treatments-xml/data/0E/68/87/0E6887CCC804FFCBFF3FFDB12495460A.xml

<document id="E9C8678D612B5F641235723ED0066424" ID="10.11646/zootaxa.4127.3.9" ID-CLB-Dataset="38722" ID-DOI="10.11646/zootaxa.4127.3.9" ID-GBIF-Dataset="d5a26107-de9b-461f-aff9-90c8e2ec9812" ID-ISSN="1175-5326" ID-Zenodo-Dep="260923" ID-ZooBank="F7E466CB-8EF0-468F-AEF6-542522233FB6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466752126263" checkinUser="plazi" docAuthor="Dronen, Norman O., Blend, Charles K., Khalifa, Refaat M. A., Mohamadain, Hoda S. &amp; Karar, Yasser F. M." docDate="2016" docId="0E6887CCC804FFCBFF3FFDB12495460A" docLanguage="en" docName="zootaxa.4127.3.9.pdf" docOrigin="Zootaxa 4127 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelopscreadium aegyptense Dronen, Blend, Khalifa, Mohamadain &amp; Karar, 2016, n. sp." docType="treatment" docUuid="877112D5-C451-41F4-9A69-994C5A7C90BB" docUuidSource="ZooBank" docVersion="8" lastPageNumber="575" masterDocId="F251FFB4C807FFC2FFA8FF95271C474A" masterDocTitle="Pelopscreadium aegyptense n. gen., n. sp. and Pelopscreadium spongiosum (Bray &amp; Cribb, 1998) n. comb., (Digenea: Lepocreadiidae), each from disjunct populations of the Yellow boxfish, Ostracion cubicus Linnaeus (Ostraciidae)" masterLastPageNumber="578" masterPageNumber="567" pageNumber="570" updateTime="1698663160537" updateUser="ExternalLinkService">
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<mods:title id="47A6EE476CBF8CE7AE0919ED234B2793">Pelopscreadium aegyptense n. gen., n. sp. and Pelopscreadium spongiosum (Bray &amp; Cribb, 1998) n. comb., (Digenea: Lepocreadiidae), each from disjunct populations of the Yellow boxfish, Ostracion cubicus Linnaeus (Ostraciidae)</mods:title>
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<mods:namePart id="E5DAE47A5C55917AE339577F5E58AFD8">Dronen, Norman O.</mods:namePart>
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<mods:namePart id="7E8A065D06BCF5FF32B0699408AA05C0">Blend, Charles K.</mods:namePart>
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<mods:namePart id="12D62B81FC19F600271F7E51B6B3EC41">Khalifa, Refaat M. A.</mods:namePart>
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<mods:namePart id="619E3B6292D5FEAC99AE154E46CE7682">Mohamadain, Hoda S.</mods:namePart>
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<mods:namePart id="50EA4B89480E5ECB08059DD68F5C84F4">Karar, Yasser F. M.</mods:namePart>
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<treatment id="0E6887CCC804FFCBFF3FFDB12495460A" ID-DOI="http://doi.org/10.5281/zenodo.5612500" ID-GBIF-Taxon="127625127" ID-Zenodo-Dep="5612500" LSID="urn:lsid:zoobank.org:act:877112D5-C451-41F4-9A69-994C5A7C90BB" httpUri="http://treatment.plazi.org/id/0E6887CCC804FFCBFF3FFDB12495460A" lastPageId="9" lastPageNumber="575" pageId="3" pageNumber="570">
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<paragraph id="867E36DAC804FFC1FF3FFDB125314574" blockId="3.[151,557,548,607]" box="[151,557,548,574]" pageId="3" pageNumber="570">
<heading id="DD3681B6C804FFC1FF3FFDB125314574" bold="true" box="[151,557,548,574]" fontSize="11" level="1" pageId="3" pageNumber="570" reason="1">
<emphasis id="B4B5EAC8C804FFC1FF3FFDB125314574" bold="true" box="[151,557,548,574]" pageId="3" pageNumber="570">
<taxonomicName id="41C14D59C804FFC1FF3FFDB126F84574" box="[151,484,548,574]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="570" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C804FFC1FF3FFDB126F84574" bold="true" box="[151,484,548,574]" italics="true" pageId="3" pageNumber="570">Pelopscreadium aegyptense</emphasis>
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<taxonomicNameLabel id="AF8657B3C804FFC1FE43FDB125314574" box="[491,557,548,574]" pageId="3" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="867E36DAC804FFC1FF3FFDD3260E4515" blockId="3.[151,557,548,607]" box="[151,274,582,607]" pageId="3" pageNumber="570">
(
<figureCitation id="1EFA2A5FC804FFC1FF37FDD326154515" box="[159,265,582,607]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURE 4" captionStartId-0="7.[151,264,1840,1862]" captionStartId-1="8.[151,250,789,811]" captionTargetBox-0="[227,1359,195,1745]" captionTargetBox-1="[400,1249,184,751]" captionTargetId-0="figure@7.[167,1405,135,1808]" captionTargetId-1="figure@8.[151,1436,175,784]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 1 – 3. Pelopscreadium aegyptense n. sp. (Lepocreadiidae) from the yellow boxfish, Ostracion cubicus Linnaeus (Ostraciidae), from the Red Sea off Egypt. 1. Adult specimen, ventral view. 2. Pharynx, lateral view. 3. Details of the female genital complex, ventral view. Abbreviations: C, ceca; D, oviduct; DL, distinct, well-differentiated pre-oral layer; E, external seminal vesicle; G, genital pore; I, internal seminal vesicle; O, ovary; OS, esophagus; OT, oötype; P, sponge-like pelops (“ shoulder pads ”); PP, prepharynx; R, seminal receptacle; T, testis; U, uterus; V, vitelline follicles; VD, vitelline ducts." captionText-1="FIGURE 4. Comparison of the male terminal genitalia of Pelopscreadium aegyptense n. sp. (left side) vs Pelopscreadium spongiosum n. gen., n. comb. (right side). Illustration of P. spongiosum n. gen., n. comb. was adapted from figure 20 of Bray &amp; Cribb (1998). Abbreviations: E, external seminal vesicle; ED, ejaculatory duct; G, genital pore; I, internal seminal vesicle; PARS, pars prostatica; U, uterus." httpUri-0="https://zenodo.org/record/260924/files/figure.png" httpUri-1="https://zenodo.org/record/260925/files/figure.png" pageId="3" pageNumber="570">Figs. 1–4</figureCitation>
)
</paragraph>
<paragraph id="867E36DAC804FFC1FF3FFD1B239745ED" blockId="3.[151,1437,654,2011]" box="[151,1163,654,679]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF3FFD1B260845ED" bold="true" box="[151,276,654,679]" pageId="3" pageNumber="570">Type-host.</emphasis>
Yellow boxfish, 
<taxonomicName id="41C14D59C804FFC1FE7BFD1A241045ED" authority="Linnaeus" authorityName="Linnaeus" box="[467,780,654,679]" class="Actinopterygii" family="Ostraciidae" genus="Ostracion" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="570" phylum="Chordata" rank="species" species="cubicus">
<emphasis id="B4B5EAC8C804FFC1FE7BFD1A25BC45ED" box="[467,672,654,679]" italics="true" pageId="3" pageNumber="570">Ostracion cubicus</emphasis>
Linnaeus
</taxonomicName>
(
<taxonomicName id="41C14D59C804FFC1FCB3FD1B24EC45ED" box="[795,1008,654,679]" class="Actinopterygii" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="570" phylum="Chordata" rank="order">Tetraodontiformes</taxonomicName>
: 
<taxonomicName id="41C14D59C804FFC1FC53FD1B239C45ED" box="[1019,1152,654,679]" class="Actinopterygii" family="Ostraciidae" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="570" phylum="Chordata" rank="family">Ostraciidae</taxonomicName>
).
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFD2724D645A5" blockId="3.[151,1437,654,2011]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFD2726744581" bold="true" box="[199,360,690,715]" pageId="3" pageNumber="570">Type-locality.</emphasis>
Northern Red Sea, off Sharm El-Naga, Makadi Bay, Southern Hurghada, 
<collectingCountry id="FED6764AC804FFC1FB14FD2623E34581" box="[1212,1279,691,715]" name="Egypt" pageId="3" pageNumber="570">Egypt</collectingCountry>
(
<geoCoordinate id="E3F5501DC804FFC1FAB8FD27228B4581" box="[1296,1431,690,715]" direction="north" orientation="latitude" pageId="3" pageNumber="570" precision="9" value="26.919333">26°55.16’N</geoCoordinate>
, 
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– 
<geoCoordinate id="E3F5501DC804FFC1FE8EFD4326B045A5" box="[294,428,726,751]" direction="north" orientation="latitude" pageId="3" pageNumber="570" precision="9" value="26.893167">26°53.59’N</geoCoordinate>
, 
<geoCoordinate id="E3F5501DC804FFC1FE1FFD43252645A5" box="[439,570,726,751]" direction="east" orientation="longitude" pageId="3" pageNumber="570" precision="9" value="33.9915">33°59.49’E</geoCoordinate>
, depth = 
<quantity id="41399B3FC804FFC1FD0AFD43240A45A4" box="[674,790,726,751]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="3" pageNumber="570" unit="m" value="1.5" valueMax="2.5" valueMin="0.5">0.5–2.5 m</quantity>
; 
<date id="F27F101AC804FFC1FC89FD4324A245A5" box="[801,958,726,751]" pageId="3" pageNumber="570" value="2012-05-26">26/May/2012</date>
).
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFD6F255A4459" blockId="3.[151,1437,654,2011]" box="[199,582,762,787]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFD6F26954459" bold="true" box="[199,393,762,787]" pageId="3" pageNumber="570">Site of infection.</emphasis>
Lower intestine.
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFC8B26554411" blockId="3.[151,1437,654,2011]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFC8B266B447D" bold="true" box="[199,375,798,823]" pageId="3" pageNumber="570">Type-material.</emphasis>
BM(NH) 
<typeStatus id="597A8878C804FFC1FDAFFC8B2575447D" box="[519,617,798,823]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
2016.4.22.1, BM(NH) 
<typeStatus id="597A8878C804FFC1FC32FC8A231A447D" box="[922,1030,799,823]" pageId="3" pageNumber="570" type="paratype">paratypes</typeStatus>
2016.4.22.2, BM(NH) vouchers 2016.4.22.3-10.
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFCF325E84435" blockId="3.[151,1437,654,2011]" box="[199,756,870,895]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFCF326584435" bold="true" box="[199,324,870,895]" pageId="3" pageNumber="570">Pevalence.</emphasis>
2 of 8 host specimens (25% infected).
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFC1F259944E9" blockId="3.[151,1437,654,2011]" box="[199,645,906,931]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFC1F262A44E9" bold="true" box="[199,310,906,931]" pageId="3" pageNumber="570">Intensity.</emphasis>
13–17 worms/host specimen.
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFC3B26E8448D" blockId="3.[151,1437,654,2011]" box="[199,500,942,967]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFC3B2663448D" bold="true" box="[199,383,942,967]" pageId="3" pageNumber="570">Mean intensity.</emphasis>
15 (30/2).
</paragraph>
<paragraph id="867E36DAC804FFC1FF6FFC4725BE44A1" blockId="3.[151,1437,654,2011]" box="[199,674,978,1003]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFC47250B44A1" bold="true" box="[199,535,978,1003]" pageId="3" pageNumber="570">Relative density/abundance.</emphasis>
3.75 (30/8).
</paragraph>
</subSubSection>
<subSubSection id="CEDB6551C804FFC1FF6FFC6326CB4379" pageId="3" pageNumber="570" type="etymology">
<paragraph id="867E36DAC804FFC1FF6FFC6326CB4379" blockId="3.[151,1437,654,2011]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFC6326524345" bold="true" box="[199,334,1014,1039]" pageId="3" pageNumber="570">Etymology.</emphasis>
The species designation “ 
<taxonomicName id="41C14D59C804FFC1FDDBFC6225F34344" box="[627,751,1015,1038]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="570" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C804FFC1FDDBFC6225F34344" box="[627,751,1015,1038]" italics="true" pageId="3" pageNumber="570">aegyptense</emphasis>
</taxonomicName>
” refers to the geographical location where this new digenean was collected, i.e. off 
<collectingCountry id="FED6764AC804FFC1FE25FB8E26C84379" box="[397,468,1051,1075]" name="Egypt" pageId="3" pageNumber="570">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CEDB6551C804FFC4FF6FFBAB25194028" lastPageId="6" lastPageNumber="573" pageId="3" pageNumber="570" type="description">
<paragraph id="867E36DAC804FFC1FF6FFBAB2652406D" blockId="3.[151,1437,654,2011]" pageId="3" pageNumber="570">
<emphasis id="B4B5EAC8C804FFC1FF6FFBAB264D431D" bold="true" box="[199,337,1086,1111]" pageId="3" pageNumber="570">Description</emphasis>
. [Based on 18 mature specimens. Measurements, morphometric percentages and morphometric ratios are given in 
<tableCitation id="CB430361C804FFC1FED1FBF726CC4331" box="[377,464,1122,1147]" captionStart="TABLE 1" captionStartId="4.[151,239,368,391]" captionTargetBox="[159,1427,486,1986]" captionTargetPageId="4" captionText="TABLE 1. Measurements, morphometric percentages and morphometric ratios of Pelopscreadium spongiosum (Bray &amp; Cribb, 1998) n. gen., n. comb. and Pelopscreadium aegyptense n. sp. from the yellow boxfish, Ostracion cubicus Linnaeus, 1758 (Tetraodontiformes: Ostraciidae), with holotype data followed by the ranges and means in parentheses." httpUri="http://table.plazi.org/id/D2BE6652C803FFC6FF3FFEE522974680" pageId="3" pageNumber="570" tableUuid="D2BE6652C803FFC6FF3FFEE522974680">Table 1</tableCitation>
.] Body elongate-oval, with maximum width in middle third of body. Anterior end broadly rounded; posterior end less broadly rounded; pre-oral lobe present. Forebody gradually narrows near anterior end; remnants of cercarial eyespots often obvious in first third of forebody. Hindbody wider than forebody, gradually narrows in posterior third of body. Large internal patches of vacuolated cells form sponge-like pelops (“shoulder pads”) laterally between anterior extremity and level 2/3 to 3/4 length of forebody. Tegument densely spined anteriorly; spines short, generally scale-like, becoming less dense posterior to sponge-like pelops, sparsely distributed from posterior extent of sponge-like pelops to near posterior extremity. Distinct, narrow, welldifferentiated layer (band) of cells immediately inside anterior margin of body extends posteriorly on both sides two-thirds length of forebody. Oral sucker spherical to transversely elongate-oval, distinctly sub-terminal, somewhat embedded in parenchyma, partially covered by tegument, unspecialized with conspicuous central mouth opening. Ventral sucker sessile, spherical to ovoid, immediately pre-equatorial, occasionally equatorial, unspecialized, smaller than oral sucker. Prepharynx short, broad and either entirely outside in 89% [n=16] or retracted within posterior concavity of oral sucker in 11% [n=2] of specimens. Pharynx muscular, subspherical, narrower than oral sucker; 5 dorsal papillae (line of 3 large middle papillate projections with 1 somewhat smaller papillate projection that abuts each end of row on dorsal rim of anterior pharyngeal aperture) and 3 ventral papillate projections on ventral side of margin of aperture (like petals of flower) with 2 extensions of pharyngeal wall inserted between dorsal and ventral rows of papillae. Esophagus distinct, short, straight, longer than prepharynx. Intestinal bifurcation in posterior forebody. Ceca 2, simple, equal in length, widest just posterior to ventral sucker, somewhat arcuate, pass along lateral margins of worm to posterior end, where they abruptly terminate near body wall; ani absent.
</paragraph>
<paragraph id="867E36DAC804FFC6FF6FF8A6230E460A" blockId="3.[151,1437,654,2011]" lastBlockId="4.[151,1437,151,320]" lastPageId="4" lastPageNumber="571" pageId="3" pageNumber="570">
Testes 2, transversely elongate, slightly to deeply lobed, median, tandem, contiguous, in posterior third of body. Distance from ventral sucker to anterior testis relatively long. External seminal vesicle long, tubular, sinuous throughout, larger than internal seminal vesicle, reaches from posterior aspect of sinistral rim of ventral sucker or slightly dorsal to it to about level of ovary. “ 
<taxonomicName id="41C14D59C804FFC1FD32F80B241140FD" box="[666,781,1950,1975]" class="Trematoda" family="Lepocreadiidae" genus="Opechona" kingdom="Animalia" order="Plagiorchiida" pageId="3" pageNumber="570" phylum="Platyhelminthes" rank="genus">
<emphasis id="B4B5EAC8C804FFC1FD32F80B241140FD" box="[666,781,1950,1975]" italics="true" pageId="3" pageNumber="570">Opechona</emphasis>
</taxonomicName>
- 
<typeStatus id="597A8878C804FFC1FCBDF80A245540FD" box="[789,841,1951,1975]" pageId="3" pageNumber="570">type</typeStatus>
” cirrus-sac (
<bibRefCitation id="E2504B2BC804FFC1FC72F80A234B40FD" author="Bray" box="[986,1111,1950,1975]" pageId="3" pageNumber="570" refString="Bray, R. A. (2005) Family Lepocreadiidae Odhner, 1905. In: Jones, A., Bray, R. A. &amp; Gibson, D. I. (Eds.), Keys to the Trematoda. Vol. 2. CABI Publishing and the Natural History Museum, Wallingford, pp. 545 - 602." type="book chapter" year="2005">Bray 2005</bibRefCitation>
; 
<bibRefCitation id="E2504B2BC804FFC1FBCCF80A225440FD" author="Bray" box="[1124,1352,1950,1975]" pageId="3" pageNumber="570" refString="Bray, R. A. &amp; Cribb, T. H. (2012) Reorganisation of the superfamily Lepocreadioidea Odhner, 1905 based on an inferred molecular phylogeny. Systematic Parasitology, 83, 169 - 177." type="journal article" year="2012">Bray &amp; Cribb 2012</bibRefCitation>
), large, claviform, sinistral to mid-line, extends posteriorly from short distance anterior to ventral sucker to short distance posterior to it, encloses ejaculatory duct, pars prostatica and internal seminal vesicle. Internal seminal vesicle oval, less extensive than pars prostatica, in posterior region of cirrus-sac. Pars prostatica conspicuous, thick-walled, almost straight, elongate-oval, much longer than ejaculatory duct, lined with anuclear cell-like bodies (anuclear blebs). Ejaculatory duct short, distinct. Genital atrium distinct. Genital pore immediately post-bifurcal, ventrosubmedian, sinistral, in forebody, overlaps left cecum or opens just medial to it.
</paragraph>
<caption id="D2BE6652C803FFC6FF3FFEE522974680" ID-Table-UUID="D2BE6652C803FFC6FF3FFEE522974680" httpUri="http://table.plazi.org/id/D2BE6652C803FFC6FF3FFEE522974680" pageId="4" pageNumber="571" targetBox="[159,1427,486,1986]" targetIsTable="true" targetPageId="4">
<paragraph id="867E36DAC803FFC6FF3FFEE522974680" blockId="4.[151,1436,367,458]" pageId="4" pageNumber="571">
<emphasis id="B4B5EAC8C803FFC6FF3FFEE5261646CD" bold="true" box="[151,266,368,391]" pageId="4" pageNumber="571">TABLE 1.</emphasis>
Measurements, morphometric percentages and morphometric ratios of 
<taxonomicName id="41C14D59C803FFC6FBA1FEFA263846ED" authority="Bray &amp; Cribb, 1998" authorityName="Bray &amp; Cribb" authorityYear="1998" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="species" species="spongiosum" status="gen. nov.">
<emphasis id="B4B5EAC8C803FFC6FBA1FEFA222046CD" box="[1033,1340,367,391]" italics="true" pageId="4" pageNumber="571">Pelopscreadium spongiosum</emphasis>
(Bray &amp; Cribb, 1998)
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C803FFC6FE87FE05266246ED" box="[303,382,400,423]" pageId="4" pageNumber="571" rank="genus">n. gen.</taxonomicNameLabel>
, n. comb. and 
<taxonomicName id="41C14D59C803FFC6FD85FE1A244746ED" box="[557,859,399,423]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C803FFC6FD85FE1A244746ED" box="[557,859,399,423]" italics="true" pageId="4" pageNumber="571">Pelopscreadium aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C803FFC6FCCEFE0524B446ED" box="[870,936,400,423]" pageId="4" pageNumber="571" rank="species">
<emphasis id="B4B5EAC8C803FFC6FCCEFE0524B446ED" bold="true" box="[870,936,400,423]" pageId="4" pageNumber="571">n. sp.</emphasis>
</taxonomicNameLabel>
from the yellow boxfish, 
<taxonomicName id="41C14D59C803FFC6FB7CFE0526274680" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Actinopterygii" family="Ostraciidae" genus="Ostracion" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="species" species="cubicus">
<emphasis id="B4B5EAC8C803FFC6FB7CFE05228746ED" box="[1236,1435,399,423]" italics="true" pageId="4" pageNumber="571">Ostracion cubicus</emphasis>
Linnaeus, 1758
</taxonomicName>
(
<taxonomicName id="41C14D59C803FFC6FEE1FE2625084680" box="[329,532,435,458]" class="Actinopterygii" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="order">Tetraodontiformes</taxonomicName>
: 
<taxonomicName id="41C14D59C803FFC6FDB5FE2625874680" box="[541,667,435,458]" class="Actinopterygii" family="Ostraciidae" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="family">Ostraciidae</taxonomicName>
), with holotype data followed by the ranges and means in parentheses.
</paragraph>
</caption>
<paragraph id="867E36DAC803FFC6FF37FE73231C4088" pageId="4" pageNumber="571">
<table id="F4C1C47AC803003DFF37FE73228F4088" box="[159,1427,486,1986]" colsContinueIn="5.[159,1427,201,1981]" gridcols="3" gridrows="21" pageId="4" pageNumber="571">
<tr id="38F13498C803003DFF37FE73228F4550" box="[159,1427,486,538]" gridrow="0" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FE7326FE4550" box="[159,482,486,538]" gridcol="0" gridrow="0" pageId="4" pageNumber="571">Species/Feature</th>
<th id="7B205DE4C803003DFDE2FE7324454550" box="[586,857,486,538]" gridcol="1" gridrow="0" pageId="4" pageNumber="571">
<taxonomicName id="41C14D59C803FFC6FDE2FE7225C846B6" box="[586,724,487,508]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C803FFC6FDE2FE72254B46B6" box="[586,599,487,508]" italics="true" pageId="4" pageNumber="571">P</emphasis>
. 
<emphasis id="B4B5EAC8C803FFC6FDCCFE7225C846B6" box="[612,724,487,508]" italics="true" pageId="4" pageNumber="571">aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C803FFC6FD72FE72240D46B6" box="[730,785,487,508]" pageId="4" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
n=18
</th>
<th id="7B205DE4C803003DFC3EFE73228F4550" box="[918,1427,486,538]" gridcol="2" gridrow="0" pageId="4" pageNumber="571">
<taxonomicName id="41C14D59C803FFC6FC3EFE72221B46B6" authority="Bray &amp; Cribb, 1998" authorityName="Bray &amp; Cribb" authorityYear="1998" box="[918,1287,486,508]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="571" phylum="Chordata" rank="species" species="spongiosum" status="gen. nov.">
<emphasis id="B4B5EAC8C803FFC6FC3EFE7224BE46B6" box="[918,930,487,508]" italics="true" pageId="4" pageNumber="571">P</emphasis>
. 
<emphasis id="B4B5EAC8C803FFC6FC07FE72233446B6" box="[943,1064,487,508]" italics="true" pageId="4" pageNumber="571">spongiosum</emphasis>
(Bray &amp; Cribb, 1998)
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C803FFC6FAA5FE72224846B6" box="[1293,1364,487,508]" pageId="4" pageNumber="571" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp. n=2
</th>
</tr>
<tr id="38F13498C803003DFF37FDBB228F4528" box="[159,1427,558,610]" gridrow="1" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FDBB26FE4528" box="[159,482,558,610]" gridcol="0" gridrow="1" pageId="4" pageNumber="571">Locality</th>
<td id="7B205DE4C803003DFDE2FDBB24454528" box="[586,857,558,610]" gridcol="1" gridrow="1" pageId="4" pageNumber="571">Red Sea, Macady Bay, Off Hurghada, Egypt</td>
<td id="7B205DE4C803003DFC3EFDBB228F4528" box="[918,1427,558,610]" gridcol="2" gridrow="1" pageId="4" pageNumber="571">Off Lizard Island, Queensland, Australia</td>
</tr>
<tr id="38F13498C803003DFF37FDE3228F45E0" box="[159,1427,630,682]" gridrow="2" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FDE326FE45E0" box="[159,482,630,682]" gridcol="0" gridrow="2" pageId="4" pageNumber="571">Body length</th>
<td id="7B205DE4C803003DFDE2FDE3244545E0" box="[586,857,630,682]" gridcol="1" gridrow="2" pageId="4" pageNumber="571">1,555 (1,160–2,642; 2,027)</td>
<td id="7B205DE4C803003DFC3EFDE3228F45E0" box="[918,1427,630,682]" gridcol="2" gridrow="2" pageId="4" pageNumber="571">(1,352–1,412)</td>
</tr>
<tr id="38F13498C803003DFF37FD2B228F45B8" box="[159,1427,702,754]" gridrow="3" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FD2B26FE45B8" box="[159,482,702,754]" gridcol="0" gridrow="3" pageId="4" pageNumber="571">Body width</th>
<td id="7B205DE4C803003DFDE2FD2B244545B8" box="[586,857,702,754]" gridcol="1" gridrow="3" pageId="4" pageNumber="571">1,312 (775–1,356; 1,014)</td>
<td id="7B205DE4C803003DFC3EFD2B228F45B8" box="[918,1427,702,754]" gridcol="2" gridrow="3" pageId="4" pageNumber="571">(438–489)</td>
</tr>
<tr id="38F13498C803003DFF37FC93228F4470" box="[159,1427,774,826]" gridrow="4" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FC9326FE4470" box="[159,482,774,826]" gridcol="0" gridrow="4" pageId="4" pageNumber="571">Forebody length</th>
<td id="7B205DE4C803003DFDE2FC9324454470" box="[586,857,774,826]" gridcol="1" gridrow="4" pageId="4" pageNumber="571">996 (581–1,054; 808)</td>
<td id="7B205DE4C803003DFC3EFC93228F4470" box="[918,1427,774,826]" gridcol="2" gridrow="4" pageId="4" pageNumber="571">(467–487)</td>
</tr>
<tr id="38F13498C803003DFF37FCDB228F44CF" box="[159,1427,846,901]" gridrow="5" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FCDB26FE44CF" box="[159,482,846,901]" gridcol="0" gridrow="5" pageId="4" pageNumber="571">[Percentage of body length]</th>
<td id="7B205DE4C803003DFDE2FCDB244544CF" box="[586,857,846,901]" gridcol="1" gridrow="5" pageId="4" pageNumber="571">64 [25–64; 39]</td>
<td id="7B205DE4C803003DFC3EFCDB228F44CF" box="[918,1427,846,901]" gridcol="2" gridrow="5" pageId="4" pageNumber="571">[34–35]1</td>
</tr>
<tr id="38F13498C803003DFF37FC0D228F4487" box="[159,1427,920,973]" gridrow="6" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FC0D26FE4487" box="[159,482,920,973]" gridcol="0" gridrow="6" pageId="4" pageNumber="571">Body spines length</th>
<td id="7B205DE4C803003DFDE2FC0D24454487" box="[586,857,920,973]" gridcol="1" gridrow="6" pageId="4" pageNumber="571">10–13 (8–13; 11)</td>
<td id="7B205DE4C803003DFC3EFC0D228F4487" box="[918,1427,920,973]" gridcol="2" gridrow="6" pageId="4" pageNumber="571">(10)</td>
</tr>
<tr id="38F13498C803003DFF37FC75228F435F" box="[159,1427,992,1045]" gridrow="7" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FC7526FE435F" box="[159,482,992,1045]" gridcol="0" gridrow="7" pageId="4" pageNumber="571">Oral sucker length</th>
<td id="7B205DE4C803003DFDE2FC752445435F" box="[586,857,992,1045]" gridcol="1" gridrow="7" pageId="4" pageNumber="571">304 (133–356; 250)</td>
<td id="7B205DE4C803003DFC3EFC75228F435F" box="[918,1427,992,1045]" gridcol="2" gridrow="7" pageId="4" pageNumber="571">(134–148)</td>
</tr>
<tr id="38F13498C803003DFF37FBBD228F4317" box="[159,1427,1064,1117]" gridrow="8" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FBBD26FE4317" box="[159,482,1064,1117]" gridcol="0" gridrow="8" pageId="4" pageNumber="571">Oral sucker width</th>
<td id="7B205DE4C803003DFDE2FBBD24454317" box="[586,857,1064,1117]" gridcol="1" gridrow="8" pageId="4" pageNumber="571">382 (162–385; 279)</td>
<td id="7B205DE4C803003DFC3EFBBD228F4317" box="[918,1427,1064,1117]" gridcol="2" gridrow="8" pageId="4" pageNumber="571">(187–193)</td>
</tr>
<tr id="38F13498C803003DFF37FBE5228F43EF" box="[159,1427,1136,1189]" gridrow="9" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FBE526FE43EF" box="[159,482,1136,1189]" gridcol="0" gridrow="9" pageId="4" pageNumber="571">[Percentage of body length]</th>
<td id="7B205DE4C803003DFDE2FBE5244543EF" box="[586,857,1136,1189]" gridcol="1" gridrow="9" pageId="4" pageNumber="571">25 [13–25; 18]</td>
<td id="7B205DE4C803003DFC3EFBE5228F43EF" box="[918,1427,1136,1189]" gridcol="2" gridrow="9" pageId="4" pageNumber="571">[13–14]1</td>
</tr>
<tr id="38F13498C803003DFF37FB2D228F43A7" box="[159,1427,1208,1261]" gridrow="10" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FB2D26FE43A7" box="[159,482,1208,1261]" gridcol="0" gridrow="10" pageId="4" pageNumber="571">Pre-oral lobe</th>
<td id="7B205DE4C803003DFDE2FB2D244543A7" box="[586,857,1208,1261]" gridcol="1" gridrow="10" pageId="4" pageNumber="571">69 (43–83; 59)</td>
<td id="7B205DE4C803003DFC3EFB2D228F43A7" box="[918,1427,1208,1261]" gridcol="2" gridrow="10" pageId="4" pageNumber="571">(19–32)</td>
</tr>
<tr id="38F13498C803003DFF37FA95228F427F" box="[159,1427,1280,1333]" gridrow="11" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FA9526FE427F" box="[159,482,1280,1333]" gridcol="0" gridrow="11" pageId="4" pageNumber="571">[Percentage of body length]</th>
<td id="7B205DE4C803003DFDE2FA952445427F" box="[586,857,1280,1333]" gridcol="1" gridrow="11" pageId="4" pageNumber="571">4 [2–5; 3]</td>
<td id="7B205DE4C803003DFC3EFA95228F427F" box="[918,1427,1280,1333]" gridcol="2" gridrow="11" pageId="4" pageNumber="571">[0.9–2.3]</td>
</tr>
<tr id="38F13498C803003DFF37FADD228F42CA" box="[159,1427,1352,1408]" gridrow="12" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FADD26FE42CA" box="[159,482,1352,1408]" gridcol="0" gridrow="12" pageId="4" pageNumber="571">Prepharynx length</th>
<td id="7B205DE4C803003DFDE2FADD244542CA" box="[586,857,1352,1408]" gridcol="1" gridrow="12" pageId="4" pageNumber="571">10 (10–78; 49)</td>
<td id="7B205DE4C803003DFC3EFADD228F42CA" box="[918,1427,1352,1408]" gridcol="2" gridrow="12" pageId="4" pageNumber="571">0 2</td>
</tr>
<tr id="38F13498C803003DFF37FA06228F4282" box="[159,1427,1427,1480]" gridrow="13" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FA0626FE4282" box="[159,482,1427,1480]" gridcol="0" gridrow="13" pageId="4" pageNumber="571">[Percentage of body length]</th>
<td id="7B205DE4C803003DFDE2FA0624454282" box="[586,857,1427,1480]" gridcol="1" gridrow="13" pageId="4" pageNumber="571">6 (6–22; 17)</td>
<td id="7B205DE4C803003DFC3EFA06228F4282" box="[918,1427,1427,1480]" gridcol="2" gridrow="13" pageId="4" pageNumber="571">-</td>
</tr>
<tr id="38F13498C803003DFF37FA4E228F415A" box="[159,1427,1499,1552]" gridrow="14" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37FA4E26FE415A" box="[159,482,1499,1552]" gridcol="0" gridrow="14" pageId="4" pageNumber="571">Pharynx length</th>
<td id="7B205DE4C803003DFDE2FA4E2445415A" box="[586,857,1499,1552]" gridcol="1" gridrow="14" pageId="4" pageNumber="571">245 (167–256; 210)</td>
<td id="7B205DE4C803003DFC3EFA4E228F415A" box="[918,1427,1499,1552]" gridcol="2" gridrow="14" pageId="4" pageNumber="571">(131–135)</td>
</tr>
<tr id="38F13498C803003DFF37F9B6228F4112" box="[159,1427,1571,1624]" gridrow="15" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37F9B626FE4112" box="[159,482,1571,1624]" gridcol="0" gridrow="15" pageId="4" pageNumber="571">Pharynx width</th>
<td id="7B205DE4C803003DFDE2F9B624454112" box="[586,857,1571,1624]" gridcol="1" gridrow="15" pageId="4" pageNumber="571">284 (180–348; 270)</td>
<td id="7B205DE4C803003DFC3EF9B6228F4112" box="[918,1427,1571,1624]" gridcol="2" gridrow="15" pageId="4" pageNumber="571">(129)</td>
</tr>
<tr id="38F13498C803003DFF37F9FE228F41EA" box="[159,1427,1643,1696]" gridrow="16" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37F9FE26FE41EA" box="[159,482,1643,1696]" gridcol="0" gridrow="16" pageId="4" pageNumber="571">Oral sucker/pharynx width ratio</th>
<td id="7B205DE4C803003DFDE2F9FE244541EA" box="[586,857,1643,1696]" gridcol="1" gridrow="16" pageId="4" pageNumber="571">1.3 (1:0.5–1.4; 1.0)</td>
<td id="7B205DE4C803003DFC3EF9FE228F41EA" box="[918,1427,1643,1696]" gridcol="2" gridrow="16" pageId="4" pageNumber="571">(1:0.7–1:0.8)</td>
</tr>
<tr id="38F13498C803003DFF37F926228F41A0" box="[159,1427,1715,1770]" gridrow="17" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37F92626FE41A0" box="[159,482,1715,1770]" gridcol="0" gridrow="17" pageId="4" pageNumber="571">[Percentage of body length]</th>
<td id="7B205DE4C803003DFDE2F926244541A0" box="[586,857,1715,1770]" gridcol="1" gridrow="17" pageId="4" pageNumber="571">18 [12–18; 13]</td>
<td id="7B205DE4C803003DFC3EF926228F41A0" box="[918,1427,1715,1770]" gridcol="2" gridrow="17" pageId="4" pageNumber="571">[about 9]1</td>
</tr>
<tr id="38F13498C803003DFF37F968228F4078" box="[159,1427,1789,1842]" gridrow="18" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37F96826FE4078" box="[159,482,1789,1842]" gridcol="0" gridrow="18" pageId="4" pageNumber="571">Esophagus length</th>
<td id="7B205DE4C803003DFDE2F96824454078" box="[586,857,1789,1842]" gridcol="1" gridrow="18" pageId="4" pageNumber="571">137 (43–151 (100)</td>
<td id="7B205DE4C803003DFC3EF968228F4078" box="[918,1427,1789,1842]" gridcol="2" gridrow="18" pageId="4" pageNumber="571">(64–71)</td>
</tr>
<tr id="38F13498C803003DFF37F8D0228F4030" box="[159,1427,1861,1914]" gridrow="19" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37F8D026FE4030" box="[159,482,1861,1914]" gridcol="0" gridrow="19" pageId="4" pageNumber="571">Ventral sucker length</th>
<td id="7B205DE4C803003DFDE2F8D024454030" box="[586,857,1861,1914]" gridcol="1" gridrow="19" pageId="4" pageNumber="571">206 (135–223; 186)</td>
<td id="7B205DE4C803003DFC3EF8D0228F4030" box="[918,1427,1861,1914]" gridcol="2" gridrow="19" pageId="4" pageNumber="571">(129–135)</td>
</tr>
<tr id="38F13498C803003DFF37F818228F4088" box="[159,1427,1933,1986]" gridrow="20" pageId="4" pageNumber="571">
<th id="7B205DE4C803003DFF37F81826FE4088" box="[159,482,1933,1986]" gridcol="0" gridrow="20" pageId="4" pageNumber="571">Ventral sucker width</th>
<td id="7B205DE4C803003DFDE2F81824454088" box="[586,857,1933,1986]" gridcol="1" gridrow="20" pageId="4" pageNumber="571">255 (151–255; 203)</td>
<td id="7B205DE4C803003DFC3EF818228F4088" box="[918,1427,1933,1986]" gridcol="2" gridrow="20" pageId="4" pageNumber="571">(136–142)</td>
</tr>
</table>
</paragraph>
<paragraph id="867E36DAC803FFC6FBF4F840228F40A6" blockId="4.[1116,1427,2005,2028]" box="[1116,1427,2005,2028]" pageId="4" pageNumber="571">
<emphasis id="B4B5EAC8C803FFC6FBF4F840228F40A6" box="[1116,1427,2005,2028]" italics="true" pageId="4" pageNumber="571">......continued on the next page</emphasis>
</paragraph>
<caption id="D2BE6652C802FFC7FF37FF35269147FC" box="[159,397,160,182]" pageId="5" pageNumber="572" targetBox="[159,1427,201,1981]" targetIsTable="true" targetPageId="5">
<paragraph id="867E36DAC802FFC7FF37FF35269147FC" blockId="5.[159,397,160,182]" box="[159,397,160,182]" pageId="5" pageNumber="572">
<emphasis id="B4B5EAC8C802FFC7FF37FF35261047FC" bold="true" box="[159,268,160,182]" pageId="5" pageNumber="572">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="867E36DAC802FFC7FF37FF5C231A40DB" pageId="5" pageNumber="572">
<table id="F4C1C47AC802003DFF37FF5C228F40F7" box="[159,1427,201,1981]" colsContinueFrom="4.[159,1427,486,1986]" colsContinueIn="6.[159,1427,201,1125]" gridcols="3" gridrows="23" pageId="5" pageNumber="572">
<tr id="38F13498C802003DFF37FF5C228F47B4" box="[159,1427,201,254]" gridrow="0" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FF5C250147B4" box="[159,541,201,254]" gridcol="0" gridrow="0" pageId="5" pageNumber="572">Species/Feature</th>
<th id="7B205DE4C802003DFDE2FF5C240D47B4" box="[586,785,201,254]" gridcol="1" gridrow="0" pageId="5" pageNumber="572">
<taxonomicName id="41C14D59C802FFC7FDE2FF5F25C84795" box="[586,724,202,223]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="572" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C802FFC7FDE2FF5F254B4795" box="[586,599,202,223]" italics="true" pageId="5" pageNumber="572">P</emphasis>
. 
<emphasis id="B4B5EAC8C802FFC7FDCCFF5F25C84795" box="[612,724,202,223]" italics="true" pageId="5" pageNumber="572">aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C802FFC7FD72FF5F240D4795" box="[730,785,202,223]" pageId="5" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
n=18
</th>
<th id="7B205DE4C802003DFC3EFF5C228F47B4" box="[918,1427,201,254]" gridcol="2" gridrow="0" pageId="5" pageNumber="572">
<taxonomicName id="41C14D59C802FFC7FC3EFF5F221B4795" authority="Bray &amp; Cribb, 1998" authorityName="Bray &amp; Cribb" authorityYear="1998" box="[918,1287,201,223]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="572" phylum="Chordata" rank="species" species="spongiosum" status="gen. nov.">
<emphasis id="B4B5EAC8C802FFC7FC3EFF5F24BE4795" box="[918,930,202,223]" italics="true" pageId="5" pageNumber="572">P</emphasis>
. 
<emphasis id="B4B5EAC8C802FFC7FC07FF5F23344795" box="[943,1064,202,223]" italics="true" pageId="5" pageNumber="572">spongiosum</emphasis>
(Bray &amp; Cribb, 1998)
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C802FFC7FAA5FF5F22484795" box="[1293,1364,202,223]" pageId="5" pageNumber="572" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp. n=2
</th>
</tr>
<tr id="38F13498C802003DFF37FE84228F460C" box="[159,1427,273,326]" gridrow="1" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FE842501460C" box="[159,541,273,326]" gridcol="0" gridrow="1" pageId="5" pageNumber="572">Ventral sucker/oral sucker width ratio</th>
<td id="7B205DE4C802003DFDE2FE84240D460C" box="[586,785,273,326]" gridcol="1" gridrow="1" pageId="5" pageNumber="572">1:0.6 (1:0.5–1:1.0; 1:0.7)</td>
<td id="7B205DE4C802003DFC3EFE84228F460C" box="[918,1427,273,326]" gridcol="2" gridrow="1" pageId="5" pageNumber="572">(1:0.73–1:0.74)</td>
</tr>
<tr id="38F13498C802003DFF37FECC228F46C4" box="[159,1427,345,398]" gridrow="2" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FECC250146C4" box="[159,541,345,398]" gridcol="0" gridrow="2" pageId="5" pageNumber="572">Cirrus-sac length</th>
<td id="7B205DE4C802003DFDE2FECC240D46C4" box="[586,785,345,398]" gridcol="1" gridrow="2" pageId="5" pageNumber="572">411 (216–616; 387)</td>
<td id="7B205DE4C802003DFC3EFECC228F46C4" box="[918,1427,345,398]" gridcol="2" gridrow="2" pageId="5" pageNumber="572">(276–277)</td>
</tr>
<tr id="38F13498C802003DFF37FE34228F4692" box="[159,1427,417,472]" gridrow="3" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FE3425014692" box="[159,541,417,472]" gridcol="0" gridrow="3" pageId="5" pageNumber="572">[Percentage of body length]</th>
<td id="7B205DE4C802003DFDE2FE34240D4692" box="[586,785,417,472]" gridcol="1" gridrow="3" pageId="5" pageNumber="572">26 [12–28; 18]</td>
<td id="7B205DE4C802003DFC3EFE34228F4692" box="[918,1427,417,472]" gridcol="2" gridrow="3" pageId="5" pageNumber="572">[19–21]1</td>
</tr>
<tr id="38F13498C802003DFF37FE79228F456A" box="[159,1427,492,544]" gridrow="4" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FE792501456A" box="[159,541,492,544]" gridcol="0" gridrow="4" pageId="5" pageNumber="572">Cirrus-sac width</th>
<td id="7B205DE4C802003DFDE2FE79240D456A" box="[586,785,492,544]" gridcol="1" gridrow="4" pageId="5" pageNumber="572">98 (63–103; 86)</td>
<td id="7B205DE4C802003DFC3EFE79228F456A" box="[918,1427,492,544]" gridcol="2" gridrow="4" pageId="5" pageNumber="572">(88–90)</td>
</tr>
<tr id="38F13498C802003DFF37FDA1228F4521" box="[159,1427,564,619]" gridrow="5" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FDA125014521" box="[159,541,564,619]" gridcol="0" gridrow="5" pageId="5" pageNumber="572">Internal seminal vesicle length</th>
<td id="7B205DE4C802003DFDE2FDA1240D4521" box="[586,785,564,619]" gridcol="1" gridrow="5" pageId="5" pageNumber="572">65 (35–105; 81)</td>
<td id="7B205DE4C802003DFC3EFDA1228F4521" box="[918,1427,564,619]" gridcol="2" gridrow="5" pageId="5" pageNumber="572">(about 75)4</td>
</tr>
<tr id="38F13498C802003DFF37FDEB228F45F9" box="[159,1427,638,691]" gridrow="6" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FDEB250145F9" box="[159,541,638,691]" gridcol="0" gridrow="6" pageId="5" pageNumber="572">[Percentage of cirrus-sac length]</th>
<td id="7B205DE4C802003DFDE2FDEB240D45F9" box="[586,785,638,691]" gridcol="1" gridrow="6" pageId="5" pageNumber="572">19 [16–20; 18]</td>
<td id="7B205DE4C802003DFC3EFDEB228F45F9" box="[918,1427,638,691]" gridcol="2" gridrow="6" pageId="5" pageNumber="572">[27]</td>
</tr>
<tr id="38F13498C802003DFF37FD53228F45B1" box="[159,1427,710,763]" gridrow="7" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FD53250145B1" box="[159,541,710,763]" gridcol="0" gridrow="7" pageId="5" pageNumber="572">Internal seminal vesicle width</th>
<td id="7B205DE4C802003DFDE2FD53240D45B1" box="[586,785,710,763]" gridcol="1" gridrow="7" pageId="5" pageNumber="572">65 (54–101; 72)</td>
<td id="7B205DE4C802003DFC3EFD53228F45B1" box="[918,1427,710,763]" gridcol="2" gridrow="7" pageId="5" pageNumber="572">NG</td>
</tr>
<tr id="38F13498C802003DFF37FC9B228F440F" box="[159,1427,782,837]" gridrow="8" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FC9B2501440F" box="[159,541,782,837]" gridcol="0" gridrow="8" pageId="5" pageNumber="572">Pars prostatica length</th>
<td id="7B205DE4C802003DFDE2FC9B240D440F" box="[586,785,782,837]" gridcol="1" gridrow="8" pageId="5" pageNumber="572">156 (89–165; 130)</td>
<td id="7B205DE4C802003DFC3EFC9B228F440F" box="[918,1427,782,837]" gridcol="2" gridrow="8" pageId="5" pageNumber="572">(about 63)4</td>
</tr>
<tr id="38F13498C802003DFF37FCCC228F44C7" box="[159,1427,857,909]" gridrow="9" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FCCC250144C7" box="[159,541,857,909]" gridcol="0" gridrow="9" pageId="5" pageNumber="572">[Percentage of cirrus-sac length]</th>
<td id="7B205DE4C802003DFDE2FCCC240D44C7" box="[586,785,857,909]" gridcol="1" gridrow="9" pageId="5" pageNumber="572">38 [26–41; 34]</td>
<td id="7B205DE4C802003DFC3EFCCC228F44C7" box="[918,1427,857,909]" gridcol="2" gridrow="9" pageId="5" pageNumber="572">[23]</td>
</tr>
<tr id="38F13498C802003DFF37FC34228F44BC" box="[159,1427,929,1014]" gridrow="10" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FC34250144BC" box="[159,541,929,1014]" gridcol="0" gridrow="10" pageId="5" pageNumber="572">Ejaculatory duct length</th>
<td id="7B205DE4C802003DFDE2FC34240D44BC" box="[586,785,929,1014]" gridcol="1" gridrow="10" pageId="5" pageNumber="572">82 (60–98; 77) n=8 5</td>
<td id="7B205DE4C802003DFC3EFC34228F44BC" box="[918,1427,929,1014]" gridcol="2" gridrow="10" pageId="5" pageNumber="572">(about 175)4</td>
</tr>
<tr id="38F13498C802003DFF37FB9F228F4374" box="[159,1427,1034,1086]" gridrow="11" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FB9F25014374" box="[159,541,1034,1086]" gridcol="0" gridrow="11" pageId="5" pageNumber="572">[Percentage of cirrus-sac length]</th>
<td id="7B205DE4C802003DFDE2FB9F240D4374" box="[586,785,1034,1086]" gridcol="1" gridrow="11" pageId="5" pageNumber="572">20 [17–25; 21]</td>
<td id="7B205DE4C802003DFC3EFB9F228F4374" box="[918,1427,1034,1086]" gridcol="2" gridrow="11" pageId="5" pageNumber="572">[63]</td>
</tr>
<tr id="38F13498C802003DFF37FBC7228F43E2" box="[159,1427,1106,1192]" gridrow="12" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FBC7250143E2" box="[159,541,1106,1192]" gridcol="0" gridrow="12" pageId="5" pageNumber="572">Inverted cirrus length</th>
<td id="7B205DE4C802003DFDE2FBC7240D43E2" box="[586,785,1106,1192]" gridcol="1" gridrow="12" pageId="5" pageNumber="572">96 (70–100; 81) n=8 5</td>
<td id="7B205DE4C802003DFC3EFBC7228F43E2" box="[918,1427,1106,1192]" gridcol="2" gridrow="12" pageId="5" pageNumber="572">(about 81)4</td>
</tr>
<tr id="38F13498C802003DFF37FB2E228F43BA" box="[159,1427,1211,1264]" gridrow="13" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FB2E250143BA" box="[159,541,1211,1264]" gridcol="0" gridrow="13" pageId="5" pageNumber="572">[Percentage of cirrus-sac length]</th>
