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<document id="241094249A5F63203D5505561415A1F3" ID-DOI="10.1093/zoolinnean/zlad143" ID-ISSN="0024-4082" ID-Zenodo-Dep="14443945" ID-ZooBank="CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734089919291" checkinUser="plazi" docAuthor="Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins &amp; Solodovnikov, Alexey" docDate="2024" docId="C60DAE358627D831D94FF9ECBDF2FABE" docLanguage="en" docName="zlad143.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlad143" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Loncovilius impunctus Reyes-Hernández, Hansen, Shaw &amp; Solodovnikov, 2024, sp.nov." docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="3A34D64D863DD82AD863FFB2BF3BFFFA" masterDocTitle="Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="27" updateTime="1734091152880" updateUser="ExternalLinkService">
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<mods:title id="33B111D59CD94945FF41E28E0A76C36A">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:role id="44D48053D7A592EC03D77F81E781B401">
<mods:roleTerm id="A7C5338A8D165D0C8A7A776CEB4C770D">Author</mods:roleTerm>
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<mods:namePart id="C71AA14191377FA7976EB063BB1B09E4">Reyes-Hernández, José L.</mods:namePart>
<mods:affiliation id="8C050C0DEAAD513CB2AAE0441D109995">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
<mods:nameIdentifier id="1D32042E20C041FD7782523E013CA3FD" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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<mods:name id="D2713E907FA14F315AAD3E3098EB7678" type="personal">
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</mods:role>
<mods:namePart id="0D93DF2E7B369DD8A3E8EC05A56BA0AE">Hansen, Aslak Kappel</mods:namePart>
<mods:affiliation id="BD975022504AD252EA3349957871505A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark &amp; Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
</mods:name>
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<mods:roleTerm id="A5865BFF4959E4BAF9B9B848A218F751">Author</mods:roleTerm>
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<mods:namePart id="6A59670E24542F717C77EF307A809594">Shaw, Josh Jenkins</mods:namePart>
<mods:affiliation id="833F1FB890D5B2929738CE8D89B9145A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:role id="3093980667CD6CA8E22C24B16602AA41">
<mods:roleTerm id="672960D4C3FF7A138F8947F06B201C47">Author</mods:roleTerm>
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<mods:namePart id="591CE30D4AB07E1BBC170F394A95F67C">Solodovnikov, Alexey</mods:namePart>
<mods:affiliation id="E7F59828838F0E4819CDF89274C36F6C">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:typeOfResource id="C5BE1DB0EECCBC5FFF284AD008239390">text</mods:typeOfResource>
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<mods:title id="F20371A2F8BC383218DE8478F0877C45">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="0D959E4CA15FA9B2CC19C4182E746D64">
<mods:date id="5630B5981BA7C26F616F2BB1312F63FB">2024</mods:date>
<mods:detail id="EA0D627D30913D3F2107EBEEECD506AA" type="series">
<mods:title id="32B53FB657D04720481143309DE6B6BA">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="7126935247B63B0D2A9FC28281793975">2023-10-28</mods:number>
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<mods:detail id="570732EB8F141424C8ED543B1088552D" type="volume">
<mods:number id="AB2134DCF0CCD04B1980E0A7D5BBA78D">202</mods:number>
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<mods:start id="CE6E433EF17BE4062EB3434F8573CD33">1</mods:start>
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<mods:classification id="951636952D4962D19CAE42F96D5DE88A">journal article</mods:classification>
<mods:identifier id="150A23BCFFB6F284FF552DF210E01AE1" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
<mods:identifier id="90DE5FF7FB0BF758BED1C7B72C49C383" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="A5A360F39D3C851BD90533D207C4E780" type="Zenodo-Dep">14443945</mods:identifier>
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<treatment id="C60DAE358627D831D94FF9ECBDF2FABE" ID-DOI="http://doi.org/10.5281/zenodo.14444227" ID-Zenodo-Dep="14444227" LSID="urn:lsid:plazi:treatment:C60DAE358627D831D94FF9ECBDF2FABE" httpUri="http://treatment.plazi.org/id/C60DAE358627D831D94FF9ECBDF2FABE" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="06BE4CA88627D830D94FF9ECBD66F982" box="[300,605,1630,1657]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="4E1B1F238627D830D94FF9ECBD66F982" blockId="26.[300,605,1630,1657]" box="[300,605,1630,1657]" pageId="26" pageNumber="27">
<heading id="1553A84F8627D830D94FF9ECBD66F982" bold="true" box="[300,605,1630,1657]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<emphasis id="7CD0C3318627D830D94FF9ECBD66F982" bold="true" box="[300,605,1630,1657]" pageId="26" pageNumber="27">
<taxonomicName id="89A464A08627D830D94FF9ECBD36F982" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[300,525,1630,1657]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus" status="sp. nov.">
<emphasis id="7CD0C3318627D830D94FF9ECBD36F982" bold="true" box="[300,525,1630,1657]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67E37E4A8627D830DA72F9ECBD66F982" box="[529,605,1630,1656]" pageId="26" pageNumber="27" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="06BE4CA88627D831D971F92EBDF2FABE" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="4E1B1F238627D830D971F92EBD43F94C" blockId="26.[274,632,1692,1719]" box="[274,632,1692,1719]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830D971F92EBD43F94C" bold="true" box="[274,632,1692,1719]" pageId="26" pageNumber="27">
(
<figureCitation id="D69F03A68627D830D97EF92FBE56F94D" box="[285,365,1693,1719]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." figureDoi="http://doi.org/10.5281/zenodo.14443961" httpUri="https://zenodo.org/record/14443961/files/figure.png" pageId="26" pageNumber="27">Figs 5C</figureCitation>
,
<figureCitation id="D69F03A68627D830D914F92FBEFEF94D" box="[375,453,1693,1719]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="26" pageNumber="27">10G, H</figureCitation>
,
<figureCitation id="D69F03A68627D830D9ACF92FBEC6F94D" box="[463,509,1693,1719]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="26" pageNumber="27">12C</figureCitation>
,
<figureCitation id="D69F03A68627D830DA64F92FBD0EF94D" box="[519,565,1693,1719]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="26" pageNumber="27">13C</figureCitation>
,
<figureCitation id="D69F03A68627D830DA5CF92FBD56F94D" box="[575,621,1693,1719]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="26" pageNumber="27">14C</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238627D830D8E2F963BED7F8F2" blockId="26.[129,777,1744,1800]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830D8E2F963BE51F912" box="[129,362,1745,1769]" italics="true" pageId="26" pageNumber="27">Zoobank registration:</emphasis>
<uri id="3A3513218627D830D9EAF962BEDCF8F2" pageId="26" pageNumber="27">
urn:lsid:zoobank.org:act:
<uuid id="3A0225F68627D830DAE9F962BEDCF8F2" pageId="26" pageNumber="27">45463DC0- 4ECA-4B24-8220-24008B5BE531</uuid>
</uri>
.
</paragraph>
<paragraph id="4E1B1F238627D830D8E2F898BA66FE99" blockId="26.[127,778,1833,1983]" lastBlockId="26.[824,1474,144,450]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830D8E2F898BE34F8BB" box="[129,271,1833,1857]" italics="true" pageId="26" pageNumber="27">Type material.</emphasis>
<materialsCitation id="FECC157E8627D830D977F89BBF9DF865" collectingDate="1954-07" collectorName="L. Pena" country="Chile" location="Cherquenco" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="7CD0C3318627D830D977F89BBE4FF8BB" box="[276,372,1833,1857]" italics="true" pageId="26" pageNumber="27">
<typeStatus id="911FA1818627D830D977F89BBE54F8BB" box="[276,367,1833,1857]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
:
</emphasis>
male, mounted, with genitalia in a separate microvial, with labels as follows:
<collectingCountry id="36B35FB38627D830DA6EF8FBBD5BF89B" box="[525,608,1865,1889]" name="Chile" pageId="26" pageNumber="27">CHILE</collectingCountry>
:
<location id="4B7B49F88627D830DA0AF8FBBDD2F89B" LSID="urn:lsid:plazi:treatment:C60DAE358627D831D94FF9ECBDF2FABE:4B7B49F88627D830DA0AF8FBBDD2F89B" box="[617,745,1865,1889]" country="Chile" name="Cherquenco" pageId="26" pageNumber="27">Cherquenco</location>
[~ -38.70, -72.01]
<date id="3A1A39E38627D830D97FF8DABE41F87A" box="[284,378,1896,1920]" pageId="26" pageNumber="27" value="1954-07">
<collectingDate id="2A5EC00B8627D830D97FF8DABE41F87A" box="[284,378,1896,1920]" pageId="26" pageNumber="27" value="1954-07">VII-1954</collectingDate>
</date>
<collectorName id="E3517AF58627D830D91CF8DABEF0F885" box="[383,459,1896,1919]" pageId="26" pageNumber="27">L. Pena</collectorName>
, leg./
<specimenCount id="58A2D4AA8627D830DA62F8D9BD20F885" box="[513,539,1899,1919]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
/FMNH-INS 0000 024 367
</materialsCitation>
/
<typeStatus id="911FA1818627D830D8CDF835BE05F865" box="[174,318,1927,1951]" pageId="26" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="89A464A08627D830D927F835BD27F865" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[324,540,1927,1951]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus">
<emphasis id="7CD0C3318627D830D927F835BD27F865" box="[324,540,1927,1951]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
<materialsCitation id="FECC157E8627D830DBF9FF23BCAEFF1C" collectingDate="1954-07" collectorName="L. Pena" country="Chile" location="Cherquenco" pageId="26" pageNumber="27" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="7CD0C3318627D830DBF9FF23BB3BFF52" box="[922,1024,145,168]" italics="true" pageId="26" pageNumber="27">
<typeStatus id="911FA1818627D830DBF9FF23BCC7FF52" box="[922,1020,145,168]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="36B35FB38627D830DC6EFF22BB5BFF52" box="[1037,1120,144,168]" name="Chile" pageId="26" pageNumber="27">CHILE</collectingCountry>
:
<location id="4B7B49F88627D830DC13FF22BBCBFF52" LSID="urn:lsid:plazi:treatment:C60DAE358627D831D94FF9ECBDF2FABE:4B7B49F88627D830DC13FF22BBCBFF52" box="[1136,1264,144,168]" country="Chile" name="Cherquenco" pageId="26" pageNumber="27">Cherquenco</location>
[~ -38.70, -72.01]
<date id="3A1A39E38627D830DB5AFF1DBCACFF3D" box="[825,919,175,199]" pageId="26" pageNumber="27" value="1954-07">
<collectingDate id="2A5EC00B8627D830DB5AFF1DBCACFF3D" box="[825,919,175,199]" pageId="26" pageNumber="27" value="1954-07">VII-1954</collectingDate>
</date>
<collectorName id="E3517AF58627D830DBFEFF02BCD0FF3D" box="[925,1003,176,199]" pageId="26" pageNumber="27">L. Pena</collectorName>
, leg./
<specimenCount id="58A2D4AA8627D830DC47FF00BB05FF3C" box="[1060,1086,178,198]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
/FMNH-INS 0000 024 368-369 [
<specimenCount id="58A2D4AA8627D830DDFBFF1DBAF9FF3C" box="[1432,1474,175,199]" count="2" pageId="26" pageNumber="27" type="female">2♀</specimenCount>
FMNH]
</materialsCitation>
;
<materialsCitation id="FECC157E8627D830DBC7FF7DBA2EFEDF" collectingDate="1976-12-19" collectingDateMax="1976-12-21" collectingDateMin="1976-12-19" collectorName="H. F. Howden" country="Chile" elevation="1100" location="Malleco Prov." pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingCountry id="36B35FB38627D830DBC7FF7DBCCCFF1D" box="[932,1015,207,231]" name="Chile" pageId="26" pageNumber="27">CHILE</collectingCountry>
:
<location id="4B7B49F88627D830DC67FF7DBBADFF1C" LSID="urn:lsid:plazi:treatment:C60DAE358627D831D94FF9ECBDF2FABE:4B7B49F88627D830DC67FF7DBBADFF1C" box="[1028,1174,207,231]" country="Chile" name="Malleco Prov." pageId="26" pageNumber="27">Malleco Prov.</location>
<locationDeviation id="5FA906018627D830DCC3FF7DBA5EFF1C" box="[1184,1381,207,231]" location="Manzanar" pageId="26" pageNumber="27">
<quantity id="895CB2C68627D830DCC3FF7DBBE0FF1D" box="[1184,1243,207,231]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="km" value="20.0">20km</quantity>
E Manzanar
</locationDeviation>
[-38.44, -71.51]
<quantity id="895CB2C68627D830DBF3FF5CBCE2FEFC" box="[912,985,238,262]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="27" unit="m" value="1100.0">
<elevation id="C589F8108627D830DBF3FF5CBCE2FEFC" box="[912,985,238,262]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="27" unit="m" value="1100.0">1100m</elevation>
</quantity>
<date id="3A1A39E38627D830DB87FF5CBB42FEFC" box="[996,1145,238,262]" pageId="26" pageNumber="27" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">
<collectingDate id="2A5EC00B8627D830DB87FF5CBB42FEFC" box="[996,1145,238,262]" pageId="26" pageNumber="27" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">19-21.xii.1976</collectingDate>
</date>
<collectorName id="E3517AF58627D830DCE7FF5CBA19FEFC" box="[1156,1314,238,262]" pageId="26" pageNumber="27">H. F. Howden</collectorName>
/CNCI L2004- 28/
<specimenCount id="58A2D4AA8627D830DB03FEA2BC41FEDE" box="[864,890,272,292]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
/FMNH-INS 0000 024 465 [
<specimenCount id="58A2D4AA8627D830DCC0FEBFBBF0FEDE" box="[1187,1227,269,293]" count="1" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
CNC]
</materialsCitation>
;
<materialsCitation id="FECC157E8627D830DD7DFEBFBA63FE99" collectorName="Coll. L. Pena" country="Chile" location="Chile" pageId="26" pageNumber="27" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<collectingCountry id="36B35FB38627D830DD7DFEBFBA60FEDF" box="[1310,1371,269,293]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
: Tolhuaco [~ -38.23, -71.71]11.I.62/
<collectorName id="E3517AF58627D830DC25FE9FBBECFEBF" box="[1094,1239,301,325]" pageId="26" pageNumber="27">Coll. L. Peña</collectorName>
/
<specimenCount id="58A2D4AA8627D830DC83FE82BBC1FEBE" box="[1248,1274,304,324]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
/FMNH-INS 0000 024 364–366 [
<specimenCount id="58A2D4AA8627D830DBADFEFEBCC3FE99" box="[974,1016,332,356]" count="2" pageId="26" pageNumber="27" type="male">2♂</specimenCount>
FMNH;
<specimenCount id="58A2D4AA8627D830DC3BFEFEBB44FE99" box="[1112,1151,332,356]" count="1" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
MNNC ex. FMNH]
</materialsCitation>
.
</paragraph>
<paragraph id="4E1B1F238627D830DB36FED9BB6EFE38" blockId="26.[824,1474,144,450]" pageId="26" pageNumber="27">
All
<typeStatus id="911FA1818627D830DBE0FEDEBCDDFE7E" box="[899,998,364,388]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
are supplied with the label: PAÞTYPE
<taxonomicName id="89A464A08627D830DB5AFE39BB2DFE58" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[825,1046,395,419]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus">
<emphasis id="7CD0C3318627D830DB5AFE39BB2DFE58" box="[825,1046,395,419]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
</paragraph>
<paragraph id="4E1B1F238627D830DB5AFE56BA3BFD2C" blockId="26.[824,1475,483,727]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830DB5AFE56BCA4FE01" box="[825,927,484,507]" italics="true" pageId="26" pageNumber="27">Diagnosis:</emphasis>
Head dorsally with meshed microsculpture; gula with isodiametric microsculpture in the middle; females with antennomere 11 (a11) distinctly longer than antennomere 10 (a10; a11:a10 ratio&gt;1.5). Elytra reddish-brown, disc glossy with only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII in both sexes with medial apical emargination. Aedeagus as in
<figureCitation id="D69F03A68627D830DC04FD0DBBC1FD2C" box="[1127,1274,703,727]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="26" pageNumber="27">Figure 10G, H</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238627D830DB5AFD4BBB0CFC57" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830DB5AFD4BBC8BFCEA" box="[825,944,761,784]" italics="true" pageId="26" pageNumber="27">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8627D830DC02FD49BB40FCF5" box="[1121,1147,763,783]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318627D830DD10FD4BBABDFCEA" box="[1395,1414,761,784]" italics="true" pageId="26" pageNumber="27">N</emphasis>
= 3]: FBL = 3.35–3.59 (3.48); HW = 1.01–1.1 (1.07); HL = 0.85– 0.93 (0.89); HW/HL = 1.18–1.21 (1.19); PW = 1.4– 1.44 (1.42); PL = 1.2–1.25 (1.23); PW/PL = 1.14–1.17 (1.15); EW = 1.67–1.7 (1.68); EL = 1.3–1.44 (1.36); PW/ HW = 1.28–1.39 (1.33).
</paragraph>
<paragraph id="4E1B1F238627D830DB36FC06BB81FB93" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
Measurements
<specimenCount id="58A2D4AA8627D830DB93FC05BB31FC31" box="[1008,1034,951,971]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318627D830DC89FC07BBC6FC36" box="[1258,1277,949,972]" italics="true" pageId="26" pageNumber="27">N</emphasis>
= 4]: FBL = 3.71– 3.93 (3.81); TL = 7.5–7.53 (7.52); HW = 1.17–1.18 (1.18); HL = 0.93–0.97 (0.95); HW/HL = 1.22–1.26 (1.24); PW = 1.54–1.59 (1.56); PL = 1.29–1.37 (1.32); PW/ PL = 1.16–1.19 (1.18); EW = 1.82–1.9 (1.87); EL = 1.46–1.59 (1.54); PW/HW = 1.32–1.35 (1.33).
</paragraph>
<paragraph id="4E1B1F238627D830DB36FBC2BCBAFB3D" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">Head, mouthparts, and legs black to brown-black; pronotum, antennae, elytra, and abdomen dark reddish-brown, elytra lighter.</paragraph>
<paragraph id="4E1B1F238627D830DB36FB7CBB62F84D" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse meshes, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes of medium size (EYL/HL = x = 0.52), from 1.51 to 1.64 times longer than temples (in lateral view); distance between eyes around 1.8 times the length of eye. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 subequal in length; antennomeres 4 to 10 gradually reduced in length; a11 from 1.54 to 1.75 times longer than a
<quantity id="895CB2C68627D830DD2EFA5ABABAF9FA" box="[1357,1409,1512,1536]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68627D830DBC4F9B5BB7CF9E5" box="[935,1095,1543,1567]" metricMagnitude="-2" metricUnit="m" metricValue="4.0893999999999995" metricValueMax="4.2672" metricValueMin="3.9116" pageId="26" pageNumber="27" unit="in" value="1.6099999999999999" valueMax="1.68" valueMin="1.54">1.54 to 1.68 in</quantity>
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical.
</paragraph>
<paragraph id="4E1B1F238626D831D8EEFF22BD10FEBF" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">Pronotum transverse, convex, evenly curved, with one PPDS and an APP, without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setose; disc with only a few rows of setiferous punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F238626D831D8EEFEFEBFFBFDC5" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrallywith apical margin slightly sinuate.</paragraph>
<paragraph id="4E1B1F238626D831D8EEFDF5BD5EFC62" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flaưened only in males. Male sternite VIII with broad V-shaped emargination medially (
<figureCitation id="D69F03A68626D831D910FD51BEEBFD01" box="[371,464,739,763]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="27" pageNumber="28">Fig. 14C</figureCitation>
); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (
<figureCitation id="D69F03A68626D831DA63FCF3BD6CFCA3" box="[512,599,833,857]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="27" pageNumber="28">Fig. 12C</figureCitation>
); female tergite X truncate to slightly arcuate medially (
<figureCitation id="D69F03A68626D831DA61FCD3BD62FC83" box="[514,601,865,889]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="27" pageNumber="28">Fig. 13C</figureCitation>
); ovipositor, its second gonocoxite without macrosetae medially.
</paragraph>
<paragraph id="4E1B1F238626D831D8EEFC2DBDAAFBAE" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">
Aedeagus as in
<figureCitation id="D69F03A68626D831D951FC2DBEF7FC4D" box="[306,460,927,951]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="27" pageNumber="28">Figure 10G, H</figureCitation>
; its total length 1.2. Median lobe bulky, in parameral view with truncated apex; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into M-shape apex, with dark peg setae, in lateral view almost straight and slightly concave apically.
</paragraph>
<paragraph id="4E1B1F238626D831D812FBC7BD11FB36" blockId="27.[113,762,1141,1228]" pageId="27" pageNumber="28">
<emphasis id="7CD0C3318626D831D812FBC7BFDAFB77" box="[113,225,1141,1165]" italics="true" pageId="27" pageNumber="28">Etymology:</emphasis>
Ŋe species epithet is derived from the Latin prefix ‘
<emphasis id="7CD0C3318626D831D814FB27BFAAFB56" box="[119,145,1173,1196]" italics="true" pageId="27" pageNumber="28">im</emphasis>
’ (not) and adjective ‘
<emphasis id="7CD0C3318626D831D90DFB27BEFAFB56" box="[366,449,1173,1196]" italics="true" pageId="27" pageNumber="28">punctate</emphasis>
’ (doưed), referring to the disc of pronotum and elytra mostly impunctate.
</paragraph>
<paragraph id="4E1B1F238626D831D812FB5FBDF2FABE" blockId="27.[112,762,1261,1348]" pageId="27" pageNumber="28">
<emphasis id="7CD0C3318626D831D812FB5FBD05FAFF" box="[113,574,1261,1285]" italics="true" pageId="27" pageNumber="28">
Distribution and habitat:
<taxonomicName id="89A464A08626D831D904FB5FBD05FAFF" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[359,574,1261,1285]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="impunctus">Loncovilius impunctus</taxonomicName>
</emphasis>
is known from the
<collectingRegion id="8C60D1C18626D831D813FABFBE1DFADF" box="[112,294,1293,1317]" country="Chile" name="Araucania" pageId="27" pageNumber="28">Araucanía Region</collectingRegion>
in
<collectingCountry id="36B35FB38626D831D927FABFBE47FADF" box="[324,380,1293,1317]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
where it is confined to Valdivian temperate forest ecoregion. It is unknown how it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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