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<document id="1316BB5B2B785EC88719B8BB057A13DE" ID-CLB-Dataset="45583" ID-DOI="10.11646/zootaxa.3811.4.4" ID-GBIF-Dataset="2b8dd933-6926-4671-9b84-c3f5a520fe51" ID-ISSN="1175-5326" ID-Zenodo-Dep="249520" ID-ZooBank="AE22B01B-B3FF-4B60-9452-A94DDDB20C2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432306852" checkinUser="plazi" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2014" docId="F53487E6FFC1D818FF7EFDB6FF125EBC" docLanguage="en" docName="zt03811p514.pdf" docOrigin="Zootaxa 3811 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enghoffosoma zebra Likhitrakarn, Golovatch &amp; Panha, 2014, sp. n." docType="treatment" docVersion="11" lastPageNumber="497" masterDocId="090DFF9EFFC3D81EFFE9FFD7FFFD593F" masterDocTitle="Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species" masterLastPageNumber="514" masterPageNumber="491" pageNumber="493" updateTime="1698331558507" updateUser="ExternalLinkService">
<mods:mods id="D69631B46865D2B96DB938B04A74AE99" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0B3C4B87429176884612D974E656BC4B">Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species</mods:title>
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<mods:namePart id="1D419ADAC2F60EB823DC4B4E7E77DBA0">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="C06C8CE153DEEE03F6202E4E95E99D0B">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="7EF9713BB3EDF5BC5B978384953AB12F">Panha, Somsak</mods:namePart>
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<mods:title id="10109BACD6FF787F88AA239CC79BD26D">Zootaxa</mods:title>
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<mods:date id="C7DCFF60F493E4D272DD180FA0EF6CBF">2014</mods:date>
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<treatment id="F53487E6FFC1D818FF7EFDB6FF125EBC" ID-DOI="http://doi.org/10.5281/zenodo.6136568" ID-GBIF-Taxon="119430662" ID-Zenodo-Dep="6136568" LSID="urn:lsid:plazi:treatment:F53487E6FFC1D818FF7EFDB6FF125EBC" httpUri="http://treatment.plazi.org/id/F53487E6FFC1D818FF7EFDB6FF125EBC" lastPageId="6" lastPageNumber="497" pageId="2" pageNumber="493">
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<emphasis id="4FE9EAE2FFC1D81CFF7EFDB6FE215B44" bold="true" box="[151,476,609,635]" pageId="2" pageNumber="493">
<taxonomicName id="BA9D4D73FFC1D81CFF7EFDB6FE6F5B44" ID-CoL="39T4M" box="[151,402,609,635]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="493" phylum="Arthropoda" rank="species" species="zebra" status="sp. nov.">
<emphasis id="4FE9EAE2FFC1D81CFF7EFDB6FE6F5B44" bold="true" box="[151,402,609,635]" italics="true" pageId="2" pageNumber="493">Enghoffosoma zebra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54DA5799FFC1D81CFE70FDB6FE215B44" box="[409,476,609,635]" pageId="2" pageNumber="493" rank="species">sp. n.</taxonomicNameLabel>
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<figureCitation id="E5A62A75FFC1D81CFF7EFD53FF045BA3" box="[151,249,644,669]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1853,1875]" captionStartId-1="4.[151,250,1632,1654]" captionStartId-2="5.[151,250,1368,1390]" captionStartId-3="6.[151,250,1380,1402]" captionStartId-4="7.[151,250,1567,1589]" captionStartId-5="8.[151,250,1552,1574]" captionStartId-6="9.[151,250,1496,1518]" captionTargetBox-0="[216,1371,193,1827]" captionTargetBox-1="[151,1436,193,1610]" captionTargetBox-2="[274,1311,199,1335]" captionTargetBox-3="[264,1315,195,1358]" captionTargetBox-4="[151,1435,237,1518]" captionTargetBox-5="[151,1435,221,1502]" captionTargetBox-6="[151,1435,193,1474]" captionTargetId-0="figure@3.[216,1371,193,1832]" captionTargetId-1="figure@4.[151,1436,193,1610]" captionTargetId-2="figure@5.[264,1323,193,1347]" captionTargetId-3="figure@6.[264,1323,193,1358]" captionTargetId-4="figure@7.[151,1436,237,1519]" captionTargetId-5="figure@8.[151,1436,221,1503]" captionTargetId-6="figure@9.[151,1436,193,1475]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. Enghoffosoma zebra sp. n., habitus, live coloration. A: holotype. B: male paratype from Numtip Temple. C: male paratype from Tadtone. D: male paratype from Ban Xanapho." captionText-1="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." captionText-2="FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text." captionText-3="FIGURE 4. Enghoffosoma zebra sp. n., male paratype from Numtip Temple (A, B), male paratype from Ban Xanapho (C, D). A, B: right gonopod, mesal and submesal views, respectively. C, D: right gonopod, mesal and submesal views, respectively. Scale bar: 0.5 mm. Designations explained in text." captionText-4="FIGURE 5. Enghoffosoma zebra sp. n., holotype, right gonopod. A, B: lateral and mesal views, respectively. C – F: distal part of left gonopod, sublateral, submesal, subcaudal and suboral views, respectively. Scale