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<document id="3AEBE078F4D8892394E2C8AFA564CEDA" ID-CLB-Dataset="306451" ID-DOI="10.11646/zootaxa.5551.1.7" ID-GBIF-Dataset="1c81b294-3149-4e1d-b6e1-83315506956f" ID-ISSN="1175-5326" ID-Zenodo-Dep="14390215" ID-ZooBank="43517112-40DA-4E69-8BFB-BEC55AC5AAF2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733924615010" checkinUser="plazi" docAuthor="Delgado-Blas, Vi ́ Ctor Hugo, Díaz-Díaz, Oscar &amp; Rozbaczylo, Nicolás" docDate="2024" docId="0381CB35A125FF8DFF50FD2BFA37A1CC" docLanguage="en" docName="zootaxa.5551.1.7.pdf" docOrigin="Zootaxa 5551 (1)" docSource="https://doi.org/10.11646/zootaxa.5551.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.20:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="20" docTitle="Laonice Malmgren 1867" docType="key" docVersion="4" lastPageNumber="165" masterDocId="FFB8B34DA12DFF85FFC7FF98FFABA45E" masterDocTitle="Laonice Malmgren, 1867 (Annelida: Spionidae) from the Aysén region in southern Chile with description of a new species" masterLastPageNumber="166" masterPageNumber="157" pageNumber="165" updateTime="1734091358508" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="814077A67571BC02252EC1B5BB0A5B88">Laonice Malmgren, 1867 (Annelida: Spionidae) from the Aysén region in southern Chile with description of a new species</mods:title>
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<mods:namePart id="D36C6A9EFFCBAC4186AD5A897F27EDCF">Delgado-Blas, Vi ́ Ctor Hugo</mods:namePart>
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<mods:affiliation id="0412488110A659FD69C8CB304ED52929">Universidad Auto ́ noma de Quintana Roo, Divisio ́ n de Ciencias, Ingenieri ́ a y Tecnologia, Chetumal, Quintana Roo, 77019, Me ́ xico</mods:affiliation>
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<mods:namePart id="34785F891A3FC4AD7D150F1928224BD9">Díaz-Díaz, Oscar</mods:namePart>
<mods:affiliation id="13330DA1437348A98B913AB1530E658F">FAUNAMAR Ltda. Consultorias Medio Ambientales e Investigación Marina, Santiago. Chile &amp; Postgrado en Ciencias Marinas, Instituto Oceanográfico de Venezuela, Venezuela</mods:affiliation>
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<mods:namePart id="79DC841DF0D34B65A3175D99594D84A7">Rozbaczylo, Nicolás</mods:namePart>
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<mods:affiliation id="6CF0A1513D35573BEA5D37D8DDEBDBF1">FAUNAMAR Ltda. Consultorias Medio Ambientales e Investigación Marina, Santiago. Chile</mods:affiliation>
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<treatment id="0381CB35A125FF8DFF50FD2BFA37A1CC" ID-DOI="http://doi.org/10.5281/zenodo.14444269" ID-Zenodo-Dep="14444269" LSID="urn:lsid:plazi:treatment:0381CB35A125FF8DFF50FD2BFA37A1CC" httpUri="http://treatment.plazi.org/id/0381CB35A125FF8DFF50FD2BFA37A1CC" lastPageNumber="165" pageId="8" pageNumber="165">
<subSubSection id="C33229A8A125FF8DFF50FD2BFD75A692" box="[151,734,690,717]" pageId="8" pageNumber="165" type="nomenclature">
<paragraph id="8B977A23A125FF8DFF50FD2BFD75A692" blockId="8.[151,734,690,717]" box="[151,734,690,717]" pageId="8" pageNumber="165">
<heading id="D0DFCD4FA125FF8DFF50FD2BFD75A692" bold="true" box="[151,734,690,717]" fontSize="11" level="1" pageId="8" pageNumber="165" reason="1">
<emphasis id="B95CA631A125FF8DFF50FD2BFD75A692" bold="true" box="[151,734,690,717]" pageId="8" pageNumber="165">
Key to all
<taxonomicName id="4C2801A0A125FF8DFED0FD2BFED2A693" ID-CoL="84PJQ" authority="Malmgren, 1867" authorityName="Malmgren" authorityYear="1867" box="[279,377,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="genus">
<emphasis id="B95CA631A125FF8DFED0FD2BFED2A693" bold="true" box="[279,377,691,717]" italics="true" pageId="8" pageNumber="165">Laonice</emphasis>
</taxonomicName>
species from South America
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A125FF8DFF50FD60FA37A1CC" pageId="8" pageNumber="165" type="key">
<keyStep id="30DC6286A125FF8DFF50FD60FA37A775" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FD60FA37A753" blockId="8.[151,1437,760,1427]" box="[151,1436,760,782]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FD60FA37A753" box="[151,1436,760,782]" pageId="8" pageNumber="165">1 Prostomium fused or partially fused to peristomium.......................................................... 2</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FC8DFA37A775" blockId="8.[151,1437,760,1427]" box="[151,1436,789,811]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FC8DFA37A775" box="[151,1436,789,811]" pageId="8" pageNumber="165">- Prostomium free from peristomium....................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DC6286A125FF8DFF50FCAAFA37A73B" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FCAAFA37A719" blockId="8.[151,1437,760,1427]" box="[151,1436,818,840]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FCAAFA37A719" box="[151,1436,818,840]" pageId="8" pageNumber="165">2 Prostomium clearly fused with peristomium along the anterior margin........................................... 3</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FCC8FA37A73B" blockId="8.[151,1437,760,1427]" box="[151,1436,847,870]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FCC8FA37A73B" box="[151,1436,847,870]" pageId="8" pageNumber="165">- Prostomium partially fused with peristomium, generally only by an inconspicuous membrane......................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DC6286A125FF8DFF50FCF5FA37A7C1" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FCF5FA37A7DD" blockId="8.[151,1437,760,1427]" box="[151,1436,876,899]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FCF5FA37A7DD" box="[151,1436,876,899]" pageId="8" pageNumber="165">
3 Branchiae with appendages on frontal basal region........................
<taxonomicName id="4C2801A0A125FF8DFC2DFCF5FB6BA7DC" ID-CoL="3S88K" authority="Nonato" authorityName="Nonato" box="[1002,1216,876,898]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="branchiata">
<emphasis id="B95CA631A125FF8DFC2DFCF5FBDAA7DC" bold="true" box="[1002,1137,876,898]" italics="true" pageId="8" pageNumber="165">L. branchiata</emphasis>
Nonato
</taxonomicName>
,
<collectingRegion id="49ECB4C1A125FF8DFB0DFCF5FAB9A7DD" box="[1226,1298,877,899]" country="Venezuela" name="Bolivar" pageId="8" pageNumber="165">Bolivar</collectingRegion>
&amp; Lana, 1986
</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FC13FA37A7C1" blockId="8.[151,1437,760,1427]" box="[151,1436,906,928]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FC13FA37A7C1" box="[151,1436,906,928]" pageId="8" pageNumber="165">- Branchiae without appendages (smooth)................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DC6286A125FF8DFF50FC30FA37A04B" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FC30FA37A785" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FC30FA37A785" pageId="8" pageNumber="165">
4 Ventrolateral pouches present from chaetigers 3–17, or absent; dorsal transverse ridges absent; hooded hooks complete; main fang of hooded hooks is accompanied at the base by apical teeth........................
<taxonomicName id="4C2801A0A125FF8DFBB4FC5DFA37A785" ID-CoL="3S88L" authority="(Kinberg, 1866)" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[1139,1436,964,987]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A125FF8DFBB4FC5DFB57A784" bold="true" box="[1139,1276,964,986]" italics="true" pageId="8" pageNumber="165">L. brevicornis</emphasis>
(Kinberg, 1866)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FC7BFA37A04B" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FC7BFA37A04B" pageId="8" pageNumber="165">
- Ventrolateral pouches present from chaetigers 35–48; dorsal transverse ridges present; hooded hooks with distal opening; main fang of hooded hooks is not accompanied at the base by apical teeth............................
<emphasis id="B95CA631A125FF8DFB78FB98FA37A04B" bold="true" box="[1215,1436,1023,1045]" pageId="8" pageNumber="165">
<taxonomicName id="4C2801A0A125FF8DFB78FB98FAE1A04A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[1215,1354,1023,1045]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFB78FB98FAE1A04A" bold="true" box="[1215,1354,1023,1045]" italics="true" pageId="8" pageNumber="165">L. ayseniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F1B4AA125FF8DFA97FB98FA37A04B" box="[1360,1436,1024,1045]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DC6286A125FF8DFF50FB85FA37A09B" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FB85FA37A033" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FB85FA37A033" pageId="8" pageNumber="165">
5 Caruncle and nuchal organs reach chaetigers 15–21; branchiae present until chaetigers 32–59; ventrolateral pouches present from chaetigers 9–24; dorsal transverse ridges present; hooded hooks from chaetigers 17–24; sabre chaetae from chaetigers 11–15..............................................................
<taxonomicName id="4C2801A0A125FF8DFBD4FBC0FA37A033" ID-CoL="85RGX" authority="Hartmann-Schroder, 1965" authorityName="Hartmann-Schroder" authorityYear="1965" box="[1043,1436,1111,1134]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="praecirrata">
<emphasis id="B95CA631A125FF8DFBD4FBC0FB36A032" bold="true" box="[1043,1181,1111,1133]" italics="true" pageId="8" pageNumber="165">L. praecirrata</emphasis>
<bibRefCitation id="EFB907D2A125FF8DFB64FBC0FA37A033" author="Hartmann-Schroder, G." box="[1187,1436,1111,1134]" pageId="8" pageNumber="165" pagination="59 - 305" refId="ref7629" refString="Hartmann-Schroder, G. (1965) Zur Kenntnis des Sublitorals der chilenischen Kuste unter besonderer Berucksichtigung der Polychaeten und Ostracoden. II Die Polychaeten des Sublitorals. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 62 (Supplement), 59 - 305." type="journal article" year="1965">Hartmann-Schröder, 1965</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FBEDFA37A09B" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FBEDFA37A09B" pageId="8" pageNumber="165">
- Caruncle and nuchal organs reach chaetigers 12–13; branchiae present until chaetiger 32; Ventrolateral pouches present from chaetigers 3–4; dorsal transverse ridges absent; hooded hooks from chaetigers 34–38; sabre chaetae from chaetigers 16–22..............................................................................
<taxonomicName id="4C2801A0A125FF8DFBBAFB28FA37A09B" ID-CoL="3S88B" authority="Hartman, 1953" authorityName="Hartman" authorityYear="1953" box="[1149,1436,1199,1221]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="antarcticae">
<emphasis id="B95CA631A125FF8DFBBAFB28FAACA09A" bold="true" box="[1149,1287,1199,1221]" italics="true" pageId="8" pageNumber="165">L. antarcticae</emphasis>
Hartman, 1953
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DC6286A125FF8DFF50FB55FA37A143" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FB55FA37A0A1" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FB55FA37A0A1" pageId="8" pageNumber="165">
6 Prostomium with short frontolateral horns; sabre chaetae from chaetigers 27–29.........................................................................................
<taxonomicName id="4C2801A0A125FF8DFCAFFB73FB4DA15E" ID-CoL="3S896" authority="Radashevsky &amp; Da" authorityName="Radashevsky &amp; Da" box="[872,1254,1258,1280]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="parvabranchiata">
<emphasis id="B95CA631A125FF8DFCAFFB73FB8DA15E" bold="true" box="[872,1062,1258,1280]" italics="true" pageId="8" pageNumber="165">L. parvabranchiata</emphasis>
Radashevsky &amp; Da
</taxonomicName>
Cunha Lana, 2009
</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FA90FA37A143" blockId="8.[151,1437,760,1427]" box="[151,1436,1287,1310]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FA90FA37A143" box="[151,1436,1287,1310]" pageId="8" pageNumber="165">- Prostomium without short fronto-lateral horns; sabre chaetae from chaetigers 5–17................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DC6286A125FF8DFF50FABDFA37A1CC" pageId="8" pageNumber="165">
<paragraph id="8B977A23A125FF8DFF50FABDFA37A106" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FABDFA37A106" pageId="8" pageNumber="165">
7 Caruncle and nuchal organs reach chaetigers 21–28; branchiae present until chaetiger 42; dorsal transverse membranes absent; hooded hooks from chaetiger 31; sabre chaetae from chaetigers 16–17............
<taxonomicName id="4C2801A0A125FF8DFBD0FADBFA37A106" ID-CoL="85RGW" authority="Hartmann-Schroder, 1965" authorityName="Hartmann-Schroder" authorityYear="1965" box="[1047,1436,1346,1368]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFBD0FADBFB36A109" bold="true" box="[1047,1181,1346,1368]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
<bibRefCitation id="EFB907D2A125FF8DFB64FADAFA37A106" author="Hartmann-Schroder, G." box="[1187,1436,1346,1368]" pageId="8" pageNumber="165" pagination="59 - 305" refId="ref7629" refString="Hartmann-Schroder, G. (1965) Zur Kenntnis des Sublitorals der chilenischen Kuste unter besonderer Berucksichtigung der Polychaeten und Ostracoden. II Die Polychaeten des Sublitorals. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 62 (Supplement), 59 - 305." type="journal article" year="1965">Hartmann-Schröder, 1965</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B977A23A125FF8DFF50FAF8FA37A1CC" blockId="8.[151,1437,760,1427]" pageId="8" pageNumber="165">
<keyLead id="30D9D916A125FF8DFF50FAF8FA37A1CC" pageId="8" pageNumber="165">
- Caruncle and nuchal organs reach chaetiger 20; branchiae present until chaetiger 50; dorsal transverse membranes present; hooded hooks from chaetigers 16–25; sabre chaetae from chaetiger 5.......................
<taxonomicName id="4C2801A0A125FF8DFB49FAE5FA37A1CC" ID-CoL="3S89J" authority="Hartman, 1978" authorityName="Hartman" authorityYear="1978" box="[1166,1436,1404,1426]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="weddellia">
<emphasis id="B95CA631A125FF8DFB49FAE5FAADA1CC" bold="true" box="[1166,1286,1404,1426]" italics="true" pageId="8" pageNumber="165">L. weddellia</emphasis>
<bibRefCitation id="EFB907D2A125FF8DFACBFAE5FA37A1CC" author="Hartman, O." box="[1292,1436,1405,1426]" pageId="8" pageNumber="165" pagination="125 - 223" refId="ref7600" refString="Hartman, O. (1978) Polychaeta from the Weddell Sea quadrant, Antarctica. Antarctic Research Series, 26 (4), 125 - 223." type="journal article" year="1978">Hartman, 1978</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

665
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<emphasis id="B95CA631A128FF80FF50FF0FFE22A4EF" bold="true" box="[151,393,151,177]" italics="true" pageId="5" pageNumber="162">Laonice ayseniensis</emphasis>
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n. sp
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<paragraph id="8B977A23A128FF80FF50FE9BFD7BA5D7" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
<emphasis id="B95CA631A128FF80FF50FE9BFE95A543" bold="true" box="[151,318,259,285]" pageId="5" pageNumber="162">Type material</emphasis>
.
<materialsCitation id="3B40707EA128FF80FE94FE9BFC0AA51C" collectedFrom="slightly gravelly sand" collectingDate="2013-12-10" collectionCode="MNHNCL" collectorName="S. Martinez" county="Pichicolo" elevation="30" location="Hualaihue" municipality="Palena" pageId="5" pageNumber="162" specimenCode="MNHNCL ANN-15311" specimenCount="1" stateProvince="Los Lagos region" typeStatus="holotype">
<collectingCounty id="62F602AFA128FF80FE94FE9BFE6BA543" box="[339,448,259,285]" pageId="5" pageNumber="162">Pichicolo</collectingCounty>
,
<collectingMunicipality id="6BF3E059A128FF80FE0CFE9BFDB2A543" box="[459,537,259,285]" pageId="5" pageNumber="162">Palena</collectingMunicipality>
,
<location id="8EF72CF8A128FF80FDE4FE9BFD32A543" LSID="urn:lsid:plazi:treatment:0381CB35A128FF8DFF50FF0FFC70A63F:8EF72CF8A128FF80FDE4FE9BFD32A543" box="[547,665,259,285]" county="Pichicolo" latitude="-44.316666" longLatPrecision="1137" longitude="-72.55" municipality="Palena" name="Hualaihue" pageId="5" pageNumber="162" stateProvince="Los Lagos region">Hualaihue</location>
,
<collectingRegion id="49ECB4C1A128FF80FD63FE9CFCC2A543" box="[676,873,259,286]" country="Chile" name="Los Lagos" pageId="5" pageNumber="162">Los Lagos region</collectingRegion>
<date id="FF965CE3A128FF80FCA8FE9BFC59A543" box="[879,1010,259,285]" pageId="5" pageNumber="162" value="2013-12-10">
<collectingDate id="EFD2A50BA128FF80FCA8FE9BFC59A543" box="[879,1010,259,285]" pageId="5" pageNumber="162" value="2013-12-10">10/12/2013</collectingDate>
</date>
,
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,
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, coll.
<collectorName id="26DD1FF5A128FF80FAD4FE9BFA33A543" box="[1299,1432,259,285]" pageId="5" pageNumber="162">S. Martínez</collectorName>
,
<collectedFrom id="383439DEA128FF80FF50FEBFFE21A51F" box="[151,394,295,321]" pageId="5" pageNumber="162">slightly gravelly sand</collectedFrom>
,
<elevation id="00059D10A128FF80FE53FEBFFDBFA51F" box="[404,532,295,322]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="162" unit="m" value="30.0">
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depth
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,
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(
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)
</materialsCitation>
and
<materialsCitation id="3B40707EA128FF80FC11FEBFFCDAA53B" collectionCode="MNHNCL" elevation="30" latitude="-44.316666" location="Canal Puyuhuapi" longLatPrecision="1137" longitude="-72.55" pageId="5" pageNumber="162" specimenCode="MNHNCL ANN-15395" specimenCount="1" stateProvince="Aysen Region" typeStatus="paratype">
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<typeStatus id="5493C481A128FF80FBCFFEB0FBD8A51C" box="[1032,1139,296,322]" pageId="5" pageNumber="162">paratypes</typeStatus>
(
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).
<location id="8EF72CF8A128FF80FF50FED3FEF2A53B" LSID="urn:lsid:plazi:treatment:0381CB35A128FF8DFF50FF0FFC70A63F:8EF72CF8A128FF80FF50FED3FEF2A53B" box="[151,345,331,357]" county="Pichicolo" latitude="-44.316666" longLatPrecision="1137" longitude="-72.55" municipality="Palena" name="Canal Puyuhuapi" pageId="5" pageNumber="162" stateProvince="Aysen Region">Canal Puyuhuapi</location>
,
<collectingRegion id="49ECB4C1A128FF80FEA6FED3FE56A53B" box="[353,509,331,357]" country="Chile" name="Aysen" pageId="5" pageNumber="162">Aysén Region</collectingRegion>
(
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),
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<quantity id="4CD0D7C6A128FF80FD35FED3FC81A538" box="[754,810,331,358]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="162" unit="m" value="30.0">30 m</quantity>
depth
</elevation>
</materialsCitation>
,
<materialsCitation id="3B40707EA128FF80FCBCFED3FD67A5D7" collectedFrom="gravelly-muddy sediments" collectingDate="2014-03-09" collectionCode="MNHNCL" collectorName="C. Carreno" pageId="5" pageNumber="162" specimenCode="MNHNCL ANN-15396" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D2EB1AAA128FF80FCBCFED3FC18A53B" box="[891,947,331,357]" pageId="5" pageNumber="162" type="generic">eight</specimenCount>
<typeStatus id="5493C481A128FF80FC7FFED4FB88A538" box="[952,1059,332,358]" pageId="5" pageNumber="162">paratypes</typeStatus>
(
<specimenCode id="DB8ED258A128FF80FBF6FED3FAEBA538" box="[1073,1344,331,358]" pageId="5" pageNumber="162">MNHNCL ANN-15396</specimenCode>
),
<date id="FF965CE3A128FF80FA95FED3FF79A5D7" pageId="5" pageNumber="162" value="2014-03-09">
<collectingDate id="EFD2A50BA128FF80FA95FED3FF79A5D7" pageId="5" pageNumber="162" value="2014-03-09">09-03- 2014</collectingDate>
</date>
, coll.
<collectorName id="26DD1FF5A128FF80FED2FEF7FE38A5D7" box="[277,403,367,393]" pageId="5" pageNumber="162">C. Carreño</collectorName>
,
<collectedFrom id="383439DEA128FF80FE59FEF7FD67A5D7" box="[414,716,367,393]" pageId="5" pageNumber="162">gravelly-muddy sediments</collectedFrom>
</materialsCitation>
.
</paragraph>
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<subSubSection id="C33229A8A128FF82FF00FE0BFBD1A48B" lastPageId="7" lastPageNumber="164" pageId="5" pageNumber="162" type="description">
<paragraph id="8B977A23A128FF80FF00FE0BFEC7A5A8" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
<emphasis id="B95CA631A128FF80FF00FE0BFEFAA5F3" bold="true" box="[199,337,403,429]" pageId="5" pageNumber="162">Description</emphasis>
.
<typeStatus id="5493C481A128FF80FEA5FE0BFE61A5F3" box="[354,458,403,429]" pageId="5" pageNumber="162" type="holotype">Holotype</typeStatus>
complete: 147 chaetigers, length
<quantity id="4CD0D7C6A128FF80FC96FE0BFC08A5F0" box="[849,931,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="162" unit="mm" value="48.0">48 mm</quantity>
and width
<quantity id="4CD0D7C6A128FF80FBE0FE0BFB29A5F0" box="[1063,1154,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="162" unit="mm" value="1.6">1.6 mm</quantity>
;
<typeStatus id="5493C481A128FF80FB57FE0CFB5BA5F0" box="[1168,1264,404,430]" pageId="5" pageNumber="162" type="paratype">paratype</typeStatus>
complete: 139 chaetigers, length
<quantity id="4CD0D7C6A128FF80FEA0FE2FFE1CA58C" box="[359,439,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="162" unit="mm" value="38.0">38 mm</quantity>
and width
<quantity id="4CD0D7C6A128FF80FDFEFE2FFD38A58C" box="[569,659,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="162" unit="mm" value="1.9">1.9 mm</quantity>
; 12 incomplete
<typeStatus id="5493C481A128FF80FC8EFE20FC12A58C" box="[841,953,440,466]" pageId="5" pageNumber="162" type="paratype">paratypes</typeStatus>
: 59–106 chaetigers, length
<quantity id="4CD0D7C6A128FF80FB36FE2FFAC7A58C" box="[1265,1388,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.6" metricValueMin="1.0" pageId="5" pageNumber="162" unit="mm" value="18.0" valueMax="26.0" valueMin="10.0">10–26 mm</quantity>
and width
<quantity id="4CD0D7C6A128FF80FF19FE43FEC3A5A8" box="[222,360,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="5" pageNumber="162" unit="mm" value="2.05" valueMax="2.5" valueMin="1.6">1.6–2.5 mm</quantity>
.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FE67FC20A6AF" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Prostomium and peristomium fused anteriorly, giving a bicornate, cap-shaped appearance (
<figureCitation id="131366A6A128FF80FB6CFE67FAA8A647" box="[1195,1283,511,537]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
); prostomium triangular in the unfused region of the prostomium and peristomium. Peristomium well developed and free from notopodial lamellae (
<figureCitation id="131366A6A128FF80FE48FDDFFE47A63F" box="[399,492,583,609]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2B</figureCitation>
). One pair of large brownish eyespots, thicker on the lateral region, giving the appearance of raccoon spots on the posterior part of the prostomium (
<figureCitation id="131366A6A128FF80FC6DFDF3FBAEA6DB" box="[938,1029,619,645]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
). Occipital tentacle short, thick, and positioned slightly beyond the notopodial lamellae. Caruncle well developed, extending together with the nuchal organs that start between the notopodium and the prostomium, and accompany the caruncle until chaetigers 14–16 (
<quantity id="4CD0D7C6A128FF80FF67FD4FFF71A6AF" box="[160,218,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="5" pageNumber="162" unit="in" value="16.0">16 in</quantity>
<typeStatus id="5493C481A128FF80FF27FD4FFEEFA6AF" box="[224,324,727,753]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) (
<figureCitation id="131366A6A128FF80FE9CFD4FFE1EA6AF" box="[347,437,727,753]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
). Palps absent in all specimens examined.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FD63FC2DA7B3" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Branchiae starting from chaetiger 2, continuing up to chaetigers 35–38 (chaetiger
<quantity id="4CD0D7C6A128FF80FB91FD63FB25A74B" box="[1110,1166,763,790]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="162" unit="in" value="35.0">35 in</quantity>
<typeStatus id="5493C481A128FF80FB53FD63FB51A74B" box="[1172,1274,763,789]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
), all branchiae basally free of the notopodial lamellae (
<figureCitation id="131366A6A128FF80FD94FC87FD4CA767" box="[595,743,799,826]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E, F, H</figureCitation>
). All branchiae digitiform, long, thick, with spear-shaped tips, and heavily ciliated; first two pairs of branchiae short (
<figureCitation id="131366A6A128FF80FCD6FCDBFC21A703" box="[785,906,835,861]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E, F</figureCitation>
); first pair slightly shorter than the notopodial postchaetal lamellae (
<figureCitation id="131366A6A128FF80FE41FCFFFE71A7DF" box="[390,474,871,897]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E</figureCitation>
), increasing in length significantly just after the third pair of notolamellae and becoming twice as long as the notopodial postchaetal lamellae from chaetiger 6 (
<figureCitation id="131366A6A128FF80FC64FC13FC57A7FB" box="[931,1020,907,933]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
). From chaetiger 10, the length of the longest branchiae equal to the total width of the body (
<figureCitation id="131366A6A128FF80FD36FC37FCC1A797" box="[753,874,943,969]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A, G</figureCitation>
), thereafter branchiae length gradually decreasing, the last three pairs so small that they are hidden in the parapodium.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FC6FFE44A16F" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Notopodial postchaetal lamellae of chaetiger 1 very small, subtriangular, shorter than neurolamellae and occipital nuchal tentacle (
<figureCitation id="131366A6A128FF80FE70FB83FDBAA06B" box="[439,529,1051,1077]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2D</figureCitation>
). From pairs 2–3, lamellae increasing in size to form a broad subtriangular shape (
<figureCitation id="131366A6A128FF80FF58FBA7FEB0A007" box="[159,283,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E, F</figureCitation>
); fourth pair and subsequent pairs even larger and wider, leaf-shaped, the fourth pair a third larger than the third pair, with the tips of the notopodial lamellae touching; fourth pair the largest notopodial lamellae, notopodial lamellae equal in size on chaetigers 13–14, thereafter gradually decreasing in size, becoming rounded from chaetiger 37; from around chaetiger 44 lamellae becoming triangular and gradually decreasing in size until the end of the body. Notopodial prechaetal lamellae rounded, moderately developed (
<figureCitation id="131366A6A128FF80FB92FB57FB5AA0B7" box="[1109,1265,1231,1257]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A, E–H</figureCitation>
), postbranchial lamellae reduced. Dorsal transverse ridges very low, connecting notopodial postchaetal lamellae on chaetigers 37– 48 (
<figureCitation id="131366A6A128FF80FF05FA8FFEB0A16F" box="[194,283,1303,1329]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2C</figureCitation>
), absent thereafter.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FAA3FD8BA1BB" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Neuropodial postchaetal lamellae of chaetiger 1 rectangular with rounded edges (
<figureCitation id="131366A6A128FF80FB98FAA3FB12A10B" box="[1119,1209,1339,1365]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2D</figureCitation>
), oval on chaetigers 2–8, with well developed ventral edges in the form of an earlobe, (
<figureCitation id="131366A6A128FF80FCA8FAC7FC4CA127" box="[879,999,1375,1401]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E–G</figureCitation>
), increasing gradually in size. Lamellae on chaetiger 9 without ventral lobe (
<figureCitation id="131366A6A128FF80FD84FA1BFD25A1C3" box="[579,654,1411,1437]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig 2I</figureCitation>
), and on chaetiger 25 lamellae subtriangular, wide, small, until the appearance of the ventrolateral pouches when they become rounded, gradually decreasing in size on posterior chaetigers, with low, rounded lobe.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FA77FD3BA20F" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Ventrolateral pouches starting from chaetigers 35–48 (
<quantity id="4CD0D7C6A128FF80FCE9FA77FCC1A257" box="[814,874,1519,1546]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="162" unit="in" value="35.0">35 in</quantity>
<typeStatus id="5493C481A128FF80FCB6FA77FC7EA257" box="[881,981,1519,1545]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) and continuing up to chaetiger 57. The first, very small, pouch emerges from the dorsal part of the prechaetal neuropodial lamellae and the posterior dorsal base of the postchaetal neuropodial lamellae.
</paragraph>
<paragraph id="8B977A23A128FF80FF00F9C3FA91A3CB" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
All anterior parapodia with capillaries arranged in two rows. Long, thin capillary chaetae in the upper part of the notopodium; notopodial chaetae from chaetiger 5 up to chaetiger 22 with chaetae arranged in two rows, anterior row with short, striate and bilimbate chaetae (
<figureCitation id="131366A6A128FF80FD5AF93BFD45A2E3" box="[669,750,1699,1725]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2J</figureCitation>
), posterior row with long, striate and bilimbate chaetae (
<figureCitation id="131366A6A128FF80FAA8F93BFF16A2BF" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2K</figureCitation>
); from chaetiger 23 capillary chaetae arranged in two rows, long, thin, striate unilimbate (
<figureCitation id="131366A6A128FF80FB61F95FFB56A2BF" box="[1190,1277,1735,1761]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2L</figureCitation>
). Neuropodial chaetae similar to notopodial chaetae, arranged in two rows, but shorter and thinner. Neuropodial hooks from chaetigers 35–38, (
<quantity id="4CD0D7C6A128FF80FEADF897FE08A377" box="[362,419,1807,1834]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="162" unit="in" value="35.0">35 in</quantity>
<typeStatus id="5493C481A128FF80FE6EF897FDA6A377" box="[425,525,1807,1833]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) with 6–12 hooks per fascicle; hooded hooks tridentate with two teeth on the main fang, giving a bidentate appearance in the lateral view, hoods with distal opening (
<figureCitation id="131366A6A128FF80FBEEF8ABFB0FA313" box="[1065,1188,1843,1869]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2N, O</figureCitation>
), companion capillary chaetae present (
<figureCitation id="131366A6A128FF80FE95F8CFFE0CA32F" box="[338,423,1879,1905]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2P</figureCitation>
). Sabre chaetae from chaetigers 24–28 (
<quantity id="4CD0D7C6A128FF80FCA2F8CFFC34A32F" box="[869,927,1879,1905]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="5" pageNumber="162" unit="in" value="28.0">28 in</quantity>
<typeStatus id="5493C481A128FF80FC62F8CFFBA2A32F" box="[933,1033,1879,1905]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) numbering up to three per fascicle; sabre chaetae heavily striate and granulate (
<figureCitation id="131366A6A128FF80FD47F8E3FD74A3CB" box="[640,735,1915,1941]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2M</figureCitation>
), from chaetiger 15 numbering up to two per fascicle.
</paragraph>
<paragraph id="8B977A23A128FF80FF00F807FDC1A383" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Pygidium with two small ventral papilliform cirri arranged closely to each other, and five long cirri: two on one side and three on the other side (
<figureCitation id="131366A6A128FF80FDC4F85BFDF6A383" box="[515,605,1987,2013]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2Q</figureCitation>
).
</paragraph>
<caption id="DF572AABA12BFF83FF50F963FE4EA3B5" ID-DOI="http://doi.org/10.5281/zenodo.14390223" ID-Zenodo-Dep="14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="6" pageNumber="163" startId="6.[151,250,1787,1811]" targetBox="[250,1337,190,1752]" targetPageId="6" targetType="figure">
<paragraph id="8B977A23A12BFF83FF50F963FE4EA3B5" blockId="6.[151,1437,1787,2027]" pageId="6" pageNumber="163">
<emphasis id="B95CA631A12BFF83FF50F963FEBEA34D" bold="true" box="[151,277,1787,1811]" pageId="6" pageNumber="163">FIGURE 2.</emphasis>
<taxonomicName id="4C2801A0A12BFF83FEDAF964FE41A34D" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[285,490,1788,1811]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="163" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12BFF83FEDAF964FE41A34D" box="[285,490,1788,1811]" italics="true" pageId="6" pageNumber="163">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12BFF83FE34F963FDEDA34D" bold="true" box="[499,582,1787,1811]" pageId="6" pageNumber="163">
<taxonomicNameLabel id="A26F1B4AA12BFF83FE34F963FDEDA34D" box="[499,582,1787,1811]" pageId="6" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp).
</paragraph>
</caption>
<paragraph id="8B977A23A12AFF82FF00FF0EFBD1A48B" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FF0EFE5FA4EE" bold="true" box="[199,500,150,177]" pageId="7" pageNumber="164">Staining in Methyl Green</emphasis>
. Peristomium stained profusely dorsally and ventrally. Anterior segments with outer margin of both notopodial and neuropodial postchaetal lamellae stained (
<figureCitation id="131366A6A12AFF82FC0EFF23FB8AA48B" box="[969,1057,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="7" pageNumber="164">Fig. 2A</figureCitation>
, A´, B).
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A12AFF82FF00FF47FE91A543" pageId="7" pageNumber="164" type="etymology">
<paragraph id="8B977A23A12AFF82FF00FF47FE91A543" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FF47FEE5A4A7" bold="true" box="[199,334,223,249]" pageId="7" pageNumber="164">Etymology.</emphasis>
The specific name
<taxonomicName id="4C2801A0A12AFF82FDEBFF47FD07A4A6" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[556,684,223,248]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis">
<emphasis id="B95CA631A12AFF82FDEBFF47FD07A4A6" box="[556,684,223,248]" italics="true" pageId="7" pageNumber="164">ayseniensis</emphasis>
</taxonomicName>
is derived from the name of a geographical zone in the
<collectingRegion id="49ECB4C1A12AFF82FAE1FF47FF40A543" country="Chile" name="Los Lagos" pageId="7" pageNumber="164">Los Lagos Region</collectingRegion>
,
<collectingCountry id="F33F3AB3A12AFF82FF31FE9BFE9DA543" box="[246,310,259,285]" name="Chile" pageId="7" pageNumber="164">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A12AFF82FF00FEBEFD6AA51F" box="[199,705,294,321]" pageId="7" pageNumber="164" type="distribution">
<paragraph id="8B977A23A12AFF82FF00FEBEFD6AA51F" blockId="7.[151,1437,150,2013]" box="[199,705,294,321]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FEBEFEF2A51E" bold="true" box="[199,345,294,320]" pageId="7" pageNumber="164">Distribution</emphasis>
Only known from
<typeStatus id="5493C481A12AFF82FDF7FEBFFDCBA51F" box="[560,608,295,321]" pageId="7" pageNumber="164">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A12AFF8DFF00FED3FC70A63F" lastPageId="8" lastPageNumber="165" pageId="7" pageNumber="164" type="discussion">
<paragraph id="8B977A23A12AFF82FF00FED3FB3EA647" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FED3FE90A53B" bold="true" box="[199,315,331,357]" pageId="7" pageNumber="164">Remarks.</emphasis>
The ‘
<emphasis id="B95CA631A12AFF82FEB8FED3FD9CA53B" box="[383,567,331,357]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FEB8FED3FE72A53A" authorityName="Malmgren" authorityYear="1867" box="[383,473,331,356]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="genus">Laonice</taxonomicName>
cirrata’
</emphasis>
group (
<bibRefCitation id="EFB907D2A12AFF82FD48FED3FC4CA53B" author="Sikorski, A. &amp; Pavlova, L. V." box="[655,999,331,357]" pageId="7" pageNumber="164" pagination="353 - 368" refId="ref8162" refString="Sikorski, A. &amp; Pavlova, L. V. (2016) Three new species of Laonice (Polychaete: Spionidae) from West and Southwest Africa. Zootaxa, 4097 (3), 353 - 368. https: // doi. org / 10.11646 / zootaxa. 4097.3.4" type="journal article" year="2016">Sikorski &amp; Pavlova 2016: 361</bibRefCitation>
) can easily be distinguished by having a prostomium clearly fused with the peristomium on the anterior margin; capillaries on the anterior part of the body arranged in two vertical rows; and by having quite variable numerical characters. Nine species belong to this group:
<taxonomicName id="4C2801A0A12AFF82FF20FE2FFDAEA58F" authority="(M. Sars, 1851)" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[231,517,438,465]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FF20FE2FFEF9A58E" box="[231,338,439,464]" italics="true" pageId="7" pageNumber="164">L. cirrata</emphasis>
(M.Sars, 1851)
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FDD7FE2FFCC2A58F" authority="(Kinberg, 1866)" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[528,873,439,465]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A12AFF82FDD7FE2FFD07A58F" box="[528,684,439,465]" italics="true" pageId="7" pageNumber="164">L. brevicornis</emphasis>
(Kinberg, 1866)
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FCB2FE2FFB4EA58F" authority="Soderstrom, 1920" authorityName="Soderstrom" authorityYear="1920" box="[885,1253,439,465]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FCB2FE2FFBBEA58F" box="[885,1045,439,465]" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
Söderström, 1920
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FB36FE2FFEB4A5AB" authority="Pillai, 1961" authorityName="Pillai" authorityYear="1961" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicristata">
<emphasis id="B95CA631A12AFF82FB36FE2FFA37A58F" box="[1265,1436,439,465]" italics="true" pageId="7" pageNumber="164">L. brevicristata</emphasis>
Pillai, 1961
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FEE9FE43FD13A5AB" authority="Nonato et al., 1986" authorityName="Nonato" authorityYear="1986" box="[302,696,475,501]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="branchiata">
<emphasis id="B95CA631A12AFF82FEE9FE43FE61A5AB" box="[302,458,475,501]" italics="true" pageId="7" pageNumber="164">L. branchiata</emphasis>
Nonato
<emphasis id="B95CA631A12AFF82FDF7FE43FDCEA5AB" box="[560,613,475,501]" italics="true" pageId="7" pageNumber="164">et al</emphasis>
., 1986
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FD00FE43FBEBA5AB" authority="Sikorski, 2003" authorityName="Sikorski" authorityYear="2003" box="[711,1088,475,501]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="shamrockensis">
<emphasis id="B95CA631A12AFF82FD00FE43FD7CA5AA" box="[711,727,475,500]" italics="true" pageId="7" pageNumber="164">L</emphasis>
.
<emphasis id="B95CA631A12AFF82FD2FFE43FC27A5AB" box="[744,908,475,501]" italics="true" pageId="7" pageNumber="164">shamrockensis</emphasis>
Sikorski, 2003
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FB88FE43FEC7A647" authority="Sigvaldadottir &amp; Desbruyeres, 2003" authorityName="Sigvaldadottir &amp; Desbruyeres" authorityYear="2003" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="asaccata">
<emphasis id="B95CA631A12AFF82FB88FE43FB7FA5AA" box="[1103,1236,475,500]" italics="true" pageId="7" pageNumber="164">L. asaccata</emphasis>
Sigvaldadottir &amp; Desbruyeres, 2003
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FEBFFE67FD0EA647" authority="Sikorski, 2011" authorityName="Sikorski" authorityYear="2011" box="[376,677,511,537]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FEBFFE67FE52A647" box="[376,505,511,537]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
<bibRefCitation id="EFB907D2A12AFF82FDC7FE67FD0EA647" author="Sikorski, A. V." box="[512,677,511,537]" pageId="7" pageNumber="164" pagination="201 - 214" refId="ref8104" refString="Sikorski, A. V. (2011) Review of Laonice (Spionidae, Annelida) with remarks on several species and a description of a new species from South Africa. Italian Journal of Zoology, 78 (S 1), 201 - 214. https: // doi. org / 10.1080 / 11250003.2011.617218" type="journal article" year="2011">Sikorski, 2011</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C2801A0A12AFF82FD26FE67FB3AA647" authority="Sikorski &amp; Pavlova, 2016" authorityName="Sikorski &amp; Pavlova" authorityYear="2016" box="[737,1169,511,537]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cricketae">
<emphasis id="B95CA631A12AFF82FD26FE67FCCEA647" box="[737,869,511,537]" italics="true" pageId="7" pageNumber="164">L. cricketae</emphasis>
<bibRefCitation id="EFB907D2A12AFF82FCABFE67FB3AA647" author="Sikorski, A. &amp; Pavlova, L. V." box="[876,1169,511,537]" pageId="7" pageNumber="164" pagination="353 - 368" refId="ref8162" refString="Sikorski, A. &amp; Pavlova, L. V. (2016) Three new species of Laonice (Polychaete: Spionidae) from West and Southwest Africa. Zootaxa, 4097 (3), 353 - 368. https: // doi. org / 10.11646 / zootaxa. 4097.3.4" type="journal article" year="2016">Sikorski &amp; Pavlova, 2016</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FDBBFE04A04F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00FDBBFE01A662" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,426,547,572]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00FDBBFE01A662" box="[199,426,547,572]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE74FDBBFDA5A663" bold="true" box="[435,526,547,573]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE74FDBBFDA5A663" box="[435,526,547,573]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, belongs to the ‘
<emphasis id="B95CA631A12AFF82FD16FDBBFC20A663" box="[721,907,547,573]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FD16FDBBFC80A662" authorityName="Malmgren" authorityYear="1867" box="[721,811,547,572]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="genus">Laonice</taxonomicName>
cirrata’
</emphasis>
group due to the anteriorly fused prostomium and peristomium, the arrangement of the capillaries in the anterior part of the body in two vertical rows. However,
<taxonomicName id="4C2801A0A12AFF82FF50FDF3FED3A6DA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,376,619,644]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF50FDF3FED3A6DA" box="[151,376,619,644]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FEB8FDF3FE72A6DB" bold="true" box="[383,473,619,645]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FEB8FDF3FE72A6DB" box="[383,473,619,645]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from all other species by having the nuchal organs and caruncle extending up to chaetigers 14–16 (in other species the nuchal organs and caruncle are either shorter (
<taxonomicName id="4C2801A0A12AFF82FB2DFD17FA3FA6F7" authorityName="Pillai" authorityYear="1961" box="[1258,1428,655,681]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicristata">
<emphasis id="B95CA631A12AFF82FB2DFD17FA3FA6F7" box="[1258,1428,655,681]" italics="true" pageId="7" pageNumber="164">L. brevicristata</emphasis>
</taxonomicName>
: to ch. 7–9,
<taxonomicName id="4C2801A0A12AFF82FED0FD2BFE05A693" authorityName="Nonato, Bolívar &amp; Lana" authorityYear="1986" box="[279,430,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="branchiate">
<emphasis id="B95CA631A12AFF82FED0FD2BFE05A693" box="[279,430,691,717]" italics="true" pageId="7" pageNumber="164">L. branchiate</emphasis>
</taxonomicName>
: ch. 12,
<taxonomicName id="4C2801A0A12AFF82FDD5FD2BFD7FA693" authorityName="Sikorski" authorityYear="2003" box="[530,724,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="shamrockensis">
<emphasis id="B95CA631A12AFF82FDD5FD2BFD7FA693" box="[530,724,691,717]" italics="true" pageId="7" pageNumber="164">L. shamrockensis</emphasis>
</taxonomicName>
: ch. 5,
<taxonomicName id="4C2801A0A12AFF82FCEEFD2BFC00A692" authorityName="Sigvaldadottir &amp; Desbruyeres" authorityYear="2003" box="[809,939,691,716]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="asaccata">
<emphasis id="B95CA631A12AFF82FCEEFD2BFC00A692" box="[809,939,691,716]" italics="true" pageId="7" pageNumber="164">L. asaccata</emphasis>
</taxonomicName>
: ch. 4,
<taxonomicName id="4C2801A0A12AFF82FBC7FD2BFB2DA693" authorityName="Sikorski &amp; Pavlova" authorityYear="2016" box="[1024,1158,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cricketae">
<emphasis id="B95CA631A12AFF82FBC7FD2BFB2DA693" box="[1024,1158,691,717]" italics="true" pageId="7" pageNumber="164">L. cricketae</emphasis>
</taxonomicName>
: caruncle to ch. 1–2 and nuchal organs to ch. 20), or longer (
<taxonomicName id="4C2801A0A12AFF82FDF3FD4FFD0AA6AE" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[564,673,727,752]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FDF3FD4FFD0AA6AE" box="[564,673,727,752]" italics="true" pageId="7" pageNumber="164">L. cirrata</emphasis>
</taxonomicName>
: to ch. 4–40,
<taxonomicName id="4C2801A0A12AFF82FC85FD4FFC74A6AF" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[834,991,727,753]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A12AFF82FC85FD4FFC74A6AF" box="[834,991,727,753]" italics="true" pageId="7" pageNumber="164">L. brevicornis</emphasis>
</taxonomicName>
: ch. 9–26,
<taxonomicName id="4C2801A0A12AFF82FBA5FD4FFAAFA6AF" authorityName="Soderstrom" authorityYear="1920" box="[1122,1284,727,753]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FBA5FD4FFAAFA6AF" box="[1122,1284,727,753]" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
</taxonomicName>
: ch. 6–35,
<taxonomicName id="4C2801A0A12AFF82FA41FD4FFF50A74B" authorityName="Sikorski" authorityYear="2011" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FA41FD4FFF50A74B" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
: ch. 34–38). In addition, the notopodial postchaetal lamellae of chaetiger 1 are very small, subtriangular, shorter than the neurolamellae; the dorsal transverse ridges connecting the notopodial postchaetal lamellae on chaetigers 37–48 are very low; the neuropodial postchaetal lamellae of chaetiger 1 are rectangular with rounded edges, oval on chaetigers 2–8 with earlobe-shaped and well developed ventral edges; the ventrolateral pouches appear from chaetigers 35–48 up to chaetiger 57 (in the other species mentioned, these pouches start on much earlier chaetigers: between chaetigers 3–17, or are absent, except in
<taxonomicName id="4C2801A0A12AFF82FC89FC37FC78A797" authorityName="Sikorski &amp; Pavlova" authorityYear="2016" box="[846,979,943,969]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cricketae">
<emphasis id="B95CA631A12AFF82FC89FC37FC78A797" box="[846,979,943,969]" italics="true" pageId="7" pageNumber="164">L. cricketae</emphasis>
</taxonomicName>
where they start from chaetiger 40); the hooded hooks have a distal opening; and the pygidium has two small ventral papilliform cirri arranged close to each other, and five long cirri.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FB83FB23A023" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00FB83FE07A06A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,428,1051,1076]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00FB83FE07A06A" box="[199,428,1051,1076]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE72FB83FDB9A06B" bold="true" box="[437,530,1051,1077]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE72FB83FDB9A06B" box="[437,530,1051,1077]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is most similar to
<emphasis id="B95CA631A12AFF82FD3EFB83FB66A06B" box="[761,1229,1051,1077]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FD3EFB83FCC1A06A" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[761,874,1051,1076]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">L. cirrata</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FCBEFB83FBB0A06B" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[889,1051,1051,1077]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">L. brevicornis</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FBEEFB83FB66A06B" authorityName="Soderstrom" authorityYear="1920" box="[1065,1229,1051,1077]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">L. bahusiensis</taxonomicName>
</emphasis>
, and
<emphasis id="B95CA631A12AFF82FAD6FB83FA37A06B" box="[1297,1436,1051,1077]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FAD6FB83FA33A06B" authorityName="Sikorski" authorityYear="2011" box="[1297,1432,1051,1077]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">L. antipoda</taxonomicName>
,
</emphasis>
because they also have a long caruncle and nuchal organs. However,
<taxonomicName id="4C2801A0A12AFF82FC54FBA7FBD8A006" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[915,1139,1087,1112]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FC54FBA7FBD8A006" box="[915,1139,1087,1112]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FBBEFBA7FB7AA007" bold="true" box="[1145,1233,1087,1113]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FBBEFBA7FB7AA007" box="[1145,1233,1087,1113]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from these species by the morphological characters mentioned above, and by those mentioned below.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FB1FFEB4A19F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00FB1FFE0DA0FE" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,422,1159,1184]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00FB1FFE0DA0FE" box="[199,422,1159,1184]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE6CFB1FFDA8A0FF" bold="true" box="[427,515,1159,1185]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE6CFB1FFDA8A0FF" box="[427,515,1159,1185]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from
<emphasis id="B95CA631A12AFF82FCF7FB1FFB40A0FF" box="[816,1259,1159,1185]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FCF7FB1FFC37A0FE" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[816,924,1159,1184]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">L. cirrata</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FC61FB1FFBE9A0FF" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[934,1090,1159,1185]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">L. brevicornis</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FB8BFB1FFB40A0FF" authorityName="Soderstrom" authorityYear="1920" box="[1100,1259,1159,1185]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">L. bahusiensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C2801A0A12AFF82FADAFB1FFA36A0FF" authorityName="Sikorski" authorityYear="2011" box="[1309,1437,1159,1185]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FADAFB1FFA36A0FF" box="[1309,1437,1159,1185]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
by the shape and number of eyes, that in
<taxonomicName id="4C2801A0A12AFF82FDA6FB33FCE9A09A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[609,834,1195,1220]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FDA6FB33FCE9A09A" box="[609,834,1195,1220]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FC8FFB33FC09A09B" bold="true" box="[840,930,1195,1221]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FC8FFB33FC09A09B" box="[840,930,1195,1221]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, have the appearance of raccoon spots on the back of the prostomium; and by the last three pairs of branchiae that are so small, they are hidden in the parapodium. In addition,
<taxonomicName id="4C2801A0A12AFF82FEEFFB6BFDA6A152" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[296,525,1267,1292]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FEEFFB6BFDA6A152" box="[296,525,1267,1292]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FDDFFB6BFDDDA153" bold="true" box="[536,630,1267,1293]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FDDFFB6BFDDDA153" box="[536,630,1267,1293]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C2801A0A12AFF82FCD0FB6BFC2CA152" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[791,903,1267,1292]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FCD0FB6BFC2CA152" box="[791,903,1267,1292]" italics="true" pageId="7" pageNumber="164">L. cirrata</emphasis>
</taxonomicName>
by not having the dorsolateral margin of the notopodial postchaetal lamellae forming a small peak, but having small, triangular posterior notopodial lamellae.
<taxonomicName id="4C2801A0A12AFF82FF50FAA3FED3A10A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,376,1339,1364]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF50FAA3FED3A10A" box="[151,376,1339,1364]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FEB9FAA3FE7CA10B" bold="true" box="[382,471,1339,1365]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FEB9FAA3FE7CA10B" box="[382,471,1339,1365]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C2801A0A12AFF82FCC2FAA3FC0AA10B" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[773,929,1339,1365]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A12AFF82FCC2FAA3FC0AA10B" box="[773,929,1339,1365]" italics="true" pageId="7" pageNumber="164">L. brevicornis</emphasis>
</taxonomicName>
by short first two pair of branchiae which are slightly shorter than the notopodial postchaetal lamellae; the presence of dorsal transverse ridges connecting the notopodial postchaetal lamellae; and because the main fang of the hooded hooks is not accompanied at the base by apical teeth.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FA53FEF7A35B" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
In addition,
<taxonomicName id="4C2801A0A12AFF82FE96FA53FD98A1BA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[337,563,1483,1508]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FE96FA53FD98A1BA" box="[337,563,1483,1508]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FDFDFA53FD3EA1BB" bold="true" box="[570,661,1483,1509]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FDFDFA53FD3EA1BB" box="[570,661,1483,1509]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C2801A0A12AFF82FC0EFA53FBC0A1BB" authorityName="Soderstrom" authorityYear="1920" box="[969,1131,1483,1509]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FC0EFA53FBC0A1BB" box="[969,1131,1483,1509]" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2801A0A12AFF82FB65FA53FA8EA1BB" authorityName="Sikorski" authorityYear="2011" box="[1186,1317,1483,1509]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FB65FA53FA8EA1BB" box="[1186,1317,1483,1509]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
by having a prostomium that is broadly rounded anteriorly, bicornate as a cap, instead of triangular, anteriorly wide, truncated or slightly incurved, sometimes broadly rounded; sabre chaetae from chaetigers 24–28
<emphasis id="B95CA631A12AFF82FB44F98BFB09A272" box="[1155,1186,1555,1580]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
from ch.
<quantity id="4CD0D7C6A12AFF82FAD0F98BFAD7A273" box="[1303,1404,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="5.334" metricValueMin="2.54" pageId="7" pageNumber="164" unit="in" value="15.5" valueMax="21.0" valueMin="10.0">10–21 in</quantity>
<emphasis id="B95CA631A12AFF82FA41F98BFE8AA20F" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FA41F98BFEB7A20F" authorityName="Soderstrom" authorityYear="1920" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">L. bahusiensis</taxonomicName>
,
</emphasis>
and ch.
<quantity id="4CD0D7C6A12AFF82FE4AF9AFFE5EA20F" box="[397,501,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.826" metricValueMin="3.556" pageId="7" pageNumber="164" unit="in" value="16.5" valueMax="19.0" valueMin="14.0">14–19 in</quantity>
<emphasis id="B95CA631A12AFF82FDC7F9AFFCD5A20E" box="[512,894,1591,1617]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FDC7F9AFFD22A20F" authorityName="Sikorski" authorityYear="2011" box="[512,649,1591,1617]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">L. antipoda</taxonomicName>
.
<taxonomicName id="4C2801A0A12AFF82FD5FF9AFFCD5A20E" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[664,894,1591,1616]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">Laonice ayseniensis</taxonomicName>
</emphasis>
<emphasis id="B95CA631A12AFF82FC4EF9AFFC4CA20F" bold="true" box="[905,999,1591,1617]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FC4EF9AFFC4CA20F" box="[905,999,1591,1617]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C2801A0A12AFF82FA41F9AFFEB1A22B" authorityName="Soderstrom" authorityYear="1920" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FA41F9AFFEB1A22B" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
</taxonomicName>
in having hooded hooks from chaetigers 35–38
<emphasis id="B95CA631A12AFF82FCEFF9C3FCECA22A" box="[808,839,1627,1652]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
from ch. 14–35, and by the number of teeth (tridentate with two teeth on the main fang, appearing bidentate in the lateral view
<emphasis id="B95CA631A12AFF82FC7CF9E7FC71A2C6" box="[955,986,1663,1688]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
five teeth (tridentate in the lateral view) with very small upper apical teeth.
<taxonomicName id="4C2801A0A12AFF82FDEFF93BFCA1A2E2" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[552,778,1699,1724]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FDEFF93BFCA1A2E2" box="[552,778,1699,1724]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FCD5F93BFCC7A2E3" bold="true" box="[786,876,1699,1725]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FCD5F93BFCC7A2E3" box="[786,876,1699,1725]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C2801A0A12AFF82FB31F93BFAD2A2E3" authorityName="Sikorski" authorityYear="2011" box="[1270,1401,1699,1725]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FB31F93BFAD2A2E3" box="[1270,1401,1699,1725]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
by the presence of nuchal tentacle, fewer pairs of branchiae (34–37
<emphasis id="B95CA631A12AFF82FCBAF95FFC30A2BE" box="[893,923,1735,1760]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
38–45); and very low (
<emphasis id="B95CA631A12AFF82FB6BF95FFB61A2BE" box="[1196,1226,1735,1760]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
prominent) dorsal transverse ridges.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00F897FD87A32F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
Hartmann-Schröder (1965) recorded three species for
<collectingCountry id="F33F3AB3A12AFF82FCF1F897FCDCA377" box="[822,887,1807,1833]" name="Chile" pageId="7" pageNumber="164">Chile</collectingCountry>
:
<taxonomicName id="4C2801A0A12AFF82FC43F897FB9DA376" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[900,1078,1807,1832]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FC43F897FB9DA376" box="[900,1078,1807,1832]" italics="true" pageId="7" pageNumber="164">Laonice cirrata</emphasis>
</taxonomicName>
, and two subspecies
<taxonomicName id="4C2801A0A12AFF82FAE9F897FEB8A312" authorityName="Hartmann-Schroder" authorityYear="1965" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="subSpecies" species="cirrata" subSpecies="postcirrata">
<emphasis id="B95CA631A12AFF82FAE9F897FEB8A312" italics="true" pageId="7" pageNumber="164">L. cirrata postcirrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2801A0A12AFF82FE8AF8ABFDE8A312" box="[333,579,1843,1868]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="subSpecies" species="cirrata" subSpecies="praecirrata">
<emphasis id="B95CA631A12AFF82FE8AF8ABFDE8A312" box="[333,579,1843,1868]" italics="true" pageId="7" pageNumber="164">L. cirrata praecirrata</emphasis>
</taxonomicName>
. The first species is already discussed above; and the other two species are very different from the new species.
</paragraph>
<paragraph id="8B977A23A12AFF8DFF00F8E3FDB9A663" blockId="7.[151,1437,150,2013]" lastBlockId="8.[151,1437,151,609]" lastPageId="8" lastPageNumber="165" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00F8E3FE02A3CA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,425,1915,1940]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00F8E3FE02A3CA" box="[199,425,1915,1940]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE77F8E3FDA1A3CB" bold="true" box="[432,522,1915,1941]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE77F8E3FDA1A3CB" box="[432,522,1915,1941]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C2801A0A12AFF82FCFBF8E3FBB2A3CA" box="[828,1049,1915,1940]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A12AFF82FCFBF8E3FBB2A3CA" box="[828,1049,1915,1940]" italics="true" pageId="7" pageNumber="164">Laonice postcirrata</emphasis>
</taxonomicName>
in that the former has an occipital tentacle situated slightly forward of the first pair of notopodial lamellae, whilst
<taxonomicName id="4C2801A0A12AFF82FBC9F807FB0CA3E6" box="[1038,1191,1951,1976]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A12AFF82FBC9F807FB0CA3E6" box="[1038,1191,1951,1976]" italics="true" pageId="7" pageNumber="164">L. postcirrata</emphasis>
</taxonomicName>
has a tentacle located between the first and second pair of notopodial lamellae; the prostomium and peristomium are completely fused in the anterior part of the prostomium, unlike
<taxonomicName id="4C2801A0A125FF8DFDB6FF0FFCA1A4EE" box="[625,778,151,176]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFDB6FF0FFCA1A4EE" box="[625,778,151,176]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
where they are not fused. Another difference is in the shape and number of eyes:
<taxonomicName id="4C2801A0A125FF8DFEB9FF23FDF5A48A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[382,606,187,212]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFEB9FF23FDF5A48A" box="[382,606,187,212]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFDA3FF23FD17A48B" bold="true" box="[612,700,187,213]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFDA3FF23FD17A48B" box="[612,700,187,213]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, has the appearance of raccoon spots on the posterior region of the prostomium, whereas
<taxonomicName id="4C2801A0A125FF8DFE4AFF47FD8DA4A6" box="[397,550,223,248]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE4AFF47FD8DA4A6" box="[397,550,223,248]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
has two or four small eyes arranged in the same row, or in a square or trapezoid.
<taxonomicName id="4C2801A0A125FF8DFF50FE9BFEDCA542" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,375,259,284]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFF50FE9BFEDCA542" box="[151,375,259,284]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFEBAFE9BFE7DA543" bold="true" box="[381,470,259,285]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFEBAFE9BFE7DA543" box="[381,470,259,285]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, also has a caruncle and branchiae that reach chaetigers 14–16 and 35–38
<emphasis id="B95CA631A125FF8DFACDFE9BFA82A542" box="[1290,1321,259,284]" italics="true" pageId="8" pageNumber="165">vs.</emphasis>
ch. 22–28 and 42, respectively, in
<taxonomicName id="4C2801A0A125FF8DFE63FEBFFD95A51E" box="[420,574,295,320]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE63FEBFFD95A51E" box="[420,574,295,320]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
. Furthermore, the shape of the anterior notopodial and neuropodial lamellae are different: in
<taxonomicName id="4C2801A0A125FF8DFE91FED3FD90A53A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[342,571,331,356]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFE91FED3FD90A53A" box="[342,571,331,356]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFD82FED3FD09A53B" bold="true" box="[581,674,331,357]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFD82FED3FD09A53B" box="[581,674,331,357]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the notopodial lamellae have a broad subtriangular shape with a blunt tip, and the postchaetal neuropodial lamellae on chaetigers 2–8 are oval with well-developed, earlobe-shaped ventral edges, whilst in
<taxonomicName id="4C2801A0A125FF8DFE61FE0BFDEAA5F2" box="[422,577,403,428]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE61FE0BFDEAA5F2" box="[422,577,403,428]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
the notopodial lamellae have pointed tips, and the neuropodial lamellae are wide and subtriangular. Ventrolateral pouches appear from from chaetigers
<quantity id="4CD0D7C6A125FF8DFC2BFE2EFBFBA58F" box="[1004,1104,438,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.2192" metricValueMin="0.889" pageId="8" pageNumber="165" unit="in" value="41.5" valueMax="48.0" valueMin="35.0">35–48 in</quantity>
<taxonomicName id="4C2801A0A125FF8DFB9FFE2FFA91A58E" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[1112,1338,439,464]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFB9FFE2FFA91A58E" box="[1112,1338,439,464]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFA85FE2FFA37A58F" bold="true" box="[1346,1436,439,465]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFA85FE2FFA37A58F" box="[1346,1436,439,465]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B95CA631A125FF8DFF50FE43FF1DA5AA" box="[151,182,475,500]" italics="true" pageId="8" pageNumber="165">vs.</emphasis>
from ch.
<quantity id="4CD0D7C6A125FF8DFEEDFE42FE25A5AB" box="[298,398,474,501]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.81" metricValueMin="2.794" pageId="8" pageNumber="165" unit="in" value="13.0" valueMax="15.0" valueMin="11.0">11–15 in</quantity>
<taxonomicName id="4C2801A0A125FF8DFE50FE43FCEEA5AB" authority="(Sirkorski et al., 2016)" baseAuthorityName="Sirkorski" baseAuthorityYear="2016" box="[407,837,475,501]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE50FE43FD98A5AA" box="[407,563,475,500]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
(Sirkorski
<emphasis id="B95CA631A125FF8DFD71FE43FD5AA5AB" box="[694,753,475,501]" italics="true" pageId="8" pageNumber="165">et al.</emphasis>
, 2016)
</taxonomicName>
, and finally,
<taxonomicName id="4C2801A0A125FF8DFC1BFE43FB6BA5AA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[988,1216,475,500]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFC1BFE43FB6BA5AA" box="[988,1216,475,500]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFB0EFE43FA8FA5AB" bold="true" box="[1225,1316,475,501]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFB0EFE43FA8FA5AB" box="[1225,1316,475,501]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has transdorsal ridges, and the neuropodial hooks and sabre chaetae appear from chaetigers 35–38 and 24–28
<emphasis id="B95CA631A125FF8DFAC4FE67FA8AA646" box="[1283,1313,511,536]" italics="true" pageId="8" pageNumber="165">vs.</emphasis>
ch. 31 and 16, respectively, in
<taxonomicName id="4C2801A0A125FF8DFEB6FDBBFDA0A662" box="[369,523,547,572]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFEB6FDBBFDA0A662" box="[369,523,547,572]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B977A23A125FF8DFF00FDDFFC70A63F" blockId="8.[151,1437,151,609]" box="[199,987,583,609]" pageId="8" pageNumber="165">Differences compared to other similar species can be found in the key.</paragraph>
</subSubSection>
</treatment>
</document>

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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9ABB5EF19F3C04F35D46B2591CCBD91C">
<mods:url id="3957C10A1B4F1C255AF8D5E995986B93">https://doi.org/10.11646/zootaxa.5551.1.8</mods:url>
</mods:location>
<mods:classification id="ABC038802E0D145BA1EDB8F247E0D6BF">journal article</mods:classification>
<mods:identifier id="2FEFFDC399104611376D3AEC654F77DB" type="DOI">10.11646/zootaxa.5551.1.8</mods:identifier>
<mods:identifier id="825950B32349B062D9E2C900BAA45B3C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B1D02296DF2477385FE965EDFF1211CE" type="Zenodo-Dep">14390234</mods:identifier>
<mods:identifier id="961326DC5ECDF9DFF38813BA0B113FE2" type="ZooBank">404750A6-70AD-4C2E-BC37-C7E49FEFF3EB</mods:identifier>
</mods:mods>
<treatment id="03E6A70A7553FFF8FF4A3997E6DAF889" LSID="urn:lsid:plazi:treatment:03E6A70A7553FFF8FF4A3997E6DAF889" httpUri="http://treatment.plazi.org/id/03E6A70A7553FFF8FF4A3997E6DAF889" lastPageId="17" lastPageNumber="184" pageId="14" pageNumber="181">
<subSubSection id="C35545977553FFE7FF4A3997E3F2FB52" box="[151,483,1219,1246]" pageId="14" pageNumber="181" type="nomenclature">
<paragraph id="8BF0161C7553FFE7FF4A3997E3F2FB52" blockId="14.[151,483,1219,1282]" box="[151,483,1219,1246]" pageId="14" pageNumber="181">
<heading id="D0B8A1707553FFE7FF4A3997E3F2FB52" bold="true" box="[151,483,1219,1246]" fontSize="11" level="1" pageId="14" pageNumber="181" reason="1">
<emphasis id="B93BCA0E7553FFE7FF4A3997E3F2FB52" bold="true" box="[151,483,1219,1246]" pageId="14" pageNumber="181">
<taxonomicName id="4C4F6D9F7553FFE7FF4A3997E36FFB52" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[151,382,1219,1246]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="181" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B93BCA0E7553FFE7FF4A3997E36FFB52" bold="true" box="[151,382,1219,1246]" italics="true" pageId="14" pageNumber="181">Sternacutus similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20877757553FFE7FE583990E3F2FB52" box="[389,483,1220,1246]" pageId="14" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35545977553FFF8FF4A39B3E712FCB5" lastPageId="17" lastPageNumber="184" pageId="14" pageNumber="181" type="description">
<paragraph id="8BF0161C7553FFE7FF4A39B3E336FA8E" blockId="14.[151,483,1219,1282]" box="[151,295,1255,1282]" pageId="14" pageNumber="181">
(
<figureCitation id="13740A997553FFE7FF4239B3E30EFA8E" box="[159,287,1255,1282]" captionStart="FIGURES 27–31" captionStartId="15.[152,270,1864,1888]" captionTargetBox="[159,1428,189,1832]" captionTargetId="figure-28@15.[151,1435,181,1840]" captionTargetPageId="15" captionText="FIGURES 27–31. Sternacutus similis sp. nov., holotype male: 27) Dorsal habitus; 28) Ventral habitus; 29) Lateral habitus; 30) Head, frontal view; 31) Centrobasal crests of elytra." figureDoi="http://doi.org/10.5281/zenodo.14390254" httpUri="https://zenodo.org/record/14390254/files/figure.png" pageId="14" pageNumber="181">Figs 27–31</figureCitation>
)
</paragraph>
<paragraph id="8BF0161C7553FFE7FF4A387BE1EAF8EA" blockId="14.[151,1437,1327,2038]" pageId="14" pageNumber="181">
<emphasis id="B93BCA0E7553FFE7FF4A387BE3FBFAC6" bold="true" box="[151,490,1327,1354]" pageId="14" pageNumber="181">
Description.
<typeStatus id="54F4A8BE7553FFE7FEEF3864E38EFAC6" box="[306,415,1328,1354]" pageId="14" pageNumber="181" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Head capsule dark brown, except gulamentum brown anteriorly, gradually lighter toward prothorax; ventral mouthparts orangish; anteclypeus and labrum mostly testaceous; scape dark brown, slightly lighter on apical quarter; pedicel orangish brown basally, brown on remaining surface; antennomere III light brown on basal 2/3, blackish on apical third; antennomeres IV–V yellowish brown on basal half, blackish on apical half; antennomeres VI–X orange on basal third, dark brown on apical 2/3; antennomere XI orange on basal half, dark brown on apical half. Pronotum and sides of prothorax blackish; prosternum brown laterally, light reddish brown centrally; prosternal process light reddish brown except dark brown apex. Ventral surface of meso- and metathorax dark brown laterally, dark reddish brown centrally. Scutellum brown centrally, dark brown laterally. Elytra dark brown, except brown base and blackish remaining basal quarter. Pro- and mesocoxae reddish brown, gradually orange toward trochanter; metacoxae dark brown on outer surface, orange on inner surface; trochanters and femoral clubs orange; pro- and mesofemoral clubs mostly dark brown; metafemoral club mostly brown; protibia reddish brown on basal half, blackish on apical half; meso- and metatibiae brown basally, reddish brown on remaining basal half, dark brown on apical half; pro- and mesotarsomeres I reddish brown on basal half, dark brown on apical half; metatarsomere I orange on basal half, blackish on apical half; tarsomeres III–V dark brown; tarsomeres V reddish brown on basal 2/3, more orangish on metatarsomere V, dark brown on apical third. Abdominal ventrites reddish brown, gradually orangish toward ventrite 5, with irregular dark brown areas.
</paragraph>
<paragraph id="8BF0161C7553FFF9FF1A3A24E36CFB9D" blockId="14.[151,1437,1327,2038]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="183" pageId="14" pageNumber="181">
<emphasis id="B93BCA0E7553FFE7FF1A3A24E31FF806" bold="true" box="[199,270,1904,1930]" pageId="14" pageNumber="181">Head.</emphasis>
Frons abundantly, minutely punctate; with abundant yellowish pubescence not obscuring integument, except glabrous median groove, pubescence yellower close to eyes; with one long, erect brown seta near eyes. Area between antennal tubercles with dense pale-yellow pubescence centrally, except glabrous median groove; remaining surface of vertex with moderately abundant pubescence not obscuring integument, pubescence pale-yellow centrally between eyes, gradually yellowish-brown toward prothorax, except dense yellow pubescence close to eyes, and glabrous median groove and narrow V-shaped transverse band between antennal tubercles and eyes. Area behind upper eye lobes with moderately abundant pale-yellow pubescence not obscuring integument. Area behind lower eye lobes with dense yellow pubescence close to eye, glabrous close to prothorax. Genae 0.57 times length of lower eye lobe; with sparse yellowish-white pubescence, except dense yellow pubescence closet to frons and clypeus and glabrous apex; with a few long, erect brown setae interspersed. Wide central area of postclypeus with pubescence as on frons, except glabrous central area; with one long, erect brown seta on each side. Sides of postclypeus glabrous. Labrum with moderately abundant yellowish-white pubescence not obscuring integument, except glabrous anterocentral third and anterior margin with short fringe of yellowish-brown setae; with a few long, erect dark-brown setae interspersed on posterior half. Gulamentum smooth, glabrous, except sparse whitish pubescence on intermaxillary process. Antennal tubercles with abundant pale-yellow pubescence partially obscuring integument, except brown pubescence partially obscuring integument dorsally, this pubescence not reaching dorsal apex. Distance between upper eye lobes 0.19 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.54 times distance between outer margins of eyes. Antennae 3.4 times elytral length, reaching elytral apex just before middle of antennomere V. Scape with abundant yellowish pubescence not obscuring integument dorsally and laterally, except sparser yellowish-brown pubescence on apical quarter, and abundant whitish pubescence not obscuring integument ventrally; with sparse, long, erect, thick black setae on apical quarter of ventral surface. Pedicel with abundant pale-yellow pubescence on dorsal base, white on base lateral and ventral surfaces, brownish on remaining surface; with a few long, erect, thick blackish setae ventrally. Antennomeres III–XI with abundant whitish pubescence not obscuring integument on light integumental area, and dark with short whitish setae interspersed on dark integumental area, whitish setae more bristly from VIII; ventral surface of III–V with sparse, moderately long, erect, thick blackish setae ventrally, gradually sparser toward V; ventral apex of VI–VII with two moderately long, erect, thick blackish setae. Antennal formula based on length of antennomere III: scape = 0.97; pedicel = 0.12; IV = 0.95; V = 0.86; VI = 0.88; VII = 0.85; VIII = 0.91; IX = 0.94; X = 0.97; XI = 1.03.
