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<document id="3E878C1C28D1113B34E3628022C68AFF" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFF67D65AC0AD0E8FD579AE9" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus moniliformis Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="A510843C-08CC-4818-A45F-E35AFDE80E71" docUuidSource="ZooBank" docVersion="8" lastPageNumber="14" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="10" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A7812CCB74D33E5469231E537E21540F">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="A7A53D69BA1081EE8D071973737FC1EC">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="AB32D2E077544332B442C7B9B4EF0D7D">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="76248229EE7480C55B4DE22C2CB1DF4B">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="31C51BBA7B2ADEB724022F7D8CD46B9D">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="3EBE6F651E83EF900C253B6329A12AD5" type="email">sgolovatch@yandex.ru&amp;sgolovatch@yandex.ru&amp;somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="22AF73770A33D8E6AAA2ABE9B52501F8">Panha, Somsak</mods:namePart>
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<mods:affiliation id="6B13266D14BA325E69E58BEC19BE3D51">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:title id="DF051BF313F19FCDCAAC17D535DC8095">European Journal of Taxonomy</mods:title>
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<mods:date id="28D0C1DB1D07FA8FCE63FFC3439E0C2F">2016</mods:date>
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<treatment id="03EB5A00FFF67D65AC0AD0E8FD579AE9" ID-DOI="http://doi.org/10.5281/zenodo.3852416" ID-GBIF-Taxon="164246088" ID-Zenodo-Dep="3852416" LSID="urn:lsid:zoobank.org:act:A510843C-08CC-4818-A45F-E35AFDE80E71" httpUri="http://treatment.plazi.org/id/03EB5A00FFF67D65AC0AD0E8FD579AE9" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10">
<subSubSection id="C358B89DFFF67D61AC0AD0E8FB179CD3" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BFDEB16FFF67D61AC0AD0E8FC1D9CF0" blockId="9.[366,1222,1209,1306]" box="[616,972,1209,1236]" pageId="9" pageNumber="10">
<heading id="D0B55C7AFFF67D61AC0AD0E8FC1D9CF0" box="[616,972,1209,1236]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="4C429095FFF67D61AC0AD0E8FCBB9CF0" ID-CoL="7CZPF" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[616,874,1209,1236]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="moniliformis" status="sp. nov.">
<emphasis id="B9363704FFF67D61AC0AD0E8FCBB9CF0" bold="true" box="[616,874,1209,1236]" italics="true" pageId="9" pageNumber="10">Tylopus moniliformis</emphasis>
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<taxonomicNameLabel id="A2058A7FFFF67D61AD10D0EBFC1D9CF0" box="[882,972,1210,1236]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="C358B89DFFF67D61ACB3D151FCB39D3D" box="[721,866,1279,1306]" pageId="9" pageNumber="10" type="description">
<paragraph id="8BFDEB16FFF67D61ACB3D151FCB39D3D" blockId="9.[366,1222,1209,1306]" box="[721,866,1279,1306]" pageId="9" pageNumber="10">
<figureCitation id="1379F793FFF67D61ACB3D151FCE89D3E" box="[721,825,1280,1306]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="10.[189,232,1858,1884]" captionStartId-1="11.[187,230,1964,1990]" captionStartId-2="12.[189,232,1930,1956]" captionTargetBox-0="[189,1398,567,1816]" captionTargetBox-1="[299,1276,1063,1923]" captionTargetBox-2="[189,1398,683,1889]" captionTargetId-0="figure@10.[189,1398,567,1816]" captionTargetId-1="figure@11.[295,1278,1054,1923]" captionTargetId-2="figure@12.[189,1398,683,1889]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." captionText-1="Fig. 8. Tylopus moniliformis sp. nov., paratype, Ƌ, left gonopod. A–B. Mesal and lateral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." captionText-2="Fig. 9. Tylopus moniliformis sp. nov., holotype, Ƌ, left gonopod. A–B. Lateral and mesal views, respectively. C–F. Distal part, lateral, mesal, subcaudal and suboral views, respectively. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837887" figureDoi-1="http://doi.org/10.5281/zenodo.3837889" figureDoi-2="http://doi.org/10.5281/zenodo.3837891" httpUri-0="https://zenodo.org/record/3837887/files/figure.png" httpUri-1="https://zenodo.org/record/3837889/files/figure.png" httpUri-2="https://zenodo.org/record/3837891/files/figure.png" pageId="9" pageNumber="10">Figs 7–9</figureCitation>
