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<document ID-DOI="http://dx.doi.org/10.3897/jhr.80.56178" ID-GBIF-Dataset="c03ce5c7-7805-4f32-95db-d4ce50995274" ID-Pensoft-Pub="1314-2607-80-99" ID-Pensoft-UUID="BCA40A8EFADF5E8A9E51A75C7C3C0D1C" ID-Zenodo-Dep="4420570" ID-ZooBank="F9DF28B5BF7545F5860B3FB234679C6D" ModsDocID="1314-2607-80-99" checkinTime="1609325082188" checkinUser="pensoft" docAuthor="Komeda, Yoto, Mita, Toshiharu, Hirose, Yoshimi &amp; Yamagishi, Kenzo" docDate="2020" docId="D55B6BB67CDA5D99B2290DE7281F23FA" docLanguage="en" docName="JourHymenoptRes 80: 99-135" docOrigin="Journal of Hymenoptera Research 80" docPubDate="2020-12-29" docSource="http://dx.doi.org/10.3897/jhr.80.56178" docTitle="Gryon yamagishii Mineo 1981" docType="treatment" docVersion="5" id="BCA40A8EFADF5E8A9E51A75C7C3C0D1C" lastPageNumber="99" masterDocId="BCA40A8EFADF5E8A9E51A75C7C3C0D1C" masterDocTitle="Taxonomic revision of charon -, floridanum - and muscaeforme - groups of Gryon Haliday, 1833 (Hymenoptera, Scelionidae) from Japan, with descriptions of two new species and host information" masterLastPageNumber="135" masterPageNumber="99" pageNumber="99" updateTime="1678754838156" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Taxonomic revision of charon -, floridanum - and muscaeforme - groups of Gryon Haliday, 1833 (Hymenoptera, Scelionidae) from Japan, with descriptions of two new species and host information</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Komeda, Yoto</mods:namePart>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kome123k123@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mita, Toshiharu</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8322-6045</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hirose, Yoshimi</mods:namePart>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yamagishi, Kenzo</mods:namePart>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Kyushu University, Motooka 744, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
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<mods:start>99</mods:start>
<mods:end>135</mods:end>
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<mods:url>http://dx.doi.org/10.3897/jhr.80.56178</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.80.56178</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-80-99</mods:identifier>
<mods:identifier type="ZooBank">F9DF28B5BF7545F5860B3FB234679C6D</mods:identifier>
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<treatment ID-GBIF-Taxon="176345122" LSID="urn:lsid:plazi:treatment:D55B6BB67CDA5D99B2290DE7281F23FA" httpUri="http://treatment.plazi.org/id/D55B6BB67CDA5D99B2290DE7281F23FA" lastPageNumber="99" pageId="0" pageNumber="99">
<subSubSection pageId="0" pageNumber="99" type="nomenclature">
<paragraph pageId="0" pageNumber="99">
<taxonomicName ID-CoL="3HHLH" LSID="D55B6BB6-7CDA-5D99-B229-0DE7281F23FA" authority="Mineo, 1981" authorityName="Mineo" authorityYear="1981" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">Gryon yamagishii Mineo, 1981</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Japanese Gryon spp., dorsal view A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., holotype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure1" httpUri="https://binary.pensoft.net/fig/490334" pageId="0" pageNumber="99">Figs 1C</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Japanese Gryon spp., lateral views A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., paratype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure2" httpUri="https://binary.pensoft.net/fig/490335" pageId="0" pageNumber="99">, 2C</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frons of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure3" httpUri="https://binary.pensoft.net/fig/490336" pageId="0" pageNumber="99">, 3C</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">, 4C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Japanese Gryon spp. A G. fulvicoxa sp. nov., female B G. japonicum, female C G. yamagishii, female D G. philippinense, female E G. shisa sp. nov., female F G. pennsylvanicum, female G G. fulvicoxa sp. nov., male H G. japonicum, male I G. yamagishii, male J G. philippinense, male K G. shisa sp. nov., male L G. pennsylvanicum, male." figureDoi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99">, 5C, I</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Forewing of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure6" httpUri="https://binary.pensoft.net/fig/490339" pageId="0" pageNumber="99">, 6C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="reference_group">
<paragraph pageId="0" pageNumber="99">
<taxonomicName authorityName="Mineo" authorityYear="1981" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">Gryon yamagishii</taxonomicName>
Mineo, 1981:
<bibRefCitation author="Johnson, NF" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="99" pagination="1 - 825" refId="B23" refString="Johnson, NF, 1992. Catalog of World Proctotrupoidea Excluding Platygastridae. Memoirs of the American Entomological Institute 51: 1 - 825" title="Catalog of World Proctotrupoidea Excluding Platygastridae." volume="51" year="1992">Johnson 1992</bibRefCitation>
;
<bibRefCitation author="Kononova, SV" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="99" pagination="53 - 59" refId="B32" refString="Kononova, SV, Petrov, SD, 2002. A review of the genera Gryon and Exon (Hymenoptera, Scelionidae) from the Palaearctic. 2. A key for identification of Gryon species and a review of the genus Exon. Zoologicheskii Zhurnal 81 (1): 53 - 59" title="A review of the genera Gryon and Exon (Hymenoptera, Scelionidae) from the Palaearctic. 2. A key for identification of Gryon species and a review of the genus Exon." volume="81" year="2002">Kononova and Petrov 2002</bibRefCitation>
;
<bibRefCitation author="Kononova, SV" journalOrPublisher="Tovarishchestvo Nauchnykh Izdanii KMK, Saint Petersburg-Moscow" pageId="0" pageNumber="99" refId="B31" refString="Kononova, SV, Kozlov, MA, 2008. Scelionids of the Palearctic (Hymenoptera, Scelionidae). Subfamily Scelioninae. Tovarishchestvo Nauchnykh Izdanii KMK, Saint Petersburg-Moscow" title="Scelionids of the Palearctic (Hymenoptera, Scelionidae). Subfamily Scelioninae." year="2008">Kononova and Kozlov 2008</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName authorityName="Mineo" authorityYear="1981" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon maruzzae" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="maruzzae">Gryon maruzzae</taxonomicName>
Mineo, 1981:
<bibRefCitation author="Johnson, NF" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="99" pagination="1 - 825" refId="B23" refString="Johnson, NF, 1992. Catalog of World Proctotrupoidea Excluding Platygastridae. Memoirs of the American Entomological Institute 51: 1 - 825" title="Catalog of World Proctotrupoidea Excluding Platygastridae." volume="51" year="1992">Johnson 1992</bibRefCitation>
;
<bibRefCitation author="Kononova, SV" journalOrPublisher="Tovarishchestvo Nauchnykh Izdanii KMK, Saint Petersburg-Moscow" pageId="0" pageNumber="99" refId="B31" refString="Kononova, SV, Kozlov, MA, 2008. Scelionids of the Palearctic (Hymenoptera, Scelionidae). Subfamily Scelioninae. Tovarishchestvo Nauchnykh Izdanii KMK, Saint Petersburg-Moscow" title="Scelionids of the Palearctic (Hymenoptera, Scelionidae). Subfamily Scelioninae." year="2008">Kononova and Kozlov 2008</bibRefCitation>
. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName authorityName="Kozlov &amp; Kononova" authorityYear="1989" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon sugonjaevi" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sugonjaevi">Gryon sugonjaevi</taxonomicName>
Kozlov &amp; Kononova, 1989:
<bibRefCitation author="Kozlov, MA" journalOrPublisher="Nauka, Leningrad" pageId="0" pageNumber="99" refId="B35" refString="Kozlov, MA, Kononova, SV, 1990. Scelioninae of the Fauna of the USSR (Hymenoptera, Scelionidae, Scelioninae). Nauka, Leningrad" title="Scelioninae of the Fauna of the USSR (Hymenoptera, Scelionidae, Scelioninae)." year="1990">Kozlov and Kononova 1990</bibRefCitation>
;
<bibRefCitation author="Johnson, NF" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="99" pagination="1 - 825" refId="B23" refString="Johnson, NF, 1992. Catalog of World Proctotrupoidea Excluding Platygastridae. Memoirs of the American Entomological Institute 51: 1 - 825" title="Catalog of World Proctotrupoidea Excluding Platygastridae." volume="51" year="1992">Johnson 1992</bibRefCitation>
;
<bibRefCitation author="Kononova, SV" editor="Lehr, PA" journalOrPublisher="vol. 4. Neuropteroidea, Mecoptera, Hymenoptera. Part 2. Dal'nauka, Vladivostok" pageId="0" pageNumber="99" pagination="57 - 121" refId="B29" refString="Kononova, SV, 1995. 25. Fam. Scelionidae. In: Lehr, PA, Ed., Key to insects of Russian Far East in six volume. vol. 4. Neuropteroidea, Mecoptera, Hymenoptera. Part 2. Dal'nauka, Vladivostok: 57 - 121" title="25. Fam. Scelionidae." volumeTitle="Key to insects of Russian Far East in six volume." year="1995">Kononova 1995</bibRefCitation>
;
<bibRefCitation author="Kononova, SV" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="99" pagination="53 - 59" refId="B32" refString="Kononova, SV, Petrov, SD, 2002. A review of the genera Gryon and Exon (Hymenoptera, Scelionidae) from the Palaearctic. 2. A key for identification of Gryon species and a review of the genus Exon. Zoologicheskii Zhurnal 81 (1): 53 - 59" title="A review of the genera Gryon and Exon (Hymenoptera, Scelionidae) from the Palaearctic. 2. A key for identification of Gryon species and a review of the genus Exon." volume="81" year="2002">Kononova and Petrov 2002</bibRefCitation>
;
<bibRefCitation author="Kononova, SV" journalOrPublisher="Tovarishchestvo Nauchnykh Izdanii KMK, Saint Petersburg-Moscow" pageId="0" pageNumber="99" refId="B31" refString="Kononova, SV, Kozlov, MA, 2008. Scelionids of the Palearctic (Hymenoptera, Scelionidae). Subfamily Scelioninae. Tovarishchestvo Nauchnykh Izdanii KMK, Saint Petersburg-Moscow" title="Scelionids of the Palearctic (Hymenoptera, Scelionidae). Subfamily Scelioninae." year="2008">Kononova and Kozlov 2008</bibRefCitation>
. syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="diagnosis">
<paragraph pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="99">Frontal depression transversely costate by strong regular carinae. Angular points of occipital carina modified as short sharp horns; horizontal portion of occipital carina straight, expanding inwardly. Coxae brown-black.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Female.</emphasis>
Length = 1.4-1.6 mm.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Color</emphasis>
.
</emphasis>
(Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Japanese Gryon spp., dorsal view A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., holotype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure1" httpUri="https://binary.pensoft.net/fig/490334" pageId="0" pageNumber="99">1C</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Japanese Gryon spp., lateral views A G. fulvicoxa sp. nov., holotype B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov., paratype F G. pennsylvanicum. Scale bars: 1 mm." figureDoi="10.3897/jhr.80.56178.figure2" httpUri="https://binary.pensoft.net/fig/490335" pageId="0" pageNumber="99">2C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="99">Body</emphasis>
mainly black. A2-6 brown. A1, mandibles, and legs (excluding coxae) yellow.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Head</emphasis>
.
</emphasis>
FCI = 1.16-1.28; LCI = 1.66-1.62; DCI = 2.03-2.23; HW/IOS = 1.64-1.78; head about 1.3 times as wide as mesosoma (HW/TSL = 1.24-1.30). Frons (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frons of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure3" httpUri="https://binary.pensoft.net/fig/490336" pageId="0" pageNumber="99">3C</figureCitation>
) reticulate with setae; central carina present ventrally; frontal depression weakly developed, transversely costate by strong regular carinae. Vertex reticulate with setae; interocellar space reticulate; hyperoccipital carina absent; POL about 4.6 times as long as OOL (POL/OOL = 3.67-4.83); OOL about 0.4 times as long as LOL (OOL/LOL = 0.33-0.50). Clypeus semi-elliptical. Gena coriaceous with setae; medial genal carina absent. Occiput (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4C</figureCitation>
) transversely costate with setae; occipital carina complete; angular points of occipital carina well-developed, as short sharp horns; horizontal portion of occipital carina straight, expanding inwardly; postoccipital carina discontinuity present; postgena weakly costate along postoccipital carina; postgenal sulcus curved toward hypostoma; postgenal bridge smooth, weakly longitudinally costate beside median sulcus. Antennae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Japanese Gryon spp. A G. fulvicoxa sp. nov., female B G. japonicum, female C G. yamagishii, female D G. philippinense, female E G. shisa sp. nov., female F G. pennsylvanicum, female G G. fulvicoxa sp. nov., male H G. japonicum, male I G. yamagishii, male J G. philippinense, male K G. shisa sp. nov., male L G. pennsylvanicum, male." figureDoi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99">5C</figureCitation>
) clavate; A1 about 3.5 times longer than radicle, as long as clava; clava with five segments; claval sensilla formula A8-12/2-2-2-2-1; claval length about 3.4 times longer than width. Mandibles thin, tridentate, anterior tooth longer than other teeth.
</paragraph>
<caption ID-Zenodo-Dep="4420586" doi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Figure 5.</emphasis>
Antennae of Japanese
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Gryon</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. fulvicoxa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fulvicoxa">
<emphasis italics="true" pageId="0" pageNumber="99">G. fulvicoxa</emphasis>
</taxonomicName>
sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. japonicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="japonicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. japonicum</emphasis>
</taxonomicName>
, female
<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
, female
<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
, female
<emphasis bold="true" pageId="0" pageNumber="99">E</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa">
<emphasis italics="true" pageId="0" pageNumber="99">G. shisa</emphasis>
</taxonomicName>
sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="99">F</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. pennsylvanicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="pennsylvanicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. pennsylvanicum</emphasis>
</taxonomicName>
, female
<emphasis bold="true" pageId="0" pageNumber="99">G</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. fulvicoxa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fulvicoxa">
<emphasis italics="true" pageId="0" pageNumber="99">G. fulvicoxa</emphasis>
</taxonomicName>
sp. nov., male
<emphasis bold="true" pageId="0" pageNumber="99">H</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. japonicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="japonicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. japonicum</emphasis>
</taxonomicName>
, male
<emphasis bold="true" pageId="0" pageNumber="99">I</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
, male
<emphasis bold="true" pageId="0" pageNumber="99">J</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
, male
<emphasis bold="true" pageId="0" pageNumber="99">K</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa">
<emphasis italics="true" pageId="0" pageNumber="99">G. shisa</emphasis>
</taxonomicName>
sp. nov., male
<emphasis bold="true" pageId="0" pageNumber="99">L</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. pennsylvanicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="pennsylvanicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. pennsylvanicum</emphasis>
</taxonomicName>
, male.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="4420588" doi="10.3897/jhr.80.56178.figure6" httpUri="https://binary.pensoft.net/fig/490339" pageId="0" pageNumber="99" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Figure 6.</emphasis>
Forewing of Japanese
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Gryon</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. fulvicoxa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fulvicoxa">
<emphasis italics="true" pageId="0" pageNumber="99">G. fulvicoxa</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. japonicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="japonicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. japonicum</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. philippinense" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="philippinense">
<emphasis italics="true" pageId="0" pageNumber="99">G. philippinense</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="99">E</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. shisa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="shisa">
<emphasis italics="true" pageId="0" pageNumber="99">G. shisa</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="99">F</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. pennsylvanicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="pennsylvanicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. pennsylvanicum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Mesosoma</emphasis>
.
</emphasis>
Cervical pronotal area granulate-punctate with dense setae; epomial carina strongly present, not reaching dorsal edge; pronotal suprahumeral sulcus foveolate, unclear mesad; lateral pronotal area smooth with transverse dense carinae. Propleuron weakly transversely costate. Mesoscutum about 1.4 times as wide as long (TSL/ML = 1.35-1.50), with dense setae, reticulate in anterior, longitudinally costate in posterior; parascutal carina absent; notaulus absent. Mesoscutellum about 2.2 times as wide as long (SW/SL = 2.07-2.35), with dense setae, longitudinally costate mesad, granulate laterad. Mesopleuron smooth with transverse dense carinae above mesopleural canina, rugulose with setae below mesopleural canina; prespecular and upper mesepisternal sulci foveolate; prespecular sulcus with setae; mesopleural carina strongly present; postacetabular sulcus foveolate. Metascutellum weakly produced, longitudinally striate. Metapleuron weakly convex, foveolate-rugulose, with dense setae posteriorly; anterior part of metapleural sulcus and upper paracoxal sulcus with setae. Propodeum foveolate with setae. Fore wing (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mesosoma of Japanese Gryon spp. A G. japonicum, dorsal view B lateral view C G. philippinense, dorsal view D lateral view E G. pennsylvanicum, dorsal view F lateral view." figureDoi="10.3897/jhr.80.56178.figure7" httpUri="https://binary.pensoft.net/fig/490340" pageId="0" pageNumber="99">7C</figureCitation>
); stigmal vein about 2.5 times longer than marginal vein; postmarginal vein about 3.7 times longer than marginal vein.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">Metasoma</emphasis>
.
</emphasis>
T1 longitudinally striate, setose laterally. S1 longitudinally striate. T2 longitudinally striate anteriorly, reticulate posteriorly, setose laterally. S2 with setae, granulate mesad, striate laterad. T3 reticulate, with setae laterad and posteriorly. S3-6 punctate with setae. T4 punctate-striate with setae. T5-6 punctate with setae.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Male.</emphasis>
Almost same as female, but antennae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Japanese Gryon spp. A G. fulvicoxa sp. nov., female B G. japonicum, female C G. yamagishii, female D G. philippinense, female E G. shisa sp. nov., female F G. pennsylvanicum, female G G. fulvicoxa sp. nov., male H G. japonicum, male I G. yamagishii, male J G. philippinense, male K G. shisa sp. nov., male L G. pennsylvanicum, male." figureDoi="10.3897/jhr.80.56178.figure5" httpUri="https://binary.pensoft.net/fig/490338" pageId="0" pageNumber="99">5I</figureCitation>
) filiform; A1 yellow, A2-11 brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="host">
<paragraph pageId="0" pageNumber="99">Host.</paragraph>
<paragraph pageId="0" pageNumber="99">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="biology_ecology">
<paragraph pageId="0" pageNumber="99">Biology.</paragraph>
<paragraph pageId="0" pageNumber="99">
In winter, the female overwinters under the bark of
<taxonomicName class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Zelkova serrata" order="Rosales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis italics="true" pageId="0" pageNumber="99">Zelkova serrata</emphasis>
</taxonomicName>
(Thunberg) Makino (
<taxonomicName authorityName="Spach" authorityYear="1841" class="Magnoliopsida" family="Ulmaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="family">Ulmaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
<paragraph pageId="0" pageNumber="99">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="99">
<materialsCitation ID-GBIF-Occurrence="3026168301" collectingDate="1958-07-25" collectingDateMax="1981-10-04" collectingDateMin="1958-07-25" collectionCode="ELKU" collectorName="K. Yamagishi, Y. Miyatake, SZ Nagoya., E. Sugonjaev" country="Japan" county="Chikuzen" location="Kooridono" municipality="Kyushu" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Niigata Pref." typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
:
<location LSID="urn:lsid:plazi:treatment:D55B6BB67CDA5D99B2290DE7281F23FA:2E1FFE7461188572F22C1C12392AF88E" country="Japan" county="Chikuzen" municipality="Kyushu" name="Kooridono" stateProvince="Niigata Pref.">
<collectingMunicipality>Kooridono</collectingMunicipality>
</location>
,
<location LSID="urn:lsid:plazi:treatment:D55B6BB67CDA5D99B2290DE7281F23FA:664AA54A742AAC5D7E5224DC72734905" country="Japan" county="Chikuzen" municipality="Kyushu" name="Ojiya" stateProvince="Niigata Pref.">Ojiya</location>
,
<collectingRegion country="Japan" name="Niigata">
<location LSID="urn:lsid:plazi:treatment:D55B6BB67CDA5D99B2290DE7281F23FA:18AD2FB718BE5925868F14C268B15088" municipality="Kooridono" name="Niigata Pref">Niigata Pref</location>
.
</collectingRegion>
[
<collectingCountry name="Japan">Japan</collectingCountry>
:
<collectingRegion country="Japan" name="Niigata">Niigata pref.</collectingRegion>
, Ojiya city, Nishi-Yoshidani,
<normalizedToken originalValue="Kôridono">Koridono</normalizedToken>
.]
<collectingDate value="1970-08-09">9.VIII.1970</collectingDate>
.
<collectorName>K. Yamagishi</collectorName>
leg.
<specimenCount type="female">♀</specimenCount>
[
<collectionCode>ELKU</collectionCode>
] (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. The holotype of G. yamagishii A dorsal view B lateral view C mesosoma, dorsal view D frons E labels." figureDoi="10.3897/jhr.80.56178.figure10" httpUri="https://binary.pensoft.net/fig/490343" pageId="0" pageNumber="99">10</figureCitation>
).
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. maruzzae" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="maruzzae">
<emphasis italics="true" pageId="0" pageNumber="99">G. maruzzae</emphasis>
</taxonomicName>
: (
<collectingMunicipality>Kyushu</collectingMunicipality>
) Okinoshima (
<collectingCounty>Chikuzen</collectingCounty>
). [
<collectingCountry name="Japan">Japan</collectingCountry>
:
<collectingRegion country="Japan" name="Fukuoka">Fukuoka pref.</collectingRegion>
, Munakata dist., Munakata town, Chikuzen Oki-no-shima Isl.]
<collectingDate value="1958-07-25" valueMax="1958-07-28" valueMin="1958-07-25">25-28.VII.1958</collectingDate>
, Hirashima, Murakami and
<collectorName>Y. Miyatake</collectorName>
leg.
<specimenCount type="male">1♂</specimenCount>
[ELKU] (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. The holotype of G. maruzzae syn. nov. A dorsal view B lateral view C mesosoma, dorsal view D frons E labels." figureDoi="10.3897/jhr.80.56178.figure11" httpUri="https://binary.pensoft.net/fig/490344" pageId="0" pageNumber="99">11</figureCitation>
).
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sugonjaevi" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sugonjaevi">
<emphasis italics="true" pageId="0" pageNumber="99">G. sugonjaevi</emphasis>
</taxonomicName>
:
<normalizedToken originalValue="Япония">Yaponiya</normalizedToken>
,
<normalizedToken originalValue="Инуяма">Inuyama</normalizedToken>
,
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.0" unit="km" value="40.0">
40
<normalizedToken originalValue="км">km</normalizedToken>
</quantity>
<collectorName>
<normalizedToken originalValue="СЗ">SZ</normalizedToken>
<normalizedToken originalValue="Нагоя">Nagoya</normalizedToken>
.
</collectorName>
[=
<collectingCountry name="Japan">Japan</collectingCountry>
:
<collectingRegion country="Japan" name="Aichi">Aichi pref.</collectingRegion>
, Inuyama city.]
<collectingDate value="1981-10-04">4.X.1981</collectingDate>
.
<collectorName>E. Sugonjaev</collectorName>
leg.
<specimenCount>♀</specimenCount>
[ZIN].
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="99">
<materialsCitation collectingDate="1970-05-05" collectingDateMax="1981-10-19" collectingDateMin="1970-05-05" collectorName="K. Yamagishi, E. Sugonjaev" country="Japan" county="Chikuzen" location="Nisshin town" municipality="Kakamigahara" specimenCount="♀" stateProvince="Aichi pref." typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="99">
<emphasis italics="true" pageId="0" pageNumber="99">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
: same locality as holotype,
<collectingDate value="1970-08-01">1.VIII.1970</collectingDate>
.
<collectorName>K. Yamagishi</collectorName>
leg.
<specimenCount>♀</specimenCount>
[ELKU]; Nisshin, Aichi-gun,
<collectingRegion country="Japan" name="Aichi">Aichi pref.</collectingRegion>
[
<collectingRegion country="Japan" name="Aichi">Aichi pref.</collectingRegion>
,
<collectingRegion country="Japan" name="Aichi">Aichi dist.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D55B6BB67CDA5D99B2290DE7281F23FA:F6D7D27BB92E98522DCC8BA26B798DAD" country="Japan" county="Chikuzen" municipality="Kakamigahara" name="Nisshin town" stateProvince="Aichi pref.">Nisshin town</location>
],
<collectingDate value="1970-05-05">5.V.1970</collectingDate>
.
<collectorName>K. Yamagishi</collectorName>
leg.
<specimenCount>♀</specimenCount>
[ELKU].
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sugonjaevi" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sugonjaevi">
<emphasis italics="true" pageId="0" pageNumber="99">G. sugonjaevi</emphasis>
</taxonomicName>
:
<normalizedToken originalValue="Япония">Yaponiya</normalizedToken>
,
<normalizedToken originalValue="Каганихара">Kaganikhara</normalizedToken>
. [=
<collectingCountry name="Japan">Japan</collectingCountry>
:
<collectingRegion country="Japan" name="Gifu">Gifu pref.</collectingRegion>
,
<collectingMunicipality>Kakamigahara</collectingMunicipality>
city]
<collectingDate value="1981-10-19">19.X.1981</collectingDate>
.
<collectorName>E. Sugonjaev</collectorName>
leg.
<specimenCount>♀</specimenCount>
[IZAN]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
<paragraph pageId="0" pageNumber="99">Other material.</paragraph>
<paragraph pageId="0" pageNumber="99">
<collectingCountry name="Japan">Japan</collectingCountry>
:
<collectingRegion country="Japan" name="Hokkaido">Hokkaido</collectingRegion>
, Sapporo city, Toyohira ward, Hitsujigaoka.
<geoCoordinate degrees="43.008" direction="north" orientation="latitude" precision="55" value="43.008">43.008°N</geoCoordinate>
,
<geoCoordinate degrees="141.415" direction="east" orientation="longitude" precision="55" value="141.415">141.415°E</geoCoordinate>
; alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
,
<date value="2010-06-08" valueMax="2010-06-15" valueMin="2010-06-08">8-15.VI.2010</date>
, Kazuhiko Konishi leg. (MT)
<specimenCount type="female">5♀</specimenCount>
[ELKU];
<date value="2010-06-15" valueMax="2010-06-22" valueMin="2010-06-15">15-22.VI.2010</date>
, Kazuhiko Konishi leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[ELKU];
<date value="2010-06-22" valueMax="2010-06-29" valueMin="2010-06-22">22-29.VI.2010</date>
, Kazuhiko Konishi leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[EUMJ];
<date value="2010-06-29" valueMax="2010-07-06" valueMin="2010-06-29">29.VI.-6.VII.2010</date>
, Kazuhiko Konishi leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[EUMJ];
<date value="2010-07-27" valueMax="2010-08-03" valueMin="2010-07-27">27.VII.-3.VIII.2010</date>
, Kazuhiko Konishi leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[EUMJ];
<date value="2010-08-10" valueMax="2010-08-17" valueMin="2010-08-10">10-17.VIII.2010</date>
, Kazuhiko Konishi leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[EUMJ];
<collectingRegion country="Japan" name="Fukushima">Fukushima pref.</collectingRegion>
, Aizu-Wakamatsu city, Monden-machi, Kuroiwa, Minami-Aoki.
<date value="2017-01-06">6.I.2017</date>
. Keisuke Narita leg. (Collected from under the bark of
<taxonomicName class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Zelkova serrata" order="Rosales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis italics="true" pageId="0" pageNumber="99">Zelkova serrata</emphasis>
</taxonomicName>
)
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Ibaraki">Ibaraki pref.</collectingRegion>
, Tsuchiura city (near Tsukuba), Pond
<normalizedToken originalValue="Shishituka-Ôike">Shishituka-Oike</normalizedToken>
.
<date value="1997-07-15">15.VII.1997</date>
. Victor Fursov leg.
<specimenCount type="female">1♀</specimenCount>
[IZAN];
<collectingRegion country="Japan" name="Tochigi">Tochigi pref.</collectingRegion>
, Nikko city, Tamozawa.
<date value="2008-06-20" valueMax="2008-06-25" valueMin="2008-06-20">20-25.VI.2008</date>
. Takeyuki Nakamura leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Niigata">Niigata pref.</collectingRegion>
, Nagaoka city, Urase-machi,
<geoCoordinate degrees="37.464" direction="north" orientation="latitude" precision="55" value="37.464">37.464°N</geoCoordinate>
,
<geoCoordinate degrees="138.907" direction="east" orientation="longitude" precision="55" value="138.907">138.907°E</geoCoordinate>
, alt.
<quantity metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">40 m</quantity>
,
<date value="2015-05-28" valueMax="2015-06-07" valueMin="2015-05-28">2015.V.28.-VI. 7</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="female">6♀</specimenCount>
[ELKU];
<date value="2015-06-07" valueMax="2015-06-18" valueMin="2015-06-07">2015. VI.7.-VI.18</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="female">8♀</specimenCount>
[ELKU];
<date value="2015-07-06" valueMax="2015-07-25" valueMin="2015-07-06">2015.VII.6-25</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="male">5♂</specimenCount>
<specimenCount type="female">17♀</specimenCount>
[ELKU];
<date value="2015-07-25" valueMax="2015-08-07" valueMin="2015-07-25">2015. VII.25-VIII.7</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="male">2♂</specimenCount>
<specimenCount type="female">16♀</specimenCount>
[ELKU];
<date value="2015-08-07" valueMax="2015-08-22" valueMin="2015-08-07">2015.VIII.7-VIII. 22</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="male">5♂</specimenCount>
<specimenCount type="female">2♀</specimenCount>
[ELKU];
<date value="2015-08-22" valueMax="2015-09-05" valueMin="2015-08-22">2015. VIII.22-IX.5</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="male">7♂</specimenCount>
<specimenCount type="female">2♀</specimenCount>
[ELKU];
<date value="2015-09-27" valueMax="2015-10-12" valueMin="2015-09-27">2015. IX.27-X. 12</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="female">6♀</specimenCount>
[ELKU];
<date value="2016-06-11" valueMax="2016-06-23" valueMin="2016-06-11">2016.VI.11-23</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<date value="2016-07-13" valueMax="2016-08-01" valueMin="2016-07-13">2016. VII.13-VIII.1</date>
, Ryo Shimizu and So Shimizu leg. (MT),
<specimenCount type="male">2♂</specimenCount>
<specimenCount type="female">12♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Ishikawa">Ishikawa pref.</collectingRegion>
, Mt. Hakusan.
<date value="1960-08-29" valueMax="1960-08-21" valueMin="1960-08-29">29-21.VIII.1960</date>
. Terunobu Hidaka leg.
<specimenCount type="male">1♂</specimenCount>
[ELKU]; Hakusan city,
<normalizedToken originalValue="San’nomiya">San'nomiya</normalizedToken>
,
<date value="2009-08-09" valueMax="2009-08-22" valueMin="2009-08-09">9-22.VIII.2009</date>
, H. Fukutomi and S. Nakagawa leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Nomi city, Mitsukuchi (paddy field)
<date value="2011-04-30" valueMax="2011-05-13" valueMin="2011-04-30">30.IV-13.V.2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2011-05-13" valueMax="2011-05-26" valueMin="2011-05-13">13-26.V.2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="female">3♀</specimenCount>
[ELMU];
<date value="2011-06-09" valueMax="2011-06-21" valueMin="2011-06-09">9-21.VI.2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="female">3♀</specimenCount>
[ELMU];
<date value="2011-07-17" valueMax="2011-08-04" valueMin="2011-07-17">17.VII.-4.VIII.2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="male">1♂</specimenCount>
[ELMU];
<date value="2011-08-19" valueMax="2011-09-07" valueMin="2011-08-19">19.VIII.-7.IX.2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="male">1♂</specimenCount>
[ELMU];
<date value="2011-09-08" valueMax="2011-09-21" valueMin="2011-09-08">8-21. IX. 2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2011-10-22" valueMax="2011-11-02" valueMin="2011-10-22">22.X.-2.XI.2011</date>
, H. Fukutomi and R. Ishiguro leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<collectingRegion country="Japan" name="Gifu">Gifu pref.</collectingRegion>
, Kani city, Katabira,
<date value="2004-05-01" valueMax="2004-05-07" valueMin="2004-05-01">1-7.V.2004</date>
, K. Ito leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[ELMU];
<date value="2004-08-06" valueMax="2004-08-12" valueMin="2004-08-06">6-12.VIII.2004</date>
, Kenzo Yamagishi leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2004-08-20" valueMax="2004-08-26" valueMin="2004-08-20">20-26.VIII.2004</date>
, Kenzo Yamagishi leg. (MT)
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2004-10-02" valueMax="2004-10-08" valueMin="2004-10-02">2-8.X.2004</date>
. Kenzo Yamagishi leg. (MT)
<specimenCount type="female">3♀</specimenCount>
[ELMU];
<collectingRegion country="Japan" name="Shizuoka">Shizuoka pref.</collectingRegion>
, Iwata city, dike of Ohta river,
<date value="2001-07-29">29.VII.2001</date>
, M. Ura leg. (YPT)
<specimenCount type="female">1♀</specimenCount>
[IZAN];
<date value="2001-08-13">13.VIII.2001</date>
, M. Ura leg. (YPT)
<specimenCount type="male">1♂</specimenCount>
[IZAN];
<date value="2001-08-27">27.VIII.2001</date>
, M. Ura leg. (YPT)
<specimenCount type="male">1♂</specimenCount>
[ELMU];
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[IZAN];
<collectingRegion country="Japan" name="Aichi">Aichi pref.</collectingRegion>
, Nagoya city, Chikusa ward, Higashiyama Park.
<date value="2001-05-01" valueMax="2001-05-10" valueMin="2001-05-01">1-10.V.2001</date>
. M. Watanabe leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[IZAN]; IV-VI. 1997, Victor Fursov leg.
<specimenCount type="female">25♀</specimenCount>
[IZAN];
<date value="1997-06-11">11.VI.1997</date>
. Victor Fursov leg.
<specimenCount type="male">1♂</specimenCount>
[IZAN]; Tempaku ward, Yagoto-Urayama,
<date value="2002-09-21">21.IX.2002</date>
. Kenzo Yamagishi leg.
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Tempaku ward,
<date value="2012-09-29">29.IX.2012</date>
. Kenzo Yamagishi leg.
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Tempaku ward, Meijo University,
<date value="2013-04-17">17.IV.2013</date>
. N. Kusuda leg.
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Tempaku ward, Meijo University,
<date value="2013-07-31">31.VII.2013</date>
. N. Kusuda leg.
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Ichinomiya city, Tomida, Kiso River,
<date value="2013-05-10">10.V.2013</date>
, Y. Miyata leg. (YPT)
<specimenCount type="female">5♀</specimenCount>
[ELMU];
<date value="2013-07-02">2.VII.2013</date>
, Y. Miyata leg. (YPT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2013-08-22">22.VIII.2013</date>
, Y. Miyata leg. (YPT)
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Kasugai city, Takagi (Grassland),
<date value="2013-05-14">14.V.2013</date>
. Y. Kamiya leg. (YPT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2013-07-31">31.VII.2013</date>
. Y. Kamiya leg. (YPT)
<specimenCount type="male">1♂</specimenCount>
[ELMU]; Nisshin city, Akaike,
<date value="2006-09-30">30.IX.2006</date>
. Kenzo Yamagishi leg.
<specimenCount type="female">1♀</specimenCount>
[ELMU]; Nisshin city, Nokata,
<date value="2011-05-28" valueMax="2011-06-03" valueMin="2011-05-28">28.V-3.VI.2011</date>
. H. Seo and R. Mizutani leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2011-06-10" valueMax="2011-06-17" valueMin="2011-06-10">10-17.VI.2011</date>
. H. Seo and R. Mizutani leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU];
<date value="2011-06-02" valueMax="2011-06-09" valueMin="2011-06-02">2-9.VI.2011</date>
. H. Seo and R. Mizutani leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELMU]; [ELMU];
<date value="2011-06-16" valueMax="2011-06-22" valueMin="2011-06-16">16-22.VI.2011</date>
. H. Seo and R. Mizutani leg. (MT)
<specimenCount type="male">1♂</specimenCount>
[ELMU]; Toyota city, Obara,
<normalizedToken originalValue="Kajiyashiki-chô">Kajiyashiki-cho</normalizedToken>
, Nishinohara, 12.VI.-6. 2014. Hiroaki Iketake leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="2014-10-12" valueMax="2014-10-26" valueMin="2014-10-12">12-26.X.2014</date>
. Hiroaki Iketake leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Osaka">Osaka pref.</collectingRegion>
,
<normalizedToken originalValue="Sen’nan">Sen'nan</normalizedToken>
Dist., Misaki town,
<normalizedToken originalValue="Kyôshi">Kyoshi</normalizedToken>
,
<date value="1975-04-16">16.IV.1975</date>
.
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Wakayama">Wakayama pref.</collectingRegion>
,
<collectingRegion country="Japan" name="Wakayama">Wakayama</collectingRegion>
city,
<normalizedToken originalValue="Sandô">Sando</normalizedToken>
,
<date value="1975-04-16">16.IV.1975</date>
.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Shimane">Shimane pref.</collectingRegion>
, Izumo city, Mt. Kita-yama,
<date value="1969-06-11">11.VI.1969</date>
. Minoru
<collectingRegion country="Japan" name="Miyazaki">Miyazaki</collectingRegion>
leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Tokushima">Tokushima pref.</collectingRegion>
,
<normalizedToken originalValue="Zen’nyuji-toh">Zen'nyuji-toh</normalizedToken>
(River island in Yoshino River),
<date value="2003-05-13" valueMax="2003-05-22" valueMin="2003-05-13">13-22.V.2003</date>
. K. Ohara and H. Otsuka leg. (MT)
<specimenCount type="female">3♀</specimenCount>
[ELMU];
<date value="2003-09-05" valueMax="2003-09-17" valueMin="2003-09-05">5-17.IX.2003</date>
. K. Ohara and H. Otsuka leg. (MT)
<specimenCount type="female">3♀</specimenCount>
[ELMU];
<collectingRegion country="Japan" name="Ehime">Ehime pref.</collectingRegion>
, Matsuyama city, Sugitake.
<date value="1976-05-26">26.V.1976</date>
. N. Takaki leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Matsuyama city, Komenono.
<date value="1978-06-01">1.VI.1978</date>
. N. Takaki leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Matsuyama city,
<normalizedToken originalValue="Jikiba-chô">Jikiba-cho</normalizedToken>
.
<geoCoordinate degrees="33.51" direction="north" orientation="latitude" precision="555" value="33.51">33.51°N</geoCoordinate>
,
<geoCoordinate degrees="132.42" direction="east" orientation="longitude" precision="555" value="132.42">132.42°E</geoCoordinate>
;
<date value="2016-05-14">14.V.2016</date>
. K. Kuroda and K. Sogoh leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Matsuyama city, Suemachi, Wakigafuchi Park.
<geoCoordinate degrees="33.52" direction="north" orientation="latitude" precision="555" value="33.52">33.52°N</geoCoordinate>
,
<geoCoordinate degrees="132.49" direction="east" orientation="longitude" precision="555" value="132.49">132.49°E</geoCoordinate>
;
<date value="2017-05-20">20.V.2017</date>
. K. Kuroda leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Ochi dist., Kamijima town, Iwagi, Akahone Isl.,
<date value="2004-11-29" valueMax="2004-11-30" valueMin="2004-11-29">29-30.XI.2004</date>
.
<normalizedToken originalValue="Jirô">Jiro</normalizedToken>
Ogawa leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Kami-Ukena dist., Yanadani vill., Yokono (alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">650 m</quantity>
).
<date value="1994-05-07">7.V.1994</date>
. M. Sakai leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Kami-Ukena dist., Kuma-Kougen town, Saragamine.
<date value="2006-06-04">4.VI.2006</date>
.
<normalizedToken originalValue="Yûgo">Yugo</normalizedToken>
<normalizedToken originalValue="Satô">Sato</normalizedToken>
leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Kami-Ukena dist., Kuma-Kougen town, Yurano.
<date value="2007-06-05">5.VI.2007</date>
. Eiji Yamamoto leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ];
<date value="2007-07-17">17.VII.2007</date>
. Eiji Yamamoto leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ];
<date value="2008-07-24" valueMax="2008-07-26" valueMin="2008-07-24">24-26.VII.2008</date>
. Eiji Yamamoto leg.
<specimenCount type="female">1♀</specimenCount>
(MT) [EUMJ]; Kita dist., Uchiko town, Hongawa, Hirose shrine.
<date value="2014-05-21">21.V.2014</date>
. Eiji Yamamoto leg.
<specimenCount type="female">2♀</specimenCount>
[EUMJ]; Kita dist., Uchiko town, Hiraoka.
<date value="2014-05-11" valueMax="2014-05-15" valueMin="2014-05-11">11-15.V.2014</date>
. Eiji Yamamoto leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[EUMJ];
<date value="2014-05-16" valueMax="2014-05-20" valueMin="2014-05-16">16-20.V.2014</date>
. Eiji Yamamoto leg. (MT)
<specimenCount type="female">2♀</specimenCount>
[EUMJ];
<collectingRegion country="Japan" name="Kochi">
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
pref.
</collectingRegion>
, Nankoku city, Estuary of Monobe river.
<date value="2002-07-01">1.VII.2002</date>
. M. Sakai leg.
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Kami city, Kami-anauchi.
<date value="2016-07-30" valueMax="2016-07-31" valueMin="2016-07-30">30-31.VII.2016</date>
. Kazuhiko Konishi leg. (MT)
<specimenCount type="female">1♀</specimenCount>
[EUMJ]; Hata dist., Nishi-Tosa vill., Oku-Yanai, Kuroson,
<date value="1956-04-29">29.IV.1956</date>
, Yozo Murakami leg.,
<specimenCount type="female">2♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Fukuoka">Fukuoka pref.</collectingRegion>
,
<collectingRegion country="Japan" name="Fukuoka">Fukuoka</collectingRegion>
city, Mt. Tachibana-yama,
<date value="1970-05-14">14.V.1970</date>
. Minoru
<collectingRegion country="Japan" name="Miyazaki">Miyazaki</collectingRegion>
leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<date value="1994-05-13">13.V.1994</date>
. Hiroshi Honda leg. (YPT)
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1994-07-02">2.VII.1994</date>
. Hiroshi Honda leg. (YPT)
<specimenCount type="male">5♂</specimenCount>
[ELKU];
<date value="1994-07-16">16.VII.1994</date>
. Hiroshi Honda leg. (YPT)
<specimenCount type="male">2♂</specimenCount>
<specimenCount type="female">4♀</specimenCount>
[ELKU];
<date value="1994-07-23">23.VII.1994</date>
. Hiroshi Honda leg. (YPT)
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1994-07-30">30.VII.1994</date>
. Hiroshi Honda leg. (YPT)
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1994-08-07">7.VIII.1994</date>
. Hiroshi Honda leg. (YPT)
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Fukuoka">Fukuoka</collectingRegion>
city, Mt. Abura-yama,
<date value="1959-06-21">21.VI.1959</date>
. Shoichi Miyamoto leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Fukuoka">Fukuoka</collectingRegion>
city, East ward, Tatara.
<date value="1996-09-08">8.IX.1996</date>
. Yoshimitsu Higashiura leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Fukuoka">Fukuoka</collectingRegion>
city, Sawara ward, Mt. Sefuri-san,
<date value="1992-08-01">1.VIII.1992</date>
, Yoshimitsu Higashiura leg.
<specimenCount type="female">2♀</specimenCount>
[ELKU]; Kasuya dist., Sasaguri town, Mt. Wakasugi-yama.
<date value="1969-05-12">12.V.1969</date>
. Masako Honda leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1970-05-13">13.V.1970</date>
. Masako Honda leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU]; Dazaifu city, Kitadani.
<date value="2016-03-05">5.III.2016</date>
. Yu Hisasue leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU]; Iizuka city, Ae,
<geoCoordinate degrees="33.564" direction="north" orientation="latitude" precision="55" value="33.564">33.564°N</geoCoordinate>
,
<geoCoordinate degrees="130.640" direction="east" orientation="longitude" precision="55" value="130.64">130.640°E</geoCoordinate>
,
<date value="2018-05-05">5.V.2018</date>
. Yu Hisasue leg., (collected from blossoms of
<taxonomicName class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acer palmatum" order="Sapindales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="palmatum">
<emphasis italics="true" pageId="0" pageNumber="99">Acer palmatum</emphasis>
</taxonomicName>
)
<specimenCount type="female">1♀</specimenCount>
[ELKU]; Tagawa dist., Soeda town, Mt. Hiko-san.
<date value="1966-09-28">28.IX.1966</date>
, Masako Honda leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<date value="1967-05-08">8.V.1967</date>
, Masako Honda leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1967-05-10">10.V.1967</date>
, Masako Honda leg.
<specimenCount type="female">2♀</specimenCount>
[ELKU];
<date value="1967-04-20">20.IV.1967</date>
, Kenkichi Kanmiya leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1967-06-20">20.VI.1967</date>
, Kenkichi Kanmiya leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1967-09-20">20.IX.1967</date>
, Kenkichi Kanmiya leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU]; 25.VII.196-,
<normalizedToken originalValue="Kôichi">Koichi</normalizedToken>
Takeno leg. (MT)
<specimenCount type="male">1♂</specimenCount>
[ELKU];
<date value="1970-06-05">5.VI.1970</date>
, Kenkichi Kanmiya leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<date value="1971-04-15">15.IV.1971</date>
, Michitaka Chujo leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1972-07-24">24.VII.1972</date>
, Michitaka Chujo leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<date value="1979-05-22" valueMax="1979-05-26" valueMin="1979-05-22">22-26.V.1979</date>
, Kaoru Maeto leg.
<specimenCount type="female">1♀</specimenCount>
[NIAES];
<date value="1979-07-25" valueMax="1979-07-26" valueMin="1979-07-25">25-26.VII.1979</date>
, Kaoru Maeto leg.
<specimenCount type="male">3♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[NIAES];
<date value="1979-10-24" valueMax="1979-10-26" valueMin="1979-10-24">24-26.X.1979</date>
, Kaoru Maeto leg.
<specimenCount type="female">3♀</specimenCount>
[NIAES]; Miyako dist., Saigawa town, Hobashira, Mt. Hiko-san,
<normalizedToken originalValue="No-tôge">No-toge</normalizedToken>
pass.
<date value="1993-08-29">29.VIII.1993</date>
, N. Wasano leg.
<specimenCount type="male">1♂</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Saga">Saga pref.</collectingRegion>
, Kishima dist., Yamauchi town.
<date value="1991-10-22">22.X.1991</date>
. Yoshimitsu Higashiura leg.
<specimenCount type="female">2♀</specimenCount>
[ELKU]; Higashi-Matsuura dist., Genkai town, Kariya.
<date value="1993-08-23">23.VIII.1993</date>
. Yoshimitsu Higashiura leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Nagasaki">Nagasaki pref.</collectingRegion>
, Minami-Takaki dist., Obama town.
<date value="1960-02-25">25.II.1960</date>
. Terunobu Hidaka leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Kumamoto">Kumamoto pref.</collectingRegion>
, Hondo city, Nishinoku Lake,
<date value="1999-06-05">5.VI.1999</date>
. Victor Fursov leg.
<specimenCount type="female">1♀</specimenCount>
[IZAN];
<collectingRegion country="Japan" name="Miyazaki">Miyazaki pref.</collectingRegion>
, Higashi-Usuki dist., Shiiba vill., Shimo-Fukura, Ohira.
<date value="1993-08-29">29.VIII.1993</date>
, N. Wasano leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Kagoshima">Kagoshima pref.</collectingRegion>
, Aira dist., Kirishima town, Taguchi,
<normalizedToken originalValue="Kirishima-jingû">Kirishima-jingu</normalizedToken>
shrine,
<date value="1969-07-15">15.VII.1969</date>
. Osamu Yata leg.
<specimenCount type="male">1♂</specimenCount>
[ELKU]; Kimotsuki dist., Sata town, Magome, Tajiri, Kape Sata,
<date value="1969-07-17">17.VII.1969</date>
. Osamu Yata leg.
<specimenCount type="female">2♀</specimenCount>
[ELKU]; Kumage Dist., Yaku-shima Isl., Yaku town, Kurio,
<date value="1975-07-08">8.VII.1975</date>
. Kenzo Yamagishi leg.
<specimenCount type="male">2♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Dist.</collectingRegion>
, Toshima vill., Tokara Isles., Kuchi-no-shima Isl.,
<date value="1993-05-02">2.V.1993</date>
, Hiroshi Honda leg.,
<specimenCount type="female">2♀</specimenCount>
[ELKU]; Satoshi Kamitani leg.,
<specimenCount type="female">2♀</specimenCount>
[ELKU];
<normalizedToken originalValue="Ôshima">Oshima</normalizedToken>
dist.,
<normalizedToken originalValue="Amami-ôshima">Amami-oshima</normalizedToken>
Isl., Uken vill., Mt. Yuwan-dake,
<date value="1963-07-30">30.VII.1963</date>
, J. L. Gressitt leg.
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Okinawa">Okinawa pref.</collectingRegion>
,
<collectingRegion country="Japan" name="Okinawa">Okinawa</collectingRegion>
Isl., Nago city, Mt. Nago-dake,
<geoCoordinate degrees="26.587" direction="north" orientation="latitude" precision="55" value="26.587">26.587°N</geoCoordinate>
,
<geoCoordinate degrees="128.000" direction="east" orientation="longitude" precision="55" value="128.0">128.000°E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">140 m</quantity>
,
<date value="2016-07-03" valueMax="2016-07-14" valueMin="2016-07-03">3-14.VII.2016</date>
, Keisuke Narita leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Tokyo">Tokyo</collectingRegion>
Met., Hachijo Isl., Hachijo town, Kamogawa.
<date value="1964-05-27">27.V.1964</date>
, Y. Hirashima and M.
<collectingRegion country="Japan" name="Shiga">Shiga</collectingRegion>
leg.
<specimenCount type="female">3♀</specimenCount>
[ELKU]; Hachijo town, Mitsune-Kantoyama.
<date value="1964-05-30">30.V.1964</date>
, Y. Hirashima and M.
<collectingRegion country="Japan" name="Shiga">Shiga</collectingRegion>
leg.
<specimenCount type="female">1♀</specimenCount>
[ELKU];
<collectingRegion country="Japan" name="Fukuoka">Fukuoka pref.</collectingRegion>
, Munakata dist., Munakata town, Chikuzen Oki-no-shima Isl.
<date value="1958-07-25" valueMax="1958-07-28" valueMin="1958-07-25">25-28.VII.1958</date>
, Yoshihiro Hirashima, Yozo Murakami and Yorio Miyatake leg.
<specimenCount type="male">1♂</specimenCount>
[ELKU].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="distribution">
<paragraph pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="99">
Japan (Hokkaido; Honshu: Fukushima, Ibaraki, Tochigi, Niigata, Ishikawa, Gifu, Shizuoka, Aichi, Osaka, Wakayama, and Shimane; Shikoku: Tokushima, Ehime, and Kochi; Kyushu: Fukuoka, Saga, Nagasaki, Kumamoto, Miyazaki, and Kagoshima; Ryukyus: Yaku-shima Isl., Naka-no-shima Isl.,
<normalizedToken originalValue="Amami-ôhshima">Amami-ohshima</normalizedToken>
Isl., and Okinawa Isl.; Hachijo Isl; and Chikuzen Oki-no-shima Isl.)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="remarks">
<paragraph pageId="0" pageNumber="99">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="99">
Among Japanese species,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
is similar to
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. japonicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="japonicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. japonicum</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. fulvicoxa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fulvicoxa">
<emphasis italics="true" pageId="0" pageNumber="99">G. fulvicoxa</emphasis>
</taxonomicName>
, but
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
differs from these species in the sculpture of the antennal depression (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frons of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure3" httpUri="https://binary.pensoft.net/fig/490336" pageId="0" pageNumber="99">3C</figureCitation>
): transversely costate by strong regular carinae;
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. fulvicoxa" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fulvicoxa">
<emphasis italics="true" pageId="0" pageNumber="99">G. fulvicoxa</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frons of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure3" httpUri="https://binary.pensoft.net/fig/490336" pageId="0" pageNumber="99">3A</figureCitation>
) and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. japonicum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="japonicum">
<emphasis italics="true" pageId="0" pageNumber="99">G. japonicum</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frons of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure3" httpUri="https://binary.pensoft.net/fig/490336" pageId="0" pageNumber="99">3B</figureCitation>
): transversely costate by strong irregular carinae) and the presence of the short sharp horn on the angular point of occipital carina (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Occiput of Japanese Gryon spp. A G. fulvicoxa sp. nov. B G. japonicum C G. yamagishii D G. philippinense E G. shisa sp. nov. F G. pennsylvanicum." figureDoi="10.3897/jhr.80.56178.figure4" httpUri="https://binary.pensoft.net/fig/490337" pageId="0" pageNumber="99">4C</figureCitation>
). Russian Far Eastern species,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. amissum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="amissum">
<emphasis italics="true" pageId="0" pageNumber="99">G. amissum</emphasis>
</taxonomicName>
Kozlov &amp; Kononova, 1990 and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sibiricum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sibiricum">
<emphasis italics="true" pageId="0" pageNumber="99">G. sibiricum</emphasis>
</taxonomicName>
Kononova, 2001 are also similar to
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
in the sculpture of the frons. In
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. amissum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="amissum">
<emphasis italics="true" pageId="0" pageNumber="99">G. amissum</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sibiricum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sibiricum">
<emphasis italics="true" pageId="0" pageNumber="99">G. sibiricum</emphasis>
</taxonomicName>
, however, the angular points of occipital carina are not developed as horns, and the mesopleural carina is weak (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. amissum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="amissum">
<emphasis italics="true" pageId="0" pageNumber="99">G. amissum</emphasis>
</taxonomicName>
) or absent (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sibiricum" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sibiricum">
<emphasis italics="true" pageId="0" pageNumber="99">G. sibiricum</emphasis>
</taxonomicName>
). We examined the holotype of
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. maruzzae" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="maruzzae">
<emphasis italics="true" pageId="0" pageNumber="99">G. maruzzae</emphasis>
</taxonomicName>
at ELKU (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. The holotype of G. maruzzae syn. nov. A dorsal view B lateral view C mesosoma, dorsal view D frons E labels." figureDoi="10.3897/jhr.80.56178.figure11" httpUri="https://binary.pensoft.net/fig/490344" pageId="0" pageNumber="99">11</figureCitation>
) and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sugonjaevi" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="sugonjaevi">
<emphasis italics="true" pageId="0" pageNumber="99">G. sugonjaevi</emphasis>
</taxonomicName>
at ZIN, and they belong to
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. yamagishii" order="Hymenoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="yamagishii">
<emphasis italics="true" pageId="0" pageNumber="99">G. yamagishii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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