</paragraph>
<caption id="DF3046947552FFE6FF453A1CE0B7F808" ID-DOI="http://doi.org/10.5281/zenodo.14390254" ID-Zenodo-Dep="14390254" httpUri="https://zenodo.org/record/14390254/files/figure.png" pageId="15" pageNumber="182" startId="15.[152,270,1864,1888]" targetBox="[159,1428,189,1832]" targetPageId="15" targetType="figure">
<paragraph id="8BF0161C7552FFE6FF453A1CE0B7F808" blockId="15.[151,1436,1864,1924]" pageId="15" pageNumber="182">
<emphasis id="B93BCA0E7552FFE6FF453A1CE34BF8EC" bold="true" box="[152,346,1864,1888]" pageId="15" pageNumber="182">FIGURES 27–31.</emphasis>
<taxonomicName id="4C4F6D9F7552FFE6FE823A1CE00FF8EC" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[351,542,1864,1888]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="182" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B93BCA0E7552FFE6FE823A1CE00FF8EC" box="[351,542,1864,1888]" italics="true" pageId="15" pageNumber="182">Sternacutus similis</emphasis>
</taxonomicName>
<emphasis id="B93BCA0E7552FFE6FDF93A1CE065F8EC" bold="true" box="[548,628,1864,1888]" pageId="15" pageNumber="182">
<taxonomicNameLabel id="A20877757552FFE6FDF93A1CE065F8EC" box="[548,628,1864,1888]" pageId="15" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male:
<emphasis id="B93BCA0E7552FFE6FCC03A1CE12EF8EC" bold="true" box="[797,831,1864,1888]" pageId="15" pageNumber="182">27)</emphasis>
Dorsal habitus;
<emphasis id="B93BCA0E7552FFE6FC393A1CE614F8EC" bold="true" box="[996,1029,1864,1888]" pageId="15" pageNumber="182">28)</emphasis>
Ventral habitus;
<emphasis id="B93BCA0E7552FFE6FB6D3A1CE6C0F8EC" bold="true" box="[1200,1233,1864,1888]" pageId="15" pageNumber="182">29)</emphasis>
Lateral habitus;
<emphasis id="B93BCA0E7552FFE6FAA73A1CE78DF8EC" bold="true" box="[1402,1436,1864,1888]" pageId="15" pageNumber="182">30)</emphasis>
Head, frontal view;
<emphasis id="B93BCA0E7552FFE6FEBF3A38E392F808" bold="true" box="[354,387,1900,1924]" pageId="15" pageNumber="182">31)</emphasis>
Centrobasal crests of elytra.
</paragraph>
</caption>
<paragraph id="8BF0161C754DFFF8FF1A394FE023FD25" blockId="16.[151,1437,151,2013]" lastBlockId="17.[151,1437,151,1797]" lastPageId="17" lastPageNumber="184" pageId="16" pageNumber="183">
<emphasis id="B93BCA0E754DFFF9FF1A394FE336FBB9" bold="true" box="[199,295,1051,1077]" pageId="16" pageNumber="183">Thorax.</emphasis>
Prothorax wider than long; anterior constriction narrow, well marked; sides slightly divergent from anterior constriction to middle, then slightly rounded widened toward lateral tubercles and subparallel-sided between them and posterolateral angles; Lateral tubercles small, conical, slightly inclined backward, located on posterior fifth. Pronotum slightly gibbose on each side of anterior half and slightly longitudinally tumid on center of posterior half; moderately abundantly, coarsely punctate near anterior and posterior margins and centrally, sparsely, coarsely punctate on remaining surface; with large, subreniform dark-brown pubescent macula on each side of middle, from anterior quarter to middle; with irregular dark-brown pubescent macula on each side of middle; with subglabrous area on each side near posterior margin; with large, rectangular area centrally on anterior third with sparse yellowish pubescence close to anterior margin, transverse, moderately pale-yellow pubescent band on middle, longitudinal yellowish pubescent band on center of anterior half, subglabrous on remaining anterior half, and sparse yellowish-brown pubescence on posterior half; remaining surface with pale-yellow pubescence, dense on some areas, abundant and not obscuring integument on others, except V-shaped whitish pubescent macula on center of posterior quarter. Sides of prothorax with abundant pale-yellow pubescence partially obscuring integument, except large dark-brown pubescent macula centrally, not reaching anterior margin, and glabrous anterior sulcus. Prosternum with abundant pale-yellow pubescence partially obscuring integument laterally and moderately abundant white pubescence on remaining surface, except glabrous anterior sulcus. Prosternal process with moderately abundant white pubescence not obscuring integument; narrowest area 0.24 times procoxal width. Mesoventrite with sparse whitish pubescence centrally and white pubescence laterally, lateral pubescence more yellowish-white toward apex. Mesanepisterna with abundant whitish pubescence close to mesoventrite, dense yellowish-brown pubescence centrally, and distinctly sparse yellowish-brown pubescence on outer side. Mesepimera with dense yellowish-brown pubescence. Metanepisterna with dense yellowish pubescence. Metaventrite with dense yellowish pubescence laterally, pubescence gradually sparser, whitish toward central region, except dense whitish pubescence close to metacoxal cavities. Mesoventral process with moderately abundant white pubescence not obscuring integument; narrowest area 0.37 times mesocoxal width. Scutellum sparse yellowish pubescence, except abundant, both yellowish and pale-yellow pubescence on close to margins, denser close to posterior margin.
<emphasis id="B93BCA0E754DFFF9FBFC3A2FE665F819" bold="true" box="[1057,1140,1915,1941]" pageId="16" pageNumber="183">Elytra.</emphasis>
Sides slightly convergent from humeri to middle, more distinctly convergent toward apex on posterior half; apex oblique, slightly concave between outer and sutural angles; centrobasal crest elevated, with dense, bristly blackish pubescence; humeral carinae absent; moderately abundantly, coarsely punctate on anterior half, punctures gradually sparser toward apex on posterior half; with abundant pubescence not obscuring integument, pubescence mostly yellowish on anterior third and laterally, mostly whitish on remaining dorsal surface, with abundant, irregular dark-brown pubescent maculae and spots interspersed, larger dark pubescent maculae oblique, located dorsally just after middle.
<emphasis id="B93BCA0E754CFFF8FA803C57E78DFE91" bold="true" box="[1373,1436,259,285]" pageId="17" pageNumber="184">Legs.</emphasis>
Femoral peduncles with abundant white pubescence not obscuring integument; femoral clubs with abundant light yellowish-brown pubescence not obscuring integument dorsally and on superior region of sides, and abundant white pubescence not obscuring integument on remaining surface. Protibiae with abundant whitish pubescence on basal half, more yellowish depending on light intensity and source, sparse yellowish-brown pubescence dorsally and laterally on anterior half, except both whitish and yellowish pubescence apically, and dense, bristly yellowish-brown pubescence on ventral apical half; meso- and metatibiae with sparse yellowish-brown pubescence basally, dense whitish pubescence on remaining basal half, and sparse, bristly, both yellowish-brown and brown setae on apical half of dorsal and lateral surfaces, and dense, bristly yellowish-brown pubescence on apical half of ventral surface; dorsal surface of mesotibiae with abundant, short, thick, erect dark-brown setae, setae gradually shorter toward apex. Dorsal surface of tarsomeres with abundant white pubescence not obscuring integument; metatarsomere I distinctly longer than II–III together.
</paragraph>
<paragraph id="8BF0161C754CFFF8FF1A3FE6E073FD7D" blockId="17.[151,1437,151,1797]" pageId="17" pageNumber="184">
<emphasis id="B93BCA0E754CFFF8FF1A3FE6E352FD40" bold="true" box="[199,323,690,716]" pageId="17" pageNumber="184">Abdomen.</emphasis>
Ventrites with abundant whitish pubescence not obscuring integument, denser apically on ventrites 1–4; apex of ventrite 5 strongly concave.
</paragraph>
<paragraph id="8BF0161C754CFFF8FF1A3FAEE712FCB5" blockId="17.[151,1437,151,1797]" pageId="17" pageNumber="184">
<emphasis id="B93BCA0E754CFFF8FF1A3FAEE3B4FC98" bold="true" box="[199,421,762,788]" pageId="17" pageNumber="184">Dimensions (mm).</emphasis>
Total length, 5.50; prothoracic length, 1.00; anterior prothoracic width, 1.25; posterior prothoracic width, 1.40; maximum prothoracic width, 1.65; humeral width, 2.15; elytral length, 4.00.
</paragraph>
</subSubSection>
<subSubSection id="C3554597754CFFF8FF1A3E17E08DFC0D" pageId="17" pageNumber="184" type="materials_examined">
<paragraph id="8BF0161C754CFFF8FF1A3E17E08DFC0D" blockId="17.[151,1437,151,1797]" pageId="17" pageNumber="184">
<emphasis id="B93BCA0E754CFFF8FF1A3E17E367FCD0" bold="true" box="[199,374,834,861]" pageId="17" pageNumber="184">Type material.</emphasis>
<materialsCitation id="3B271C41754CFFF8FEA03E17E08DFC0D" collectingDate="2003-04-01" collectingDateMax="2003-04-06" collectingDateMin="2003-04-01" collectionCode="MNKM" collectorName="R. Clarke" country="Bolivia" location="Flora &amp; Fauna Hotel" municipality="Buena Vista" pageId="17" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="54F4A8BE754CFFF8FEA03E17E3F4FCD1" box="[381,485,835,861]" pageId="17" pageNumber="184" type="holotype">Holotype</typeStatus>
<specimenCount id="9D49DD95754CFFF8FE303E17E032FCD1" box="[493,547,835,861]" pageId="17" pageNumber="184" type="male">male</specimenCount>
from
<collectingCountry id="F358568C754CFFF8FDB43E17E0CEFCD1" box="[617,735,835,861]" name="Bolivia" pageId="17" pageNumber="184">BOLIVIA</collectingCountry>
,
<emphasis id="B93BCA0E754CFFF8FD373E17E179FCD1" box="[746,872,835,861]" italics="true" pageId="17" pageNumber="184">
<collectingRegion id="498BD8FE754CFFF8FD373E17E179FCD1" box="[746,872,835,861]" country="Bolivia" name="Santa Cruz" pageId="17" pageNumber="184">Santa Cruz</collectingRegion>
</emphasis>
:
<locationDeviation id="9A420F3E754CFFF8FCA53E17E615FCD1" bearing="157.5" box="[888,1028,835,861]" distance="5000.0" pageId="17" pageNumber="184">
<quantity id="4CB7BBF9754CFFF8FCA53E17E1DCFCD1" box="[888,973,835,861]" metricMagnitude="3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="184" unit="km" value="5.0" valueMax="6.0" valueMin="4.0">4–6 km</quantity>
SSE
</locationDeviation>
<collectingMunicipality id="6B948C66754CFFF8FBD13E17E686FCD1" box="[1036,1175,835,861]" country="Bolivia" name="Buena Vista" pageId="17" pageNumber="184" stateProvince="Santa Cruz">Buena Vista</collectingMunicipality>
, F &amp; F [
<location id="8E9040C7754CFFF8FB283E17E2CAFC0D" LSID="urn:lsid:plazi:treatment:03E6A70A7553FFF8FF4A3997E6DAF889:8E9040C7754CFFF8FB283E17E2CAFC0D" country="Bolivia" municipality="Buena Vista" name="Flora &amp; Fauna Hotel" pageId="17" pageNumber="184" stateProvince="Santa Cruz">Flora &amp; Fauna Hotel</location>
],
<date id="FFF130DC754CFFF8FF303E33E368FC0D" box="[237,377,871,897]" pageId="17" pageNumber="184" value="2003-04-01" valueMax="2003-04-06" valueMin="2003-04-01">
<collectingDate id="EFB5C934754CFFF8FF303E33E368FC0D" box="[237,377,871,897]" pageId="17" pageNumber="184" value="2003-04-01" valueMax="2003-04-06" valueMin="2003-04-01">1–6.IV.2003</collectingDate>
</date>
,
<collectorName id="26BA73CA754CFFF8FE583E33E3E1FC0D" box="[389,496,871,897]" pageId="17" pageNumber="184">R. Clarke</collectorName>
leg. (
<collectionCode id="ED5E8ED9754CFFF8FDF23E33E081FC0D" box="[559,656,871,897]" collectionName="MNKM" pageId="17" pageNumber="184">MNKM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3554597754CFFF8FF1A3EDFE0AFFC45" pageId="17" pageNumber="184" type="etymology">
<paragraph id="8BF0161C754CFFF8FF1A3EDFE0AFFC45" blockId="17.[151,1437,151,1797]" pageId="17" pageNumber="184">
<emphasis id="B93BCA0E754CFFF8FF1A3EDFE35FFC29" bold="true" box="[199,334,907,933]" pageId="17" pageNumber="184">Etymology.</emphasis>
The species name
<taxonomicName id="4C4F6D9F754CFFF8FDFE3EDFE068FC29" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[547,633,907,933]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="similis">
“
<emphasis id="B93BCA0E754CFFF8FDED3EDFE068FC29" box="[560,633,907,933]" italics="true" pageId="17" pageNumber="184">similis</emphasis>
</taxonomicName>
” (Latin, meaning “similar”) refers to the fact that this species is similar to several already described species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3554597754CFFF8FF1A3E87E6DAF889" pageId="17" pageNumber="184" type="discussion">
<paragraph id="8BF0161C754CFFF8FF1A3E87E6DAF889" blockId="17.[151,1437,151,1797]" pageId="17" pageNumber="184">
<emphasis id="B93BCA0E754CFFF8FF1A3E87E32AFC61" bold="true" box="[199,315,979,1005]" pageId="17" pageNumber="184">Remarks.</emphasis>
<taxonomicName id="4C4F6D9F754CFFF8FE9C3E87E004FC61" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[321,533,979,1005]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B93BCA0E754CFFF8FE9C3E87E004FC61" box="[321,533,979,1005]" italics="true" pageId="17" pageNumber="184">Sternacutus similis</emphasis>
</taxonomicName>
<emphasis id="B93BCA0E754CFFF8FDC63E87E065FC61" bold="true" box="[539,628,979,1005]" pageId="17" pageNumber="184">
<taxonomicNameLabel id="A2087775754CFFF8FDC63E87E065FC61" box="[539,628,979,1005]" pageId="17" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C4F6D9F754CFFF8FCDE3E87E65EFC61" authority="(Bates, 1885)" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[771,1103,978,1005]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="angulistigma">
<emphasis id="B93BCA0E754CFFF8FCDE3E87E1A1FC61" box="[771,944,979,1005]" italics="true" pageId="17" pageNumber="184">S. angulistigma</emphasis>
(Bates, 1885)
</taxonomicName>
,
<taxonomicName id="4C4F6D9F754CFFF8FB873E87E786FC61" authority="(Bates, 1863)" baseAuthorityName="Bates" baseAuthorityYear="1863" box="[1114,1431,979,1005]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cinerascens">
<emphasis id="B93BCA0E754CFFF8FB873E87E6E8FC60" box="[1114,1273,979,1005]" italics="true" pageId="17" pageNumber="184">S. cinerascens</emphasis>
(Bates, 1863)
</taxonomicName>
,
<taxonomicName id="4C4F6D9F754CFFF8FF4A3EA3E3C2FB9D" authority="Roguet, 2023" authorityName="Roguet" authorityYear="2023" box="[151,467,1015,1041]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93BCA0E754CFFF8FF4A3EA3E33CFB9C" box="[151,301,1015,1041]" italics="true" pageId="17" pageNumber="184">S. compactus</emphasis>
Roguet, 2023
</taxonomicName>
,
<taxonomicName id="4C4F6D9F754CFFF8FE3D3EA3E13AFB9D" authority="(Bates, 1885)" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[480,811,1014,1041]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="B93BCA0E754CFFF8FE3D3EA3E096FB9D" box="[480,647,1015,1041]" italics="true" pageId="17" pageNumber="184">S. cribripennis</emphasis>
(Bates, 1885)
</taxonomicName>
,
<taxonomicName id="4C4F6D9F754CFFF8FCE43EA3E6BFFB9D" authority="(Aurivillius, 1922)" baseAuthorityName="Aurivillius" baseAuthorityYear="1922" box="[825,1198,1015,1041]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B93BCA0E754CFFF8FCE43EA3E1DFFB9D" box="[825,974,1015,1041]" italics="true" pageId="17" pageNumber="184">S. dimidiatus</emphasis>
(Aurivillius, 1922)
</taxonomicName>
, and
<taxonomicName id="4C4F6D9F754CFFF8FB303EA3E3AEFBB9" authority="(Monne &amp; Monne, 2012)" baseAuthorityName="Monne &amp; Monne" baseAuthorityYear="2012" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="griseostigma">
<emphasis id="B93BCA0E754CFFF8FB303EA3E78CFB9C" box="[1261,1437,1015,1041]" italics="true" pageId="17" pageNumber="184">S. griseostigma</emphasis>
(Monné &amp; Monné, 2012)
</taxonomicName>
. It differs from
<taxonomicName id="4C4F6D9F754CFFF8FDA7394FE13AFBB9" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[634,811,1051,1077]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="angulistigma">
<emphasis id="B93BCA0E754CFFF8FDA7394FE13AFBB9" box="[634,811,1051,1077]" italics="true" pageId="17" pageNumber="184">S. angulistigma</emphasis>
</taxonomicName>
(see photographs on
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and drawing in the original description) by the body stouter, width about 0.4 times the total length (slender, with width almost 0.3 times the total length in
<taxonomicName id="4C4F6D9F754CFFF8FEB83937E005FBF1" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[357,532,1123,1149]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="angulistigma">
<emphasis id="B93BCA0E754CFFF8FEB83937E005FBF1" box="[357,532,1123,1149]" italics="true" pageId="17" pageNumber="184">S. angulistigma</emphasis>
</taxonomicName>
, the difference much more conspicuous when comparing the two species); from
<taxonomicName id="4C4F6D9F754CFFF8FF4A39D3E327FB2C" baseAuthorityName="Bates" baseAuthorityYear="1863" box="[151,310,1159,1185]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cinerascens">
<emphasis id="B93BCA0E754CFFF8FF4A39D3E327FB2C" box="[151,310,1159,1185]" italics="true" pageId="17" pageNumber="184">S. cinerascens</emphasis>
</taxonomicName>
(see photographs on
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) especially by the upper eye lobes distinctly wider and the distance between them slightly wider than the width of one upper lobe (distinctly narrower and the distance between then larger than twice the width of one upper lobe); from
<taxonomicName id="4C4F6D9F754CFFF8FD2F399BE196FB64" authorityName="Roguet" authorityYear="2023" box="[754,903,1231,1257]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93BCA0E754CFFF8FD2F399BE196FB64" box="[754,903,1231,1257]" italics="true" pageId="17" pageNumber="184">S. compactus</emphasis>
</taxonomicName>
(see photographs on
<bibRefCitation id="EFDE6BED754CFFF8FBA1399BE71DFB65" author="Bezark, L. G." box="[1148,1292,1231,1257]" pageId="17" pageNumber="184" refId="ref10785" refString="Bezark, L. G. (2024) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 8 June 2024)" type="url" year="2024">Bezark 2024</bibRefCitation>
and original description) by the lateral tubercles of the prothorax smaller (moderately large in
<taxonomicName id="4C4F6D9F754CFFF8FBED39A7E6D4FA80" authorityName="Roguet" authorityYear="2023" box="[1072,1221,1267,1293]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93BCA0E754CFFF8FBED39A7E6D4FA80" box="[1072,1221,1267,1293]" italics="true" pageId="17" pageNumber="184">S. compactus</emphasis>
</taxonomicName>
), the elytral length about 1.9 times the humeral width (about 1.6 times in
<taxonomicName id="4C4F6D9F754CFFF8FD233843E182FABC" authorityName="Roguet" authorityYear="2023" box="[766,915,1303,1329]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93BCA0E754CFFF8FD233843E182FABC" box="[766,915,1303,1329]" italics="true" pageId="17" pageNumber="184">S. compactus</emphasis>
</taxonomicName>
), outer apex of elytra projected (not projected in
<taxonomicName id="4C4F6D9F754CFFF8FF68386FE35BFAD8" authorityName="Roguet" authorityYear="2023" box="[181,330,1339,1365]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93BCA0E754CFFF8FF68386FE35BFAD8" box="[181,330,1339,1365]" italics="true" pageId="17" pageNumber="184">S. compactus</emphasis>
</taxonomicName>
), and prosternal and mesoventral processes wider (slender in
<taxonomicName id="4C4F6D9F754CFFF8FBD4386FE68FFAD8" authorityName="Roguet" authorityYear="2023" box="[1033,1182,1339,1365]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B93BCA0E754CFFF8FBD4386FE68FFAD8" box="[1033,1182,1339,1365]" italics="true" pageId="17" pageNumber="184">S. compactus</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C4F6D9F754CFFF8FB2B386FE78DFAD9" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[1270,1436,1339,1365]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="B93BCA0E754CFFF8FB2B386FE78DFAD9" box="[1270,1436,1339,1365]" italics="true" pageId="17" pageNumber="184">S. cribripennis</emphasis>
</taxonomicName>
(see photographs on
<bibRefCitation id="EFDE6BED754CFFF8FE5C380BE01EFAF5" author="Bezark, L. G." box="[385,527,1375,1401]" pageId="17" pageNumber="184" refId="ref10785" refString="Bezark, L. G. (2024) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 8 June 2024)" type="url" year="2024">Bezark 2024</bibRefCitation>
and
<bibRefCitation id="EFDE6BED754CFFF8FD9B380BE19AFAF5" author="Santos-Silva, A. &amp; Devesa, S." box="[582,907,1375,1401]" pageId="17" pageNumber="184" pagination="54 - 71" refId="ref11527" refString="Santos-Silva, A., &amp; Devesa, S. (2021) Descriptions, transference, and new records in Lamiinae from Nicaragua (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, 39, 54 - 71. https: // doi. org / 10.11646 / zootaxa. 4942.4.5" type="journal article" year="2021">Santos-Silva &amp; Devesa 2021</bibRefCitation>
) by the stouter body, width about 0.4 times the total length (slender, about
<quantity id="4CB7BBF9754CFFF8FE0C38D6E002FA11" box="[465,531,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="17" pageNumber="184" unit="in" value="0.5">0.5 in</quantity>
<taxonomicName id="4C4F6D9F754CFFF8FDC138D7E0D2FA11" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[540,707,1411,1437]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="B93BCA0E754CFFF8FDC138D7E0D2FA11" box="[540,707,1411,1437]" italics="true" pageId="17" pageNumber="184">S. cribripennis</emphasis>
</taxonomicName>
), elytra with a short and longitudinal yellowish pubescent band between base and centrobasal crest slightly contrasting with the pubescence on its sides (distinctly contrasting with the pubescence on its sides in
<taxonomicName id="4C4F6D9F754CFFF8FE2C389FE086FA69" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[497,663,1483,1509]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="B93BCA0E754CFFF8FE2C389FE086FA69" box="[497,663,1483,1509]" italics="true" pageId="17" pageNumber="184">S. cribripennis</emphasis>
</taxonomicName>
), with large dark macula on the posterior half of elytra oblique (not oblique in
<taxonomicName id="4C4F6D9F754CFFF8FECE38BBE3AAF985" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[275,443,1519,1545]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="B93BCA0E754CFFF8FECE38BBE3AAF985" box="[275,443,1519,1545]" italics="true" pageId="17" pageNumber="184">S. cribripennis</emphasis>
</taxonomicName>
), and by the metafemoral clubs distinctly stouter in males (slender in
<taxonomicName id="4C4F6D9F754CFFF8FB3938BBE79AF985" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[1252,1419,1519,1545]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="B93BCA0E754CFFF8FB3938BBE79AF985" box="[1252,1419,1519,1545]" italics="true" pageId="17" pageNumber="184">S. cribripennis</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C4F6D9F754CFFF8FF0B3B47E378F9A1" baseAuthorityName="Aurivillius" baseAuthorityYear="1922" box="[214,361,1555,1581]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B93BCA0E754CFFF8FF0B3B47E378F9A1" box="[214,361,1555,1581]" italics="true" pageId="17" pageNumber="184">S. dimidiatus</emphasis>
</taxonomicName>
(see photographs on
<bibRefCitation id="EFDE6BED754CFFF8FD803B47E0FFF9A1" author="Bezark, L. G." box="[605,750,1555,1581]" pageId="17" pageNumber="184" refId="ref10785" refString="Bezark, L. G. (2024) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 8 June 2024)" type="url" year="2024">Bezark 2024</bibRefCitation>
) by the stouter body (slender in
<taxonomicName id="4C4F6D9F754CFFF8FBBC3B47E6E5F9A1" baseAuthorityName="Aurivillius" baseAuthorityYear="1922" box="[1121,1268,1555,1581]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B93BCA0E754CFFF8FBBC3B47E6E5F9A1" box="[1121,1268,1555,1581]" italics="true" pageId="17" pageNumber="184">S. dimidiatus</emphasis>
</taxonomicName>
, about as in
<taxonomicName id="4C4F6D9F754CFFF8FA5A3B47E338F9DD" baseAuthorityName="Bates" baseAuthorityYear="1885" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="angulistigma">
<emphasis id="B93BCA0E754CFFF8FA5A3B47E338F9DD" italics="true" pageId="17" pageNumber="184">S. angulistigma</emphasis>
</taxonomicName>
), distance between upper eye lobes slightly larger than the width of one upper lobe (wider than twice the width of one upper lobe in
<taxonomicName id="4C4F6D9F754CFFF8FE1E3B0FE045F9F9" baseAuthorityName="Aurivillius" baseAuthorityYear="1922" box="[451,596,1627,1653]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B93BCA0E754CFFF8FE1E3B0FE045F9F9" box="[451,596,1627,1653]" italics="true" pageId="17" pageNumber="184">S. dimidiatus</emphasis>
</taxonomicName>
), and elytra without transverse dark macula on middle of the dorsal surface (present in
<taxonomicName id="4C4F6D9F754CFFF8FEC73B2BE3BFF915" baseAuthorityName="Aurivillius" baseAuthorityYear="1922" box="[282,430,1663,1689]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B93BCA0E754CFFF8FEC73B2BE3BFF915" box="[282,430,1663,1689]" italics="true" pageId="17" pageNumber="184">S. dimidiatus</emphasis>
</taxonomicName>
); and from
<taxonomicName id="4C4F6D9F754CFFF8FDE63B2BE0FBF914" baseAuthorityName="Monne &amp; Monne" baseAuthorityYear="2012" box="[571,746,1663,1689]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="griseostigma">
<emphasis id="B93BCA0E754CFFF8FDE63B2BE0FBF914" box="[571,746,1663,1689]" italics="true" pageId="17" pageNumber="184">S. griseostigma</emphasis>
</taxonomicName>
(see photographs on
<bibRefCitation id="EFDE6BED754CFFF8FC3E3B2BE665F915" author="Bezark, L. G." box="[995,1140,1663,1689]" pageId="17" pageNumber="184" refId="ref10785" refString="Bezark, L. G. (2024) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 8 June 2024)" type="url" year="2024">Bezark 2024</bibRefCitation>
and original description), by the pronotum with large subreniform dark macula on each side of middle (absent in
<taxonomicName id="4C4F6D9F754CFFF8FBB63BF7E709F930" baseAuthorityName="Monne &amp; Monne" baseAuthorityYear="2012" box="[1131,1304,1699,1725]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="griseostigma">
<emphasis id="B93BCA0E754CFFF8FBB63BF7E709F930" box="[1131,1304,1699,1725]" italics="true" pageId="17" pageNumber="184">S. griseostigma</emphasis>
</taxonomicName>
), pronotum without narrow longitudinal light pubescent band on each side of the posterior half (present in
<taxonomicName id="4C4F6D9F754CFFF8FB6C3B93E74CF96C" baseAuthorityName="Monne &amp; Monne" baseAuthorityYear="2012" box="[1201,1373,1735,1761]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="griseostigma">
<emphasis id="B93BCA0E754CFFF8FB6C3B93E74CF96C" box="[1201,1373,1735,1761]" italics="true" pageId="17" pageNumber="184">S. griseostigma</emphasis>
</taxonomicName>
), and elytra without wide and large dark zig-zag macula on posterior half (present in
<taxonomicName id="4C4F6D9F754CFFF8FBD33BBFE6AAF888" baseAuthorityName="Monne &amp; Monne" baseAuthorityYear="2012" box="[1038,1211,1771,1797]" class="Insecta" family="Cerambycidae" genus="Sternacutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="184" phylum="Arthropoda" rank="species" species="griseostigma">
<emphasis id="B93BCA0E754CFFF8FBD33BBFE6AAF888" box="[1038,1211,1771,1797]" italics="true" pageId="17" pageNumber="184">S. griseostigma</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3F136271FCB78740180BC5E9D06E9092">New species, notes and record of New World Acanthocinini (Coleoptera: Cerambycidae, Lamiinae)</mods:title>
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<mods:part id="B4918752ACAC722626EF7B7FFFE63346">
<mods:date id="F1FC22DE0260B1947F53E06AE9A050EA">2024</mods:date>
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<mods:number id="3BBC000D8396357F171882ABB46567DD">2024-12-11</mods:number>
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<mods:detail id="4EAD579AF38C4A3344BBC22B74EB8200" type="volume">
<mods:number id="FC30A0CE907AD8A5C32A2008A70FDACD">5551</mods:number>
</mods:detail>
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<mods:end id="72C30F643355DC65F1497CCC22AABB8C">185</mods:end>
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<mods:url id="0982CD32A53534776826AFCC23D88159">https://doi.org/10.11646/zootaxa.5551.1.8</mods:url>
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<mods:classification id="37EF889C11313B88E566FFF7408B7381">journal article</mods:classification>
<mods:identifier id="68CD2F1FEED154095ECAEBED8AC15841" type="DOI">10.11646/zootaxa.5551.1.8</mods:identifier>
<mods:identifier id="E19D8AECA254DD0078C50EF40B862320" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E6A70A7557FFE2FF4A389FE78CFB56" LSID="urn:lsid:plazi:treatment:03E6A70A7557FFE2FF4A389FE78CFB56" httpUri="http://treatment.plazi.org/id/03E6A70A7557FFE2FF4A389FE78CFB56" lastPageId="11" lastPageNumber="178" pageId="10" pageNumber="177">
<subSubSection id="C35545977557FFE3FF4A389FE386FA69" box="[151,407,1483,1509]" pageId="10" pageNumber="177" type="nomenclature">
<paragraph id="8BF0161C7557FFE3FF4A389FE386FA69" blockId="10.[151,407,1483,1509]" box="[151,407,1483,1509]" pageId="10" pageNumber="177">
<taxonomicName id="4C4F6D9F7557FFE3FF4A389FE333FA69" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[151,290,1483,1509]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93BCA0E7557FFE3FF4A389FE333FA69" box="[151,290,1483,1509]" italics="true" pageId="10" pageNumber="177">Quissunego</emphasis>
</taxonomicName>
<emphasis id="B93BCA0E7557FFE3FEF4389FE386FA69" bold="true" box="[297,407,1483,1509]" pageId="10" pageNumber="177">
<taxonomicNameLabel id="A20877757557FFE3FEF4389FE386FA69" box="[297,407,1483,1509]" pageId="10" pageNumber="177" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35545977557FFE3FF4A3B47E15DF9A1" box="[151,844,1554,1581]" pageId="10" pageNumber="177" type="type_taxon">
<paragraph id="8BF0161C7557FFE3FF4A3B47E15DF9A1" blockId="10.[151,1437,1554,2013]" box="[151,844,1554,1581]" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF4A3B47E33AF9A0" bold="true" box="[151,299,1554,1581]" pageId="10" pageNumber="177">
<typeStatus id="54F4A8BE7557FFE3FF4A3B47E2C3F9A1" box="[151,210,1555,1581]" pageId="10" pageNumber="177">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C4F6D9F7557FFE3FEE73B47E036F9A1" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[314,551,1555,1581]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
<emphasis id="B93BCA0E7557FFE3FEE73B47E036F9A1" box="[314,551,1555,1581]" italics="true" pageId="10" pageNumber="177">Quissunego confusus</emphasis>
</taxonomicName>
<emphasis id="B93BCA0E7557FFE3FDF33B47E099F9A1" bold="true" box="[558,648,1555,1581]" pageId="10" pageNumber="177">
<taxonomicNameLabel id="A20877757557FFE3FDF33B47E099F9A1" box="[558,648,1555,1581]" pageId="10" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="C35545977557FFE3FF1A3B63E08EF9F9" pageId="10" pageNumber="177" type="etymology">
<paragraph id="8BF0161C7557FFE3FF1A3B63E08EF9F9" blockId="10.[151,1437,1554,2013]" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF1A3B63E35FF9DD" bold="true" box="[199,334,1591,1617]" pageId="10" pageNumber="177">Etymology.</emphasis>
Latin, “
<taxonomicName id="4C4F6D9F7557FFE3FE783B63E03FF9DD" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[421,558,1591,1617]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="genus">Quissunego</taxonomicName>
” (Quis sun ego?) (Who am I?), allusive to the complicated process of including the type-species in a genus. Masculine gender.
</paragraph>
</subSubSection>
<subSubSection id="C35545977557FFE2FF1A3B2AE61DFE5D" lastPageId="11" lastPageNumber="178" pageId="10" pageNumber="177" type="description">
<paragraph id="8BF0161C7557FFE2FF1A3B2AE61DFE5D" blockId="10.[151,1437,1554,2013]" lastBlockId="11.[151,1437,151,1041]" lastPageId="11" lastPageNumber="178" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF1A3B2AE3B4F915" bold="true" box="[199,421,1662,1689]" pageId="10" pageNumber="177">Description. Male.</emphasis>
Body widened, slightly flattened, less than
<quantity id="4CB7BBF97557FFE3FC433B2BE1FEF915" box="[926,1007,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="177" unit="mm" value="10.0">10 mm</quantity>
in the
<typeStatus id="54F4A8BE7557FFE3FB9F3B2BE663F915" box="[1090,1138,1663,1689]" pageId="10" pageNumber="177">type</typeStatus>
species. Frons subsquare. Eyes moderately coarsely faceted; upper eye lobes wide, distance between them slightly larger than width of one upper lobe; lower eye lobes distinctly longer than genae; area of connection between eye lobes distinctly narrower than upper lobe.Antennae 11-segmented, distinctly longer than body; scape gradually widened from base to middle, then subparallel-sided slightly narrowed on apical fifth without apical modification, not reaching posterior margin of prothorax, with one long, erect dark seta near apex of ventral surface; pedicel cylindrical, slightly longer than wide; antennomeres III–XI filiform, without tufts of setae; III and IV similar same length, the longest, with a few short, erect, fine setae ventrally. Prothorax wider than long; anterior constriction narrow, well marked; sides slightly rounded and divergent from anterior constriction to lateral tubercles, then convergent to posterolateral angles; lateral tubercles large, as a continuation of the lateral margin, located slightly before posterior quarter. Pronotum without tubercles; sparsely, coarsely punctate on anterior 3/4, abundantly punctate on posterior quarter, punctures not following toward sides of prothorax. Prosternal process strongly narrowed centrally, then strongly widened posteriorly; narrowest area distinctly shorter than one-tenth of procoxal width. Mesoventral process narrow, gradually narrowed from base to posterior fifth, then moderately widened; narrowest area about one-eight mesocoxal width. Scutellum semicircle shaped. Elytra without tubercles and erect setae; subparallel-sided on anterior half, slightly narrowed just after humeri, then gradually convergent toward apex; apex slightly oblique and concave, with rounded outer and sutural angles; humeral angles not projected; humeral and dorsal carinae and centrobasal crest absent. Femora pedunculate-clavate; femoral clubs strongly widened; metafemora reaching apex of elytra. Tibiae cylindrical, without modifications. Metatarsomere I longer than II–III together.
</paragraph>
</subSubSection>
<subSubSection id="C35545977556FFE2FF1A3C8FE3F8FB9D" pageId="11" pageNumber="178" type="discussion">
<paragraph id="8BF0161C7556FFE2FF1A3C8FE3F9FC0D" blockId="11.[151,1437,151,1041]" pageId="11" pageNumber="178">
<emphasis id="B93BCA0E7556FFE2FF1A3C8FE32AFE79" bold="true" box="[199,315,475,501]" pageId="11" pageNumber="178">Remarks.</emphasis>
The key from
<bibRefCitation id="EFDE6BED7556FFE2FE343C8FE0CAFE79" author="Monne, M. A. &amp; Santos-Silva, A. &amp; Monne, M. L." box="[489,731,475,501]" pageId="11" pageNumber="178" pagination="302 - 337" refId="ref11322" refString="Monne, M. A., Santos-Silva, A. &amp; Monne, M. L. (2020 a) Key to Mexico and Central America genera of Acanthocinini (Coleoptera, Cerambycidae, Lamiinae) without erect setae on elytral surface, excluding the Caribbean Islands. Zootaxa, 4861 (3), 302 - 337. https: // doi. org / 10.11646 / zootaxa. 4861.3.1" type="journal article" year="2020">
Monné
<emphasis id="B93BCA0E7556FFE2FD9C3C8FE06AFE79" box="[577,635,475,501]" italics="true" pageId="11" pageNumber="178">et al.</emphasis>
(2020a)
</bibRefCitation>
leads
<taxonomicName id="4C4F6D9F7556FFE2FCFB3C8FE1BBFE79" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[806,938,475,501]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93BCA0E7556FFE2FCFB3C8FE1BBFE79" box="[806,938,475,501]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20877757556FFE2FC6E3C8FE60AFE79" box="[947,1051,475,501]" pageId="11" pageNumber="178" rank="genus">gen. nov.</taxonomicNameLabel>
to the alternative of couplet “40” with
<taxonomicName id="4C4F6D9F7556FFE2FF0C3CABE049FD95" authority="Bates, 1863" authorityName="Bates" authorityYear="1863" box="[209,600,511,537]" class="Insecta" family="Cerambycidae" genus="Lepturges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subGenus" subGenus="Lepturges">
<emphasis id="B93BCA0E7556FFE2FF0C3CABE351FD94" box="[209,320,511,536]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
(
<emphasis id="B93BCA0E7556FFE2FE8C3CABE3D1FD94" box="[337,448,511,536]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
) Bates, 1863
</taxonomicName>
. However, the lateral tubercles of the prothorax are distinctly distant from the posterolateral angles of the prothorax, making it very artificial to consider the new species as belonging to
<taxonomicName id="4C4F6D9F7556FFE2FF4A3F13E383FDED" authorityName="Bates" authorityYear="1863" box="[151,402,583,609]" class="Insecta" family="Cerambycidae" genus="Lepturges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subGenus" subGenus="Lepturges">
<emphasis id="B93BCA0E7556FFE2FF4A3F13E317FDEC" box="[151,262,583,608]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
(
<emphasis id="B93BCA0E7556FFE2FEC53F13E396FDEC" box="[280,391,583,608]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
)
</taxonomicName>
. Furthermore, the femoral clubs are much stouter than in the species currently included in
<taxonomicName id="4C4F6D9F7556FFE2FF4A3F3FE381FD09" authorityName="Bates" authorityYear="1863" box="[151,400,619,645]" class="Insecta" family="Cerambycidae" genus="Lepturges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subGenus" subGenus="Lepturges">
<emphasis id="B93BCA0E7556FFE2FF4A3F3FE317FD08" box="[151,262,619,644]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
(
<emphasis id="B93BCA0E7556FFE2FECB3F3FE394FD08" box="[278,389,619,644]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
)
</taxonomicName>
. The key from
<bibRefCitation id="EFDE6BED7556FFE2FDE63F3FE13BFD09" author="Monne, M. A. &amp; Santos-Silva, A. &amp; Monne, M. L." box="[571,810,619,645]" pageId="11" pageNumber="178" pagination="1 - 65" refId="ref11395" refString="Monne, M. A., Santos-Silva, A. &amp; Monne, M. L. (2020 b) Key to South American genera of Acanthocinini (Coleoptera, Cerambycidae, Lamiinae) without erect setae on elytra; synonymies, transferences, revalidation, and notes on genera with erect setae on elytra. Zootaxa, 4863 (1), 1 - 65. https: // doi. org / 10.11646 / zootaxa. 4863.1.1" type="journal article" year="2020">
Monné
<emphasis id="B93BCA0E7556FFE2FD4F3F3FE0DBFD09" box="[658,714,619,645]" italics="true" pageId="11" pageNumber="178">et al.</emphasis>
(2020b)
</bibRefCitation>
leads
<taxonomicName id="4C4F6D9F7556FFE2FCAC3F3FE1E4FD09" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[881,1013,619,645]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93BCA0E7556FFE2FCAC3F3FE1E4FD09" box="[881,1013,619,645]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20877757556FFE2FC213F3FE673FD09" box="[1020,1122,619,645]" pageId="11" pageNumber="178" rank="genus">gen. nov.</taxonomicNameLabel>
to the alternative of couplet “56,” and in it to
<taxonomicName id="4C4F6D9F7556FFE2FEB33FDBE0DAFD25" authority="Melzer, 1931" authorityName="Melzer" authorityYear="1931" box="[366,715,655,681]" class="Insecta" family="Cerambycidae" genus="Lophopoenopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FEB33FDBE037FD25" box="[366,550,655,681]" italics="true" pageId="11" pageNumber="178">Lophopoenopsis</emphasis>
Melzer, 1931
</taxonomicName>
. Although included in a key to genera without erect setae on elytra, we now know that the
<typeStatus id="54F4A8BE7556FFE2FE3F3FE7E003FD41" box="[482,530,691,717]" pageId="11" pageNumber="178">type</typeStatus>
species of
<taxonomicName id="4C4F6D9F7556FFE2FD523FE7E156FD41" authorityName="Melzer" authorityYear="1931" box="[655,839,691,717]" class="Insecta" family="Cerambycidae" genus="Lophopoenopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FD523FE7E156FD41" box="[655,839,691,717]" italics="true" pageId="11" pageNumber="178">Lophopoenopsis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4F6D9F7556FFE2FC893FE7E66EFD41" authority="Melzer, 1931" authorityName="Melzer" authorityYear="1931" box="[852,1151,691,717]" class="Insecta" family="Cerambycidae" genus="Lophopoenopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B93BCA0E7556FFE2FC893FE7E1F3FD41" box="[852,994,691,717]" italics="true" pageId="11" pageNumber="178">L. singularis</emphasis>
Melzer, 1931
</taxonomicName>
, has erect setae on elytra, which are absent in
<taxonomicName id="4C4F6D9F7556FFE2FEA03F83E010FD7D" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[381,513,727,753]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FEA03F83E010FD7D" box="[381,513,727,753]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
. Using the key by Dillon (1956), which includes only a few American genera of Acanthocinini,
<taxonomicName id="4C4F6D9F7556FFE2FE993FAFE3D9FC99" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[324,456,763,789]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FE993FAFE3D9FC99" box="[324,456,763,789]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
is included in the alternative of couplet “31” with
<taxonomicName id="4C4F6D9F7556FFE2FC223FAFE6E9FC99" authorityName="Bates" authorityYear="1863" box="[1023,1272,763,789]" class="Insecta" family="Cerambycidae" genus="Lepturges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subGenus" subGenus="Lepturges">
<emphasis id="B93BCA0E7556FFE2FC223FAFE67FFC98" box="[1023,1134,763,788]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
(
<emphasis id="B93BCA0E7556FFE2FBA33FAFE6FCFC98" box="[1150,1261,763,788]" italics="true" pageId="11" pageNumber="178">Lepturges</emphasis>
)
</taxonomicName>
. Finally, using the key to North American genera of
<taxonomicName id="4C4F6D9F7556FFE2FDEE3E4BE0C3FCB5" authorityName="Blanchard" authorityYear="1845" box="[563,722,799,825]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="185" phylum="Arthropoda" rank="tribe" tribe="Acanthocinini">Acanthocinini</taxonomicName>
from
<bibRefCitation id="EFDE6BED7556FFE2FCC83E4BE659FCB5" author="Linsley, E. G. &amp; Chemsak, J. A." box="[789,1096,798,825]" pageId="11" pageNumber="178" pagination="1 - 292" refId="ref11079" refString="Linsley, E. G. &amp; Chemsak, J. A. (1995) The Cerambycidae of North America. Part VII. No. 2. Taxonomy and classification of the subfamily Lamiinae, Tribes Acanthocinini through Hemilophini. In: Daly, H. V., Doyen, J., Hespenheide, H., Lane, R. S., Pinto, J., Powell, J. A., Shapiro, A, Ward, P. S. &amp; Smiley, J. (Eds.), University of California Publications in Entomology. Vol. 114. University of California Press, Berkeley, Los Angeles, and London, pp. i - xii + 1 - 292." type="book chapter" year="1995">Linsley &amp; Chemsak (1995)</bibRefCitation>
,
<taxonomicName id="4C4F6D9F7556FFE2FB8E3E4BE6C6FCB5" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[1107,1239,799,825]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FB8E3E4BE6C6FCB5" box="[1107,1239,799,825]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
is included in the alternative of couplet “10” with
<taxonomicName id="4C4F6D9F7556FFE2FDDB3E17E13BFCD1" authority="LeConte, 1853" authorityName="LeConte" authorityYear="1853" box="[518,810,834,861]" class="Insecta" family="Cerambycidae" genus="Sternidius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FDDB3E17E069FCD1" box="[518,632,835,861]" italics="true" pageId="11" pageNumber="178">Sternidius</emphasis>
LeConte, 1853
</taxonomicName>
, which has the mesoventral process very wide, while it is very narrow in
<taxonomicName id="4C4F6D9F7556FFE2FE803E33E3F0FC0D" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[349,481,871,897]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FE803E33E3F0FC0D" box="[349,481,871,897]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF0161C7556FFE2FF1A3EDFE3F8FB9D" blockId="11.[151,1437,151,1041]" pageId="11" pageNumber="178">
<taxonomicName id="4C4F6D9F7556FFE2FF1A3EDFE35AFC29" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[199,331,907,933]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FF1A3EDFE35AFC29" box="[199,331,907,933]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C4F6D9F7556FFE2FE343EDFE1D7FC29" authority="Audinet-Serville, 1835" authorityName="Audinet-Serville" authorityYear="1835" box="[489,966,906,933]" class="Insecta" family="Cerambycidae" genus="Leiopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subGenus" subGenus="Leiopus">
<emphasis id="B93BCA0E7556FFE2FE343EDFE053FC28" box="[489,578,907,932]" italics="true" pageId="11" pageNumber="178">Leiopus</emphasis>
(
<emphasis id="B93BCA0E7556FFE2FD883EDFE0BFFC28" box="[597,686,907,932]" italics="true" pageId="11" pageNumber="178">Leiopus</emphasis>
) Audinet-Serville, 1835
</taxonomicName>
, but differs by the upper eye lobes wide (narrow or moderately narrow in
<taxonomicName id="4C4F6D9F7556FFE2FDD33EFBE076FC44" box="[526,615,943,968]" class="Insecta" family="Cerambycidae" genus="Leiopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FDD33EFBE076FC44" box="[526,615,943,968]" italics="true" pageId="11" pageNumber="178">Leiopus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4F6D9F7556FFE2FDAA3EFBE0C1FC44" box="[631,720,943,968]" class="Insecta" family="Cerambycidae" genus="Leiopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FDAA3EFBE0C1FC44" box="[631,720,943,968]" italics="true" pageId="11" pageNumber="178">Leiopus</emphasis>
</taxonomicName>
)), and the punctures on the posterior region of the pronotum do not follow toward sides of the prothorax (following toward sides of the prothorax in
<taxonomicName id="4C4F6D9F7556FFE2FB883E87E733FC61" authorityName="Audinet-Serville" authorityYear="1835" box="[1109,1314,979,1005]" class="Insecta" family="Cerambycidae" genus="Leiopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subGenus" subGenus="Leiopus">
<emphasis id="B93BCA0E7556FFE2FB883E87E6BFFC60" box="[1109,1198,979,1004]" italics="true" pageId="11" pageNumber="178">Leiopus</emphasis>
(
<emphasis id="B93BCA0E7556FFE2FB623E87E709FC60" box="[1215,1304,979,1004]" italics="true" pageId="11" pageNumber="178">Leiopus</emphasis>
)
</taxonomicName>
, at least in the
<typeStatus id="54F4A8BE7556FFE2FF1D3EA3E2E1FB9D" box="[192,240,1015,1041]" pageId="11" pageNumber="178">type</typeStatus>
species of the genus).
</paragraph>
</subSubSection>
<subSubSection id="C35545977556FFE2FF4A3937E77EFBF1" box="[151,1391,1122,1149]" pageId="11" pageNumber="178" type="multiple">
<paragraph id="8BF0161C7556FFE2FF4A3937E77EFBF1" blockId="11.[151,1391,1122,1149]" box="[151,1391,1122,1149]" pageId="11" pageNumber="178">
<emphasis id="B93BCA0E7556FFE2FF4A3937E38EFBF1" bold="true" box="[151,415,1122,1149]" pageId="11" pageNumber="178">
The key from
<bibRefCitation id="EFDE6BED7556FFE2FE9A3936E38EFBF1" author="Monne, M. A. &amp; Santos-Silva, A. &amp; Monne, M. L." box="[327,415,1122,1149]" pageId="11" pageNumber="178" pagination="1 - 65" refId="ref11395" refString="Monne, M. A., Santos-Silva, A. &amp; Monne, M. L. (2020 b) Key to South American genera of Acanthocinini (Coleoptera, Cerambycidae, Lamiinae) without erect setae on elytra; synonymies, transferences, revalidation, and notes on genera with erect setae on elytra. Zootaxa, 4863 (1), 1 - 65. https: // doi. org / 10.11646 / zootaxa. 4863.1.1" type="journal article" year="2020">Monné</bibRefCitation>
</emphasis>
<emphasis id="B93BCA0E7556FFE2FE7A3937E3F3FBF1" box="[423,482,1123,1149]" italics="true" pageId="11" pageNumber="178">et al.</emphasis>
<emphasis id="B93BCA0E7556FFE2FE343936E68CFBF0" bold="true" box="[489,1181,1122,1149]" pageId="11" pageNumber="178">(2020b) can be updated as follows, with the exclusion of</emphasis>
<taxonomicName id="4C4F6D9F7556FFE2FB783937E774FBF1" authorityName="Melzer" authorityYear="1931" box="[1189,1381,1123,1149]" class="Insecta" family="Cerambycidae" genus="Lophopoenopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FB783937E774FBF1" box="[1189,1381,1123,1149]" italics="true" pageId="11" pageNumber="178">Lophopoenopsis</emphasis>
</taxonomicName>
:
</paragraph>
</subSubSection>
<subSubSection id="C35545977556FFE2FF4A39F3E78CFB56" pageId="11" pageNumber="178" type="key">
<keyStep id="30BB0EB97556FFE2FF4A39F3E78CFB56" pageId="11" pageNumber="178">
<paragraph id="8BF0161C7556FFE2FF4A39F3E78DFB31" blockId="11.[151,1437,1191,1243]" box="[151,1436,1191,1214]" pageId="11" pageNumber="178">
<keyLead id="30BEB5297556FFE2FF4A39F3E78DFB31" box="[151,1436,1191,1214]" pageId="11" pageNumber="178">
56(55). Distance between upper eye lobes larger than three times width of one upper lobe....
<taxonomicName id="4C4F6D9F7556FFE2FBF939FCE78DFB31" authority="Monne &amp; Monne, 2012" authorityName="Monne &amp; Monne" authorityYear="2012" box="[1060,1436,1192,1213]" class="Insecta" family="Cerambycidae" genus="Luteolepturges" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E7556FFE2FBF939FCE6A2FB31" box="[1060,1203,1192,1213]" italics="true" pageId="11" pageNumber="178">Luteolepturges</emphasis>
Monné &amp; Monné, 2012
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF0161C7556FFE2FF4A3991E78CFB56" blockId="11.[151,1437,1191,1243]" box="[151,1437,1221,1243]" pageId="11" pageNumber="178">
<keyLead id="30BEB5297556FFE2FF4A3991E78BFB56" box="[151,1434,1221,1243]" pageId="11" pageNumber="178">
– Distance between upper eye lobes slightly larger than width of one upper lobe.....................
<taxonomicName id="4C4F6D9F7556FFE2FB133991E72EFB56" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[1230,1343,1221,1242]" class="Insecta" family="Cerambycidae" genus="Quissunego" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93BCA0E7556FFE2FB133991E72EFB56" box="[1230,1343,1221,1242]" italics="true" pageId="11" pageNumber="178">Quissunego</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20877757556FFE2FA983991E78CFB56" box="[1349,1437,1221,1242]" pageId="11" pageNumber="178" rank="genus">gen. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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178
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<mods:title id="7A74985D9B12B64EA6D2BCA8683BEE32">New species, notes and record of New World Acanthocinini (Coleoptera: Cerambycidae, Lamiinae)</mods:title>
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<subSubSection id="C3554597755AFFEEFF4A3EBFE008FB8A" box="[151,537,1003,1030]" pageId="7" pageNumber="174" type="nomenclature">
<paragraph id="8BF0161C755AFFEEFF4A3EBFE008FB8A" blockId="7.[151,537,1003,1066]" box="[151,537,1003,1030]" pageId="7" pageNumber="174">
<heading id="D0B8A170755AFFEEFF4A3EBFE008FB8A" bold="true" box="[151,537,1003,1030]" fontSize="11" level="1" pageId="7" pageNumber="174" reason="1">
<emphasis id="B93BCA0E755AFFEEFF4A3EBFE008FB8A" bold="true" box="[151,537,1003,1030]" pageId="7" pageNumber="174">
<taxonomicName id="4C4F6D9F755AFFEEFF4A3EBFE3A5FB8A" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[151,436,1003,1030]" class="Insecta" family="Cerambycidae" genus="Pseudastylopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="morrisi" status="sp. nov.">
<emphasis id="B93BCA0E755AFFEEFF4A3EBFE3A5FB8A" bold="true" box="[151,436,1003,1030]" italics="true" pageId="7" pageNumber="174">Pseudastylopsis morrisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2087775755AFFEEFE663EB8E008FB8A" box="[443,537,1004,1030]" pageId="7" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3554597755AFFE3FF4A3944E712FC29" lastPageId="10" lastPageNumber="177" pageId="7" pageNumber="174" type="description">
<paragraph id="8BF0161C755AFFEEFF4A3944E336FBA6" blockId="7.[151,537,1003,1066]" box="[151,295,1040,1066]" pageId="7" pageNumber="174">(Figs 17–21)</paragraph>
<paragraph id="8BF0161C755AFFEEFF4A390CE735FAAA" blockId="7.[151,1437,1112,2039]" pageId="7" pageNumber="174">
<emphasis id="B93BCA0E755AFFEEFF4A390CE3ECFBFE" bold="true" box="[151,509,1112,1138]" pageId="7" pageNumber="174">
Description.
<typeStatus id="54F4A8BE755AFFEEFEEC390CE38FFBFE" box="[305,414,1112,1138]" pageId="7" pageNumber="174" type="holotype">Holotype</typeStatus>
female.
</emphasis>
Head capsule black except gulamentum partially brown; mentum and ligula light brown; palpomeres dark brown, except yellowish apex of maxillary palpomeres I–III and labial palpomeres I–II; anterior half of anteclypeus yellowish brown; labrum dark brown; scape black; pedicel, antennomeres III–IV, and XI dark brown; antennomeres V–X with wide reddish-brown ring between base and apex. Pronotum and sides of prothorax black; ventral surface of pro-, meso-, and metathorax dark brown. Scutellum, elytra, and legs black. Abdominal ventrites dark brown, except dark reddish-brown apex of ventrites 2–4, less so on ventrite 3.
</paragraph>
<paragraph id="8BF0161C755AFFE0FF1A3864E05EFE79" blockId="7.[151,1437,1112,2039]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="176" pageId="7" pageNumber="174">
<emphasis id="B93BCA0E755AFFEEFF1A3864E31FFAC6" bold="true" box="[199,270,1328,1354]" pageId="7" pageNumber="174">Head.</emphasis>
Frons densely, finely punctate, punctures slightly sparser close to eyes; with brown pubescence not obscuring integument, with short, decumbent white setae interspersed, except pale-yellow pubescence close to median groove superiorly, and pale-yellow pubescent macula close to eyes; with one long, erect dark brown seta close to eyes. Area between antennal tubercles with dense pale-yellow pubescence centrally, except glabrous median groove, and sparse yellowish-white pubescence laterally. Remaining surface of vertex with dense pale-yellow pubescence except two triangular subglabrous areas close to prothorax; with a few long, erect dark-brown setae close to eyes. Area behind upper eye lobes with sparse yellowish-white pubescence. Area behind lower eye lobes with sparse yellowish-white pubescence close to eye, glabrous on remaining surface. Genae 1.1 times length of lower eye lobe; with moderately sparse yellowish-white pubescence, except glabrous apex. Antennal tubercles with abundant yellowish pubescence with white pubescence interspersed. Wide central area of postclypeus with abundant, bristly yellowish-white pubescence not obscuring integument, with long white setae interspersed; sides with one long, erect dark brown seta. Sides of postclypeus glabrous. Base of anteclypeus with sparse, moderately long yellowish-white setae forming tufts, setae yellower laterally. Labrum with abundant yellowish-white pubescence not obscuring integument, pubescence denser laterally, except glabrous central area of anterior third; with long, erect dark-brown setae interspersed on pubescent area; anterior margin with fringe of golden setae. Gulamentum smooth, glabrous, except sparse whitish pubescence on intermaxillary process. Distance between upper eye lobes 0.23 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.57 times distance between outer margins of eyes. Antennae 1.65 times elytral length, reaching elytral apex at base of antennomere VIII. Scape with dense, irregular pubescent ring on posterior half, pubescence dense, pale-yellow dorsally and laterally, sparser, whitish ventrally, and sparse, thick, decumbent yellowish-white setae throughout, setae whiter ventrally. Pedicel with brown pubescence not obscuring integument and a few short, decumbent white setae interspersed. Dorsal surface of antennomeres III–IV with dark-brown pubescence not obscuring integument, irregular yellowish pubescent macula on basal 5/6 of III and basal 4/5 of IV, and sparse, thick, decumbent white setae interspersed, except on apical area; sides and ventral surface mostly with abundant whitish pubescence partially obscuring integument, except apical sixth of III and apical fifth of IV with dark-brown pubescence not obscuring integument. Reddish-brown area of antennomeres V–X with dense yellowish-white pubescence, whiter laterally and ventrally, and dark-brown pubescence not obscuring integument on remaining surface; antennomere XI with dark-brown pubescence not obscuring integument, and short, bristly whitish setae interspersed. Antennal formula based on length of antennomere III: scape = 0.87; pedicel = 0.10; IV = 0.77; V = 0.53; VI = 0.43; VII = 0.39; VIII = 0.36; IX = 0.33; X = 0.30; XI = 0.28.
</paragraph>
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<paragraph id="8BF0161C7555FFE1FF453A11E19AF80D" blockId="8.[151,1436,1861,1922]" pageId="8" pageNumber="175">
<emphasis id="B93BCA0E7555FFE1FF453A11E346F8D1" bold="true" box="[152,343,1861,1885]" pageId="8" pageNumber="175">FIGURES 17–21</emphasis>
<taxonomicName id="4C4F6D9F7555FFE1FEBC3A12E049F8D1" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[353,600,1862,1886]" class="Insecta" family="Cerambycidae" genus="Pseudastylopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="morrisi" status="sp. nov.">
<emphasis id="B93BCA0E7555FFE1FEBC3A12E049F8D1" box="[353,600,1862,1886]" italics="true" pageId="8" pageNumber="175">Pseudastylopsis morrisi</emphasis>
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<taxonomicNameLabel id="A20877757555FFE1FDBC3A12E0A4F8D2" box="[609,693,1862,1886]" pageId="8" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female:
<emphasis id="B93BCA0E7555FFE1FCA33A11E1B1F8D1" bold="true" box="[894,928,1861,1885]" pageId="8" pageNumber="175">17)</emphasis>
Dorsal habitus;
<emphasis id="B93BCA0E7555FFE1FB8C3A11E662F8D1" bold="true" box="[1105,1139,1861,1885]" pageId="8" pageNumber="175">18)</emphasis>
Ventral habitus;
<emphasis id="B93BCA0E7555FFE1FAF43A11E75AF8D1" bold="true" box="[1321,1355,1861,1885]" pageId="8" pageNumber="175">19)</emphasis>
Lateral habitus;
<emphasis id="B93BCA0E7555FFE1FF333A3DE31EF80D" bold="true" box="[238,271,1897,1921]" pageId="8" pageNumber="175">20)</emphasis>
Head, frontal view;
<emphasis id="B93BCA0E7555FFE1FE3D3A3DE013F80D" bold="true" box="[480,514,1897,1921]" pageId="8" pageNumber="175">21)</emphasis>
Prosternal and mesoventral processes.
</paragraph>
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<paragraph id="8BF0161C7554FFE3FF1A3CABE0F1FC99" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,1401]" lastPageId="10" lastPageNumber="177" pageId="9" pageNumber="176">
<emphasis id="B93BCA0E7554FFE0FF1A3CABE336FD95" bold="true" box="[199,295,511,537]" pageId="9" pageNumber="176">Thorax.</emphasis>
Prothorax wider than long; anterior constriction narrow, well marked; sides with large, rounded protuberance about middle. Pronotum with five gibbosities, one subcircular, loctated on each side of anterior third, one oblique, elongated, located on each side of posterior third, another elongated, longitudinal, located on center of posterior half; sparsely, coarsely punctate, punctures denser centrally on posterior 2/3, forming transverse row on posterior quarter, except smooth central gibbosity; with dense yellowish-white pubescence, pubescence yellower on some irregular areas, except sparse brown pubescence with short, decumbent white setae interspersed on sides of anterior quarter, one subtriangular short macula on each side of central area close to anterior margin with dark brown pubescence not obscuring integument, and a few short, thick white setae interspersed, and wide, longitudinal, sinuous black pubescent band on each side of middle close to posterior margin, most glabrous, longitudinal, sinuous area centrally on posterior half, and small, irregular glabrous areas on remaining central area; with a few long, erect dark-brown setae on posterior half. Sides of prothorax sparsely, coarsely punctate; with dense yellowish pubescence centrally close to pronotum, irregular areas with moderately abundant yellowish-white pubescence partially obscuring integument, with irregular glabrous areas interspersed, except posterior area close to pronotum with large comma-shaped area with sparse dark-brown pubescence and short, decumbent white setae interspersed, and glabrous anterior sulcus. Prosternum and prosternal process with abundant white pubescence not obscuring integument, except glabrous anterior sulcus; narrowest area of prosternal process 0.45 times procoxal width. Sides of ventral surface of meso- and metathorax with abundant yellowish pubescence not obscuring integument, absent or almost absent on some areas; central area with abundant white pubescence not obscuring integument, pubescence sparser on center of mesoventrite and denser on mesoventral process; sides of mesoventral process convergent from base to apical fifth, then distinctly widened; narrowest area of mesoventral process 0.72 times mesocoxal width; apex of mesoventral process almost as wide as one mesocoxa. Scutellum with sparse brownish pubescence and short, thick, decumbent yellowish-white setae interspersed.
<emphasis id="B93BCA0E7554FFE0FCBA39A7E1ABFA81" bold="true" box="[871,954,1267,1293]" pageId="9" pageNumber="176">Elytra.</emphasis>
Subparallel-sided on anterior half, slightly constricted on center of this area, then gradually narrowed toward apex, more distinctly on posterior quarter; apex slightly obliquely truncate; centrobasal crest moderately elevated, elongated, irregularly tuberculate dorsally; moderately abundantly, coarsely punctate on anterior half, punctures slightly finer and sparser toward apex on posterior half; with small, moderately abundant tubercles throughout, nearly all tubercle with short blackish seta on apex; with dense yellowish-white pubescence, irregularly yellowish on some areas, except: humeri subglabrous, with short, thick, decumbent whitish setae; sides of basal half with abundant brownish pubescence not obscuring integument, and short, decumbent, thick white setae interspersed; some tubercles with dense, bristly pale-yellow pubescence; sides of posterior half with dense yellowish-white pubescence, pubescence yellower on irregular areas, and subglabrous areas with short, decumbent, thick white setae interspersed; circum-scutellar area with rectangular area with minute dark brown pubescence not obscuring integument and a few short, decumbent, thick white setae interspersed; wide, longitudinal band on dorsal surface of anterior third with minute dark-brown pubescence not obscuring integument, fused with circum-scutellar area; large, irregular area laterally before middle of dorsal surface with minute dark-brown pubescence not obscuring integument; longitudinal, wide band with minute dark-brown pubescence not obscuring integument, fused to anterior longitudinal band and irregular lateral areas with similar pubescence, tubercles on this band with yellowish pubescence; narrow band with minute dark-brown pubescence not obscuring integument close to suture, from scutellum to middle, then distinctly widened, J-shaped just after middle of elytra, apical side of this area acutely, slightly projected forward, area before apical expansion with short, triangular projections, and tubercles on expanded apical area with dense yellowish pubescence; apical quarter of dorsal surface with irregular, wide, transverse band with minute dark pubescence not obscuring integument, arc-shaped considering both elytra; irregular, arc-shaped band with sparse, short, thick, decumbent yellowish-white setae on side of posterior quarter, subfused to the previous band, and fused with the area with the same setae close to epipleural margin; another macula with sparse, short, thick, decumbent yellowish-white setae on side near apex, fused with the area with the same setae close to epipleural margin.
<emphasis id="B93BCA0E7557FFE3FCB33D8BE1BCFF75" bold="true" box="[878,941,223,249]" pageId="10" pageNumber="177">Legs.</emphasis>
Femoral peduncles with areas with abundant white pubescence and areas with sparse white pubescence, and short, thick, decumbent white setae interspersed throughout; femoral clubs with dense pale-yellow pubescence on part of dorsal surface and inner surface, remaining surface mostly with sparse white pubescence, and with short, thick, decumbent white setae throughout. Protibiae with two dense pale-yellow pubescent arc covering dorsal and lateral surfaces, one basally, another wider located about central third; remaining dorsal and lateral surfaces on basal half with sparse, thick, decumbent pale-yellow setae; ventral surface on basal half with moderately abundant whitish pubescence not obscuring integument; apical third of dorsal and lateral surfaces with sparse dark-brown pubescence and short, thick, decumbent yellowish setae interspersed; apical half of ventral surface with abundant, bristly, brown pubescence. Meso- and metatibiae with dense pale-yellow pubescent arc basally covering dorsal and lateral surfaces; with dense pale-yellow pubescent ring about central third; remaining surface of basal third with sparse whitish pubescence and short, thick, decumbent yellowish setae interspersed; apical third with abundant dark-brown pubescence partially obscuring integument, denser and bristly ventrally, and short, thick, decumbent yellowish setae interspersed dorsally and laterally; dorsal surface of mesotibiae with short, erect, thick dark-brown setae. Dorsal surface of tarsi with dark-brown pubescence not obscuring integument and short, thick, decumbent yellowish-white setae interspersed on some tarsomeres; metatarsomere I slightly shorter than II–III together.
</paragraph>
<paragraph id="8BF0161C7557FFE3FF1A3E4AE736FCD1" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF1A3E4AE352FCB4" bold="true" box="[199,323,798,824]" pageId="10" pageNumber="177">Abdomen.</emphasis>
Ventrites with abundant white pubescence not obscuring integument, sparser on center of ventrites 1–2 and posterior half of ventrite 5, absent on apex of ventrites 1–4. Apex of ventrite 5 slightly concave.
</paragraph>
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<emphasis id="B93BCA0E7557FFE3FF1A3E32E3B5FC0C" bold="true" box="[199,420,870,896]" pageId="10" pageNumber="177">Dimensions (mm).</emphasis>
Total length, 11.15; prothoracic length, 2.10; anterior prothoracic width, 2.15; posterior prothoracic width, 2.70; maximum prothoracic width, 2.90; humeral width, 4.30; elytral length, 7.85.
</paragraph>
</subSubSection>
<subSubSection id="C35545977557FFE3FF1A3EFBE0E5FC61" pageId="10" pageNumber="177" type="materials_examined">
<paragraph id="8BF0161C7557FFE3FF1A3EFBE0E5FC61" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF1A3EFBE367FC44" bold="true" box="[199,374,942,969]" pageId="10" pageNumber="177">Type material.</emphasis>
<materialsCitation id="3B271C417557FFE3FEA33EFBE0E5FC61" collectingDate="2004-06-17" collectionCode="FSCA" collectorName="D. T. Homas &amp; D. Robacker" country="Honduras" location="Nat. [National] Park" municipality="Pico Pijol" pageId="10" pageNumber="177" specimenCount="1" stateProvince="Yoro" typeStatus="holotype">
<typeStatus id="54F4A8BE7557FFE3FEA33EFBE3F7FC45" box="[382,486,943,969]" pageId="10" pageNumber="177" type="holotype">Holotype</typeStatus>
female from
<collectingCountry id="F358568C7557FFE3FD5F3EFBE131FC45" box="[642,800,943,969]" name="Honduras" pageId="10" pageNumber="177">HONDURAS</collectingCountry>
,
<emphasis id="B93BCA0E7557FFE3FCF03EFBE171FC44" box="[813,864,943,968]" italics="true" pageId="10" pageNumber="177">
<collectingRegion id="498BD8FE7557FFE3FCF03EFBE171FC44" box="[813,864,943,968]" country="Honduras" name="Yoro" pageId="10" pageNumber="177">Yoro</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B948C667557FFE3FCB23EFBE1F3FC45" box="[879,994,943,969]" pageId="10" pageNumber="177">Pico Pijol</collectingMunicipality>
,
<location id="8E9040C77557FFE3FC333EFBE6C5FC45" LSID="urn:lsid:plazi:treatment:03E6A70A755AFFE3FF4A3EBFE3E5FAF5:8E9040C77557FFE3FC333EFBE6C5FC45" box="[1006,1236,943,969]" country="Honduras" municipality="Pico Pijol" name="Nat. [National] Park" pageId="10" pageNumber="177" stateProvince="Yoro">Nat. [National] Park</location>
<date id="FFF130DC7557FFE3FB013EFBE74FFC45" box="[1244,1374,943,969]" pageId="10" pageNumber="177" value="2004-06-17">
<collectingDate id="EFB5C9347557FFE3FB013EFBE74FFC45" box="[1244,1374,943,969]" pageId="10" pageNumber="177" value="2004-06-17">17.VI.2004</collectingDate>
</date>
,
<collectorName id="26BA73CA7557FFE3FAB63EFBE2F7FC61" pageId="10" pageNumber="177">D.T. Homas</collectorName>
&amp;
<collectorName id="26BA73CA7557FFE3FED73E87E389FC61" box="[266,408,979,1005]" pageId="10" pageNumber="177">D. Robacker</collectorName>
leg. (
<collectionCode id="ED5E8ED97557FFE3FE0A3E87E030FC61" box="[471,545,979,1005]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" pageId="10" pageNumber="177">FSCA</collectionCode>
, formerly RFMC).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35545977557FFE3FF1A3EA3E190FBB9" pageId="10" pageNumber="177" type="etymology">
<paragraph id="8BF0161C7557FFE3FF1A3EA3E190FBB9" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF1A3EA3E35FFB9D" bold="true" box="[199,334,1015,1041]" pageId="10" pageNumber="177">Etymology.</emphasis>
The new species is dedicated to Roy F. Morris II for his friendship, constant sending of specimens for study, and the donation of the
<typeStatus id="54F4A8BE7557FFE3FDCC394FE062FBB9" box="[529,627,1051,1077]" pageId="10" pageNumber="177" type="holotype">holotype</typeStatus>
to the FSCA collection.
</paragraph>
</subSubSection>
<subSubSection id="C35545977557FFE3FF1A396BE3E5FAF5" pageId="10" pageNumber="177" type="discussion">
<paragraph id="8BF0161C7557FFE3FF1A396BE3E5FAF5" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="177">
<emphasis id="B93BCA0E7557FFE3FF1A396BE32AFBD5" bold="true" box="[199,315,1087,1113]" pageId="10" pageNumber="177">Remarks.</emphasis>
<taxonomicName id="4C4F6D9F7557FFE3FE95396BE048FBD4" authorityName="Santos-Silva &amp; Bezark" authorityYear="2024" box="[328,601,1087,1113]" class="Insecta" family="Cerambycidae" genus="Pseudastylopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="species" species="morrisi" status="sp. nov.">
<emphasis id="B93BCA0E7557FFE3FE95396BE048FBD4" box="[328,601,1087,1113]" italics="true" pageId="10" pageNumber="177">Pseudastylopsis morrisi</emphasis>
</taxonomicName>
<emphasis id="B93BCA0E7557FFE3FDBB396BE0D4FBD5" bold="true" box="[614,709,1087,1113]" pageId="10" pageNumber="177">
<taxonomicNameLabel id="A20877757557FFE3FDBB396BE0D4FBD5" box="[614,709,1087,1113]" pageId="10" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C4F6D9F7557FFE3FCB3396BE772FBD5" authority="Perez-Flores &amp; Santos-Silva, 2021" authorityName="Perez-Flores &amp; Santos-Silva" authorityYear="2021" box="[878,1379,1087,1113]" class="Insecta" family="Cerambycidae" genus="Pseudastylopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="species" species="alba">
<emphasis id="B93BCA0E7557FFE3FCB3396BE1D0FBD5" box="[878,961,1087,1113]" italics="true" pageId="10" pageNumber="177">P. alba</emphasis>
Pérez-Flores &amp;
<bibRefCitation id="EFDE6BED7557FFE3FB54396BE772FBD5" author="Santos-Silva, A. &amp; Devesa, S." box="[1161,1379,1087,1113]" pageId="10" pageNumber="177" pagination="54 - 71" refId="ref11527" refString="Santos-Silva, A., &amp; Devesa, S. (2021) Descriptions, transference, and new records in Lamiinae from Nicaragua (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, 39, 54 - 71. https: // doi. org / 10.11646 / zootaxa. 4942.4.5" type="journal article" year="2021">Santos-Silva, 2021</bibRefCitation>
</taxonomicName>
(see photographs on
<bibRefCitation id="EFDE6BED7557FFE3FE923937E3CEFBF1" author="Bezark, L. G." box="[335,479,1123,1149]" pageId="10" pageNumber="177" refId="ref10785" refString="Bezark, L. G. (2024) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / byciddb / wdefault. asp? w = n / (accessed 8 June 2024)" type="url" year="2024">Bezark 2024</bibRefCitation>
and original description), but differs as follows: pubescence on frons not obscuring integument; scape with short area with dense light pubescence; scape, femora, and tibiae with abundant, short, and tick yellowish-white setae throughout; circum-scutellar area and areas on center of the dorsal surface without dense light pubescence; and narrowest area of mesoventral process about 0.7 times mesocoxal width. In
<taxonomicName id="4C4F6D9F7557FFE3FAFB399BE762FB65" authorityName="Perez-Flores &amp; Santos-Silva" authorityYear="2021" box="[1318,1395,1231,1257]" class="Insecta" family="Cerambycidae" genus="Pseudastylopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="species" species="alba">
<emphasis id="B93BCA0E7557FFE3FAFB399BE762FB65" box="[1318,1395,1231,1257]" italics="true" pageId="10" pageNumber="177">P. alba</emphasis>
</taxonomicName>
the pubescence on frons partially obscuring the integument, scape has large areas with dense light pubescence, scape, femora, and tibia do not have abundant, short, and tick yellowish-white setae, circum-scutellar area and the entire center of the dorsal surface with dense light pubescence, and the narrowest area of the mesoventral process about 0.5 times the mesocoxal width.
</paragraph>
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</treatment>
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<paragraph id="8BF0161C755CFFE8FF4A39F0E049FB32" blockId="1.[151,600,1188,1251]" box="[151,600,1188,1215]" pageId="1" pageNumber="168">
<heading id="D0B8A170755CFFE8FF4A39F0E049FB32" bold="true" box="[151,600,1188,1215]" fontSize="11" level="1" pageId="1" pageNumber="168" reason="1">
<taxonomicName id="4C4F6D9F755CFFE8FF4A39F0E049FB32" ID-CoL="3TX96" authority="Wickham, 1914" authorityName="Wickham" authorityYear="1914" box="[151,600,1188,1215]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="scudderi">
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<emphasis id="B93BCA0E755CFFE8FF4A39F0E39FFB33" bold="true" box="[151,398,1188,1215]" italics="true" pageId="1" pageNumber="168">Leptostylus scudderi</emphasis>
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<paragraph id="8BF0161C755CFFE8FF4A399CE2FAFB6F" blockId="1.[151,600,1188,1251]" box="[151,235,1224,1251]" pageId="1" pageNumber="168">
(
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)
</paragraph>
</subSubSection>
<subSubSection id="C3554597755CFFE8FF4A3859E058FAA9" box="[151,585,1293,1317]" pageId="1" pageNumber="168" type="reference_group">
<paragraph id="8BF0161C755CFFE8FF4A3859E058FAA9" blockId="1.[151,585,1293,1317]" box="[151,585,1293,1317]" pageId="1" pageNumber="168">
<treatmentCitationGroup id="AB5F3132755CFFE8FF4A3859E058FAA9" box="[151,585,1293,1317]" pageId="1" pageNumber="168">
<treatmentCitation id="0AEE300D755CFFE8FF4A3859E054FAA9" author="Wickham, H. F." box="[151,581,1293,1317]" page="469" pageId="1" pageNumber="168" year="1914">
<taxonomicName id="4C4F6D9F755CFFE8FF4A3859E054FAA9" ID-CoL="3TX96" authority="Wickham, 1914: 469" authorityName="Wickham" authorityPageNumber="469" authorityYear="1914" box="[151,581,1293,1317]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="scudderi">
<emphasis id="B93BCA0E755CFFE8FF4A3859E379FAA9" box="[151,360,1293,1317]" italics="true" pageId="1" pageNumber="168">Leptostylus scudderi</emphasis>
<bibRefCitation id="EFDE6BED755CFFE8FEB33859E054FAA9" author="Wickham, H. F." box="[366,581,1293,1317]" pageId="1" pageNumber="168" pagination="423 - 494" refId="ref11581" refString="Wickham, H. F. (1914) New Miocene Coleoptera from the Florissant. Bulletin of the Museum of Comparative Zoology, 58 (11), 423 - 494. https: // doi. org / 10.5962 / bhl. title. 28703" type="journal article" year="1914">Wickham, 1914: 469</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3554597755CFFE8FF4A3807E14EF8CD" pageId="1" pageNumber="168" type="description">
<paragraph id="8BF0161C755CFFE8FF4A3807E14EF8CD" blockId="1.[151,1437,1363,2038]" pageId="1" pageNumber="168">
Original description: “Form moderately elongated. Head with prominent jaws. Antennae slender, about one and three times the length of the body, first joint long, subcylindrical, second short, third barely longer than the first, fourth subequal to the third, fifth, and sixth a little shorter, the remainder not distinctly separable. There is no visible antennal vestiture. Prothorax nearly twice as broad as long, sides arcuate with no defined spine nor tubercle. Elytra long, four and a half times the prothoracic length, strongly tapering from about the middle to the apices which are moderately sharply pointed but unarmed. Legs lacking except one which probably belongs to the middle pair; this is of moderate length and stoutness, tarsal joints ill defined. Length, from front of head to tip of abdomen,
<quantity id="4CB7BBF9755CFFE8FAF83B7FE788F9CA" box="[1317,1433,1579,1606]" metricMagnitude="-2" metricUnit="m" metricValue="1.135" pageId="1" pageNumber="168" unit="mm" value="11.35">11.35 mm</quantity>
. Described from a single specimen.
<typeStatus id="54F4A8BE755CFFE8FDFB3B04E073F9E6" box="[550,610,1616,1642]" pageId="1" pageNumber="168">Type</typeStatus>
.—No. 2,598 M. C. Z. Florissant, Col. (No. 929 S. H. Scudder Col.). The facies of this insect are entirely that of a
<taxonomicName id="4C4F6D9F755CFFE8FDBB3B27E0F8F901" ID-CoL="5DF7" authorityName="LeConte" authorityYear="1852" box="[614,745,1651,1677]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">Leptostylus</taxonomicName>
[
<emphasis id="B93BCA0E755CFFE8FD273B20E108F901" box="[762,793,1652,1677]" italics="true" pageId="1" pageNumber="168">sic</emphasis>
,
<taxonomicName id="4C4F6D9F755CFFE8FCF53B27E1B9F901" ID-CoL="5DF7" authorityName="LeConte" authorityYear="1852" box="[808,936,1651,1677]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E755CFFE8FCF53B27E1B9F901" box="[808,936,1651,1677]" italics="true" pageId="1" pageNumber="168">Leptostylus</emphasis>
</taxonomicName>
], with which genus it agrees in the antennal and thoracic structures. The long antennae are in accord with those of
<taxonomicName id="4C4F6D9F755CFFE8FC603BCCE63AF93D" ID-CoL="3TWSM" box="[957,1067,1687,1713]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="biustus">
<emphasis id="B93BCA0E755CFFE8FC603BCCE63AF93D" box="[957,1067,1687,1713]" italics="true" pageId="1" pageNumber="168">L. biustus</emphasis>
</taxonomicName>
[currently in
<taxonomicName id="4C4F6D9F755CFFE8FB173BC3E2C3F95A" ID-CoL="7PS4" authority="Dillon, 1956" authorityName="Dillon" authorityYear="1956" class="Insecta" family="Cerambycidae" genus="Styloleptus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E755CFFE8FB173BC3E754F93D" box="[1226,1349,1687,1713]" italics="true" pageId="1" pageNumber="168">Styloleptus</emphasis>
Dillon, 1956
</taxonomicName>
] and
<taxonomicName id="4C4F6D9F755CFFE8FECD3BE8E3BFF959" ID-CoL="6PJQV" box="[272,430,1723,1749]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="terraecolor">
<emphasis id="B93BCA0E755CFFE8FECD3BE8E3BFF959" box="[272,430,1723,1749]" italics="true" pageId="1" pageNumber="168">L. terraecolor</emphasis>
</taxonomicName>
[currently in
<taxonomicName id="4C4F6D9F755CFFE8FD953BEFE195F95A" ID-CoL="5DF5" authority="Dillon, 1956" authorityName="Dillon" authorityYear="1956" box="[584,900,1723,1750]" class="Insecta" family="Cerambycidae" genus="Leptostylopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E755CFFE8FD953BEFE0F9F959" box="[584,744,1723,1749]" italics="true" pageId="1" pageNumber="168">Leptostylopsis</emphasis>
Dillon, 1956
</taxonomicName>
]. The coloration seems to have been a mottling, arranged in irregular transverse bands, the best marked of which are in the form of two rather broad postmedian fasciae. It will be noted that the thoracic sides, as preserved, are not symmetrical, the left one showing about the same shape as in
<taxonomicName id="4C4F6D9F755CFFE8FE873A7CE3D7F8CD" ID-CoL="3TWSM" box="[346,454,1831,1857]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="biustus">
<emphasis id="B93BCA0E755CFFE8FE873A7CE3D7F8CD" box="[346,454,1831,1857]" italics="true" pageId="1" pageNumber="168">L. biustus</emphasis>
</taxonomicName>
, while the other is simply rounded.”
</paragraph>
</subSubSection>
<subSubSection id="C3554597755CFFEBFF1A3A1FE702FECD" lastPageId="2" lastPageNumber="169" pageId="1" pageNumber="168" type="discussion">
<paragraph id="8BF0161C755CFFEBFF1A3A1FE702FECD" blockId="1.[151,1437,1363,2038]" lastBlockId="2.[151,1437,151,321]" lastPageId="2" lastPageNumber="169" pageId="1" pageNumber="168">
<emphasis id="B93BCA0E755CFFE8FF1A3A1FE32AF8E9" bold="true" box="[199,315,1867,1893]" pageId="1" pageNumber="168">Remarks.</emphasis>
The original description and drawing (
<figureCitation id="13740A99755CFFE8FCDD3A1FE152F8E9" box="[768,835,1867,1893]" captionStart="FIGURE 1" captionStartId="2.[152,255,1485,1509]" captionTargetBox="[451,1136,342,1458]" captionTargetId="figure-142@2.[451,1136,342,1458]" captionTargetPageId="2" captionText="FIGURE 1. Leptostylus scudderi Wickham, 1914, holotype, dorsal habitus. From Wickham (1914)." figureDoi="http://doi.org/10.5281/zenodo.14390238" httpUri="https://zenodo.org/record/14390238/files/figure.png" pageId="1" pageNumber="168">Fig. 1</figureCitation>
) do not allow considering
<taxonomicName id="4C4F6D9F755CFFE8FBA23A18E710F8E9" authorityName="Wickham" authorityYear="1914" box="[1151,1281,1867,1894]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="169" phylum="Arthropoda" rank="species" species="scudderi">
L.
<emphasis id="B93BCA0E755CFFE8FB7C3A1FE710F8E9" box="[1185,1281,1867,1893]" italics="true" pageId="1" pageNumber="168">scudderi</emphasis>
</taxonomicName>
as belonging to
<taxonomicName id="4C4F6D9F755CFFE8FF683A3BE324F805" authorityName="LeConte" authorityYear="1852" box="[181,309,1903,1929]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E755CFFE8FF683A3BE324F805" box="[181,309,1903,1929]" italics="true" pageId="1" pageNumber="168">Leptostylus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4F6D9F755CFFE8FE983A3BE3F4F805" authorityName="Dillon" authorityYear="1956" box="[325,485,1903,1929]" class="Insecta" family="Cerambycidae" genus="Leptostylopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E755CFFE8FE983A3BE3F4F805" box="[325,485,1903,1929]" italics="true" pageId="1" pageNumber="168">Leptostylopsis</emphasis>
</taxonomicName>
, or
<taxonomicName id="4C4F6D9F755CFFE8FDC93A3BE09EF805" authorityName="Dillon" authorityYear="1956" box="[532,655,1903,1929]" class="Insecta" family="Cerambycidae" genus="Styloleptus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B93BCA0E755CFFE8FDC93A3BE09EF805" box="[532,655,1903,1929]" italics="true" pageId="1" pageNumber="168">Styloleptus</emphasis>
</taxonomicName>
. This is because currently, there are no species in these three genera with the apex of the elytra uniformly acute and divergent from apical fifth. The drawing in the original description shows a single femur on the right side and it is not clavate, while the femora are pedunculate-clavate in these genera. However, photographs of the
<typeStatus id="54F4A8BE755CFFE8FE3D3A8FE053F879" box="[480,578,2011,2037]" pageId="1" pageNumber="168" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFDE6BED755CFFE8FD8D3A8FE107F879" author="MCZBASE" box="[592,790,2011,2037]" pageId="1" pageNumber="168" refId="ref11213" refString="MCZBASE, (2024) The database of the zoological collections. Museum of Comparative Zoology - Harvard Universty. Available from: https: // mczbase. mcz. harvard. edu / SpecimenSearch. cfm? collection _ id = 1 (accessed 14 June 2024)" type="url" year="2024">MCZBASE 2024</bibRefCitation>
) show that the
<typeStatus id="54F4A8BE755CFFE8FC633A8FE631F879" box="[958,1056,2011,2037]" pageId="1" pageNumber="168" type="holotype">holotype</typeStatus>
has two femora, one on each side, and that in the left side appears to be clavate. Additionally, the original description does not inform if there are small tubercles on the elytra, which are present in the three genera mentioned by
<bibRefCitation id="EFDE6BED755FFFEBFC0B3DEFE687FF59" author="Wickham, H. F." box="[982,1174,187,213]" pageId="2" pageNumber="169" pagination="423 - 494" refId="ref11581" refString="Wickham, H. F. (1914) New Miocene Coleoptera from the Florissant. Bulletin of the Museum of Comparative Zoology, 58 (11), 423 - 494. https: // doi. org / 10.5962 / bhl. title. 28703" type="journal article" year="1914">Wickham (1914)</bibRefCitation>
. It is not possible to see if the tubercles are or are not present in the
<typeStatus id="54F4A8BE755FFFEBFD553D8BE0FFFF75" box="[648,750,223,249]" pageId="2" pageNumber="169" type="holotype">holotype</typeStatus>
. Lateral tubercles of the prothorax rounded as in
<taxonomicName id="4C4F6D9F755FFFEBFAC03D8BE78DFF75" authorityName="Wickham" authorityYear="1914" box="[1309,1436,223,249]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="scudderi">
<emphasis id="B93BCA0E755FFFEBFAC03D8BE78DFF75" box="[1309,1436,223,249]" italics="true" pageId="2" pageNumber="169">L. scudderi</emphasis>
</taxonomicName>
are not rare in
<taxonomicName id="4C4F6D9F755FFFEBFEE53C57E3CBFE91" authorityName="Blanchard" authorityYear="1845" box="[312,474,259,285]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="185" phylum="Arthropoda" rank="tribe" tribe="Acanthocinini">Acanthocinini</taxonomicName>
. Therefore, we are transferring
<taxonomicName id="4C4F6D9F755FFFEBFCE43C57E1A7FE91" authorityName="Wickham" authorityYear="1914" box="[825,950,259,285]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="scudderi">
<emphasis id="B93BCA0E755FFFEBFCE43C57E1A7FE91" box="[825,950,259,285]" italics="true" pageId="2" pageNumber="169">L. scudderi</emphasis>
</taxonomicName>
to
<emphasis id="B93BCA0E755FFFEBFC073C57E665FE91" box="[986,1140,259,285]" italics="true" pageId="2" pageNumber="169">incertae sedis</emphasis>
in
<taxonomicName id="4C4F6D9F755FFFEBFB4B3C57E724FE91" authorityName="Blanchard" authorityYear="1845" box="[1174,1333,259,285]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="185" phylum="Arthropoda" rank="tribe" tribe="Acanthocinini">Acanthocinini</taxonomicName>
since we have not been able to find a reliable feature that allows us to be certain about the genus of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

116
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<subSubSection id="06BE4CA88618D80FDB06FE1CBA43FE32" box="[869,1400,430,457]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="4E1B1F238618D80FDB06FE1CBA43FE32" blockId="37.[869,1400,430,457]" box="[869,1400,430,457]" pageId="37" pageNumber="38">
<heading id="1553A84F8618D80FDB06FE1CBA43FE32" bold="true" box="[869,1400,430,457]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="3">
<emphasis id="7CD0C3318618D80FDB06FE1CBA43FE32" bold="true" box="[869,1400,430,457]" pageId="37" pageNumber="38">
Genus
<taxonomicName id="89A464A08618D80FDBCCFE1CBA22FE32" ID-CoL="8XPYL" authority="Bernhauer, 1941" authorityName="Bernhauer" authorityYear="1941" box="[943,1305,430,457]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="stat. nov.">
<emphasis id="7CD0C3318618D80FDBCCFE1CBB59FE33" bold="true" box="[943,1122,430,457]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
Bernhauer, 1941
</taxonomicName>
<taxonomicNameLabel id="67E37E4A8618D80FDD7EFE1CBA43FE32" box="[1309,1400,430,456]" pageId="37" pageNumber="38">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="06BE4CA88618D80CDC54FE50BEC6FA5B" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="4E1B1F238618D80FDC54FE50BB9DFE06" blockId="37.[1079,1190,482,508]" box="[1079,1190,482,508]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDC54FE50BB9DFE06" bold="true" box="[1079,1190,482,508]" pageId="37" pageNumber="38">
(
<figureCitation id="D69F03A68618D80FDC21FE50BBA0FE06" box="[1090,1179,482,508]" captionStart="Figure 15" captionStartId="38.[129,194,1066,1090]" captionTargetBox="[129,1472,144,1039]" captionTargetId="figure-496@38.[129,1473,144,1039]" captionTargetPageId="38" captionText="Figure 15. Habitus of Amblyopinini genera.A, Apoquedius aeneipennis (Fairmaire and Germain, 1862). B, Lienturius leiocephalus (Solier, 1849). C, Sphingoquedius discoideus (Fairmaire and Germain, 1862)." figureDoi="http://doi.org/10.5281/zenodo.14443987" httpUri="https://zenodo.org/record/14443987/files/figure.png" pageId="37" pageNumber="38">Fig. 15C</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238618D80FDB49FDABBCB8FC90" blockId="37.[809,1460,537,874]" pageId="37" pageNumber="38">
<taxonomicName id="89A464A08618D80FDB49FDABBB81FDCB" authority="Bernhauer 1941: 27" authorityName="Bernhauer" authorityPageNumber="27" authorityYear="1941" box="[810,1210,537,561]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDB49FDABBCF9FDCB" box="[810,962,537,561]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
Bernhauer 1941: 27
</taxonomicName>
(original description; species included:
<taxonomicName id="89A464A08618D80FDB8DFD8ABB09FDAA" authorityName="Bernhauer" authorityYear="1941" box="[1006,1074,568,592]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="7CD0C3318618D80FDB8DFD8ABB09FDAA" box="[1006,1074,568,592]" italics="true" pageId="37" pageNumber="38">strandi</emphasis>
</taxonomicName>
); Klimaszewski
<emphasis id="7CD0C3318618D80FDC89FD8BBA21FDAA" box="[1258,1306,568,592]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. (1996): 151 (
<taxonomicName id="89A464A08618D80FDB56FDE5BCF6FD95" authorityName="Bernhauer" authorityYear="1941" box="[821,973,599,623]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDB56FDE5BCF6FD95" box="[821,973,599,623]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
</taxonomicName>
; endemic to
<collectingCountry id="36B35FB38618D80FDC35FDEABBDDFD95" box="[1110,1254,599,623]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
); Solodovnikov and Schomann 2009: 34 (included in phylogenetic analysis); Jenkins Shaw and Solodovnikov (2016): 22 (absent on Lord Howe Island); Solodovnikov and Brunke (2016): 43 (new combination); Solodovnikov and Jenkins Shaw (2017): 318 (notes on relationship); Jenkins Shaw
<emphasis id="7CD0C3318618D80FDC2AFD47BB4AFCF6" box="[1097,1137,756,780]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2017: 714 (included in phylogenetic analysis); Jenkins Shaw
<emphasis id="7CD0C3318618D80FDC15FCA6BBA5FCD1" box="[1142,1182,787,811]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2020a: 45 (notes on relationship); Jenkins Shaw
<emphasis id="7CD0C3318618D80FDC7CFC81BB73FCB1" box="[1055,1096,819,843]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2020b: 2 (description of new fossil species).
</paragraph>
<paragraph id="4E1B1F238618D80FDB49FC27BA7AFC56" blockId="37.[810,1345,916,940]" box="[810,1345,916,940]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDB49FC27BC9DFC56" box="[810,934,917,940]" italics="true" pageId="37" pageNumber="38">
<typeStatus id="911FA1818618D80FDB49FC27BC61FC56" box="[810,858,917,940]" pageId="37" pageNumber="38">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="89A464A08618D80FDBC8FC26BA07FC56" authority="Bernhauer, 1941" authorityName="Bernhauer" authorityYear="1941" box="[939,1340,916,940]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="7CD0C3318618D80FDBC8FC26BBB6FC56" box="[939,1165,916,940]" italics="true" pageId="37" pageNumber="38">Sphingoquedius strandi</emphasis>
Bernhauer, 1941
</taxonomicName>
.
</paragraph>
<paragraph id="4E1B1F238618D80FDB49FC7FBAB6FBB9" blockId="37.[810,1459,973,1091]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDB49FC7FBCFCFC1F" box="[810,967,973,997]" italics="true" pageId="37" pageNumber="38">Included species:</emphasis>
<taxonomicName id="89A464A08618D80FDBB2FC7FBA0FFC1F" authority="(Saiz 1971)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" box="[977,1332,973,997]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="brevis" status="comb. nov.">
<emphasis id="7CD0C3318618D80FDBB2FC7FBB83FC1F" bold="true" box="[977,1208,973,997]" italics="true" pageId="37" pageNumber="38">Sphingoquedius brevis</emphasis>
(Sáiz 1971)
</taxonomicName>
<emphasis id="7CD0C3318618D80FDD5AFC7FBA95FC1F" bold="true" box="[1337,1454,973,997]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="67E37E4A8618D80FDD5AFC7FBA95FC1F" box="[1337,1454,973,997]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="89A464A08618D80FDB49FC5FBBD7FBFF" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[810,1260,1004,1029]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="discoideus" status="comb. nov.">
<emphasis id="7CD0C3318618D80FDB49FC5FBC96FBFE" bold="true" box="[810,941,1004,1029]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDB49FC5FBC0CFBFF" bold="true" box="[810,823,1005,1029]" italics="true" pageId="37" pageNumber="38">S</emphasis>
.
<emphasis id="7CD0C3318618D80FDB21FC5EBC96FBFE" bold="true" box="[834,941,1004,1028]" italics="true" pageId="37" pageNumber="38">discoideus</emphasis>
</emphasis>
(Fairmaire and Germain, 1862)
</taxonomicName>
<emphasis id="7CD0C3318618D80FDC93FC5FBA51FBFF" bold="true" box="[1264,1386,1005,1029]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="67E37E4A8618D80FDC93FC5FBA5EFBFF" box="[1264,1381,1005,1029]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
,
</emphasis>
<taxonomicName id="89A464A08618D80FDD0DFC5FBB0FFBDE" authority="Jenkins Shaw et al., 2020 b" authorityName="Jenkins Shaw" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="meto">
<emphasis id="7CD0C3318618D80FDD0DFC5FBA40FBFE" box="[1390,1403,1005,1028]" italics="true" pageId="37" pageNumber="38">S</emphasis>
.
<emphasis id="7CD0C3318618D80FDDE7FC5FBA88FBFE" box="[1412,1459,1005,1028]" italics="true" pageId="37" pageNumber="38">meto</emphasis>
Jenkins Shaw et al., 2020b
</taxonomicName>
,
<taxonomicName id="89A464A08618D80FDC5DFBBEBA22FBDE" authority="(Saiz, 1971)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" box="[1086,1305,1036,1060]" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nanus" status="comb. nov.">
<emphasis id="7CD0C3318618D80FDC5DFBBEBBADFBD9" bold="true" box="[1086,1174,1036,1060]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDC5DFBBEBB71FBDE" bold="true" box="[1086,1098,1036,1060]" italics="true" pageId="37" pageNumber="38">S</emphasis>
.
<emphasis id="7CD0C3318618D80FDC34FBBEBBADFBD9" bold="true" box="[1111,1174,1036,1059]" italics="true" pageId="37" pageNumber="38">nanus</emphasis>
</emphasis>
(Sáiz, 1971)
</taxonomicName>
<emphasis id="7CD0C3318618D80FDD7CFBBEBAADFBDE" bold="true" box="[1311,1430,1036,1060]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="67E37E4A8618D80FDD7CFBBEBAADFBDE" box="[1311,1430,1036,1060]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="89A464A08618D80FDDC2FBBEBB4CFBB9" authority="(Duvivier, 1883)" baseAuthorityName="Duvivier" baseAuthorityYear="1883" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="novaezeelandiae">
<emphasis id="7CD0C3318618D80FDDC2FBBEBA95FBD9" box="[1441,1454,1036,1059]" italics="true" pageId="37" pageNumber="38">S</emphasis>
.
<emphasis id="7CD0C3318618D80FDB49FB99BCFEFBB9" box="[810,965,1067,1091]" italics="true" pageId="37" pageNumber="38">novaezeelandiae</emphasis>
(Duvivier, 1883)
</taxonomicName>
,
<taxonomicName id="89A464A08618D80FDCE3FB9EBAB2FBB9" authority="Bernhauer, 1941" authorityName="Bernhauer" authorityYear="1941" box="[1152,1417,1067,1091]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="7CD0C3318618D80FDCE3FB9EBBB6FBB9" box="[1152,1165,1068,1091]" italics="true" pageId="37" pageNumber="38">S</emphasis>
.
<emphasis id="7CD0C3318618D80FDCFBFB99BBE0FBB9" box="[1176,1243,1067,1091]" italics="true" pageId="37" pageNumber="38">strandi</emphasis>
Bernhauer, 1941
</taxonomicName>
.
</paragraph>
<paragraph id="4E1B1F238618D80FDB49FBD4BA2FFA4D" blockId="37.[810,1460,1125,1463]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDB49FBD4BCABFB87" box="[810,912,1126,1149]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
Ŋe diagnosis of
<taxonomicName id="89A464A08618D80FDC36FBD7BBD6FB87" authorityName="Bernhauer" authorityYear="1941" box="[1109,1261,1125,1149]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDC36FBD7BBD6FB87" box="[1109,1261,1125,1149]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
</taxonomicName>
is likely to change pending phylogenomic results; however, these species can be tentatively placed in
<taxonomicName id="89A464A08618D80FDB9DFB16BBADFB46" authorityName="Bernhauer" authorityYear="1941" box="[1022,1174,1188,1212]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDB9DFB16BBADFB46" box="[1022,1174,1188,1212]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
</taxonomicName>
based on the large eyes and presence of patches of pale radiating setae on abdominal tergites III and IV. American
<taxonomicName id="89A464A08618D80FDC7BFB51BB8BFB01" authorityName="Bernhauer" authorityYear="1941" box="[1048,1200,1251,1275]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDC7BFB51BB8BFB01" box="[1048,1200,1251,1275]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
</taxonomicName>
are easily distinguished from
<taxonomicName id="89A464A08618D80FDB07FAB0BCE9FAE0" box="[868,978,1282,1306]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDB07FAB0BCE9FAE0" box="[868,978,1282,1306]" italics="true" pageId="37" pageNumber="38">Loncovilius</emphasis>
</taxonomicName>
by empodial setae as long as the tarsal claws; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of patches of pale radiating setae on abdominal tergites; sternite III with basal transverse carina converges abruptly at an acute angle with rounded tip (projection sides invaginated).
</paragraph>
<paragraph id="4E1B1F238618D80FDB49FA69BC40F98B" blockId="37.[810,1459,1499,1649]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDB49FA69BC93FA09" box="[810,936,1499,1523]" italics="true" pageId="37" pageNumber="38">Distribution:</emphasis>
Asser the taxonomic changes implemented here,
<taxonomicName id="89A464A08618D80FDB49FA49BCF9F9E9" authorityName="Bernhauer" authorityYear="1941" box="[810,962,1531,1555]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDB49FA49BCF9F9E9" box="[810,962,1531,1555]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
</taxonomicName>
occurs in
<collectingCountry id="36B35FB38618D80FDC5CFA49BBF5F9E9" box="[1087,1230,1531,1555]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
and southern South America. A single fossil species (
<taxonomicName id="89A464A08618D80FDC1BF9A8BA7EF9C8" box="[1144,1349,1562,1586]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="meto">
<emphasis id="7CD0C3318618D80FDC1BF9A8BA7EF9C8" box="[1144,1349,1562,1586]" italics="true" pageId="37" pageNumber="38">Sphingoquedius meto</emphasis>
</taxonomicName>
) is known from the earliest Miocene of
<collectingCountry id="36B35FB38618D80FDC34F988BBDBF9AB" box="[1111,1248,1593,1617]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
(Jenkins Shaw
<emphasis id="7CD0C3318618D80FDDE6F988BA96F9AB" box="[1413,1453,1593,1617]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2020b).
</paragraph>
<paragraph id="4E1B1F238618D80CDB49F922BFE8FB3C" blockId="37.[809,1460,1680,1986]" lastBlockId="38.[129,778,1166,1441]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FDB49F922BCB3F952" box="[810,904,1680,1704]" italics="true" pageId="37" pageNumber="38">Remarks:</emphasis>
Ŋe new combinations implemented here [
<taxonomicName id="89A464A08618D80FDB56F91DBBA9F93D" authority="(Saiz, 1971)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" box="[821,1170,1711,1735]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="7CD0C3318618D80FDB56F91DBB36F93D" box="[821,1037,1711,1735]" italics="true" pageId="37" pageNumber="38">Sphingoquedius nanus</emphasis>
(Sáiz, 1971)
</taxonomicName>
;
<taxonomicName id="89A464A08618D80FDCFDF91DBC51F91D" authority="(Saiz, 1971)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="7CD0C3318618D80FDCFDF91DBA4FF93D" box="[1182,1396,1711,1735]" italics="true" pageId="37" pageNumber="38">Sphingoquedius brevis</emphasis>
(Sáiz, 1971)
</taxonomicName>
;
<taxonomicName id="89A464A08618D80FDBE1F97DBC51F8FC" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="7CD0C3318618D80FDBE1F97DBBB7F91D" box="[898,1164,1743,1767]" italics="true" pageId="37" pageNumber="38">Sphingoquedius discoideus</emphasis>
(Fairmaire and Germain, 1862)
</taxonomicName>
] are supported by the phylogenetic analysis where species of
<taxonomicName id="89A464A08618D80FDB16F8BFBB36F8DF" authorityName="Bernhauer" authorityYear="1941" box="[885,1037,1805,1829]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDB16F8BFBB36F8DF" box="[885,1037,1805,1829]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
</taxonomicName>
are united based on two unique synapomorphies: empodial setae longer or at least as long as tarsal claws (character 62, state 1; empodial setae missing in
<taxonomicName id="89A464A08618D80FDD23F8FEBA9AF89E" authorityName="Bernhauer" authorityYear="1941" box="[1344,1441,1868,1892]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="7CD0C3318618D80FDD23F8FEBA76F899" box="[1344,1357,1868,1891]" italics="true" pageId="37" pageNumber="38">S</emphasis>
.
<emphasis id="7CD0C3318618D80FDD3EF8FEBA9AF89E" box="[1373,1441,1868,1892]" italics="true" pageId="37" pageNumber="38">strandi</emphasis>
</taxonomicName>
); abdomen with tergites III and IV bearing patches of pale radiating setae (character 93, state 2; also see: Jenkins Shaw
<emphasis id="7CD0C3318618D80FDDE2F839BA96F859" box="[1409,1453,1931,1955]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2020b: fig. 2.3); and a single homoplasious synapomorphy: postmandibular ridge located close to eye margin (character 16, state 1).
</paragraph>
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<emphasis id="7CD0C331861BD80CD8E2FB98BFDEFBB9" bold="true" box="[129,229,1066,1091]" pageId="38" pageNumber="39">Figure 15.</emphasis>
Habitus of
<taxonomicName id="89A464A0861BD80CD931FB99BEEDFBB9" authorityName="Seevers" authorityYear="1944" box="[338,470,1067,1091]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="tribe" tribe="Amblyopinini">Amblyopinini</taxonomicName>
genera. A,
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<emphasis id="7CD0C331861BD80CDA5DFB99BC2FFBB8" box="[574,788,1067,1091]" italics="true" pageId="38" pageNumber="39">Apoquedius aeneipennis</emphasis>
(Fairmaire and Germain, 1862)
</taxonomicName>
. B,
<taxonomicName id="89A464A0861BD80CDC06FB99BA87FBB9" authority="(Solier, 1849)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" box="[1125,1468,1067,1091]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="leiocephalus">
<emphasis id="7CD0C331861BD80CDC06FB99BA08FBB9" box="[1125,1331,1067,1091]" italics="true" pageId="38" pageNumber="39">Lienturius leiocephalus</emphasis>
(Solier, 1849)
</taxonomicName>
. C,
<taxonomicName id="89A464A0861BD80CD8F8FBF5BD83FBA5" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[155,696,1095,1119]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="7CD0C331861BD80CD8F8FBF5BEBCFBA5" box="[155,391,1095,1119]" italics="true" pageId="38" pageNumber="39">Sphingoquedius discoideus</emphasis>
(Fairmaire and Germain, 1862)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="4E1B1F23861BD80CD8FFFB7CBEC6FA5B" blockId="38.[129,778,1166,1441]" pageId="38" pageNumber="39">
Numerous described and undescribed species from
<collectingCountry id="36B35FB3861BD80CDABAFB7CBFEFFAFF" name="New Zealand" pageId="38" pageNumber="39">New Zealand</collectingCountry>
and
<collectingCountry id="36B35FB3861BD80CD96FFB5FBE5CFAFF" box="[268,359,1261,1285]" name="Australia" pageId="38" pageNumber="39">Australia</collectingCountry>
may also belong to
<taxonomicName id="89A464A0861BD80CDA21FB5FBDE1FAFF" authorityName="Bernhauer" authorityYear="1941" box="[578,730,1261,1285]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C331861BD80CDA21FB5FBDE1FAFF" box="[578,730,1261,1285]" italics="true" pageId="38" pageNumber="39">Sphingoquedius</emphasis>
</taxonomicName>
and will be dealt with separately pending further study. Even though the generic placement of the new combinations implemented here may change further, their placement in
<taxonomicName id="89A464A0861BD80CDA3BFAF9BDCBFA99" authorityName="Bernhauer" authorityYear="1941" box="[600,752,1355,1379]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C331861BD80CDA3BFAF9BDCBFA99" box="[600,752,1355,1379]" italics="true" pageId="38" pageNumber="39">Sphingoquedius</emphasis>
</taxonomicName>
is supported by the phylogenetic analysis and can at least be regarded as a step in the right direction.
</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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Coiffait and Sáiz, 1966
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(
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)
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Coiffait and Sáiz 1966: 403 [original description; as subgenus of
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<emphasis id="7CD0C3318618D80FD8E8FCD3BFC2FC83" box="[139,249,865,889]" italics="true" pageId="37" pageNumber="38">Loncovilius</emphasis>
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; species included:
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<emphasis id="7CD0C3318618D80FD9D4FCD0BD1CFC83" box="[439,551,866,889]" italics="true" pageId="37" pageNumber="38">aeneipennis</emphasis>
</taxonomicName>
(as
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<emphasis id="7CD0C3318618D80FDA2DFCD3BD46FC83" box="[590,637,865,889]" italics="true" pageId="37" pageNumber="38">heeri</emphasis>
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),
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<emphasis id="7CD0C3318618D80FDAF0FCD3BDCFFC83" box="[659,756,865,889]" italics="true" pageId="37" pageNumber="38">discoideus</emphasis>
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,
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<emphasis id="7CD0C3318618D80FD812FC33BFFCFC62" box="[113,199,897,920]" italics="true" pageId="37" pageNumber="38">germaini</emphasis>
</taxonomicName>
, and
<taxonomicName id="89A464A08618D80FD89FFC33BE4BFC63" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" box="[252,368,897,921]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="leiocephalus">
<emphasis id="7CD0C3318618D80FD89FFC33BE4BFC63" box="[252,368,897,921]" italics="true" pageId="37" pageNumber="38">leiocephalus</emphasis>
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]; Sáiz 1969: 9 (subgenus of
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<emphasis id="7CD0C3318618D80FDAECFC33BDC8FC62" box="[655,755,897,920]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
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; notes); Sáiz 1971: 382, 383 (subgenus of
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<emphasis id="7CD0C3318618D80FDA58FC13BDA4FC42" box="[571,671,929,952]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
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; characters;
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); Newton, 2022 [synonym of
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<emphasis id="7CD0C3318618D80FDA7FFC0DBD57FC2D" box="[540,620,959,983]" italics="true" pageId="37" pageNumber="38">Quedius</emphasis>
(
<emphasis id="7CD0C3318618D80FDA1EFC0DBDD4FC2D" box="[637,751,959,983]" italics="true" pageId="37" pageNumber="38">Apoquedius</emphasis>
) Scheerpeltz, 1972
</taxonomicName>
].
</paragraph>
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<emphasis id="7CD0C3318618D80FD812FBABBFD6FBCA" box="[113,237,1049,1072]" italics="true" pageId="37" pageNumber="38">
<typeStatus id="911FA1818618D80FD812FBABBF9AFBCA" box="[113,161,1049,1072]" pageId="37" pageNumber="38">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="89A464A08618D80FD894FBAABD45FBCA" authority="(Solier, 1849)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" box="[247,638,1048,1072]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="leiocephalus" status="comb. nov.">
<emphasis id="7CD0C3318618D80FD894FBAABED1FBCA" bold="true" box="[247,490,1048,1072]" italics="true" pageId="37" pageNumber="38">Lienturius leiocephalus</emphasis>
(Solier, 1849)
</taxonomicName>
<emphasis id="7CD0C3318618D80FDAE0FBAABDC1FBCA" bold="true" box="[643,762,1048,1072]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="67E37E4A8618D80FDAE0FBAABDC1FBCA" box="[643,762,1048,1072]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="4E1B1F238618D80FD812FBE0BEF7F967" blockId="37.[112,764,1105,1693]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FD812FBE0BFECFB93" box="[113,215,1106,1129]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
Ŋe phylogenetic relationship of
<taxonomicName id="89A464A08618D80FDA4EFBE0BDAAFB93" authorityName="Coiffait and Saiz" authorityYear="1966" box="[557,657,1106,1129]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDA4EFBE0BDAAFB93" box="[557,657,1106,1129]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
</taxonomicName>
to the rest of the
<taxonomicName id="89A464A08618D80FD8D2FBC3BE04FB73" authorityName="Seevers" authorityYear="1944" box="[177,319,1137,1161]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="tribe" tribe="Amblyopinini">Amblyopinini</taxonomicName>
genera remains unknown, so it is likely that the diagnosis of this genus will change as we wait for the results of our pending phylogenomic research. For the moment the only species of this genus is distinguished based on the combination of the following characters: head with three or more paraocular punctures on each side (not in a row), of which at least one is close to the eye margin and one is far from the eye margin and the remaining paraocular punctures, posterior frontal puncture anterior to temporal puncture; flexible postcoxal hypomeral extension (process) rounded and not interrupted by inferior line; elytra without humeral spines or spine-like setae; two short empodial setae on each tarsomere 5; tergites III–VI with PTBC, sternite III with basal transverse carina converging abruptly at an acute angle with rounded tip.
<taxonomicName id="89A464A08618D80FD9CFF9BBBD2BF9DA" authorityName="Coiffait and Saiz" authorityYear="1966" box="[428,528,1545,1568]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FD9CFF9BBBD2BF9DA" box="[428,528,1545,1568]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="89A464A08618D80FD8C9F995BE23F9C5" box="[170,280,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FD8C9F995BE23F9C5" box="[170,280,1575,1599]" italics="true" pageId="37" pageNumber="38">Loncovilius</emphasis>
</taxonomicName>
by the absence of a complete frontoclypeal suture; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of PTBC in tergites III to VI.
</paragraph>
<paragraph id="4E1B1F238618D80FD812F90DBDEBF92D" blockId="37.[113,720,1727,1751]" box="[113,720,1727,1751]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FD812F90DBFD4F92D" box="[113,239,1727,1751]" italics="true" pageId="37" pageNumber="38">Distribution:</emphasis>
Only known from the Southern zone of
<collectingCountry id="36B35FB38618D80FDAF2F90DBDF0F92D" box="[657,715,1727,1751]" name="Chile" pageId="37" pageNumber="38">Chile</collectingCountry>
.
</paragraph>
<paragraph id="4E1B1F238618D80FD812F95CBBB0FE79" blockId="37.[113,763,1774,1986]" lastBlockId="37.[810,1459,144,387]" pageId="37" pageNumber="38">
<emphasis id="7CD0C3318618D80FD812F95CBFF4F8FC" box="[113,207,1774,1798]" italics="true" pageId="37" pageNumber="38">Remarks:</emphasis>
Ŋe new combination implemented here [
<taxonomicName id="89A464A08618D80FDAF4F95DBE43F8DF" authority="(Solier, 1849)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="leiocephalus">
<emphasis id="7CD0C3318618D80FDAF4F95DBFDEF8DF" italics="true" pageId="37" pageNumber="38">Lienturius leiocephalus</emphasis>
(Solier, 1849)
</taxonomicName>
] is supported by the phylogenetic analysis based on eight homoplasious synapomorphies: right mandible with proximal tooth and distal tooth, distal tooth not bifid, space between proximal and distal teeth narrow and smooth (character 24, state 2); labrum, shape of anterior margin (only sclerotized part, excluding apical membrane where present) entire, without emargination (character 28, state 1); three or more paraocular punctures (character 41, state 2); posterior frontal puncture anterior to temporal puncture (character 42, state 0); pronotum with SLSP (character 55, state 1); protarsi with tarsomeres 1–3 as long as wide (character 63, state 0); tergite VI with PTBC (character 89, state 1); sternite III with basal transverse carina converging abruptly at an acute angle with rounded tip (character 99, state 3).
</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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<emphasis id="7CD0C3318619D80EDBACFDCCBB6EFD63" bold="true" box="[975,1109,638,665]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
Scheerpeltz, 1972
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<emphasis id="7CD0C3318619D80EDC24FD1FBB8FFD3D" bold="true" box="[1095,1204,685,711]" pageId="36" pageNumber="37">
(
<figureCitation id="D69F03A68619D80EDC31FD1FBB92FD3D" box="[1106,1193,685,711]" captionStart="Figure 15" captionStartId="38.[129,194,1066,1090]" captionTargetBox="[129,1472,144,1039]" captionTargetId="figure-496@38.[129,1473,144,1039]" captionTargetPageId="38" captionText="Figure 15. Habitus of Amblyopinini genera.A, Apoquedius aeneipennis (Fairmaire and Germain, 1862). B, Lienturius leiocephalus (Solier, 1849). C, Sphingoquedius discoideus (Fairmaire and Germain, 1862)." figureDoi="http://doi.org/10.5281/zenodo.14443987" httpUri="https://zenodo.org/record/14443987/files/figure.png" pageId="36" pageNumber="37">Fig. 15A</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238619D80EDB5AFD56BA70FCA0" blockId="36.[825,1474,740,858]" pageId="36" pageNumber="37">
<taxonomicName id="89A464A08619D80EDB5AFD56BB93FD06" authority="Scheerpeltz, 1972: 24" authorityName="Scheerpeltz" authorityPageNumber="24" authorityYear="1972" box="[825,1192,740,764]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDB5AFD56BC90FD06" box="[825,939,740,764]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
Scheerpeltz, 1972: 24
</taxonomicName>
, 25 (original description; as subgenus of
<taxonomicName id="89A464A08619D80EDB8CFCB1BB04FCE1" box="[1007,1087,771,795]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDB8CFCB1BB04FCE1" box="[1007,1087,771,795]" italics="true" pageId="36" pageNumber="37">Quedius</emphasis>
</taxonomicName>
; included species:
<taxonomicName id="89A464A08619D80EDD7EFCB6BAB6FCE1" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[1309,1421,772,795]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeneipennis">
<emphasis id="7CD0C3318619D80EDD7EFCB6BAB6FCE1" box="[1309,1421,772,795]" italics="true" pageId="36" pageNumber="37">aeneipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="89A464A08619D80EDB5AFC91BCA3FCC1" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[825,920,803,827]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="piciformis">
<emphasis id="7CD0C3318619D80EDB5AFC91BCA3FCC1" box="[825,920,803,827]" italics="true" pageId="36" pageNumber="37">piciformis</emphasis>
</taxonomicName>
); Herman 2001a: 24 (synonym of
<taxonomicName id="89A464A08619D80EDD21FC91BA8BFCC1" box="[1346,1456,803,827]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDD21FC91BA8BFCC1" box="[1346,1456,803,827]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
</taxonomicName>
); Newton, 2022 (synonym of
<taxonomicName id="89A464A08619D80EDC3AFCF0BA7CFCA0" authorityName="Coiffait &amp; Saiz" authorityYear="1966" box="[1113,1351,834,858]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="subGenus" subGenus="Lienturius">
<emphasis id="7CD0C3318619D80EDC3AFCF0BBFCFCA0" box="[1113,1223,834,858]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318619D80EDCB4FCF1BA00FCA0" box="[1239,1339,835,858]" italics="true" pageId="36" pageNumber="37">Lienturius</emphasis>
)
</taxonomicName>
.
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<paragraph id="4E1B1F238619D80EDB5AFCCEBCCFFC49" blockId="36.[825,1474,891,947]" pageId="36" pageNumber="37">
<emphasis id="7CD0C3318619D80EDB5AFCCEBC95FC69" box="[825,942,892,915]" italics="true" pageId="36" pageNumber="37">
<typeStatus id="911FA1818619D80EDB5AFCCEBC52FC69" box="[825,873,892,915]" pageId="36" pageNumber="37">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="89A464A08619D80EDBD5FCC9BC43FC49" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeneipennis" status="comb. nov.">
<emphasis id="7CD0C3318619D80EDBD5FCC9BB8DFC69" bold="true" box="[950,1206,891,915]" italics="true" pageId="36" pageNumber="37">Apoquedius aeneipennis</emphasis>
(Fairmaire and Germain, 1862)
</taxonomicName>
<emphasis id="7CD0C3318619D80EDB1DFC29BCCFFC49" bold="true" box="[894,1012,923,947]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="67E37E4A8619D80EDB1DFC29BCCFFC49" box="[894,1012,923,947]" pageId="36" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="4E1B1F238619D80EDB5AFC66BC8BFBD1" blockId="36.[825,1474,980,1067]" pageId="36" pageNumber="37">
<emphasis id="7CD0C3318619D80EDB5AFC66BCDFFC16" box="[825,996,980,1004]" italics="true" pageId="36" pageNumber="37">Included species:</emphasis>
<taxonomicName id="89A464A08619D80EDC65FC66BCDAFBF1" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeneipennis" status="comb. nov.">
<emphasis id="7CD0C3318619D80EDC65FC66BA35FC16" bold="true" box="[1030,1294,980,1004]" italics="true" pageId="36" pageNumber="37">Apoquedius aeneipennis</emphasis>
(Fairmaire and Germain, 1862)
</taxonomicName>
<emphasis id="7CD0C3318619D80EDB8AFC41BB59FBF1" bold="true" box="[1001,1122,1011,1035]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="67E37E4A8619D80EDB8AFC41BB59FBF1" box="[1001,1122,1011,1035]" pageId="36" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="89A464A08619D80EDC0CFC41BAF9FBF1" authority="(Bernhauer, 1912)" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[1135,1474,1011,1035]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="piciformis" status="comb. nov.">
<emphasis id="7CD0C3318619D80EDC0CFC41BBC3FBF1" bold="true" box="[1135,1272,1011,1035]" italics="true" pageId="36" pageNumber="37">A. piciformis</emphasis>
(Bernhauer, 1912)
</taxonomicName>
<emphasis id="7CD0C3318619D80EDB5AFBA1BC8BFBD1" bold="true" box="[825,944,1043,1067]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="67E37E4A8619D80EDB5AFBA1BC8BFBD1" box="[825,944,1043,1067]" pageId="36" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="4E1B1F238619D80EDB5AFBFFBB4CF982" blockId="36.[825,1476,1100,1657]" pageId="36" pageNumber="37">
<emphasis id="7CD0C3318619D80EDB5AFBFFBB13FB9E" box="[825,1064,1100,1124]" italics="true" pageId="36" pageNumber="37">
Diagnosis:
<taxonomicName id="89A464A08619D80EDBD5FBFEBB13FB9E" authorityName="Scheerpeltz" authorityYear="1972" box="[950,1064,1100,1124]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Apoquedius</taxonomicName>
</emphasis>
can be diagnosed based on the head with frontoclypeal suture distinct only laterally; wide neck; paraocular punctures present; apical segment of maxillary palpi fusiform; apical segment of labial palpi fusiform, widest at the middle portion; apical labial palpomere distinctly longer than penultimate labial palpomere; flexible postcoxal hypomeral extension or process elongated and thin; two short empodial setae on each tarsal 5 segment; metacoxae with more than four spines on the posterior surface; abdomen with apical row of setae at the anterior margin of tergites and sternites III–VI; anterior margin of tergites III–VI distinctly crenulate.
<taxonomicName id="89A464A08619D80EDCA4FA37BA02FA67" authorityName="Scheerpeltz" authorityYear="1972" box="[1223,1337,1413,1437]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDCA4FA37BA02FA67" box="[1223,1337,1413,1437]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="89A464A08619D80EDB86FA17BB68FA47" box="[997,1107,1445,1469]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDB86FA17BB68FA47" box="[997,1107,1445,1469]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
</taxonomicName>
by the shape of the maxillary palps, which are fusiform; the shape of the postcoxal process which is thin and elongated; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of PTBC in tergites III to V; the most apical row of setae at the apical margin of tergites and sternites III to VI giving a crenulated appearance.
</paragraph>
<paragraph id="4E1B1F238619D80EDB5AF928BC9EF92B" blockId="36.[825,1474,1690,1745]" pageId="36" pageNumber="37">
<emphasis id="7CD0C3318619D80EDB5AF928BC8CF948" box="[825,951,1690,1714]" italics="true" pageId="36" pageNumber="37">Distribution:</emphasis>
As of now,
<taxonomicName id="89A464A08619D80EDC5EF928BB94F948" authorityName="Scheerpeltz" authorityYear="1972" box="[1085,1199,1690,1714]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDC5EF928BB94F948" box="[1085,1199,1690,1714]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
</taxonomicName>
is restricted to
<collectingCountry id="36B35FB38619D80EDD39F928BAA9F948" box="[1370,1426,1690,1714]" name="Chile" pageId="36" pageNumber="37">Chile</collectingCountry>
and
<collectingCountry id="36B35FB38619D80EDB5AF90BBC9AF92B" box="[825,929,1721,1745]" name="Argentina" pageId="36" pageNumber="37">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="4E1B1F238619D80FDB5AF941BEA6FD47" blockId="36.[825,1474,1779,1959]" lastBlockId="37.[112,764,144,701]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="7CD0C3318619D80EDB5AF941BCACF8F1" box="[825,919,1779,1803]" italics="true" pageId="36" pageNumber="37">Remarks:</emphasis>
Ŋe monophyly of
<taxonomicName id="89A464A08619D80EDCE7F941BBCDF8F1" authorityName="Scheerpeltz" authorityYear="1972" box="[1156,1270,1779,1803]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318619D80EDCE7F941BBCDF8F1" box="[1156,1270,1779,1803]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
</taxonomicName>
is supported by a unique synapomorphy: flexible postcoxal hypomeral extension or process elongated and thin (character 58, state 1); and 11 homoplastic synapomorphies: head with frontoclypeal suture distinct only laterally (character 1, state 1); infraorbital ridge (IOR) not merged with postgenal ridge (PGR) (character 17, state 1); apical segment of maxillary palpi more or less fusiform, widest at its middle portion (character 29, state 0); apical segment of labial palpi more or less fusiform, widest at the middle portion (character 32, state 0); apical labial palpomere distinctly longer than penultimate labial palpomere (character 33, state 1); two parocular punctures on each side (character 41, state 1); basisternum from about as long as to moderately longer than furcasternum (character 46, state 0); protibiae, apical tibial spur distinctly outside tibial margin (character 66, state 1); more than four spines on the posterior surface of metacoxae (character 73, state 1); mesothorax with sternopleural (anapleural) suture transverse, or nearly transverse (very slightly oblique) (character 79, state 0); abdomen with apical row of setae at the anterior margin of tergites and sternites III–VI and anterior margin of tergites III–VI distinctly crenulate (character 90, state 1). Several undescribed species from southern South America and one described species from
<collectingRegion id="8C60D1C18618D80FD90EFD37BEF4FD67" box="[365,463,645,669]" country="Australia" name="Tasmania" pageId="37" pageNumber="38">Tasmania</collectingRegion>
likely belong to
<taxonomicName id="89A464A08618D80FDAEBFD37BDC1FD67" authorityName="Scheerpeltz" authorityYear="1972" box="[648,762,645,669]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318618D80FDAEBFD37BDC1FD67" box="[648,762,645,669]" italics="true" pageId="37" pageNumber="38">Apoquedius</emphasis>
</taxonomicName>
pending further examination.
</paragraph>
</subSubSection>
</treatment>
</document>

312
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@ -0,0 +1,312 @@
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<taxonomicName id="89A464A0861DD80AD95BFC44BD3AFBEB" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[312,513,1014,1041]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="7CD0C331861DD80AD95BFC44BD3AFBEB" bold="true" box="[312,513,1014,1041]" italics="true" pageId="32" pageNumber="33">Loncovilius barclayi</emphasis>
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(
<figureCitation id="D69F03A6861DD80AD978FB87BE57FBB5" box="[283,364,1077,1103]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." figureDoi="http://doi.org/10.5281/zenodo.14443963" httpUri="https://zenodo.org/record/14443963/files/figure.png" pageId="32" pageNumber="33">Figs 6D</figureCitation>
, 11D–F, 12H, 13H, 14D)
</emphasis>
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urn:lsid:zoobank.org:act:
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.
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<emphasis id="7CD0C331861DD80AD8E2FB7DBE2AFB1D" box="[129,273,1231,1255]" italics="true" pageId="32" pageNumber="33">Type material.</emphasis>
<materialsCitation id="FECC157E861DD80AD97AFB7DBD4AFABE" collectingDate="1986-01-10" collectorName="L. E. Pena" country="Chile" location="Valdivia" municipality="Purulon" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="7CD0C331861DD80AD97AFB7DBE42FB1D" box="[281,377,1231,1255]" italics="true" pageId="32" pageNumber="33">
<typeStatus id="911FA181861DD80AD97AFB7DBE4FFB1D" box="[281,372,1231,1255]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
:
</emphasis>
male, pinned, with genitalia in a separate microvial, with labels as follows:
<collectingCountry id="36B35FB3861DD80ADA29FB5CBDA6FAFC" box="[586,669,1262,1286]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
:
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:
<collectingMunicipality id="AE7F8559861DD80AD8E2FABFBFEEFADF" box="[129,213,1293,1317]" pageId="32" pageNumber="33">Purulón</collectingMunicipality>
[-39.47, -72.69],
<date id="3A1A39E3861DD80AD9E5FABFBED3FADF" box="[390,488,1293,1317]" pageId="32" pageNumber="33" value="1986-01-10">
<collectingDate id="2A5EC00B861DD80AD9E5FABFBED3FADF" box="[390,488,1293,1317]" pageId="32" pageNumber="33" value="1986-01-10">10.I.1986</collectingDate>
</date>
<collectorName id="E3517AF5861DD80AD98FFABCBD74FADF" box="[492,591,1294,1317]" pageId="32" pageNumber="33">L.E. Peña</collectorName>
, leg. FIELD MUS. NAT. HIST./
<specimenCount id="58A2D4AA861DD80AD977FA82BE15FABE" box="[276,302,1328,1348]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
/FMNH-INS 0000 023 665
</materialsCitation>
/
<typeStatus id="911FA181861DD80ADA1AFA9EBC32FABE" box="[633,777,1324,1348]" pageId="32" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="89A464A0861DD80AD8E2FAFEBE71FA9E" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[129,330,1356,1380]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">
<emphasis id="7CD0C331861DD80AD8E2FAFEBE71FA9E" box="[129,330,1356,1380]" italics="true" pageId="32" pageNumber="33">Loncovilius barclayi</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
<materialsCitation id="FECC157E861DD80ADAC1FADEBD2CFA3B" collectingDate="1986-01-10" collectorName="Valdivia &amp; Purulon &amp; Pena" country="Chile" location="CHILE" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="7CD0C331861DD80ADAC1FADEBC32FA79" box="[674,777,1388,1411]" italics="true" pageId="32" pageNumber="33">
<typeStatus id="911FA181861DD80ADAC1FADEBC3FFA79" box="[674,772,1388,1411]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="36B35FB3861DD80AD8E2FA39BFEFFA59" box="[129,212,1419,1443]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
:
<collectorName id="E3517AF5861DD80AD880FA39BE02FA59" box="[227,313,1419,1443]" pageId="32" pageNumber="33">Valdivia</collectorName>
:
<collectorName id="E3517AF5861DD80AD92BFA39BEA7FA59" box="[328,412,1419,1443]" pageId="32" pageNumber="33">Purulón</collectorName>
[-39.47, -72.69],
<date id="3A1A39E3861DD80ADA06FA39BDFCFA59" box="[613,711,1419,1443]" pageId="32" pageNumber="33" value="1986-01-10">
<collectingDate id="2A5EC00B861DD80ADA06FA39BDFCFA59" box="[613,711,1419,1443]" pageId="32" pageNumber="33" value="1986-01-10">10.I.1986</collectingDate>
</date>
L. E.
<collectorName id="E3517AF5861DD80AD8E2FA18BF8EFA3B" box="[129,181,1450,1473]" pageId="32" pageNumber="33">Peña</collectorName>
, leg. FIELD MUS. NAT. HIST./
<specimenCount id="58A2D4AA861DD80AD99EFA1FBD2CFA3B" box="[509,535,1453,1473]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
</materialsCitation>
/FMNH-INS 0000 023 667/
<taxonomicName id="89A464A0861DD80AD8CCFA7BBD57FA1B" authority="J. L. Reyes-Hernandez" authorityName="J. L. Reyes-Hernandez" box="[175,620,1481,1505]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">Loncovilius barclayi J. L. Reyes-Hernández</taxonomicName>
[
<specimenCount id="58A2D4AA861DD80ADA1FFA7BBD98FA1A" box="[636,675,1481,1505]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
FMNH];
<materialsCitation id="FECC157E861DD80AD8E2FA5BBD2CF9E5" collectingDate="1986-01-10" collectorName="Valdivia &amp; Purulon &amp; Pena" country="Chile" location="CHILE" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="36B35FB3861DD80AD8E2FA5BBFEFF9FB" box="[129,212,1513,1537]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
:
<collectorName id="E3517AF5861DD80AD880FA5BBE02F9FB" box="[227,313,1513,1537]" pageId="32" pageNumber="33">Valdivia</collectorName>
:
<collectorName id="E3517AF5861DD80AD92AFA5BBEA6F9FB" box="[329,413,1513,1537]" pageId="32" pageNumber="33">Purulón</collectorName>
[-39.47, -72.69],
<date id="3A1A39E3861DD80ADA06FA5BBDFCF9FB" box="[613,711,1513,1537]" pageId="32" pageNumber="33" value="1986-01-10">
<collectingDate id="2A5EC00B861DD80ADA06FA5BBDFCF9FB" box="[613,711,1513,1537]" pageId="32" pageNumber="33" value="1986-01-10">10.I.1986</collectingDate>
</date>
L. E.
<collectorName id="E3517AF5861DD80AD8E2F9BABF8EF9E5" box="[129,181,1544,1567]" pageId="32" pageNumber="33">Peña</collectorName>
, leg. FIELD MUS. NAT. HIST./
<specimenCount id="58A2D4AA861DD80AD99EF9B9BD2CF9E5" box="[509,535,1547,1567]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
</materialsCitation>
/
<materialsCitation id="FECC157E861DD80ADA43F9BABD1AF967" collectingDate="1993-01-20" collectorName="Coll. T Curkovic" country="Chile" determinerName="Newton" location="Region-Chile" municipality="Nahuelbuta" pageId="32" pageNumber="33" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
FMNH-INS 0000 023 666/
<taxonomicName id="89A464A0861DD80AD8D3F995BDBAF9C5" authority="J. L. Reyes-Hernandez" authorityName="J. L. Reyes-Hernandez" box="[176,641,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">Loncovilius barclayi J. L. Reyes-Hernández</taxonomicName>
[
<specimenCount id="58A2D4AA861DD80ADAF5F995BD85F9C4" box="[662,702,1575,1599]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
NHM ex. FMNH];
<collectingMunicipality id="AE7F8559861DD80AD96DF9F5BEBDF9A5" box="[270,390,1607,1631]" pageId="32" pageNumber="33">Nahuelbuta</collectingMunicipality>
IX
<location id="4B7B49F8861DD80AD9D2F9F5BD06F9A5" LSID="urn:lsid:plazi:treatment:C60DAE35861DD80BD95BFC44BE97FCB5:4B7B49F8861DD80AD9D2F9F5BD06F9A5" box="[433,573,1607,1631]" country="Chile" municipality="Nahuelbuta" name="Region-Chile" pageId="32" pageNumber="33">Region-Chile</location>
,
<date id="3A1A39E3861DD80ADA29F9F5BD8AF9A5" box="[586,689,1607,1631]" pageId="32" pageNumber="33" value="1993-01-20">
<collectingDate id="2A5EC00B861DD80ADA29F9F5BD8AF9A5" box="[586,689,1607,1631]" pageId="32" pageNumber="33" value="1993-01-20">20-I-1993</collectingDate>
</date>
,
<collectorName id="E3517AF5861DD80ADADEF9F5BFD9F984" pageId="32" pageNumber="33">Coll. T Curkovic</collectorName>
/
<taxonomicName id="89A464A0861DD80AD889F9D4BEDEF984" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[234,485,1638,1662]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semiflavus">Loncovilius? semiflavus</taxonomicName>
det.
<determinerName id="BBC7F3E7861DD80ADA46F9D4BD42F987" box="[549,633,1638,1661]" pageId="32" pageNumber="33">Newton</determinerName>
2000 [
<specimenCount id="58A2D4AA861DD80ADAB3F9D4BDCCF987" box="[720,759,1638,1662]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
]; [
<collectingCountry id="36B35FB3861DD80AD8E4F937BFDBF967" box="[135,224,1669,1693]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
:] COLL. CERDA MNHN
</materialsCitation>
<materialsCitation id="FECC157E861DD80ADA54F937BE21F921" collectingDate="1981-01-22" collectorName="Nahuelbuta Parque &amp; Mus" country="Chile" location="Santiago" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Region Metropolitana" typeStatus="paratype">
<collectingCountry id="36B35FB3861DD80ADA54F937BDBDF967" box="[567,646,1669,1693]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
/
<collectorName id="E3517AF5861DD80ADAEEF937BFF2F946" pageId="32" pageNumber="33">Nahuelbuta Parque</collectorName>
[-37.80, -73.01]
<date id="3A1A39E3861DD80AD910F917BEE7F947" box="[371,476,1701,1725]" pageId="32" pageNumber="33" value="1981-01-22">
<collectingDate id="2A5EC00B861DD80AD910F917BEE7F947" box="[371,476,1701,1725]" pageId="32" pageNumber="33" value="1981-01-22">22.1.1981</collectingDate>
</date>
. P. [illegible] B/
<collectingRegion id="8C60D1C1861DD80ADA18F917BDECF947" box="[635,727,1701,1725]" country="Chile" name="Region Metropolitana" pageId="32" pageNumber="33">Santiago</collectingRegion>
<collectorName id="E3517AF5861DD80ADABFF917BC32F946" box="[732,777,1701,1724]" pageId="32" pageNumber="33">Mus</collectorName>
[MNNC
<specimenCount id="58A2D4AA861DD80AD884F976BE35F921" box="[231,270,1732,1756]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
]
</materialsCitation>
;
<materialsCitation id="FECC157E861DD80AD948F976BEA4F84D" collectingDate="2005-01-25" collectingDateMax="2005-01-31" collectingDateMin="2005-01-25" collectorName="P. N. Nahuelbula &amp; Fairmaire" country="Chile" elevation="1200" location="Sv Bily" pageId="32" pageNumber="33" specimenCount="2" specimenCount-male="2" stateProvince="Araucania" typeStatus="paratype">
<collectingCountry id="36B35FB3861DD80AD948F976BE40F926" box="[299,379,1732,1756]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
, IX r.,
<date id="3A1A39E3861DD80AD9A8F976BD75F926" box="[459,590,1732,1756]" pageId="32" pageNumber="33" value="2005-01-25" valueMax="2005-01-31" valueMin="2005-01-25">
<collectingDate id="2A5EC00B861DD80AD9A8F976BD75F926" box="[459,590,1732,1756]" pageId="32" pageNumber="33" value="2005-01-25" valueMax="2005-01-31" valueMin="2005-01-25">25-31.i.2005</collectingDate>
</date>
,
<collectorName id="E3517AF5861DD80ADA3EF976BC32F926" box="[605,777,1732,1756]" pageId="32" pageNumber="33">P.N. Nahuelbula</collectorName>
[
<collectingRegion id="8C60D1C1861DD80AD8E5F951BFC8F901" box="[134,243,1763,1787]" country="Chile" name="Araucania" pageId="32" pageNumber="33">Araucanía</collectingRegion>
, Nahuelbuta National Park] 37°48
<emphasis id="7CD0C331861DD80ADA0AF950BD54F901" box="[617,623,1762,1787]" italics="true" pageId="32" pageNumber="33">ʹ</emphasis>
530 73°00
<emphasis id="7CD0C331861DD80ADABEF950BDD8F901" box="[733,739,1762,1787]" italics="true" pageId="32" pageNumber="33">ʹ</emphasis>
954 [-37.80, -73.01],
<quantity id="895CB2C6861DD80AD95AF8B1BEB5F8E0" box="[313,398,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="33" unit="m" value="1200.0">
<elevation id="C589F810861DD80AD95AF8B1BEB5F8E0" box="[313,398,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="33" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<location id="4B7B49F8861DD80AD9FEF8B1BED3F8E1" LSID="urn:lsid:plazi:treatment:C60DAE35861DD80BD95BFC44BE97FCB5:4B7B49F8861DD80AD9FEF8B1BED3F8E1" box="[413,488,1795,1819]" country="Chile" name="Sv Bily" pageId="32" pageNumber="33" stateProvince="Araucania">Sv Bíly</location>
lgt./NHMD 916914 [
<specimenCount id="58A2D4AA861DD80ADA82F8B1BC32F8E0" box="[737,777,1795,1819]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
NMPC]; [
<collectingCountry id="36B35FB3861DD80AD892F890BE7CF8C0" box="[241,327,1826,1850]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
: no exact location] col.
<collectorName id="E3517AF5861DD80ADA39F890BDFDF8C0" box="[602,710,1826,1850]" pageId="32" pageNumber="33">Fairmaire</collectorName>
/1251/
<taxonomicName id="89A464A0861DD80AD8E2F8F3BE6AF8A3" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[129,337,1857,1881]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semiflavus" status="sp. nov.">Quedius semiflavus</taxonomicName>
<taxonomicNameLabel id="67E37E4A861DD80AD903F8F3BEB7F8A3" box="[352,396,1857,1881]" pageId="32" pageNumber="33" rank="species">n.sp.</taxonomicNameLabel>
/FMNH-INS 0000 024 190 [
<specimenCount id="58A2D4AA861DD80ADA82F8F3BC32F8A2" box="[737,777,1857,1881]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
MNNC ex. FMNH]. All
<typeStatus id="911FA181861DD80AD9FEF8D3BD3BF883" box="[413,512,1889,1913]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
with label: PAÞTYPE
<taxonomicName id="89A464A0861DD80AD8E2F832BE71F862" box="[129,330,1920,1944]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">
<emphasis id="7CD0C331861DD80AD8E2F832BE71F862" box="[129,330,1920,1944]" italics="true" pageId="32" pageNumber="33">Loncovilius barclayi</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
</materialsCitation>
.
</paragraph>
<paragraph id="4E1B1F23861DD80ADB5AFF23BBD9FEFF" blockId="32.[825,1474,144,262]" pageId="32" pageNumber="33">
<emphasis id="7CD0C331861DD80ADB5AFF23BCA4FF52" box="[825,927,145,168]" italics="true" pageId="32" pageNumber="33">Diagnosis:</emphasis>
Elytra yellow or yellowish-brown with black marks around mesoscutellum and on suture. Male tergite X trilobed (
<figureCitation id="D69F03A6861DD80ADB27FF7DBCA4FF1D" box="[836,927,207,231]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="32" pageNumber="33">Fig. 12H</figureCitation>
); female tergite X deeply emarginate medio-apically (
<figureCitation id="D69F03A6861DD80ADB27FF5CBCA6FEFC" box="[836,925,238,262]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="32" pageNumber="33">Fig. 13H</figureCitation>
). Aedeagus as in Figure 11D–F.
</paragraph>
<paragraph id="4E1B1F23861DD80ADB5AFE89BBD7FE15" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">
<emphasis id="7CD0C331861DD80ADB5AFE89BC8BFEA8" box="[825,944,315,338]" italics="true" pageId="32" pageNumber="33">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA861DD80ADC02FE8CBB40FEA8" box="[1121,1147,318,338]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C331861DD80ADD10FE89BABDFEA8" box="[1395,1414,315,338]" italics="true" pageId="32" pageNumber="33">N</emphasis>
= 6]: FBL = 2.42–3.86 (3.56); TL = 7.13–7.5 (7.32); HW = 0.9– 0.98 (0.95); HL = 0.9–0.98 (0.95); HW/HL = 0.99–1.03 (1); PW = 1.19–1.31 (1.25); PL = 1.25–1.3 (1.28); PW/ PL = 0.93–1.02 (0.98); EW = 1.54–1.75 (1.64); EL = 1.45– 1.65 (1.56); PW/HW = 1.28–1.34 (1.31).
</paragraph>
<paragraph id="4E1B1F23861DD80ADB36FE45BA6CFDB7" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">
Measurements
<specimenCount id="58A2D4AA861DD80ADC68FE48BB1EFDF4" box="[1035,1061,506,526]" count="1" pageId="32" pageNumber="33" type="female">♀</specimenCount>
(
<emphasis id="7CD0C331861DD80ADC32FE45BB5FFDF4" box="[1105,1124,503,526]" italics="true" pageId="32" pageNumber="33">N</emphasis>
= 1): FBL = 4.03; TL = 7.5; HW = 1.05; HL = 1; HW/HL = 1.05; PW = 1.38; PL = 1.38; PW/PL = 1; EW = 1.7; EL = 1.65; PW/HW = 1.31.
</paragraph>
<paragraph id="4E1B1F23861DD80ADB36FDE7BCFDFCF3" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">Head and pronotum yellow-orange; antennae, mouthparts, and legs yellow or yellowish-brown; elytra yellow or yellowish-brown with dark mark around mesoscutellum and on suture; meso- and metasternum, all coxae, femora and abdomen dark reddish-brown; abdominal tergites and sternites with pale posterior margin.</paragraph>
<paragraph id="4E1B1F23861DD80ADB36FCA3BCD6FA41" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">
Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small-size. Eyes medium size (EYL/HL = x = 0.52), from 1.4 to 1.55 times longer than length of temples (in lateral view); distance between eyes about equal to length of eye. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 6 subequal in length; antennomeres 6 to 10 gradually reduced in length; a11 1.75 to 1.88 times as long as a
<quantity id="895CB2C6861DD80ADBFBFBF8BCF5FB98" box="[920,974,1098,1122]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="32" pageNumber="33" unit="in" value="10.0">10 in</quantity>
males and
<quantity id="895CB2C6861DD80ADC2BFBF8BBAAFB98" box="[1096,1169,1098,1122]" metricMagnitude="-2" metricUnit="m" metricValue="4.2926" pageId="32" pageNumber="33" unit="in" value="1.69">1.69 in</quantity>
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, fused with infraorbital ridge; gular sutures moderately separated. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
</paragraph>
<paragraph id="4E1B1F23861DD80ADB36FA70BA80F98D" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">Pronotum about as wide as long, strongly convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (or process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or four times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F23861DD80ADB36F9CCBA7EF8A8" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate.</paragraph>
<paragraph id="4E1B1F23861DD80BDB36F8EBBEC0FE59" blockId="32.[823,1476,315,1968]" lastBlockId="33.[113,763,144,607]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with strong, V-shaped emargination medially (
<figureCitation id="D69F03A6861CD80BD8CAFF5CBE3DFEFC" box="[169,262,238,262]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="33" pageNumber="34">Fig. 14H</figureCitation>
); female sternite VIII with weak apical medial emargination; male sternite IX not emarginate medio-apically, its basal portion distinctly longer than distal portion; male tergite X trilobed (
<figureCitation id="D69F03A6861CD80BD978FEFEBE4DFE9E" box="[283,374,332,356]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="33" pageNumber="34">Fig. 12H</figureCitation>
); female tergite X deeply emarginate medio-apically (
<figureCitation id="D69F03A6861CD80BD97DFED9BE46FE79" box="[286,381,363,387]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="33" pageNumber="34">Fig. 13H</figureCitation>
); ovipositor, its second gonocoxite with one macroseta medially (
<figureCitation id="D69F03A6861CD80BD9C6FE39BED1FE59" box="[421,490,395,419]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="33" pageNumber="34">Fig 9G</figureCitation>
).
</paragraph>
<paragraph id="4E1B1F23861CD80BD8EEFE18BDC0FDA5" blockId="33.[113,763,144,607]" pageId="33" pageNumber="34">Aedeagus as in Figure 11D–F, its length ~ 1.46; median lobe rod-like, in parameral view apex deeply emarginate; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into a deeply emarginate apex, with dark peg setae, in lateral view almost straight and slightly concave apically.</paragraph>
<paragraph id="4E1B1F23861CD80BD812FD32BDC1FD4D" blockId="33.[113,763,640,695]" pageId="33" pageNumber="34">
<emphasis id="7CD0C331861CD80BD812FD32BFE5FD62" box="[113,222,640,664]" italics="true" pageId="33" pageNumber="34">Etymology:</emphasis>
Ŋe species epithet is a patronym in recognition of Maxwell V.L.Barclay, in particular for his earlier mentoring of J.J.S..
</paragraph>
<paragraph id="4E1B1F23861CD80BD812FD6BBE97FCB5" blockId="33.[113,763,729,847]" pageId="33" pageNumber="34">
<emphasis id="7CD0C331861CD80BD812FD6BBD15FD0B" box="[113,558,729,753]" italics="true" pageId="33" pageNumber="34">
Distribution and habitat:
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</emphasis>
is known only from
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, from the
<collectingRegion id="8C60D1C1861CD80BD979FD4ABEBAFCEA" box="[282,385,760,784]" country="Chile" name="Araucania" pageId="33" pageNumber="34">Araucanía</collectingRegion>
and
<collectingRegion id="8C60D1C1861CD80BD9D4FD4ABD2FFCEA" box="[439,532,760,784]" country="Chile" name="Los Rios" pageId="33" pageNumber="34">Los Ríos</collectingRegion>
regions, in the north-central portion of the Valdivian temperate forest ecoregion. It is unknown how it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>

319
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<emphasis id="7CD0C331861FD808DB8AFE10BBFAFE47" bold="true" box="[1001,1217,418,445]" italics="true" pageId="34" pageNumber="35">Loncovilius carlsbergi</emphasis>
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<emphasis id="7CD0C331861FD808DBA5FE60BA0DFE16" bold="true" box="[966,1334,466,492]" pageId="34" pageNumber="35">
(
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,
<figureCitation id="D69F03A6861FD808DC22FE60BB5AFE16" box="[1089,1121,466,492]" captionStart="Figure 8" captionStartId="16.[129,194,1829,1853]" captionTargetBox="[183,1419,150,1799]" captionTargetId="figure-8@16.[183,1419,144,1799]" captionTargetPageId="16" captionText="Figure 8. A, pronotum of L. carlsbergi. AMRSP = anterior marginal row of setiferous punctures, APP = additional paired punctures adjacent to AMRSP, LLSP = large lateral setiferous puncture, LPSP = large posterior setiferous puncture, PPDS = paired punctures on dorsal serial, SLSP = sublateral setiferous puncture.B, L. cantharoides ventral view of mesotarsomere 3 with pale adhesive setae with terminal plate.C, L. cantharoides ventral view of metatarsomere 4, apical margin emarginated (bilobed), with pale adhesive setae without terminal plate. D, L. cantharoides tergites VI and VII.Black arrow up = anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, white arrow down = ATBC descending laterally without reaching tergite margin.Head in ventral view of Loncovilius species. E, L. hammondi. F, L. lividipennis. PMP = postmandibular puncture, PMR = postmandibular ridge, EM = eye margin." figureDoi="http://doi.org/10.5281/zenodo.14443971" httpUri="https://zenodo.org/record/14443971/files/figure.png" pageId="34" pageNumber="35">8A</figureCitation>
, 11I, J, 12J, 13J, 14J)
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urn:lsid:zoobank.org:act:
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.
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<emphasis id="7CD0C331861FD808DB5AFDEFBCF0FD8E" box="[825,971,604,628]" italics="true" pageId="34" pageNumber="35">Type material.</emphasis>
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male, pinned, with labels as follows: [
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:
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and Chilean
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[-49.11, -74.40]
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/
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/NHM [blue label]/NHMD 916967
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/
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<emphasis id="7CD0C331861FD808DB58FD6BBB2AFD0B" box="[827,1041,729,753]" italics="true" pageId="34" pageNumber="35">Loncovilius carlsbergi</emphasis>
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des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM.
<materialsCitation id="FECC157E861FD808DD38FD4BBB3FF9FB" collectingDate="1958-11-20" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectorName="Wellington Is. Pto. Eden &amp; I. Puerto Eden &amp; Pto. Eden Wellington Is. &amp; Pena. Munoz, HM &amp; Fr. Kuschel &amp; P. Voillians" country="Chile" determinerName="A. Newton" elevation="12" location="Isla Wellington" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="39" specimenCount-female="18" specimenCount-male="21" stateProvince="Magallanes y de la Antartica Chilena" typeStatus="paratype">
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<typeStatus id="911FA181861FD808DD38FD4BBA86FCEA" box="[1371,1469,761,784]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
:
</emphasis>
[
<collectingCountry id="36B35FB3861FD808DB23FCAABCADFCCA" box="[832,918,792,816]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C1861FD808DBC6FCAABB23FCCA" box="[933,1048,792,816]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
and Chilean
<taxonomicName id="89A464A0861FD808DCD6FCAABA86FCCA" authority=", Puerto Eden" authorityName="Puerto Eden" box="[1205,1469,792,816]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">Antarctica, Puerto Edén</taxonomicName>
]
<collectorName id="E3517AF5861FD808DB5AFC85BB09FCB5" box="[825,1074,823,847]" pageId="34" pageNumber="35">Wellington Is. Pto. Eden</collectorName>
<date id="3A1A39E3861FD808DC54FC85BB9DFCB5" box="[1079,1190,823,847]" pageId="34" pageNumber="35" value="1958-12-04">
<collectingDate id="2A5EC00B861FD808DC54FC85BB9DFCB5" box="[1079,1190,823,847]" pageId="34" pageNumber="35" value="1958-12-04">4.XII.1958</collectingDate>
</date>
/
<collectingCountry id="36B35FB3861FD808DCCDFC85BA3AFCB5" box="[1198,1281,823,847]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
<collectingMunicipality id="AE7F8559861FD808DD64FC8ABA40FCB5" box="[1287,1403,823,847]" pageId="34" pageNumber="35">Fr Kuschel</collectingMunicipality>
/NHM [blue label] [
<specimenCount id="58A2D4AA861FD808DBD8FCE5BCDFFC94" box="[955,996,855,879]" count="2" pageId="34" pageNumber="35" type="male">2♂</specimenCount>
<specimenCount id="58A2D4AA861FD808DB8BFCE5BB34FC94" box="[1000,1039,855,879]" count="6" pageId="34" pageNumber="35" type="female">6♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA861FD808DC3CFCE5BBBDFC94" box="[1119,1158,855,879]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
NHMD ex. NHM;
<specimenCount id="58A2D4AA861FD808DD2DFCE5BA4EFC94" box="[1358,1397,855,879]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
CZUG ex. NHM;
<specimenCount id="58A2D4AA861FD808DBC5FCC4BCF4FC77" box="[934,975,886,910]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
FMNH ex. NHM]; [same data as above except:]
<date id="3A1A39E3861FD808DB5AFC27BC96FC57" box="[825,941,917,941]" pageId="34" pageNumber="35" value="1958-11-20">
<collectingDate id="2A5EC00B861FD808DB5AFC27BC96FC57" box="[825,941,917,941]" pageId="34" pageNumber="35" value="1958-11-20">20.XI.1958</collectingDate>
</date>
[
<specimenCount id="58A2D4AA861FD808DBD8FC27BCDEFC56" box="[955,997,917,941]" count="5" pageId="34" pageNumber="35" type="male">5♂</specimenCount>
<specimenCount id="58A2D4AA861FD808DB88FC27BB29FC56" box="[1003,1042,917,941]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA861FD808DC05FC27BBB6FC56" box="[1126,1165,917,941]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
FMNH ex. NHM]; [
<collectingCountry id="36B35FB3861FD808DD0AFC27BA84FC57" box="[1385,1471,917,941]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C1861FD808DB59FC07BC96FC37" box="[826,941,949,973]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
and Chilean Antarctica]
<taxonomicName id="89A464A0861FD808DCAEFC07BA80FC37" box="[1229,1467,949,973]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">Nothofagus antarctica</taxonomicName>
/
<collectingCountry id="36B35FB3861FD808DB5AFC66BCBCFC16" box="[825,903,980,1004]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
HE
<collectorName id="E3517AF5861FD808DBA3FC66BB6EFC16" box="[960,1109,980,1004]" pageId="34" pageNumber="35">I.Puerto Eden</collectorName>
,
<location id="4B7B49F8861FD808DC07FC66BA38FC16" LSID="urn:lsid:plazi:treatment:C60DAE35861FD809DB8AFE10BBDFFE67:4B7B49F8861FD808DC07FC66BA38FC16" box="[1124,1283,980,1004]" country="Chile" municipality="Fr Kuschel" name="Isla Wellington" pageId="34" pageNumber="35" stateProvince="Magallanes y de la Antartica Chilena">Isla Wellington</location>
[-49.126, -74.40] 490S 40 ss.
<date id="3A1A39E3861FD808DBCFFC41BB22FBF1" box="[940,1049,1011,1035]" pageId="34" pageNumber="35" value="1958-11-29">
<collectingDate id="2A5EC00B861FD808DBCFFC41BB22FBF1" box="[940,1049,1011,1035]" pageId="34" pageNumber="35" value="1958-11-29">29.xi.1958</collectingDate>
</date>
/NHM [
<specimenCount id="58A2D4AA861FD808DC17FC41BBA0FBF0" box="[1140,1179,1011,1035]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
NHM]; [same data as above except:]
<collectingCountry id="36B35FB3861FD808DBF6FBA1BCD8FBD1" box="[917,995,1043,1067]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
HE6. [
<specimenCount id="58A2D4AA861FD808DC50FBA1BB61FBD0" box="[1075,1114,1043,1067]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB3861FD808DCA2FBA1BA2CFBD1" box="[1217,1303,1043,1067]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C1861FD808DD41FBA1BAAEFBD1" box="[1314,1429,1043,1067]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
and Chilean Antarctica] S.
<collectingCountry id="36B35FB3861FD808DC78FB80BB52FBB0" box="[1051,1129,1074,1098]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
<collectorName id="E3517AF5861FD808DC0EFB80BA5BFBB3" box="[1133,1376,1074,1098]" pageId="34" pageNumber="35">Pto. Edén Wellington Is.</collectorName>
[-49.126, -74.40] Dec.7-15,’62 PJDarlinton/FIELD MUSEUM ex MCZ (retained duplic.)/
<taxonomicName id="89A464A0861FD808DC64FBC3BBCCFB73" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[1031,1271,1137,1161]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lividipennis">
<emphasis id="7CD0C331861FD808DC64FBC3BBCCFB73" box="[1031,1271,1137,1161]" italics="true" pageId="34" pageNumber="35">Loncovilius lividipennis</emphasis>
</taxonomicName>
(F + G) det.
<determinerName id="BBC7F3E7861FD808DDC8FBC3BCB6FB5D" pageId="34" pageNumber="35">A. Newton</determinerName>
1993 [
<specimenCount id="58A2D4AA861FD808DBB5FB22BCC6FB5D" box="[982,1021,1168,1192]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB3861FD808DC0CFB22BBFEFB52" box="[1135,1221,1168,1192]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C1861FD808DCAEFB22BA7BFB52" box="[1229,1344,1168,1192]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
and Chilean Antarctica]
<taxonomicName id="89A464A0861FD808DBA9FB1DBBF3FB3D" box="[970,1224,1199,1223]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">Nothofagus antarctica</taxonomicName>
/
<collectingCountry id="36B35FB3861FD808DCB3FB1DBA25FB3D" box="[1232,1310,1199,1223]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
<collectorName id="E3517AF5861FD808DD5FFB02BC5EFB1D" pageId="34" pageNumber="35">Pena.Munoz, HM</collectorName>
3.Gamero. 20–40 ss. [-52.33, -73.53,
<quantity id="895CB2C6861FD808DC9DFB7DBA14FB1D" box="[1278,1327,1231,1255]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="m" value="12.0">
<elevation id="C589F810861FD808DC9DFB7DBA14FB1D" box="[1278,1327,1231,1255]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="m" value="12.0">12m</elevation>
</quantity>
]
<date id="3A1A39E3861FD808DD20FB7CBA81FB1C" box="[1347,1466,1230,1254]" pageId="34" pageNumber="35" value="1958-12-27">
<collectingDate id="2A5EC00B861FD808DD20FB7CBA81FB1C" box="[1347,1466,1230,1254]" pageId="34" pageNumber="35" value="1958-12-27">27.xii.1958</collectingDate>
</date>
/ NHM [
<specimenCount id="58A2D4AA861FD808DBEBFB5CBC8AFAFF" box="[904,945,1262,1286]" count="3" pageId="34" pageNumber="35" type="male">3♂</specimenCount>
<specimenCount id="58A2D4AA861FD808DBDBFB5CBCE4FAFF" box="[952,991,1262,1286]" count="2" pageId="34" pageNumber="35" type="female">2♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA861FD808DC57FB5CBB60FAFF" box="[1076,1115,1262,1286]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA861FD808DC02FB5CBBB2FAFF" box="[1121,1161,1262,1286]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
MNNC ex. NHM]; [
<collectingCountry id="36B35FB3861FD808DD0AFB5CBA84FAFC" box="[1385,1471,1262,1286]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C1861FD808DB58FABFBC95FADF" box="[827,942,1293,1317]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
and Chilean Antarctica] Munoz Gamero [-52.33, -73.53]
<date id="3A1A39E3861FD808DBF1FA9EBB35FABE" box="[914,1038,1324,1348]" pageId="34" pageNumber="35" value="1958-12-27">
<collectingDate id="2A5EC00B861FD808DBF1FA9EBB35FABE" box="[914,1038,1324,1348]" pageId="34" pageNumber="35" value="1958-12-27">27.XII.1958</collectingDate>
</date>
/
<collectingCountry id="36B35FB3861FD808DC75FA9EBB52FABE" box="[1046,1129,1324,1348]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
<collectorName id="E3517AF5861FD808DC16FA9FBBCEFABF" box="[1141,1269,1325,1349]" pageId="34" pageNumber="35">Fr. Kuschel</collectorName>
/Shore/NHM [
<specimenCount id="58A2D4AA861FD808DDFBFA9FBAF9FABE" box="[1432,1474,1325,1349]" count="6" pageId="34" pageNumber="35" type="male">6♂</specimenCount>
<specimenCount id="58A2D4AA861FD808DB5AFAFEBC5BFA99" box="[825,864,1356,1380]" count="3" pageId="34" pageNumber="35" type="female">3♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA861FD808DBA0FAFEBCD1FA99" box="[963,1002,1356,1380]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
NHMD ex. NHM]; [
<collectingCountry id="36B35FB3861FD808DC8AFAFEBA04FA9E" box="[1257,1343,1356,1380]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C1861FD808DD2CFAFEBAF9FA9E" box="[1359,1474,1356,1380]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
and Chilean Antarctica]
<collectorName id="E3517AF5861FD808DC20FAD9BB90FA79" box="[1091,1195,1387,1411]" pageId="34" pageNumber="35">P.Voillians</collectorName>
[Puerto Williams, -54.93, -67.619] 10.1-1959/
<collectingCountry id="36B35FB3861FD808DC69FA38BB66FA58" box="[1034,1117,1418,1442]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
Fr Kuschel/4/NHM [
<specimenCount id="58A2D4AA861FD808DD29FA39BA49FA58" box="[1354,1394,1419,1443]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
CZUG ex. NHM]. All
<typeStatus id="911FA181861FD808DB86FA18BB73FA38" box="[997,1096,1450,1474]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
with label: PAÞTYPE
<taxonomicName id="89A464A0861FD808DD37FA18BCAEFA1B" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carlsbergi">
<emphasis id="7CD0C331861FD808DD37FA18BCAEFA1B" italics="true" pageId="34" pageNumber="35">Loncovilius carlsbergi</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
</materialsCitation>
.
</paragraph>
<paragraph id="4E1B1F23861FD808DB5AF991BBACF982" blockId="34.[824,1474,1570,1657]" pageId="34" pageNumber="35">
<emphasis id="7CD0C331861FD808DB5AF991BCA4F9C0" box="[825,927,1571,1594]" italics="true" pageId="34" pageNumber="35">Diagnosis:</emphasis>
Pronotum with APP. Elytra yellowish-brown. Sternite III with basal transverse carina evenly converging at an obtuse angle. Aedeagus as in Figure 11I, J.
</paragraph>
<paragraph id="4E1B1F23861FD808DB5AF929BB81F8B5" blockId="34.[825,1475,1690,1996]" pageId="34" pageNumber="35">
<emphasis id="7CD0C331861FD808DB5AF929BC8BF948" box="[825,944,1691,1714]" italics="true" pageId="34" pageNumber="35">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA861FD808DC02F92FBB40F94B" box="[1121,1147,1693,1713]" count="1" pageId="34" pageNumber="35" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C331861FD808DD10F929BABDF948" box="[1395,1414,1691,1714]" italics="true" pageId="34" pageNumber="35">N</emphasis>
= 6]: FBL = 3.5–3.95 (3.69); TL = 6.47–7.17 (6.71); HW = 0.87– 0.95 (0.9); HL = 0.89–0.95 (0.91); HW/HL = 0.95–1.03 (0.99); PW = 1.19–1.3 (1.25); PL = 1.11–1.2 (1.16); PW/ PL = 1.03–1.13 (1.08); EW = 1.75–1.86 (1.81); EL = 1.46–1.8 (1.61); PW/HW = 1.36–1.44 (1.38).
</paragraph>
<paragraph id="4E1B1F23861FD809DB36F8E4BE54FF3D" blockId="34.[825,1475,1690,1996]" lastBlockId="35.[112,764,144,1985]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Measurements
<specimenCount id="58A2D4AA861FD808DB93F8EBBB31F897" box="[1008,1034,1881,1901]" count="1" pageId="34" pageNumber="35" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C331861FD808DC89F8E5BBC6F894" box="[1258,1277,1879,1902]" italics="true" pageId="34" pageNumber="35">N</emphasis>
= 6]: FBL = 3.73– 3.94 (3.85); TL = 6.82–8.54 (7.92); HW = 0.91–0.96 (0.94); HL = 0.93–0.97 (0.95); HW/HL = 0.98–1 (0.99); PW = 1.2– 1.36 (1.28); PL = 1.2–1.26 (1.23); PW/PL = 1–1.09 (1.04); EW = 1.71–1.96 (1.84); EL = 1.59–1.74 (1.67); PW/ HW = 1.32–1.42 (1.36).
</paragraph>
<paragraph id="4E1B1F23861ED809D8EEFF7DBE69FEBF" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">Head, mandibles, legs, last antennomeres, and last palpomeres dark-brown; first palpomeres and antennomeres, and elytra yellow; pronotum yellowish-brown; tergites and sternites with pale posterior margin.</paragraph>
<paragraph id="4E1B1F23861ED809D8EEFEFEBEAAFB89" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">
Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small-size. Eyes small size (EYL/HL = x = 0.45), from 1.27 to 1.35 times as long as temples (in lateral view) in males and from 1 to 1.16 times in females; distance between eyes about 1.24 times as long as length of eye in males and 1.41 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 7 subequal in length; antennomeres 8 antennomeres 10 subequal in length; a11 from 1.62 to 2 times as long as a
<quantity id="895CB2C6861ED809D9F7FD17BEF2FD47" box="[404,457,677,701]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C6861ED809DA1FFD16BFBEFD26" metricMagnitude="-2" metricUnit="m" metricValue="4.1275" metricValueMax="4.2926" metricValueMin="3.9624" pageId="35" pageNumber="36" unit="in" value="1.625" valueMax="1.69" valueMin="1.56">1.56 to 1.69 in</quantity>
females. Basal and parocular punctures usually single; posterior frontal puncture located posterior to temporal puncture; without small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight, ends distinctly far from gular sutures; postgenal ridge present; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated in males and widely in females. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical.
</paragraph>
<paragraph id="4E1B1F23861ED809D8EEFBC9BD45FAD5" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">Pronotum slightly transverse, convex, evenly curved, with two PPDS, with an APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three or five times as long as, diameter of punctures. Mesosternum with five macrosetae arranged in two rows medially.</paragraph>
<paragraph id="4E1B1F23861ED809D8EEFA85BFFEF9D0" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
<paragraph id="4E1B1F23861ED809D8EEF983BDF5F858" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">
Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with rectangular V-shaped emargination medially (
<figureCitation id="D69F03A6861ED809D81FF95FBFF1F8FF" box="[124,202,1773,1797]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="35" pageNumber="36">Fig. 14J</figureCitation>
); female sternite VIII with arcuate emargination medially; male sternite IX with obtuse emargination medially, its basal portion moderately longer than distal portion; male tergite X with U-shaped emargination medially (
<figureCitation id="D69F03A6861ED809DA2DF8F9BDA6F899" box="[590,669,1867,1891]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="35" pageNumber="36">Fig. 12J</figureCitation>
); female tergite X with U-shaped emargination medially (
<figureCitation id="D69F03A6861ED809DA09F8D9BD81F879" box="[618,698,1899,1923]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="35" pageNumber="36">Fig. 13J</figureCitation>
); ovipositor, its second gonocoxite with one macroseta medially.
</paragraph>
<paragraph id="4E1B1F23861ED809D8EEF81BBD8DF83B" blockId="35.[112,764,144,1985]" box="[141,694,1961,1985]" pageId="35" pageNumber="36">Aedeagus as in Figure 11I, J; its total length 1.35–1.45.</paragraph>
<paragraph id="4E1B1F23861ED809DB49FF22BA6FFEFF" blockId="35.[810,1459,144,262]" pageId="35" pageNumber="36">
<emphasis id="7CD0C331861ED809DB49FF22BCA2FF52" box="[810,921,144,168]" italics="true" pageId="35" pageNumber="36">Etymology:</emphasis>
Ŋe species epithet is in honour of the Denmarkbased Carlsberg beer brewery, whose generous grants and beverages have facilitated much research in the Solodovnikov Lab. In addition, the male aedeagus resembles a boưle opener.
</paragraph>
<paragraph id="4E1B1F23861ED809DB49FE95BBDFFE67" blockId="35.[809,1459,295,413]" pageId="35" pageNumber="36">
<emphasis id="7CD0C331861ED809DB49FE95BBD7FEC5" box="[810,1260,295,319]" italics="true" pageId="35" pageNumber="36">
Distribution and habitat:
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</emphasis>
is known only from the Magallanes and Chilean
<collectingCountry id="36B35FB3861ED809DC24FEF5BB8AFEA5" box="[1095,1201,327,351]" name="Antarctica" pageId="35" pageNumber="36">Antarctica</collectingCountry>
Region of
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, where it occurs in lowlands confined to Magellanic subpolar forest ecoregions. It is unknown how it was collected.
</paragraph>
</subSubSection>
</treatment>
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539
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<taxonomicName id="89A464A08620D837D971F935BD32F958" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[274,521,1671,1698]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides" status="sp. nov.">
<emphasis id="7CD0C3318620D837D971F935BD32F958" bold="true" box="[274,521,1671,1698]" italics="true" pageId="29" pageNumber="30">Loncovilius cantharoides</emphasis>
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<emphasis id="7CD0C3318620D837D894F974BD4EF91A" bold="true" box="[247,629,1734,1760]" pageId="29" pageNumber="30">
(
<figureCitation id="D69F03A68620D837D961F974BE74F91A" box="[258,335,1734,1760]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." figureDoi="http://doi.org/10.5281/zenodo.14443963" httpUri="https://zenodo.org/record/14443963/files/figure.png" pageId="29" pageNumber="30">Figs 6B</figureCitation>
,
<figureCitation id="D69F03A68620D837D93AF974BE4FF91A" box="[345,372,1734,1760]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." figureDoi="http://doi.org/10.5281/zenodo.14443965" httpUri="https://zenodo.org/record/14443965/files/figure.png" pageId="29" pageNumber="30">7F</figureCitation>
,
<figureCitation id="D69F03A68620D837D91DF974BEE9F91A" box="[382,466,1734,1760]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="29" pageNumber="30">10O–Q</figureCitation>
,
<figureCitation id="D69F03A68620D837D9BFF974BD3EF91A" box="[476,517,1734,1760]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="29" pageNumber="30">12F</figureCitation>
,
<figureCitation id="D69F03A68620D837DA6DF974BD0CF91A" box="[526,567,1734,1760]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="29" pageNumber="30">13F</figureCitation>
,
<figureCitation id="D69F03A68620D837DA23F974BD51F91A" box="[576,618,1734,1760]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="29" pageNumber="30">14F</figureCitation>
)
</emphasis>
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<emphasis id="7CD0C3318620D837D812F8B5BE5BF8E4" box="[113,352,1799,1823]" italics="true" pageId="29" pageNumber="30">Zoobank registration:</emphasis>
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urn:lsid:zoobank.org:act:
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<emphasis id="7CD0C3318620D837D812F8D2BE3EF88D" box="[113,261,1887,1911]" italics="true" pageId="29" pageNumber="30">Type material.</emphasis>
<materialsCitation id="FECC157E8620D837D96CF8EDBDC1F84C" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectorName="Km E Malalcahuello" country="Chile" elevation="1080" location="Malleco Prov." municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
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:
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male, mounted, with labels as follows:
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:
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6.5
<collectorName id="E3517AF58620D837D9AEF8CDBD98F86D" box="[461,675,1919,1943]" pageId="29" pageNumber="30">
Km E
<collectingMunicipality id="AE7F85598620D837DA70F8CDBD98F86D" box="[531,675,1919,1943]" pageId="29" pageNumber="30">Malalcahuello</collectingMunicipality>
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, trap site 651
<date id="3A1A39E38620D837D961F82CBEACF84C" box="[258,407,1950,1974]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">
<collectingDate id="2A5EC00B8620D837D961F82CBEACF84C" box="[258,407,1950,1974]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</collectingDate>
</date>
North. Dombeyi with Chusquea
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</paragraph>
<paragraph id="4E1B1F238620D834DB49FF22BDEEFEEB" blockId="29.[809,1461,144,1985]" lastBlockId="30.[128,780,155,368]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
A. Newton and M. Ŋayer/window trap/
<taxonomicName id="89A464A08620D837DC93FF22BAA1FF52" authority="Sp." authorityName="Sp." box="[1264,1434,144,168]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius Sp.</taxonomicName>
5 Solodovnikov det. 2002/
<taxonomicName id="89A464A08620D837DC55FF02BABAFF3D" authority="J. Jenkins Shaw" authorityName="J. Jenkins Shaw" box="[1078,1409,175,199]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides">
<emphasis id="7CD0C3318620D837DC55FF02BB7EFF3D" box="[1078,1093,176,199]" italics="true" pageId="29" pageNumber="30">L</emphasis>
.
<emphasis id="7CD0C3318620D837DC35FF1DBBEAFF3D" box="[1110,1233,175,199]" italics="true" pageId="29" pageNumber="30">cantharoides</emphasis>
J. Jenkins Shaw
</taxonomicName>
det. 2016/NHMD 916904/
<typeStatus id="911FA1818620D837DC40FF7DBB8AFF1D" box="[1059,1201,207,231]" pageId="29" pageNumber="30" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="89A464A08620D837DCDDFF7DBA88FF1D" box="[1214,1459,207,231]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides">
<emphasis id="7CD0C3318620D837DCDDFF7DBA88FF1D" box="[1214,1459,207,231]" italics="true" pageId="29" pageNumber="30">Loncovilius cantharoides</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNM.
<emphasis id="7CD0C3318620D837DC3BFEBCBB85FEDF" box="[1112,1214,270,293]" italics="true" pageId="29" pageNumber="30">
<typeStatus id="911FA1818620D837DC3BFEBCBB81FEDF" box="[1112,1210,270,293]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="36B35FB38620D837DCA4FEBFBA21FEDF" box="[1223,1306,269,293]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: Malleco Prov.