,
<figureCitation id="1379F793FFF67D61AD27D0AEFCB39D3D" box="[837,866,1279,1305]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="9" pageNumber="10">26</figureCitation>
</paragraph>
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<subSubSection id="C358B89DFFF67D61AEDFD110FC189DF4" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BFDEB16FFF67D61AEDFD110FEE79D7F" blockId="9.[189,310,1345,1371]" box="[189,310,1345,1371]" pageId="9" pageNumber="10">
<heading id="D0B55C7AFFF67D61AEDFD110FEE79D7F" bold="true" box="[189,310,1345,1371]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9363704FFF67D61AEDFD110FEE79D7F" bold="true" box="[189,310,1345,1371]" pageId="9" pageNumber="10">Diagnosis</emphasis>
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<paragraph id="8BFDEB16FFF67D61AEDFD121FC189DF4" blockId="9.[189,1398,1392,1489]" pageId="9" pageNumber="10">
Differs from all known congeners by the almost missing paraterga, much like in
<taxonomicName id="4C429095FFF67D61AA1ED121FA9D9DAE" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[1148,1356,1392,1418]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hongkhraiensis" status="sp. nov.">
<emphasis id="B9363704FFF67D61AA1ED121FA9D9DAE" box="[1148,1356,1392,1418]" italics="true" pageId="9" pageNumber="10">T. hongkhraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF67D61AB37D121FF209D89" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, coupled with the gonopod structure being similar to that of
<taxonomicName id="4C429095FFF67D61ADB0D1C5FAF99D89" authority="Golovatch, 2013" authorityName="Golovatch" authorityYear="2013" box="[978,1320,1427,1453]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B9363704FFF67D61ADB0D1C5FB879D89" box="[978,1110,1427,1453]" italics="true" pageId="9" pageNumber="10">T. reductus</emphasis>
<bibRefCitation id="EFD396E7FFF67D61AA3DD1C2FAF99D89" author="Golovatch S. I." box="[1119,1320,1427,1453]" pageId="9" pageNumber="10" pagination="1 - 31" refId="ref19577" refString="Golovatch S. I. 2013. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIII. Arthropoda Selecta 22 (1): 1 - 31." type="journal article" year="2013">Golovatch, 2013</bibRefCitation>
</taxonomicName>
, but it differs in the solenophore being more strongly twisted and thicker.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF67D61AEDFD1A9FB9A9E64" box="[189,1099,1528,1601]" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BFDEB16FFF67D61AEDFD1A9FE959E36" blockId="9.[189,324,1528,1554]" box="[189,324,1528,1554]" pageId="9" pageNumber="10">
<heading id="D0B55C7AFFF67D61AEDFD1A9FE959E36" bold="true" box="[189,324,1528,1554]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9363704FFF67D61AEDFD1A9FE959E36" bold="true" box="[189,324,1528,1554]" pageId="9" pageNumber="10">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF67D61AEDFD276FB9A9E64" blockId="9.[189,1099,1574,1601]" box="[189,1099,1574,1601]" pageId="9" pageNumber="10">To emphasize the body being strongly bead-shaped, or moniliform; adjective.</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF67D61AEDFD236FE209F5A" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BFDEB16FFF67D61AEDFD236FE7B9EA5" blockId="9.[189,426,1639,1665]" box="[189,426,1639,1665]" pageId="9" pageNumber="10">
<heading id="D0B55C7AFFF67D61AEDFD236FE7B9EA5" bold="true" box="[189,426,1639,1665]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9363704FFF67D61AEDFD236FE7B9EA5" bold="true" box="[189,426,1639,1665]" pageId="9" pageNumber="10">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B2AE14BFFF67D61AE88D2C7FCEF9ED3" ID-GBIF-Occurrence="2621711302" collectingDate="2013-07-20" collectionCode="CUMZ" collectorName="S. Panha &amp; Panha, C &amp; Sutcharit, W" country="Laos" elevation="970" latitude="15.180556" location="Paksong" longLatPrecision="21" longitude="106.138885" municipality="Tad Fane Waterfall" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
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<typeStatus id="54F955B4FFF67D61AE88D2C7FE8D9E94" box="[234,348,1686,1712]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
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<collectingCountry id="F355AB86FFF67D61AEDFD2E8FEDF9EF7" box="[189,270,1721,1747]" name="Laos" pageId="9" pageNumber="10">LAOS</collectingCountry>
: Ƌ,
<collectingMunicipality id="6B99716CFFF67D61AF59D2E8FDCD9EF7" box="[315,540,1721,1748]" pageId="9" pageNumber="10">Tad Fane Waterfall</collectingMunicipality>
,
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,
<collectingRegion id="498625F4FFF67D61ACF5D2E8FCF29EF7" box="[663,803,1721,1747]" country="Laos" name="Champasak" pageId="9" pageNumber="10">Champasak</collectingRegion>
,
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,
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, ca
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a.s.l.
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,
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<collectingDate id="EFB8343EFFF67D61AB4BD2E8FF2B9ED3" pageId="9" pageNumber="10" value="2013-07-20">20 Jul. 2013</collectingDate>
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, leg.
<collectorName id="26B78EC0FFF67D61AF5BD28CFE729ED2" box="[313,419,1756,1783]" pageId="9" pageNumber="10">S. Panha</collectorName>
, C.
<collectorName id="26B78EC0FFF67D61AFB2D28DFDB49ED3" box="[464,613,1756,1783]" pageId="9" pageNumber="10">Sutcharit, W</collectorName>
. Siriwut (
<collectionCode id="ED5373D3FFF67D61ACBAD28CFCE39ED3" box="[728,818,1757,1783]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9" pageNumber="10">CUMZ</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BFDEB16FFF67D61AE88D34FFEB69F1C" blockId="9.[189,1398,1822,1919]" box="[234,359,1822,1848]" pageId="9" pageNumber="10">
<emphasis id="B9363704FFF67D61AE88D34FFEB69F1C" bold="true" box="[234,359,1822,1848]" pageId="9" pageNumber="10">
<typeStatus id="54F955B4FFF67D61AE88D34FFEB69F1C" box="[234,359,1822,1848]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFDEB16FFF67D61AEDFD310FE209F5A" blockId="9.[189,1398,1822,1919]" pageId="9" pageNumber="10">
<collectingCountry id="F355AB86FFF67D61AEDFD310FEDF9F7F" box="[189,270,1857,1883]" name="Laos" pageId="9" pageNumber="10">LAOS</collectingCountry>
: 2 ƋƋ,
<specimenCount id="9D44209FFFF67D61AF0FD310FE609F78" box="[365,433,1857,1884]" pageId="9" pageNumber="10" type="female">8 ♀♀</specimenCount>
(
<collectionCode id="ED5373D3FFF67D61AFA3D310FDCD9F7F" box="[449,540,1857,1883]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9" pageNumber="10">CUMZ</collectionCode>
); 1 Ƌ,
<specimenCount id="9D44209FFFF67D61AC0DD310FD4C9F78" box="[623,669,1857,1884]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFF67D61ACCFD310FCDB9F7F" box="[685,778,1857,1883]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="9" pageNumber="10">ZMUM</collectionCode>
ρ3061); 1 Ƌ,
<specimenCount id="9D44209FFFF67D61ADD6D310FC309F78" box="[948,993,1857,1884]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFF67D61AD90D310FB9C9F7F" box="[1010,1101,1857,1883]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="10">ZMUC</collectionCode>
); 1 Ƌ,
<specimenCount id="9D44209FFFF67D61AAC2D310FB1C9F78" box="[1184,1229,1857,1884]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFF67D61AABFD310FA929F7F" box="[1245,1347,1857,1883]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="10" type="Museum">NHMW</collectionCode>
), all same data as for
<typeStatus id="54F955B4FFF67D61AFE0D335FE3C9F5A" box="[386,493,1892,1918]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF57D65AEDFD55AFAE699F5" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
<paragraph id="8BFDEB16FFF57D62AEDFD55AFE9F9901" blockId="10.[189,334,267,293]" box="[189,334,267,293]" pageId="10" pageNumber="11">
<heading id="D0B55C7AFFF57D62AEDFD55AFE9F9901" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B9363704FFF57D62AEDFD55AFE9F9901" bold="true" box="[189,334,267,293]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF57D62AEDFD56BFE9B9A20" blockId="10.[189,1399,313,516]" pageId="10" pageNumber="11">
MEASUREMENTS AND COLOUR. Length 14–16 (Ƌ) or
<quantity id="4CBA46F3FFF57D62AD62D568FCAC9970" box="[768,893,313,340]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="10" pageNumber="11" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16–18 mm</quantity>
(
<specimenCount id="9D44209FFFF57D62ADEDD56BFC739970" box="[911,930,314,340]" pageId="10" pageNumber="11" type="female">♀</specimenCount>
), width of midbody pro- and metazonae 0.6–0.9 and
<quantity id="4CBA46F3FFF57D62AF33D50CFE319953" box="[337,480,349,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="10" pageNumber="11" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
(Ƌ) or 0.9–1.2 and
<quantity id="4CBA46F3FFF57D62ACB3D50CFCB19953" box="[721,864,349,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="10" pageNumber="11" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
(
<specimenCount id="9D44209FFFF57D62AD16D50CFC569953" box="[884,903,349,375]" pageId="10" pageNumber="11" type="female">♀</specimenCount>
), respectively. Coloration of live animals dark castaneous brown (
<figureCitation id="1379F793FFF57D62AF82D5D1FD9099BE" box="[480,577,384,410]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="10" pageNumber="11">Fig. 7A</figureCitation>
); legs light brown, venter and a few basal podomeres light brown to yellow-brown. Coloration of alcohol material after three months of preservation faded to light brown; antennae and epiproct light brown to pallid, venter and a few basal podomeres light brown to pallid (
<figureCitation id="1379F793FFF57D62AEA4D5B8FEDD9A20" box="[198,268,489,516]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
B–J).
</paragraph>
<caption id="DF3DBB9EFFF57D62AEDFD313FD5D9FCE" ID-DOI="http://doi.org/10.5281/zenodo.3837887" ID-Zenodo-Dep="3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1858,1884]" targetBox="[189,1398,567,1816]" targetPageId="10">
<paragraph id="8BFDEB16FFF57D62AEDFD313FD5D9FCE" blockId="10.[189,1399,1858,2026]" pageId="10" pageNumber="11">
<emphasis id="B9363704FFF57D62AEDFD313FEC09F79" bold="true" box="[189,273,1858,1885]" pageId="10" pageNumber="11">Fig. 7.</emphasis>
<taxonomicName id="4C429095FFF57D62AF7DD312FDCF9F79" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[287,542,1859,1885]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="moniliformis" status="sp. nov.">
<emphasis id="B9363704FFF57D62AF7DD312FDCF9F79" box="[287,542,1859,1885]" italics="true" pageId="10" pageNumber="11">Tylopus moniliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF57D62AC4ED312FD419F79" box="[556,656,1859,1885]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ (
<emphasis id="B9363704FFF57D62AD2ED312FCAC9F79" bold="true" box="[844,893,1859,1885]" pageId="10" pageNumber="11">B–J</emphasis>
), paratypes, Ƌ, ♀ (
<emphasis id="B9363704FFF57D62AA19D313FB419F78" bold="true" box="[1147,1168,1858,1884]" pageId="10" pageNumber="11">A</emphasis>
).
<emphasis id="B9363704FFF57D62AACDD313FB159F78" bold="true" box="[1199,1220,1858,1884]" pageId="10" pageNumber="11">A</emphasis>
. Habitus, live coloration.
<emphasis id="B9363704FFF57D62AF27D337FEAD9FA4" bold="true" box="[325,380,1894,1920]" pageId="10" pageNumber="11">B–C</emphasis>
. Anterior part of body, dorsal and lateral views, respectively.
<emphasis id="B9363704FFF57D62AA00D337FB489FA4" bold="true" box="[1122,1177,1894,1920]" pageId="10" pageNumber="11">D–E</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively.
<emphasis id="B9363704FFF57D62ACC5D3D8FD0F9F87" bold="true" box="[679,734,1929,1955]" pageId="10" pageNumber="11">F–H</emphasis>
. Posterior part of body, subdorsal, ventral and sublateral views, respectively.
<emphasis id="B9363704FFF57D62AFCFD3FDFE079FE2" bold="true" box="[429,470,1964,1990]" pageId="10" pageNumber="11">I–J</emphasis>
. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFF47D63AEDFD55CFD0B99F3" blockId="11.[189,1399,268,471]" pageId="11" pageNumber="12">
HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (
<figureCitation id="1379F793FFF47D63AF62D57EFE8C996E" box="[256,349,303,330]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
), surpassing body segment 5 (Ƌ) or reaching body segment 4 (
<specimenCount id="9D44209FFFF47D63AA55D561FB9B996E" box="[1079,1098,304,330]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
) when stretched dorsally. In width, head &lt;segment 4 &lt;2 = 3 &lt;collum &lt;segment 5 &lt;6–17 (Ƌ,
<specimenCount id="9D44209FFFF47D63AA6ED502FBF09949" box="[1036,1057,339,365]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 5+5 anterior, 2+2 intermediate and 1+1 posterior; a very faint marginal incision laterally in posterior 1/3; caudal corner very narrowly rounded, not surpassing rear tergal margin (
<figureCitation id="1379F793FFF47D63AC2CD5EDFD1B99F2" box="[590,714,444,471]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">
Fig. 7B
<emphasis id="B9363704FFF47D63ACC5D5ECFD6799F3" bold="true" box="[679,694,445,471]" pageId="11" pageNumber="12">–</emphasis>
C
</figureCitation>
).
</paragraph>
<paragraph id="8BFDEB16FFF47D64AEDFD5AFFB549996" blockId="11.[189,1399,510,994]" lastBlockId="12.[189,1398,267,435]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
BODY. Tegument smooth and shining, prozonae very finely shagreened, metaterga almost smooth, delicately rugulose, leathery (
<figureCitation id="1379F793FFF47D63AC7CD670FDB59A18" box="[542,612,545,572]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
B–F, H). Postcollum metaterga with two transverse rows of setae: 2+2 anterior, always abraded, and 1+1 posterior row, setae traceable at least as insertion points. Tergal setae simple, strong, slender, about 1/3 of metatergal length. Axial line visible, but barely traceable on prozonae (Ƌ). Paraterga 2 well-developed (
<figureCitation id="1379F793FFF47D63ACD9D6DAFCEB9A81" box="[699,826,651,677]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">Fig. 7B, C</figureCitation>
), horizontal, anterior edges protruded anteriorly, fore corner bent ventrad, pointed; lateral edge with two minute incisions in anterior half; caudal corner very narrowly rounded. Paraterga 3 and 4 rounded, expanded laterally, with two minute incisions only in segment 3. Following paraterga virtually missing (Ƌ) or traceable as small, rounded, laterally expanded bulges (
<specimenCount id="9D44209FFFF47D63AF42D749FEE29B16" box="[288,307,792,818]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), in pore-bearing segments with ozopores set at about half of midbody height. Ozopores (
<emphasis id="B9363704FFF47D63AEA5D76BFF3F9B70" bold="true" box="[199,238,826,852]" pageId="11" pageNumber="12">Op</emphasis>
) evident (
<figureCitation id="1379F793FFF47D63AF09D76AFE189B71" box="[363,457,827,853]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">Fig. 7E</figureCitation>
), lateral, lying in an ovoid groove at about 1/3 of metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct (
<figureCitation id="1379F793FFF47D63ADD7D70FFC2D9B5C" box="[949,1020,862,888]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
B–F, H), slightly incomplete on segment 3, complete on segments 4–18 (Ƌ) or complete on segments 5–18 (
<specimenCount id="9D44209FFFF47D63AA25D7D0FB8B9BBF" box="[1095,1114,897,923]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), always incomplete on segment 19 (Ƌ,
<specimenCount id="9D44209FFFF47D63AF18D7F4FE5F9B9B" box="[378,398,933,959]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
), narrow, not reaching bases of paraterga, at most faintly beaded at bottom. Stricture between pro- and metazonae wide, evidently ribbed at bottom down to base of paraterga (
<figureCitation id="1379F793FFF47D63AB69D796FA859BC6" box="[1291,1364,967,994]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
B– F, H). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3, reduced and remaining a sharp caudal tooth on segment 4, thereafter missing (Ƌ) (
<figureCitation id="1379F793FFF37D64AD80D57FFBEC996D" box="[994,1085,302,329]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="12" pageNumber="13">Fig. 7C</figureCitation>
), or thereafter increasingly reduced and remaining a front bulge until segment 18 (
<specimenCount id="9D44209FFFF37D64AD2ED503FCB19948" box="[844,864,338,364]" pageId="12" pageNumber="13" type="female">♀</specimenCount>
). Epiproct (
<figureCitation id="1379F793FFF37D64AD8DD503FBE59948" box="[1007,1076,338,364]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
F–H) rather short, flattened dorsoventrally, tip subtruncate, subapical lateral papillae small, but visible, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge well-separated and evident.
</paragraph>
<caption id="DF3DBB9EFFF47D63AED9D3FDFBC69FCE" ID-DOI="http://doi.org/10.5281/zenodo.3837889" ID-Zenodo-Dep="3837889" httpUri="https://zenodo.org/record/3837889/files/figure.png" pageId="11" pageNumber="12" startId="11.[187,230,1964,1990]" targetBox="[299,1276,1063,1923]" targetPageId="11">
<paragraph id="8BFDEB16FFF47D63AED9D3FDFBC69FCE" blockId="11.[187,1399,1964,2026]" pageId="11" pageNumber="12">
<emphasis id="B9363704FFF47D63AED9D3FDFEDE9FE2" bold="true" box="[187,271,1964,1990]" pageId="11" pageNumber="12">Fig. 8.</emphasis>
<taxonomicName id="4C429095FFF47D63AF7CD3FDFDCC9FE2" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[286,541,1964,1990]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moniliformis" status="sp. nov.">
<emphasis id="B9363704FFF47D63AF7CD3FDFDCC9FE2" box="[286,541,1964,1990]" italics="true" pageId="11" pageNumber="12">Tylopus moniliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF47D63AC49D3FCFD5F9FE3" box="[555,654,1965,1991]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, Ƌ, left gonopod.
<emphasis id="B9363704FFF47D63AD9AD3FDFBFE9FE2" bold="true" box="[1016,1071,1964,1990]" pageId="11" pageNumber="12">A–B</emphasis>
. Mesal and lateral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFF37D64AEDFD58BFAA69A1E" blockId="12.[189,1399,473,570]" pageId="12" pageNumber="13">
STERNA. Moderately setose, without modifications; an entire, high, inverted funnel-shaped, sternal lobe each between Ƌ coxae 3 and 4 (
<figureCitation id="1379F793FFF37D64ACECD5ACFD2F9A33" box="[654,766,509,535]" captionStart="Fig" captionStartId="10.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@10.[189,1398,567,1816]" captionTargetPageId="10" captionText="Fig. 7. Tylopus moniliformis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, subdorsal, ventral and sublateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837887" httpUri="https://zenodo.org/record/3837887/files/figure.png" pageId="12" pageNumber="13">
Fig. 7I
<emphasis id="B9363704FFF37D64AC83D5ACFD219A33" bold="true" box="[737,752,509,535]" pageId="12" pageNumber="13">–</emphasis>
J
</figureCitation>
), lobe being larger between coxae 4. A paramedian pair of evident tubercles in front of gonopod aperture. Legs very long and slender, slightly incrassate
</paragraph>
<caption id="DF3DBB9EFFF37D64AEDFD3DBFECF9FCF" ID-DOI="http://doi.org/10.5281/zenodo.3837891" ID-Zenodo-Dep="3837891" httpUri="https://zenodo.org/record/3837891/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1930,1956]" targetBox="[189,1398,683,1889]" targetPageId="12">
<paragraph id="8BFDEB16FFF37D64AEDFD3DBFECF9FCF" blockId="12.[189,1399,1930,2027]" pageId="12" pageNumber="13">
<emphasis id="B9363704FFF37D64AEDFD3DBFEC19F80" bold="true" box="[189,272,1930,1956]" pageId="12" pageNumber="13">Fig. 9.</emphasis>
<taxonomicName id="4C429095FFF37D64AF7CD3DBFDCD9F80" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[286,540,1930,1956]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="moniliformis" status="sp. nov.">
<emphasis id="B9363704FFF37D64AF7CD3DBFDCD9F80" box="[286,540,1930,1956]" italics="true" pageId="12" pageNumber="13">Tylopus moniliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF37D64AC4BD3DAFD5C9F81" box="[553,653,1931,1957]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, left gonopod.
<emphasis id="B9363704FFF37D64AD96D3DBFBFA9F80" bold="true" box="[1012,1067,1930,1956]" pageId="12" pageNumber="13">A–B</emphasis>
. Lateral and mesal views, respectively.
<emphasis id="B9363704FFF37D64AF38D3FCFE419FE3" bold="true" box="[346,400,1965,1991]" pageId="12" pageNumber="13">C–F</emphasis>
. Distal part, lateral, mesal, subcaudal and suboral views, respectively. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFF27D65AEDFD55AFD52996C" blockId="13.[189,1399,267,329]" pageId="13" pageNumber="14">
in Ƌ, midbody ones ca 1.5–1.8 (Ƌ) or 1.1–1.4 times (
<specimenCount id="9D44209FFFF27D65AD2FD55AFCB19901" box="[845,864,267,293]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
) as long as body height, prefemora without modifications, Ƌ tarsal brushes absent.
</paragraph>
<paragraph id="8BFDEB16FFF27D65AEDFD521FAE699F5" blockId="13.[189,1398,368,465]" pageId="13" pageNumber="14">
GONOPOD. Very simple (
<figureCitation id="1379F793FFF27D65AFB3D521FDCD99AE" box="[465,540,368,394]" captionStart="Fig" captionStartId="11.[187,230,1964,1990]" captionTargetBox="[299,1276,1063,1923]" captionTargetId="figure@11.[295,1278,1054,1923]" captionTargetPageId="11" captionText="Fig. 8. Tylopus moniliformis sp. nov., paratype, Ƌ, left gonopod. A–B. Mesal and lateral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837889" httpUri="https://zenodo.org/record/3837889/files/figure.png" pageId="13" pageNumber="14">Figs 8</figureCitation>
,
<figureCitation id="1379F793FFF27D65AC4BD521FDEB99AE" box="[553,570,368,394]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1398,683,1889]" captionTargetId="figure@12.[189,1398,683,1889]" captionTargetPageId="12" captionText="Fig. 9. Tylopus moniliformis sp. nov., holotype, Ƌ, left gonopod. A–B. Lateral and mesal views, respectively. C–F. Distal part, lateral, mesal, subcaudal and suboral views, respectively. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837891" httpUri="https://zenodo.org/record/3837891/files/figure.png" pageId="13" pageNumber="14">9</figureCitation>
); fermorite straight and rather stout, slightly expanded distad, showing a distinct mesal groove, without outgrowths except for a low, sometimes poorly delimited lobe
<emphasis id="B9363704FFF27D65AB04D5C2FABF9989" bold="true" box="[1382,1390,403,429]" pageId="13" pageNumber="14">l</emphasis>
; solenophore (
<emphasis id="B9363704FFF27D65AF3FD5E6FE5899F5" bold="true" box="[349,393,439,465]" pageId="13" pageNumber="14">sph</emphasis>
) expanded and flattened, typically coiled, suberect distally, devoid of processes.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF27D65AEDFD5A9FD579AE9" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BFDEB16FFF27D65AEDFD5A9FEFE9A36" blockId="13.[189,303,504,530]" box="[189,303,504,530]" pageId="13" pageNumber="14">
<heading id="D0B55C7AFFF27D65AEDFD5A9FEFE9A36" bold="true" box="[189,303,504,530]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9363704FFF27D65AEDFD5A9FEFE9A36" bold="true" box="[189,303,504,530]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF27D65AEDFD677FD579AE9" blockId="13.[189,1399,550,718]" pageId="13" pageNumber="14">
This species shows the gonopod with lobe
<emphasis id="B9363704FFF27D65ACA2D676FD199A65" bold="true" box="[704,712,551,577]" pageId="13" pageNumber="14">l</emphasis>
separated from the femorite by a rather indistinct sulcus, the presence of the latter lobe being one of the main characteristics of
<emphasis id="B9363704FFF27D65AA62D61BFBB29A40" box="[1024,1123,586,612]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C429095FFF27D65AA62D61BFB8E9A40" authorityName="Jeekel" authorityYear="1968" box="[1024,1119,586,612]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Tylopus</taxonomicName>
.
</emphasis>
Since this locality also yielded the
<typeStatus id="54F955B4FFF27D65AF21D63CFE7B9AA3" box="[323,426,621,647]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C429095FFF27D65AFACD63FFB619AA3" authority="Likhitrakarn, Golovatch &amp; Panha, 2014" authorityName="Likhitrakarn, Golovatch &amp; Panha" authorityYear="2014" box="[462,1200,621,647]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rhinoceros">
<emphasis id="B9363704FFF27D65AFACD63FFD0C9AA3" box="[462,733,621,647]" italics="true" pageId="13" pageNumber="14">Desmoxytes rhinoceros</emphasis>
Likhitrakarn,
<bibRefCitation id="EFD396E7FFF27D65ADE1D63CFB619AA3" author="Golovatch S. I." box="[899,1200,621,647]" pageId="13" pageNumber="14" pagination="1 - 19" refId="ref19610" refString="Golovatch S. I. 2014. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XV. Arthropoda Selecta 23 (1): 1 - 19." type="journal article" year="2014">Golovatch &amp; Panha, 2014</bibRefCitation>
</taxonomicName>
(cf.
<bibRefCitation id="EFD396E7FFF27D65AA86D63CFE9D9A8F" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Panha S." pageId="13" pageNumber="14" pagination="483 - 504" refId="ref19913" refString="Likhitrakarn N., Golovatch S. I. &amp; Panha S. 2014 b. Two new species of dragon millipedes, genus Desmoxytes Chamnerlin, 1923, from Laos (Diplopoda: Polydesmida: Paradoxosomatidae), with redescriptions of all four species of Attems from Vietnam. Zootaxa 3931 (4): 483 - 504. http: // dx. doi. org / 10.11646 / zootaxa. 3931.4.2" type="journal article" year="2014">
Likhitrakarn
<emphasis id="B9363704FFF27D65AEDFD6C0FF219A8E" box="[189,240,656,682]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. 2014b
</bibRefCitation>
), Tad Fane Waterfall becomes a site quite important for nature conservation, being the type locality of these two millipede species.
</paragraph>
</subSubSection>
</treatment>
</document>
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