363 lines
40 KiB
XML
363 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1187.111101" ID-Pensoft-Pub="1313-2970-1187-223" ID-Pensoft-UUID="CF4334073C9858E9B3B5804A795B3FBF" ID-ZooBank="9B1CA3A4435B417982DE204771324B1C" ModsDocID="1313-2970-1187-223" checkinTime="1703228390790" checkinUser="pensoft" docAuthor="Li, Yange, Jing, Wenqing, Hao, Shulian & Yu, Haili" docDate="2023" docId="E0B73F79767759A0883172C2F333903F" docLanguage="en" docName="ZooKeys 1187: 223-236" docOrigin="ZooKeys 1187" docPubDate="2023-12-21" docSource="http://dx.doi.org/10.3897/zookeys.1187.111101" docTitle="Phaecadophora dactylina Li & Yu 2023, sp. nov." docType="treatment" docUuid="5FDA1766-F739-4A04-8E08-F56DCA536F9E" docUuidSource="ZooBank" docVersion="1" id="CF4334073C9858E9B3B5804A795B3FBF" lastPageNumber="223" masterDocId="CF4334073C9858E9B3B5804A795B3FBF" masterDocTitle="Descriptions of two new species of Phaecadophora Walsingham, 1900 (Lepidoptera, Tortricidae, Olethreutinae) from China" masterLastPageNumber="236" masterPageNumber="223" pageNumber="223" updateTime="1703228390790" updateUser="pensoft">
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<mods:titleInfo id="ABC3AD3A6BB26B96AA8FC2D49C0B7297">
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<mods:title id="741B5B16A2D31BC240FEEC2221CBA400">Descriptions of two new species of Phaecadophora Walsingham, 1900 (Lepidoptera, Tortricidae, Olethreutinae) from China</mods:title>
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<mods:name id="ECB40F168721225DC2AFD26E9B9951AA" type="personal">
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<mods:namePart id="9A44A958F3C88C2A2F44D4A326B6FF13">Li, Yange</mods:namePart>
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<mods:nameIdentifier id="EF5C705CE47D7BF57B87598C510227B1" type="ORCID">https://orcid.org/0009-0002-9188-2178</mods:nameIdentifier>
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<mods:affiliation id="AAB7296097FFCE73F40A1B8B033FFE7D">Shaanxi Key Laboratory for Animal Conservation, Northwest University, Xi'an, Shaanxi Province, 710069, China</mods:affiliation>
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<mods:roleTerm id="509059AD1A56A4DF61945A3402007D01">Author</mods:roleTerm>
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<mods:namePart id="907350EB73FF5B5E56F2A646E232C1A1">Jing, Wenqing</mods:namePart>
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<mods:nameIdentifier id="5937E2EB1B3B0AD181B3ED0E24BC1151" type="ORCID">https://orcid.org/0009-0007-9306-8854</mods:nameIdentifier>
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<mods:affiliation id="089CF7092F5872FB146A30D79671FE74">Shaanxi Key Laboratory for Animal Conservation, Northwest University, Xi'an, Shaanxi Province, 710069, China</mods:affiliation>
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<mods:name id="7DEE209450FBEFA02BE7DE1B500359CB" type="personal">
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<mods:roleTerm id="E4430B1D3EDA6D4158F2F5227FB48A87">Author</mods:roleTerm>
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<mods:namePart id="9BF5D6C3284E20633CE7D7E9F8D6198C">Hao, Shulian</mods:namePart>
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<mods:nameIdentifier id="096D29316150DE172A765208F44E8BE0" type="ORCID">https://orcid.org/0009-0005-6556-4151</mods:nameIdentifier>
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<mods:affiliation id="549333658E6DBD4FA1951D3A6A38E951">Tianjin Natural History Museum, Tianjin, 300201, China</mods:affiliation>
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<mods:namePart id="12E3B4120387D31102CF206FD3E5B96E">Yu, Haili</mods:namePart>
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<mods:nameIdentifier id="68BB8AB8CFE5318E0CFC3A1BE4EBF448" type="ORCID">https://orcid.org/0000-0003-1249-5461</mods:nameIdentifier>
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<mods:affiliation id="0C460CCC80E229AF12A41B009FEB940E">Shaanxi Key Laboratory for Animal Conservation, Northwest University, Xi'an, Shaanxi Province, 710069, China</mods:affiliation>
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<mods:nameIdentifier id="E23CA12975C2B985CE422D6A56016944" type="email">yuhaili@nwu.edu.cn</mods:nameIdentifier>
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<mods:typeOfResource id="C357963779205DA7ACAF5DD70723B15B">text</mods:typeOfResource>
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<mods:title id="4D95E58E78846937E1F62ECF2DB600BC">ZooKeys</mods:title>
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<mods:date id="2A0E2FE0FE16BC7110C123CD3F86F9A8">2023</mods:date>
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<mods:number id="E871227ED9B326771D61E91F9B000C78">2023-12-21</mods:number>
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<mods:number id="46977E452D28781121E37185FC45302C">1187</mods:number>
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<mods:start id="FEE1EF30556CDF4DE2C1488F1F077EBC">223</mods:start>
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<mods:url id="7B24EB69AD99957A134D927E6033DD99">http://dx.doi.org/10.3897/zookeys.1187.111101</mods:url>
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<mods:classification id="C14317BAEEBD5E4570F05B7172DDE35A">journal article</mods:classification>
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<mods:identifier id="F2872F0E3DE2547F4D56DD21EEF480CD" type="DOI">http://dx.doi.org/10.3897/zookeys.1187.111101</mods:identifier>
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<mods:identifier id="B61DD072772EDC18BF3B921A86DA3775" type="Pensoft-Pub">1313-2970-1187-223</mods:identifier>
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<mods:identifier id="1BAC15FB08657AE0A2C6695DDBA963D0" type="ZooBank">9B1CA3A4435B417982DE204771324B1C</mods:identifier>
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<treatment id="E0B73F79767759A0883172C2F333903F" LSID="urn:lsid:zoobank.org:act:5FDA1766-F739-4A04-8E08-F56DCA536F9E" httpUri="http://treatment.plazi.org/id/E0B73F79767759A0883172C2F333903F" lastPageNumber="223" pageId="0" pageNumber="223">
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<subSubSection id="1F079353BD2D55CBD93BB435A8D887EF" pageId="0" pageNumber="223" type="nomenclature">
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<paragraph id="01406C4A92E2C1C6E9586A4E6F8282B9" pageId="0" pageNumber="223">
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<taxonomicName id="3EBFDFCE73A03E79E75DF154B35DC52C" LSID="https://zoobank.org/5FDA1766-F739-4A04-8E08-F56DCA536F9E" authority="Li & Yu" authorityName="Li & Yu" authorityYear="2023" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora dactylina" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="dactylina" status="sp. nov.">Phaecadophora dactylina Li & Yu</taxonomicName>
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<taxonomicNameLabel id="553DEC00E0D6A1FFA95490BE6143ADAF" pageId="0" pageNumber="223">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="B5CAC4E16F2ECFC427D45CEB2D22CCF3" pageId="0" pageNumber="223" type="description">
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<paragraph id="80F80CAC03938E437FE3830713E7BA6E" pageId="0" pageNumber="223">
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<figureCitation id="91DC436D47F7DD6953C1E7F633C77FF4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adults of Phaecadophora spp. A P. dactylina sp. nov. (holotype, male) B P. dactylina sp. nov. (paratype, female) C P. vascularis sp. nov. (holotype, male). Scale bars: 2 mm." figureDoi="10.3897/zookeys.1187.111101.figure1" httpUri="https://binary.pensoft.net/fig/956181" pageId="0" pageNumber="223">Figs 1A, B</figureCitation>
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<figureCitation id="EC1DB4ECDCAFC7C957DFB69AF4C44658" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Phaecadophora spp. A, B P. dactylina sp. nov. (holotype, male) A lateral view B dorsal view C, D P. dactylina sp. nov. (paratype, female) C lateral view D dorsal view E, F P. vascularis sp. nov. (holotype, male) E lateral view F dorsal view." figureDoi="10.3897/zookeys.1187.111101.figure2" httpUri="https://binary.pensoft.net/fig/956182" pageId="0" pageNumber="223">, 2A-D</figureCitation>
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<figureCitation id="8B4342CFD023AAEDCA4816AFC2CCB0D0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hind tibiae in male of Phaecadophora spp. A P. dactylina sp. nov. (paratype) B P. vascularis sp. nov. (paratype)." figureDoi="10.3897/zookeys.1187.111101.figure3" httpUri="https://binary.pensoft.net/fig/956183" pageId="0" pageNumber="223">, 3A</figureCitation>
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<figureCitation id="0B9B66E6401C8AE2F7EDD901435C0EF4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male hindwings of Phaecadophora spp. A-C P. dactylina sp. nov. (paratype) A hindwing B cubital pecten C axillary hair pencils D-F P. vascularis sp. nov. (holotype) D hindwing E cubital pecten F axillary hair pencils." figureDoi="10.3897/zookeys.1187.111101.figure4" httpUri="https://binary.pensoft.net/fig/956184" pageId="0" pageNumber="223">, 4A-C</figureCitation>
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<figureCitation id="1CCB51D462B3EC5EADB53088136005E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Phaecadophora dactylina sp. nov. (holotype) A male genitalia B costal prominence of valva C ventral prominence of cucullus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure5" httpUri="https://binary.pensoft.net/fig/956185" pageId="0" pageNumber="223">, 5</figureCitation>
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<figureCitation id="94C09ED4367BF871856378A2C22D2288" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of Phaecadophora dactylina sp. nov. (paratype) A female genitalia B sterigma and colliculum C signum. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure6" httpUri="https://binary.pensoft.net/fig/956186" pageId="0" pageNumber="223">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6CF64706EB176F8AAC7E6F405E7B5707" pageId="0" pageNumber="223" type="type material">
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<paragraph id="738C05E115ED2162D3E16327E33B994D" pageId="0" pageNumber="223">Type materials.</paragraph>
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<paragraph id="75E14789BD07F8C876B8F1C1EC6A8483" pageId="0" pageNumber="223">
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<emphasis id="EE944913C055EB3CEFFF330A3E49F584" bold="true" italics="true" pageId="0" pageNumber="223">Holotype</emphasis>
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: ♂, China, Tibet: Motuo County, Beibengxiang,
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<geoCoordinate id="92FBEB44AB54E67AF939EF831DB9BF00" degrees="29" direction="north" minutes="19.00" orientation="latitude" precision="9" value="29.316668">29°19.00'N</geoCoordinate>
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,
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<geoCoordinate id="2B9B3DBE022ADC4C812BB849AEC80B49" degrees="95" direction="east" minutes="10.80" orientation="longitude" precision="9" value="95.18">95°10.80'E</geoCoordinate>
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, alt. 810 m, 13 Aug. 2017, Mujie Qi and Xiaofei Yang leg., genitalia slide no. YWX18220.
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<emphasis id="94B271DF0018C610D138A8403BAD1730" bold="true" italics="true" pageId="0" pageNumber="223">Paratypes</emphasis>
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: China, Tibet: 3♂, same data as holotype except
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<geoCoordinate id="2393C649E2789D88AB9504E1AF3A388B" degrees="29" direction="north" minutes="14.40" orientation="latitude" precision="9" value="29.24">29°14.40'N</geoCoordinate>
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,
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<geoCoordinate id="C9565CA04DC58B10ED1ABCF37887B26C" degrees="95" direction="east" minutes="19.20" orientation="longitude" precision="9" value="95.32">95°19.20'E</geoCoordinate>
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, alt. 810-990 m, 12-13 Aug. 2017; 1♂, same data as holotype except
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<geoCoordinate id="FB428C07B88D993FE4B697B87D1442F4" degrees="29" direction="north" minutes="19.20" orientation="latitude" precision="9" value="29.32">29°19.20'N</geoCoordinate>
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,
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<geoCoordinate id="7F59EB7BDC6D259FB1607A7C95241336" degrees="95" direction="east" minutes="19.20" orientation="longitude" precision="9" value="95.32">95°19.20'E</geoCoordinate>
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, alt. 1100 m, 10 Aug. 2017; 1♀, Nielamu County,
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<geoCoordinate id="31F79C20CED1055E12C82EA7553C78BA" degrees="27" direction="north" minutes="58.80" orientation="latitude" precision="9" value="27.98">27°58.80'N</geoCoordinate>
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,
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<geoCoordinate id="83C307339C940FC9DE709328F4996E97" degrees="85" direction="east" minutes="58.20" orientation="longitude" precision="9" value="85.97">85°58.20'E</geoCoordinate>
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, alt. 1960 m, 6 Jul. 2019, Mujie Qi and Jiaqi Deng leg.; Yunnan Prov.: 1♂, 3♀, Tengchong County, Linjiapuzi,
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<geoCoordinate id="D33E141F7967B9746AC64075BC9DBE83" degrees="25" direction="north" minutes="17.40" orientation="latitude" precision="9" value="25.29">25°17.40'N</geoCoordinate>
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,
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<geoCoordinate id="8AB3D661E7ED8DE771FF918474516B3D" degrees="98" direction="east" minutes="42.00" orientation="longitude" precision="9" value="98.7">98°42.00'E</geoCoordinate>
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, alt. 2140 m, 15 Aug. 2014, Kaijian Teng, Shurong Liu and Hua Rong leg.
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</paragraph>
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</subSubSection>
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<subSubSection id="3DFB9C3B39A400705EA540BFEB4A0EA5" pageId="0" pageNumber="223" type="diagnosis">
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<paragraph id="26DBB70EF7F554B1EE0A58AC767FA7F3" pageId="0" pageNumber="223">Diagnosis.</paragraph>
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<paragraph id="39515D28024B640706D7133836B08B4C" pageId="0" pageNumber="223">
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The male of
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<taxonomicName id="A498BC002341D96699B47332ED3B37B0" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
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<emphasis id="476B60432B3AE15B1853CA41293151B2" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
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</taxonomicName>
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sp. nov. resembles
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<taxonomicName id="07E2B1D57E7AA399AF42FA5D6A0903E2" lsidName="P. fimbriata" pageId="0" pageNumber="223" rank="species" species="fimbriata">
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<emphasis id="5B0066421CF515EFC5458EC3481C5F18" italics="true" pageId="0" pageNumber="223">P. fimbriata</emphasis>
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</taxonomicName>
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in appearance in having darker scaling in the forewing pattern, two hair pencils and long scales in the anal roll of hindwing, and the hindleg broadened. Dissection of the genitalia is necessary for identification. Conversely, the female can be readily separated from
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<taxonomicName id="60BD9F276D4FE04EA9769ACCCA12DE87" lsidName="P. fimbriata" pageId="0" pageNumber="223" rank="species" species="fimbriata">
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<emphasis id="1D3D35C77880C8CD706B28018C14279C" italics="true" pageId="0" pageNumber="223">P. fimbriata</emphasis>
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</taxonomicName>
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in having the forewing pale brown suffused with tawny longitudinal markings. More diagnostic characters are found in the male and female genitalia. The male of
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<taxonomicName id="C466E7388FCBCDD7276BB094AFF66A45" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
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<emphasis id="F79604444B0A715992991A14A67EF00A" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished by the apically furcated uncus, the valva adorned with a short finger-like prominence below the costa, a nearly bare basal region of the cucullus, and the absence of cornuti in the phallus. In contrast,
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<taxonomicName id="40DF77A4E2C38C2D4EC53687C17D155A" lsidName="P. fimbriata" pageId="0" pageNumber="223" rank="species" species="fimbriata">
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<emphasis id="BF14472C0707607E4F2D024D3454ED3F" italics="true" pageId="0" pageNumber="223">P. fimbriata</emphasis>
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</taxonomicName>
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presents a hooked uncus, the valva devoid of prominences below the costa, and a spined ridge across the base of the cucullus, with the phallus bearing a short spine on the vesica. In the female genitalia,
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<taxonomicName id="DA661DF2279C968EEE43AA57897652D3" lsidName="P. fimbriata" pageId="0" pageNumber="223" rank="species" species="fimbriata">
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<emphasis id="7D864A40FDB0490C5E62081C0DC8B81E" italics="true" pageId="0" pageNumber="223">P. fimbriata</emphasis>
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</taxonomicName>
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exhibits the sterigma lacking posterior extensions, whereas in
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<taxonomicName id="29E56A17550A121251514DC1F29AF8C3" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
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<emphasis id="BBB1B2D6C184FD0F88A70AF90EE74691" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
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</taxonomicName>
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sp. nov., this structure manifests as two broad plates.
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</paragraph>
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</subSubSection>
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<subSubSection id="CC867026F1CCE02C32E90D52D936E97B" pageId="0" pageNumber="223" type="description">
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<paragraph id="1C659E0490D37147449744BBDB6F4385" pageId="0" pageNumber="223">Description.</paragraph>
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<paragraph id="14ED7D900CB7BFA0190C7E80FEEDCFF7" pageId="0" pageNumber="223">
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<emphasis id="3AB7641DA035CA7E6ADAB03CC1079D5B" bold="true" pageId="0" pageNumber="223">Male</emphasis>
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(Fig.
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<figureCitation id="99509890EF1E8EF97BEA5C710E5CCFB8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adults of Phaecadophora spp. A P. dactylina sp. nov. (holotype, male) B P. dactylina sp. nov. (paratype, female) C P. vascularis sp. nov. (holotype, male). Scale bars: 2 mm." figureDoi="10.3897/zookeys.1187.111101.figure1" httpUri="https://binary.pensoft.net/fig/956181" pageId="0" pageNumber="223">1A</figureCitation>
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) with forewing length 8.0-9.0 mm.
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<emphasis id="537E2A2E3291ECCCCB9110994328FDEF" bold="true" italics="true" pageId="0" pageNumber="223">Head</emphasis>
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(Fig.
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<figureCitation id="0700483CA33177101C31CF2BC9EA49AA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Phaecadophora spp. A, B P. dactylina sp. nov. (holotype, male) A lateral view B dorsal view C, D P. dactylina sp. nov. (paratype, female) C lateral view D dorsal view E, F P. vascularis sp. nov. (holotype, male) E lateral view F dorsal view." figureDoi="10.3897/zookeys.1187.111101.figure2" httpUri="https://binary.pensoft.net/fig/956182" pageId="0" pageNumber="223">2A, B</figureCitation>
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): vertex and upper frons with shaggy, pale gray-tawny scales (shiny gray distally), lower frons with gray-white appressed scales. Antenna gray-tawny, extending to middle of forewing costa. Ocellus well developed; chaetosema present. Labial palpus ascending, basal half white, distal part gray-white, gray-tawny to gray, medially dusted with a few black scales; median segment broadened distally; terminal segment a little slender, porrect.
|
|
</paragraph>
|
|
<caption id="A6CA371B4F7350E8071BE83B8F8A4E8D" doi="10.3897/zookeys.1187.111101.figure1" httpUri="https://binary.pensoft.net/fig/956181" pageId="0" pageNumber="223" start="Figure 1" startId="F1">
|
|
<paragraph id="6EE9B3ECF8DB3B586A93F56A83BD8E14" pageId="0" pageNumber="223">
|
|
<emphasis id="58C9EB1BCBDF84A61CF63BE73CCA7994" bold="true" pageId="0" pageNumber="223">Figure 1.</emphasis>
|
|
Adults of
|
|
<taxonomicName id="4BEBF2CD33B43DCFAD45A701EF1F2C45" authorityName="Walsingham" authorityYear="1900" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="FE854C6F77A4DEBDFE8002DBF8F9F49A" italics="true" pageId="0" pageNumber="223">Phaecadophora</emphasis>
|
|
</taxonomicName>
|
|
spp.
|
|
<emphasis id="1F85D084D2B1F69ADE45B90631516926" bold="true" pageId="0" pageNumber="223">A</emphasis>
|
|
<taxonomicName id="2FDFD2D0192D5BE66A2B70948C4D36E9" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
|
|
<emphasis id="A7CA92A34BDB75BCDD38023BC2891707" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype, male)
|
|
<emphasis id="BB518D8E487D21D23FFB05092593C078" bold="true" pageId="0" pageNumber="223">B</emphasis>
|
|
<taxonomicName id="1A6C74CFEB85BC403C7AE4F5444588AA" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
|
|
<emphasis id="EB76AFE5D01C437EA08A9E0B3BB55679" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype, female)
|
|
<emphasis id="F1B3C298EE2DBB299B1720BA9A0FEB48" bold="true" pageId="0" pageNumber="223">C</emphasis>
|
|
<taxonomicName id="0B2AA8C5E1945E20FC6FA262A058D45B" lsidName="P. vascularis" pageId="0" pageNumber="223" rank="species" species="vascularis">
|
|
<emphasis id="6D5620FCAC12FAEB1C8DA4E4261804E3" italics="true" pageId="0" pageNumber="223">P. vascularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype, male). Scale bars: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="E80BA83C533F7B1E67103E1B97674814" doi="10.3897/zookeys.1187.111101.figure2" httpUri="https://binary.pensoft.net/fig/956182" pageId="0" pageNumber="223" start="Figure 2" startId="F2">
|
|
<paragraph id="F5323D41FF01209FFA3DA1C84FBF64BF" pageId="0" pageNumber="223">
|
|
<emphasis id="8565803CEE270D87961F45E584FD3E31" bold="true" pageId="0" pageNumber="223">Figure 2.</emphasis>
|
|
Heads of
|
|
<taxonomicName id="A40162CFD9EEE54348AEE731C1E8E2DA" authorityName="Walsingham" authorityYear="1900" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C40CAACF489805A9A5A1E6C1E0A7580E" italics="true" pageId="0" pageNumber="223">Phaecadophora</emphasis>
|
|
</taxonomicName>
|
|
spp.
|
|
<emphasis id="211E801A291BF806168F53B039F5407A" bold="true" pageId="0" pageNumber="223">A, B</emphasis>
|
|
<taxonomicName id="C7E711AABE5B7F9D4267CB5B8901BE1E" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
|
|
<emphasis id="0AA1271466067A52B70EBE2C226D49B1" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype, male)
|
|
<emphasis id="3A2F1E0A0EC2166BEB316FC9C4753EE2" bold="true" pageId="0" pageNumber="223">A</emphasis>
|
|
lateral view
|
|
<emphasis id="9498A2378D6F1AF057E22F60CB162EFA" bold="true" pageId="0" pageNumber="223">B</emphasis>
|
|
dorsal view
|
|
<emphasis id="A99D67E47130FC0F2D529D451AD967B0" bold="true" pageId="0" pageNumber="223">C, D</emphasis>
|
|
<taxonomicName id="5658E4A1B441B04F0BB1B4D37E5C01D7" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
|
|
<emphasis id="D081B41A594FC2016C0235594103F181" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype, female)
|
|
<emphasis id="AC7AFFB9C11C240FEF246FFACE304E71" bold="true" pageId="0" pageNumber="223">C</emphasis>
|
|
lateral view
|
|
<emphasis id="013C93235CD9322A40910270BED0257A" bold="true" pageId="0" pageNumber="223">D</emphasis>
|
|
dorsal view
|
|
<emphasis id="92B8081003EDDC8094283D9CDAD5D53B" bold="true" pageId="0" pageNumber="223">E, F</emphasis>
|
|
<taxonomicName id="A224688B74454F0E547FBB6FB3C81F6C" lsidName="P. vascularis" pageId="0" pageNumber="223" rank="species" species="vascularis">
|
|
<emphasis id="F87535C9BD6783F98D5485F98F9C69CD" italics="true" pageId="0" pageNumber="223">P. vascularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype, male)
|
|
<emphasis id="ECD600327E681D99F7ED5CAF51212D0F" bold="true" pageId="0" pageNumber="223">E</emphasis>
|
|
lateral view
|
|
<emphasis id="E287ED11AB3EDE39168BD5C862CC1E95" bold="true" pageId="0" pageNumber="223">F</emphasis>
|
|
dorsal view.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="B66F1E7CE51B829CE8298C0B824C44D9" pageId="0" pageNumber="223">
|
|
<emphasis id="B1614C62E563FD6230E6DCF1589D194D" bold="true" italics="true" pageId="0" pageNumber="223">Thorax</emphasis>
|
|
: fuscous basally, suffused with gray-white posteriorly. Hind tibia in male short, distally dilated by dense, long scales, creamy white, with a concolorous hair tuft on apical inner surface (Fig.
|
|
<figureCitation id="CF1C6BF57E6996575DA29F8C01B550FA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hind tibiae in male of Phaecadophora spp. A P. dactylina sp. nov. (paratype) B P. vascularis sp. nov. (paratype)." figureDoi="10.3897/zookeys.1187.111101.figure3" httpUri="https://binary.pensoft.net/fig/956183" pageId="0" pageNumber="223">3A</figureCitation>
|
|
); tarsus strongly broadened by dense scales dorsally; inner side of first segment forming a short, fuscous suffused with brown, scaled cavity; other tarsal segments creamy. Forewing subrectangular, slightly dilated towards termen, costa curved evenly, apex slightly produced, termen weakly oblique, tornus rounded; upperside fuscous, dusted with brown; pairs of strigulae on costa creamy, well-defined striae from them concolorous, extending longitudinally to termen and occupying halfway across the wing, partly confluent below distal half of costa; a double creamy streaks rising from base of wing, zigzagging between cell and 1A+2A to termen above tornus; cilia pale gray on upper part of termen, fuscous on lower part of termen, gray-white on tornus; underside brown, paler on costa, pairs of strigulae creamy, suffused with pale tawny, area of hindwing overlap white. Hindwing (Fig.
|
|
<figureCitation id="3C4C6139EB6CCFC38B8B322AEEC5A1BC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male hindwings of Phaecadophora spp. A-C P. dactylina sp. nov. (paratype) A hindwing B cubital pecten C axillary hair pencils D-F P. vascularis sp. nov. (holotype) D hindwing E cubital pecten F axillary hair pencils." figureDoi="10.3897/zookeys.1187.111101.figure4" httpUri="https://binary.pensoft.net/fig/956184" pageId="0" pageNumber="223">4A</figureCitation>
|
|
) fuscous, paler basally, costal area of forewing overlap white; pecten (Fig.
|
|
<figureCitation id="07D07A02E3B7912C22513DFED7142610" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male hindwings of Phaecadophora spp. A-C P. dactylina sp. nov. (paratype) A hindwing B cubital pecten C axillary hair pencils D-F P. vascularis sp. nov. (holotype) D hindwing E cubital pecten F axillary hair pencils." figureDoi="10.3897/zookeys.1187.111101.figure4" httpUri="https://binary.pensoft.net/fig/956184" pageId="0" pageNumber="223">4B</figureCitation>
|
|
) distinct; with a series of long hair-scales between CuA1 and distal half of CuA2; anal region triangularly expanded, with pale tawny and fuscous long hair pencils (Fig.
|
|
<figureCitation id="6D65E95B247272831A630DCCB8FFBA29" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male hindwings of Phaecadophora spp. A-C P. dactylina sp. nov. (paratype) A hindwing B cubital pecten C axillary hair pencils D-F P. vascularis sp. nov. (holotype) D hindwing E cubital pecten F axillary hair pencils." figureDoi="10.3897/zookeys.1187.111101.figure4" httpUri="https://binary.pensoft.net/fig/956184" pageId="0" pageNumber="223">4C</figureCitation>
|
|
) in anal roll; anal roll narrowly folded upward, bearing long hair-scales on margin; cilia gray-white; underside brown.
|
|
</paragraph>
|
|
<caption id="FD616B3F9022D1B1CC44EDEE5EF5B88F" doi="10.3897/zookeys.1187.111101.figure3" httpUri="https://binary.pensoft.net/fig/956183" pageId="0" pageNumber="223" start="Figure 3" startId="F3">
|
|
<paragraph id="28C30D199B59DED9EC14CB60F3DBB750" pageId="0" pageNumber="223">
|
|
<emphasis id="613797A7ACA7C7288D29989026676DD4" bold="true" pageId="0" pageNumber="223">Figure 3.</emphasis>
|
|
Hind tibiae in male of
|
|
<taxonomicName id="9FB0F79FC313CFF57B3BAAB9B9CEE3AD" authorityName="Walsingham" authorityYear="1900" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="8ED8928A6A7B8B80FCD3B81D5EA219C4" italics="true" pageId="0" pageNumber="223">Phaecadophora</emphasis>
|
|
</taxonomicName>
|
|
spp.
|
|
<emphasis id="14C39767DC193AC7877E224FD8BEF217" bold="true" pageId="0" pageNumber="223">A</emphasis>
|
|
<taxonomicName id="4465D4F1915355BC0987977EAAC2F441" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
|
|
<emphasis id="840CFB68D79AF84681E1CE7AAB12D3D8" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype)
|
|
<emphasis id="92E25458C96E50E540B7DFDEF2DA1966" bold="true" pageId="0" pageNumber="223">B</emphasis>
|
|
<taxonomicName id="7B83862DAD5EA71E33E3CFA84F61386F" lsidName="P. vascularis" pageId="0" pageNumber="223" rank="species" species="vascularis">
|
|
<emphasis id="599CA0CE0CC9E83699957AB606E87310" italics="true" pageId="0" pageNumber="223">P. vascularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="34327F0D820AD81D5809133DD0E85385" doi="10.3897/zookeys.1187.111101.figure4" httpUri="https://binary.pensoft.net/fig/956184" pageId="0" pageNumber="223" start="Figure 4" startId="F4">
|
|
<paragraph id="073E46548062587DE241F031E2D89AAC" pageId="0" pageNumber="223">
|
|
<emphasis id="80B51E9727CADDC72841519E67EEC91E" bold="true" pageId="0" pageNumber="223">Figure 4.</emphasis>
|
|
Male hindwings of
|
|
<taxonomicName id="6E518A071D8132D76FB487346BEC7FF0" authorityName="Walsingham" authorityYear="1900" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="1A5B0437B6085AF6565F0DC927C05DEE" italics="true" pageId="0" pageNumber="223">Phaecadophora</emphasis>
|
|
</taxonomicName>
|
|
spp.
|
|
<emphasis id="E5F0D95607EFB91ED70AAF63EECE6063" bold="true" pageId="0" pageNumber="223">A-C</emphasis>
|
|
<taxonomicName id="9DEEC0E0E74A35CCB57AA01E05A6A0B9" lsidName="P. dactylina" pageId="0" pageNumber="223" rank="species" species="dactylina">
|
|
<emphasis id="9441A5864569891004F35BAC7E795457" italics="true" pageId="0" pageNumber="223">P. dactylina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype)
|
|
<emphasis id="0CB2083A45B53D0CC177F92213073D0A" bold="true" pageId="0" pageNumber="223">A</emphasis>
|
|
hindwing
|
|
<emphasis id="E932D45066DE5E4691916718C01357A1" bold="true" pageId="0" pageNumber="223">B</emphasis>
|
|
cubital pecten
|
|
<emphasis id="2F47A2FDCAF8892C39C93D55CAFC0708" bold="true" pageId="0" pageNumber="223">C</emphasis>
|
|
axillary hair pencils
|
|
<emphasis id="4FDDB8BCDF56C593B106DBAA24430E7F" bold="true" pageId="0" pageNumber="223">D-F</emphasis>
|
|
<taxonomicName id="1E2E1350A89E08A7C0806BBAA728B4B6" lsidName="P. vascularis" pageId="0" pageNumber="223" rank="species" species="vascularis">
|
|
<emphasis id="E63CFC0D36F43D9EBA6F5E0A48464652" italics="true" pageId="0" pageNumber="223">P. vascularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype)
|
|
<emphasis id="C041428B4744588D0F1B377A0BF029C3" bold="true" pageId="0" pageNumber="223">D</emphasis>
|
|
hindwing
|
|
<emphasis id="1DD9506B0271A6E4C26155BA6B2A3B3A" bold="true" pageId="0" pageNumber="223">E</emphasis>
|
|
cubital pecten
|
|
<emphasis id="A775126299C1057CB4D8C8F68D9D51F6" bold="true" pageId="0" pageNumber="223">F</emphasis>
|
|
axillary hair pencils.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="91416F991F8408756C5ED78D6E9A28F3" pageId="0" pageNumber="223">
|
|
<emphasis id="CBA98AE303E38C907D45162E55BCD9B6" bold="true" pageId="0" pageNumber="223">Female</emphasis>
|
|
(Fig.
|
|
<figureCitation id="5507CCC122FF8B9FB0FA3691E20B0457" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Adults of Phaecadophora spp. A P. dactylina sp. nov. (holotype, male) B P. dactylina sp. nov. (paratype, female) C P. vascularis sp. nov. (holotype, male). Scale bars: 2 mm." figureDoi="10.3897/zookeys.1187.111101.figure1" httpUri="https://binary.pensoft.net/fig/956181" pageId="0" pageNumber="223">1B</figureCitation>
|
|
) with forewing length 9.0-10.0 mm.
|
|
<emphasis id="78E50EE6F8945D75E7508E0800A86513" bold="true" italics="true" pageId="0" pageNumber="223">Head</emphasis>
|
|
(Fig.
|
|
<figureCitation id="60901AACB32C2826601899ECDF8273C1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Phaecadophora spp. A, B P. dactylina sp. nov. (holotype, male) A lateral view B dorsal view C, D P. dactylina sp. nov. (paratype, female) C lateral view D dorsal view E, F P. vascularis sp. nov. (holotype, male) E lateral view F dorsal view." figureDoi="10.3897/zookeys.1187.111101.figure2" httpUri="https://binary.pensoft.net/fig/956182" pageId="0" pageNumber="223">2C, D</figureCitation>
|
|
): vertex and upper frons rough, paler brown; scales shiny gray distally; lower frons with tawny appressed scales. Antenna brown, extending to middle of forewing costa. Ocellus well developed; chaetosema present. Labial palpus ascending, mostly pale brown, paler on inner surface and base; median segment expanded distally, terminal segment porrect, rather slender.
|
|
</paragraph>
|
|
<paragraph id="9653701BE9F94A4286CEF826E1D0C6DC" pageId="0" pageNumber="223">
|
|
<emphasis id="36A6799B0881424D8E52D0ECD2084D6E" bold="true" italics="true" pageId="0" pageNumber="223">Thorax</emphasis>
|
|
: brown-fuscous. Legs normal. Forewing subrectangular, slightly dilated towards termen, costa curved evenly, termen straight, tornus rounded; upperside with upper 3/4 longitudinally, finely striped with tawny striae from concolorous pairs of costal strigulae and pale brown broken markings, slightly mottled, except a short streak on outer edge of cell; area below 1A+2A and CuA1 fuscous, suffused with blackish fuscous, upper edge wavy, produced at middle of fold and base of CuA1; cilia fuscous, suffused with brown; underside tawny, pairs of strigulae on costa tawny, area of hindwing overlap white. Hindwing brown-fuscous; costa area of forewing overlap white; pecten distinct; inner side unmodified; cilia pale brown, with brown-fuscous baseline; underside brown.
|
|
</paragraph>
|
|
<paragraph id="54FDA3AFD7995D717FF099F2965FF832" pageId="0" pageNumber="223">
|
|
<emphasis id="D519CDEC895A2C17572A687EF3EF43F5" bold="true" pageId="0" pageNumber="223">
|
|
<emphasis id="ED8597A04B026AFAD311E1D2CFB5E610" italics="true" pageId="0" pageNumber="223">Abdomen</emphasis>
|
|
:
|
|
<emphasis id="6ADC5954F986CB746B963F4B52D05647" italics="true" pageId="0" pageNumber="223">male genitalia</emphasis>
|
|
</emphasis>
|
|
(Fig.
|
|
<figureCitation id="9492C6B55BC9CB0E332CCA5E44D1468A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Phaecadophora dactylina sp. nov. (holotype) A male genitalia B costal prominence of valva C ventral prominence of cucullus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure5" httpUri="https://binary.pensoft.net/fig/956185" pageId="0" pageNumber="223">5A</figureCitation>
|
|
) with tegumen high and narrow. Uncus slender, densely covered with spines; apex bifurcated, with short spines. Socius small, oval, densely covered with spines. Gnathos membranous, forming a broad band. Valva robust, curved, constricted beyond basal excavation, sacculus nearly half length of valva; a short prominence (Fig.
|
|
<figureCitation id="416177626FA63A62F73C625B61B598E8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Phaecadophora dactylina sp. nov. (holotype) A male genitalia B costal prominence of valva C ventral prominence of cucullus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure5" httpUri="https://binary.pensoft.net/fig/956185" pageId="0" pageNumber="223">5B</figureCitation>
|
|
) above the rim of basal excavation below costa, about half of uncus in length, finger-like, apex broadened, with short, dense spines; sacclus well defined, weakly spined, with only sparse, fine hairs beyond basal excavation and along ventral edge, and bearing a spine cluster on midlength of ventral edge; cucullus somewhat elongately triangular, basal 1/3 naked except for strongly protruding ventroproximal base which bears dense, short bristles and a tuft of long bristles apically (Fig.
|
|
<figureCitation id="2C24E7574719233AC38B7762177E294E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Phaecadophora dactylina sp. nov. (holotype) A male genitalia B costal prominence of valva C ventral prominence of cucullus. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure5" httpUri="https://binary.pensoft.net/fig/956185" pageId="0" pageNumber="223">5C</figureCitation>
|
|
), these long bristles longer than uncus; distal 2/3 of cucullus with dense spines. Phallus short, straight, without cornuti.
|
|
<emphasis id="C644E91983099A4366F58F94CF0322FD" bold="true" italics="true" pageId="0" pageNumber="223">Female genitalia</emphasis>
|
|
(Fig.
|
|
<figureCitation id="44D71A6C7B2E5D80982592F119A6F5CC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of Phaecadophora dactylina sp. nov. (paratype) A female genitalia B sterigma and colliculum C signum. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure6" httpUri="https://binary.pensoft.net/fig/956186" pageId="0" pageNumber="223">6A</figureCitation>
|
|
) with papillae anales narrow, densely setose. Anterior apophysis a little longer than posterior apophysis. Sterigma (Fig.
|
|
<figureCitation id="EC2856AF22C973EE5CDEBFA086AA44C8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of Phaecadophora dactylina sp. nov. (paratype) A female genitalia B sterigma and colliculum C signum. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure6" httpUri="https://binary.pensoft.net/fig/956186" pageId="0" pageNumber="223">6B</figureCitation>
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) derived from a raised spinulose fold encircling ostium, with a dorsal notch and posterior portion on each side produced into a broad plate. Colliculum about 1/3 times of length of ductus bursae, strongly sclerotized, caudally wrench-like, abruptly narrowed below, the rest of ductus bursae membranous, ductus seminalis originating posterior to midlength. Corpus bursae ovoid, granulated; signa (Fig.
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<figureCitation id="6F5C6D71AE59ECAE95C0344B071B22F7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genitalia of Phaecadophora dactylina sp. nov. (paratype) A female genitalia B sterigma and colliculum C signum. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1187.111101.figure6" httpUri="https://binary.pensoft.net/fig/956186" pageId="0" pageNumber="223">6C</figureCitation>
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) two, double-folded, roughly equal in size, generally leaf-like.
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</paragraph>
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<caption id="E0B9EDC3F65210F9D2075C8AEDDE3946" doi="10.3897/zookeys.1187.111101.figure5" httpUri="https://binary.pensoft.net/fig/956185" pageId="0" pageNumber="223" start="Figure 5" startId="F5">
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<paragraph id="64E6D496DD9ED0466F4428C4913BE8BF" pageId="0" pageNumber="223">
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<emphasis id="FC72666E259F807726972DDEA2993261" bold="true" pageId="0" pageNumber="223">Figure 5.</emphasis>
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Male genitalia of
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<taxonomicName id="1C30BB0E833DF935004EB617305AFDE0" authorityName="Li & Yu" authorityYear="2023" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora dactylina" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="dactylina">
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<emphasis id="BFB8E1130C4620F196143755B537C515" italics="true" pageId="0" pageNumber="223">Phaecadophora dactylina</emphasis>
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</taxonomicName>
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sp. nov. (holotype)
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<emphasis id="6348B1EC24F2A5FD298A1A0D938095BB" bold="true" pageId="0" pageNumber="223">A</emphasis>
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male genitalia
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<emphasis id="6218F357CF9541D5ADBDE85AACA63B61" bold="true" pageId="0" pageNumber="223">B</emphasis>
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costal prominence of valva
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<emphasis id="B7434F7DE9CA42ABCDC4E8598ADE40A6" bold="true" pageId="0" pageNumber="223">C</emphasis>
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ventral prominence of cucullus. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption id="D8CDDDB29A9414120ACDF06FA90F0709" doi="10.3897/zookeys.1187.111101.figure6" httpUri="https://binary.pensoft.net/fig/956186" pageId="0" pageNumber="223" start="Figure 6" startId="F6">
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<paragraph id="77929720D4A18211B888A283018DA1A0" pageId="0" pageNumber="223">
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<emphasis id="54B51A00650EEAFA934C8C43E69216E0" bold="true" pageId="0" pageNumber="223">Figure 6.</emphasis>
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Female genitalia of
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<taxonomicName id="73FECB680C1C7080A7ED7D356385BB23" authorityName="Li & Yu" authorityYear="2023" class="Insecta" family="Tortricidae" genus="Phaecadophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaecadophora dactylina" order="Lepidoptera" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="dactylina">
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<emphasis id="85CBF9AAEF7912CC5499869079502A1B" italics="true" pageId="0" pageNumber="223">Phaecadophora dactylina</emphasis>
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</taxonomicName>
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sp. nov. (paratype)
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<emphasis id="7760FED0F704924E2B7F061FBB1FBFB9" bold="true" pageId="0" pageNumber="223">A</emphasis>
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female genitalia
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<emphasis id="3F408853E5613B3043B3ED4FD068AB72" bold="true" pageId="0" pageNumber="223">B</emphasis>
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sterigma and colliculum
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<emphasis id="BC7A91E8B783418C5726D3391351C531" bold="true" pageId="0" pageNumber="223">C</emphasis>
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signum. Scale bars: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="F2F0D62441E942F4A5366424F2379D17" pageId="0" pageNumber="223" type="etymology">
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<paragraph id="474D1D39A9F2092B06BA91BB6723B7C8" pageId="0" pageNumber="223">Etymology.</paragraph>
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<paragraph id="AE3BA52DD03F413BB466A84642978F62" pageId="0" pageNumber="223">
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The specific name is derived from the Latin
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<emphasis id="98369B73DC71836D4505EF3208CE8308" italics="true" pageId="0" pageNumber="223">dactylinus</emphasis>
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(= finger-like), referring to the shape of costal prominence of the valva in the male genitalia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |