The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution Author López-García, Margarita M. Author Gasca-Álvarez, Héctor J. Author Amat-García, Germán text Zootaxa 2015 4048 4 451 492 journal article 10.11646/zootaxa.4048.4.1 bbb51c92-e184-489d-aba8-18d1dc9a25cd 1175-5326 234023 71E2B8DA-825E-429F-BAA0-F582702B4A80 Tomarus cicatricosus ( Prell, 1937 ) ( Figs. 102 – 108 ) Ligyrus cicatricosus Prell, 1937 : 90 . FIGURES 102–108. Tomarus cicatricosus . 102) habitus in dorsal view, 103) habitus in frontal view, 104) pygidium , 105) parameres in frontal view, 106) parameres in lateral view, 107) spiculum gastrale, 108) locality records in Colombia. Description. Habitus as in Fig. 102 . Length 24 mm . Width 13 mm (♂). Color black, procoxae lighter. Head : Frons slightly concave between eyes and behind frontoclypeal tubercles; surface rugose and grooved, nearly smooth on vertex and only with deep, sparse punctures. Frontoclypeal region with 2 transverse tubercles broadly separated from one another (about 9 tubercle diameters). Frontoclypeal suture present, as a weak carina connecting frontal tubercles. Clypeus with surface transversely rugose; apex broadly truncate, with 2 small teeth; teeth reflexed, widely separated (about 5 tooth diameters). Interocular distance equals 3.3 times the transverse ocular diameter. Antennal club subequal in length to antennomeres 2–7. Mandibles with 2 apical and one lateral tooth. Pronotum: Surface with punctures evenly sparse, deep, large, weakly umbilicale punctures. Apical tubercle small, only as a small tumescence ( Fig. 103 ). Subapical fovea narrow, elongate, shallow, rugose and finely punctate. Scutellum: Surface with 2 deep and some minute, sparse punctures. Elytra: Surface with deep, large, ocellate punctures, some smaller between intervals; 3 pairs of double rows of punctures distinct; elytral suture present. Pygidium: Surface completely punctate; punctures subequal in size to those of apex elytra, ocellate, minutely setigerous ( Fig.104 ). Surface evenly convex in males. Legs: Protibia tridentate, basal tooth distinctly removed from others. Male protarsus simple, not enlarged. Metatibia with 7 slender, long spinules on the apical margin. Apex of first metatarsomere triangularly expanded. Venter: Prosternal process long, extending beyond procoxae; apex spherical and completely cover by long yellow setae. Parameres : Base broad; lateral teeth large and with apex acute; apical third narrowed, apices strongly expanded outwards ( Figs. 105–106 ). Spiculum gastrale: Base longer than lateral branches, apex truncate ( Fig. 107 ). Diagnosis. This species can be separated from other Tomarus species by a strongly punctate dorsal surface, frontal tubercles and clypeal teeth widely separated, pronotal tubercle and fovea small, surface of pygidium strongly and evenly punctate, and protarsal claws not enlarged in males. Males can be easily identified by the shape of the parameres and spiculum gastrale. Locality records. ( Fig. 108 ) 1♂ . The single specimen was seen from ICN. Valle del Cauca (1): Buenaventura, río San Juan (1). Temporal distribution. December (1). Distribution . Mexico , Guatemala ( Endrödi 1969 ), Costa Rica ( Ratcliffe 2003 ), El Salvador , Honduras ( Ratcliffe & Cave 2006 ), and Colombia . The specimen listed represents a NEW COUNTRY RECORD. Natural history. The specimen examined was collected in a mangrove from the Choco biogeographic region, at about 30 m elevation.