130 lines
17 KiB
XML
130 lines
17 KiB
XML
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<mods:title id="B089B099A35B279B6B617C92FBD52789">The weevil genera Nyphaeba Pascoe and Pantiala Pascoe and the problems of an unstable nomenclature in orphaned taxa</mods:title>
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<mods:name id="4E681A4E133540F5137AAA3EE7565D31" type="personal">
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<mods:namePart id="84FBB97FE42215B93658425C0C46F4B6">Riedel, Alexander</mods:namePart>
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<mods:title id="0E834EFFAE572559837FB23484E69243">Zootaxa</mods:title>
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<mods:date id="A1E54A7951E27287BA8CDC6CD60C74F4">2017</mods:date>
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<mods:number id="5396764CE6042B78E76C37E2ABC38DC2">4244</mods:number>
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<mods:number id="549EDFFE4DA11EEA3908BCB5B9E7F762">3</mods:number>
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<mods:identifier id="DEE866DE44759C5B87539855DA583F74" type="DOI">10.11646/zootaxa.4244.3.6</mods:identifier>
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<treatment id="03F287B6067AFF9C54F3CDB9FB392DE3" ID-DOI="http://doi.org/10.5281/zenodo.6041777" ID-GBIF-Taxon="127848133" ID-Zenodo-Dep="6041777" LSID="urn:lsid:plazi:treatment:03F287B6067AFF9C54F3CDB9FB392DE3" httpUri="http://treatment.plazi.org/id/03F287B6067AFF9C54F3CDB9FB392DE3" lastPageNumber="384" pageId="7" pageNumber="384">
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<subSubSection id="C341652B067AFF9C54F3CDB9FEF528CE" box="[151,345,151,178]" pageId="7" pageNumber="384" type="nomenclature">
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<paragraph id="8BE436A0067AFF9C54F3CDB9FEF528CE" blockId="7.[151,345,151,178]" box="[151,345,151,178]" pageId="7" pageNumber="384">
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<heading id="D0AC81CC067AFF9C54F3CDB9FEF528CE" bold="true" box="[151,345,151,178]" fontSize="11" level="1" pageId="7" pageNumber="384" reason="1">
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<taxonomicName id="4C5B4D23067AFF9C54F3CDB9FEF528CE" ID-CoL="8HNRV" authority="Pascoe" authorityName="Pascoe" box="[151,345,151,178]" class="Insecta" family="Curculionidae" genus="Pantiala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067AFF9C54F3CDB9FEF528CE" bold="true" box="[151,345,151,178]" pageId="7" pageNumber="384">
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<emphasis id="B92FEAB2067AFF9C54F3CDB9FF5228CD" bold="true" box="[151,254,151,177]" italics="true" pageId="7" pageNumber="384">Pantiala</emphasis>
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Pascoe
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C341652B067AFF9C54F3CDF2FE18288E" box="[151,436,220,242]" pageId="7" pageNumber="384" type="reference_group">
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<paragraph id="8BE436A0067AFF9C54F3CDF2FE18288E" blockId="7.[151,788,220,272]" box="[151,436,220,242]" pageId="7" pageNumber="384">
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<treatmentCitationGroup id="AB4B118E067AFF9C54F3CDF2FE18288E" box="[151,436,220,242]" pageId="7" pageNumber="384">
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<taxonomicName id="4C5B4D23067AFF9C54F3CDF2FE1C288E" ID-CoL="8HNRV" authority="Pascoe, 1885: 284" authorityName="Pascoe" authorityPageNumber="284" authorityYear="1885" box="[151,432,220,242]" class="Insecta" family="Curculionidae" genus="Pantiala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067AFF9C54F3CDF2FF42288E" box="[151,238,220,242]" italics="true" pageId="7" pageNumber="384">Pantiala</emphasis>
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<treatmentCitation id="0AFA10B1067AFF9C5490CDF3FE1C288E" author="Pascoe" box="[244,432,220,242]" page="284" pageId="7" pageNumber="384" year="1885">
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<bibRefCitation id="EFCA4B51067AFF9C5490CDF3FED1288E" author="Pascoe" box="[244,381,220,242]" pageId="7" pageNumber="384" refString="Pascoe, F. P. (1885) List of the Curculionidae of the Malay Archipelago collected by Dr. Odoardo Beccari, L. M. D'Albertis, and others. Annali del Museo Civico di Storia Naturale di Genova, 22, 201 - 332, pls. I - III." type="journal article" year="1885">Pascoe, 1885</bibRefCitation>
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: 284
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C341652B067AFF9C54F3CDD5FCB8296C" box="[151,788,250,272]" pageId="7" pageNumber="384" type="materials_examined">
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<paragraph id="8BE436A0067AFF9C54F3CDD5FCB8296C" blockId="7.[151,788,220,272]" box="[151,788,250,272]" pageId="7" pageNumber="384">
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Type species:
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<taxonomicName id="4C5B4D23067AFF9C5542CDD4FDD2296C" authority="Pascoe, 1885: 284" authorityName="Pascoe" authorityPageNumber="284" authorityYear="1885" box="[294,638,250,272]" class="Insecta" family="Curculionidae" genus="Pantiala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="illusa">
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<emphasis id="B92FEAB2067AFF9C5542CDD4FE17296C" box="[294,443,250,272]" italics="true" pageId="7" pageNumber="384">Pantiala illusa</emphasis>
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<bibRefCitation id="EFCA4B51067AFF9C55A6CDD5FDE7296C" author="Pascoe" box="[450,587,250,272]" pageId="7" pageNumber="384" refString="Pascoe, F. P. (1885) List of the Curculionidae of the Malay Archipelago collected by Dr. Odoardo Beccari, L. M. D'Albertis, and others. Annali del Museo Civico di Storia Naturale di Genova, 22, 201 - 332, pls. I - III." type="journal article" year="1885">Pascoe, 1885</bibRefCitation>
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: 284
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C341652B067AFF9C54F3CC14FB392DE3" pageId="7" pageNumber="384" type="description">
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<paragraph id="8BE436A0067AFF9C54F3CC14FD2B2C8B" blockId="7.[151,1437,314,1439]" pageId="7" pageNumber="384">
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<emphasis id="B92FEAB2067AFF9C54F3CC14FE91292F" bold="true" box="[151,317,314,339]" pageId="7" pageNumber="384">Redescription</emphasis>
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. Length
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<quantity id="4CA39B45067AFF9C55C2CC14FDE2292E" box="[422,590,314,339]" metricMagnitude="-3" metricUnit="m" metricValue="4.22" metricValueMax="4.64" metricValueMin="3.8" pageId="7" pageNumber="384" unit="mm" value="4.22" valueMax="4.64" valueMin="3.8">3.80–4.64 mm</quantity>
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. Habitus with sides of pronotum and elytra converging to apex; in profile relatively broad, dorsally weakly convex, elytral apex steeply declivous. Integument largely subglabrous, with small patches of scales. Rostrum short, 1.3 X as long as wide before apex; ca. 0.46 X length of pronotum; sides diverging to shortly before apex; in profile weakly curved, weakly widening to shortly before apex; in both sexes dorsal surface without carinae, densely punctate. Antennae inserted at middle. Scrobe leading ventrad, keeping distance from eye. Scape shorter than funicle, subglabrous; funicle with sparse thin setae, articles 6–7 each wider than long. Club subovate. Eyes with subrotund outline, in dorsal position, medially approximate, separated 0.4 X basal width of rostrum. Pronotum 1.06 X wider than long; widest at base, sides subparallel in basal half, markedly converging in apical half; without subapical constriction. Postocular lobe indistinct, with fringe of setae. Scutellar shield distinct, subglabrous. Elytron with 10 distinct striae; stria 10 complete; intervals subglabrous except subapical patch of white scales; base markedly sinuate; subapically on ventral side with stridulatory file. Wings fully developed. Thoracic venter (
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<figureCitation id="13602A25067AFF9C5651CF87FD3D2ABE" box="[565,657,681,706]" captionStart="FIGURE 5" captionStartId="9.[151,250,1927,1949]" captionTargetBox="[195,1349,303,1889]" captionTargetId="figure@9.[195,1392,303,1902]" captionTargetPageId="9" captionText="FIGURE 5 a, c – d. Nyphaeba mimica sp. n., holotype; (a) venter (c) detail of ventrite 1 with row of short ridges marked by arrows (d) posterior surface of metafemur with patch of ridges, presumably acting as stridulatory device together with structures shown in (c). Fig. 5 b. Pantiala illusa Pascoe, venter." httpUri="https://zenodo.org/record/428754/files/figure.png" pageId="7" pageNumber="384">Fig. 5b</figureCitation>
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). Pronotum forming acute process above procoxa. Mesothoracic receptacle ca.