<td id="7B205DE4C802003DFDE2FB2E240D43BA" box="[586,785,1211,1264]" gridcol="1" gridrow="13" pageId="5" pageNumber="572">23 [18–28; 22]</td>
<td id="7B205DE4C802003DFC3EFB2E228F43BA" box="[918,1427,1211,1264]" gridcol="2" gridrow="13" pageId="5" pageNumber="572">[29]</td>
</tr>
<tr id="38F13498C802003DFF37FA96228F4213" box="[159,1427,1283,1369]" gridrow="14" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FA9625014213" box="[159,541,1283,1369]" gridcol="0" gridrow="14" pageId="5" pageNumber="572">External seminal vesicle length</th>
<td id="7B205DE4C802003DFDE2FA96240D4213" box="[586,785,1283,1369]" gridcol="1" gridrow="14" pageId="5" pageNumber="572">540 (485–540; 512) n=8 5</td>
<td id="7B205DE4C802003DFC3EFA96228F4213" box="[918,1427,1283,1369]" gridcol="2" gridrow="14" pageId="5" pageNumber="572">(about 150)4</td>
</tr>
<tr id="38F13498C802003DFF37FAF9228F42EB" box="[159,1427,1388,1441]" gridrow="15" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FAF9250142EB" box="[159,541,1388,1441]" gridcol="0" gridrow="15" pageId="5" pageNumber="572">External seminal vesicle width</th>
<td id="7B205DE4C802003DFDE2FAF9240D42EB" box="[586,785,1388,1441]" gridcol="1" gridrow="15" pageId="5" pageNumber="572">48 (45–94: 63)</td>
<td id="7B205DE4C802003DFC3EFAF9228F42EB" box="[918,1427,1388,1441]" gridcol="2" gridrow="15" pageId="5" pageNumber="572">NG</td>
</tr>
<tr id="38F13498C802003DFF37FA21228F42A3" box="[159,1427,1460,1513]" gridrow="16" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FA21250142A3" box="[159,541,1460,1513]" gridcol="0" gridrow="16" pageId="5" pageNumber="572">Distance from ventral sucker to ovary</th>
<td id="7B205DE4C802003DFDE2FA21240D42A3" box="[586,785,1460,1513]" gridcol="1" gridrow="16" pageId="5" pageNumber="572">304 (160–321; 243)</td>
<td id="7B205DE4C802003DFC3EFA21228F42A3" box="[918,1427,1460,1513]" gridcol="2" gridrow="16" pageId="5" pageNumber="572">(119–140)</td>
</tr>
<tr id="38F13498C802003DFF37FA69228F417B" box="[159,1427,1532,1585]" gridrow="17" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37FA692501417B" box="[159,541,1532,1585]" gridcol="0" gridrow="17" pageId="5" pageNumber="572">[Percentage of body length]</th>
<td id="7B205DE4C802003DFDE2FA69240D417B" box="[586,785,1532,1585]" gridcol="1" gridrow="17" pageId="5" pageNumber="572">20 [10–20; 12]</td>
<td id="7B205DE4C802003DFC3EFA69228F417B" box="[918,1427,1532,1585]" gridcol="2" gridrow="17" pageId="5" pageNumber="572">[8–10]</td>
</tr>
<tr id="38F13498C802003DFF37F9D1228F4133" box="[159,1427,1604,1657]" gridrow="18" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37F9D125014133" box="[159,541,1604,1657]" gridcol="0" gridrow="18" pageId="5" pageNumber="572">Ovary length</th>
<td id="7B205DE4C802003DFDE2F9D1240D4133" box="[586,785,1604,1657]" gridcol="1" gridrow="18" pageId="5" pageNumber="572">127 (108–303; 170)</td>
<td id="7B205DE4C802003DFC3EF9D1228F4133" box="[918,1427,1604,1657]" gridcol="2" gridrow="18" pageId="5" pageNumber="572">(122–142)</td>
</tr>
<tr id="38F13498C802003DFF37F919228F418B" box="[159,1427,1676,1729]" gridrow="19" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37F9192501418B" box="[159,541,1676,1729]" gridcol="0" gridrow="19" pageId="5" pageNumber="572">Ovary width</th>
<td id="7B205DE4C802003DFDE2F919240D418B" box="[586,785,1676,1729]" gridcol="1" gridrow="19" pageId="5" pageNumber="572">205 (127–301; 203)</td>
<td id="7B205DE4C802003DFC3EF919228F418B" box="[918,1427,1676,1729]" gridcol="2" gridrow="19" pageId="5" pageNumber="572">(122–135)</td>
</tr>
<tr id="38F13498C802003DFF37F941228F4041" box="[159,1427,1748,1803]" gridrow="20" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37F94125014041" box="[159,541,1748,1803]" gridcol="0" gridrow="20" pageId="5" pageNumber="572">Seminal receptacle length</th>
<td id="7B205DE4C802003DFDE2F941240D4041" box="[586,785,1748,1803]" gridcol="1" gridrow="20" pageId="5" pageNumber="572">284 (114–310; 233)</td>
<td id="7B205DE4C802003DFC3EF941228F4041" box="[918,1427,1748,1803]" gridcol="2" gridrow="20" pageId="5" pageNumber="572">(about 116)4</td>
</tr>
<tr id="38F13498C802003DFF37F88A228F4019" box="[159,1427,1823,1875]" gridrow="21" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37F88A25014019" box="[159,541,1823,1875]" gridcol="0" gridrow="21" pageId="5" pageNumber="572">Seminal receptacle width</th>
<td id="7B205DE4C802003DFDE2F88A240D4019" box="[586,785,1823,1875]" gridcol="1" gridrow="21" pageId="5" pageNumber="572">128 (56–131; 94)</td>
<td id="7B205DE4C802003DFC3EF88A228F4019" box="[918,1427,1823,1875]" gridcol="2" gridrow="21" pageId="5" pageNumber="572">NG</td>
</tr>
<tr id="38F13498C802003DFF37F8F2228F40F7" box="[159,1427,1895,1981]" gridrow="22" pageId="5" pageNumber="572">
<th id="7B205DE4C802003DFF37F8F2250140F7" box="[159,541,1895,1981]" gridcol="0" gridrow="22" pageId="5" pageNumber="572">Metraterm length</th>
<td id="7B205DE4C802003DFDE2F8F2240D40F7" box="[586,785,1895,1981]" gridcol="1" gridrow="22" pageId="5" pageNumber="572">276 (180–335; 243) n=8 5</td>
<td id="7B205DE4C802003DFC3EF8F2228F40F7" box="[918,1427,1895,1981]" gridcol="2" gridrow="22" pageId="5" pageNumber="572">(about 75)3</td>
</tr>
</table>
</paragraph>
<paragraph id="867E36DAC802FFC4FBF4F84525E843D7" blockId="5.[1116,1427,2000,2022]" lastBlockId="6.[151,1121,1158,1352]" lastPageId="6" lastPageNumber="573" pageId="5" pageNumber="572">
<emphasis id="B4B5EAC8C802FFC7FBF4F845228F40AC" box="[1116,1427,2000,2022]" italics="true" pageId="5" pageNumber="572">......continued on the next page</emphasis>
n = Sample size. NG = Not given in original description.
</paragraph>
<caption id="D2BE6652C801FFC4FF37FF35269147FC" box="[159,397,160,182]" pageId="6" pageNumber="573" targetBox="[159,1427,201,1125]" targetIsTable="true" targetPageId="6">
<paragraph id="867E36DAC801FFC4FF37FF35269147FC" blockId="6.[159,397,160,182]" box="[159,397,160,182]" pageId="6" pageNumber="573">
<emphasis id="B4B5EAC8C801FFC4FF37FF35261047FC" bold="true" box="[159,268,160,182]" pageId="6" pageNumber="573">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="867E36DAC801FFC4FF37FF5C2311432F" pageId="6" pageNumber="573">
<table id="F4C1C47AC801003DFF37FF5C228F432F" box="[159,1427,201,1125]" colsContinueFrom="5.[159,1427,201,1981]" gridcols="3" gridrows="13" pageId="6" pageNumber="573">
<tr id="38F13498C801003DFF37FF5C228F47B4" box="[159,1427,201,254]" gridrow="0" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FF5C26D047B4" box="[159,460,201,254]" gridcol="0" gridrow="0" pageId="6" pageNumber="573">Species/Feature</th>
<th id="7B205DE4C801003DFDE2FF5C240D47B4" box="[586,785,201,254]" gridcol="1" gridrow="0" pageId="6" pageNumber="573">
<taxonomicName id="41C14D59C801FFC4FDE2FF5F25C84795" box="[586,724,202,223]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="6" pageNumber="573" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C801FFC4FDE2FF5F254B4795" box="[586,599,202,223]" italics="true" pageId="6" pageNumber="573">P</emphasis>
. 
<emphasis id="B4B5EAC8C801FFC4FDCCFF5F25C84795" box="[612,724,202,223]" italics="true" pageId="6" pageNumber="573">aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C801FFC4FD72FF5F240D4795" box="[730,785,202,223]" pageId="6" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
n=18
</th>
<th id="7B205DE4C801003DFC3EFF5C228F47B4" box="[918,1427,201,254]" gridcol="2" gridrow="0" pageId="6" pageNumber="573">
<taxonomicName id="41C14D59C801FFC4FC3EFF5F221B4795" authority="Bray &amp; Cribb, 1998" authorityName="Bray &amp; Cribb" authorityYear="1998" box="[918,1287,201,223]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="6" pageNumber="573" phylum="Chordata" rank="species" species="spongiosum" status="gen. nov.">
<emphasis id="B4B5EAC8C801FFC4FC3EFF5F24BE4795" box="[918,930,202,223]" italics="true" pageId="6" pageNumber="573">P</emphasis>
. 
<emphasis id="B4B5EAC8C801FFC4FC07FF5F23344795" box="[943,1064,202,223]" italics="true" pageId="6" pageNumber="573">spongiosum</emphasis>
(Bray &amp; Cribb, 1998)
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C801FFC4FAA5FF5F22484795" box="[1293,1364,202,223]" pageId="6" pageNumber="573" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp. n=2
</th>
</tr>
<tr id="38F13498C801003DFF37FE84228F460C" box="[159,1427,273,326]" gridrow="1" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FE8426D0460C" box="[159,460,273,326]" gridcol="0" gridrow="1" pageId="6" pageNumber="573">[Percentage of body length]</th>
<td id="7B205DE4C801003DFDE2FE84240D460C" box="[586,785,273,326]" gridcol="1" gridrow="1" pageId="6" pageNumber="573">7 [7–16; 11]</td>
<td id="7B205DE4C801003DFC3EFE84228F460C" box="[918,1427,273,326]" gridcol="2" gridrow="1" pageId="6" pageNumber="573">[5]</td>
</tr>
<tr id="38F13498C801003DFF37FECC228F46C4" box="[159,1427,345,398]" gridrow="2" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FECC26D046C4" box="[159,460,345,398]" gridcol="0" gridrow="2" pageId="6" pageNumber="573">Anterior testis length</th>
<td id="7B205DE4C801003DFDE2FECC240D46C4" box="[586,785,345,398]" gridcol="1" gridrow="2" pageId="6" pageNumber="573">323 (147–323; 206)</td>
<td id="7B205DE4C801003DFC3EFECC228F46C4" box="[918,1427,345,398]" gridcol="2" gridrow="2" pageId="6" pageNumber="573">(138–155)</td>
</tr>
<tr id="38F13498C801003DFF37FE34228F469C" box="[159,1427,417,470]" gridrow="3" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FE3426D0469C" box="[159,460,417,470]" gridcol="0" gridrow="3" pageId="6" pageNumber="573">Anterior testis width</th>
<td id="7B205DE4C801003DFDE2FE34240D469C" box="[586,785,417,470]" gridcol="1" gridrow="3" pageId="6" pageNumber="573">441 (233–463; 345)</td>
<td id="7B205DE4C801003DFC3EFE34228F469C" box="[918,1427,417,470]" gridcol="2" gridrow="3" pageId="6" pageNumber="573">(129–148)</td>
</tr>
<tr id="38F13498C801003DFF37FE7C228F4554" box="[159,1427,489,542]" gridrow="4" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FE7C26D04554" box="[159,460,489,542]" gridcol="0" gridrow="4" pageId="6" pageNumber="573">Posterior testis length</th>
<td id="7B205DE4C801003DFDE2FE7C240D4554" box="[586,785,489,542]" gridcol="1" gridrow="4" pageId="6" pageNumber="573">412 (137–412; 225)</td>
<td id="7B205DE4C801003DFC3EFE7C228F4554" box="[918,1427,489,542]" gridcol="2" gridrow="4" pageId="6" pageNumber="573">(155–180)</td>
</tr>
<tr id="38F13498C801003DFF37FDA4228F452C" box="[159,1427,561,614]" gridrow="5" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FDA426D0452C" box="[159,460,561,614]" gridcol="0" gridrow="5" pageId="6" pageNumber="573">Posterior testis width</th>
<td id="7B205DE4C801003DFDE2FDA4240D452C" box="[586,785,561,614]" gridcol="1" gridrow="5" pageId="6" pageNumber="573">412 (199–412; 323)</td>
<td id="7B205DE4C801003DFC3EFDA4228F452C" box="[918,1427,561,614]" gridcol="2" gridrow="5" pageId="6" pageNumber="573">(116–155)</td>
</tr>
<tr id="38F13498C801003DFF37FDEC228F45FA" box="[159,1427,633,688]" gridrow="6" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FDEC26D045FA" box="[159,460,633,688]" gridcol="0" gridrow="6" pageId="6" pageNumber="573">Mean testes/ovary width ratio</th>
<td id="7B205DE4C801003DFDE2FDEC240D45FA" box="[586,785,633,688]" gridcol="1" gridrow="6" pageId="6" pageNumber="573">1:2.1 (1:1.4–1:2.1; 1:1.6)</td>
<td id="7B205DE4C801003DFC3EFDEC228F45FA" box="[918,1427,633,688]" gridcol="2" gridrow="6" pageId="6" pageNumber="573">(1:1.0–1:1.2)1</td>
</tr>
<tr id="38F13498C801003DFF37FD51228F45B1" box="[159,1427,708,763]" gridrow="7" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FD5126D045B1" box="[159,460,708,763]" gridcol="0" gridrow="7" pageId="6" pageNumber="573">Intertesticular space</th>
<td id="7B205DE4C801003DFDE2FD51240D45B1" box="[586,785,708,763]" gridcol="1" gridrow="7" pageId="6" pageNumber="573">0 (0–17; 7)</td>
<td id="7B205DE4C801003DFC3EFD51228F45B1" box="[918,1427,708,763]" gridcol="2" gridrow="7" pageId="6" pageNumber="573">(about 13)3</td>
</tr>
<tr id="38F13498C801003DFF37FC9B228F4409" box="[159,1427,782,835]" gridrow="8" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FC9B26D04409" box="[159,460,782,835]" gridcol="0" gridrow="8" pageId="6" pageNumber="573">[Percentage of body length]</th>
<td id="7B205DE4C801003DFDE2FC9B240D4409" box="[586,785,782,835]" gridcol="1" gridrow="8" pageId="6" pageNumber="573">0 [0–0.8; 0.3]</td>
<td id="7B205DE4C801003DFC3EFC9B228F4409" box="[918,1427,782,835]" gridcol="2" gridrow="8" pageId="6" pageNumber="573">[0.9]</td>
</tr>
<tr id="38F13498C801003DFF37FCC3228F44C1" box="[159,1427,854,907]" gridrow="9" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FCC326D044C1" box="[159,460,854,907]" gridcol="0" gridrow="9" pageId="6" pageNumber="573">Post-testicular space</th>
<td id="7B205DE4C801003DFDE2FCC3240D44C1" box="[586,785,854,907]" gridcol="1" gridrow="9" pageId="6" pageNumber="573">304 (170–390; 259)</td>
<td id="7B205DE4C801003DFC3EFCC3228F44C1" box="[918,1427,854,907]" gridcol="2" gridrow="9" pageId="6" pageNumber="573">(236–245)</td>
</tr>
<tr id="38F13498C801003DFF37FC0B228F449F" box="[159,1427,926,981]" gridrow="10" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FC0B26D0449F" box="[159,460,926,981]" gridcol="0" gridrow="10" pageId="6" pageNumber="573">[Percentage of body length]</th>
<td id="7B205DE4C801003DFDE2FC0B240D449F" box="[586,785,926,981]" gridcol="1" gridrow="10" pageId="6" pageNumber="573">20 [8–20; 13]</td>
<td id="7B205DE4C801003DFC3EFC0B228F449F" box="[918,1427,926,981]" gridcol="2" gridrow="10" pageId="6" pageNumber="573">[17–18]1</td>
</tr>
<tr id="38F13498C801003DFF37FC7C228F4357" box="[159,1427,1001,1053]" gridrow="11" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FC7C26D04357" box="[159,460,1001,1053]" gridcol="0" gridrow="11" pageId="6" pageNumber="573">Egg length</th>
<td id="7B205DE4C801003DFDE2FC7C240D4357" box="[586,785,1001,1053]" gridcol="1" gridrow="11" pageId="6" pageNumber="573">57–62 (56–65; 60)</td>
<td id="7B205DE4C801003DFC3EFC7C228F4357" box="[918,1427,1001,1053]" gridcol="2" gridrow="11" pageId="6" pageNumber="573">(60–63; 62)</td>
</tr>
<tr id="38F13498C801003DFF37FBA4228F432F" box="[159,1427,1073,1125]" gridrow="12" pageId="6" pageNumber="573">
<th id="7B205DE4C801003DFF37FBA426D0432F" box="[159,460,1073,1125]" gridcol="0" gridrow="12" pageId="6" pageNumber="573">Egg width</th>
<td id="7B205DE4C801003DFDE2FBA4240D432F" box="[586,785,1073,1125]" gridcol="1" gridrow="12" pageId="6" pageNumber="573">38–40 (36–45; 41)</td>
<td id="7B205DE4C801003DFC3EFBA4228F432F" box="[918,1427,1073,1125]" gridcol="2" gridrow="12" pageId="6" pageNumber="573">(40–45; 42)</td>
</tr>
</table>
</paragraph>
<paragraph id="867E36DAC801FFC4FF3FFB3225AA43A9" blockId="6.[151,1121,1158,1352]" pageId="6" pageNumber="573">1 Calculated from measurements given in the original description. 2 Entirely within posterior concavity of oral sucker.</paragraph>
<paragraph id="867E36DAC801FFC4FF3FFB7825A3424C" blockId="6.[151,1121,1158,1352]" box="[151,703,1261,1286]" pageId="6" pageNumber="573">3Calculated from Fig. 18 of the original description.</paragraph>
<paragraph id="867E36DAC801FFC4FF3FFA8525DE4263" blockId="6.[151,1121,1158,1352]" box="[151,706,1296,1321]" pageId="6" pageNumber="573">4 Calculated from Fig. 20 of the original description.</paragraph>
<paragraph id="867E36DAC801FFC4FF3FFABA237D4202" blockId="6.[151,1121,1158,1352]" box="[151,1121,1327,1352]" pageId="6" pageNumber="573">
5 Based on 
<specimenCount id="90C7FD53C801FFC4FEA3FAA426924202" box="[267,398,1329,1352]" pageId="6" pageNumber="573" type="generic">8 specimens</specimenCount>
exclusive of the 
<typeStatus id="597A8878C801FFC4FDECFAA425BD4202" box="[580,673,1329,1352]" pageId="6" pageNumber="573" type="holotype">holotype</typeStatus>
where the terminal genitalia was clearest.
</paragraph>
<paragraph id="867E36DAC801FFC4FF6FFAE323C94050" blockId="6.[151,1437,1397,2034]" pageId="6" pageNumber="573">
Ovary immediately pre-testicular, sometimes contiguous with anterior testis, dextral to nearly medial, multilobed, consists of about 9–16 separate sub-globular follicles. Seminal receptacle canalicular, large, elongate-oval, sinistral to ovary, located beside or ventral to sinistral rim of anterior testis, narrows anteriorly to form Laurer’s canal which winds back posteriorly along sinistral side of seminal receptacle and opens dorsally at level of anterior third of seminal receptacle. Uterus pre-testicular, tubular, intercecal, mainly in anterior hindbody, begins at level of ovary and passes antero-sinistrally and laterally to ventral sucker, may overlap anterior margin of ovary and posterior margin of ventral sucker. Metraterm distinct, thick-walled, lined by cells with large nuclei (i.e. “muscular” wall of 
<bibRefCitation id="E2504B2BC801FFC4FED1F9E7254741C0" author="Bray" box="[377,603,1649,1674]" pageId="6" pageNumber="573" refString="Bray, R. A. &amp; Cribb, T. H. (1998) Lepocreadiidae (Digenea) of Australian coastal fishes: new species of Opechona Looss, 1907, Lepotrema Ozaki, 1932 and Bianium Stunkard, 1930 and comments on other species reported for the first time or poorly known in Australian waters. Systematic Parasitology, 41, 123 - 148." type="journal article" year="1998">Bray &amp; Cribb 1998</bibRefCitation>
), extends posteriorly about 3/4 length of cirrus-sac, enters genital atrium from left. Vitellarium follicular; fields extend along lateral margins from near level of intestinal bifurcation to posterior extremity, completely confluent in forebody and in post-testicular region, often encroaches into immediate pre-ovarian space. Follicles numerous, subspherical to elongate-oval or irregular in shape, both ventral and lateral to ceca but not dorsal. Eggs relatively numerous, oval, small, operculate, thin-shelled.
</paragraph>
<paragraph id="867E36DAC801FFC4FF6FF8B225194028" blockId="6.[151,1437,1397,2034]" pageId="6" pageNumber="573">Excretory vesicle I-shaped, wide, appears to reach almost to level of intestinal bifurcation. Excretory pore terminal to dorsally subterminal.</paragraph>
</subSubSection>
<subSubSection id="CEDB6551C801FFCAFF6FF8F825DA435E" lastPageId="8" lastPageNumber="575" pageId="6" pageNumber="573" type="discussion">
<paragraph id="867E36DAC801FFCAFF6FF8F825DA435E" blockId="6.[151,1437,1397,2034]" lastBlockId="8.[151,1436,948,2017]" lastPageId="8" lastPageNumber="575" pageId="6" pageNumber="573">
<emphasis id="B4B5EAC8C801FFC4FF6FF8F8262740CC" bold="true" box="[199,315,1901,1926]" pageId="6" pageNumber="573">Remarks.</emphasis>
The new species has an “ 
<taxonomicName id="41C14D59C801FFC4FDCDF8F825C440CC" box="[613,728,1901,1926]" class="Trematoda" family="Lepocreadiidae" genus="Opechona" kingdom="Animalia" order="Plagiorchiida" pageId="6" pageNumber="573" phylum="Platyhelminthes" rank="genus">
<emphasis id="B4B5EAC8C801FFC4FDCDF8F825C440CC" box="[613,728,1901,1926]" italics="true" pageId="6" pageNumber="573">Opechona</emphasis>
</taxonomicName>
- 
<typeStatus id="597A8878C801FFC4FD48F8FB240840CC" box="[736,788,1902,1926]" pageId="6" pageNumber="573">type</typeStatus>
” cirrus-sac and is placed in the 
<taxonomicName id="41C14D59C801FFC4FB2BF8F8222C40CC" box="[1155,1328,1901,1926]" class="Trematoda" family="Lepocreadiidae" kingdom="Animalia" order="Plagiorchiida" pageId="6" pageNumber="573" phylum="Platyhelminthes" rank="family">Lepocreadiidae</taxonomicName>
based on the reorganization of the Lepocreadioidea by 
<bibRefCitation id="E2504B2BC801FFC4FD30F807249540E0" author="Bray" box="[664,905,1937,1962]" pageId="6" pageNumber="573" refString="Bray, R. A. &amp; Cribb, T. H. (2012) Reorganisation of the superfamily Lepocreadioidea Odhner, 1905 based on an inferred molecular phylogeny. Systematic Parasitology, 83, 169 - 177." type="journal article" year="2012">Bray &amp; Cribb (2012)</bibRefCitation>
. In addition, the presence of a genital pore that is ventral and submedian, two testes, sponge-like lateral patches (pelops) in the anterior portion of the forebody instead of the lateral margins in the anterior portion of the body forming an incomplete scoop (i.e. not joined posteriorly), and ceca that extend close to the posterior body wall place this new species in 
<taxonomicName id="41C14D59C80FFFCAFB26FC21225F4487" box="[1166,1347,948,973]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B4B5EAC8C80FFFCAFB26FC21225F4487" box="[1166,1347,948,973]" italics="true" pageId="8" pageNumber="575">Pelopscreadium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C80FFFCAFAE1FC20228B4487" box="[1353,1431,949,973]" pageId="8" pageNumber="575" rank="genus">n. gen.</taxonomicNameLabel>
; however, there are distinct differences between the new species and the type-species of the new genus, 
<taxonomicName id="41C14D59C80FFFCAFA2FFC4C2601435E" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum" status="comb. nov.">
<emphasis id="B4B5EAC8C80FFFCAFA2FFC4C2601435E" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C80FFFCAFE8CFC68269E435F" box="[292,386,1020,1045]" pageId="8" pageNumber="575">n. comb</taxonomicNameLabel>
. (syn. 
<taxonomicName id="41C14D59C80FFFCAFE62FC6825AA435E" box="[458,694,1021,1044]" class="Trematoda" family="Lepocreadiidae" genus="Bianium" kingdom="Animalia" order="Plagiorchiida" pageId="8" pageNumber="575" phylum="Platyhelminthes" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFE62FC6825AA435E" box="[458,694,1021,1044]" italics="true" pageId="8" pageNumber="575">Bianium spongiosum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="D2BE6652C800FFC5FF3FF8A52259408A" httpUri="https://zenodo.org/record/260924/files/figure.png" pageId="7" pageNumber="574" targetBox="[227,1359,195,1745]" targetPageId="7">
<paragraph id="867E36DAC800FFC5FF3FF8A52259408A" blockId="7.[151,1436,1840,1984]" pageId="7" pageNumber="574">
<emphasis id="B4B5EAC8C800FFC5FF3FF8A52621400C" bold="true" box="[151,317,1840,1862]" pageId="7" pageNumber="574">FIGURES 1–3.</emphasis>
<taxonomicName id="41C14D59C800FFC5FEEFF8A52579400C" box="[327,613,1840,1862]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="7" pageNumber="574" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C800FFC5FEEFF8A52579400C" box="[327,613,1840,1862]" italics="true" pageId="7" pageNumber="574">Pelopscreadium aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C800FFC5FDC6F8A525B7400C" box="[622,683,1840,1862]" pageId="7" pageNumber="574" rank="species">
<emphasis id="B4B5EAC8C800FFC5FDC6F8A525B7400C" bold="true" box="[622,683,1840,1862]" pageId="7" pageNumber="574">n. sp.</emphasis>
</taxonomicNameLabel>
(
<taxonomicName id="41C14D59C800FFC5FD13F8A52446400C" box="[699,858,1840,1862]" class="Trematoda" family="Lepocreadiidae" kingdom="Animalia" order="Plagiorchiida" pageId="7" pageNumber="574" phylum="Platyhelminthes" rank="family">Lepocreadiidae</taxonomicName>
) from the yellow boxfish, 
<taxonomicName id="41C14D59C800FFC5FBD1F8A42602402E" authority="Linnaeus" authorityName="Linnaeus" class="Actinopterygii" family="Ostraciidae" genus="Ostracion" kingdom="Animalia" order="Tetraodontiformes" pageId="7" pageNumber="574" phylum="Chordata" rank="species" species="cubicus">
<emphasis id="B4B5EAC8C800FFC5FBD1F8A42229400C" box="[1145,1333,1840,1862]" italics="true" pageId="7" pageNumber="574">Ostracion cubicus</emphasis>
Linnaeus (Ostraciidae)
</taxonomicName>
, from the Red Sea off Egypt. 
<emphasis id="B4B5EAC8C800FFC5FDFCF8DB257B402E" bold="true" box="[596,615,1870,1892]" pageId="7" pageNumber="574">1.</emphasis>
Adult specimen, ventral view. 
<emphasis id="B4B5EAC8C800FFC5FC01F8DB24A7402E" bold="true" box="[937,955,1870,1892]" pageId="7" pageNumber="574">2.</emphasis>
Pharynx, lateral view. 
<emphasis id="B4B5EAC8C800FFC5FB02F8DB23A0402E" bold="true" box="[1194,1212,1870,1892]" pageId="7" pageNumber="574">3.</emphasis>
Details of the female genital complex, ventral view. 
<emphasis id="B4B5EAC8C800FFC5FE7DF8F8257E40C9" box="[469,610,1901,1923]" italics="true" pageId="7" pageNumber="574">Abbreviations</emphasis>
: C, ceca; D, oviduct; DL, distinct, well-differentiated pre-oral layer; E, external seminal vesicle; G, genital pore; I, internal seminal vesicle; O, ovary; OS, esophagus; OT, oötype; P, sponge-like pelops (“shoulder pads”); PP, prepharynx; R, seminal receptacle; T, testis; U, uterus; V, vitelline follicles; VD, vitelline ducts.
</paragraph>
</caption>
<caption id="D2BE6652C80FFFCAFF3FFC8026F944CD" httpUri="https://zenodo.org/record/260925/files/figure.png" pageId="8" pageNumber="575" targetBox="[400,1249,184,751]" targetPageId="8">
<paragraph id="867E36DAC80FFFCAFF3FFC8026F944CD" blockId="8.[151,1436,789,903]" pageId="8" pageNumber="575">
<emphasis id="B4B5EAC8C80FFFCAFF3FFC8026094461" bold="true" box="[151,277,789,811]" pageId="8" pageNumber="575">FIGURE 4.</emphasis>
Comparison of the male terminal genitalia of 
<taxonomicName id="41C14D59C80FFFCAFCA8FC8023014461" box="[768,1053,789,811]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C80FFFCAFCA8FC8023014461" box="[768,1053,789,811]" italics="true" pageId="8" pageNumber="575">Pelopscreadium aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C80FFFCAFB8EFC80237E4461" box="[1062,1122,789,811]" pageId="8" pageNumber="575" rank="species">
<emphasis id="B4B5EAC8C80FFFCAFB8EFC80237E4461" bold="true" box="[1062,1122,789,811]" pageId="8" pageNumber="575">n. sp.</emphasis>
</taxonomicNameLabel>
(left side) vs 
<taxonomicName id="41C14D59C80FFFCAFB50FC80260C4400" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum" status="gen. nov.">
<emphasis id="B4B5EAC8C80FFFCAFB50FC80260C4400" italics="true" pageId="8" pageNumber="575">Pelopscreadium spongiosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C80FFFCAFEBEFCA026414400" box="[278,349,821,842]" pageId="8" pageNumber="575" rank="genus">n. gen.</taxonomicNameLabel>
, n. comb. (right side). Illustration of 
<taxonomicName id="41C14D59C80FFFCAFD7AFCA024784400" box="[722,868,821,842]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum" status="gen. nov.">
<emphasis id="B4B5EAC8C80FFFCAFD7AFCA025C34400" box="[722,735,821,842]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFD43FCA024784400" box="[747,868,821,842]" italics="true" pageId="8" pageNumber="575">spongiosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C80FFFCAFCC2FCA024AD4400" box="[874,945,821,842]" pageId="8" pageNumber="575" rank="genus">n. gen.</taxonomicNameLabel>
, n. comb. was adapted from figure 20 of Bray &amp; Cribb (1998). 
<emphasis id="B4B5EAC8C80FFFCAFE85FCC626A64423" box="[301,442,851,873]" italics="true" pageId="8" pageNumber="575">Abbreviations</emphasis>
: E, external seminal vesicle; ED, ejaculatory duct; G, genital pore; I, internal seminal vesicle; PARS, pars prostatica; U, uterus.
</paragraph>
</caption>
<subSubSection id="CEDB6551C80FFFCBFF6FFBB52495460A" lastPageId="9" lastPageNumber="576" pageId="8" pageNumber="575" type="reference_group">
<paragraph id="867E36DAC80FFFCBFF6FFBB52495460A" blockId="8.[151,1436,948,2017]" lastBlockId="9.[151,1436,151,320]" lastPageId="9" lastPageNumber="576" pageId="8" pageNumber="575">
<taxonomicName id="41C14D59C80FFFCAFF6FFBB525164372" box="[199,522,1056,1081]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFF6FFBB525164372" box="[199,522,1056,1081]" italics="true" pageId="8" pageNumber="575">Pelopscreadium spongiosum</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="41C14D59C80FFFCAFD24FBB4243B4372" box="[652,807,1057,1080]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense" status="sp. nov.">
<emphasis id="B4B5EAC8C80FFFCAFD24FBB425864372" box="[652,666,1057,1080]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFD02FBB4243B4372" box="[682,807,1057,1080]" italics="true" pageId="8" pageNumber="575">aegyptense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657B3C80FFFCAFC87FBB5246D4373" box="[815,881,1056,1081]" pageId="8" pageNumber="575" rank="species">
<emphasis id="B4B5EAC8C80FFFCAFC87FBB5246D4373" bold="true" box="[815,881,1056,1081]" pageId="8" pageNumber="575">n. sp.</emphasis>
</taxonomicNameLabel>
in lacking a scoop and instead possessing large, internal patches of voluminous vacuolated cells which form symmetrical, sponge-like, lateral pelops in the forebody of the worm in the region between the anterior margin of the oral sucker and about the level of the intestinal bifurcation (
<bibRefCitation id="E2504B2BC80FFFCAFE32FB19259643EE" author="Bray" box="[410,650,1164,1189]" pageId="8" pageNumber="575" refString="Bray, R. A. &amp; Cribb, T. H. (1998) Lepocreadiidae (Digenea) of Australian coastal fishes: new species of Opechona Looss, 1907, Lepotrema Ozaki, 1932 and Bianium Stunkard, 1930 and comments on other species reported for the first time or poorly known in Australian waters. Systematic Parasitology, 41, 123 - 148." type="journal article" year="1998">Bray &amp; Cribb 1998</bibRefCitation>
). Along with the sponge-like, lateral pelops both species are also somewhat similar in overall appearance, egg size (60–63 × 40–45 µm in 
<taxonomicName id="41C14D59C80FFFCAFC6DFB24237A4382" box="[965,1126,1201,1224]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFC6DFB24237A4382" box="[965,1126,1201,1224]" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
vs 56–65 × 38–45 µm in 
<taxonomicName id="41C14D59C80FFFCAFA2FFB24260F43A6" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80FFFCAFA2FFB2422894382" box="[1415,1429,1201,1224]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFF3FFB40260F43A6" box="[151,275,1237,1260]" italics="true" pageId="8" pageNumber="575">aegyptense</emphasis>
</taxonomicName>
), sucker ratio (1:0.73–1:0.74 vs 1:0.59–1:0.94), possessing a sinistrally submedian and post-bifurcal genital pore, and both have a similar vitelline distribution (
<tableCitation id="CB430361C80FFFCAFCC0FB6D24DA425A" box="[872,966,1272,1297]" captionStart="TABLE 1" captionStartId="4.[151,239,368,391]" captionTargetBox="[159,1427,486,1986]" captionTargetPageId="4" captionText="TABLE 1. Measurements, morphometric percentages and morphometric ratios of Pelopscreadium spongiosum (Bray &amp; Cribb, 1998) n. gen., n. comb. and Pelopscreadium aegyptense n. sp. from the yellow boxfish, Ostracion cubicus Linnaeus, 1758 (Tetraodontiformes: Ostraciidae), with holotype data followed by the ranges and means in parentheses." httpUri="http://table.plazi.org/id/D2BE6652C803FFC6FF3FFEE522974680" pageId="8" pageNumber="575" tableUuid="D2BE6652C803FFC6FF3FFEE522974680">Table 1</tableCitation>
; 
<bibRefCitation id="E2504B2BC80FFFCAFC72FB6C23D1425A" author="Bray" box="[986,1229,1272,1297]" pageId="8" pageNumber="575" refString="Bray, R. A. &amp; Cribb, T. H. (1998) Lepocreadiidae (Digenea) of Australian coastal fishes: new species of Opechona Looss, 1907, Lepotrema Ozaki, 1932 and Bianium Stunkard, 1930 and comments on other species reported for the first time or poorly known in Australian waters. Systematic Parasitology, 41, 123 - 148." type="journal article" year="1998">Bray &amp; Cribb 1998</bibRefCitation>
). 
<taxonomicName id="41C14D59C80FFFCAFB4FFB6D2608427E" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80FFFCAFB4FFB6D2608427E" italics="true" pageId="8" pageNumber="575">Pelopscreadium aegyptense</emphasis>
</taxonomicName>
and 
<taxonomicName id="41C14D59C80FFFCAFEFEFA8826E2427E" box="[342,510,1309,1332]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFEFEFA8826E2427E" box="[342,510,1309,1332]" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
differ in a number of features, the most striking of which is the extent and appearance of the external seminal vesicle and the distribution of the contents of the cirrus-sac. Within the cirrussac, the pars prostatica of 
<taxonomicName id="41C14D59C80FFFCAFE63FAF0256C4236" box="[459,624,1381,1404]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFE63FAF0256C4236" box="[459,624,1381,1404]" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
was described as “oval, vesicular”; whereas, the pars prostatica of 
<taxonomicName id="41C14D59C80FFFCAFA2FFAF0260842EA" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80FFFCAFA2FFAF022894236" box="[1415,1429,1381,1404]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFF3FFA1C260842EA" box="[151,276,1417,1440]" italics="true" pageId="8" pageNumber="575">aegyptense</emphasis>
</taxonomicName>
is more elongate with a conspicuously longer longitudinal axis (about 63 vs 89–165 µm) and it generally occupies a greater portion of the cirrus-sac (23% vs 26–41% of the cirrus-sac length, respectively). The external seminal vesicle of 
<taxonomicName id="41C14D59C80FFFCAFE70FA44256142A2" box="[472,637,1489,1512]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFE70FA44256142A2" box="[472,637,1489,1512]" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="E2504B2BC80FFFCAFCF2FA44221D42A2" author="Bray" box="[858,1281,1488,1513]" pageId="8" pageNumber="575" refString="Bray, R. A. &amp; Cribb, T. H. (1998) Lepocreadiidae (Digenea) of Australian coastal fishes: new species of Opechona Looss, 1907, Lepotrema Ozaki, 1932 and Bianium Stunkard, 1930 and comments on other species reported for the first time or poorly known in Australian waters. Systematic Parasitology, 41, 123 - 148." type="journal article" year="1998">Bray &amp; Cribb (1998, figs. 18 &amp; 20)</bibRefCitation>
as “saccular, reaching to about level of ovary”, and it is illustrated as a simple, moderately-wide, oblong pouch (about 150 µm long—see 
<tableCitation id="CB430361C80FFFCAFEB8F98D267E417A" box="[272,354,1560,1585]" captionStart="TABLE 1" captionStartId="4.[151,239,368,391]" captionTargetBox="[159,1427,486,1986]" captionTargetPageId="4" captionText="TABLE 1. Measurements, morphometric percentages and morphometric ratios of Pelopscreadium spongiosum (Bray &amp; Cribb, 1998) n. gen., n. comb. and Pelopscreadium aegyptense n. sp. from the yellow boxfish, Ostracion cubicus Linnaeus, 1758 (Tetraodontiformes: Ostraciidae), with holotype data followed by the ranges and means in parentheses." httpUri="http://table.plazi.org/id/D2BE6652C803FFC6FF3FFEE522974680" pageId="8" pageNumber="575" tableUuid="D2BE6652C803FFC6FF3FFEE522974680">Table 1</tableCitation>
). However, the external seminal vesicle of 
<taxonomicName id="41C14D59C80FFFCAFCE6F98C24FB417A" box="[846,999,1561,1584]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80FFFCAFCE6F98C2440417A" box="[846,860,1561,1584]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFCC2F98C24FB417A" box="[874,999,1561,1584]" italics="true" pageId="8" pageNumber="575">aegyptense</emphasis>
</taxonomicName>
is tubular, considerably more elongate and sinuous, and longer than the cirrus-sac (485–540 compared to 150 µm). Additionally, the cirrus-sac of 
<taxonomicName id="41C14D59C80FFFCAFA2FF9A826084132" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80FFFCAFA2FF9A82289411E" box="[1415,1429,1597,1620]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFF3FF9F426084132" box="[151,276,1633,1656]" italics="true" pageId="8" pageNumber="575">aegyptense</emphasis>
</taxonomicName>
ranges from terminating at the level of the posterior margin of the ventral sucker to surpassing the ventral sucker a relatively short distance into the hindbody; whereas, in 
<taxonomicName id="41C14D59C80FFFCAFC7FF910236641D6" box="[983,1146,1669,1692]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFC7FF910236641D6" box="[983,1146,1669,1692]" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
it extends some distance into the hindbody (about half the length of the distance between the ventral sucker and the ovary). Another difference between these two species is the appearance of the testes. In 
<taxonomicName id="41C14D59C80FFFCAFC7BF958237341AE" box="[979,1135,1741,1764]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80FFFCAFC7BF95824FD41AE" box="[979,993,1741,1764]" italics="true" pageId="8" pageNumber="575">P</emphasis>
. 
<emphasis id="B4B5EAC8C80FFFCAFC5AF958237341AE" box="[1010,1135,1741,1764]" italics="true" pageId="8" pageNumber="575">aegyptense</emphasis>
</taxonomicName>
the testes are consistently irregular in shape and slightly to deeply lobed, but in 
<taxonomicName id="41C14D59C80FFFCAFD42F96424974042" box="[746,907,1777,1800]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="8" pageNumber="575" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80FFFCAFD42F96424974042" box="[746,907,1777,1800]" italics="true" pageId="8" pageNumber="575">P. spongiosum</emphasis>
</taxonomicName>
they are subglobular and entire (smooth) (
<bibRefCitation id="E2504B2BC80FFFCAFACCF96526834066" author="Bray" pageId="8" pageNumber="575" refString="Bray, R. A. &amp; Cribb, T. H. (1998) Lepocreadiidae (Digenea) of Australian coastal fishes: new species of Opechona Looss, 1907, Lepotrema Ozaki, 1932 and Bianium Stunkard, 1930 and comments on other species reported for the first time or poorly known in Australian waters. Systematic Parasitology, 41, 123 - 148." type="journal article" year="1998">Bray &amp; Cribb 1998, fig. 18</bibRefCitation>
). Although any differences in testicular shape (entire vs lobed) may simply reflect the potentially “plastic” nature of this morphological feature within many groups of digeneans (e.g. see 
<bibRefCitation id="E2504B2BC80FFFCAFAAFF8AC27CE403E" author="Dronen" pageId="8" pageNumber="575" refString="Dronen, N. O., Blend, C. K., Ostrowski de Nunez, M. C., Malhotra, S. K. &amp; Jaiswal, N. (2014) Keys to the species of Neolebouria Gibson, 1976 (Digenea: Opecoelidae: Plagioporinae), with a redescription of Neolebouria truncata (Linton, 1940). Systematic Parasitology, 88, 213 - 225." type="journal article" year="2014">
Dronen 
<emphasis id="B4B5EAC8C80FFFCAFACBF8AC2289401B" box="[1379,1429,1848,1873]" italics="true" pageId="8" pageNumber="575">et al</emphasis>
. 2014
</bibRefCitation>
, for a discussion of the difficulties in using testicular shape as a distinguishing taxonomic feature for species within the opecoelid genus 
<taxonomicName id="41C14D59C80FFFCAFE67F81525E140D3" ID-CoL="7PQ3Q" authority="Gibson, 1976" authorityName="Gibson" authorityYear="1976" box="[463,765,1920,1945]" class="Trematoda" family="Opecoelidae" genus="Neolebouria" kingdom="Animalia" order="Plagiorchiida" pageId="8" pageNumber="575" phylum="Platyhelminthes" rank="genus">
<emphasis id="B4B5EAC8C80FFFCAFE67F815254740D3" box="[463,603,1920,1945]" italics="true" pageId="8" pageNumber="575">Neolebouria</emphasis>
<bibRefCitation id="E2504B2BC80FFFCAFDCBF81525E140D3" author="Gibson" box="[611,765,1920,1945]" pageId="8" pageNumber="575" refString="Gibson, D. I. (1976) Monogenea and Digenea from fishes. Discovery Reports, 36, 179 - 266." type="journal article" year="1976">Gibson, 1976</bibRefCitation>
</taxonomicName>
) and on its own may not be a “strong enough character” to distinguish species, it is likely that in this particular instance, this feature may be more taxonomically (phylogenetically) informative than in some other digenean groups. We also observed that although the body lengths of 
<taxonomicName id="41C14D59C80EFFCBFEA5FF0C26BB47FA" box="[269,423,152,176]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="9" pageNumber="576" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80EFFCBFEA5FF0C260747FA" box="[269,283,153,176]" italics="true" pageId="9" pageNumber="576">P</emphasis>
. 
<emphasis id="B4B5EAC8C80EFFCBFE82FF0C26BB47FA" box="[298,423,153,176]" italics="true" pageId="9" pageNumber="576">aegyptense</emphasis>
</taxonomicName>
and 
<taxonomicName id="41C14D59C80EFFCBFE75FF0C256347FA" box="[477,639,153,176]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="9" pageNumber="576" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80EFFCBFE75FF0C256347FA" box="[477,639,153,176]" italics="true" pageId="9" pageNumber="576">P. spongiosum</emphasis>
</taxonomicName>
overlap (
<tableCitation id="CB430361C80EFFCBFD41FF02242147FA" box="[745,829,151,176]" captionStart="TABLE 1" captionStartId="4.[151,239,368,391]" captionTargetBox="[159,1427,486,1986]" captionTargetPageId="4" captionText="TABLE 1. Measurements, morphometric percentages and morphometric ratios of Pelopscreadium spongiosum (Bray &amp; Cribb, 1998) n. gen., n. comb. and Pelopscreadium aegyptense n. sp. from the yellow boxfish, Ostracion cubicus Linnaeus, 1758 (Tetraodontiformes: Ostraciidae), with holotype data followed by the ranges and means in parentheses." httpUri="http://table.plazi.org/id/D2BE6652C803FFC6FF3FFEE522974680" pageId="9" pageNumber="576" tableUuid="D2BE6652C803FFC6FF3FFEE522974680">Table 1</tableCitation>
), specimens of 
<taxonomicName id="41C14D59C80EFFCBFC58FF0C239547FA" box="[1008,1161,153,176]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="9" pageNumber="576" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80EFFCBFC58FF0C239547FA" box="[1008,1161,153,176]" italics="true" pageId="9" pageNumber="576">P. aegyptense</emphasis>
</taxonomicName>
tend to be larger overall (1,352–1,412 × 438–489 vs 1,160–2,642 × 775–1,356 µm). The extent of the metraterm relative to body length differed between these two species (about 75 µm, or 5% the length of the body in 
<taxonomicName id="41C14D59C80EFFCBFB80FF7423D547B2" box="[1064,1225,225,248]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="9" pageNumber="576" phylum="Chordata" rank="species" species="spongiosum">
<emphasis id="B4B5EAC8C80EFFCBFB80FF7423D547B2" box="[1064,1225,225,248]" italics="true" pageId="9" pageNumber="576">P. spongiosum</emphasis>
</taxonomicName>
vs 180–335 µm or 7–16% in 
<taxonomicName id="41C14D59C80EFFCBFEBAFE9026B14656" box="[274,429,261,284]" class="Actinopterygii" family="Ostraciidae" genus="Pelopscreadium" kingdom="Animalia" order="Tetraodontiformes" pageId="9" pageNumber="576" phylum="Chordata" rank="species" species="aegyptense">
<emphasis id="B4B5EAC8C80EFFCBFEBAFE9026B14656" box="[274,429,261,284]" italics="true" pageId="9" pageNumber="576">P. aegyptense</emphasis>
</taxonomicName>
), and while both species parasitize the yellow boxfish, 
<taxonomicName id="41C14D59C80EFFCBFB97FE9023A64657" ID-CoL="6TD5R" box="[1087,1210,260,285]" class="Actinopterygii" family="Ostraciidae" genus="Ostracion" kingdom="Animalia" order="Tetraodontiformes" pageId="9" pageNumber="576" phylum="Chordata" rank="species" species="cubicus">
<emphasis id="B4B5EAC8C80EFFCBFB97FE9023A64657" box="[1087,1210,260,285]" italics="true" pageId="9" pageNumber="576">
<collectingCountry id="FED6764AC80EFFCBFB97FE90234A4656" box="[1087,1110,261,284]" name="Iceland" pageId="9" pageNumber="576">O</collectingCountry>
. cubicus
</emphasis>
</taxonomicName>
, their geographical localities are considerably distant (
<collectingCountry id="FED6764AC80EFFCBFDB3FEB22598460A" box="[539,644,295,320]" name="Australia" pageId="9" pageNumber="576">Australia</collectingCountry>
vs Red Sea off 
<collectingCountry id="FED6764AC80EFFCBFC9EFEBD2462460A" box="[822,894,296,320]" name="Egypt" pageId="9" pageNumber="576">Egypt</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>