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<mods:title id="74D40F57649C59E2F7829A66285E96C5">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="993F5AE98AC88D40FB25E3AF28F0238B">Frisch, Johannes</mods:namePart>
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<mods:namePart id="5B997F15910E23F659D0754CDF5522A0">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="B6B9A028AF9DC6BE40D64E397EC30686">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:date id="CD313D18C460AFD229816D3761FE357F">2022</mods:date>
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<treatment id="03B7C77AFFC8FFFEBF70FA960FD30561" ID-DOI="http://doi.org/10.5281/zenodo.10724260" ID-Zenodo-Dep="10724260" LSID="urn:lsid:plazi:treatment:03B7C77AFFC8FFFEBF70FA960FD30561" httpUri="http://treatment.plazi.org/id/03B7C77AFFC8FFFEBF70FA960FD30561" lastPageId="10" lastPageNumber="33" pageId="8" pageNumber="31">
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<paragraph id="8BA1766CFFC8FFFCBF70FA9609A10185" blockId="8.[811,1212,1390,1450]" box="[811,1212,1390,1416]" pageId="8" pageNumber="31">
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<taxonomicName id="4C1E0DEFFFC8FFFCBF70FA9609A10185" ID-CoL="6XYZZ" authority="Cameron, 1941" authorityName="Cameron" authorityYear="1941" box="[811,1212,1390,1416]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B96AAA7EFFC8FFFCBF70FA9609A10185" bold="true" box="[811,1212,1390,1416]" pageId="8" pageNumber="31">
<emphasis id="B96AAA7EFFC8FFFCBF70FA960EE50185" bold="true" box="[811,1016,1390,1416]" italics="true" pageId="8" pageNumber="31">Scopaeus mixtus</emphasis>
<bibRefCitation id="EF8F0B9DFFC8FFFCB85BFA9609A10185" author="Cameron, M." box="[1024,1212,1390,1416]" pageId="8" pageNumber="31" pagination="203 - 230" refId="ref31151" refString="Cameron, M. (1941): New Species of Staphylinidae (Col.) from Borneo. - The Entomologist's Monthly Magazine 77 (Fourth Series, Vol. 2): 203 - 230." type="journal article" year="1941">Cameron, 1941</bibRefCitation>
</emphasis>
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<subSubSection id="C30425E7FFC8FFFCBF70FA7709AC01A7" box="[811,1201,1423,1450]" pageId="8" pageNumber="31" type="description">
<paragraph id="8BA1766CFFC8FFFCBF70FA7709AC01A7" blockId="8.[811,1212,1390,1450]" box="[811,1201,1423,1450]" pageId="8" pageNumber="31">
(
<figureCitation id="13256AE9FFC8FFFCBF68FA770E6301A4" box="[819,894,1423,1449]" captionStart="Figures 17" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 17. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="8" pageNumber="31">Figs 6</figureCitation>
,
<figureCitation id="13256AE9FFC8FFFCBFD0FA680EBA01A7" box="[907,935,1424,1450]" captionStart="Figures 2734" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 2734. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="8" pageNumber="31">31</figureCitation>
,
<figureCitation id="13256AE9FFC8FFFCBFE8FA77091B01A4" box="[947,1030,1423,1450]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="8" pageNumber="31">62 64</figureCitation>
,
<figureCitation id="13256AE9FFC8FFFCB849FA68092201A7" box="[1042,1087,1424,1450]" captionStart="Figures 119133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119133. Abdominal sternite VII (126) and abdominal sternite VIII (119125, 127 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="8" pageNumber="31">123</figureCitation>
,
<figureCitation id="13256AE9FFC8FFFCB811FA6809B401A7" box="[1098,1193,1423,1450]" captionStart="Figures 146154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146154. Sperm pump (146154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="8" pageNumber="31">149, 151</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB0E5142FFC8FFFCBF70FA2A083F01E6" box="[811,1314,1489,1516]" pageId="8" pageNumber="31">
<treatmentCitation id="0ABF507DFFC8FFFCBF70FA2A09F801E6" author="Cameron, M." box="[811,1253,1489,1516]" page="229" pageId="8" pageNumber="31" year="1941">
<taxonomicName id="4C1E0DEFFFC8FFFCBF70FA2A09F801E6" ID-CoL="6XYZZ" authority="Cameron, 1941: 229" authorityName="Cameron" authorityPageNumber="229" authorityYear="1941" box="[811,1253,1489,1516]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B96AAA7EFFC8FFFCBF70FA2A0EED01E6" box="[811,1008,1490,1516]" italics="true" pageId="8" pageNumber="31">Scopaeus mixtus</emphasis>
<bibRefCitation id="EF8F0B9DFFC8FFFCBFA3FA2909F801E6" author="Cameron, M." box="[1016,1253,1489,1515]" pageId="8" pageNumber="31" pagination="203 - 230" refId="ref31151" refString="Cameron, M. (1941): New Species of Staphylinidae (Col.) from Borneo. - The Entomologist's Monthly Magazine 77 (Fourth Series, Vol. 2): 203 - 230." type="journal article" year="1941">Cameron, 1941: 229</bibRefCitation>
</taxonomicName>
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, 230.
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</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC8FFFDBF70F9ED0FA005FD" lastPageId="9" lastPageNumber="32" pageId="8" pageNumber="31" type="materials_examined">
<paragraph id="8BA1766CFFC8FFFCBF70F9ED0EC7037B" blockId="8.[788,1399,1556,1975]" pageId="8" pageNumber="31">
<emphasis id="B96AAA7EFFC8FFFCBF70F9ED09770221" bold="true" box="[811,1130,1556,1581]" pageId="8" pageNumber="31">Type specimens examined:</emphasis>
<typeStatus id="54A5C8CEFFC8FFFCB82CF9ED09C60220" box="[1143,1243,1557,1581]" pageId="8" pageNumber="31" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFC8FFFCB8B3F9ED09E10221" box="[1256,1276,1557,1580]" count="1" pageId="8" pageNumber="31" type="male"></specimenCount>
,
<collectingCountry id="F30936FCFFC8FFFCB955F9ED086E0220" box="[1294,1395,1557,1581]" name="Malaysia" pageId="8" pageNumber="31">Malaysia</collectingCountry>
,
<collectingRegion id="49DAB88EFFC8FFFCBF4FF9CE0E470243" box="[788,858,1590,1614]" country="Malaysia" name="Sabah" pageId="8" pageNumber="31">Sabah</collectingRegion>
: Sandakan, leg. Cameron; labeled Type (red edged, round, printed label), Sandakan, / N. Borneo. / Dr. Cameron. (printed), S. / mixtus / TYPE Cam. (handwritten), M. Cameron / Bequest / B.M.1955- 147. (printed),
<typeStatus id="54A5C8CEFFC8FFFCBF9AF941093702DC" box="[961,1066,1721,1745]" pageId="8" pageNumber="31" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFC8FFFCB818F94109E702DC" authorityName="Cameron" authorityYear="1941" box="[1091,1274,1721,1745]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B96AAA7EFFC8FFFCB818F94109E702DC" box="[1091,1274,1721,1745]" italics="true" pageId="8" pageNumber="31">Scopaeus mixtus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFC8FFFCB94FF9410E5502FF" author="Cameron, M." pageId="8" pageNumber="31" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron 1930</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFC8FFFCBF80F922097A02FF" author="Frisch, J. &amp; T. Mainda" box="[987,1127,1754,1778]" pageId="8" pageNumber="31" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
(red, printed) (
<collectionCode id="ED0FEEA9FFC8FFFCB94DF922087602FF" box="[1302,1387,1754,1778]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="8" pageNumber="31">NHML</collectionCode>
).
<typeStatus id="54A5C8CEFFC8FFFCBF4FF9030E9F031E" box="[788,898,1787,1811]" pageId="8" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D18BDE5FFC8FFFCBFD7F9030EA9031F" box="[908,948,1787,1811]" count="2" pageId="8" pageNumber="31" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFC8FFFCBF9BF9030EFA031F" box="[960,999,1787,1811]" count="2" pageId="8" pageNumber="31" type="female">2 ♀</specimenCount>
, same labels as
<typeStatus id="54A5C8CEFFC8FFFCB8CEF90309E9031E" box="[1173,1268,1787,1811]" pageId="8" pageNumber="31" type="holotype">holotype</typeStatus>
except type label, subsequent
<typeStatus id="54A5C8CEFFC8FFFCBF85F8E409200339" box="[990,1085,1820,1844]" pageId="8" pageNumber="31" type="paratype">paratype</typeStatus>
label
<typeStatus id="54A5C8CEFFC8FFFCB8D7F8E409ED0339" box="[1164,1264,1820,1844]" pageId="8" pageNumber="31" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFC8FFFCB954F8E40E460359" authorityName="Cameron" authorityYear="1941" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B96AAA7EFFC8FFFCB954F8E40E460359" italics="true" pageId="8" pageNumber="31">Scopaeus mixtus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFC8FFFCBF2DF8C5090B0358" author="Cameron, M." box="[886,1046,1853,1877]" pageId="8" pageNumber="31" pagination="325 - 348" refId="ref31003" refString="Cameron, M. (1930): Fauna Sumatrensis. Bijdrage No. 67, Staphylinidae (Col.). - Tijdschrift voor Entomologie 73: 325 - 348." type="journal article" year="1930">Cameron 1930</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFC8FFFCB8F0F8C5082A0358" author="Frisch, J. &amp; T. Mainda" box="[1195,1335,1853,1877]" pageId="8" pageNumber="31" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
(red, printed) (
<collectionCode id="ED0FEEA9FFC8FFFCBF22F8A60ED3037B" box="[889,974,1886,1910]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="8" pageNumber="31">NHML</collectionCode>
).
</paragraph>
<paragraph id="8BA1766CFFC8FFFDBF70F8860F930561" blockId="8.[788,1399,1556,1975]" lastBlockId="9.[189,801,241,857]" lastPageId="9" lastPageNumber="32" pageId="8" pageNumber="31">
The type specimen represents a
<typeStatus id="54A5C8CEFFC8FFFCB8CEF88609E9039B" box="[1173,1268,1918,1942]" pageId="8" pageNumber="31" type="holotype">holotype</typeStatus>
by original designation (
<specimenCode id="DBB8DE17FFC8FFFCBFF5F867092303BA" box="[942,1086,1951,1975]" collectionCode="ICZN" pageId="8" pageNumber="31">ICZN 1999</specimenCode>
: Article 73.1.1.), because
<bibRefCitation id="EF8F0B9DFFC9FFFDBCE6FF0A0CB10507" author="Cameron, M." box="[189,428,242,266]" pageId="9" pageNumber="32" pagination="203 - 230" refId="ref31151" refString="Cameron, M. (1941): New Species of Staphylinidae (Col.) from Borneo. - The Entomologist's Monthly Magazine 77 (Fourth Series, Vol. 2): 203 - 230." type="journal article" year="1941">Cameron (1941: 230)</bibRefCitation>
stated Type in my collection. and labeled the specimen accordingly. Four
<typeStatus id="54A5C8CEFFC9FFFDBEEEFEEA0C330546" isOtherTypeStatus="true" pageId="9" pageNumber="32">additional specimens</typeStatus>
from the Cameron collection at NHML are rated as
<typeStatus id="54A5C8CEFFC9FFFDBD4CFEAD0C9C0560" box="[279,385,341,365]" pageId="9" pageNumber="32" type="paratype">paratypes</typeStatus>
and labeled accordingly.
</paragraph>
<paragraph id="8BA1766CFFC9FFFDBC88FE8D0FA005FD" blockId="9.[189,801,241,857]" pageId="9" pageNumber="32">
<emphasis id="B96AAA7EFFC9FFFDBC88FE8D0C720580" bold="true" box="[211,367,373,397]" pageId="9" pageNumber="32">New records:</emphasis>
<materialsCitation id="3B767C31FFC9FFFDBD22FE8D0FA405FD" collectionCode="NHML, MFNB, HECO" collectorName="S Lokan &amp; Chung" country="Malaysia" elevation="300" latitude="9.1996" location="Sandakan" longLatPrecision="7" longitude="3.1997" pageId="9" pageNumber="32" specimenCount="1" stateProvince="Sabah">
<collectingCountry id="F30936FCFFC9FFFDBD22FE8D0CC30580" box="[377,478,373,397]" name="Malaysia" pageId="9" pageNumber="32">Malaysia</collectingCountry>
,
<collectingRegion id="49DAB88EFFC9FFFDBDB7FE8D0F2F0580" box="[492,562,373,397]" country="Malaysia" name="Sabah" pageId="9" pageNumber="32">Sabah</collectingRegion>
:
<location id="8EC120B7FFC9FFFDBE1BFE8D0FB00580" LSID="urn:lsid:plazi:treatment:03B7C77AFFC8FFFEBF70FA960FD30561:8EC120B7FFC9FFFDBE1BFE8D0FB00580" box="[576,685,373,397]" country="Malaysia" latitude="9.1996" longLatPrecision="7" longitude="3.1997" name="Sandakan" pageId="9" pageNumber="32" stateProvince="Sabah">Sandakan</location>
,
<collectorName id="26EB13BAFFC9FFFDBEE0FE8D0E000580" box="[699,797,373,397]" pageId="9" pageNumber="32">S Lokan</collectorName>
,
<geoCoordinate id="EE2A10ABFFC9FFFDBCE6FE6E0C1805A3" box="[189,261,406,430]" degrees="9.1996" direction="north" orientation="latitude" pageId="9" pageNumber="32" precision="5" value="9.1996">9.1996</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFC9FFFDBD4BFE6E0C4A05A3" box="[272,343,406,430]" degrees="3.1997" direction="east" orientation="longitude" pageId="9" pageNumber="32" precision="5" value="3.1997">3.1997</geoCoordinate>
, leg.
<collectorName id="26EB13BAFFC9FFFDBDD4FE6E0CCA05A3" box="[399,471,406,430]" pageId="9" pageNumber="32">Chung</collectorName>
(
<collectionCode id="ED0FEEA9FFC9FFFDBDBEFE6E0F2605A3" box="[485,571,406,430]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="9" pageNumber="32">NHML</collectionCode>
,
<collectionCode id="ED0FEEA9FFC9FFFDBE1DFE6E0F8A05A3" box="[582,663,406,430]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="32">MFNB</collectionCode>
); (04°6350 - 04°7716N, 117°4383 - 117°7031E),
<quantity id="4CE6DB89FFC9FFFDBE92FE4F0E0105C3" box="[713,796,439,463]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="32" unit="m" value="300.0">
<elevation id="0033915FFFC9FFFDBE92FE4F0E0105C3" box="[713,796,439,463]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="32" unit="m" value="300.0">300 m</elevation>
</quantity>
, 5. 6.2012, leg. SAFE Project (
<collectionCode id="ED0FEEA9FFC9FFFDBE52FE200F4B05FD" box="[521,598,472,496]" pageId="9" pageNumber="32">HECO</collectionCode>
,
<collectionCode id="ED0FEEA9FFC9FFFDBE3AFE200FAF05FD" box="[609,690,472,496]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="32">MFNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC9FFFDBC88FE000EB50754" pageId="9" pageNumber="32" type="description">
<paragraph id="8BA1766CFFC9FFFDBC88FE0009600527" blockId="9.[189,801,241,857]" lastBlockId="9.[824,1437,241,857]" pageId="9" pageNumber="32">
<emphasis id="B96AAA7EFFC9FFFDBC88FE000C9D061D" bold="true" box="[211,384,504,528]" pageId="9" pageNumber="32">Redescription:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFC9FFFDBE8DFE000E00061D" box="[726,797,504,528]" captionStart="Figures 17" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 17. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="9" pageNumber="32">Fig. 6</figureCitation>
. Head subquadrate. Antennae with penultimate segment quadrate. Mesotibia slender. Surface of head and elytra matt due to dense setose punctation; pronotum shiny with very fine, sparse punctation; forebody without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body reddish medium brown to dark brown except for elytra gradually blackened from about longitudinal middle towards contrasted, yellow-brown posterior fifth and yellow-brown tip of abdomen; maxillary palpi, antennae and legs yellow brown. Total body length
<quantity id="4CE6DB89FFC9FFFDB951FF0908850504" box="[1290,1432,241,266]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="9" pageNumber="32" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5 2.9 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFC9FFFDBFB5FEEA09640527" box="[1006,1145,274,298]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="9" pageNumber="32" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4 1.5 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFC9FFFDBF14FECB09F906FA" blockId="9.[824,1437,241,857]" pageId="9" pageNumber="32">
Male: Abdominal sternite VII with unmodified posterior margin.AbdominalsterniteVIIIwithconcaveemargination in almost posterior third (
<figureCitation id="13256AE9FFC9FFFDB808FE8D09AE0580" box="[1107,1203,373,397]" captionStart="Figures 119133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119133. Abdominal sternite VII (126) and abdominal sternite VIII (119125, 127 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="9" pageNumber="32">Fig. 123</figureCitation>
). Aedeagus (
<figureCitation id="13256AE9FFC9FFFDB919FE8D08850580" box="[1346,1432,373,397]" captionStart="Figures 2734" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 2734. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="9" pageNumber="32">Figs 31</figureCitation>
,
<figureCitation id="13256AE9FFC9FFFDBF63FE6E0E9F05A3" box="[824,898,406,430]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="9" pageNumber="32">62 64</figureCitation>
) about
<quantity id="4CE6DB89FFC9FFFDBF8EFE6E093B05A0" box="[981,1062,406,430]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="32" unit="mm" value="0.7">0.7 mm</quantity>
long; apical lobes narrow, hooked ventrally with slender, emarginate apices (
<figureCitation id="13256AE9FFC9FFFDB8A3FE4F085605C2" box="[1272,1355,439,463]" captionStart="Figures 2734" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 2734. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="9" pageNumber="32">Figs 31</figureCitation>
,
<figureCitation id="13256AE9FFC9FFFDB90CFE4F086905C2" box="[1367,1396,439,463]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="9" pageNumber="32">62</figureCitation>
), in ventral and dorsal view (
<figureCitation id="13256AE9FFC9FFFDB86DFE2009B105FD" box="[1078,1196,472,496]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="9" pageNumber="32">Figs 63, 64</figureCitation>
) subparallel with acute ends, together only third as wide as phallobase; dorsal lobe as long as apical lobes, strongly projecting from apical lobes ventrally, semicircularly curved dorsad, with conspicuous ventral dilatation evenly tapered towards acute end; lateral lobes subrectangular, somewhat projecting laterad, each with spacious row of few setae (
<figureCitation id="13256AE9FFC9FFFDB8CFFD6509FB06B8" box="[1172,1254,669,693]" captionStart="Figures 2734" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 2734. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="9" pageNumber="32">Figs 31</figureCitation>
,
<figureCitation id="13256AE9FFC9FFFDB8A8FD65082706B8" box="[1267,1338,669,693]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="9" pageNumber="32">62, 63</figureCitation>
); median foramen transverse, short longitudinally, limited distally by strongly sclerotized, transverse ridge.
</paragraph>
<paragraph id="8BA1766CFFC9FFFDBF14FCF80EB50754" blockId="9.[824,1437,241,857]" pageId="9" pageNumber="32">
Female: Sperm pump with long, slender segments (
<figureCitation id="13256AE9FFC9FFFDBF1BFCD80E840735" box="[832,921,800,824]" captionStart="Figures 146154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146154. Sperm pump (146154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="9" pageNumber="32">Fig. 151</figureCitation>
); bursa hyaline with small, apical sclerotization (
<figureCitation id="13256AE9FFC9FFFDBF1BFCB90E800754" box="[832,925,833,857]" captionStart="Figures 146154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146154. Sperm pump (146154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="9" pageNumber="32">Fig. 149</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6126E4FFC9FFFDBCE6F8720EC103BA" ID-DOI="http://doi.org/10.5281/zenodo.10724246" ID-Zenodo-Dep="10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="9" pageNumber="32" startId="9.[189,262,1930,1952]" targetBox="[262,1384,910,1880]" targetPageId="9" targetType="figure">
<paragraph id="8BA1766CFFC9FFFDBCE6F8720EC103BA" blockId="9.[189,1437,1930,1975]" pageId="9" pageNumber="32">
<emphasis id="B96AAA7EFFC9FFFDBCE6F8720C570392" bold="true" box="[189,330,1930,1952]" pageId="9" pageNumber="32">Figures 1416.</emphasis>
Habitus of
<taxonomicName id="4C1E0DEFFFC9FFFDBDECF8730F7A0392" authorityName="Frisch" authorityYear="2022" box="[439,615,1930,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="heronifer">
<emphasis id="B96AAA7EFFC9FFFDBDECF8730F7A0392" box="[439,615,1930,1952]" italics="true" pageId="9" pageNumber="32">Scopaeus heronifer</emphasis>
</taxonomicName>
, paratype, Jawa Barat: Mt Salak (
<emphasis id="B96AAA7EFFC9FFFDBFFBF8720EAA0392" bold="true" box="[928,951,1930,1951]" pageId="9" pageNumber="32">14</emphasis>
);
<taxonomicName id="4C1E0DEFFFC9FFFDBF90F873097F0392" authorityName="Frisch" authorityYear="2022" box="[971,1122,1930,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="diversilobatus">
<emphasis id="B96AAA7EFFC9FFFDBF90F8730ECB03AD" box="[971,982,1931,1952]" italics="true" pageId="9" pageNumber="32">S</emphasis>
.
<emphasis id="B96AAA7EFFC9FFFDBFB9F872097F0392" box="[994,1122,1930,1951]" italics="true" pageId="9" pageNumber="32">diversilobatus</emphasis>
</taxonomicName>
, paratype, Jawa Barat, Sukabumi: Djampang Kulon (
<emphasis id="B96AAA7EFFC9FFFDBD3CF8590C6303BB" bold="true" box="[359,382,1953,1974]" pageId="9" pageNumber="32">15</emphasis>
);
<taxonomicName id="4C1E0DEFFFC9FFFDBDCAF8590F0603BB" authorityName="Frisch" authorityYear="2022" box="[401,539,1953,1975]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="cuspilobatus">
<emphasis id="B96AAA7EFFC9FFFDBDCAF8590C8103BB" box="[401,412,1953,1974]" italics="true" pageId="9" pageNumber="32">S</emphasis>
.
<emphasis id="B96AAA7EFFC9FFFDBDF3F8590F0603BB" box="[424,539,1953,1974]" italics="true" pageId="9" pageNumber="32">cuspilobatus</emphasis>
</taxonomicName>
, paratype, Jawa Barat, Sukabumi: Sirnarasa (
<emphasis id="B96AAA7EFFC9FFFDBFE3F8590ED203BB" bold="true" box="[952,975,1953,1974]" pageId="9" pageNumber="32">16</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C30425E7FFCAFFFEBCF5FF090FD30561" pageId="10" pageNumber="33" type="distribution">
<paragraph id="8BA1766CFFCAFFFEBCF5FF090FD30561" blockId="10.[151,764,241,364]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBCF5FF090C590504" bold="true" box="[174,324,241,265]" pageId="10" pageNumber="33">Distribution:</emphasis>
Examined specimens of
<taxonomicName id="4C1E0DEFFFCAFFFEBECFFF0A0DC30527" authorityName="Cameron" authorityYear="1941" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B96AAA7EFFCAFFFEBECFFF0A0DC30527" italics="true" pageId="10" pageNumber="33">Scopaeus mixtus</emphasis>
</taxonomicName>
originate from
<collectingRegion id="49DAB88EFFCAFFFEBDC3FEEA0CC70527" box="[408,474,274,298]" country="Malaysia" name="Sabah" pageId="10" pageNumber="33">Sabah</collectingRegion>
in the Malaysian part of Borneo. I expect this species to occur in
<collectingRegion id="49DAB88EFFCAFFFEBE06FECB0FC00546" box="[605,733,307,331]" country="Indonesia" name="Kalimantan Utara" pageId="10" pageNumber="33">Kalimantan</collectingRegion>
as well, which is why it is included in this contribution.
</paragraph>
</subSubSection>
</treatment>
</document>