Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species Author Frisch, Johannes Author Narakusumo, Raden Pramesa MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia text Soil Organisms 2022 2022-04-01 95 1 23 73 http://dx.doi.org/10.25674/so95iss1id311 journal article 10.25674/so95iss1id311 2509-9523 10724236 Scopaeus mixtus Cameron, 1941 ( Figs 6 , 31 , 62– 64 , 123 , 149, 151 ) Scopaeus mixtus Cameron, 1941: 229 , 230. Type specimens examined: Holotype ♂ , Malaysia , Sabah : Sandakan, leg. Cameron; labeled ‘Type’ (red edged, round, printed label), ‘Sandakan, / N. Borneo. / Dr. Cameron.’ (printed), ‘S. / mixtus / TYPE Cam.’ (handwritten), ‘M. Cameron / Bequest / B.M.1955- 147.’ (printed), ‘ Holotype / Scopaeus mixtus / Cameron 1930 / label by J. Frisch, 2022 ’ (red, printed) ( NHML ). Paratypes : 2 ♂ , 2 ♀ , same labels as holotype except type label, subsequent paratype label ‘ Paratype / Scopaeus mixtus / Cameron 1930 / label by J. Frisch, 2022 ’ (red, printed) ( NHML ). The type specimen represents a holotype by original designation ( ICZN 1999 : Article 73.1.1.), because Cameron (1941: 230) stated ‘Type in my collection.’ and labeled the specimen accordingly. Four additional specimens from the Cameron collection at NHML are rated as paratypes and labeled accordingly. New records: Malaysia , Sabah : Sandakan , S Lokan , 9.1996 , 3.1997 , leg. Chung ( NHML , MFNB ); (04°63’50’’ - 04°77’16’’N, 117°43’83’’ - 117°70’31’’E), 300 m , 5. – 6.2012, leg. SAFE Project ( HECO , MFNB ) . Redescription: Habitus and coloring as in Fig. 6 . Head subquadrate. Antennae with penultimate segment quadrate. Mesotibia slender. Surface of head and elytra matt due to dense setose punctation; pronotum shiny with very fine, sparse punctation; forebody without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body reddish medium brown to dark brown except for elytra gradually blackened from about longitudinal middle towards contrasted, yellow-brown posterior fifth and yellow-brown tip of abdomen; maxillary palpi, antennae and legs yellow brown. Total body length 2.5 – 2.9 mm ; forebody length 1.4 – 1.5 mm . Male: Abdominal sternite VII with unmodified posterior margin.AbdominalsterniteVIIIwithconcaveemargination in almost posterior third ( Fig. 123 ). Aedeagus ( Figs 31 , 62 – 64 ) about 0.7 mm long; apical lobes narrow, hooked ventrally with slender, emarginate apices ( Figs 31 , 62 ), in ventral and dorsal view ( Figs 63, 64 ) subparallel with acute ends, together only third as wide as phallobase; dorsal lobe as long as apical lobes, strongly projecting from apical lobes ventrally, semicircularly curved dorsad, with conspicuous ventral dilatation evenly tapered towards acute end; lateral lobes subrectangular, somewhat projecting laterad, each with spacious row of few setae ( Figs 31 , 62, 63 ); median foramen transverse, short longitudinally, limited distally by strongly sclerotized, transverse ridge. Female: Sperm pump with long, slender segments ( Fig. 151 ); bursa hyaline with small, apical sclerotization ( Fig. 149 ). Figures 14–16. Habitus of Scopaeus heronifer , paratype, Jawa Barat: Mt Salak ( 14 ); S . diversilobatus , paratype, Jawa Barat, Sukabumi: Djampang Kulon ( 15 ); S . cuspilobatus , paratype, Jawa Barat, Sukabumi: Sirnarasa ( 16 ). Distribution: Examined specimens of Scopaeus mixtus originate from Sabah in the Malaysian part of Borneo. I expect this species to occur in Kalimantan as well, which is why it is included in this contribution.