bar: 0.2 mm." captionText-5="FIGURE 6. Enghoffosoma zebra sp. n., male paratype from Numtip Temple (A, B), male paratype from Ban Tadtone (C, D). A, B: right gonopod, mesal and lateral views, respectively. C, D: right gonopod, mesal and lateral views, respectively. Scale bar: 0.2 mm." captionText-6="FIGURE 7. Enghoffosoma zebra sp. n., male paratype from Ban Xanapho, right gonopod. A, B: lateral and mesal views, respectively. C – F: distal part of left gonopod, lateral, mesal, subcaudal and suboral views, respectively. Scale bar: 0.2 mm." httpUri-0="https://zenodo.org/record/249521/files/figure.png" httpUri-1="https://zenodo.org/record/249522/files/figure.png" httpUri-2="https://zenodo.org/record/249523/files/figure.png" httpUri-3="https://zenodo.org/record/249524/files/figure.png" httpUri-4="https://zenodo.org/record/249525/files/figure.png" httpUri-5="https://zenodo.org/record/249526/files/figure.png" httpUri-6="https://zenodo.org/record/249527/files/figure.png" pageId="2" pageNumber="493">Figs 1–7</figureCitation>
</paragraph>
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<emphasis id="4FE9EAE2FFC1D81CFF7EFD1BFEF95BDA" bold="true" box="[151,260,716,741]" pageId="2" pageNumber="493">Holotype</emphasis>
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male (
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),
<collectingCountry id="058A7660FFC1D81CFE34FD1BFDBE5BDA" box="[477,579,716,741]" name="Thailand" pageId="2" pageNumber="493">Thailand</collectingCountry>
, Nakhon Ratchasima Province, Wang Nam Khiao District, Sakaerat Environmental Research Station, ca
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a.s.l.,
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,
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, 0
<date id="09231030FFC1D81CFBE3FD27FB805A37" box="[1034,1149,752,776]" pageId="2" pageNumber="493" value="2013-03-10">3.10.2013</date>
, leg. C. Sutcharit and W. Siriwut.
</paragraph>
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<typeStatus id="A2268852FFC1D81CFF2EFCE0FEBC5A6F" box="[199,321,823,848]" pageId="2" pageNumber="493" type="paratype">Paratypes</typeStatus>
.
</emphasis>
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,
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(
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),
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,
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(
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), same data, together with
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.
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,
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(
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),
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,
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(
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),
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,
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(
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), same locality,
<date id="09231030FFC1D81CFAFEFC8BFA655A4B" box="[1303,1432,860,884]" pageId="2" pageNumber="493" value="2009-04-24">24.04.2009</date>
, leg. S. Panha, C. Sutcharit and N. Likhitrakarn.
<specimenCount id="6B9BFD79FFC1D81CFD56FC57FCE75AA7" box="[703,794,895,920]" pageId="2" pageNumber="493" type="male">3 males</specimenCount>
,
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(
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), Ubon Ratchathani Province, Khong Chiam District, Tadtone, ca
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a.s.l.,
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,
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,
<date id="09231030FFC1D81CFC7FFC73FBEB5A82" box="[918,1046,932,957]" pageId="2" pageNumber="493" value="2011-05-14">14.05.2011</date>
, leg. S. Panha, C. Sutcharit and N. Likhitrakarn.
<specimenCount id="6B9BFD79FFC1D81CFEDDFC1FFE785ADF" box="[308,389,967,992]" pageId="2" pageNumber="493" type="male">1 male</specimenCount>
,
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(
<collectionCode id="1B8CAE35FFC1D81CFDFCFC1FFD915ADF" box="[533,620,968,992]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="2" pageNumber="493">CUMZ</collectionCode>
), same Province, Nam Yuen District, Numtip Temple, ca
<quantity id="BA659B15FFC1D81CFAE4FC1FFAAA5AE0" box="[1293,1367,968,992]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="493" unit="m" value="220.0">220 m</quantity>
a.s.l.,
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,
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,
<date id="09231030FFC1D81CFE2CFC3BFDB85D3B" box="[453,581,1004,1028]" pageId="2" pageNumber="493" value="2010-10-10">10.10.2010</date>
, leg. S. Panha, C. Sutcharit and N. Likhitrakarn.