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<date id="FFE51060067AFF9C5553CFE5FECD2A9E" box="[311,353,715,739]" pageId="7" pageNumber="384">2 X</date>
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wider than long, cavernous, internally glabrous; ventral rim produced, covered with white erect scales, anterior margin subtruncate. Metaventrite at middle concave; sublaterally with oblique intercoxal ridge continuing in front of metacoxa, forming glabrous cavity. Metanepisternum distinct, squamose unless abraded; anapleural suture complete, without sclerolepidia. Legs. Femora ventrally sulcate; anteroventral ridge with sharp tooth slightly apicad of middle. Tibiae with distinct dorsal carina terminating at subbasal angulation; apically with strong uncus, without premucro. Tarsi with third tarsomere 2.0 X as wide as second tarsomere. Abdominal venter. Ventrite 1 anteriorly concave; intercoxal process subangulate, wider than long; posteriorly markedly convex together with ventrite 2. Ventrite 5 ca. 2.4 X as wide as long.
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<emphasis id="B92FEAB2067AFF9C573ACE9AFB9C2BB1" box="[862,1072,948,973]" italics="true" pageId="7" pageNumber="384">Female terminalia</emphasis>
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. Tergite VII slightly wider than long, subtrapezoid; basal margin weakly convex; apical margin subtruncate; with one subapical pair of microsetae. Tergite VIII ca. as long as wide; in apical half subangulate, margin simple, not crenulate. Sternite VIII with apical plate wider than long. Styli long, slender. Bursa without sclerite. Spermathecal duct arising on bursa away from junction with oviduct.
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<emphasis id="B92FEAB2067AFF9C55CCC917FDCF2C2E" box="[424,611,1081,1106]" italics="true" pageId="7" pageNumber="384">Male terminalia</emphasis>
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. Tergite VII wider than long, subtrapezoid; subapically with pair of microsetae surrounded by clusters of other setae. Tergite VIII apically subtruncate, outline subquadrate. Sternum VIII with hemisternites large, medially connected by narrow, weakly sclerotized part. Spiculum gastrale subequal in length to apodemes of penis, straight. Tegmen with parameres very short; apodeme slightly longer than width of tegminal ring. Body of penis subequal to length of apodemes, lanceolate in dorsal aspect. Endophallus without sclerites; without distinct transfer apparatus.
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</paragraph>
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<paragraph id="8BE436A0067AFF9C54A3C82FFB392DE3" blockId="7.[151,1437,314,1439]" pageId="7" pageNumber="384">
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<emphasis id="B92FEAB2067AFF9C54A3C82FFEA62D66" bold="true" box="[199,266,1281,1306]" pageId="7" pageNumber="384">Notes</emphasis>
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.
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<taxonomicName id="4C5B4D23067AFF9C5578C82FFED12D66" box="[284,381,1281,1306]" class="Insecta" family="Curculionidae" genus="Pantiala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067AFF9C5578C82FFED12D66" box="[284,381,1281,1306]" italics="true" pageId="7" pageNumber="384">Pantiala</emphasis>
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</taxonomicName>
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has well-developed wings and metanepisterna and therefore it needs to be removed from
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<taxonomicName id="4C5B4D23067AFF9C54F3C80CFE2F2D47" authority="Lacordaire" authorityName="Lacordaire" box="[151,387,1314,1339]" class="Gastropoda" family="Tylodinidae" genus="Tylodina" kingdom="Animalia" order="Umbraculida" pageId="7" pageNumber="384" phylum="Mollusca" rank="genus">Tylodina Lacordaire</taxonomicName>
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of
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<taxonomicName id="4C5B4D23067AFF9C55DEC80CFCBF2D47" authority="Schoenherr" authorityName="Schoenherr" box="[442,787,1314,1339]" class="Insecta" family="Curculionidae" genus="Cryptorhynchinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">Cryptorhynchinae Schoenherr</taxonomicName>
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and placed in
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<taxonomicName id="4C5B4D23067AFF9C57BEC80CFB082D47" box="[986,1188,1314,1339]" class="Insecta" family="Curculionidae" genus="Cryptorhynchinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="genus" status="incertae sedis">Cryptorhynchinae</taxonomicName>
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<taxonomicNameLabel id="A21C57C9067AFF9C50D0C80CFAF82D47" box="[1204,1364,1314,1339]" pageId="7" pageNumber="384">incertae sedis</taxonomicNameLabel>
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(new placement). In his original description,
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<bibRefCitation id="EFCA4B51067AFF9C5629C86BFD402D21" author="Pascoe" box="[589,748,1348,1373]" pageId="7" pageNumber="384" refString="Pascoe, F. P. (1885) List of the Curculionidae of the Malay Archipelago collected by Dr. Odoardo Beccari, L. M. D'Albertis, and others. Annali del Museo Civico di Storia Naturale di Genova, 22, 201 - 332, pls. I - III." type="journal article" year="1885">Pascoe (1885)</bibRefCitation>
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stresses the short, thick rostrum and this concept of the genus is consistent with the present lectotype designation. In the phylogeny of
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<bibRefCitation id="EFCA4B51067AFF9C57B8C84BFB112D02" box="[988,1213,1381,1406]" pageId="7" pageNumber="384" refString="Riedel, A., Tanzler, R., Pons, J., Suhardjono, Y. R. & Balke, M. (2016) Large-scale molecular phylogeny of Cryptorhynchinae (Coleoptera, Curculionidae) from multiple genes suggests American origin and later Australian radiation. Systematic Entomology, 41 (2), 492 - 503." type="journal article">
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Riedel
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<emphasis id="B92FEAB2067AFF9C504AC848FBC52D02" box="[1070,1129,1381,1406]" italics="true" pageId="7" pageNumber="384">et al.</emphasis>
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(2016)
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</bibRefCitation>
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<taxonomicName id="4C5B4D23067AFF9C50A2C84BFA8B2D02" box="[1222,1319,1381,1406]" class="Insecta" family="Curculionidae" genus="Pantiala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067AFF9C50A2C84BFA8B2D02" box="[1222,1319,1381,1406]" italics="true" pageId="7" pageNumber="384">Pantiala</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName id="4C5B4D23067AFF9C5563C8A9FE652DE2" authority="Pascoe" authorityName="Pascoe" box="[263,457,1415,1438]" class="Insecta" family="Curculionidae" genus="Perissops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067AFF9C5563C8A9FED82DE2" box="[263,372,1415,1438]" italics="true" pageId="7" pageNumber="384">Perissops</emphasis>
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Pascoe
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</taxonomicName>
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and
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<taxonomicName id="4C5B4D23067AFF9C559BC8A9FD1E2DE2" authority="Lea" authorityName="Lea" box="[511,690,1415,1438]" class="Insecta" family="Curculionidae" genus="Critomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAB2067AFF9C559BC8A9FDD32DE2" box="[511,639,1415,1438]" italics="true" pageId="7" pageNumber="384">Critomerus</emphasis>
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Lea
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</taxonomicName>
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, genera that share its subovate body shape.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |