207 lines
24 KiB
XML
207 lines
24 KiB
XML
<document id="19D2F7913C2FBF529A3CBC85380CAB1D" ID-DOI="10.11646/zootaxa.4048.4.1" ID-GBIF-Dataset="bbb51c92-e184-489d-aba8-18d1dc9a25cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="234023" ID-ZooBank="71E2B8DA-825E-429F-BAA0-F582702B4A80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461348015003" checkinUser="plazi" docAuthor="López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán" docDate="2015" docId="990DA529FF82FF9AFF46405F17FFFD2B" docLanguage="en" docName="zt04048p492.pdf" docOrigin="Zootaxa 4048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomarus cicatricosus Prell 1937" docType="treatment" docVersion="11" lastPageNumber="480" masterDocId="6534DD51FF9EFF87FFD1425F1659FFE6" masterDocTitle="The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution" masterLastPageNumber="492" masterPageNumber="451" pageNumber="479" updateTime="1698621022009" updateUser="plazi">
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<mods:title id="5031E99E6C13E0B514F2F8E9C6B6377E">The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution</mods:title>
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<mods:name id="F221923DA610CEF75BD159089B04CACB" type="personal">
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<mods:namePart id="8281BD21AAF4CDC2DC6415D4BD30D6E9">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="A88B48A0D8C07FABF093CCAEA15FA6B6">Gasca-Álvarez, Héctor J.</mods:namePart>
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<mods:roleTerm id="CC4B08369DC53431D2F22F2D609FECE7">Author</mods:roleTerm>
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<mods:namePart id="A9FE6A39FD594113A065711BFA007D08">Amat-García, Germán</mods:namePart>
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<mods:title id="BF634E175A91636EE19C95F7C5F0F145">Zootaxa</mods:title>
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<mods:date id="13C8CEB90EDA22B76E4FB91378FFB589">2015</mods:date>
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<mods:number id="23CACFD4EEE2042A3320E2DF30870269">4048</mods:number>
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<mods:identifier id="DF13D7C624EB49238D684B91FA22F8B2" type="DOI">10.11646/zootaxa.4048.4.1</mods:identifier>
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<treatment id="990DA529FF82FF9AFF46405F17FFFD2B" ID-DOI="http://doi.org/10.5281/zenodo.5699386" ID-GBIF-Taxon="119645674" ID-Zenodo-Dep="5699386" LSID="urn:lsid:plazi:treatment:990DA529FF82FF9AFF46405F17FFFD2B" httpUri="http://treatment.plazi.org/id/990DA529FF82FF9AFF46405F17FFFD2B" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479">
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<subSubSection id="59BE47B4FF82FF9BFF46405F1710FDDD" pageId="28" pageNumber="479" type="nomenclature">
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<paragraph id="111B143FFF82FF9BFF46405F146EFDFC" blockId="28.[151,567,512,571]" box="[151,567,512,538]" pageId="28" pageNumber="479">
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<heading id="4A53A353FF82FF9BFF46405F146EFDFC" bold="true" box="[151,567,512,538]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
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<taxonomicName id="D6A46FBCFF82FF9BFF46405F146EFDFC" authority="Prell, 1937" authorityName="Prell" authorityYear="1937" box="[151,567,512,538]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="cicatricosus">
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<emphasis id="23D0C82DFF82FF9BFF46405F146EFDFC" bold="true" box="[151,567,512,538]" pageId="28" pageNumber="479">
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<emphasis id="23D0C82DFF82FF9BFF46405F17C1FDFC" bold="true" box="[151,408,512,538]" italics="true" pageId="28" pageNumber="479">Tomarus cicatricosus</emphasis>
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(
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<bibRefCitation id="753569CEFF82FF9BFE76405F1476FDFC" author="Prell" box="[423,559,512,538]" pageId="28" pageNumber="479" refString="Prell, H. (1937) Beitrage zur Kenntnis der Dynastiden. XV. 3. Neue Arten und Rassen. Entomologische Zeitschrift, Frankfurt, 51, 89 - 90." type="journal article" year="1937">Prell, 1937</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="111B143FFF82FF9BFF46407D1710FDDD" blockId="28.[151,567,512,571]" box="[151,329,546,571]" pageId="28" pageNumber="479">
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(
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<figureCitation id="899F08BAFF82FF9BFF4E407D1718FDDD" box="[159,321,546,571]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="28" pageNumber="479">
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Figs. 102
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<emphasis id="23D0C82DFF82FF9BFED9407D174FFDDD" bold="true" box="[264,278,546,571]" pageId="28" pageNumber="479">–</emphasis>
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||
108
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||
</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59BE47B4FF82FF9BFF4640341451FD66" box="[151,520,618,640]" pageId="28" pageNumber="479" type="reference_group">
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||
<paragraph id="111B143FFF82FF9BFF4640341451FD66" blockId="28.[151,520,618,640]" box="[151,520,618,640]" pageId="28" pageNumber="479">
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<treatmentCitationGroup id="31B43311FF82FF9BFF4640341451FD66" box="[151,520,618,640]" pageId="28" pageNumber="479">
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<taxonomicName id="D6A46FBCFF82FF9BFF464034145DFD66" ID-CoL="7ZP6V" authority="Prell, 1937: 90" authorityName="Prell" authorityPageNumber="90" authorityYear="1937" box="[151,516,618,640]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="cicatricosus">
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<emphasis id="23D0C82DFF82FF9BFF464034173CFD66" box="[151,357,619,640]" italics="true" pageId="28" pageNumber="479">Ligyrus cicatricosus</emphasis>
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<treatmentCitation id="9005322EFF82FF9BFEBA4035145DFD66" author="Prell" box="[363,516,618,640]" page="90" pageId="28" pageNumber="479" year="1937">
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<bibRefCitation id="753569CEFF82FF9BFEBA40351787FD66" author="Prell" box="[363,478,618,640]" pageId="28" pageNumber="479" refString="Prell, H. (1937) Beitrage zur Kenntnis der Dynastiden. XV. 3. Neue Arten und Rassen. Entomologische Zeitschrift, Frankfurt, 51, 89 - 90." type="journal article" year="1937">Prell, 1937</bibRefCitation>
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: 90
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</treatmentCitation>
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</taxonomicName>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<caption id="45DB44B7FF82FF9BFF4647E4136DFA16" httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="28" pageNumber="479" targetBox="[157,1429,699,1441]" targetPageId="28">
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<paragraph id="111B143FFF82FF9BFF4647E4136DFA16" blockId="28.[151,1435,1467,1520]" pageId="28" pageNumber="479">
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<emphasis id="23D0C82DFF82FF9BFF4647E41729FA37" bold="true" box="[151,368,1467,1490]" pageId="28" pageNumber="479">FIGURES 102–108.</emphasis>
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<taxonomicName id="D6A46FBCFF82FF9BFEA847E2140DFA34" box="[377,596,1469,1490]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="cicatricosus">
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<emphasis id="23D0C82DFF82FF9BFEA847E2140DFA34" box="[377,596,1469,1490]" italics="true" pageId="28" pageNumber="479">Tomarus cicatricosus</emphasis>
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</taxonomicName>
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. 102) habitus in dorsal view, 103) habitus in frontal view, 104)
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<taxonomicName id="D6A46FBCFF82FF9BFB2C47E31338FA34" box="[1277,1377,1468,1490]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="488" phylum="Chordata" rank="genus">pygidium</taxonomicName>
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, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia.
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</paragraph>
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</caption>
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<subSubSection id="59BE47B4FF82FF9AFF46444215F9FEA6" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479" type="description">
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<paragraph id="111B143FFF82FF9AFF46444215F9FEA6" blockId="28.[151,1437,1564,2022]" lastBlockId="29.[151,1437,151,717]" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479">
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<emphasis id="23D0C82DFF82FF9BFF4644421770F9D0" bold="true" box="[151,297,1565,1590]" pageId="28" pageNumber="479">Description.</emphasis>
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Habitus as in
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<figureCitation id="899F08BAFF82FF9BFE1644421471F9D0" box="[455,552,1565,1590]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="28" pageNumber="479">Fig. 102</figureCitation>
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. Length
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<quantity id="D65CB9DAFF82FF9BFD5944411483F9D3" box="[648,730,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="479" unit="mm" value="24.0">24 mm</quantity>
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. Width
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<quantity id="D65CB9DAFF82FF9BFCE144411527F9D3" box="[816,894,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="28" pageNumber="479" unit="mm" value="13.0">13 mm</quantity>
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(♂). Color black, procoxae lighter.
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<emphasis id="23D0C82DFF82FF9BFADC4443130CF9D0" bold="true" box="[1293,1365,1564,1590]" pageId="28" pageNumber="479">
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||
<emphasis id="23D0C82DFF82FF9BFADC44431312F9D3" bold="true" box="[1293,1355,1564,1589]" italics="true" pageId="28" pageNumber="479">Head</emphasis>
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:
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</emphasis>
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Frons slightly concave between eyes and behind frontoclypeal tubercles; surface rugose and grooved, nearly smooth on vertex and only with deep, sparse punctures. Frontoclypeal region with 2 transverse tubercles broadly separated from one another (about 9 tubercle diameters). Frontoclypeal suture present, as a weak carina connecting frontal tubercles. Clypeus with surface transversely rugose; apex broadly truncate, with 2 small teeth; teeth reflexed, widely separated (about 5 tooth diameters). Interocular distance equals 3.3 times the transverse ocular diameter. Antennal club subequal in length to antennomeres 2–7. Mandibles with 2 apical and one lateral tooth.
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<emphasis id="23D0C82DFF82FF9BFF0F45461705F8D6" bold="true" box="[222,348,1817,1840]" italics="true" pageId="28" pageNumber="479">Pronotum:</emphasis>
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Surface with punctures evenly sparse, deep, large, weakly umbilicale punctures. Apical tubercle small, only as a small tumescence (
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<figureCitation id="899F08BAFF82FF9BFD96456214F6F8B0" box="[583,687,1853,1878]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="28" pageNumber="479">Fig. 103</figureCitation>
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). Subapical fovea narrow, elongate, shallow, rugose and finely punctate.
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<taxonomicName id="D6A46FBCFF82FF9BFED6453F17B8F89C" authority="Surface" authorityName="Surface" box="[263,481,1888,1914]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF82FF9BFED6453F17DCF89F" bold="true" box="[263,389,1888,1913]" italics="true" pageId="28" pageNumber="479">Scutellum:</emphasis>
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Surface
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</taxonomicName>
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with 2 deep and some minute, sparse punctures.
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<emphasis id="23D0C82DFF82FF9BFBF2453F122DF89F" bold="true" box="[1059,1140,1888,1913]" italics="true" pageId="28" pageNumber="479">Elytra:</emphasis>
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Surface with deep, large, ocellate punctures, some smaller between intervals; 3 pairs of double rows of punctures distinct; elytral suture present.
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<taxonomicName id="D6A46FBCFF82FF9BFF2445F71790F824" authority="Surface" authorityName="Surface" box="[245,457,1960,1986]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="479" phylum="Chordata" rank="genus">
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<emphasis id="23D0C82DFF82FF9BFF2445F71734F827" bold="true" box="[245,365,1960,1985]" italics="true" pageId="28" pageNumber="479">Pygidium:</emphasis>
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Surface
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</taxonomicName>
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completely punctate; punctures subequal in size to those of apex elytra, ocellate, minutely setigerous (
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<figureCitation id="899F08BAFF82FF9BFE5A459217BEF800" box="[395,487,1997,2022]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="28" pageNumber="479">Fig.104</figureCitation>
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). Surface evenly convex in males.
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<emphasis id="23D0C82DFF82FF9BFC534591159BF803" bold="true" box="[898,962,1998,2021]" italics="true" pageId="28" pageNumber="479">Legs:</emphasis>
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Protibia tridentate, basal tooth distinctly removed from others. Male protarsus simple, not enlarged. Metatibia with 7 slender, long spinules on the apical margin. Apex of first metatarsomere triangularly expanded.
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<emphasis id="23D0C82DFF83FF9AFC8542E315F1FF35" bold="true" box="[852,936,188,211]" italics="true" pageId="29" pageNumber="480">Venter:</emphasis>
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Prosternal process long, extending beyond procoxae; apex spherical and completely cover by long yellow setae.
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<emphasis id="23D0C82DFF83FF9AFC6642BF1264FF1E" bold="true" box="[951,1085,223,248]" pageId="29" pageNumber="480">
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<emphasis id="23D0C82DFF83FF9AFC6642BF126AFF11" bold="true" box="[951,1075,224,247]" italics="true" pageId="29" pageNumber="480">Parameres</emphasis>
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:
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</emphasis>
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Base broad; lateral teeth large and with apex acute; apical third narrowed, apices strongly expanded outwards (
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<figureCitation id="899F08BAFF83FF9AFBBF435B1342FEFB" box="[1134,1307,260,285]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="29" pageNumber="480">Figs. 105–106</figureCitation>
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).
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<emphasis id="23D0C82DFF83FF9AFAFC435C16A5FEA6" bold="true" italics="true" pageId="29" pageNumber="480">Spiculum gastrale:</emphasis>
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Base longer than lateral branches, apex truncate (
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<figureCitation id="899F08BAFF83FF9AFCE0437815CAFEA6" box="[817,915,295,320]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="29" pageNumber="480">Fig. 107</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF83FF9AFF1643131426FDDA" pageId="29" pageNumber="480" type="diagnosis">
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<paragraph id="111B143FFF83FF9AFF164313140DFE36" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="480">
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<emphasis id="23D0C82DFF83FF9AFF1643131718FE83" bold="true" box="[199,321,332,357]" pageId="29" pageNumber="480">Diagnosis.</emphasis>
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This species can be separated from other
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<taxonomicName id="D6A46FBCFF83FF9AFCF9431215D0FE82" box="[808,905,333,356]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23D0C82DFF83FF9AFCF9431215D0FE82" box="[808,905,333,356]" italics="true" pageId="29" pageNumber="480">Tomarus</emphasis>
|
||
</taxonomicName>
|
||
species by a strongly punctate dorsal surface, frontal tubercles and clypeal teeth widely separated, pronotal tubercle and fovea small, surface of
|
||
<taxonomicName id="D6A46FBCFF83FF9AFAE0433013C5FE6E" box="[1329,1436,367,392]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="488" phylum="Chordata" rank="genus">pygidium</taxonomicName>
|
||
strongly and evenly punctate, and protarsal claws not enlarged in males. Males can be easily identified by the shape of the parameres and spiculum gastrale.
|
||
</paragraph>
|
||
<paragraph id="111B143FFF83FF9AFF16438317A0FDFE" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="480">
|
||
<emphasis id="23D0C82DFF83FF9AFF16438317C2FE13" bold="true" box="[199,411,476,501]" pageId="29" pageNumber="480">Locality records.</emphasis>
|
||
(
|
||
<figureCitation id="899F08BAFF83FF9AFE6443831446FE12" box="[437,543,476,501]" captionStart="FIGURES 102 – 108" captionStartId="28.[151,264,1467,1490]" captionTargetBox="[157,1429,699,1441]" captionTargetId="figure@28.[151,1436,690,1446]" captionTargetPageId="28" captionText="FIGURES 102 – 108. Tomarus cicatricosus. 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium, 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia." httpUri="https://zenodo.org/record/234037/files/figure.png" pageId="29" pageNumber="480">Fig. 108</figureCitation>
|
||
)
|
||
<specimenCount id="07A2DFB6FF83FF9AFDE843831407FE12" box="[569,606,476,500]" pageId="29" pageNumber="480" type="generic">1♂</specimenCount>
|
||
. The single specimen was seen from ICN.
|
||
<emphasis id="23D0C82DFF83FF9AFB43438313C5FE13" bold="true" box="[1170,1436,476,501]" pageId="29" pageNumber="480">Valle del Cauca (1):</emphasis>
|
||
Buenaventura, río San Juan (1).
|
||
</paragraph>
|
||
<paragraph id="111B143FFF83FF9AFF16407B1426FDDA" blockId="29.[151,1437,151,717]" box="[199,639,548,573]" pageId="29" pageNumber="480">
|
||
<emphasis id="23D0C82DFF83FF9AFF16407B178FFDDB" bold="true" box="[199,470,548,573]" pageId="29" pageNumber="480">Temporal distribution.</emphasis>
|
||
December (1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59BE47B4FF83FF9AFF16401817FFFD2B" pageId="29" pageNumber="480" type="distribution">
|
||
<paragraph id="111B143FFF83FF9AFF164018136BFD63" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="480">
|
||
<emphasis id="23D0C82DFF83FF9AFF1640181700FD86" bold="true" box="[199,345,583,608]" pageId="29" pageNumber="480">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="69B354AFFF83FF9AFEBE4018179EFD86" box="[367,455,583,608]" name="Mexico" pageId="29" pageNumber="480">Mexico</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF83FF9AFE0A4018140DFD86" box="[475,596,583,608]" name="Guatemala" pageId="29" pageNumber="480">Guatemala</collectingCountry>
|
||
(
|
||
<bibRefCitation id="753569CEFF83FF9AFDBB4018154BFD86" author="Endrodi" box="[618,786,583,608]" pageId="29" pageNumber="480" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Museum fur Tierkunde, Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
|
||
),
|
||
<collectingCountry id="69B354AFFF83FF9AFCFC401715F6FD86" box="[813,943,583,608]" name="Costa Rica" pageId="29" pageNumber="480">Costa Rica</collectingCountry>
|
||
(
|
||
<bibRefCitation id="753569CEFF83FF9AFC144018122AFD86" author="Ratcliffe" box="[965,1139,583,608]" pageId="29" pageNumber="480" refString="Ratcliffe, B. C. (2003) The Dynastinae scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum, 16, 1 - 506." type="journal article" year="2003">Ratcliffe 2003</bibRefCitation>
|
||
),
|
||
<collectingCountry id="69B354AFFF83FF9AFB5E40181345FD86" box="[1167,1308,583,608]" name="El Salvador" pageId="29" pageNumber="480">El Salvador</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF83FF9AFAFE401813C5FD86" box="[1327,1436,583,608]" name="Honduras" pageId="29" pageNumber="480">Honduras</collectingCountry>
|
||
(
|
||
<bibRefCitation id="753569CEFF83FF9AFF4F403317FBFD63" author="Ratcliffe" box="[158,418,620,645]" pageId="29" pageNumber="480" refString="Ratcliffe, B. C. & Cave, R. D. (2006) The Dynastinae Scarab Beetles of Honduras, Nicaragua and El Salvador. Bulletin of the University of Nebraska State Museum, 21, 1 - 424." type="journal article" year="2006">Ratcliffe & Cave 2006</bibRefCitation>
|
||
), and
|
||
<collectingCountry id="69B354AFFF83FF9AFE344033140EFD63" box="[485,599,620,645]" name="Colombia" pageId="29" pageNumber="480">Colombia</collectingCountry>
|
||
. The specimen listed represents a
|
||
<emphasis id="23D0C82DFF83FF9AFC054033136BFD63" bold="true" box="[980,1330,620,645]" pageId="29" pageNumber="480">NEW COUNTRY RECORD.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="111B143FFF83FF9AFF1640D017FFFD2B" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="480">
|
||
<emphasis id="23D0C82DFF83FF9AFF1640D017DCFD4E" bold="true" box="[199,389,655,680]" pageId="29" pageNumber="480">Natural history.</emphasis>
|
||
The specimen examined was collected in a mangrove from the Choco biogeographic region, at about
|
||
<quantity id="D65CB9DAFF83FF9AFF2640EB1769FD2A" box="[247,304,692,716]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="480" unit="m" value="30.0">30 m</quantity>
|
||
elevation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |