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101 lines
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<document id="6917ADC4239D10D23DCD43D649CC97E9" ID-DOI="10.2478/if-2018-0019" ID-ISSN="2533-4069" ID-Zenodo-Dep="4586600" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614971146920" checkinUser="felipe" docAuthor="Mendes, Mário Miguel, Dinis, Pedro & Kvaček, Jiří" docDate="2018" docId="0227EE3A684CFFAD91052489FC27FB35" docLanguage="en" docName="FossilImprint.74.3-4.317-326.pdf" docOrigin="Fossil Imprint 74 (3 - 4)" docSource="http://dx.doi.org/10.2478/if-2018-0019" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Pagiophyllum HEER 1881" docType="treatment" docVersion="3" lastPageNumber="321" masterDocId="FE1E96426848FFA9952F2552FFF8FFEE" masterDocTitle="Some Conifers From The Early Cretaceous (Late Aptian - Early Albian) Of Catefica, Lusitanian Basin, Western Portugal" masterLastPageNumber="326" masterPageNumber="317" pageNumber="321" updateTime="1698881519349" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:titleInfo id="47C2749B245D5FB7A49B91EF0EE38D2C">
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<mods:title id="2E3A42B453CD08E7CC951F9F6E4CE8C0">Some Conifers From The Early Cretaceous (Late Aptian - Early Albian) Of Catefica, Lusitanian Basin, Western Portugal</mods:title>
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<mods:name id="12B5CE5D3E3B81499260DBEAEE10581A" type="personal">
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<mods:namePart id="1DA59EFACE07697BCA20461B9DB9C0EA">Mendes, Mário Miguel</mods:namePart>
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<mods:affiliation id="78D4F30FB60F3D5DD07C09EE67445B8A">CIMA - Centre for Marine and Environmental Research, Universidade do Algarve, Campus Universitário de Gambelas, 8005 - 139 Faro, Portugal; e-mail: mmmendes @ mail. telepac. pt</mods:affiliation>
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<mods:nameIdentifier id="6890BD3AA3A272DFA43F6B4A26C11F2E" type="email">mmmendes@mail.telepac.pt</mods:nameIdentifier>
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<mods:namePart id="D5EC25A29D49F362CE41AF34FC1E6D6F">Dinis, Pedro</mods:namePart>
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<mods:affiliation id="719F301CA83DB236995B28C443BE8E46">MARE - Marine and Environmental Sciences Centre, Departamento de Ciências da Terra da Universidade de Coimbra, Largo Marquês de Pombal, 3000 - 272 Coimbra, Portugal; e-mail: pdinis @ dct. uc. pt.</mods:affiliation>
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<mods:nameIdentifier id="E1C1729C72D06D38EB4EE68DE6F23FD5" type="email">pdinis@dct.uc.pt.</mods:nameIdentifier>
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<mods:roleTerm id="AC5E4559BFCD2BEB979E1DE108AB370D">Author</mods:roleTerm>
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<mods:namePart id="F9972C105B8105704D4AD038A265075E">Kvaček, Jiří</mods:namePart>
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<mods:affiliation id="23302AFBD571185A0BF0538D001C585D">Department of Palaeontology, National Museum, Václavské nám. 68, 110 00 Praha 1, the Czech Republic; e-mail: jiri _ kvacek @ nm. cz.</mods:affiliation>
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<mods:nameIdentifier id="B85AA256B384CD8C58F6B2AB747CC5B7" type="email">jiri_kvacek@nm.cz.</mods:nameIdentifier>
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<mods:typeOfResource id="D6C229F1E77C947E644F2A411338B379">text</mods:typeOfResource>
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<mods:title id="7ED139946503ABDA6CE32824F9C6677A">Fossil Imprint</mods:title>
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<mods:date id="38AD932B20E9E614DE75BB1A882880F7">2018</mods:date>
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<mods:number id="E3E649FA270F957671583EBEA7833D53">2018-12-28</mods:number>
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<mods:number id="CBDF6F6376EAE92B99DD66CE07463E37">74</mods:number>
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<mods:number id="7BC610E7CEEDF9067F15F8B05ADDDDD1">3 - 4</mods:number>
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<mods:url id="C20A82C6372FBFA3068567494B07A9C8">http://dx.doi.org/10.2478/if-2018-0019</mods:url>
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<mods:identifier id="7E9E98CAFC4E94EF5C836C1467B32640" type="DOI">10.2478/if-2018-0019</mods:identifier>
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<mods:identifier id="AB68BBC9574EFD6F71BDE73AFFCC39CE" type="ISSN">2533-4069</mods:identifier>
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<treatment id="0227EE3A684CFFAD91052489FC27FB35" LSID="urn:lsid:plazi:treatment:0227EE3A684CFFAD91052489FC27FB35" httpUri="http://treatment.plazi.org/id/0227EE3A684CFFAD91052489FC27FB35" lastPageNumber="321" pageId="4" pageNumber="321">
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<subSubSection id="C2940CA7684CFFAD91052489FB2AFDFA" pageId="4" pageNumber="321" type="nomenclature">
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<paragraph id="8A315F2C684CFFAD91052489FB12FE1D" blockId="4.[1066,1258,475,499]" box="[1066,1258,475,499]" pageId="4" pageNumber="321">
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<heading id="D179E840684CFFAD91052489FB12FE1D" box="[1066,1258,475,499]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="321" reason="2">
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<emphasis id="B8FA833E684CFFAD91052489FB12FE1D" bold="true" box="[1066,1258,475,499]" pageId="4" pageNumber="321">
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<taxonomicName id="4D8E24AF684CFFAD91052489FB3CFE1D" authorityName="HEER" authorityYear="1881" box="[1066,1220,475,499]" class="Pinopsida" family="Araucariaceae" genus="Pagiophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="321" phylum="Tracheophyta" rank="genus">
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<emphasis id="B8FA833E684CFFAD91052489FB3CFE1D" bold="true" box="[1066,1220,475,499]" italics="true" pageId="4" pageNumber="321">Pagiophyllum</emphasis>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8A315F2C684CFFAD916D24AFFB2AFDFA" blockId="4.[1090,1234,509,532]" box="[1090,1234,509,532]" pageId="4" pageNumber="321">
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<figureCitation id="12B543A9684CFFAD916D24AFFB2AFDFA" box="[1090,1234,509,532]" captionStart="Text-fig" captionStartId="5.[125,206,1799,1821]" captionTargetBox="[125,1438,151,1780]" captionTargetId="figure-2@5.[124,1441,150,1781]" captionTargetPageId="5" captionText="Text-fig. 4. SEM micrographs of two types of conifers from the Early Cretaceous mesofossil flora of Catefica, Portugal. (a) Leafy twig of Watsoniocladus cf. virginiensis with opposite-decussate leaves showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0127, sample Catefica 316). (b), (c) Details of the leaf apex showing amphistomatic abaxial and adaxial surfaces (P0127, sample Catefica 316). (d) Detail of stomata surrounded by a raised rim with finger-like outer papillae showing star-shaped stomatal apertures (arrows) (P0127, sample Catefica 316). (e), (f) Leafy twig of Watsoniocladus cf. virginiensis showing hook-shaped leaves in outline (P0129, sample Catefica 317). (g) Detail of the abaxial surface showing stomata arranged in ill-defined rows (P0129, sample Catefica 317). (h) Leafy twig of Pagiophyllum sp. showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0126, sample Catefica 318). (i) Detail of the abaxial leaf surface showing serrate margin and scattered stomata (P0126, sample Catefica 318). Scale bars = 1 mm (a, e, f, h); 100 μm (b, c, g, i); 50 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.4586608" httpUri="https://zenodo.org/record/4586608/files/figure.png" pageId="4" pageNumber="321">Text-fig. 4h, i</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C2940CA7684CFFAD965E277EFC27FB35" pageId="4" pageNumber="321" type="description">
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<paragraph id="8A315F2C684CFFAD965E277EFC22FC89" blockId="4.[843,1482,556,871]" pageId="4" pageNumber="321">
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D e s c r i p t i o n. The fragmentary shoot specimens consist of long, slender leaves arranged helically along unbranched axes (
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<figureCitation id="12B543A9684CFFAD9136273FFB59FD6A" box="[1049,1185,621,644]" captionStart="Text-fig" captionStartId="5.[125,206,1799,1821]" captionTargetBox="[125,1438,151,1780]" captionTargetId="figure-2@5.[124,1441,150,1781]" captionTargetPageId="5" captionText="Text-fig. 4. SEM micrographs of two types of conifers from the Early Cretaceous mesofossil flora of Catefica, Portugal. (a) Leafy twig of Watsoniocladus cf. virginiensis with opposite-decussate leaves showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0127, sample Catefica 316). (b), (c) Details of the leaf apex showing amphistomatic abaxial and adaxial surfaces (P0127, sample Catefica 316). (d) Detail of stomata surrounded by a raised rim with finger-like outer papillae showing star-shaped stomatal apertures (arrows) (P0127, sample Catefica 316). (e), (f) Leafy twig of Watsoniocladus cf. virginiensis showing hook-shaped leaves in outline (P0129, sample Catefica 317). (g) Detail of the abaxial surface showing stomata arranged in ill-defined rows (P0129, sample Catefica 317). (h) Leafy twig of Pagiophyllum sp. showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0126, sample Catefica 318). (i) Detail of the abaxial leaf surface showing serrate margin and scattered stomata (P0126, sample Catefica 318). Scale bars = 1 mm (a, e, f, h); 100 μm (b, c, g, i); 50 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.4586608" httpUri="https://zenodo.org/record/4586608/files/figure.png" pageId="4" pageNumber="321">Text-fig. 4h</figureCitation>
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). The leaves are up to
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<quantity id="4D76F2C9684CFFAD9093273FFC8CFD4A" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="321" unit="mm" value="3.0">3 mm</quantity>
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long and
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<quantity id="4D76F2C9684CFFAD96CC27DFFBCEFD4A" box="[995,1078,653,676]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="321" unit="mm" value="0.6">0.6 mm</quantity>
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wide. Their leaf cushion is narrowly elongate. They represent a continuation of a basal part of the leaf, without any particular broadened part (
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<figureCitation id="12B543A9684CFFAD9042279CFC91FCE8" captionStart="Text-fig" captionStartId="5.[125,206,1799,1821]" captionTargetBox="[125,1438,151,1780]" captionTargetId="figure-2@5.[124,1441,150,1781]" captionTargetPageId="5" captionText="Text-fig. 4. SEM micrographs of two types of conifers from the Early Cretaceous mesofossil flora of Catefica, Portugal. (a) Leafy twig of Watsoniocladus cf. virginiensis with opposite-decussate leaves showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0127, sample Catefica 316). (b), (c) Details of the leaf apex showing amphistomatic abaxial and adaxial surfaces (P0127, sample Catefica 316). (d) Detail of stomata surrounded by a raised rim with finger-like outer papillae showing star-shaped stomatal apertures (arrows) (P0127, sample Catefica 316). (e), (f) Leafy twig of Watsoniocladus cf. virginiensis showing hook-shaped leaves in outline (P0129, sample Catefica 317). (g) Detail of the abaxial surface showing stomata arranged in ill-defined rows (P0129, sample Catefica 317). (h) Leafy twig of Pagiophyllum sp. showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0126, sample Catefica 318). (i) Detail of the abaxial leaf surface showing serrate margin and scattered stomata (P0126, sample Catefica 318). Scale bars = 1 mm (a, e, f, h); 100 μm (b, c, g, i); 50 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.4586608" httpUri="https://zenodo.org/record/4586608/files/figure.png" pageId="4" pageNumber="321">Text-fig. 4h</figureCitation>
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). The abaxial surfaces are convex and keeled, whilst the adaxial surfaces are concave. Leaf margins show tiny, regular teeth (
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<figureCitation id="12B543A9684CFFAD96C72662FB9FFCA9" box="[1000,1127,816,839]" captionStart="Text-fig" captionStartId="5.[125,206,1799,1821]" captionTargetBox="[125,1438,151,1780]" captionTargetId="figure-2@5.[124,1441,150,1781]" captionTargetPageId="5" captionText="Text-fig. 4. SEM micrographs of two types of conifers from the Early Cretaceous mesofossil flora of Catefica, Portugal. (a) Leafy twig of Watsoniocladus cf. virginiensis with opposite-decussate leaves showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0127, sample Catefica 316). (b), (c) Details of the leaf apex showing amphistomatic abaxial and adaxial surfaces (P0127, sample Catefica 316). (d) Detail of stomata surrounded by a raised rim with finger-like outer papillae showing star-shaped stomatal apertures (arrows) (P0127, sample Catefica 316). (e), (f) Leafy twig of Watsoniocladus cf. virginiensis showing hook-shaped leaves in outline (P0129, sample Catefica 317). (g) Detail of the abaxial surface showing stomata arranged in ill-defined rows (P0129, sample Catefica 317). (h) Leafy twig of Pagiophyllum sp. showing convex, keeled, abaxial surface and concave, unkeeled, adaxial surface (P0126, sample Catefica 318). (i) Detail of the abaxial leaf surface showing serrate margin and scattered stomata (P0126, sample Catefica 318). Scale bars = 1 mm (a, e, f, h); 100 μm (b, c, g, i); 50 μm (d)." figureDoi="http://doi.org/10.5281/zenodo.4586608" httpUri="https://zenodo.org/record/4586608/files/figure.png" pageId="4" pageNumber="321">Text-fig. 4i</figureCitation>
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). Leaves hypostomatic, stomata lack papillae.
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</paragraph>
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<paragraph id="8A315F2C684CFFAD965E26D2FC27FB35" blockId="4.[843,1482,896,1243]" pageId="4" pageNumber="321">
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D i s c u s s i o n. This taxon shows helically arranged leaves with stomata lacking papillae. It provides completely different characteristics than the first described group of conifers belonging to the
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<taxonomicName id="4D8E24AF684CFFAD915626B3FAC5FC16" box="[1145,1341,993,1016]" class="Pinopsida" family="Cheirolepidiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="321" phylum="Tracheophyta" rank="family">Cheirolepidiaceae</taxonomicName>
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. Its stomata do not show any papillae. This
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<typeStatus id="5535E18E684CFFAD91B02153FB35FBF6" box="[1183,1229,1025,1048]" pageId="4" pageNumber="321">type</typeStatus>
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of foliage with falcate, helically arranged leaves may indicate
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<taxonomicName id="4D8E24AF684CFFAD91CA2170FA8EFBD7" box="[1253,1398,1058,1081]" class="Pinopsida" family="Cupressaceae" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="321" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
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s.l., but its stomata are not arranged in bands. Due to its fragmentary preservation, the material is left in open nomenclature and its systematic affinity remains questionable. Additional studies should provide more details to its more precise identification.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |