164 lines
22 KiB
XML
164 lines
22 KiB
XML
<document id="D5A9103EF30DAD8FE513B24174747469" ID-CLB-Dataset="20365" ID-DOI="10.11646/zootaxa.5104.3.1" ID-GBIF-Dataset="b05ecf1d-1574-4c8a-8df7-12fdd1f7ab22" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332042" ID-ZooBank="F081D236-D592-4829-A3E9-0CA58EF39E43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594429951" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="D31B87CF1E65FFF8CBA9F905EDD0F412" docLanguage="en" docName="zootaxa.5104.3.1.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium umbrinum Shavrin 2022, sp. n." docType="treatment" docVersion="8" lastPageNumber="336" masterDocId="2F22FFB71E47FFDBCB3EFFFDED05F16A" masterDocTitle="A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records" masterLastPageNumber="346" masterPageNumber="301" pageNumber="335" updateTime="1699316848316" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="33F9CA49094BB4A2C9767C224BA856EC">
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<mods:title id="EC9848F4C0245B864BF7C56DC14E41F7">A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records</mods:title>
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<mods:namePart id="7353EF569DDA5B895E39DACAB6A0ED07">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="BCAC2B29EA38385227A5A91AAE035CA7">text</mods:typeOfResource>
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<mods:title id="A4DE13EAE3F12909D3352948A24AD6C7">Zootaxa</mods:title>
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<mods:part id="9380793A20B543905722398587F442EC">
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<mods:date id="5125AA90D329E8429AF7A7DCE2D801B5">2022</mods:date>
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<mods:number id="B936CC2E2730C770E35DC7D2F98542CB">2022-02-28</mods:number>
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<mods:number id="7F41EC13FE3C5CF122B28A02E763F64D">5104</mods:number>
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<mods:identifier id="E27607C663C1BA93DDF3FE611D38EE11" type="DOI">10.11646/zootaxa.5104.3.1</mods:identifier>
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<treatment id="D31B87CF1E65FFF8CBA9F905EDD0F412" ID-DOI="http://doi.org/10.5281/zenodo.6333138" ID-GBIF-Taxon="193878639" ID-Zenodo-Dep="6333138" LSID="urn:lsid:plazi:treatment:D31B87CF1E65FFF8CBA9F905EDD0F412" httpUri="http://treatment.plazi.org/id/D31B87CF1E65FFF8CBA9F905EDD0F412" lastPageId="35" lastPageNumber="336" pageId="34" pageNumber="335">
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<subSubSection id="13A865521E65FFF9CBA9F905ECF7F679" box="[151,498,1784,1811]" pageId="34" pageNumber="335" type="nomenclature">
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<paragraph id="5B0D36D91E65FFF9CBA9F905ECF7F679" blockId="34.[151,498,1784,1847]" box="[151,498,1784,1811]" pageId="34" pageNumber="335">
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<heading id="004581B51E65FFF9CBA9F905ECF7F679" bold="true" box="[151,498,1784,1811]" fontSize="11" level="1" pageId="34" pageNumber="335" reason="1">
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<emphasis id="69C6EACB1E65FFF9CBA9F905ECF7F679" bold="true" box="[151,498,1784,1811]" pageId="34" pageNumber="335">
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<taxonomicName id="9CB24D5A1E65FFF9CBA9F905ECADF679" ID-CoL="9GCV3" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,424,1784,1811]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="umbrinum" status="sp. nov.">
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<emphasis id="69C6EACB1E65FFF9CBA9F905ECADF679" bold="true" box="[151,424,1784,1811]" italics="true" pageId="34" pageNumber="335">Anthobium umbrinum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72F557B01E65FFF9CA8EF904ECF7F679" box="[432,498,1785,1811]" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="13A865521E65FFF9CBA9F8E1EC9BF65D" box="[151,414,1820,1847]" pageId="34" pageNumber="335" type="description">
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<paragraph id="5B0D36D91E65FFF9CBA9F8E1EC9BF65D" blockId="34.[151,498,1784,1847]" box="[151,414,1820,1847]" pageId="34" pageNumber="335">
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(
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<figureCitation id="C3892A5C1E65FFF9CBA1F8E1EDF8F65D" box="[159,253,1820,1847]" captionStart="FIGURE 93" captionStartId="33.[151,250,1706,1730]" captionTargetBox="[151,1437,885,1683]" captionTargetId="figure-299@33.[151,1437,885,1683]" captionTargetPageId="33" captionText="FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6332076" httpUri="https://zenodo.org/record/6332076/files/figure.png" pageId="34" pageNumber="335">Figs. 93</figureCitation>
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,
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<figureCitation id="C3892A5C1E65FFF9CA37F8E1EC22F65C" box="[265,295,1820,1846]" captionStart="FIGURES 94–98" captionStartId="36.[151,264,1931,1955]" captionTargetBox="[171,1408,280,1896]" captionTargetId="figure-71@36.[159,1429,273,1908]" captionTargetPageId="36" captionText="FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332079" httpUri="https://zenodo.org/record/6332079/files/figure.png" pageId="34" pageNumber="335">96</figureCitation>
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,
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<figureCitation id="C3892A5C1E65FFF9CA0DF8E0EC93F65D" box="[307,406,1821,1847]" captionStart="FIGURES 99–104" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[183,1403,190,1910]" captionTargetId="figure-16@37.[174,1413,181,1919]" captionTargetPageId="37" captionText="FIGURES 99–104. Aedeagus of Anthobium tichomirovae (Figs. 99–100; holotype), A. umbrinum (Figs. 101–102; holotype) and A. vitreum (Figs. 103–104; holotype): 99, 101, 103—parameral view, 100, 102, 104—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332081" httpUri="https://zenodo.org/record/6332081/files/figure.png" pageId="34" pageNumber="335">101–102</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="13A865521E65FFF9CBA9F898EFCFF681" pageId="34" pageNumber="335" type="materials_examined">
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<paragraph id="5B0D36D91E65FFF9CBA9F898EFCFF681" blockId="34.[151,1437,1892,2027]" pageId="34" pageNumber="335">
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<emphasis id="69C6EACB1E65FFF9CBA9F898ECC3F614" bold="true" box="[151,454,1892,1919]" pageId="34" pageNumber="335">Type material examined:</emphasis>
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<materialsCitation id="EBDA3C841E65FFF9CAF1F899EFCFF681" ID-GBIF-Occurrence="3697351446" collectingDate="16. - 19. JUN" collectionCode="cSch" latitude="27.1" location="YUNNAN" longLatPrecision="743" longitude="100.13" pageId="34" pageNumber="335" specimenCount="1" specimenCount-male="1" stateProvince="YUNNAN" typeStatus="holotype">
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<typeStatus id="8409887B1E65FFF9CAF1F899EF32F614" box="[463,567,1892,1918]" pageId="34" pageNumber="335" type="holotype">Holotype</typeStatus>
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<specimenCount id="4DB4FD501E65FFF9C97EF898EF50F615" box="[576,597,1893,1919]" pageId="34" pageNumber="335" type="male">♂</specimenCount>
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[dissected; left antennomeres 4–11 are glued on the same card under the specimen; right apical antennomere missing; left paramere without apical portion]: ‘
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<collectingRegion id="9976F83B1E65FFF9CF53F874E9EBF6C9" box="[1133,1262,1929,1955]" country="China" name="Yunnan" pageId="34" pageNumber="335">YUNNAN</collectingRegion>
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,
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<date id="2F0C10191E65FFF9CFC3F875E88EF6C9" box="[1277,1419,1928,1955]" pageId="34" pageNumber="335">
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<collectingDate id="3F48E9F11E65FFF9CFC3F875E88EF6C9" box="[1277,1419,1928,1955]" pageId="34" pageNumber="335">16.-19. JUN</collectingDate>
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</date>
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| YULONG Mts., 1993 |
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<geoCoordinate id="3E86501E1E65FFF9CA9CF850ECF0F6AD" box="[418,501,1965,1991]" degrees="27.10" direction="north" orientation="latitude" pageId="34" pageNumber="335" precision="555" value="27.1">27.10N</geoCoordinate>
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<geoCoordinate id="3E86501E1E65FFF9CAC2F850EF5FF6AD" box="[508,602,1965,1991]" degrees="100.13" direction="east" orientation="longitude" pageId="34" pageNumber="335" precision="555" value="100.13">100.13E</geoCoordinate>
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| Bolm lgt. 3900 m’ <printed>, ‘HOLOTYPE |
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<taxonomicName id="9CB24D5A1E65FFF9CF4FF851E9E9F6AC" authorityName="Leach" authorityYear="1819" box="[1137,1260,1964,1990]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="genus">
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<emphasis id="69C6EACB1E65FFF9CF4FF851E9E9F6AC" box="[1137,1260,1964,1990]" italics="true" pageId="34" pageNumber="335">Anthobium</emphasis>
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</taxonomicName>
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<taxonomicName id="9CB24D5A1E65FFF9CE3EF851E876F6AC" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[1280,1395,1964,1990]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="umbrinum" status="sp. nov.">
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<emphasis id="69C6EACB1E65FFF9CE3EF851E876F6AC" box="[1280,1395,1964,1990]" italics="true" pageId="34" pageNumber="335">umbrinum</emphasis>
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</taxonomicName>
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<emphasis id="69C6EACB1E65FFF9CE45F850EDABF681" bold="true" pageId="34" pageNumber="335">
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<taxonomicNameLabel id="72F557B01E65FFF9CE45F850EDABF681" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2022’ <red, printed> (
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<collectionCode id="3DA3AE1C1E65FFF9C9BFF82DEFB8F681" box="[641,701,2000,2027]" pageId="34" pageNumber="335">cSch</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="13A865521E64FFF8CBF9FF6BEC49F5CA" pageId="35" pageNumber="336" type="description">
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<paragraph id="5B0D36D91E64FFF8CBF9FF6BE9E7F1BE" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
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<emphasis id="69C6EACB1E64FFF8CBF9FF6BEC5CF1DA" bold="true" box="[199,345,150,176]" pageId="35" pageNumber="336">Description.</emphasis>
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Measurements: HL: 0.35; HW: 0.67; AL: 0.96; OL: 0.17; PL: 0.55; PW: 0.97; ESL: 1.30; EW: 1.42; AW: 1.19; MTbL: 0.60; MTrL: 0.39 (MTrL 1–4: 0.24; MTrL 5: 0.15); AedL: 0.52; TL: 3.00.
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</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FF23E99CF356" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
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Habitus as in
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<figureCitation id="C3892A5C1E64FFF8CA60FF23ECB5F192" box="[350,432,222,248]" captionStart="FIGURES 94–98" captionStartId="36.[151,264,1931,1955]" captionTargetBox="[171,1408,280,1896]" captionTargetId="figure-71@36.[159,1429,273,1908]" captionTargetPageId="36" captionText="FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332079" httpUri="https://zenodo.org/record/6332079/files/figure.png" pageId="35" pageNumber="336">Fig. 96</figureCitation>
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. Body and antennomeres 4–11 brown; lateral portions of pronotum and elytra, paratergites and apical part of abdomen yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Head with distinct microsculpture, transverse and moderately dense on clypeus, sparser, more irregular and diagonal in middle and transverse on infraorbital ridges, moderately wide mediobasal portion between ocelli without sculpture; pronotum with distinct transverse microreticulation; scutellum without meshes; abdomen with dense transverse microsculpture. Head with irregular, moderately large and deep punctation, sparser in middle, denser and coarser on infraorbital ridges; middle part of neck with sparse and fine punctation; pronotum with moderately dense punctation, coarser and deeper than that in middle of head, sparser in middle and mediobasal third; punctation of elytra about as that on pronotum, but slightly denser, markedly finer in median portion, on each elytron forming very tangled and vague six longitudinal rows of punctures; abdominal tergites with very fine and sparse punctation.
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</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FDBBE9B7F39A" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">Head slightly less than twice as broad as long; middle portion with indistinct semicircular impression between level of posterior margins of eyes and long and very deep anteocellar fovea, reaching level of apical margin of eye; postocular ridges obtuse. Apical segment of maxillary palpomere twice as long as preceding segment. Antennomere 2 about as broad as basal antennomere, 3 slightly narrower than 2, 4 shorter and slightly broader than 3, 5 longer and broader than 4, 6–9 slightly broader than 5, 10 about as long as broad, slightly broader than 9.</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FD06E9F7F252" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">Pronotum 1.7 times as broad as long, 1.4 times as broad as head, more or less evenly narrowed both anteriad and posteriad, with widely rounded anterior angles not protruded anteriad, without median impressions.</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FCBFEF6FF2EA" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">Elytra slightly broader than long, more than twice as long as pronotum, reaching apical margin of abdominal tergite IV, with rounded posterior margin.</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FC76EF99F2CE" blockId="35.[151,1437,150,1400]" box="[199,668,906,933]" pageId="35" pageNumber="336">Metatarsus 1.5 times as long as metatibia.</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FC53E846F55E" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
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Male. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus very wide, suboval, with wide basal bulb, gradually narrowed toward widely rounded apex; parameres moderately short, narrowed apically, exceeding apex of median lobe, with short apical and two preapical setae; internal sac very long, spirally folded in basal portion (
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<figureCitation id="C3892A5C1E64FFF8C9C0FBE7EE5AF55E" box="[766,863,1050,1076]" captionStart="FIGURES 99–104" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[183,1403,190,1910]" captionTargetId="figure-16@37.[174,1413,181,1919]" captionTargetPageId="37" captionText="FIGURES 99–104. Aedeagus of Anthobium tichomirovae (Figs. 99–100; holotype), A. umbrinum (Figs. 101–102; holotype) and A. vitreum (Figs. 103–104; holotype): 99, 101, 103—parameral view, 100, 102, 104—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332081" httpUri="https://zenodo.org/record/6332081/files/figure.png" pageId="35" pageNumber="336">Fig. 101</figureCitation>
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). Lateral aspect of aedeagus as in
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<figureCitation id="C3892A5C1E64FFF8CFE0FBE7E83BF55E" box="[1246,1342,1050,1076]" captionStart="FIGURES 99–104" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[183,1403,190,1910]" captionTargetId="figure-16@37.[174,1413,181,1919]" captionTargetPageId="37" captionText="FIGURES 99–104. Aedeagus of Anthobium tichomirovae (Figs. 99–100; holotype), A. umbrinum (Figs. 101–102; holotype) and A. vitreum (Figs. 103–104; holotype): 99, 101, 103—parameral view, 100, 102, 104—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332081" httpUri="https://zenodo.org/record/6332081/files/figure.png" pageId="35" pageNumber="336">Fig. 102</figureCitation>
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.
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</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FBC3EC94F532" blockId="35.[151,1437,150,1400]" box="[199,401,1086,1112]" pageId="35" pageNumber="336">Female unknown.</paragraph>
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<paragraph id="5B0D36D91E64FFF8CBF9FB9FEC49F5CA" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
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<emphasis id="69C6EACB1E64FFF8CBF9FB9FECB5F516" bold="true" box="[199,432,1122,1148]" pageId="35" pageNumber="336">Comparative notes.</emphasis>
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<taxonomicName id="9CB24D5A1E64FFF8CA88FB9FEFAFF516" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[438,682,1122,1148]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="umbrinum" status="sp. nov.">
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<emphasis id="69C6EACB1E64FFF8CA88FB9FEFAFF516" box="[438,682,1122,1148]" italics="true" pageId="35" pageNumber="336">Anthobium umbrinum</emphasis>
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</taxonomicName>
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<emphasis id="69C6EACB1E64FFF8C98FFB9FEFF5F516" bold="true" box="[689,752,1122,1148]" pageId="35" pageNumber="336">
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<taxonomicNameLabel id="72F557B01E64FFF8C98FFB9FEFF5F516" box="[689,752,1122,1148]" pageId="35" pageNumber="336" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
|
||
differs from the other species of the
|
||
<taxonomicName id="9CB24D5A1E64FFF8CFB9FB9EE9D7F516" authorityName=": Curtis" authorityYear="1829" box="[1159,1234,1123,1148]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="69C6EACB1E64FFF8CFB9FB9EE9D7F516" box="[1159,1234,1123,1148]" italics="true" pageId="35" pageNumber="336">tectum</emphasis>
|
||
</taxonomicName>
|
||
group by the very broad aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A865521E64FFF8CBF9FB57EDEEF45A" pageId="35" pageNumber="336" type="distribution">
|
||
<paragraph id="5B0D36D91E64FFF8CBF9FB57EC05F582" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
|
||
<emphasis id="69C6EACB1E64FFF8CBF9FB57EC64F5AE" bold="true" box="[199,353,1194,1220]" pageId="35" pageNumber="336">Distribution.</emphasis>
|
||
The species is known only from the
|
||
<typeStatus id="8409887B1E64FFF8C830FB56EE3BF5AF" box="[782,830,1195,1221]" pageId="35" pageNumber="336">type</typeStatus>
|
||
locality in Yulong mountain range in
|
||
<collectingRegion id="9976F83B1E64FFF8CFCAFB56E848F5AF" box="[1268,1357,1195,1221]" country="China" name="Yunnan" pageId="35" pageNumber="336">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCountry id="23A576491E64FFF8CE67FB57E899F5AE" box="[1369,1436,1194,1220]" name="China" pageId="35" pageNumber="336">China</collectingCountry>
|
||
(
|
||
<figureCitation id="C3892A5C1E64FFF8CB9EFB33EDF1F582" box="[160,244,1230,1256]" captionStart="FIGURE 93" captionStartId="33.[151,250,1706,1730]" captionTargetBox="[151,1437,885,1683]" captionTargetId="figure-299@33.[151,1437,885,1683]" captionTargetPageId="33" captionText="FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6332076" httpUri="https://zenodo.org/record/6332076/files/figure.png" pageId="35" pageNumber="336">Fig. 93</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5B0D36D91E64FFF8CBF9FB0FEDEEF45A" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
|
||
<emphasis id="69C6EACB1E64FFF8CBF9FB0FEC83F466" bold="true" box="[199,390,1266,1292]" pageId="35" pageNumber="336">Natural history.</emphasis>
|
||
The specimen was collected at an elevation of
|
||
<quantity id="9C4A9B3C1E64FFF8C8A3FB0FEEF7F467" box="[925,1010,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="35" pageNumber="336" unit="m" value="3900.0">3900 m</quantity>
|
||
a.s.l. Detailed ecological data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A865521E64FFF8CBF9FAC7EDD0F412" pageId="35" pageNumber="336" type="etymology">
|
||
<paragraph id="5B0D36D91E64FFF8CBF9FAC7EDD0F412" blockId="35.[151,1437,150,1400]" pageId="35" pageNumber="336">
|
||
<emphasis id="69C6EACB1E64FFF8CBF9FAC7EC4BF43E" bold="true" box="[199,334,1338,1364]" pageId="35" pageNumber="336">Etymology.</emphasis>
|
||
The specific epithet (Latin adjective: brown, like earth of
|
||
<collectingRegion id="9976F83B1E64FFF8C8D7FAC7E945F43F" box="[1001,1088,1338,1365]" country="Italy" name="Umbria" pageId="35" pageNumber="336">Umbria</collectingRegion>
|
||
) refers to the coloration of the body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |