176 lines
20 KiB
XML
176 lines
20 KiB
XML
<document id="63CA8A20B0D60E45D0DADE338944C148" ID-CLB-Dataset="27392" ID-DOI="10.5852/ejt.2019.512" ID-GBIF-Dataset="85c6241d-5e5a-4022-945f-e207d31458ef" ID-Zenodo-Dep="2628794" ID-ZooBank="urn:lsid:zoobank.org:pub:8DF89CFD-A549-4C69-8FBC-90EC8DEA9A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554393989085" checkinUser="plazi" docAuthor="Darby, Michael" docDate="2019" docId="BE2487B5FF87FFC26D3B79F74C32FEC6" docLanguage="en" docName="ejt-512_darby.pdf.imf" docOrigin="European Journal of Taxonomy 512" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Discheramocephalus nigritus Darby, 2019, sp. nov." docType="treatment" docUuid="9FAADB43-B9A8-4B23-8223-16A129D2D088" docUuidSource="ZooBank" docVersion="9" lastPageNumber="22" masterDocId="421DFFCDFF93FFD46F0C7D524D30FFAA" masterDocTitle="New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London" masterLastPageNumber="50" masterPageNumber="1" pageNumber="21" updateTime="1698718664008" updateUser="ExternalLinkService">
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<mods:titleInfo id="316395D364CBDB706249FF2B8FB5FD37">
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<mods:title id="8724191315BC5D91BDC32F94B67A5633">New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London</mods:title>
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<mods:name id="B2ACB5240B22CC3601BA183B0E93FACE" type="personal">
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<mods:namePart id="44E1C6C7C10D01D474EAA9CB980E8B5D">Darby, Michael</mods:namePart>
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<mods:typeOfResource id="B7CE8189DD4CB211DCB13D90B2666CEB">text</mods:typeOfResource>
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<mods:title id="6D26D8C42A8048827F809B76C9141BBA">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="A2BBD8823F7A4697F158573B7CB65384">
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<mods:date id="A4399D5950DF273C019F3B2C5EEF4D89">2019</mods:date>
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<mods:number id="3D2A98016D03D91D28CD2833B6DE1FFF">2019-04-04</mods:number>
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<mods:number id="793041C4E3D88C4D4B2405612BD29B68">512</mods:number>
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<mods:identifier id="828D48FEB5882645979CCB88200684B5" type="DOI">10.5852/ejt.2019.512</mods:identifier>
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<treatment id="BE2487B5FF87FFC26D3B79F74C32FEC6" ID-DOI="http://doi.org/10.5281/zenodo.5941290" ID-GBIF-Taxon="154555286" ID-Zenodo-Dep="5941290" LSID="urn:lsid:zoobank.org:act:9FAADB43-B9A8-4B23-8223-16A129D2D088" httpUri="http://treatment.plazi.org/id/BE2487B5FF87FFC26D3B79F74C32FEC6" lastPageId="22" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="7E976528FF87FFC06D3B79F749F9FB48" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="363236A3FF87FFC06D3B79F74ECDFB6A" blockId="20.[363,1225,1189,1286]" box="[567,1021,1189,1216]" pageId="20" pageNumber="21">
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<heading id="6D7A81CFFF87FFC06D3B79F74ECDFB6A" box="[567,1021,1189,1216]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
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<taxonomicName id="F18D4D20FF87FFC06D3B79F74EABFB15" ID-CoL="8XZQF" box="[567,923,1189,1216]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nigritus" status="sp. nov.">
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<emphasis id="04F9EAB1FF87FFC06D3B79F74EABFB15" bold="true" box="[567,923,1189,1216]" italics="true" pageId="20" pageNumber="21">Discheramocephalus nigritus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1FCA57CAFF87FFC06CAE79F44ECDFB6A" box="[930,1021,1190,1216]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="363236A3FF87FFC06E67799A49F9FB48" blockId="20.[363,1225,1189,1286]" box="[363,1225,1224,1250]" pageId="20" pageNumber="21">
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<uri id="421C3AA1FF87FFC06E67799A49F9FB48" box="[363,1225,1224,1250]" pageId="20" pageNumber="21">
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urn:lsid:zoobank.org:act:
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<uuid id="422B0C76FF87FFC06D81799A49F9FB48" box="[653,1225,1224,1250]" pageId="20" pageNumber="21">9FAADB43-B9A8-4B23-8223-16A129D2D088</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="7E976528FF87FFC06DFC79BE4E73FAAC" box="[752,835,1260,1286]" pageId="20" pageNumber="21" type="description">
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<paragraph id="363236A3FF87FFC06DFC79BE4E73FAAC" blockId="20.[363,1225,1189,1286]" box="[752,835,1260,1286]" pageId="20" pageNumber="21">
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<figureCitation id="AEB62A26FF87FFC06DFC79BE4E73FAAC" box="[752,835,1260,1286]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="7E976528FF87FFC06FB1787F4C02FA16" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="363236A3FF87FFC06FB1787F4C06FAED" blockId="20.[189,310,1325,1351]" box="[189,310,1325,1351]" pageId="20" pageNumber="21">
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<heading id="6D7A81CFFF87FFC06FB1787F4C06FAED" bold="true" box="[189,310,1325,1351]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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||
<emphasis id="04F9EAB1FF87FFC06FB1787F4C06FAED" bold="true" box="[189,310,1325,1351]" pageId="20" pageNumber="21">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="363236A3FF87FFC06FB1780E4C02FA16" blockId="20.[189,1399,1372,1468]" pageId="20" pageNumber="21">
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Easily distinguished from all other
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<taxonomicName id="F18D4D20FF87FFC06D5A780E4F88FADC" authorityName="Erichson" authorityYear="1845" box="[598,696,1372,1398]" class="Insecta" family="Ptiliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Ptiliidae</taxonomicName>
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in
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<collectingCountry id="4E9A7633FF87FFC06DD7780E4E70FADC" box="[731,832,1372,1398]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
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by the longitudinal foveae on the pronotum, and by the pair of deep sulci on abdominal ventrite VIII (most easily seen in dissections but not showing in
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<figureCitation id="AEB62A26FF87FFC06FB178F04C15FA16" box="[189,293,1442,1468]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="20" pageNumber="21">Fig. 10F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="7E976528FF87FFC06FB178B6481DF986" box="[189,1325,1508,1581]" pageId="20" pageNumber="21" type="etymology">
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<paragraph id="363236A3FF87FFC06FB178B64C74FA54" blockId="20.[189,324,1508,1534]" box="[189,324,1508,1534]" pageId="20" pageNumber="21">
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<heading id="6D7A81CFFF87FFC06FB178B64C74FA54" bold="true" box="[189,324,1508,1534]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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<emphasis id="04F9EAB1FF87FFC06FB178B64C74FA54" bold="true" box="[189,324,1508,1534]" pageId="20" pageNumber="21">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="363236A3FF87FFC06FB17B40481DF986" blockId="20.[189,1325,1554,1581]" box="[189,1325,1554,1581]" pageId="20" pageNumber="21">
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Named after the Latin adjective ‘
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<emphasis id="04F9EAB1FF87FFC06D4D7B414F4FF986" box="[577,639,1555,1580]" italics="true" pageId="20" pageNumber="21">niger</emphasis>
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’, meaning ‘dark’, and referring to the insect's colouration.
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</paragraph>
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</subSubSection>
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<subSubSection id="7E976528FF87FFC06FB17B0649D0F949" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="363236A3FF87FFC06FB17B064C5CF9C7" blockId="20.[189,364,1619,1646]" box="[189,364,1619,1646]" pageId="20" pageNumber="21">
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<heading id="6D7A81CFFF87FFC06FB17B064C5CF9C7" bold="true" box="[189,364,1619,1646]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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<emphasis id="04F9EAB1FF87FFC06FB17B064C5CF9C7" bold="true" box="[189,364,1619,1646]" pageId="20" pageNumber="21">Type material</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="363236A3FF87FFC06FE67BD049D0F949" blockId="20.[189,1399,1666,1763]" pageId="20" pageNumber="21">
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<materialsCitation id="86E53CFEFF87FFC06FE67BD049D0F949" ID-GBIF-Occurrence="2234828132" collectingDate="1978-04-26" collectionCode="BMNH" collectorName="I. Hanski" country="Malaysia" elevation="1250" location="Gunung Mulu National Park" pageId="20" pageNumber="21" specimenCode="BM 1978-524" specimenCount="1" typeStatus="holotype">
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<typeStatus id="E9368801FF87FFC06FE67BD04C6CF936" box="[234,348,1666,1692]" pageId="20" pageNumber="21" type="holotype">
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<emphasis id="04F9EAB1FF87FFC06FE67BD04C6CF936" bold="true" box="[234,348,1666,1692]" pageId="20" pageNumber="21">Holotype</emphasis>
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</typeStatus>
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MALAYSIA-BORNEO •
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<collectingCountry id="4E9A7633FF87FFC06EE17BF74F65F915" box="[493,597,1701,1727]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
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, 4
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<superScript id="C1F89BEBFF87FFC06D627BF64F4BF918" attach="left" box="[622,635,1700,1714]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
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division,
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<location id="33526078FF87FFC06DE17BF7490BF915" LSID="urn:lsid:plazi:treatment:BE2487B5FF87FFC26D3B79F74C32FEC6:33526078FF87FFC06DE17BF7490BF915" box="[749,1083,1701,1727]" country="Malaysia" name="Gunung Mulu National Park" pageId="20" pageNumber="21">Gunung Mulu National Park</location>
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;
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<elevation id="BDA0D190FF87FFC06B497BF74826F915" box="[1093,1302,1701,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="20" pageNumber="21" unit="m" value="1250.0" valueMax="1700.0" valueMin="800.0">
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<quantity id="F1759B46FF87FFC06B497BF749E7F96A" box="[1093,1239,1701,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="20" pageNumber="21" unit="m" value="1250.0" valueMax="1700.0" valueMin="800.0">800–1700 m</quantity>
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a.s.l.
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</elevation>
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;
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<collectingDate id="5277E98BFF87FFC06A2E7BF74DCBF949" pageId="20" pageNumber="21" value="1978-04-26">
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<date id="42331063FF87FFC06A2E7BF74DCBF949" pageId="20" pageNumber="21" value="1978-04-26">26 Apr. 1978</date>
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</collectingDate>
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;
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<collectorName id="9B785375FF87FFC06E0B7B9B4C43F948" box="[263,371,1736,1763]" pageId="20" pageNumber="21">I. Hanski</collectorName>
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leg.,
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<specimenCode id="662B9ED8FF87FFC06EB87B9B4F6CF949" box="[436,604,1736,1763]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="20" pageNumber="21">BM 1978-524</specimenCode>
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;
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<collectingMethod id="EFCC4EB4FF87FFC06D657B9A4FD4F949" box="[617,740,1736,1763]" pageId="20" pageNumber="21">pitfall trap</collectingMethod>
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fish baited, LMR forest slope 4-6;
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<collectionCode id="509CAE66FF87FFC06B8C7B9B49ECF949" box="[1152,1244,1737,1763]" pageId="20" pageNumber="21">BMNH</collectionCode>
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||
.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7E976528FF87FFC26FB17A584C32FEC6" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
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||
<paragraph id="363236A3FF87FFC06FB17A584C7EF88E" blockId="20.[189,334,1802,1828]" box="[189,334,1802,1828]" pageId="20" pageNumber="21">
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<heading id="6D7A81CFFF87FFC06FB17A584C7EF88E" bold="true" box="[189,334,1802,1828]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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||
<emphasis id="04F9EAB1FF87FFC06FB17A584C7EF88E" bold="true" box="[189,334,1802,1828]" pageId="20" pageNumber="21">Description</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="363236A3FF87FFC06FB17A6B4F9AF8F9" blockId="20.[189,682,1848,1875]" box="[189,682,1848,1875]" pageId="20" pageNumber="21">
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SIZE. Habitus (
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||
<figureCitation id="AEB62A26FF87FFC06E6B7A6A4CE4F8F9" box="[359,468,1848,1875]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="20" pageNumber="21">Fig. 10A</figureCitation>
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||
), length
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||
<quantity id="F1759B46FF87FFC06D367A6A4F95F8F9" box="[570,677,1848,1875]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="20" pageNumber="21" unit="mm" value="0.55">0.55 mm</quantity>
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||
.
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||
</paragraph>
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||
<paragraph id="363236A3FF87FFC16FB17A284F17FEC6" blockId="20.[189,944,1914,1940]" lastBlockId="21.[189,1399,267,364]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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COLOUR. Almost black, shining, antennae and legs dusky yellow. HEAD. With a deep transverse fossa behind the eyes extending laterally to the ventral surface and a group of setae in clearly defined foveae behind each eye (
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||
<figureCitation id="AEB62A26FF86FFC16C2D7C7C4EA0FEE3" box="[801,912,302,329]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="21" pageNumber="22">Fig. 10D</figureCitation>
|
||
). Antennomeres III–XI, length
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||
<quantity id="F1759B46FF86FFC16A097C7D4842FEE3" box="[1285,1394,303,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="22" unit="mm" value="0.21">0.21 mm</quantity>
|
||
. Distance across eyes
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||
<quantity id="F1759B46FF86FFC16EBB7C004F13FEC6" box="[439,547,338,364]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="mm" value="0.17">0.17 mm</quantity>
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||
.
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||
</paragraph>
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||
<paragraph id="363236A3FF86FFC16FB17CC64933FE5E" blockId="21.[189,1399,403,500]" pageId="21" pageNumber="22">
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||
PRONOTUM. Length
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<quantity id="F1759B46FF86FFC16E927CC14F3AFE04" box="[414,522,403,430]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="mm" value="0.12">0.12 mm</quantity>
|
||
, width
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||
<quantity id="F1759B46FF86FFC16D537CC14FFBFE04" box="[607,715,403,430]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="22" unit="mm" value="0.19">0.19 mm</quantity>
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||
, with a pair of longitudinal foveae on the disk and another pair close to the lateral margins, in between these two much smaller foveae close to the posterior margin on each side (
|
||
<figureCitation id="AEB62A26FF86FFC16E527C8B4CF9FE5E" box="[350,457,473,500]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="21" pageNumber="22">Fig. 10B</figureCitation>
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||
). Scutellum with a sharp central keel (
|
||
<figureCitation id="AEB62A26FF86FFC16C877C8B4EC6FE5E" box="[907,1014,473,500]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="21" pageNumber="22">Fig. 10E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="363236A3FF86FFC16FB17F4E4F9CFD9C" blockId="21.[189,684,539,566]" box="[189,684,539,566]" pageId="21" pageNumber="22">
|
||
ELYTRA. Length
|
||
<quantity id="F1759B46FF86FFC16E767F494CD6FD9C" box="[378,486,539,566]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="21" pageNumber="22" unit="mm" value="0.33">0.33 mm</quantity>
|
||
, width
|
||
<quantity id="F1759B46FF86FFC16D377F494F97FD9C" box="[571,679,539,566]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="21" pageNumber="22" unit="mm" value="0.27">0.27 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="363236A3FF86FFC16FB17F0F495EFCA9" blockId="21.[189,1399,604,772]" pageId="21" pageNumber="22">VENTRUM. Proventrum depressed in front of procoxae and with raised angular humeri, extending posteriorly. Mesoventrum with a broad strongly reticulated medial keel and concave lateral margins before posterior termination, keels present extending to sides of mesocoxae, mesocoxae separated by their width, posterior margins obliterated. Metaventrum with scattered pubescence, anterior margins extending in form of a lobe between the mesocoxae, metacoxae almost adjacent.</paragraph>
|
||
<paragraph id="363236A3FF86FFC26FB17E794C32FEC6" blockId="21.[189,1399,811,872]" lastBlockId="22.[189,1398,267,364]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
ABDOMEN. Depressed laterally apparently for reception of metatibiae and tarsi, first visible segment with a serrated posterior margin, ventrite VIII with a pair of deep sulci. Pygidium with ± 10 spines and two coarse setae near the anterior corners (
|
||
<figureCitation id="AEB62A26FF85FFC26D8C7C594FD8FE8F" box="[640,744,267,293]" captionStart="Fig" captionStartId="21.[189,232,1951,1977]" captionTargetBox="[189,1398,973,1897]" captionTargetId="figure@21.[189,1399,973,1897]" captionTargetPageId="21" captionText="Fig. 10. Discheramocephalus nigritus sp. nov. A. Habitus. B. Pronotum ×625. C. Back of head ×600. D. Fovea behind eye ×1800. E. Scutellum ×1500. F. Pygidium." httpUri="https://zenodo.org/record/2628816/files/figure.png" pageId="22" pageNumber="23">Fig. 10F</figureCitation>
|
||
). Wings of usual ptiliid
|
||
<typeStatus id="E9368801FF85FFC26B0F7C5E4909FE8C" box="[1027,1081,268,294]" pageId="22" pageNumber="23">type</typeStatus>
|
||
. Dissection failed to reveal any genitalia to determine sex, but size of eyes suggests male (females in this genus usually have smaller eyes).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |