAFB01AF2D9CE0CA32" author="Peterson, R" box="[1239,1411,1890,1914]" pageId="8" pageNumber="81" pagination="26 - 41" refId="ref23034" refString="Peterson, R. L. 1991. Systematic variation in the Megachiropteran Tube-nosed Bats Nyctimene cyclotis and N. certans. Bulletin of the American Museum of Natural History 206: 26 - 41." type="journal article" year="1991">Peterson (1991)</bibRefCitation>
who identified this specimen as
<taxonomicName id="3FC419F4D44BFFEAFBBEAFCF9DB5CADF" authorityName=", Andersen" authorityYear="1912" box="[1128,1238,1920,1943]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFBBEAFCF9DB5CADF" box="[1128,1238,1920,1943]" italics="true" pageId="8" pageNumber="81">N. certans</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F87B6277D44BFFE9FCB3AFD2988DC8C0" blockId="8.[837,1468,1831,2002]" lastBlockId="11.[119,750,923,1710]" lastPageId="11" lastPageNumber="84" pageId="8" pageNumber="81">
<taxonomicName id="3FC419F4D44BFFEAFCB3AFD29D46CAFC" authorityName=", Andersen" authorityYear="1912" box="[869,1061,1949,1972]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="81" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44BFFEAFCB3AFD29D46CAFC" box="[869,1061,1949,1972]" italics="true" pageId="8" pageNumber="81">Nyctimene certans</emphasis>
</taxonomicName>
is the only species within the subfamily to have round palatal fenestrations (vacuities) at the distal end of the palate. However, as noted by Smith &amp; Hood (1983), this character is variable, sometimes with two present as in BPBM 28398 (Smith &amp; Hood, 1983) and AM M.7908 (as illustrated in
<bibRefCitation id="9C551F86D448FFE9FE5BABBC9B5CC943" author="Flannery, T" box="[397,575,1011,1035]" pageId="11" pageNumber="84" refId="ref21280" refString="Flannery, T. F. 1995 b. Mammals of New Guinea. Second edition. Chatswood, New South Wales: Australian Museum / Reed Books. 568 pp." type="book" year="1995">Flannery, 1995b</bibRefCitation>
), or none as for AM M.17888 (
<figureCitation id="60FF7EF2D448FFE9FEF7AC5F9806C960" box="[289,357,1040,1064]" captionStart="Figure 4" captionStartId="9.[121,184,1872,1894]" captionTargetBox="[121,1421,151,1857]" captionTargetId="figure-9@9.[121,1421,151,1857]" captionTargetPageId="9" captionText="Figure 4. Dorsal and ventral views of skulls and mandibles of Nyctimene certans (young adult) (AM M.17888), N.certans (old, showing extensive tooth wear) (AM M.30648), N. wrightae sp. nov. (holotype) (AM M.16423) and N. a. papuanus (AM M.16425)." figureDoi="http://doi.org/10.5281/zenodo.5237691" httpUri="https://zenodo.org/record/5237691/files/figure.png" pageId="11" pageNumber="84">Fig. 4</figureCitation>
). The only other recorded case of a palatal fenestration in another species of the subfamily is an AMNH specimen of
<taxonomicName id="3FC419F4D448FFE9FE5CAC03988DC92B" baseAuthorityName="Dobson" baseAuthorityYear="1877" box="[394,494,1100,1123]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="major">
<emphasis id="CAB0BE65D448FFE9FE5CAC03988DC92B" box="[394,494,1100,1123]" italics="true" pageId="11" pageNumber="84">N. major</emphasis>
</taxonomicName>
from Kiriwina Island. Koopman (1982) discussed this specimen and concluded that it was very aged and somewhat pathological, adding that there was secondary bone deposition lateral to the molars. It is unlikely that these fenestrations have any function, as they are so variable between individuals. They could therefore represent a vestigial pattern of growth or possibly an indication of masticator stress on the palate, arising from eating hard fruit during growth as is thought to occur in other mammals (Moss &amp; Feliciano, 1977). They remain a feature generally found only in
<taxonomicName id="3FC419F4D448FFE9FEAFAD3E9884C8C0" authorityName=", Andersen" authorityYear="1912" box="[377,487,1393,1416]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D448FFE9FEAFAD3E9884C8C0" box="[377,487,1393,1416]" italics="true" pageId="11" pageNumber="84">N. certans</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="ACBB32FFD44AFFEBFFAFAF1F9C77CAC8" ID-DOI="http://doi.org/10.5281/zenodo.5237691" ID-Zenodo-Dep="5237691" httpUri="https://zenodo.org/record/5237691/files/figure.png" pageId="9" pageNumber="82" startId="9.[121,184,1872,1894]" targetBox="[121,1421,151,1857]" targetPageId="9">
<paragraph id="F87B6277D44AFFEBFFAFAF1F9C77CAC8" blockId="9.[120,1420,1872,1921]" pageId="9" pageNumber="82">
Figure 4. Dorsal and ventral views of skulls and mandibles of
<taxonomicName id="3FC419F4D44AFFEBFD1FAF1E9A19CA2E" authorityName=", Andersen" authorityYear="1912" box="[713,890,1873,1894]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="82" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44AFFEBFD1FAF1E9A19CA2E" box="[713,890,1873,1894]" italics="true" pageId="9" pageNumber="82">Nyctimene certans</emphasis>
</taxonomicName>
(young adult) (AM M.17888),
<taxonomicName id="3FC419F4D44AFFEBFB75AF1E9C66CA2E" authorityName=", Andersen" authorityYear="1912" box="[1187,1285,1873,1894]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="82" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D44AFFEBFB75AF1E9C66CA2E" box="[1187,1285,1873,1894]" italics="true" pageId="9" pageNumber="82">N. certans</emphasis>
</taxonomicName>
(old, showing extensive tooth wear) (AM M.30648),
<taxonomicName id="3FC419F4D44AFFEBFE38AF249B3CCAC8" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[494,607,1899,1920]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="82" phylum="Chordata" rank="species" species="wrightae" status="sp. nov.">
<emphasis id="CAB0BE65D44AFFEBFE38AF249B3CCAC8" box="[494,607,1899,1920]" italics="true" pageId="9" pageNumber="82">N. wrightae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D183031ED44AFFEBFDB3AF249BD3CAC8" box="[613,688,1899,1920]" pageId="9" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
(holotype) (AM M.16423) and
<emphasis id="CAB0BE65D44AFFEBFC37AF249D11CAC8" box="[993,1138,1899,1920]" italics="true" pageId="9" pageNumber="82">
N. a.
<taxonomicName id="3FC419F4D44AFFEBFBC3AF249D11CAC8" authorityName="Andersen" authorityYear="1910" box="[1045,1138,1899,1920]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="82" phylum="Chordata" rank="species" species="papuanus">papuanus</taxonomicName>
</emphasis>
(AM M.16425).
</paragraph>
</caption>
<caption id="ACBB32FFD449FFE8FF70AEB998F6CAD9" ID-DOI="http://doi.org/10.5281/zenodo.5237693" ID-Zenodo-Dep="5237693" httpUri="https://zenodo.org/record/5237693/files/figure.png" pageId="10" pageNumber="83" startId="10.[166,229,1782,1804]" targetBox="[167,1467,151,1766]" targetPageId="10">
<paragraph id="F87B6277D449FFE8FF70AEB998F6CAD9" blockId="10.[165,1467,1781,1937]" pageId="10" pageNumber="83">
Figure 5. Dorsal and ventral photographs of the holotypes of
<emphasis id="CAB0BE65D449FFE8FD22AEB99D1ACA43" box="[756,1145,1782,1803]" italics="true" pageId="10" pageNumber="83">
<taxonomicName id="3FC419F4D449FFE8FD22AEB99AC6CA43" authorityName=", Andersen" authorityYear="1912" box="[756,933,1782,1803]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="certans">Nyctimene certans</taxonomicName>
and
<taxonomicName id="3FC419F4D449FFE8FC02AEB99D5FCA43" box="[980,1084,1782,1803]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="cyclotis">N. cyclotis</taxonomicName>
. (a, b)
</emphasis>
Holotype of
<taxonomicName id="3FC419F4D449FFE8FB21AEB99CD9CA43" authority="NHMUK" authorityName="NHMUK" box="[1271,1466,1782,1803]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D449FFE8FB21AEB99C39CA43" box="[1271,1370,1782,1803]" italics="true" pageId="10" pageNumber="83">N. certans</emphasis>
NHMUK
</taxonomicName>
1911.11.29.1; photographs courtesy of Harry Taylor (NHMUK).
<emphasis id="CAB0BE65D449FFE8FCD5AF5E9A57CA6D" box="[771,820,1808,1830]" italics="true" pageId="10" pageNumber="83">(c, d)</emphasis>
Holotype of
<taxonomicName id="3FC419F4D449FFE8FC7BAF5E9D0CCA6E" authority="NHMUK" authorityName="NHMUK" box="[941,1135,1808,1830]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D449FFE8FC7BAF5E9D73CA6D" box="[941,1040,1808,1830]" italics="true" pageId="10" pageNumber="83">N. cyclotis</emphasis>
NHMUK
</taxonomicName>
1910.7.16.9; photographs courtesy of Jeff Streicher (NHMUK). The ears are short and rounded in
<taxonomicName id="3FC419F4D449FFE8FCD8AF639A11CA09" box="[782,882,1836,1857]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D449FFE8FCD8AF639A11CA09" box="[782,882,1836,1857]" italics="true" pageId="10" pageNumber="83">N. certans</emphasis>
</taxonomicName>
and
<emphasis id="CAB0BE65D449FFE8FC74AF639D73CA08" box="[930,1040,1835,1857]" italics="true" pageId="10" pageNumber="83">
<taxonomicName id="3FC419F4D449FFE8FC74AF639D68CA08" box="[930,1035,1835,1857]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="cyclotis">N. cyclotis</taxonomicName>
.
</emphasis>
Both FAs of
<taxonomicName id="3FC419F4D449FFE8FB4FAF639D9CCA08" box="[1177,1279,1835,1857]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D449FFE8FB4FAF639D9CCA08" box="[1177,1279,1835,1857]" italics="true" pageId="10" pageNumber="83">N. cyclotis</emphasis>
</taxonomicName>
are clearly broken. The dorsal stripe is difficult to distinguish in the base of the dorsal in
<taxonomicName id="3FC419F4D449FFE8FC90AF099ACACA13" box="[838,937,1862,1883]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D449FFE8FC90AF099ACACA13" box="[838,937,1862,1883]" italics="true" pageId="10" pageNumber="83">N. certans</emphasis>
</taxonomicName>
and
<taxonomicName id="3FC419F4D449FFE8FC0FAF099D5DCA13" box="[985,1086,1862,1883]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D449FFE8FC0FAF099D5DCA13" box="[985,1086,1862,1883]" italics="true" pageId="10" pageNumber="83">N. cyclotis</emphasis>
</taxonomicName>
. Overall there is a grizzled grey-brown appearance in the fur in the dorsal of
<taxonomicName id="3FC419F4D449FFE8FDC7AF2E9B17CA3E" box="[529,628,1889,1910]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D449FFE8FDC7AF2E9B17CA3E" box="[529,628,1889,1910]" italics="true" pageId="10" pageNumber="83">N. certans</emphasis>
</taxonomicName>
and
<emphasis id="CAB0BE65D449FFE8FD75AF2E9A6CCA3D" box="[675,783,1888,1910]" italics="true" pageId="10" pageNumber="83">
<taxonomicName id="3FC419F4D449FFE8FD75AF2E9A68CA3D" box="[675,779,1888,1910]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="cyclotis">N. cyclotis</taxonomicName>
,
</emphasis>
compared to a more even brown colour in
<taxonomicName id="3FC419F4D449FFE8FB67AF2E9C41CA3D" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[1201,1314,1888,1910]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="wrightae" status="sp. nov.">
<emphasis id="CAB0BE65D449FFE8FB67AF2E9C41CA3D" box="[1201,1314,1888,1910]" italics="true" pageId="10" pageNumber="83">N. wrightae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D183031ED449FFE8FAFFAF2E9C16CA3E" box="[1321,1397,1889,1910]" pageId="10" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
and
<emphasis id="CAB0BE65D449FFE8FA72AF2E987FCAD9" italics="true" pageId="10" pageNumber="83">
N. a.
<taxonomicName id="3FC419F4D449FFE8FF69AF33987FCAD9" authorityName="Andersen" authorityYear="1910" box="[191,284,1916,1937]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="83" phylum="Chordata" rank="species" species="papuanus">papuanus</taxonomicName>
</emphasis>
(see Fig. 6).
</paragraph>
</caption>
<caption id="ACBB32FFD448FFE9FFAFA8E89D0DCDF7" box="[121,1134,166,191]" pageId="11" pageNumber="84" startId="11.[121,184,167,191]">
<paragraph id="F87B6277D448FFE9FFAFA8E89D0DCDF7" blockId="11.[121,1134,166,191]" box="[121,1134,166,191]" pageId="11" pageNumber="84">
<emphasis id="CAB0BE65D448FFE9FFAFA8E899A8CDF6" bold="true" box="[121,203,166,191]" pageId="11" pageNumber="84">Table 3</emphasis>
. Welch Two Sample
<emphasis id="CAB0BE65D448FFE9FE79A8E89890CDF6" box="[431,499,167,190]" italics="true" pageId="11" pageNumber="84">ttests</emphasis>
for each pair of species, on 24 variables used in the LDFA.
</paragraph>
</caption>
<paragraph id="F87B6277D448FFE9FF29A8939C1FCE2B" pageId="11" pageNumber="84">
<table id="8AC490D7D448001DFF53A8949CEDCE2C" box="[133,1422,219,868]" gridcols="13" gridrows="26" pageId="11" pageNumber="84">
<tr id="46F46035D448001DFF53A8949CEDCDB9" box="[133,1422,219,241]" gridrow="0" pageId="11" pageNumber="84" rowspan-0="1">
<th id="05250949D448001DFF29A8949B01CDB9" box="[255,610,219,241]" colspan="4" colspanRight="3" gridcol="1" gridrow="0" pageId="11" pageNumber="84">
<emphasis id="CAB0BE65D448FFE9FF29A8939B01CDB9" box="[255,610,220,241]" italics="true" pageId="11" pageNumber="84">
<taxonomicName id="3FC419F4D448FFE9FF29A89398D1CDB9" box="[255,434,220,241]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="certans">Nyctimene certans</taxonomicName>
vs N. a.
<taxonomicName id="3FC419F4D448FFE9FDD3A8939B01CDB9" authorityName="Andersen" authorityYear="1910" box="[517,610,220,241]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="papuanus">papuanus</taxonomicName>
</emphasis>
</th>
<th id="05250949D448001DFD7FA8949A9BCDB9" box="[681,1016,219,241]" colspan="4" colspanRight="3" gridcol="5" gridrow="0" pageId="11" pageNumber="84">
<emphasis id="CAB0BE65D448FFE9FD7AA8939AC3CDB8" box="[684,928,219,241]" italics="true" pageId="11" pageNumber="84">
<taxonomicName id="3FC419F4D448FFE9FD7AA8939A6CCDB9" box="[684,783,220,241]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="certans">N. certans</taxonomicName>
vs
<taxonomicName id="3FC419F4D448FFE9FCF9A8939AC3CDB8" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[815,928,219,241]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="wrightae" status="sp. nov.">N. wrightae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="D183031ED448FFE9FC70A8939A92CDB9" box="[934,1009,220,241]" pageId="11" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="05250949D448001DFBCEA8949CEDCDB9" box="[1048,1422,219,241]" colspan="4" colspanRight="3" gridcol="9" gridrow="0" pageId="11" pageNumber="84">
<taxonomicName id="3FC419F4D448FFE9FBCEA8939DEACDB8" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[1048,1161,219,241]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="wrightae" status="sp. nov.">
<emphasis id="CAB0BE65D448FFE9FBCEA8939DEACDB8" box="[1048,1161,219,241]" italics="true" pageId="11" pageNumber="84">N. wrightae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D183031ED448FFE9FB59A8939DB9CDB9" box="[1167,1242,220,241]" pageId="11" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="CAB0BE65D448FFE9FB09A8939CE9CDB9" box="[1247,1418,220,241]" italics="true" pageId="11" pageNumber="84">
vs N. a.
<taxonomicName id="3FC419F4D448FFE9FAFBA8939CE9CDB9" authorityName="Andersen" authorityYear="1910" box="[1325,1418,220,241]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="papuanus">papuanus</taxonomicName>
</emphasis>
</th>
</tr>
<tr id="46F46035D448001DFF53A8B79CEDCC46" box="[133,1422,248,270]" gridrow="1" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A8B799B7CC46" box="[133,212,248,270]" gridcol="0" gridrow="1" pageId="11" pageNumber="84">variable</th>
<td id="05250949D448001DFF29A8B7983ACC46" box="[255,345,248,270]" gridcol="1" gridrow="1" pageId="11" pageNumber="84">t</td>
<td id="05250949D448001DFEA2A8B79B01CC46" box="[372,610,248,270]" colspan="3" colspanRight="2" gridcol="2" gridrow="1" pageId="11" pageNumber="84">df p-value σ level</td>
<td id="05250949D448001DFD7FA8B79B88CC46" box="[681,747,248,270]" gridcol="5" gridrow="1" pageId="11" pageNumber="84">t</td>
<td id="05250949D448001DFCDAA8B79A26CC46" box="[780,837,248,270]" gridcol="6" gridrow="1" pageId="11" pageNumber="84">df</td>
<td id="05250949D448001DFC83A8B79A9BCC46" box="[853,1016,248,270]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="11" pageNumber="84">p-value σ level</td>
<td id="05250949D448001DFBCEA8B79DE4CC46" box="[1048,1159,248,270]" gridcol="9" gridrow="1" pageId="11" pageNumber="84">t</td>
<td id="05250949D448001DFB72A8B79DBECC46" box="[1188,1245,248,270]" gridcol="10" gridrow="1" pageId="11" pageNumber="84">df</td>
<td id="05250949D448001DFB2EA8B79CEDCC46" box="[1272,1422,248,270]" colspan="2" colspanRight="1" gridcol="11" gridrow="1" pageId="11" pageNumber="84">p-value σ level</td>
</tr>
<tr id="46F46035D448001DFF53A9669CEDCC73" box="[133,1422,297,315]" gridrow="2" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A96699B7CC73" box="[133,212,297,315]" gridcol="0" gridrow="2" pageId="11" pageNumber="84">FA</th>
<td id="05250949D448001DFF29A966983ACC73" box="[255,345,297,315]" gridcol="1" gridrow="2" pageId="11" pageNumber="84">10.3759</td>
<td id="05250949D448001DFEA2A96698C0CC73" box="[372,419,297,315]" gridcol="2" gridrow="2" pageId="11" pageNumber="84">43.30</td>
<td id="05250949D448001DFE6FA9669B6ECC73" box="[441,525,297,315]" gridcol="3" gridrow="2" pageId="11" pageNumber="84">2.562e-13</td>
<td id="05250949D448001DFDFAA9669B01CC73" box="[556,610,297,315]" gridcol="4" gridrow="2" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA9669B88CC73" box="[681,747,297,315]" gridcol="5" gridrow="2" pageId="11" pageNumber="84">3.4479</td>
<td id="05250949D448001DFCDAA9669A26CC73" box="[780,837,297,315]" gridcol="6" gridrow="2" pageId="11" pageNumber="84">46.65</td>
<td id="05250949D448001DFC83A9669AD5CC73" box="[853,950,297,315]" gridcol="7" gridrow="2" pageId="11" pageNumber="84">0.001</td>
<td id="05250949D448001DFC18A9669A9BCC73" box="[974,1016,297,315]" gridcol="8" gridrow="2" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEA9669DE4CC73" box="[1048,1159,297,315]" gridcol="9" gridrow="2" pageId="11" pageNumber="84">13.1333</td>
<td id="05250949D448001DFB72A9669DBECC73" box="[1188,1245,297,315]" gridcol="10" gridrow="2" pageId="11" pageNumber="84">223.63</td>
<td id="05250949D448001DFB2EA9669C2DCC73" box="[1272,1358,297,315]" gridcol="11" gridrow="2" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFAB0A9669CEDCC73" box="[1382,1422,297,315]" gridcol="12" gridrow="2" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A90E9CEDCC1B" box="[133,1422,321,339]" gridrow="3" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A90E99B7CC1B" box="[133,212,321,339]" gridcol="0" gridrow="3" pageId="11" pageNumber="84">5-MET</th>
<td id="05250949D448001DFF29A90E983ACC1B" box="[255,345,321,339]" gridcol="1" gridrow="3" pageId="11" pageNumber="84">4.7578</td>
<td id="05250949D448001DFEA2A90E98C0CC1B" box="[372,419,321,339]" gridcol="2" gridrow="3" pageId="11" pageNumber="84">46.90</td>
<td id="05250949D448001DFE6FA90E9B6ECC1B" box="[441,525,321,339]" gridcol="3" gridrow="3" pageId="11" pageNumber="84">1.905e-05</td>
<td id="05250949D448001DFDFAA90E9B01CC1B" box="[556,610,321,339]" gridcol="4" gridrow="3" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA90E9B88CC1B" box="[681,747,321,339]" gridcol="5" gridrow="3" pageId="11" pageNumber="84">-1.1760</td>
<td id="05250949D448001DFCDAA90E9A26CC1B" box="[780,837,321,339]" gridcol="6" gridrow="3" pageId="11" pageNumber="84">49.75</td>
<td id="05250949D448001DFC83A90E9AD5CC1B" box="[853,950,321,339]" gridcol="7" gridrow="3" pageId="11" pageNumber="84">0.245</td>
<td id="05250949D448001DFC18A90E9A9BCC1B" box="[974,1016,321,339]" gridcol="8" gridrow="3" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFBCEA90E9DE4CC1B" box="[1048,1159,321,339]" gridcol="9" gridrow="3" pageId="11" pageNumber="84">9.8496</td>
<td id="05250949D448001DFB72A90E9DBECC1B" box="[1188,1245,321,339]" gridcol="10" gridrow="3" pageId="11" pageNumber="84">215.45</td>
<td id="05250949D448001DFB2EA90E9C2DCC1B" box="[1272,1358,321,339]" gridcol="11" gridrow="3" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFAB0A90E9CEDCC1B" box="[1382,1422,321,339]" gridcol="12" gridrow="3" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A9169CEDCC23" box="[133,1422,345,363]" gridrow="4" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A91699B7CC23" box="[133,212,345,363]" gridcol="0" gridrow="4" pageId="11" pageNumber="84">4-MET</th>
<td id="05250949D448001DFF29A916983ACC23" box="[255,345,345,363]" gridcol="1" gridrow="4" pageId="11" pageNumber="84">8.6278</td>
<td id="05250949D448001DFEA2A91698C0CC23" box="[372,419,345,363]" gridcol="2" gridrow="4" pageId="11" pageNumber="84">41.84</td>
<td id="05250949D448001DFE6FA9169B6ECC23" box="[441,525,345,363]" gridcol="3" gridrow="4" pageId="11" pageNumber="84">7.820e-11</td>
<td id="05250949D448001DFDFAA9169B01CC23" box="[556,610,345,363]" gridcol="4" gridrow="4" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA9169B88CC23" box="[681,747,345,363]" gridcol="5" gridrow="4" pageId="11" pageNumber="84">1.8121</td>
<td id="05250949D448001DFCDAA9169A26CC23" box="[780,837,345,363]" gridcol="6" gridrow="4" pageId="11" pageNumber="84">47.11</td>
<td id="05250949D448001DFC83A9169AD5CC23" box="[853,950,345,363]" gridcol="7" gridrow="4" pageId="11" pageNumber="84">0.076</td>
<td id="05250949D448001DFC18A9169A9BCC23" box="[974,1016,345,363]" gridcol="8" gridrow="4" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFBCEA9169DE4CC23" box="[1048,1159,345,363]" gridcol="9" gridrow="4" pageId="11" pageNumber="84">11.3896</td>
<td id="05250949D448001DFB72A9169DBECC23" box="[1188,1245,345,363]" gridcol="10" gridrow="4" pageId="11" pageNumber="84">203.93</td>
<td id="05250949D448001DFB2EA9169C2DCC23" box="[1272,1358,345,363]" gridcol="11" gridrow="4" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFAB0A9169CEDCC23" box="[1382,1422,345,363]" gridcol="12" gridrow="4" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A93E9CEDCCCB" box="[133,1422,369,387]" gridrow="5" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A93E99B7CCCB" box="[133,212,369,387]" gridcol="0" gridrow="5" pageId="11" pageNumber="84">3-MET</th>
<td id="05250949D448001DFF29A93E983ACCCB" box="[255,345,369,387]" gridcol="1" gridrow="5" pageId="11" pageNumber="84">7.8199</td>
<td id="05250949D448001DFEA2A93E98C0CCCB" box="[372,419,369,387]" gridcol="2" gridrow="5" pageId="11" pageNumber="84">44.85</td>
<td id="05250949D448001DFE6FA93E9B6ECCCB" box="[441,525,369,387]" gridcol="3" gridrow="5" pageId="11" pageNumber="84">6.456e-10</td>
<td id="05250949D448001DFDFAA93E9B01CCCB" box="[556,610,369,387]" gridcol="4" gridrow="5" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA93E9B88CCCB" box="[681,747,369,387]" gridcol="5" gridrow="5" pageId="11" pageNumber="84">2.0823</td>
<td id="05250949D448001DFCDAA93E9A26CCCB" box="[780,837,369,387]" gridcol="6" gridrow="5" pageId="11" pageNumber="84">51.22</td>
<td id="05250949D448001DFC83A93E9AD5CCCB" box="[853,950,369,387]" gridcol="7" gridrow="5" pageId="11" pageNumber="84">0.042</td>
<td id="05250949D448001DFC18A93E9A9BCCCB" box="[974,1016,369,387]" gridcol="8" gridrow="5" pageId="11" pageNumber="84">*</td>
<td id="05250949D448001DFBCEA93E9DE4CCCB" box="[1048,1159,369,387]" gridcol="9" gridrow="5" pageId="11" pageNumber="84">9.3295</td>
<td id="05250949D448001DFB72A93E9DBECCCB" box="[1188,1245,369,387]" gridcol="10" gridrow="5" pageId="11" pageNumber="84">199.57</td>
<td id="05250949D448001DFB2EA93E9C2DCCCB" box="[1272,1358,369,387]" gridcol="11" gridrow="5" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFAB0A93E9CEDCCCB" box="[1382,1422,369,387]" gridcol="12" gridrow="5" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A9C69CEDCCD3" box="[133,1422,393,411]" gridrow="6" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A9C699B7CCD3" box="[133,212,393,411]" gridcol="0" gridrow="6" pageId="11" pageNumber="84">2-MET</th>
<td id="05250949D448001DFF29A9C6983ACCD3" box="[255,345,393,411]" gridcol="1" gridrow="6" pageId="11" pageNumber="84">8.1459</td>
<td id="05250949D448001DFEA2A9C698C0CCD3" box="[372,419,393,411]" gridcol="2" gridrow="6" pageId="11" pageNumber="84">47.23</td>
<td id="05250949D448001DFE6FA9C69B6ECCD3" box="[441,525,393,411]" gridcol="3" gridrow="6" pageId="11" pageNumber="84">1.492e-10</td>
<td id="05250949D448001DFDFAA9C69B01CCD3" box="[556,610,393,411]" gridcol="4" gridrow="6" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA9C69B88CCD3" box="[681,747,393,411]" gridcol="5" gridrow="6" pageId="11" pageNumber="84">0.2553</td>
<td id="05250949D448001DFCDAA9C69A26CCD3" box="[780,837,393,411]" gridcol="6" gridrow="6" pageId="11" pageNumber="84">69.19</td>
<td id="05250949D448001DFC83A9C69AD5CCD3" box="[853,950,393,411]" gridcol="7" gridrow="6" pageId="11" pageNumber="84">0.799</td>
<td id="05250949D448001DFC18A9C69A9BCCD3" box="[974,1016,393,411]" gridcol="8" gridrow="6" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFBCEA9C69DE4CCD3" box="[1048,1159,393,411]" gridcol="9" gridrow="6" pageId="11" pageNumber="84">10.1930</td>
<td id="05250949D448001DFB72A9C69DBECCD3" box="[1188,1245,393,411]" gridcol="10" gridrow="6" pageId="11" pageNumber="84">166.62</td>
<td id="05250949D448001DFB2EA9C69C2DCCD3" box="[1272,1358,393,411]" gridcol="11" gridrow="6" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFAB0A9C69CEDCCD3" box="[1382,1422,393,411]" gridcol="12" gridrow="6" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A9EE9CEDCCFB" box="[133,1422,417,435]" gridrow="7" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A9EE99B7CCFB" box="[133,212,417,435]" gridcol="0" gridrow="7" pageId="11" pageNumber="84">1-MET</th>
<td id="05250949D448001DFF29A9EE983ACCFB" box="[255,345,417,435]" gridcol="1" gridrow="7" pageId="11" pageNumber="84">5.5086</td>
<td id="05250949D448001DFEA2A9EE98C0CCFB" box="[372,419,417,435]" gridcol="2" gridrow="7" pageId="11" pageNumber="84">6.08</td>
<td id="05250949D448001DFE6FA9EE9B6ECCFB" box="[441,525,417,435]" gridcol="3" gridrow="7" pageId="11" pageNumber="84">9.399e-07</td>
<td id="05250949D448001DFDFAA9EE9B01CCFB" box="[556,610,417,435]" gridcol="4" gridrow="7" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA9EE9B88CCFB" box="[681,747,417,435]" gridcol="5" gridrow="7" pageId="11" pageNumber="84">2.7283</td>
<td id="05250949D448001DFCDAA9EE9A26CCFB" box="[780,837,417,435]" gridcol="6" gridrow="7" pageId="11" pageNumber="84">71.44</td>
<td id="05250949D448001DFC83A9EE9AD5CCFB" box="[853,950,417,435]" gridcol="7" gridrow="7" pageId="11" pageNumber="84">0.008</td>
<td id="05250949D448001DFC18A9EE9A9BCCFB" box="[974,1016,417,435]" gridcol="8" gridrow="7" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEA9EE9DE4CCFB" box="[1048,1159,417,435]" gridcol="9" gridrow="7" pageId="11" pageNumber="84">3.1572</td>
<td id="05250949D448001DFB72A9EE9DBECCFB" box="[1188,1245,417,435]" gridcol="10" gridrow="7" pageId="11" pageNumber="84">192.50</td>
<td id="05250949D448001DFB2EA9EE9C2DCCFB" box="[1272,1358,417,435]" gridcol="11" gridrow="7" pageId="11" pageNumber="84">1.849e-03</td>
<td id="05250949D448001DFAB0A9EE9CEDCCFB" box="[1382,1422,417,435]" gridcol="12" gridrow="7" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A9F69CEDCC83" box="[133,1422,441,459]" gridrow="8" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A9F699B7CC83" box="[133,212,441,459]" gridcol="0" gridrow="8" pageId="11" pageNumber="84">2M-2P</th>
<td id="05250949D448001DFF29A9F6983ACC83" box="[255,345,441,459]" gridcol="1" gridrow="8" pageId="11" pageNumber="84">12.0540</td>
<td id="05250949D448001DFEA2A9F698C0CC83" box="[372,419,441,459]" gridcol="2" gridrow="8" pageId="11" pageNumber="84">74.41</td>
<td id="05250949D448001DFE6FA9F69B6ECC83" box="[441,525,441,459]" gridcol="3" gridrow="8" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFDFAA9F69B01CC83" box="[556,610,441,459]" gridcol="4" gridrow="8" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA9F69B88CC83" box="[681,747,441,459]" gridcol="5" gridrow="8" pageId="11" pageNumber="84">11.5190</td>
<td id="05250949D448001DFCDAA9F69A26CC83" box="[780,837,441,459]" gridcol="6" gridrow="8" pageId="11" pageNumber="84">93.95</td>
<td id="05250949D448001DFC83A9F69AD5CC83" box="[853,950,441,459]" gridcol="7" gridrow="8" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFC18A9F69A9BCC83" box="[974,1016,441,459]" gridcol="8" gridrow="8" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEA9F69DE4CC83" box="[1048,1159,441,459]" gridcol="9" gridrow="8" pageId="11" pageNumber="84">-0.8151</td>
<td id="05250949D448001DFB72A9F69DBECC83" box="[1188,1245,441,459]" gridcol="10" gridrow="8" pageId="11" pageNumber="84">183.81</td>
<td id="05250949D448001DFB2EA9F69C2DCC83" box="[1272,1358,441,459]" gridcol="11" gridrow="8" pageId="11" pageNumber="84">0.416</td>
<td id="05250949D448001DFAB0A9F69CEDCC83" box="[1382,1422,441,459]" gridcol="12" gridrow="8" pageId="11" pageNumber="84">ns</td>
</tr>
<tr id="46F46035D448001DFF53A99E9CEDCCAB" box="[133,1422,465,483]" gridrow="9" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A99E99B7CCAB" box="[133,212,465,483]" gridcol="0" gridrow="9" pageId="11" pageNumber="84">5M-1P</th>
<td id="05250949D448001DFF29A99E983ACCAB" box="[255,345,465,483]" gridcol="1" gridrow="9" pageId="11" pageNumber="84">16.9670</td>
<td id="05250949D448001DFEA2A99E98C0CCAB" box="[372,419,465,483]" gridcol="2" gridrow="9" pageId="11" pageNumber="84">42.29</td>
<td id="05250949D448001DFE6FA99E9B6ECCAB" box="[441,525,465,483]" gridcol="3" gridrow="9" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFDFAA99E9B01CCAB" box="[556,610,465,483]" gridcol="4" gridrow="9" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA99E9B88CCAB" box="[681,747,465,483]" gridcol="5" gridrow="9" pageId="11" pageNumber="84">9.8638</td>
<td id="05250949D448001DFCDAA99E9A26CCAB" box="[780,837,465,483]" gridcol="6" gridrow="9" pageId="11" pageNumber="84">49.32</td>
<td id="05250949D448001DFC83A99E9AD5CCAB" box="[853,950,465,483]" gridcol="7" gridrow="9" pageId="11" pageNumber="84">2.949e-13</td>
<td id="05250949D448001DFC18A99E9A9BCCAB" box="[974,1016,465,483]" gridcol="8" gridrow="9" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEA99E9DE4CCAB" box="[1048,1159,465,483]" gridcol="9" gridrow="9" pageId="11" pageNumber="84">11.7104</td>
<td id="05250949D448001DFB72A99E9DBECCAB" box="[1188,1245,465,483]" gridcol="10" gridrow="9" pageId="11" pageNumber="84">187.49</td>
<td id="05250949D448001DFB2EA99E9C2DCCAB" box="[1272,1358,465,483]" gridcol="11" gridrow="9" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFAB0A99E9CEDCCAB" box="[1382,1422,465,483]" gridcol="12" gridrow="9" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53A9A69CEDCCB3" box="[133,1422,489,507]" gridrow="10" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53A9A699B7CCB3" box="[133,212,489,507]" gridcol="0" gridrow="10" pageId="11" pageNumber="84">Tibia</th>
<td id="05250949D448001DFF29A9A6983ACCB3" box="[255,345,489,507]" gridcol="1" gridrow="10" pageId="11" pageNumber="84">6.1387</td>
<td id="05250949D448001DFEA2A9A698C0CCB3" box="[372,419,489,507]" gridcol="2" gridrow="10" pageId="11" pageNumber="84">39.38</td>
<td id="05250949D448001DFE6FA9A69B6ECCB3" box="[441,525,489,507]" gridcol="3" gridrow="10" pageId="11" pageNumber="84">3.201e-07</td>
<td id="05250949D448001DFDFAA9A69B01CCB3" box="[556,610,489,507]" gridcol="4" gridrow="10" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FA9A69B88CCB3" box="[681,747,489,507]" gridcol="5" gridrow="10" pageId="11" pageNumber="84">2.2659</td>
<td id="05250949D448001DFCDAA9A69A26CCB3" box="[780,837,489,507]" gridcol="6" gridrow="10" pageId="11" pageNumber="84">52.72</td>
<td id="05250949D448001DFC83A9A69AD5CCB3" box="[853,950,489,507]" gridcol="7" gridrow="10" pageId="11" pageNumber="84">0.028</td>
<td id="05250949D448001DFC18A9A69A9BCCB3" box="[974,1016,489,507]" gridcol="8" gridrow="10" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEA9A69DE4CCB3" box="[1048,1159,489,507]" gridcol="9" gridrow="10" pageId="11" pageNumber="84">6.0383</td>
<td id="05250949D448001DFB72A9A69DBECCB3" box="[1188,1245,489,507]" gridcol="10" gridrow="10" pageId="11" pageNumber="84">165.00</td>
<td id="05250949D448001DFB2EA9A69C2DCCB3" box="[1272,1358,489,507]" gridcol="11" gridrow="10" pageId="11" pageNumber="84">9.983e-09</td>
<td id="05250949D448001DFAB0A9A69CEDCCB3" box="[1382,1422,489,507]" gridcol="12" gridrow="10" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AA4E9CEDCF5B" box="[133,1422,513,531]" gridrow="11" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA4E99B7CF5B" box="[133,212,513,531]" gridcol="0" gridrow="11" pageId="11" pageNumber="84">Ear</th>
<td id="05250949D448001DFF29AA4E983ACF5B" box="[255,345,513,531]" gridcol="1" gridrow="11" pageId="11" pageNumber="84">-1.3502</td>
<td id="05250949D448001DFEA2AA4E98C0CF5B" box="[372,419,513,531]" gridcol="2" gridrow="11" pageId="11" pageNumber="84">36.74</td>
<td id="05250949D448001DFE6FAA4E9B6ECF5B" box="[441,525,513,531]" gridcol="3" gridrow="11" pageId="11" pageNumber="84">0.185</td>
<td id="05250949D448001DFDFAAA4E9B01CF5B" box="[556,610,513,531]" gridcol="4" gridrow="11" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFD7FAA4E9B88CF5B" box="[681,747,513,531]" gridcol="5" gridrow="11" pageId="11" pageNumber="84">-3.6779</td>
<td id="05250949D448001DFCDAAA4E9A26CF5B" box="[780,837,513,531]" gridcol="6" gridrow="11" pageId="11" pageNumber="84">37.33</td>
<td id="05250949D448001DFC83AA4E9AD5CF5B" box="[853,950,513,531]" gridcol="7" gridrow="11" pageId="11" pageNumber="84">0.001</td>
<td id="05250949D448001DFC18AA4E9A9BCF5B" box="[974,1016,513,531]" gridcol="8" gridrow="11" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA4E9DE4CF5B" box="[1048,1159,513,531]" gridcol="9" gridrow="11" pageId="11" pageNumber="84">4.8329</td>
<td id="05250949D448001DFB72AA4E9DBECF5B" box="[1188,1245,513,531]" gridcol="10" gridrow="11" pageId="11" pageNumber="84">223.55</td>
<td id="05250949D448001DFB2EAA4E9C2DCF5B" box="[1272,1358,513,531]" gridcol="11" gridrow="11" pageId="11" pageNumber="84">2.500e-06</td>
<td id="05250949D448001DFAB0AA4E9CEDCF5B" box="[1382,1422,513,531]" gridcol="12" gridrow="11" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AA579CEDCF63" box="[133,1422,536,555]" gridrow="12" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA5799B7CF63" box="[133,212,536,555]" gridcol="0" gridrow="12" pageId="11" pageNumber="84">GSL</th>
<td id="05250949D448001DFF29AA57983ACF63" box="[255,345,536,555]" gridcol="1" gridrow="12" pageId="11" pageNumber="84">17.0682</td>
<td id="05250949D448001DFEA2AA5798C0CF63" box="[372,419,536,555]" gridcol="2" gridrow="12" pageId="11" pageNumber="84">38.52</td>
<td id="05250949D448001DFE6FAA579B6ECF63" box="[441,525,536,555]" gridcol="3" gridrow="12" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFDFAAA579B01CF63" box="[556,610,536,555]" gridcol="4" gridrow="12" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAA579B88CF63" box="[681,747,536,555]" gridcol="5" gridrow="12" pageId="11" pageNumber="84">9.1484</td>
<td id="05250949D448001DFCDAAA579A26CF63" box="[780,837,536,555]" gridcol="6" gridrow="12" pageId="11" pageNumber="84">27.34</td>
<td id="05250949D448001DFC83AA579AD5CF63" box="[853,950,536,555]" gridcol="7" gridrow="12" pageId="11" pageNumber="84">8.251e-10</td>
<td id="05250949D448001DFC18AA579A9BCF63" box="[974,1016,536,555]" gridcol="8" gridrow="12" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA579DE4CF63" box="[1048,1159,536,555]" gridcol="9" gridrow="12" pageId="11" pageNumber="84">4.7336</td>
<td id="05250949D448001DFB72AA579DBECF63" box="[1188,1245,536,555]" gridcol="10" gridrow="12" pageId="11" pageNumber="84">17.21</td>
<td id="05250949D448001DFB2EAA579C2DCF63" box="[1272,1358,536,555]" gridcol="11" gridrow="12" pageId="11" pageNumber="84">1.859e-04</td>
<td id="05250949D448001DFAB0AA579CEDCF63" box="[1382,1422,536,555]" gridcol="12" gridrow="12" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AA7F9CEDCF0B" box="[133,1422,560,579]" gridrow="13" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA7F99B7CF0B" box="[133,212,560,579]" gridcol="0" gridrow="13" pageId="11" pageNumber="84">CL</th>
<td id="05250949D448001DFF29AA7F983ACF0B" box="[255,345,560,579]" gridcol="1" gridrow="13" pageId="11" pageNumber="84">12.7717</td>
<td id="05250949D448001DFEA2AA7F98C0CF0B" box="[372,419,560,579]" gridcol="2" gridrow="13" pageId="11" pageNumber="84">48.41</td>
<td id="05250949D448001DFE6FAA7F9B6ECF0B" box="[441,525,560,579]" gridcol="3" gridrow="13" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFDFAAA7F9B01CF0B" box="[556,610,560,579]" gridcol="4" gridrow="13" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAA7F9B88CF0B" box="[681,747,560,579]" gridcol="5" gridrow="13" pageId="11" pageNumber="84">2.7974</td>
<td id="05250949D448001DFCDAAA7F9A26CF0B" box="[780,837,560,579]" gridcol="6" gridrow="13" pageId="11" pageNumber="84">10.54</td>
<td id="05250949D448001DFC83AA7F9AD5CF0B" box="[853,950,560,579]" gridcol="7" gridrow="13" pageId="11" pageNumber="84">0.018</td>
<td id="05250949D448001DFC18AA7F9A9BCF0B" box="[974,1016,560,579]" gridcol="8" gridrow="13" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA7F9DE4CF0B" box="[1048,1159,560,579]" gridcol="9" gridrow="13" pageId="11" pageNumber="84">3.1863</td>
<td id="05250949D448001DFB72AA7F9DBECF0B" box="[1188,1245,560,579]" gridcol="10" gridrow="13" pageId="11" pageNumber="84">9.55</td>
<td id="05250949D448001DFB2EAA7F9C2DCF0B" box="[1272,1358,560,579]" gridcol="11" gridrow="13" pageId="11" pageNumber="84">0.010</td>
<td id="05250949D448001DFAB0AA7F9CEDCF0B" box="[1382,1422,560,579]" gridcol="12" gridrow="13" pageId="11" pageNumber="84">ns</td>
</tr>
<tr id="46F46035D448001DFF53AA069CEDCF13" box="[133,1422,585,603]" gridrow="14" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA0699B7CF13" box="[133,212,585,603]" gridcol="0" gridrow="14" pageId="11" pageNumber="84">R-B</th>
<td id="05250949D448001DFF29AA06983ACF13" box="[255,345,585,603]" gridcol="1" gridrow="14" pageId="11" pageNumber="84">-0.0698</td>
<td id="05250949D448001DFEA2AA0698C0CF13" box="[372,419,585,603]" gridcol="2" gridrow="14" pageId="11" pageNumber="84">67.99</td>
<td id="05250949D448001DFE6FAA069B6ECF13" box="[441,525,585,603]" gridcol="3" gridrow="14" pageId="11" pageNumber="84">0.945</td>
<td id="05250949D448001DFDFAAA069B01CF13" box="[556,610,585,603]" gridcol="4" gridrow="14" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFD7FAA069B88CF13" box="[681,747,585,603]" gridcol="5" gridrow="14" pageId="11" pageNumber="84">-2.9867</td>
<td id="05250949D448001DFCDAAA069A26CF13" box="[780,837,585,603]" gridcol="6" gridrow="14" pageId="11" pageNumber="84">17.13</td>
<td id="05250949D448001DFC83AA069AD5CF13" box="[853,950,585,603]" gridcol="7" gridrow="14" pageId="11" pageNumber="84">0.008</td>
<td id="05250949D448001DFC18AA069A9BCF13" box="[974,1016,585,603]" gridcol="8" gridrow="14" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA069DE4CF13" box="[1048,1159,585,603]" gridcol="9" gridrow="14" pageId="11" pageNumber="84">2.9625</td>
<td id="05250949D448001DFB72AA069DBECF13" box="[1188,1245,585,603]" gridcol="10" gridrow="14" pageId="11" pageNumber="84">16.98</td>
<td id="05250949D448001DFB2EAA069C2DCF13" box="[1272,1358,585,603]" gridcol="11" gridrow="14" pageId="11" pageNumber="84">0.009</td>
<td id="05250949D448001DFAB0AA069CEDCF13" box="[1382,1422,585,603]" gridcol="12" gridrow="14" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AA2E9CEDCF3B" box="[133,1422,609,627]" gridrow="15" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA2E99B7CF3B" box="[133,212,609,627]" gridcol="0" gridrow="15" pageId="11" pageNumber="84">R-L</th>
<td id="05250949D448001DFF29AA2E983ACF3B" box="[255,345,609,627]" gridcol="1" gridrow="15" pageId="11" pageNumber="84">7.1231</td>
<td id="05250949D448001DFEA2AA2E98C0CF3B" box="[372,419,609,627]" gridcol="2" gridrow="15" pageId="11" pageNumber="84">49.18</td>
<td id="05250949D448001DFE6FAA2E9B6ECF3B" box="[441,525,609,627]" gridcol="3" gridrow="15" pageId="11" pageNumber="84">4.195e-09</td>
<td id="05250949D448001DFDFAAA2E9B01CF3B" box="[556,610,609,627]" gridcol="4" gridrow="15" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAA2E9B88CF3B" box="[681,747,609,627]" gridcol="5" gridrow="15" pageId="11" pageNumber="84">5.0585</td>
<td id="05250949D448001DFCDAAA2E9A26CF3B" box="[780,837,609,627]" gridcol="6" gridrow="15" pageId="11" pageNumber="84">14.99</td>
<td id="05250949D448001DFC83AA2E9AD5CF3B" box="[853,950,609,627]" gridcol="7" gridrow="15" pageId="11" pageNumber="84">1.418e-04</td>
<td id="05250949D448001DFC18AA2E9A9BCF3B" box="[974,1016,609,627]" gridcol="8" gridrow="15" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA2E9DE4CF3B" box="[1048,1159,609,627]" gridcol="9" gridrow="15" pageId="11" pageNumber="84">-0.3671</td>
<td id="05250949D448001DFB72AA2E9DBECF3B" box="[1188,1245,609,627]" gridcol="10" gridrow="15" pageId="11" pageNumber="84">11.65</td>
<td id="05250949D448001DFB2EAA2E9C2DCF3B" box="[1272,1358,609,627]" gridcol="11" gridrow="15" pageId="11" pageNumber="84">0.720</td>
<td id="05250949D448001DFAB0AA2E9CEDCF3B" box="[1382,1422,609,627]" gridcol="12" gridrow="15" pageId="11" pageNumber="84">ns</td>
</tr>
<tr id="46F46035D448001DFF53AA379CEDCFC3" box="[133,1422,632,651]" gridrow="16" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA3799B7CFC3" box="[133,212,632,651]" gridcol="0" gridrow="16" pageId="11" pageNumber="84">P3-B</th>
<td id="05250949D448001DFF29AA37983ACFC3" box="[255,345,632,651]" gridcol="1" gridrow="16" pageId="11" pageNumber="84">13.8054</td>
<td id="05250949D448001DFEA2AA3798C0CFC3" box="[372,419,632,651]" gridcol="2" gridrow="16" pageId="11" pageNumber="84">31.41</td>
<td id="05250949D448001DFE6FAA379B6ECFC3" box="[441,525,632,651]" gridcol="3" gridrow="16" pageId="11" pageNumber="84">6.985e-15</td>
<td id="05250949D448001DFDFAAA379B01CFC3" box="[556,610,632,651]" gridcol="4" gridrow="16" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAA379B88CFC3" box="[681,747,632,651]" gridcol="5" gridrow="16" pageId="11" pageNumber="84">5.4119</td>
<td id="05250949D448001DFCDAAA379A26CFC3" box="[780,837,632,651]" gridcol="6" gridrow="16" pageId="11" pageNumber="84">35.24</td>
<td id="05250949D448001DFC83AA379AD5CFC3" box="[853,950,632,651]" gridcol="7" gridrow="16" pageId="11" pageNumber="84">4.515e-06</td>
<td id="05250949D448001DFC18AA379A9BCFC3" box="[974,1016,632,651]" gridcol="8" gridrow="16" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA379DE4CFC3" box="[1048,1159,632,651]" gridcol="9" gridrow="16" pageId="11" pageNumber="84">12.7710</td>
<td id="05250949D448001DFB72AA379DBECFC3" box="[1188,1245,632,651]" gridcol="10" gridrow="16" pageId="11" pageNumber="84">20.90</td>
<td id="05250949D448001DFB2EAA379C2DCFC3" box="[1272,1358,632,651]" gridcol="11" gridrow="16" pageId="11" pageNumber="84">2.456e-11</td>
<td id="05250949D448001DFAB0AA379CEDCFC3" box="[1382,1422,632,651]" gridcol="12" gridrow="16" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AADF9CEDCFEB" box="[133,1422,656,675]" gridrow="17" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AADF99B7CFEB" box="[133,212,656,675]" gridcol="0" gridrow="17" pageId="11" pageNumber="84">P4-B</th>
<td id="05250949D448001DFF29AADF983ACFEB" box="[255,345,656,675]" gridcol="1" gridrow="17" pageId="11" pageNumber="84">13.9676</td>
<td id="05250949D448001DFEA2AADF98C0CFEB" box="[372,419,656,675]" gridcol="2" gridrow="17" pageId="11" pageNumber="84">30.78</td>
<td id="05250949D448001DFE6FAADF9B6ECFEB" box="[441,525,656,675]" gridcol="3" gridrow="17" pageId="11" pageNumber="84">7.281e-15</td>
<td id="05250949D448001DFDFAAADF9B01CFEB" box="[556,610,656,675]" gridcol="4" gridrow="17" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAADF9B88CFEB" box="[681,747,656,675]" gridcol="5" gridrow="17" pageId="11" pageNumber="84">5.0534</td>
<td id="05250949D448001DFCDAAADF9A26CFEB" box="[780,837,656,675]" gridcol="6" gridrow="17" pageId="11" pageNumber="84">35.85</td>
<td id="05250949D448001DFC83AADF9AD5CFEB" box="[853,950,656,675]" gridcol="7" gridrow="17" pageId="11" pageNumber="84">1.289e-05</td>
<td id="05250949D448001DFC18AADF9A9BCFEB" box="[974,1016,656,675]" gridcol="8" gridrow="17" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAADF9DE4CFEB" box="[1048,1159,656,675]" gridcol="9" gridrow="17" pageId="11" pageNumber="84">10.8670</td>
<td id="05250949D448001DFB72AADF9DBECFEB" box="[1188,1245,656,675]" gridcol="10" gridrow="17" pageId="11" pageNumber="84">16.31</td>
<td id="05250949D448001DFB2EAADF9C2DCFEB" box="[1272,1358,656,675]" gridcol="11" gridrow="17" pageId="11" pageNumber="84">6.993e-09</td>
<td id="05250949D448001DFAB0AADF9CEDCFEB" box="[1382,1422,656,675]" gridcol="12" gridrow="17" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AAE69CEDCFF4" box="[133,1422,681,700]" gridrow="18" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AAE699B7CFF4" box="[133,212,681,700]" gridcol="0" gridrow="18" pageId="11" pageNumber="84">M1-B</th>
<td id="05250949D448001DFF29AAE6983ACFF4" box="[255,345,681,700]" gridcol="1" gridrow="18" pageId="11" pageNumber="84">6.1484</td>
<td id="05250949D448001DFEA2AAE698C0CFF4" box="[372,419,681,700]" gridcol="2" gridrow="18" pageId="11" pageNumber="84">28.38</td>
<td id="05250949D448001DFE6FAAE69B6ECFF4" box="[441,525,681,700]" gridcol="3" gridrow="18" pageId="11" pageNumber="84">1.161e-06</td>
<td id="05250949D448001DFDFAAAE69B01CFF4" box="[556,610,681,700]" gridcol="4" gridrow="18" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAAE69B88CFF4" box="[681,747,681,700]" gridcol="5" gridrow="18" pageId="11" pageNumber="84">2.3401</td>
<td id="05250949D448001DFCDAAAE69A26CFF4" box="[780,837,681,700]" gridcol="6" gridrow="18" pageId="11" pageNumber="84">30.91</td>
<td id="05250949D448001DFC83AAE69AD5CFF4" box="[853,950,681,700]" gridcol="7" gridrow="18" pageId="11" pageNumber="84">0.026</td>
<td id="05250949D448001DFC18AAE69A9BCFF4" box="[974,1016,681,700]" gridcol="8" gridrow="18" pageId="11" pageNumber="84">*</td>
<td id="05250949D448001DFBCEAAE69DE4CFF4" box="[1048,1159,681,700]" gridcol="9" gridrow="18" pageId="11" pageNumber="84">8.6007</td>
<td id="05250949D448001DFB72AAE69DBECFF4" box="[1188,1245,681,700]" gridcol="10" gridrow="18" pageId="11" pageNumber="84">23.46</td>
<td id="05250949D448001DFB2EAAE69C2DCFF4" box="[1272,1358,681,700]" gridcol="11" gridrow="18" pageId="11" pageNumber="84">1.033e-08</td>
<td id="05250949D448001DFAB0AAE69CEDCFF4" box="[1382,1422,681,700]" gridcol="12" gridrow="18" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AA8E9CEDCF9C" box="[133,1422,705,724]" gridrow="19" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA8E99B7CF9C" box="[133,212,705,724]" gridcol="0" gridrow="19" pageId="11" pageNumber="84">M2-B</th>
<td id="05250949D448001DFF29AA8E983ACF9C" box="[255,345,705,724]" gridcol="1" gridrow="19" pageId="11" pageNumber="84">1.7237</td>
<td id="05250949D448001DFEA2AA8E98C0CF9C" box="[372,419,705,724]" gridcol="2" gridrow="19" pageId="11" pageNumber="84">26.12</td>
<td id="05250949D448001DFE6FAA8E9B6ECF9C" box="[441,525,705,724]" gridcol="3" gridrow="19" pageId="11" pageNumber="84">0.097</td>
<td id="05250949D448001DFDFAAA8E9B01CF9C" box="[556,610,705,724]" gridcol="4" gridrow="19" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFD7FAA8E9B88CF9C" box="[681,747,705,724]" gridcol="5" gridrow="19" pageId="11" pageNumber="84">-0.9012</td>
<td id="05250949D448001DFCDAAA8E9A26CF9C" box="[780,837,705,724]" gridcol="6" gridrow="19" pageId="11" pageNumber="84">32.59</td>
<td id="05250949D448001DFC83AA8E9AD5CF9C" box="[853,950,705,724]" gridcol="7" gridrow="19" pageId="11" pageNumber="84">0.374</td>
<td id="05250949D448001DFC18AA8E9A9BCF9C" box="[974,1016,705,724]" gridcol="8" gridrow="19" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFBCEAA8E9DE4CF9C" box="[1048,1159,705,724]" gridcol="9" gridrow="19" pageId="11" pageNumber="84">4.1265</td>
<td id="05250949D448001DFB72AA8E9DBECF9C" box="[1188,1245,705,724]" gridcol="10" gridrow="19" pageId="11" pageNumber="84">16.58</td>
<td id="05250949D448001DFB2EAA8E9C2DCF9C" box="[1272,1358,705,724]" gridcol="11" gridrow="19" pageId="11" pageNumber="84">0.001</td>
<td id="05250949D448001DFAB0AA8E9CEDCF9C" box="[1382,1422,705,724]" gridcol="12" gridrow="19" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AA969CEDCFA3" box="[133,1422,729,747]" gridrow="20" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AA9699B7CFA3" box="[133,212,729,747]" gridcol="0" gridrow="20" pageId="11" pageNumber="84">Brain-W</th>
<td id="05250949D448001DFF29AA96983ACFA3" box="[255,345,729,747]" gridcol="1" gridrow="20" pageId="11" pageNumber="84">9.7852</td>
<td id="05250949D448001DFEA2AA9698C0CFA3" box="[372,419,729,747]" gridcol="2" gridrow="20" pageId="11" pageNumber="84">43.31</td>
<td id="05250949D448001DFE6FAA969B6ECFA3" box="[441,525,729,747]" gridcol="3" gridrow="20" pageId="11" pageNumber="84">1.538e-12</td>
<td id="05250949D448001DFDFAAA969B01CFA3" box="[556,610,729,747]" gridcol="4" gridrow="20" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAA969B88CFA3" box="[681,747,729,747]" gridcol="5" gridrow="20" pageId="11" pageNumber="84">4.2857</td>
<td id="05250949D448001DFCDAAA969A26CFA3" box="[780,837,729,747]" gridcol="6" gridrow="20" pageId="11" pageNumber="84">18.82</td>
<td id="05250949D448001DFC83AA969AD5CFA3" box="[853,950,729,747]" gridcol="7" gridrow="20" pageId="11" pageNumber="84">0.000</td>
<td id="05250949D448001DFC18AA969A9BCFA3" box="[974,1016,729,747]" gridcol="8" gridrow="20" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAA969DE4CFA3" box="[1048,1159,729,747]" gridcol="9" gridrow="20" pageId="11" pageNumber="84">4.1300</td>
<td id="05250949D448001DFB72AA969DBECFA3" box="[1188,1245,729,747]" gridcol="10" gridrow="20" pageId="11" pageNumber="84">28.07</td>
<td id="05250949D448001DFB2EAA969C2DCFA3" box="[1272,1358,729,747]" gridcol="11" gridrow="20" pageId="11" pageNumber="84">0.000</td>
<td id="05250949D448001DFAB0AA969CEDCFA3" box="[1382,1422,729,747]" gridcol="12" gridrow="20" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AABE9CEDCE4B" box="[133,1422,753,771]" gridrow="21" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AABE99B7CE4B" box="[133,212,753,771]" gridcol="0" gridrow="21" pageId="11" pageNumber="84">Brain-H</th>
<td id="05250949D448001DFF29AABE983ACE4B" box="[255,345,753,771]" gridcol="1" gridrow="21" pageId="11" pageNumber="84">4.2623</td>
<td id="05250949D448001DFEA2AABE98C0CE4B" box="[372,419,753,771]" gridcol="2" gridrow="21" pageId="11" pageNumber="84">40.96</td>
<td id="05250949D448001DFE6FAABE9B6ECE4B" box="[441,525,753,771]" gridcol="3" gridrow="21" pageId="11" pageNumber="84">1.158e-04</td>
<td id="05250949D448001DFDFAAABE9B01CE4B" box="[556,610,753,771]" gridcol="4" gridrow="21" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAABE9B88CE4B" box="[681,747,753,771]" gridcol="5" gridrow="21" pageId="11" pageNumber="84">-0.8322</td>
<td id="05250949D448001DFCDAAABE9A26CE4B" box="[780,837,753,771]" gridcol="6" gridrow="21" pageId="11" pageNumber="84">26.17</td>
<td id="05250949D448001DFC83AABE9AD5CE4B" box="[853,950,753,771]" gridcol="7" gridrow="21" pageId="11" pageNumber="84">0.413</td>
<td id="05250949D448001DFC18AABE9A9BCE4B" box="[974,1016,753,771]" gridcol="8" gridrow="21" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFBCEAABE9DE4CE4B" box="[1048,1159,753,771]" gridcol="9" gridrow="21" pageId="11" pageNumber="84">4.7784</td>
<td id="05250949D448001DFB72AABE9DBECE4B" box="[1188,1245,753,771]" gridcol="10" gridrow="21" pageId="11" pageNumber="84">18.36</td>
<td id="05250949D448001DFB2EAABE9C2DCE4B" box="[1272,1358,753,771]" gridcol="11" gridrow="21" pageId="11" pageNumber="84">1.428e-04</td>
<td id="05250949D448001DFAB0AABE9CEDCE4B" box="[1382,1422,753,771]" gridcol="12" gridrow="21" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AB469CEDCE53" box="[133,1422,777,795]" gridrow="22" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AB4699B7CE53" box="[133,212,777,795]" gridcol="0" gridrow="22" pageId="11" pageNumber="84">Zygo-B</th>
<td id="05250949D448001DFF29AB46983ACE53" box="[255,345,777,795]" gridcol="1" gridrow="22" pageId="11" pageNumber="84">9.8423</td>
<td id="05250949D448001DFEA2AB4698C0CE53" box="[372,419,777,795]" gridcol="2" gridrow="22" pageId="11" pageNumber="84">50.00</td>
<td id="05250949D448001DFE6FAB469B6ECE53" box="[441,525,777,795]" gridcol="3" gridrow="22" pageId="11" pageNumber="84">2.738e-13</td>
<td id="05250949D448001DFDFAAB469B01CE53" box="[556,610,777,795]" gridcol="4" gridrow="22" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAB469B88CE53" box="[681,747,777,795]" gridcol="5" gridrow="22" pageId="11" pageNumber="84">2.8808</td>
<td id="05250949D448001DFCDAAB469A26CE53" box="[780,837,777,795]" gridcol="6" gridrow="22" pageId="11" pageNumber="84">20.65</td>
<td id="05250949D448001DFC83AB469AD5CE53" box="[853,950,777,795]" gridcol="7" gridrow="22" pageId="11" pageNumber="84">0.009</td>
<td id="05250949D448001DFC18AB469A9BCE53" box="[974,1016,777,795]" gridcol="8" gridrow="22" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAB469DE4CE53" box="[1048,1159,777,795]" gridcol="9" gridrow="22" pageId="11" pageNumber="84">4.1662</td>
<td id="05250949D448001DFB72AB469DBECE53" box="[1188,1245,777,795]" gridcol="10" gridrow="22" pageId="11" pageNumber="84">16.91</td>
<td id="05250949D448001DFB2EAB469C2DCE53" box="[1272,1358,777,795]" gridcol="11" gridrow="22" pageId="11" pageNumber="84">6.537e-04</td>
<td id="05250949D448001DFAB0AB469CEDCE53" box="[1382,1422,777,795]" gridcol="12" gridrow="22" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AB6E9CEDCE7B" box="[133,1422,801,819]" gridrow="23" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AB6E99B7CE7B" box="[133,212,801,819]" gridcol="0" gridrow="23" pageId="11" pageNumber="84">Zygo-L</th>
<td id="05250949D448001DFF29AB6E983ACE7B" box="[255,345,801,819]" gridcol="1" gridrow="23" pageId="11" pageNumber="84">19277</td>
<td id="05250949D448001DFEA2AB6E98C0CE7B" box="[372,419,801,819]" gridcol="2" gridrow="23" pageId="11" pageNumber="84">70.43</td>
<td id="05250949D448001DFE6FAB6E9B6ECE7B" box="[441,525,801,819]" gridcol="3" gridrow="23" pageId="11" pageNumber="84">&lt;2.2e-16</td>
<td id="05250949D448001DFDFAAB6E9B01CE7B" box="[556,610,801,819]" gridcol="4" gridrow="23" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAB6E9B88CE7B" box="[681,747,801,819]" gridcol="5" gridrow="23" pageId="11" pageNumber="84">6.0478</td>
<td id="05250949D448001DFCDAAB6E9A26CE7B" box="[780,837,801,819]" gridcol="6" gridrow="23" pageId="11" pageNumber="84">13.86</td>
<td id="05250949D448001DFC83AB6E9AD5CE7B" box="[853,950,801,819]" gridcol="7" gridrow="23" pageId="11" pageNumber="84">3.134e-05</td>
<td id="05250949D448001DFC18AB6E9A9BCE7B" box="[974,1016,801,819]" gridcol="8" gridrow="23" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAB6E9DE4CE7B" box="[1048,1159,801,819]" gridcol="9" gridrow="23" pageId="11" pageNumber="84">0.2211</td>
<td id="05250949D448001DFB72AB6E9DBECE7B" box="[1188,1245,801,819]" gridcol="10" gridrow="23" pageId="11" pageNumber="84">14.21</td>
<td id="05250949D448001DFB2EAB6E9C2DCE7B" box="[1272,1358,801,819]" gridcol="11" gridrow="23" pageId="11" pageNumber="84">0.828</td>
<td id="05250949D448001DFAB0AB6E9CEDCE7B" box="[1382,1422,801,819]" gridcol="12" gridrow="23" pageId="11" pageNumber="84">ns</td>
</tr>
<tr id="46F46035D448001DFF53AB779CEDCE04" box="[133,1422,824,844]" gridrow="24" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AB7799B7CE04" box="[133,212,824,844]" gridcol="0" gridrow="24" pageId="11" pageNumber="84">C1-M2</th>
<td id="05250949D448001DFF29AB77983ACE04" box="[255,345,824,844]" gridcol="1" gridrow="24" pageId="11" pageNumber="84">8.6045</td>
<td id="05250949D448001DFEA2AB7798C0CE04" box="[372,419,824,844]" gridcol="2" gridrow="24" pageId="11" pageNumber="84">38.50</td>
<td id="05250949D448001DFE6FAB779B6ECE04" box="[441,525,824,844]" gridcol="3" gridrow="24" pageId="11" pageNumber="84">1.677e-10</td>
<td id="05250949D448001DFDFAAB779B01CE04" box="[556,610,824,844]" gridcol="4" gridrow="24" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAB779B88CE04" box="[681,747,824,844]" gridcol="5" gridrow="24" pageId="11" pageNumber="84">4.5861</td>
<td id="05250949D448001DFCDAAB779A26CE04" box="[780,837,824,844]" gridcol="6" gridrow="24" pageId="11" pageNumber="84">32.99</td>
<td id="05250949D448001DFC83AB779AD5CE04" box="[853,950,824,844]" gridcol="7" gridrow="24" pageId="11" pageNumber="84">6.216e-05</td>
<td id="05250949D448001DFC18AB779A9BCE04" box="[974,1016,824,844]" gridcol="8" gridrow="24" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFBCEAB779DE4CE04" box="[1048,1159,824,844]" gridcol="9" gridrow="24" pageId="11" pageNumber="84">4.6721</td>
<td id="05250949D448001DFB72AB779DBECE04" box="[1188,1245,824,844]" gridcol="10" gridrow="24" pageId="11" pageNumber="84">28.61</td>
<td id="05250949D448001DFB2EAB779C2DCE04" box="[1272,1358,824,844]" gridcol="11" gridrow="24" pageId="11" pageNumber="84">6.487e-05</td>
<td id="05250949D448001DFAB0AB779CEDCE04" box="[1382,1422,824,844]" gridcol="12" gridrow="24" pageId="11" pageNumber="84">***</td>
</tr>
<tr id="46F46035D448001DFF53AB1F9CEDCE2C" box="[133,1422,848,868]" gridrow="25" pageId="11" pageNumber="84">
<th id="05250949D448001DFF53AB1F99B7CE2C" box="[133,212,848,868]" gridcol="0" gridrow="25" pageId="11" pageNumber="84">C1-C1</th>
<td id="05250949D448001DFF29AB1F983ACE2C" box="[255,345,848,868]" gridcol="1" gridrow="25" pageId="11" pageNumber="84">12.1752</td>
<td id="05250949D448001DFEA2AB1F98C0CE2C" box="[372,419,848,868]" gridcol="2" gridrow="25" pageId="11" pageNumber="84">40.66</td>
<td id="05250949D448001DFE6FAB1F9B6ECE2C" box="[441,525,848,868]" gridcol="3" gridrow="25" pageId="11" pageNumber="84">3.83e-15</td>
<td id="05250949D448001DFDFAAB1F9B01CE2C" box="[556,610,848,868]" gridcol="4" gridrow="25" pageId="11" pageNumber="84">***</td>
<td id="05250949D448001DFD7FAB1F9B88CE2C" box="[681,747,848,868]" gridcol="5" gridrow="25" pageId="11" pageNumber="84">1.4792</td>
<td id="05250949D448001DFCDAAB1F9A26CE2C" box="[780,837,848,868]" gridcol="6" gridrow="25" pageId="11" pageNumber="84">16.876</td>
<td id="05250949D448001DFC83AB1F9AD5CE2C" box="[853,950,848,868]" gridcol="7" gridrow="25" pageId="11" pageNumber="84">0.1575</td>
<td id="05250949D448001DFC18AB1F9A9BCE2C" box="[974,1016,848,868]" gridcol="8" gridrow="25" pageId="11" pageNumber="84">ns</td>
<td id="05250949D448001DFBCEAB1F9DE4CE2C" box="[1048,1159,848,868]" gridcol="9" gridrow="25" pageId="11" pageNumber="84">4.4133</td>
<td id="05250949D448001DFB72AB1F9DBECE2C" box="[1188,1245,848,868]" gridcol="10" gridrow="25" pageId="11" pageNumber="84">13.451</td>
<td id="05250949D448001DFB2EAB1F9C2DCE2C" box="[1272,1358,848,868]" gridcol="11" gridrow="25" pageId="11" pageNumber="84">0.0006</td>
<td id="05250949D448001DFAB0AB1F9CEDCE2C" box="[1382,1422,848,868]" gridcol="12" gridrow="25" pageId="11" pageNumber="84">***</td>
</tr>
</table>
</paragraph>
<paragraph id="F87B6277D448FFE9FF41ADC19BEFC8B5" blockId="11.[119,750,923,1710]" pageId="11" pageNumber="84">
The yellow colour of the ear and wing spots fades on fixation in alcohol in
<taxonomicName id="3FC419F4D448FFE9FE82ADE39BE2C88B" authority="(Flannery, 1995 b)" baseAuthorityName="Flannery" baseAuthorityYear="1995" box="[340,641,1451,1475]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D448FFE9FE82ADE398DCC88B" box="[340,447,1452,1475]" italics="true" pageId="11" pageNumber="84">N. certans</emphasis>
(
<bibRefCitation id="9C551F86D448FFE9FE1CADE49B1AC88B" author="Flannery, T" box="[458,633,1451,1475]" pageId="11" pageNumber="84" refId="ref21280" refString="Flannery, T. F. 1995 b. Mammals of New Guinea. Second edition. Chatswood, New South Wales: Australian Museum / Reed Books. 568 pp." type="book" year="1995">Flannery, 1995b</bibRefCitation>
)
</taxonomicName>
, similar to
<taxonomicName id="3FC419F4D448FFE9FFA1AD869986C8A8" box="[119,229,1480,1504]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="CAB0BE65D448FFE9FFA1AD869986C8A8" box="[119,229,1480,1504]" italics="true" pageId="11" pageNumber="84">N. cyclotis</emphasis>
</taxonomicName>
, and therefore colour assessment from specimens preserved in alcohol may not be entirely accurate.
</paragraph>
<paragraph id="F87B6277D448FFE9FF41AE4C98F9CBE6" blockId="11.[119,750,923,1710]" pageId="11" pageNumber="84">
Little is known of the biology of
<taxonomicName id="3FC419F4D448FFE9FDDEAE4B9B1ACB53" baseAuthorityName="Flannery" baseAuthorityYear="1995" box="[520,633,1540,1563]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="84" phylum="Chordata" rank="species" species="certans">
<emphasis id="CAB0BE65D448FFE9FDDEAE4B9B1ACB53" box="[520,633,1540,1563]" italics="true" pageId="11" pageNumber="84">N. certans</emphasis>
</taxonomicName>
, including its diet, population density, home range, or life expectancy. What natural history is reported is usually from short-term expeditions of less than a few weeks and is summarized in the excellent mammal and field guides of the region (
<bibRefCitation id="9C551F86D448FFE9FD53AE3799DCCBE5" author="Flannery, T" pageId="11" pageNumber="84" refId="ref21280" refString="Flannery, T. F. 1995 b. Mammals of New Guinea. Second edition. Chatswood, New South Wales: Australian Museum / Reed Books. 568 pp." type="book" year="1995">Flannery, 1995b</bibRefCitation>
;
<bibRefCitation id="9C551F86D448FFE9FF1CAED998EDCBE6" author="Bonaccorso, F" box="[202,398,1686,1710]" pageId="11" pageNumber="84" refId="ref20829" refString="Bonaccorso, F. J. 1998. Bats of Papua New Guinea. Washington, D. C.: Conservation International. 489 pp." type="book" year="1998">Bonaccorso, 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>