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<document id="C3A0DBBAEA0B6EF546A51C943400667C" ID-CLB-Dataset="20365" ID-DOI="10.11646/zootaxa.5104.3.1" ID-GBIF-Dataset="b05ecf1d-1574-4c8a-8df7-12fdd1f7ab22" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332042" ID-ZooBank="F081D236-D592-4829-A3E9-0CA58EF39E43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594429951" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="D31B87CF1E4EFFD6CBA9FA13EC95F583" docLanguage="en" docName="zootaxa.5104.3.1.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium morosum Shavrin 2022, sp. n." docType="treatment" docVersion="8" lastPageNumber="314" masterDocId="2F22FFB71E47FFDBCB3EFFFDED05F16A" masterDocTitle="A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records" masterLastPageNumber="346" masterPageNumber="301" pageNumber="310" updateTime="1699316848316" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="70306C7705CEDD805C20170FDF9B3690" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="01392DCB3090A28106A544AC2520F4F7">
<mods:title id="C9233A5B20C5830770BB418202D923A0">A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records</mods:title>
</mods:titleInfo>
<mods:name id="0B9A1DA0844AD3317D2F2DCE0B46E7DB" type="personal">
<mods:role id="F7FFDD0792D5DC7417031F35CAFA9DFF">
<mods:roleTerm id="4F7215457AB7CC7ECF27EE8BCC0D4C01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4561DBC963AED1260515F5889E323409">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="47EB7550ABE1690C99BB853C76991F40">text</mods:typeOfResource>
<mods:relatedItem id="03973C09B3657BE5F64E7B266F0CF876" type="host">
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<mods:title id="0A3D4C83D95ED2248323EA239574DC41">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="ED12E72738EB597B30E88CFD0B19EC0B">
<mods:date id="A30BF4D752FCD9B1D85807A42FBA8A3C">2022</mods:date>
<mods:detail id="9DA64878D6888430A96C7BB45D0E237D" type="pubDate">
<mods:number id="78BF466CFB10F1DEEE9DAE2314A19960">2022-02-28</mods:number>
</mods:detail>
<mods:detail id="DCBE523D4F7E435959DC836E9F202FAC" type="volume">
<mods:number id="592B4BA34313F51E2390A73C9F6B2ACD">5104</mods:number>
</mods:detail>
<mods:detail id="4B61D07516F2E7D4496394B85F31382E" type="issue">
<mods:number id="DB6D526A671F38105520E21CDEF1487C">3</mods:number>
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<mods:start id="3B442F11D5ADAB05E05B5376F0B699A8">301</mods:start>
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<mods:classification id="FA8D3484441AA2F924940C1B910541EE">journal article</mods:classification>
<mods:identifier id="56734D3D0471BEC2F117A7EFC6ECC9A1" type="CLB-Dataset">20365</mods:identifier>
<mods:identifier id="19E3DECF28E90EFB6BFA531629110018" type="DOI">10.11646/zootaxa.5104.3.1</mods:identifier>
<mods:identifier id="61C0A611323F0F7BFE5E81D9CE30AD46" type="GBIF-Dataset">b05ecf1d-1574-4c8a-8df7-12fdd1f7ab22</mods:identifier>
<mods:identifier id="F9636CFA94D41F0D5A8BD067E417E1A3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="906E4D1F30CCDCB9BBF73CBC937F87ED" type="Zenodo-Dep">6332042</mods:identifier>
<mods:identifier id="3296E735A395A17A446A155094861FFB" type="ZooBank">F081D236-D592-4829-A3E9-0CA58EF39E43</mods:identifier>
</mods:mods>
<treatment id="D31B87CF1E4EFFD6CBA9FA13EC95F583" ID-DOI="http://doi.org/10.5281/zenodo.6335816" ID-GBIF-Taxon="193878647" ID-Zenodo-Dep="6335816" LSID="urn:lsid:plazi:treatment:D31B87CF1E4EFFD6CBA9FA13EC95F583" httpUri="http://treatment.plazi.org/id/D31B87CF1E4EFFD6CBA9FA13EC95F583" lastPageId="13" lastPageNumber="314" pageId="9" pageNumber="310">
<subSubSection id="13A865521E4EFFD2CBA9FA13ECE6F763" box="[151,483,1518,1545]" pageId="9" pageNumber="310" type="nomenclature">
<paragraph id="5B0D36D91E4EFFD2CBA9FA13ECE6F763" blockId="9.[151,483,1518,1581]" box="[151,483,1518,1545]" pageId="9" pageNumber="310">
<heading id="004581B51E4EFFD2CBA9FA13ECE6F763" bold="true" box="[151,483,1518,1545]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
<emphasis id="69C6EACB1E4EFFD2CBA9FA13ECE6F763" bold="true" box="[151,483,1518,1545]" pageId="9" pageNumber="310">
<taxonomicName id="9CB24D5A1E4EFFD2CBA9FA13EC9CF763" ID-CoL="9GCTS" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,409,1518,1545]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="morosum" status="sp. nov.">
<emphasis id="69C6EACB1E4EFFD2CBA9FA13EC9CF763" bold="true" box="[151,409,1518,1545]" italics="true" pageId="9" pageNumber="310">Anthobium morosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72F557B01E4EFFD2CA9FFA12ECE6F763" box="[417,483,1519,1545]" pageId="9" pageNumber="310" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13A865521E4EFFD2CBA9F9EEECA9F747" box="[151,428,1554,1581]" pageId="9" pageNumber="310" type="description">
<paragraph id="5B0D36D91E4EFFD2CBA9F9EEECA9F747" blockId="9.[151,483,1518,1581]" box="[151,428,1554,1581]" pageId="9" pageNumber="310">
(
<figureCitation id="C3892A5C1E4EFFD2CBA1F9EEEC22F747" box="[159,295,1555,1581]" captionStart="FIGURES 26–28" captionStartId="10.[151,264,1582,1606]" captionTargetBox="[161,1425,578,1546]" captionTargetId="figure-193@10.[151,1436,561,1558]" captionTargetPageId="10" captionText="FIGURES 26–28. Habitus (Fig. 26; paratype) and aedeagus (Figs. 27—28) of Anthobium morosum: 27—parameral view, 28—lateral view. Scale bar: 1.0 mm (Fig. 26), 0.1 mm (Figs. 27–28)." figureDoi="http://doi.org/10.5281/zenodo.6332054" httpUri="https://zenodo.org/record/6332054/files/figure.png" pageId="9" pageNumber="310">Figs. 26–28</figureCitation>
,
<figureCitation id="C3892A5C1E4EFFD2CA0DF9EFEC79F747" box="[307,380,1554,1581]" captionStart-0="FIGURES 29–33" captionStart-1="FIGURES 34–45" captionStartId-0="11.[151,264,1929,1953]" captionStartId-1="12.[151,264,1648,1672]" captionTargetBox-0="[166,1424,650,1899]" captionTargetBox-1="[163,1428,187,1615]" captionTargetId-0="figure-232@11.[151,1436,633,1905]" captionTargetId-1="figure-21@12.[151,1436,181,1625]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 29–33. Mouthparts (dorsal view) of Anthobium morosum: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm." captionText-1="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6332056" figureDoi-1="http://doi.org/10.5281/zenodo.6332058" httpUri-0="https://zenodo.org/record/6332056/files/figure.png" httpUri-1="https://zenodo.org/record/6332058/files/figure.png" pageId="9" pageNumber="310">29–45</figureCitation>
,
<figureCitation id="C3892A5C1E4EFFD2CAB9F9EEECA1F747" box="[391,420,1555,1581]" captionStart="FIGURE 93" captionStartId="33.[151,250,1706,1730]" captionTargetBox="[151,1437,885,1683]" captionTargetId="figure-299@33.[151,1437,885,1683]" captionTargetPageId="33" captionText="FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6332076" httpUri="https://zenodo.org/record/6332076/files/figure.png" pageId="9" pageNumber="310">93</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13A865521E4EFFD2CBA9F9A6E812F66F" pageId="9" pageNumber="310" type="materials_examined">
<paragraph id="5B0D36D91E4EFFD2CBA9F9A6ECDFF7D7" blockId="9.[151,1437,1626,2013]" pageId="9" pageNumber="310">
<materialsCitation id="EBDA3C841E4EFFD2CBA9F9A6ECDFF7D7" ID-GBIF-Occurrence="3697351307" collectingDate="16. IV." collectionCode="CNN" country="Nepal" elevation="2800" location="Lobl" pageId="9" pageNumber="310" specimenCount="1" specimenCount-male="1" stateProvince="Bagmati" typeStatus="holotype">
<emphasis id="69C6EACB1E4EFFD2CBA9F9A6ECC2F71E" bold="true" box="[151,455,1626,1653]" pageId="9" pageNumber="310">Type material examined:</emphasis>
<typeStatus id="8409887B1E4EFFD2CAEFF9A6EF3CF71F" box="[465,569,1627,1653]" pageId="9" pageNumber="310" type="holotype">Holotype</typeStatus>
<specimenCount id="4DB4FD501E4EFFD2C97DF9A6EF5DF71F" box="[579,600,1627,1653]" pageId="9" pageNumber="310" type="male">♂</specimenCount>
[dissected]: ‘
<collectingCountry id="23A576491E4EFFD2C9CEF9A6EE49F71F" box="[752,844,1627,1653]" name="Nepal" pageId="9" pageNumber="310">NEPAL</collectingCountry>
(Prov.
<collectingRegion id="9976F83B1E4EFFD2C89DF9A6E901F71C" box="[931,1028,1627,1654]" country="Nepal" name="Bagmati" pageId="9" pageNumber="310">Bagmati</collectingRegion>
) | Malemchi [Melamchi],
<quantity id="9C4A9B3C1E4EFFD2CE0BF9A6E888F71F" box="[1333,1421,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="310" unit="m" value="2800.0">
<elevation id="D09FD1EA1E4EFFD2CE0BF9A6E888F71F" box="[1333,1421,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="310" unit="m" value="2800.0">2800 m</elevation>
</quantity>
|
<date id="2F0C10191E4EFFD2CBA9F983EDDBF7F3" box="[151,222,1662,1689]" pageId="9" pageNumber="310">
<collectingDate id="3F48E9F11E4EFFD2CBA9F983EDDBF7F3" box="[151,222,1662,1689]" pageId="9" pageNumber="310">16.IV.</collectingDate>
</date>
[19]81 | [I.]
<location id="5E6D60021E4EFFD2CA5DF982EC9DF7F3" LSID="urn:lsid:plazi:treatment:D31B87CF1E4EFFD6CBA9FA13EC95F583:5E6D60021E4EFFD2CA5DF982EC9DF7F3" box="[355,408,1663,1689]" country="Nepal" name="Lobl" pageId="9" pageNumber="310" stateProvince="Bagmati">Löbl</location>
&amp; [A.] Smetana’ &lt;printed&gt;, ‘HOLOTYPE |
<taxonomicName id="9CB24D5A1E4EFFD2C8B5F982E903F7F3" authorityName="Leach" authorityYear="1819" box="[907,1030,1663,1689]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="69C6EACB1E4EFFD2C8B5F982E903F7F3" box="[907,1030,1663,1689]" italics="true" pageId="9" pageNumber="310">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="9CB24D5A1E4EFFD2CF24F982E984F7F2" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[1050,1153,1663,1688]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="morosum" status="sp. nov.">
<emphasis id="69C6EACB1E4EFFD2CF24F982E984F7F2" box="[1050,1153,1663,1688]" italics="true" pageId="9" pageNumber="310">morosum</emphasis>
</taxonomicName>
<emphasis id="69C6EACB1E4EFFD2CFB6F982E9C2F7F3" bold="true" box="[1160,1223,1663,1689]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="72F557B01E4EFFD2CFB6F982E9C2F7F3" box="[1160,1223,1663,1689]" pageId="9" pageNumber="310" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2022’ &lt;red, printed&gt; (
<collectionCode id="3DA3AE1C1E4EFFD2CAB3F95EECCBF7D7" box="[397,462,1699,1725]" collectionName="CNN" pageId="9" pageNumber="310">CNN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5B0D36D91E4EFFD2CBF9F93AE812F66F" blockId="9.[151,1437,1626,2013]" pageId="9" pageNumber="310">
<materialsCitation id="EBDA3C841E4EFFD2CBF9F93AE812F66F" ID-GBIF-Occurrence="3697351384" collectionCode="CNN" location="Anthobium" municipality="All" pageId="9" pageNumber="310" specimenCount="6" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="8409887B1E4EFFD2CBF9F93AEC31F78B" box="[199,308,1735,1761]" pageId="9" pageNumber="310" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="4DB4FD501E4EFFD2CA7DF93AECD6F788" box="[323,467,1735,1762]" pageId="9" pageNumber="310" type="generic">2 specimens</specimenCount>
):
<specimenCount id="4DB4FD501E4EFFD2CADBF93AEF26F78B" box="[485,547,1735,1761]" pageId="9" pageNumber="310" type="female">2 ♀♀</specimenCount>
[
<specimenCount id="4DB4FD501E4EFFD2C90EF93AEFC9F78B" box="[560,716,1735,1761]" pageId="9" pageNumber="310" type="generic">one specimen</specimenCount>
dissected]: same data as the
<typeStatus id="8409887B1E4EFFD2CF33F93AE96AF78B" box="[1037,1135,1735,1761]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
(cSh,
<collectionCode id="3DA3AE1C1E4EFFD2CF88F93AE9F3F78B" box="[1206,1270,1735,1761]" collectionName="CNN" pageId="9" pageNumber="310">CNN</collectionCode>
).
<collectingMunicipality id="BB69ACA31E4EFFD2CE39F93AE82EF78B" box="[1287,1323,1735,1761]" pageId="9" pageNumber="310">All</collectingMunicipality>
<typeStatus id="8409887B1E4EFFD2CE0FF93AE899F78B" box="[1329,1436,1735,1761]" pageId="9" pageNumber="310" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="8409887B1E4EFFD2C92FF916EFA4F66F" box="[529,673,1771,1797]" pageId="9" pageNumber="310" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="9CB24D5A1E4EFFD2C98BF916EE35F66F" authorityName="Leach" authorityYear="1819" box="[693,816,1771,1797]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="69C6EACB1E4EFFD2C98BF916EE35F66F" box="[693,816,1771,1797]" italics="true" pageId="9" pageNumber="310">
<location id="5E6D60021E4EFFD2C98BF916EE35F66F" LSID="urn:lsid:plazi:treatment:D31B87CF1E4EFFD6CBA9FA13EC95F583:5E6D60021E4EFFD2C98BF916EE35F66F" box="[693,816,1771,1797]" municipality="All" name="Anthobium" pageId="9" pageNumber="310">Anthobium</location>
</emphasis>
</taxonomicName>
|
<taxonomicName id="9CB24D5A1E4EFFD2C87AF916EEAEF66E" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[836,939,1771,1796]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="morosum" status="sp. nov.">
<emphasis id="69C6EACB1E4EFFD2C87AF916EEAEF66E" box="[836,939,1771,1796]" italics="true" pageId="9" pageNumber="310">morosum</emphasis>
</taxonomicName>
<emphasis id="69C6EACB1E4EFFD2C88CF916EEF4F66F" bold="true" box="[946,1009,1771,1797]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="72F557B01E4EFFD2C88CF916EEF4F66F" box="[946,1009,1771,1797]" pageId="9" pageNumber="310" rank="species">
<specimenCount id="4DB4FD501E4EFFD2C88CF916EED6F66F" box="[946,979,1771,1797]" pageId="9" pageNumber="310" type="generic">sp.</specimenCount>
n.
</taxonomicNameLabel>
</emphasis>
|
<location id="5E6D60021E4EFFD2CF3AF916E991F66F" LSID="urn:lsid:plazi:treatment:D31B87CF1E4EFFD6CBA9FA13EC95F583:5E6D60021E4EFFD2CF3AF916E991F66F" box="[1028,1172,1771,1797]" municipality="All" name="Shavrin A. V." pageId="9" pageNumber="310">Shavrin A.V.</location>
des. 2022’.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="13A865521E4EFFD6CBF9F8F3E81BF55F" lastPageId="13" lastPageNumber="314" pageId="9" pageNumber="310" type="description">
<paragraph id="5B0D36D91E4EFFD2CBF9F8F3EEBEF61B" blockId="9.[151,1437,1626,2013]" pageId="9" pageNumber="310">
<emphasis id="69C6EACB1E4EFFD2CBF9F8F3EC5CF642" bold="true" box="[199,345,1806,1832]" pageId="9" pageNumber="310">Description.</emphasis>
Measurements (n=3): HL: 0.66–0.70; HW: 1.00–1.05; AL (
<typeStatus id="8409887B1E4EFFD2CF3FF8F2E96FF640" box="[1025,1130,1807,1834]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
): 2.20; OL: 0.27–0.30; PL: 0.90–0.95; PW: 1.50–1.60; ESL: 2.20–2.30; EW: 2.01–2.09; AW: 1.65–1.85; MTbL (
<typeStatus id="8409887B1E4EFFD2CF62F8CEE9C0F624" box="[1116,1221,1843,1870]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
): 1.45, MTrL: 0.60 (MTrL 1–4: 0.37; MTrL 5: 0.23); AedL: 1.46; TL: 5.65(
<typeStatus id="8409887B1E4EFFD2C836F8ABEE6BF61B" box="[776,878,1878,1905]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
)–5.80.
</paragraph>
<paragraph id="5B0D36D91E4EFFD1CBF9F886EF7DF373" blockId="9.[151,1437,1626,2013]" lastBlockId="10.[151,1437,151,537]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
Body moderately large, elongate, convex (
<figureCitation id="C3892A5C1E4EFFD2C9A3F886EFF5F6FC" box="[669,752,1915,1942]" captionStart="FIGURES 26–28" captionStartId="10.[151,264,1582,1606]" captionTargetBox="[161,1425,578,1546]" captionTargetId="figure-193@10.[151,1436,561,1558]" captionTargetPageId="10" captionText="FIGURES 26–28. Habitus (Fig. 26; paratype) and aedeagus (Figs. 27—28) of Anthobium morosum: 27—parameral view, 28—lateral view. Scale bar: 1.0 mm (Fig. 26), 0.1 mm (Figs. 27–28)." figureDoi="http://doi.org/10.5281/zenodo.6332054" httpUri="https://zenodo.org/record/6332054/files/figure.png" pageId="9" pageNumber="310">Fig. 26</figureCitation>
). Body brown; antennomeres 3–11, lateral and apical margins of scutellum, lateral margins of pronotum and legs yellow-brown; mouthparts, antennomeres 1–2 and apical tarsomeres yellow. Forebody glossy, with rudiments of indistinct striate microsculpture in middle of head (
<specimenCount id="4DB4FD501E4EFFD2CFC5F83EE894F6B7" box="[1275,1425,1987,2013]" pageId="9" pageNumber="310" type="generic" typeStatus="paratype">one paratype</specimenCount>
); posterio-lateral parts of head and portion between antennal insertion and anterior margin of eye, and abdominal tergites with distinct isodiametric microsculpture. Head and neck with sparse, irregular and moderately deep punctation, sparser between anteocellar foveae and margins of eye, with impunctate longitudinal portions of inner margins of median impression; middle of pronotum with irregular punctation, slightly denser than that on head, distinctly sparser in latero-apical portions, with narrow impunctate portions behind lateral pits; scutellum with several moderately large and deep punctures; punctation of elytra denser and coarser than that on pronotum, each elytron with tangled and vague seven to eight longitudinal rows of serial punctures, apical portion of each elytron finer and sparser; abdominal tergites IV–VI without visible punctures, apical part of abdominal tergite VII and tergite VIII with fine and irregular punctation. Apical and middle parts of head, middle of pronotum and elytra, paratergites and abdominal tergites with very fine and sparse, semierect pubescence; abdominal tergites with additional very short and sparse setae, denser on apical margins.
</paragraph>
<caption id="0FCD66511E4DFFD1CBA9F9D3EE56F700" ID-DOI="http://doi.org/10.5281/zenodo.6332054" ID-Zenodo-Dep="6332054" httpUri="https://zenodo.org/record/6332054/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,264,1582,1606]" targetBox="[161,1425,578,1546]" targetPageId="10">
<paragraph id="5B0D36D91E4DFFD1CBA9F9D3EE56F700" blockId="10.[151,1436,1582,1642]" pageId="10" pageNumber="311">
<emphasis id="69C6EACB1E4DFFD1CBA9F9D3EC5DF72C" bold="true" box="[151,344,1582,1606]" pageId="10" pageNumber="311">FIGURES 26–28.</emphasis>
Habitus (Fig. 26; paratype) and aedeagus (Figs. 27—28) of
<taxonomicName id="9CB24D5A1E4DFFD1C8E0F9D3E9B6F72C" authorityName="Shavrin" authorityYear="2022" box="[990,1203,1582,1606]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="morosum">
<emphasis id="69C6EACB1E4DFFD1C8E0F9D3E9B6F72C" box="[990,1203,1582,1606]" italics="true" pageId="10" pageNumber="311">Anthobium morosum</emphasis>
</taxonomicName>
: 27—parameral view, 28—lateral view. Scale bar: 1.0 mm (Fig. 26), 0.1 mm (Figs. 27–28).
</paragraph>
</caption>
<paragraph id="5B0D36D91E4DFFD0CBF9F965EC79F30B" blockId="10.[151,1437,1688,2039]" lastBlockId="11.[151,1437,151,609]" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311">
Head 1.5 times as broad as long, with slightly convex middle portion; middle portion with wide and moderately deep semicircular impression; anteocellar foveae narrow, diagonally directed toward about level of anterior margins of eye or slightly shorter; postocular ridges distinct, smooth; anterior portion between antennal insertion and anterior margin of eye widely concave. Ocelli large, situated at level of postocular ridge, distance between ocelli distinctly longer than distance between ocellus and posterior margin of eye. Eyes large, convex. Gular sutures with rounded apical parts, widely divergent posteriad; distance between sutures moderately narrow, shortest distance located at level about posterior margins of eyes (
<figureCitation id="C3892A5C1E4DFFD1C96DF88DEFACF6E1" box="[595,681,1904,1931]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="10" pageNumber="311">Fig. 34</figureCitation>
). Labrum transverse, with widely rounded and strongly protruded apicad membranous latero-apical portions, with wide and very deep apical emargination, and long laterobasal projections (internally, under the clypeus) (
<figureCitation id="C3892A5C1E4DFFD1C96CF845EFA3F6B9" box="[594,678,1976,2003]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[166,1424,650,1899]" captionTargetId="figure-232@11.[151,1436,633,1905]" captionTargetPageId="11" captionText="FIGURES 29–33. Mouthparts (dorsal view) of Anthobium morosum: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332056" httpUri="https://zenodo.org/record/6332056/files/figure.png" pageId="10" pageNumber="311">Fig. 29</figureCitation>
). Mandibles long, distinctly asymmetric, with narrow basal part and elongate, moderately narrow and strongly curved apical portions (larger and more curved in lefte mandible), with large subtriangular tooth below middle of right mandible, with apex slightly protruded apicad; molar lobes moderately small and narrow (
<figureCitation id="C3892A5C1E4CFFD0CA9BFF46ECFFF1BF" box="[421,506,187,213]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[166,1424,650,1899]" captionTargetId="figure-232@11.[151,1436,633,1905]" captionTargetPageId="11" captionText="FIGURES 29–33. Mouthparts (dorsal view) of Anthobium morosum: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332056" httpUri="https://zenodo.org/record/6332056/files/figure.png" pageId="11" pageNumber="312">Fig. 30</figureCitation>
). Labium moderately narrow, with apical labial palpomere about four times as long as preceding segment (
<figureCitation id="C3892A5C1E4CFFD0CAABFF22ECECF193" box="[405,489,223,249]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[166,1424,650,1899]" captionTargetId="figure-232@11.[151,1436,633,1905]" captionTargetPageId="11" captionText="FIGURES 29–33. Mouthparts (dorsal view) of Anthobium morosum: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332056" httpUri="https://zenodo.org/record/6332056/files/figure.png" pageId="11" pageNumber="312">Fig. 32</figureCitation>
). Mentum narrow, strongly transverse, with two pairs of long latero-apical setae (
<figureCitation id="C3892A5C1E4CFFD0CE51FF22EDB3F077" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[166,1424,650,1899]" captionTargetId="figure-232@11.[151,1436,633,1905]" captionTargetPageId="11" captionText="FIGURES 29–33. Mouthparts (dorsal view) of Anthobium morosum: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332056" httpUri="https://zenodo.org/record/6332056/files/figure.png" pageId="11" pageNumber="312">Fig. 33</figureCitation>
). Maxilla as in
<figureCitation id="C3892A5C1E4CFFD0CA52FEFEECC5F077" box="[364,448,259,285]" captionStart="FIGURES 29–33" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[166,1424,650,1899]" captionTargetId="figure-232@11.[151,1436,633,1905]" captionTargetPageId="11" captionText="FIGURES 29–33. Mouthparts (dorsal view) of Anthobium morosum: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332056" httpUri="https://zenodo.org/record/6332056/files/figure.png" pageId="11" pageNumber="312">Fig. 31</figureCitation>
; maxillary palpomere long, with elongate maxillary palpomere 2, slightly shorter than apical palpomere, and apical palpomere very long, slightly more than three times as long as preapical segment, narrow in basal portion, gradually broadened toward apical third, with rounded apex; galea moderately wide, distinctly longer than lacinia; lacinia with narrow and curved apical portion, with unarticulated spine-like apex and numerous spines and setae along medial margin. Antennae moderately long, with elongate antennomeres, reaching basal third of elytra when reclined; basal antennomere long, about three times as broad as long, antennomere 2 slightly narrower and 1.3 times as long as basal antennomere, 3 distinctly longer and slightly narrower than 2, 4–6 distinctly shorter than 3, 7 slightly longer and broader than 6, 8–9 slightly shorter than 7, 10 slightly shorter and broader than 7, apical antennomere broader and 1.4 times as long as preceding segment, from apical third sharply narrowed toward subacute apex.
</paragraph>
<caption id="0FCD66511E4CFFD0CBA9F874EFABF6AE" ID-DOI="http://doi.org/10.5281/zenodo.6332056" ID-Zenodo-Dep="6332056" httpUri="https://zenodo.org/record/6332056/files/figure.png" pageId="11" pageNumber="312" startId="11.[151,264,1929,1953]" targetBox="[166,1424,650,1899]" targetPageId="11">
<paragraph id="5B0D36D91E4CFFD0CBA9F874EFABF6AE" blockId="11.[151,1436,1929,1989]" pageId="11" pageNumber="312">
<emphasis id="69C6EACB1E4CFFD0CBA9F874EC51F6CB" bold="true" box="[151,340,1929,1953]" pageId="11" pageNumber="312">FIGURES 29–33.</emphasis>
Mouthparts (dorsal view) of
<taxonomicName id="9CB24D5A1E4CFFD0C947F874EE4EF6CB" authorityName="Shavrin" authorityYear="2022" box="[633,843,1929,1953]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="morosum">
<emphasis id="69C6EACB1E4CFFD0C947F874EE4EF6CB" box="[633,843,1929,1953]" italics="true" pageId="11" pageNumber="312">Anthobium morosum</emphasis>
</taxonomicName>
: 29—labrum, 30—mandibles (left and right), 31—maxilla (left), 32—labium, 33—mentum. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="0FCD66511E4BFFD7CBA9F98DE931F672" ID-DOI="http://doi.org/10.5281/zenodo.6332058" ID-Zenodo-Dep="6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="12" pageNumber="313" startId="12.[151,264,1648,1672]" targetBox="[163,1428,187,1615]" targetPageId="12">
<paragraph id="5B0D36D91E4BFFD7CBA9F98DE931F672" blockId="12.[151,1436,1648,1817]" pageId="12" pageNumber="313">
<emphasis id="69C6EACB1E4BFFD7CBA9F98DEC53F7E3" bold="true" box="[151,342,1648,1673]" pageId="12" pageNumber="313">FIGURES 34–45.</emphasis>
<taxonomicName id="9CB24D5A1E4BFFD7CA60F98CEF37F7E2" authorityName="Shavrin" authorityYear="2022" box="[350,562,1649,1673]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="morosum">
<emphasis id="69C6EACB1E4BFFD7CA60F98CEF37F7E2" box="[350,562,1649,1673]" italics="true" pageId="12" pageNumber="313">Anthobium morosum</emphasis>
</taxonomicName>
: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="5B0D36D91E4BFFD6CBF9F8BAE9CCF02B" blockId="12.[151,1437,1863,2033]" lastBlockId="13.[151,1437,151,1257]" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
Pronotum transverse, 1.6 times as broad as long, 1.5–1.6 times as broad as head, widest about middle, distinctly more narrowed posteriad than anteriad, with widely rounded and slightly protruded apical angles; posterior angles obtuse, with slightly concave latero-basal margins; apical margin widely rounded, distinctly shorter than moderately straight or rounded posterior margin; middle portion evenly elevated, with or without indistinct longitudinal and distinct, wide semicircular impression in mediobasal third; lateral edges with distinct, irregular, small crenulation; lateral portions slightly explanate and widely impressed, each with deep oval pit slightly in front of middle. Prothorax with moderately long and narrow interocoxal process and widely protruded mesosternal processes (
<figureCitation id="C3892A5C1E4AFFD6CE02FF46E88AF1BF" box="[1340,1423,186,213]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 35</figureCitation>
). Mesoventrite moderately narrow, with acute intercoxal process, reaching apex of posterior margin of procoxae (
<figureCitation id="C3892A5C1E4AFFD6CE51FF22EDB2F077" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 38</figureCitation>
). Scutellum large, with widely rounded apical portion (
<figureCitation id="C3892A5C1E4AFFD6C825FEFEEE6BF074" box="[795,878,259,286]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 36</figureCitation>
). Metaventrite with very wide and deep intercoxal cavities and wide intercoxal process with rounded apex (
<figureCitation id="C3892A5C1E4AFFD6C82CFEDAEE62F02B" box="[786,871,295,321]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 39</figureCitation>
). Metendosternite as in
<figureCitation id="C3892A5C1E4AFFD6CF4CFEDAE9C1F02B" box="[1138,1220,295,321]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 40</figureCitation>
.
</paragraph>
<paragraph id="5B0D36D91E4AFFD6CBF9FEB6EFC3F09F" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">
Elytra slightly longer than broad and slightly broadened posteriad, reaching apical margin of abdominal tergite IV, with widely rounded apical margins, distinctly more than twice as long as pronotum; humeri widely rounded, not protruded anteriad (
<figureCitation id="C3892A5C1E4AFFD6CA9CFE6EECF3F0C7" box="[418,502,403,429]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 37</figureCitation>
); middle surface of each elytron with two indistinct longitudinal elevations; lateral portions narrow, slightly explanate; latero-apical edges with small and acute crenulation, more flattened and reduced in about apical third. Hind wings fully developed.
</paragraph>
<paragraph id="5B0D36D91E4AFFD6CBF9FE02E9E7F3C3" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">Legs long and slender, with very dense and long setation on pro- and mesotibiae, with additional long and moderately dense setae on inner sides, setation of metatibiae somewhat finer and sparser, with several stout setae around apical margin of each mesotibiae; middle and apical portions of inner sides of pro- and mesotibiae with two long rows of short peg-setae; pro- and mesotarsi twice longer than tibiae, metatarsi two and a half times as long as metatibia; apical metatarsomere of all tarsi about as long as three preceding combined tarsomeres.</paragraph>
<paragraph id="5B0D36D91E4AFFD6CBF9FD4EE9B0F27F" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">
Abdomen convex, distinctly narrower than elytra, with a pair of small and oval tomentose spots in middle of tergire V, with narrow palisade fringe on margin of abdominal tergite VII; intersegmental membranes between tergites IV–VI with brickwall-like sculpture; tergites and sternites VIII elongate (
<figureCitation id="C3892A5C1E4AFFD6C8F5FD06E9ACF27F" box="[971,1193,762,789]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Figs. 41–42, 44–45</figureCitation>
).
</paragraph>
<paragraph id="5B0D36D91E4AFFD6CBF9FCE2E9F3F2A3" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">
Male. Protarsomeres 1–4 wide. Apical margins of abdominal tergite VIII (
<figureCitation id="C3892A5C1E4AFFD6CF2CFCE2E960F253" box="[1042,1125,799,825]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 41</figureCitation>
) and sternite VIII (
<figureCitation id="C3892A5C1E4AFFD6CE7FFCE2E891F253" box="[1345,1428,799,825]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 42</figureCitation>
) slightly sinuate. Aedeagus narrow and very long, with small basal bulb, gradually narrowed toward median lobe with widely rounded apex; parameres very long, significantly exceeding apex of median lobe, significantly broadened in preapical portion and strongly narrowed toward slightly curved apices; each paramere with three short latero-apical setae; internal sac very narrow, long (
<figureCitation id="C3892A5C1E4AFFD6C9F3FC52EE27F2A3" box="[717,802,943,969]" captionStart="FIGURES 26–28" captionStartId="10.[151,264,1582,1606]" captionTargetBox="[161,1425,578,1546]" captionTargetId="figure-193@10.[151,1436,561,1558]" captionTargetPageId="10" captionText="FIGURES 26–28. Habitus (Fig. 26; paratype) and aedeagus (Figs. 27—28) of Anthobium morosum: 27—parameral view, 28—lateral view. Scale bar: 1.0 mm (Fig. 26), 0.1 mm (Figs. 27–28)." figureDoi="http://doi.org/10.5281/zenodo.6332054" httpUri="https://zenodo.org/record/6332054/files/figure.png" pageId="13" pageNumber="314">Fig. 27</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="C3892A5C1E4AFFD6CFA1FC52E9F7F2A3" box="[1183,1266,943,969]" captionStart="FIGURES 26–28" captionStartId="10.[151,264,1582,1606]" captionTargetBox="[161,1425,578,1546]" captionTargetId="figure-193@10.[151,1436,561,1558]" captionTargetPageId="10" captionText="FIGURES 26–28. Habitus (Fig. 26; paratype) and aedeagus (Figs. 27—28) of Anthobium morosum: 27—parameral view, 28—lateral view. Scale bar: 1.0 mm (Fig. 26), 0.1 mm (Figs. 27–28)." figureDoi="http://doi.org/10.5281/zenodo.6332054" httpUri="https://zenodo.org/record/6332054/files/figure.png" pageId="13" pageNumber="314">Fig. 28</figureCitation>
.
</paragraph>
<paragraph id="5B0D36D91E4AFFD6CBF9FC2EE81BF55F" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">
Female. Protarsomeres 1–4 moderately narrow. Apical margin of abdominal tergite VIII straight (
<figureCitation id="C3892A5C1E4AFFD6CE3AFC2EE85DF287" box="[1284,1368,979,1005]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 43</figureCitation>
). Apical margin of abdominal sternite VIII (
<figureCitation id="C3892A5C1E4AFFD6C968FC0AEFACF57B" box="[598,681,1015,1041]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 44</figureCitation>
) rounded. Genital segment with elongate sternite IX, gonocoxites moderately narrow and long, subparallel-sided, styli elongate and narrow, each with long seta (
<figureCitation id="C3892A5C1E4AFFD6CF80FBE6E817F55F" box="[1214,1298,1050,1077]" captionStart="FIGURES 34–45" captionStartId="12.[151,264,1648,1672]" captionTargetBox="[163,1428,187,1615]" captionTargetId="figure-21@12.[151,1436,181,1625]" captionTargetPageId="12" captionText="FIGURES 34–45. Anthobium morosum: 34—ventral aspect of head (ventral view), 35—ventral aspect of prothorax (ventral view), 36—scutellum (dorsal view), 37—ventral aspect of elytron, 38—ventral aspect of mesoventrite, 39—ventral aspect of metaventrite (without metanepisternum), 40—metendosternite (ventral view), 41—male abdominal tergite VIII (dorsal view), 42—male abdominal sternite VIII (ventral view), 43—female abdominal tergite VIII (dorsal view), 44—female abdominal sternite VIII (ventral view), 45—female genital segment (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332058" httpUri="https://zenodo.org/record/6332058/files/figure.png" pageId="13" pageNumber="314">Fig. 45</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="13A865521E4AFFD6CBF9FBC3EDEEF5CB" pageId="13" pageNumber="314" type="distribution">
<paragraph id="5B0D36D91E4AFFD6CBF9FBC3E859F533" blockId="13.[151,1437,151,1257]" box="[199,1372,1086,1113]" pageId="13" pageNumber="314">
<emphasis id="69C6EACB1E4AFFD6CBF9FBC3EC64F532" bold="true" box="[199,353,1086,1112]" pageId="13" pageNumber="314">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="8409887B1E4AFFD6C899FBC2EED2F533" box="[935,983,1087,1113]" pageId="13" pageNumber="314">type</typeStatus>
locality in central
<collectingCountry id="23A576491E4AFFD6CF97FBC2E9E9F533" box="[1193,1260,1087,1113]" name="Nepal" pageId="13" pageNumber="314">Nepal</collectingCountry>
(
<figureCitation id="C3892A5C1E4AFFD6CFC5FBC2E855F533" box="[1275,1360,1087,1113]" captionStart="FIGURE 93" captionStartId="33.[151,250,1706,1730]" captionTargetBox="[151,1437,885,1683]" captionTargetId="figure-299@33.[151,1437,885,1683]" captionTargetPageId="33" captionText="FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6332076" httpUri="https://zenodo.org/record/6332076/files/figure.png" pageId="13" pageNumber="314">Fig. 93</figureCitation>
).
</paragraph>
<paragraph id="5B0D36D91E4AFFD6CBF9FB9EEDEEF5CB" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">
<emphasis id="69C6EACB1E4AFFD6CBF9FB9EEC83F516" bold="true" box="[199,390,1122,1149]" pageId="13" pageNumber="314">Natural history.</emphasis>
The specimens were collected at an elevation
<quantity id="9C4A9B3C1E4AFFD6C8A7FB9EEEEAF517" box="[921,1007,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="314" unit="m" value="2800.0">2800 m</quantity>
a.s.l. Detailed ecological data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="13A865521E4AFFD6CBF9FB56EC95F583" pageId="13" pageNumber="314" type="etymology">
<paragraph id="5B0D36D91E4AFFD6CBF9FB56EC95F583" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="314">
<emphasis id="69C6EACB1E4AFFD6CBF9FB56EC4BF5AF" bold="true" box="[199,334,1195,1221]" pageId="13" pageNumber="314">Etymology.</emphasis>
The specific epithet (Latin adjective: strange) alludes to the unusual shapes of some morphological structures of the body.
</paragraph>
</subSubSection>
</treatment>
</document>
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