<specimenCount id="6B9BFD79FFC1D81CFB8FFC3BFB425D3A" box="[1126,1215,1004,1029]" pageId="2" pageNumber="493" type="male">2 males</specimenCount>
,
<specimenCount id="6B9BFD79FFC1D81CFB23FC3BFAD65D3A" box="[1226,1323,1004,1029]" pageId="2" pageNumber="493" type="female">1 female</specimenCount>
(
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), same District, Yod Dom Forest Reserve,
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a.s.l.,
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,
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,
<date id="09231030FFC1D81CFBC1FBC7FB545D17" box="[1064,1193,1040,1064]" pageId="2" pageNumber="493" value="2010-10-10">10.10.2010</date>
, leg. C. Sutcharit and N. Likhitrakarn.
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(
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),
<collectingCountry id="058A7660FFC1D81CFDC2FBE2FD995D73" box="[555,612,1077,1100]" name="Laos" pageId="2" pageNumber="493">Laos</collectingCountry>
, Champasak Province, Pathoumphone District, Ban Xanapho, ca
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a.s.l.,
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,
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,
<date id="09231030FFC1D81CFDEEFB8FFD745D4F" box="[519,649,1112,1136]" pageId="2" pageNumber="493" value="2013-07-22">22.07.2013</date>
, leg. C. Sutcharit.
</paragraph>
<paragraph id="7D2236F0FFC1D81CFF2EFBABFBEB5DAA" blockId="2.[151,1437,716,2037]" box="[199,1046,1148,1173]" pageId="2" pageNumber="493">
<emphasis id="4FE9EAE2FFC1D81CFF2EFBABFEF05DAA" bold="true" box="[199,269,1148,1173]" pageId="2" pageNumber="493">Name</emphasis>
. To emphasize a stripy, zebra-like colour pattern; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFC1D81CFF2EFB48FB895C3F" pageId="2" pageNumber="493" type="diagnosis">
<paragraph id="7D2236F0FFC1D81CFF2EFB48FB895C3F" blockId="2.[151,1437,716,2037]" pageId="2" pageNumber="493">
<emphasis id="4FE9EAE2FFC1D81CFF2EFB48FEBC5D87" bold="true" box="[199,321,1183,1208]" pageId="2" pageNumber="493">Diagnosis.</emphasis>
Differs from congeners in the characteristic colour pattern of blackish to blackish brown bands with contrasting dark brown to whitish posterior halves of meta- and paraterga, coupled with the gonopod showing a prominent process
<emphasis id="4FE9EAE2FFC1D81CFE85FB30FE855C3F" bold="true" box="[364,376,1255,1280]" pageId="2" pageNumber="493">e</emphasis>
and an even stronger, twisted solenomere (
<emphasis id="4FE9EAE2FFC1D81CFCB7FB30FC8C5C3F" bold="true" box="[862,881,1255,1280]" pageId="2" pageNumber="493">sl</emphasis>
) (see also Key below).
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFC1D818FF2EFADBFADB5F94" lastPageId="6" lastPageNumber="497" pageId="2" pageNumber="493" type="description">
<paragraph id="7D2236F0FFC1D81CFF2EFADBFBD25C77" blockId="2.[151,1437,716,2037]" pageId="2" pageNumber="493">
<emphasis id="4FE9EAE2FFC1D81CFF2EFADBFEA45C1A" bold="true" box="[199,345,1292,1317]" pageId="2" pageNumber="493">Description.</emphasis>
Length
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(male) or
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(female), width of midbody pro- and metazonae 2.9–4.5 and
<quantity id="BA659B15FFC1D81CFEC8FAF8FE555C77" box="[289,424,1327,1352]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="5.2" metricValueMin="3.5" pageId="2" pageNumber="493" unit="mm" value="4.35" valueMax="5.2" valueMin="3.5">3.5–5.2 mm</quantity>
(male) or 3.5–5.0 and
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(female), respectively.
</paragraph>
<paragraph id="7D2236F0FFC1D81CFF2EFA83FDDF5F7B" blockId="2.[151,1437,716,2037]" pageId="2" pageNumber="493">
Coloration of live animals blackish to blackish brown with a pattern of contrasting dark brown to whitish posterior halves of collum, metaterga, paraterga and epiproct, of dark brown to light brown (
<figureCitation id="E5A62A75FFC1D81CFB74FAA0FB275CAF" box="[1181,1242,1399,1424]" captionStart="FIGURE 1" captionStartId="3.[151,250,1853,1875]" captionTargetBox="[216,1371,193,1827]" captionTargetId="figure@3.[216,1371,193,1832]" captionTargetPageId="3" captionText="FIGURE 1. Enghoffosoma zebra sp. n., habitus, live coloration. A: holotype. B: male paratype from Numtip Temple. C: male paratype from Tadtone. D: male paratype from Ban Xanapho." httpUri="https://zenodo.org/record/249521/files/figure.png" pageId="2" pageNumber="493">Fig. 1</figureCitation>
A–C) or carmine to light pink legs and antennae (
<figureCitation id="E5A62A75FFC1D81CFE15FA4BFDC65C8B" box="[508,571,1436,1461]" captionStart="FIGURE 1" captionStartId="3.[151,250,1853,1875]" captionTargetBox="[216,1371,193,1827]" captionTargetId="figure@3.[216,1371,193,1832]" captionTargetPageId="3" captionText="FIGURE 1. Enghoffosoma zebra sp. n., habitus, live coloration. A: holotype. B: male paratype from Numtip Temple. C: male paratype from Tadtone. D: male paratype from Ban Xanapho." httpUri="https://zenodo.org/record/249521/files/figure.png" pageId="2" pageNumber="493">Fig. 1</figureCitation>
D); coloration in alcohol, after a long time of preservation, blackish brown to light brown, posterior halves of collum, metaterga, paraterga and epiproct mostly light brown to whitish (
<figureCitation id="E5A62A75FFC1D81CFA83FA68FF595CC3" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Figs 2</figureCitation>
A–F &amp; J–L), antennae and legs dark brown to light brown (
<figureCitation id="E5A62A75FFC1D81CFCADFA33FC775CC3" box="[836,906,1508,1533]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Figs 2</figureCitation>
A–F, J &amp; K) or carmine to light pink (
<figureCitation id="E5A62A75FFC1D81CFAD1FA33FA815CC3" box="[1336,1404,1508,1533]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Fig. 2</figureCitation>
L), venter and a few basal podomeres light yellow to whitish (
<figureCitation id="E5A62A75FFC1D81CFCDCF9D0FC805F1F" box="[821,893,1543,1568]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Figs 2</figureCitation>
A–F, J &amp; K) or light crimson to light pink, legs growing crimson distally (
<figureCitation id="E5A62A75FFC1D81CFE57F9FBFDFF5F7B" box="[446,514,1580,1605]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Fig. 2</figureCitation>
L).
</paragraph>
<paragraph id="7D2236F0FFC1D81CFF2EF998FEE25E23" blockId="2.[151,1437,716,2037]" pageId="2" pageNumber="493">
Clypeolabral region sparsely setose, vertex bare, epicranial suture distinct. Antennae rather short (
<figureCitation id="E5A62A75FFC1D81CFAFEF998FAA75F57" box="[1303,1370,1615,1640]" captionStart="FIGURE 1" captionStartId="3.[151,250,1853,1875]" captionTargetBox="[216,1371,193,1827]" captionTargetId="figure@3.[216,1371,193,1832]" captionTargetPageId="3" captionText="FIGURE 1. Enghoffosoma zebra sp. n., habitus, live coloration. A: holotype. B: male paratype from Numtip Temple. C: male paratype from Tadtone. D: male paratype from Ban Xanapho." httpUri="https://zenodo.org/record/249521/files/figure.png" pageId="2" pageNumber="493">Figs 1</figureCitation>
A–D, 2A &amp; L), clavate (antennomere 6 highest), extending behind body segment 3 (male) or 2 (female) when stretched dorsally. In width, head &lt;segment 3 = 4 &lt;collum &lt;segment 5 &lt;2 &lt;6–17 (both sexes), gently and gradually tapering thereafter. Collum with three transverse rows of setae: 4+
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anterior, 3+
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intermediate, and 5+
<quantity id="BA659B15FFC1D81CFA86F96BFA615FEA" box="[1391,1436,1724,1749]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="493" unit="in" value="5.0">5 in</quantity>
posterior row; caudal corner of paraterga very broadly rounded, declined, not extending behind rear margin (
<figureCitation id="E5A62A75FFC1D81CFA83F908FF585E23" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Figs 2</figureCitation>
A, B &amp; L).
</paragraph>
<paragraph id="7D2236F0FFC1D81BFF2EF8FEFE8A5E0A" blockId="2.[151,1437,716,2037]" lastBlockId="5.[151,1436,1460,2025]" lastPageId="5" lastPageNumber="496" pageId="2" pageNumber="493">
Tegument smooth and shining, prozonae finely shagreened, metaterga smooth and leathery, posterior halves delicately rugulose; surface below paraterga finely microgranulate (
<figureCitation id="E5A62A75FFC1D81CFC79F89BFC2B5E5B" box="[912,982,1868,1893]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Figs 2</figureCitation>
A–F &amp; J–L). Postcollum metaterga with two transverse rows of setae traceable at least as insertion points when setae broken off: 3(4)+3(4) in anterior (presulcus), 4+
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posterior (post-sulcus) row, caudal row barely traceable as insertion points. Tergal setae long and slender, about 1/3 of metatergal length. Axial line invisible. Paraterga rather well developed (
<figureCitation id="E5A62A75FFC1D81CFB29F86FFAF55EEF" box="[1216,1288,1976,2001]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="2" pageNumber="493">Figs 2</figureCitation>
A–F &amp; J–L), mostly slightly upturned, all lying below dorsum, set at about half of body height, subhorizontal, caudal corner almost or fully pointed, produced beyond rear tergal margin until segment 15, increasingly rounded and smaller on following segments; posterior edge nearly straight (
<figureCitation id="E5A62A75FFC6D81BFD33FA0FFCDC5CCF" box="[730,801,1496,1521]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
A–F &amp; J–L). Calluses on paraterga narrow, delimited by a sulcus only dorsally. Paraterga 2 broad, anterior edge angular to rounded, lateral edge without incision (
<figureCitation id="E5A62A75FFC6D81BFAD3FA2BFA7C5F2A" box="[1338,1409,1532,1557]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
A, B &amp; L). Ozopore evident, lateral, lying in an ovoid groove at about 1/4 of metatergite’s length in front of caudal corner. Transverse sulcus at most faint and abbreviated (
<figureCitation id="E5A62A75FFC6D81BFCFBF993FCA55F62" box="[786,856,1604,1629]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
A–F &amp; J–L). Stricture between pro- and metazonae narrow, deep, line-shaped, clearly beaded at bottom down to base of paraterga (
<figureCitation id="E5A62A75FFC6D81BFBD3F9BFFB7E5FBF" box="[1082,1155,1640,1665]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
A–E &amp; J). Pleurosternal carinae strongly developed, complete crests with a very sharp caudal tooth in segments 2–17, a very small caudal tooth in segment 18, absent from segment 19 (both sexes) (
<figureCitation id="E5A62A75FFC6D81BFCB5F967FC545FF7" box="[860,937,1712,1737]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
B, D &amp; E). Epiproct (
<figureCitation id="E5A62A75FFC6D81BFB59F967FB065FF7" box="[1200,1275,1712,1737]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
E–G) conical, flattened dorsoventrally, with two small apical papillae; tip subtruncate; pre-apical papillae very small, but visible, lying rather close to tip. Hypoproct roundly subtriangular, setiferous knobs at caudal edge very small and wellseparated (
<figureCitation id="E5A62A75FFC6D81BFEF9F8CBFEAE5E0A" box="[272,339,1820,1845]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Fig. 2</figureCitation>
G).
</paragraph>
<caption id="29E26678FFC0D81DFF7EF8EAFCF55E4D" httpUri="https://zenodo.org/record/249521/files/figure.png" pageId="3" pageNumber="494" targetBox="[216,1371,193,1827]" targetPageId="3">
<paragraph id="7D2236F0FFC0D81DFF7EF8EAFCF55E4D" blockId="3.[151,1436,1853,1906]" pageId="3" pageNumber="494">
<emphasis id="4FE9EAE2FFC0D81DFF7EF8EAFEEE5E6C" bold="true" box="[151,275,1853,1875]" pageId="3" pageNumber="494">FIGURE 1.</emphasis>
<taxonomicName id="BA9D4D73FFC0D81DFEF3F8EAFE175E6C" box="[282,490,1853,1875]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="zebra" status="sp. nov.">
<emphasis id="4FE9EAE2FFC0D81DFEF3F8EAFE175E6C" box="[282,490,1853,1875]" italics="true" pageId="3" pageNumber="494">Enghoffosoma zebra</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFC0D81DFE18F8E9FDCC5E6C" bold="true" box="[497,561,1854,1875]" pageId="3" pageNumber="494">
<taxonomicNameLabel id="54DA5799FFC0D81DFE18F8E9FDD65E6C" box="[497,555,1854,1875]" pageId="3" pageNumber="494" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
habitus, live coloration. A: holotype. B: male paratype from Numtip Temple. C: male paratype from Tadtone. D: male paratype from Ban Xanapho.
</paragraph>
</caption>
<caption id="29E26678FFC7D81AFF7EF9B7FEEF5FCF" httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="4" pageNumber="495" targetBox="[151,1436,193,1610]" targetPageId="4">
<paragraph id="7D2236F0FFC7D81AFF7EF9B7FEEF5FCF" blockId="4.[151,1436,1632,1776]" pageId="4" pageNumber="495">
<emphasis id="4FE9EAE2FFC7D81AFF7EF9B7FEE95F4A" bold="true" box="[151,276,1632,1654]" pageId="4" pageNumber="495">FIGURE 2.</emphasis>
<taxonomicName id="BA9D4D73FFC7D81AFEF2F9B7FE105F49" box="[283,493,1632,1654]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="495" phylum="Arthropoda" rank="species" species="zebra" status="sp. nov.">
<emphasis id="4FE9EAE2FFC7D81AFEF2F9B7FE105F49" box="[283,493,1632,1654]" italics="true" pageId="4" pageNumber="495">Enghoffosoma zebra</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFC7D81AFE1DF9B7FDC85F4A" bold="true" box="[500,565,1632,1653]" pageId="4" pageNumber="495">
<taxonomicNameLabel id="54DA5799FFC7D81AFE1DF9B7FDD25F4A" box="[500,559,1632,1653]" pageId="4" pageNumber="495" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
holotype (A–I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9–11, lateral view. E–G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view.
</paragraph>
</caption>
<caption id="29E26678FFC6D81BFF7EFA8FFB025CB2" httpUri="https://zenodo.org/record/249523/files/figure.png" pageId="5" pageNumber="496" targetBox="[274,1311,199,1335]" targetPageId="5">
<paragraph id="7D2236F0FFC6D81BFF7EFA8FFB025CB2" blockId="5.[151,1436,1368,1421]" pageId="5" pageNumber="496">
<emphasis id="4FE9EAE2FFC6D81BFF7EFA8FFEEE5C52" bold="true" box="[151,275,1368,1390]" pageId="5" pageNumber="496">FIGURE 3.</emphasis>
<taxonomicName id="BA9D4D73FFC6D81BFEF3FA8FFE165C51" box="[282,491,1368,1390]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="zebra" status="sp. nov.">
<emphasis id="4FE9EAE2FFC6D81BFEF3FA8FFE165C51" box="[282,491,1368,1390]" italics="true" pageId="5" pageNumber="496">Enghoffosoma zebra</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFC6D81BFE1BFA8FFDCF5C52" bold="true" box="[498,562,1368,1389]" pageId="5" pageNumber="496">
<taxonomicNameLabel id="54DA5799FFC6D81BFE1BFA8FFDD15C52" box="[498,556,1368,1389]" pageId="5" pageNumber="496" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
holotype. A–C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text.
</paragraph>
</caption>
<paragraph id="7D2236F0FFC6D81BFF2EF897FB0D5ED7" blockId="5.[151,1436,1460,2025]" pageId="5" pageNumber="496">
Sterna very densely setose, with a small cone caudally near each coxa, rear cones being a little larger than front ones (
<figureCitation id="E5A62A75FFC6D81BFF32F8B3FEE35E42" box="[219,286,1892,1917]" captionStart="FIGURE 3" captionStartId="5.[151,250,1368,1390]" captionTargetBox="[274,1311,199,1335]" captionTargetId="figure@5.[264,1323,193,1347]" captionTargetPageId="5" captionText="FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text." httpUri="https://zenodo.org/record/249523/files/figure.png" pageId="5" pageNumber="496">Fig. 3</figureCitation>
D); a rather large, linguiform, densely setose, sternal lobe between male coxae 4 (
<figureCitation id="E5A62A75FFC6D81BFB28F8B3FAF45E42" box="[1217,1289,1892,1917]" captionStart="FIGURE 2" captionStartId="4.[151,250,1632,1654]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@4.[151,1436,193,1610]" captionTargetPageId="4" captionText="FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view." httpUri="https://zenodo.org/record/249522/files/figure.png" pageId="5" pageNumber="496">Figs 2</figureCitation>
H &amp; I). Legs moderately long and slender, midbody ones ca 1.0–1.2 (male) or 1.1–1.3 times (female) as long as body height, prefemora with a lateral bulge growing gradually reduced towards telson (
<figureCitation id="E5A62A75FFC6D81BFC0DF87BFBDA5EFA" box="[996,1063,1964,1989]" captionStart="FIGURE 3" captionStartId="5.[151,250,1368,1390]" captionTargetBox="[274,1311,199,1335]" captionTargetId="figure@5.[264,1323,193,1347]" captionTargetPageId="5" captionText="FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text." httpUri="https://zenodo.org/record/249523/files/figure.png" pageId="5" pageNumber="496">Fig. 3</figureCitation>
E), these bulges being especially strong in male; ventral brushes on male tibiae and tarsi traceable until legs of segment 16 (
<figureCitation id="E5A62A75FFC6D81BFB65F807FB335ED7" box="[1164,1230,2000,2025]" captionStart="FIGURE 3" captionStartId="5.[151,250,1368,1390]" captionTargetBox="[274,1311,199,1335]" captionTargetId="figure@5.[264,1323,193,1347]" captionTargetPageId="5" captionText="FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text." httpUri="https://zenodo.org/record/249523/files/figure.png" pageId="5" pageNumber="496">Fig. 3</figureCitation>
E).
</paragraph>
<caption id="29E26678FFC5D818FF7EFAB3FD6B5C88" httpUri="https://zenodo.org/record/249524/files/figure.png" pageId="6" pageNumber="497" targetBox="[264,1315,195,1358]" targetPageId="6">
<paragraph id="7D2236F0FFC5D818FF7EFAB3FD6B5C88" blockId="6.[151,1437,1380,1463]" pageId="6" pageNumber="497">
<emphasis id="4FE9EAE2FFC5D818FF7EFAB3FEEE5C46" bold="true" box="[151,275,1380,1402]" pageId="6" pageNumber="497">FIGURE 4.</emphasis>
<taxonomicName id="BA9D4D73FFC5D818FEF0FAB3FE175C45" box="[281,490,1380,1402]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="497" phylum="Arthropoda" rank="species" species="zebra" status="sp. nov.">
<emphasis id="4FE9EAE2FFC5D818FEF0FAB3FE175C45" box="[281,490,1380,1402]" italics="true" pageId="6" pageNumber="497">Enghoffosoma zebra</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFC5D818FE19FAB3FDCD5C46" bold="true" box="[496,560,1380,1401]" pageId="6" pageNumber="497">
<taxonomicNameLabel id="54DA5799FFC5D818FE19FAB3FDD75C46" box="[496,554,1380,1401]" pageId="6" pageNumber="497" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
male paratype from Numtip Temple (A, B), male paratype from Ban Xanapho (C, D). A, B: right gonopod, mesal and submesal views, respectively. C, D: right gonopod, mesal and submesal views, respectively. Scale bar: 0.5 mm. Designations explained in text.
</paragraph>
</caption>
<paragraph id="7D2236F0FFC5D818FF2EFA09FADB5F94" blockId="6.[151,1437,1502,1923]" pageId="6" pageNumber="497">
Gonopods (
<figureCitation id="E5A62A75FFC5D818FEA5FA09FE695CC8" box="[332,404,1502,1527]" captionStart="FIGURE 3" captionStartId="5.[151,250,1368,1390]" captionTargetBox="[274,1311,199,1335]" captionTargetId="figure@5.[264,1323,193,1347]" captionTargetPageId="5" captionText="FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text." httpUri="https://zenodo.org/record/249523/files/figure.png" pageId="6" pageNumber="497">Figs 3</figureCitation>
A–C &amp; 4–7) with subcylindrical and long coxae, sparsely setose distoventrally. Femorite about 3 times as long as prefemoral (= strongly setose) part. Femorite rather stout and long, slightly curved and enlarged distad, parabasal subventral process
<emphasis id="4FE9EAE2FFC5D818FD7CF9F1FD595F00" bold="true" box="[661,676,1574,1599]" pageId="6" pageNumber="497">p</emphasis>
pointed, rather short or long (
<figureCitation id="E5A62A75FFC5D818FC1BF9F1FBCA5F00" box="[1010,1079,1574,1599]" captionStart="FIGURE 3" captionStartId="5.[151,250,1368,1390]" captionTargetBox="[274,1311,199,1335]" captionTargetId="figure@5.[264,1323,193,1347]" captionTargetPageId="5" captionText="FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text." httpUri="https://zenodo.org/record/249523/files/figure.png" pageId="6" pageNumber="497">Figs 3</figureCitation>
A–C, 5, 6C &amp; D), up to broadly lobe-shaped (
<figureCitation id="E5A62A75FFC5D818FEC7F99DFE8A5F5C" box="[302,375,1610,1635]" captionStart="FIGURE 4" captionStartId="6.[151,250,1380,1402]" captionTargetBox="[264,1315,195,1358]" captionTargetId="figure@6.[264,1323,193,1358]" captionTargetPageId="6" captionText="FIGURE 4. Enghoffosoma zebra sp. n., male paratype from Numtip Temple (A, B), male paratype from Ban Xanapho (C, D). A, B: right gonopod, mesal and submesal views, respectively. C, D: right gonopod, mesal and submesal views, respectively. Scale bar: 0.5 mm. Designations explained in text." httpUri="https://zenodo.org/record/249524/files/figure.png" pageId="6" pageNumber="497">Figs 4</figureCitation>
C, D &amp; 7) or unciform (
<figureCitation id="E5A62A75FFC5D818FD60F99DFD2F5F5C" box="[649,722,1610,1635]" captionStart="FIGURE 4" captionStartId="6.[151,250,1380,1402]" captionTargetBox="[264,1315,195,1358]" captionTargetId="figure@6.[264,1323,193,1358]" captionTargetPageId="6" captionText="FIGURE 4. Enghoffosoma zebra sp. n., male paratype from Numtip Temple (A, B), male paratype from Ban Xanapho (C, D). A, B: right gonopod, mesal and submesal views, respectively. C, D: right gonopod, mesal and submesal views, respectively. Scale bar: 0.5 mm. Designations explained in text." httpUri="https://zenodo.org/record/249524/files/figure.png" pageId="6" pageNumber="497">Figs 4</figureCitation>
A, B &amp; 6A, B); seminal groove running entirely mesally along fermorite, “postfemoral” portion demarcated by a lateral sulcus. Process
<emphasis id="4FE9EAE2FFC5D818FC0CF9B9FC0C5FB8" bold="true" box="[997,1009,1646,1671]" pageId="6" pageNumber="497">e</emphasis>
prominent, stout, expanded apically, about as long as solenomere (
<emphasis id="4FE9EAE2FFC5D818FE0AF945FE0B5F94" bold="true" box="[483,502,1682,1707]" pageId="6" pageNumber="497">sl</emphasis>
), denticulate near middle. Solenomere (
<emphasis id="4FE9EAE2FFC5D818FC51F945FC365F94" bold="true" box="[952,971,1682,1707]" pageId="6" pageNumber="497">sl</emphasis>
) thick, slightly coiled apically.
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFC5D818FF2EF961FF125EBC" pageId="6" pageNumber="497" type="discussion">
<paragraph id="7D2236F0FFC5D818FF2EF961FF125EBC" blockId="6.[151,1437,1502,1923]" pageId="6" pageNumber="497">
<emphasis id="4FE9EAE2FFC5D818FF2EF961FEC65FF0" bold="true" box="[199,315,1718,1743]" pageId="6" pageNumber="497">Remarks.</emphasis>
This species appears to be quite widespread. It has been found in several places in eastern
<collectingCountry id="058A7660FFC5D818FAD0F961FA605FF0" box="[1337,1437,1718,1743]" name="Thailand" pageId="6" pageNumber="497">Thailand</collectingCountry>
to southern
<collectingCountry id="058A7660FFC5D818FEC8F90CFEAA5FCD" box="[289,343,1755,1778]" name="Laos" pageId="6" pageNumber="497">Laos</collectingCountry>
(
<figureCitation id="E5A62A75FFC5D818FE83F90DFE3C5FCC" box="[362,449,1754,1779]" captionStart="FIGURE 20" captionStartId="22.[151,250,1695,1717]" captionTargetBox="[151,1436,195,1673]" captionTargetId="figure@22.[151,1436,195,1674]" captionTargetPageId="22" captionText="FIGURE 20. Distribution of Enghoffosoma species. Open diamond: Enghoffosoma longipes Golovatch, 2011. Open square: Enghoffosoma spinipleurum (Carl, 1941). Open triangle: Enghoffosoma zebra sp. n. Open circle: Enghoffosoma funda sp. n. Asterisk: Enghoffosoma lanceolatum sp. n. and E. anchoriforme sp. n. Crossed square: Enghoffosoma bispinum sp. n." httpUri="https://zenodo.org/record/249540/files/figure.png" pageId="6" pageNumber="497">Fig. 20</figureCitation>
). The animals reveal the same morphological characters in all populations except the population from Ban Xanapho,
<collectingCountry id="058A7660FFC5D818FDE9F928FDC45E29" box="[512,569,1791,1814]" name="Laos" pageId="6" pageNumber="497">Laos</collectingCountry>
, in which the legs and antennae are carmine to light pink while their bodies larger (length
<quantity id="BA659B15FFC5D818FED3F8F5FE1E5E04" box="[314,483,1826,1851]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="4.72" metricValueMin="4.58" pageId="6" pageNumber="497" unit="mm" value="46.5" valueMax="47.2" valueMin="45.8">45.8–47.2 mm</quantity>
). However, the width of midbody pro- and metazonae lies well within the species’ variation range (3.9 and
<quantity id="BA659B15FFC5D818FE5EF890FDEE5E61" box="[439,531,1863,1887]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="497" unit="mm" value="4.7">4.7 mm</quantity>
, 4.2 and
<quantity id="BA659B15FFC5D818FD6BF890FD205E61" box="[642,733,1863,1887]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="497" unit="mm" value="4.9">4.9 mm</quantity>
, respectively), so the Laotian population is attributed to this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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