<quantity id="895CB2C68620D837DB49FE9EBC49FEBF" box="[810,882,300,325]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="30" unit="km" value="6.5">6.5 km</quantity>
E Malalcahuello [-38.46, -71.51],
<quantity id="895CB2C68620D837DCB4FE9FBA1AFEBF" box="[1239,1313,301,325]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">1080m</quantity>
, trap site 651,
<date id="3A1A39E38620D837DB4FFEFEBCFBFE9E" box="[812,960,332,356]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</date>
<taxonomicName id="89A464A08620D837DBB1FEFEBB41FE9E" box="[978,1146,332,356]" pageId="29" pageNumber="30" rank="species" species="dombeyi">Noth. dombeyi</taxonomicName>
[
<taxonomicName id="89A464A08620D837DCF6FEFEBA52FE9E" box="[1173,1385,332,356]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="dombeyi">
<emphasis id="7CD0C3318620D837DCF6FEFEBA52FE9E" box="[1173,1385,332,356]" italics="true" pageId="29" pageNumber="30">Nothofagus dombeyi</emphasis>
</taxonomicName>
] with Chusquea A.Newton and M.Ŋayer/window trap/FMNH-INS 0000 023 458–463, 465–466, 467–471, 472–474, 476–477, 478, 481, 484–485 [
<specimenCount id="58A2D4AA8620D837DC71FE18BB72FE3B" box="[1042,1097,426,450]" count="13" pageId="29" pageNumber="30" type="male">13♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DC3BFE18BB44FE3B" box="[1112,1151,426,450]" count="9" pageId="29" pageNumber="30" type="female">9♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DC9FFE18BA74FE38" box="[1276,1359,426,450]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: Malleco Prov.,
<quantity id="895CB2C68620D837DB0FFE7BBC94FE1B" box="[876,943,457,481]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">40 km</quantity>
W [
<quantity id="895CB2C68620D837DBBCFE7BBB1DFE1B" box="[991,1062,457,481]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">40 km</quantity>
E] Curacautin [-38.44, -71.50],
<quantity id="895CB2C68620D837DD1CFE7BBC7BFDFA" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</quantity>
,
<date id="3A1A39E38620D837DB3AFE5BBCEEFDFB" box="[857,981,489,513]" pageId="29" pageNumber="30" value="1984-12-12">12.XII.1984</date>
-
<date id="3A1A39E38620D837DC64FE5ABB55FDFA" box="[1031,1134,488,512]" pageId="29" pageNumber="30" value="1985-02-16">16.II.1985</date>
/FMHD#85-905,
<emphasis id="7CD0C3318620D837DD58FE5BBA88FDFA" box="[1339,1459,489,513]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="89A464A08620D837DD58FE5BBA90FDFB" box="[1339,1451,489,513]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
-
</emphasis>
<taxonomicName id="89A464A08620D837DB49FDBABB3EFDDA" authority=", S. Peck, P" authorityName="S. Peck, P" box="[810,1029,520,544]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="7CD0C3318620D837DB49FDBABCB7FDE5" box="[810,908,520,543]" italics="true" pageId="29" pageNumber="30">Araucaria</emphasis>
, S. Peck, P
</taxonomicName>
#85-19, Malaise FIELD MUSEUM NAT HIST/FMNH-INS 0000 023 490–532, 602–613 [
<specimenCount id="58A2D4AA8620D837DD5DFD95BA4EFDC4" box="[1342,1397,551,575]" count="35" pageId="29" pageNumber="30" type="male">35♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DD1CFD95BA88FDC4" box="[1407,1459,551,575]" count="24" pageId="29" pageNumber="30" type="female">24♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DBEDFDF5BCDAFDA5" box="[910,993,583,607]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: Malleco Prov.
<quantity id="895CB2C68620D837DC14FDF5BB8DFDA5" box="[1143,1206,583,607]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="km" value="14.0">14 km</quantity>
E Malalcahuello [-38.43, -71.48]
<quantity id="895CB2C68620D837DBE3FDD4BCF1FD84" box="[896,970,614,638]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="29" pageNumber="30" unit="m" value="1570.0">1570m</quantity>
, trap site 649,
<date id="3A1A39E38620D837DC10FDD4BA3CFD84" box="[1139,1287,614,638]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</date>
Noth. pumilio-
<taxonomicName id="89A464A08620D837DB49FD37BCB7FD67" box="[810,908,645,669]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
f. [
<emphasis id="7CD0C3318620D837DBDAFD37BBD2FD67" box="[953,1257,645,669]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="89A464A08620D837DBDAFD37BB45FD67" box="[953,1150,645,669]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="pumilio">Nothofagus pumilio</taxonomicName>
-
<taxonomicName id="89A464A08620D837DCE4FD34BBD2FD67" box="[1159,1257,646,669]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
</emphasis>
forest], A.Newton and M.Ŋayer/window trap 649/FMNH-INS 0000 023 475, 478, 480, 482, 486–489 [
<specimenCount id="58A2D4AA8620D837DC54FD76BB5BFD21" box="[1079,1120,708,732]" count="3" pageId="29" pageNumber="30" type="male">3♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DC0AFD76BBABFD21" box="[1129,1168,708,732]" count="5" pageId="29" pageNumber="30" type="female">5♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DD61FD76BA6EFD26" box="[1282,1365,708,732]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: Malleco Prov.
<quantity id="895CB2C68620D837DB04FD51BC99FD01" box="[871,930,739,763]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="km" value="12.0">12km</quantity>
E Malacahuello [-38.43, -71.49]
<quantity id="895CB2C68620D837DD65FD51BA6BFD01" box="[1286,1360,739,763]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="29" pageNumber="30" unit="m" value="1350.0">1350m</quantity>
, trap site 650
<date id="3A1A39E38620D837DB3AFCB1BCD6FCE1" box="[857,1005,771,795]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</date>
Noth. dombeyi-Araucaria f. A.Newton and M.Ŋayer/carrion trap (squid)/
<specimenCount id="58A2D4AA8620D837DCE8FC97BB9EFCC3" box="[1163,1189,805,825]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
/FMNH-INS 0000 023 595 [
<specimenCount id="58A2D4AA8620D837DB06FCF3BCB6FCA2" box="[869,909,833,857]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DC62FCF3BB6AFCA3" box="[1025,1105,833,857]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
, MallecoProv.
<quantity id="895CB2C68620D837DC8DFCF3BA11FCA3" box="[1262,1322,833,857]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="km" value="20.0">20km</quantity>
E Manzanar [-38.44, -71.51]
<quantity id="895CB2C68620D837DBBAFCD3BB1AFC83" box="[985,1057,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">1100m</quantity>
<date id="3A1A39E38620D837DC49FCD3BB84FC83" box="[1066,1215,865,889]" pageId="29" pageNumber="30" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">19-21.xii.1976</date>
H.F.Howden/Beating/ CNCI L2004-28/
<specimenCount id="58A2D4AA8620D837DB8FFC31BB3DFC6D" box="[1004,1030,899,919]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
/FMNH-INS 0000 023 593-594 [
<specimenCount id="58A2D4AA8620D837DDEAFC32BA88FC6D" box="[1417,1459,896,920]" count="2" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
CNC];
<collectingCountry id="36B35FB38620D837DB1FFC2DBCF4FC4D" box="[892,975,927,951]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: Malleco: Las Raices (W of) [-38.54, -71.45],
<date id="3A1A39E38620D837DB49FC0DBCF2FC2D" box="[810,969,959,983]" pageId="29" pageNumber="30" value="1976-12-21" valueMax="1976-12-22" valueMin="1976-12-21">21-22.XII.1976</date>
L.E.Peña leg. FIELD MUSE. NAT. HIST./ FMNH-INS 0000 023 596 [
<specimenCount id="58A2D4AA8620D837DC2AFC6CBB49FC0F" box="[1097,1138,990,1014]" count="2" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38620D837DC87FC6CBA01FC0C" box="[1252,1338,990,1014]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18620D837DD21FC6CBA96FC0C" box="[1346,1453,990,1014]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
] Rio Blanco Malleco Prov. [-39.10, -71.61]/I-27-59 Pena/ CNC 928331-928333 [
<specimenCount id="58A2D4AA8620D837DC47FBAFBB70FBCE" box="[1060,1099,1053,1077]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DC36FBAFBB47FBCE" box="[1109,1148,1053,1077]" count="2" pageId="29" pageNumber="30" type="female">2♀</specimenCount>
CNC]; [
<collectingCountry id="36B35FB38620D837DC83FBAFBA0DFBCF" box="[1248,1334,1053,1077]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18620D837DD21FBAFBA96FBCF" box="[1346,1453,1053,1077]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
] Cabrerias 1100 metro Cord. Nahuelbuta [-37.71, -73.03]
<date id="3A1A39E38620D837DB49FBE9BCB4FB89" box="[810,911,1115,1139]" pageId="29" pageNumber="30" value="1977-01-18">18.I.1977</date>
/
<taxonomicName id="89A464A08620D837DBFAFBE9BB02FB89" authority="sp." authorityName="Sp." box="[921,1081,1115,1139]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318620D837DBFAFBE9BB3CFB89" box="[921,1031,1115,1139]" italics="true" pageId="29" pageNumber="30">Loncovilius</emphasis>
sp.
</taxonomicName>
det. Newton 2000/
<specimenCount id="58A2D4AA8620D837DD55FBECBA6BFB88" box="[1334,1360,1118,1138]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
/FMNH-INS 0000 023 577 [
<specimenCount id="58A2D4AA8620D837DC69FBC9BB0AFB68" box="[1034,1073,1147,1171]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DC5FFBC9BB58FB68" box="[1084,1123,1147,1171]" count="2" pageId="29" pageNumber="30" type="female">2♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DCBBFBC9BA10FB69" box="[1240,1323,1147,1171]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18620D837DD5DFBC9BA96FB69" box="[1342,1453,1147,1171]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
] Malleco: Cord. Nahuelbuta, Cabreria [-37.71, -73.03]
<quantity id="895CB2C68620D837DD06FB28BA94FB48" box="[1381,1455,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">1100m</quantity>
,
<date id="3A1A39E38620D837DB49FB0BBC9CFB2B" box="[810,935,1209,1233]" pageId="29" pageNumber="30" value="1976-12-14">14.XII.1976</date>
L.E.Peña leg. FIELD MUS. NAT. HIST./FMNH-INS 0000 023 585-590, 592, 600 [
<specimenCount id="58A2D4AA8620D837DCCBFB6BBBE9FB0A" box="[1192,1234,1241,1265]" count="6" pageId="29" pageNumber="30" type="male">6♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DCBEFB6BBA3FFB0A" box="[1245,1284,1241,1265]" count="2" pageId="29" pageNumber="30" type="female">2♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DD18FB6BBA88FB0B" box="[1403,1459,1241,1265]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
[
<collectingRegion id="8C60D1C18620D837DB53FB4ABCA6FAEA" box="[816,925,1272,1296]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
, Curacautín]: P.N.Lonquimay,
<date id="3A1A39E38620D837DD4FFB4ABA95FAEA" box="[1324,1454,1272,1296]" pageId="29" pageNumber="30" value="2013-12-23">23/12/2013</date>
,
<geoCoordinate id="2B9079E48620D837DB49FAA5BC42FAD5" box="[810,889,1303,1327]" degrees="38.44" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="-38.44">38.44°S</geoCoordinate>
–
<geoCoordinate id="2B9079E48620D837DBE4FAA5BCDAFAD5" box="[903,993,1303,1327]" degrees="71.51" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-71.51">71.51°W</geoCoordinate>
[
<geoCoordinate id="2B9079E48620D837DB8FFAA5BB75FAD5" box="[1004,1102,1303,1327]" degrees="38.4400" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="-38.44">-38.4400</geoCoordinate>
,
<geoCoordinate id="2B9079E48620D837DC34FAA5BB82FAD5" box="[1111,1209,1303,1327]" degrees="71.5100" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="-71.51">-71.5100</geoCoordinate>
], liưer layer small stream [
<specimenCount id="58A2D4AA8620D837DB57FA85BC67FAB4" box="[820,860,1335,1359]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
TSC];
<collectingCountry id="36B35FB38620D837DBD2FA85BCD2FAB5" box="[945,1001,1335,1359]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
[
<collectingRegion id="8C60D1C18620D837DB9AFA85BB5DFAB5" box="[1017,1126,1335,1359]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
, Pucón]: Palguin,
<date id="3A1A39E38620D837DD4FFA84BA94FAB4" box="[1324,1455,1334,1358]" pageId="29" pageNumber="30" value="2013-12-05">05/12/2013</date>
,
<geoCoordinate id="2B9079E48620D837DB49FAE4BC4CFA94" box="[810,887,1366,1390]" degrees="39.43" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="-39.43">39.43°S</geoCoordinate>
–
<geoCoordinate id="2B9079E48620D837DBE6FAE4BCDAFA94" box="[901,993,1366,1390]" degrees="71.79" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-71.79">71.79°W</geoCoordinate>
[
<geoCoordinate id="2B9079E48620D837DB8CFAE4BB6AFA94" box="[1007,1105,1366,1390]" degrees="39.4300" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="-39.43">-39.4300</geoCoordinate>
,
<geoCoordinate id="2B9079E48620D837DC3FFAE4BB85FA94" box="[1116,1214,1366,1390]" degrees="71.7900" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="-71.79">-71.7900</geoCoordinate>
], liưer layer/
<taxonomicName id="89A464A08620D837DD26FAE4BA88FA94" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[1349,1459,1366,1390]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" status="n. sp.">
<emphasis id="7CD0C3318620D837DD26FAE4BA88FA94" box="[1349,1459,1366,1390]" italics="true" pageId="29" pageNumber="30">Loncovilius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67E37E4A8620D837DB49FAC4BC5FFA77" box="[810,868,1398,1421]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
[
<specimenCount id="58A2D4AA8620D837DB1BFAC7BC9BFA76" box="[888,928,1397,1421]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DBC8FAC7BCE9FA76" box="[939,978,1397,1421]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
TSC]; [
<collectingCountry id="36B35FB38620D837DC4CFAC7BBBEFA77" box="[1071,1157,1397,1421]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18620D837DCF0FAC7BBC7FA77" box="[1171,1276,1397,1421]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
, Melipeuco]
<specimenCount id="58A2D4AA8620D837DDEFFACABA9DFA76" box="[1420,1446,1400,1420]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
/
<collectingCountry id="36B35FB38620D837DB49FA27BC5FFA57" box="[810,868,1429,1453]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
, La
<collectingRegion id="8C60D1C18620D837DBFBFA27BCC4FA57" box="[920,1023,1429,1453]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucania</collectingRegion>
(IX Región),
<quantity id="895CB2C68620D837DCFDFA26BBDFFA57" box="[1182,1252,1428,1453]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="29" pageNumber="30" unit="km" value="85.0">85 km</quantity>
E Temuco,
<quantity id="895CB2C68620D837DD0EFA27BA88FA57" box="[1389,1459,1429,1453]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="km" value="26.0">26 km</quantity>
NNE Melipeuco, Parque Nacional Corguillío, NE coast of Laguna Conguillío,
<geoCoordinate id="2B9079E48620D837DC6EFA61BB47FA11" box="[1037,1148,1490,1515]" degrees="38" direction="south" minutes="38.24" orientation="latitude" pageId="29" pageNumber="30" precision="9" value="-38.637333">
38°38.24
<emphasis id="7CD0C3318620D837DC0BFA60BB55FA11" box="[1128,1134,1490,1515]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
S
</geoCoordinate>
, W
<geoCoordinate id="2B9079E48620D837DCDFFA61BA0DFA11" box="[1212,1334,1490,1515]" degrees="71" direction="west" minutes="36.74" orientation="longitude" pageId="29" pageNumber="30" precision="9" value="-71.612335">
71°36.74
<emphasis id="7CD0C3318620D837DD7BFA60BA25FA11" box="[1304,1310,1490,1515]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
W
</geoCoordinate>
[
<geoCoordinate id="2B9079E48620D837DD2EFA61BA94FA11" box="[1357,1455,1491,1515]" degrees="38.6291" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="-38.6291">-38.6291</geoCoordinate>
,
<geoCoordinate id="2B9079E48620D837DB49FA41BCB7F9F1" box="[810,908,1523,1547]" degrees="71.6203" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="-71.6203">-71.6203</geoCoordinate>
],
<quantity id="895CB2C68620D837DBC0FA41BCC1F9F0" box="[931,1018,1523,1547]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="m" value="1200.0">1200 m</quantity>
V.I.Gusarov
<date id="3A1A39E38620D837DCEFFA41BBD5F9F1" box="[1164,1262,1523,1547]" pageId="29" pageNumber="30" value="1999-12-09">9.xii.1999</date>
/
<taxonomicName id="89A464A08620D837DC95FA41BA99F9F0" authority="sp." authorityName="Sp." box="[1270,1442,1523,1547]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
Loncovilius
<taxonomicNameLabel id="67E37E4A8620D837DDE6FA41BA99F9F0" box="[1413,1442,1523,1546]" pageId="29" pageNumber="30" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
n. A.Solodovnikov det. 2004 [
<specimenCount id="58A2D4AA8620D837DC01F9A0BBB1F9D3" box="[1122,1162,1554,1578]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
NHMO];
<collectingCountry id="36B35FB38620D837DD6DF9A0BA5AF9D0" box="[1294,1377,1554,1578]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
: Nuble [Ñuble] Prov.,
<quantity id="895CB2C68620D837DBAAF983BB37F9B3" box="[969,1036,1585,1609]" metricMagnitude="4" metricUnit="m" metricValue="7.2" pageId="29" pageNumber="30" unit="km" value="72.0">72 km</quantity>
SE Chillan, Trancas nr Termas [-36.90, -71.44],
<quantity id="895CB2C68620D837DBF9F9E3BCDFF993" box="[922,996,1617,1641]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="29" pageNumber="30" unit="m" value="1700.0">1700m</quantity>
,
<date id="3A1A39E38620D837DC64F9E2BB43F992" box="[1031,1144,1616,1640]" pageId="29" pageNumber="30" value="1984-12-06">6.XII.1984</date>
-
<date id="3A1A39E38620D837DCE3F9E2BBDDF992" box="[1152,1254,1616,1640]" pageId="29" pageNumber="30" value="1985-02-19">19.II.1985</date>
/FMNHD#85-893,
<taxonomicName id="89A464A08620D837DB49F9C2BC98F972" box="[810,931,1648,1672]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
forest, S.Peck, P#85-8 FIT FIELD MUSEUM NAT HIST/FMNH-INS 0000 023 578–584 [
<specimenCount id="58A2D4AA8620D837DC84F93DBA2BF95C" box="[1255,1296,1679,1703]" count="3" pageId="29" pageNumber="30" type="male">3♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DD7DF93DBA7EF95C" box="[1310,1349,1679,1703]" count="4" pageId="29" pageNumber="30" type="female">4♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38620D837DB49F91DBCFFF93D" box="[810,964,1711,1735]" name="Argentina" pageId="29" pageNumber="30">ARGENTINA</collectingCountry>
,
<collectingRegion id="8C60D1C18620D837DBACF91DBB06F93D" box="[975,1085,1711,1735]" country="Argentina" name="Rio Negro" pageId="29" pageNumber="30">Rio Negro</collectingRegion>
8: Lago Nahuel Huapi Puerto Blest [-41.02, -71.81],
<quantity id="895CB2C68620D837DB86F97CBB17F91F" box="[997,1068,1742,1766]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="29" pageNumber="30" unit="m" value="770.0">770 m</quantity>
<date id="3A1A39E38620D837DC54F97CBBA7F91C" box="[1079,1180,1742,1766]" pageId="29" pageNumber="30" value="1978-12-04">4.xii.1978</date>
Mision Cientifica Danesa [pale orange label
<specimenCount id="58A2D4AA8620D837DB9EF95FBB1FF8FE" box="[1021,1060,1773,1797]" count="2" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DC52F95FBB63F8FE" box="[1073,1112,1773,1797]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
NHMD;
<specimenCount id="58A2D4AA8620D837DCADF95FBBCEF8FE" box="[1230,1269,1773,1797]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DD60F95FBA11F8FE" box="[1283,1322,1773,1797]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
MNNC ex. NHMD;
<specimenCount id="58A2D4AA8620D837DBEEF8BFBC8FF8DE" box="[909,948,1805,1829]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DBDFF8BFBCD8F8DE" box="[956,995,1805,1829]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
CZUG ex. NHMD]; [
<collectingCountry id="36B35FB38620D837DCB0F8BFBA4FF8DF" box="[1235,1396,1805,1829]" name="Argentina" pageId="29" pageNumber="30">ARGENTINA</collectingCountry>
: same locality, only the date different (
<date id="3A1A39E38620D837DC17F89EBBD6F8BE" box="[1140,1261,1836,1860]" pageId="29" pageNumber="30" value="1978-12-16">16.xii.1978</date>
)
<specimenCount id="58A2D4AA8620D837DC9FF89EBA18F8B9" box="[1276,1315,1836,1860]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DD49F89EBA6AF8B9" box="[1322,1361,1836,1860]" count="3" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
NHMD;
<specimenCount id="58A2D4AA8620D837DB4FF8F9BC6AF898" box="[812,849,1867,1891]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8620D837DB3CF8F9BCBDF898" box="[863,902,1867,1891]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
NHM ex. NHMD];
<collectingCountry id="36B35FB38620D837DC1CF8F9BBDCF899" box="[1151,1255,1867,1891]" name="Argentina" pageId="29" pageNumber="30">Argentina</collectingCountry>
,
<collectingRegion id="8C60D1C18620D837DC9BF8F9BA8BF899" box="[1272,1456,1867,1892]" country="Argentina" name="Rio Negro" pageId="29" pageNumber="30">Rio Negro Prov.</collectingRegion>
, Bariloche forest,
<taxonomicName id="89A464A08620D837DB88F8D9BB5FF879" box="[1003,1124,1899,1923]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
antartica, unburnt,
<geoCoordinate id="2B9079E48620D837DD2FF8D9BA88F879" box="[1356,1459,1897,1923]" degrees="41" direction="south" minutes="21" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="9" value="-41.352497">
41°21
<emphasis id="7CD0C3318620D837DDEBF8DBBAB5F878" box="[1416,1422,1897,1922]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
9
<emphasis id="7CD0C3318620D837DDF9F8DBBA9EF878" box="[1434,1445,1897,1922]" italics="true" pageId="29" pageNumber="30">ʹʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="2B9079E48620D837DB49F838BCEAF85B" box="[810,977,1928,1954]" degrees="71" direction="west" minutes="37" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="34.5" value="-71.62625">
71°37
<emphasis id="7CD0C3318620D837DB05F83ABC57F85B" box="[870,876,1928,1953]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
34.5
<emphasis id="7CD0C3318620D837DBFAF83ABC9FF85B" box="[921,932,1928,1953]" italics="true" pageId="29" pageNumber="30">ʹʹ</emphasis>
W
</geoCoordinate>
, [
<geoCoordinate id="2B9079E48620D837DB8CF838BB50F858" box="[1007,1131,1930,1954]" degrees="41.352500" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="-41.3525">-41.352500</geoCoordinate>
,
<geoCoordinate id="2B9079E48620D837DCE7F838BBC4F858" box="[1156,1279,1930,1954]" degrees="71.626250" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="-71.62625">-71.626250</geoCoordinate>
] 823 ±
<quantity id="895CB2C68620D837DD3EF838BA94F85B" box="[1373,1455,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="150.0">150 m</quantity>
, ABCONF
<specimenCount id="58A2D4AA8620D837DBF4F81BBCF5F83B" box="[919,974,1961,1985]" count="17" pageId="29" pageNumber="30" type="generic">17 P.</specimenCount>
Sackmann/
<collectingCountry id="36B35FB38620D837DC21F81BBB90F83B" box="[1090,1195,1961,1985]" name="Argentina" pageId="29" pageNumber="30">Argentina</collectingCountry>
, Patagonia ABCONF17 Paula Sackmann leg./
<taxonomicName id="89A464A08623D834D939FF29BEC0FF49" authority="sp." authorityName="Sp." box="[346,507,155,179]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp.</taxonomicName>
<specimenCount id="58A2D4AA8623D834DA60FF2DBD26FF49" box="[515,541,159,179]" count="1" pageId="30" pageNumber="31" type="female">♀</specimenCount>
V.I. Gusarov det. 2002 [
<specimenCount id="58A2D4AA8623D834D8E8FF09BF89FF28" box="[139,178,187,211]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
NHMO];
<collectingCountry id="36B35FB38623D834D944FF09BEFAFF29" box="[295,449,187,211]" name="Argentina" pageId="30" pageNumber="31">ARGENTINA</collectingCountry>
,
<collectingRegion id="8C60D1C18623D834D9AFFF09BD0BFF29" box="[460,560,187,211]" country="Argentina" name="Neuquen" pageId="30" pageNumber="31">Neuquén</collectingRegion>
: Cerro Malo,
<quantity id="895CB2C68623D834DAA3FF09BDD5FF29" box="[704,750,187,211]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="km" value="6.0">6km</quantity>
N Pucara,
<date id="3A1A39E38623D834D8B0FF69BE6DFF08" box="[211,342,218,242]" pageId="30" pageNumber="31" value="1972-01-22">Jan. 22, 1972</date>
L.Herman 874/AMNH 1489 [pale orange label]/FMNH-INS 0000 023 597–599 [
<specimenCount id="58A2D4AA8623D834DA7FFF48BD7FFEEB" box="[540,580,250,274]" count="1" pageId="30" pageNumber="31" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8623D834DA2AFF48BD4BFEEB" box="[585,624,250,274]" count="2" pageId="30" pageNumber="31" type="female">2♀</specimenCount>
FMNH].
</paragraph>
<paragraph id="4E1B1F238623D834D8FFFEABBF80FE8A" blockId="30.[128,780,155,368]" pageId="30" pageNumber="31">
All
<typeStatus id="911FA1818623D834D8DCFEABBE19FECB" box="[191,290,281,305]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
with label: PAÞTYPE
<taxonomicName id="89A464A08623D834DA7FFEABBC31FECB" box="[540,778,281,305]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="cantharoides">
<emphasis id="7CD0C3318623D834DA7FFEABBC31FECB" box="[540,778,281,305]" italics="true" pageId="30" pageNumber="31">Loncovilius cantharoides</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
</paragraph>
<paragraph id="4E1B1F238623D834D8E2FE20BDB3FDDC" blockId="30.[128,778,401,550]" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834D8E2FE20BFDCFE53" box="[129,231,402,425]" italics="true" pageId="30" pageNumber="31">Diagnosis:</emphasis>
Nuchal ridge with small portion absent dorsally. Pronotum outline evenly curved. Elytra yellow or yellowish-brown with brown-black mark around mesoscutellum and suture. Protibiae sexually dimorphic, with laterodorsal row of thick spines only females. Aedeagus as in
<figureCitation id="D69F03A68623D834D988FDBCBDB9FDDC" box="[491,642,526,550]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="30" pageNumber="31">Figure 10O–Q</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238623D834D8E2FDFABECEFD06" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834D8E2FDFABFCCFDA5" box="[129,247,584,607]" italics="true" pageId="30" pageNumber="31">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8623D834D9CBFDF9BEF9FDA5" box="[424,450,587,607]" count="1" pageId="30" pageNumber="31" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318623D834DAD9FDFABDF6FDA5" box="[698,717,584,607]" italics="true" pageId="30" pageNumber="31">N</emphasis>
= 5]: FBL = 3.98–4.34 (4.22); TL = 7.18–8.4 (7.59); HW = 0.93– 1.08 (1); HL = 1.03–1.23 (1.16); HW/HL = 0.88–0.91 (0.89); PW = 1.28–1.36 (1.32); PL = 1.25–1.36 (1.31); PW/ PL = 1–1.04 (1.02); EW = 1.74–1.92 (1.84); EL = 1.59–1.81 (1.7); PW/HW = 1.26–1.28 (1.27).
</paragraph>
<paragraph id="4E1B1F238623D834D8FFFCB6BE44FC42" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">
Measurements
<specimenCount id="58A2D4AA8623D834D954FCB5BE6AFCE1" box="[311,337,775,795]" count="1" pageId="30" pageNumber="31" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318623D834DA52FCB6BD7FFCE1" box="[561,580,772,795]" italics="true" pageId="30" pageNumber="31">N</emphasis>
= 5]: FBL = 4.43– 4.67 (4.55); TL = 8.13–8.47 (8.3); HW = 1.18–1.22 (1.20); HL = 1.25–1.33 (1.29); HW/HL = 0.92–0.94 (0.93); PW = 1.44–1.49 (1.47);PL = 1.4–1.44 (1.42);PW/PL = 1.03– 1.03 (1.03); EW = 2–2.1 (2.05); EL = 1.78–1.9 (1.84); PW/ HW = 4.43–4.67 (4.55).
</paragraph>
<paragraph id="4E1B1F238623D834D8FFFC0DBDD1FBAF" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">Head and pronotum brown-black with slight greenish, bluish, or golden overtones, with yellow margins; antennae, mouthparts, and legs yellow or yellowish-brown; elytra yellow with brown-black mark around mesoscutellum and suture; tergites and sternites dark reddish-brown with pale posterior margin.</paragraph>
<paragraph id="4E1B1F238623D834D8FFFBEEBD5CF89E" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">
Head elongate; dorsally and ventrally dull with metallic iridescence; with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.44), from 1.05 to 1.11 times as long as temples (in lateral view); distance between eyes about 1.24 times as long as length of eye in both sexes. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 and 5 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.52 to 1.7 times long as a
<quantity id="895CB2C68623D834D9E9FA24BE87FA57" box="[394,444,1429,1454]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="30" pageNumber="31" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68623D834DA06FA24BDD1FA57" box="[613,746,1429,1454]" metricMagnitude="-2" metricUnit="m" metricValue="3.6576" metricValueMax="3.7592" metricValueMin="3.5559999999999996" pageId="30" pageNumber="31" unit="in" value="1.44" valueMax="1.48" valueMin="1.4">1.4 to 1.48 in</quantity>
females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; with small setiferous punctures between frontoclypeal and anterior frontal punctures: ventral basal ridge in checkmark shape, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and a liưle dorsally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures widely and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
</paragraph>
<paragraph id="4E1B1F238623D834D8FFF8DEBA1DFF1D" blockId="30.[128,780,583,1986]" lastBlockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">Pronotum about as wide as long, convex, evenly curved, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to four or five times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F238623D834DB36FF5CBCA8FE1B" blockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">Protibiae sexually dimorphic, with laterodorsal row of thick spines only females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 1–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
<paragraph id="4E1B1F238623D834DB36FE5BBB35FCA3" blockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">
Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large, impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina evenly converging at obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with U-shaped emargination medially (
<figureCitation id="D69F03A68623D834DCF7FD37BBDDFD67" box="[1172,1254,645,669]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="30" pageNumber="31">Fig. 14F</figureCitation>
); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X with anterior margin subtruncate (
<figureCitation id="D69F03A68623D834DDFFFD51BC5AFCE1" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="30" pageNumber="31">Fig. 12F</figureCitation>
); female tergite X medio-apically distinctly subacute to acute (
<figureCitation id="D69F03A68623D834DBE0FC90BCE1FCC0" box="[899,986,802,826]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="30" pageNumber="31">Fig. 13F</figureCitation>
); ovipositor, its second gonocoxite with three macrosetae medially.
</paragraph>
<paragraph id="4E1B1F238623D834DB36FCD3BAF8FBEF" blockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">
Aedeagus as in
<figureCitation id="D69F03A68623D834DB9CFCD3BBA6FC83" box="[1023,1181,865,889]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="30" pageNumber="31">Figure 10O–Q</figureCitation>
; its total length 1.25–1.31. Median lobe rod-like; internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a subrectangular apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically.
</paragraph>
<paragraph id="4E1B1F238623D834DB5AFB85BC85FB57" blockId="30.[825,1474,1079,1197]" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834DB5AFB85BC93FBB5" box="[825,936,1079,1103]" italics="true" pageId="30" pageNumber="31">Etymology:</emphasis>
Ŋe species epithet is derived from the Greek κανθαΡίς (
<emphasis id="7CD0C3318623D834DBCBFBE4BB3FFB94" box="[936,1028,1110,1134]" italics="true" pageId="30" pageNumber="31">kantharís</emphasis>
; soldier beetles) and suffix -ειδής (
<emphasis id="7CD0C3318623D834DDE7FBE4BA81FB94" box="[1412,1466,1106,1134]" italics="true" pageId="30" pageNumber="31">
-eid
<emphasis id="7CD0C3318623D834DDCBFBE0BA88FB94" bold="true" box="[1448,1459,1106,1134]" italics="true" pageId="30" pageNumber="31">ḗ</emphasis>
s
</emphasis>
; like), as the habitus resembles some species of the beetle family
<taxonomicName id="89A464A08623D834DB59FB27BC81FB57" authorityName="Imhoff" authorityYear="1856" box="[826,954,1173,1197]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
.
</paragraph>
<paragraph id="4E1B1F238623D834DB5AFB7CBA3EFA3B" blockId="30.[825,1477,1230,1473]" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834DB5AFB7CBA13FB1C" box="[825,1320,1230,1254]" italics="true" pageId="30" pageNumber="31">
Distribution and habitat:
<taxonomicName id="89A464A08623D834DC5BFB7CBA13FB1C" box="[1080,1320,1230,1254]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="cantharoides">Loncovilius cantharoides</taxonomicName>
</emphasis>
is known from
<collectingCountry id="36B35FB38623D834DB58FB5FBC4EFAFF" box="[827,885,1261,1285]" name="Chile" pageId="30" pageNumber="31">Chile</collectingCountry>
, from the
<collectingRegion id="8C60D1C18623D834DBBBFB5FBB04FAFF" box="[984,1087,1261,1285]" country="Chile" name="Araucania" pageId="30" pageNumber="31">Araucanía</collectingRegion>
and Ñuble regions, and from the westernmost portions of the
<collectingRegion id="8C60D1C18623D834DC59FABFBBA1FADF" box="[1082,1178,1293,1317]" country="Argentina" name="Neuquen" pageId="30" pageNumber="31">Neuquén</collectingRegion>
and
<collectingRegion id="8C60D1C18623D834DCADFABFBA07FADF" box="[1230,1340,1293,1317]" country="Argentina" name="Rio Negro" pageId="30" pageNumber="31">Río Negro</collectingRegion>
provinces of
<collectingCountry id="36B35FB38623D834DB5FFA9EBC9FFABE" box="[828,932,1324,1348]" name="Argentina" pageId="30" pageNumber="31">Argentina</collectingCountry>
. Ŋe species occurs from
<quantity id="895CB2C68623D834DCCAFA9EBA7BFAB9" box="[1193,1344,1324,1348]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="30" pageNumber="31" unit="m" value="1200.0" valueMax="1600.0" valueMin="800.0">800 to 1600 m</quantity>
of elevation; reported in the
<emphasis id="7CD0C3318623D834DB86FAFEBB84FA99" box="[997,1215,1355,1379]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="89A464A08623D834DB86FAFEBB7CFA99" box="[997,1095,1356,1379]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
-
<taxonomicName id="89A464A08623D834DC2CFAF9BB84FA99" box="[1103,1215,1355,1379]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
</emphasis>
forests from the central and north-central portions of the Valdivian temperate forest ecoregion. It has been collected using Malaise, window, carrion and dung-baited pitfall traps, and sissing leaf liưer.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="89A464A08623D834DBD3FA56BA77FA04" ID-CoL="3VYHP" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[944,1356,1508,1535]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
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<emphasis id="7CD0C3318623D834DBD3FA56BB94FA05" bold="true" box="[944,1199,1508,1535]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsianus</emphasis>
(Korge, 1963)
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<emphasis id="7CD0C3318623D834DBDEF9ACBA04F9C2" bold="true" box="[957,1343,1566,1592]" pageId="30" pageNumber="31">
(
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,
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,
<figureCitation id="D69F03A68623D834DC29F9ACBBACF9C2" box="[1098,1175,1566,1592]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="30" pageNumber="31">10L–N</figureCitation>
,
<figureCitation id="D69F03A68623D834DCC2F9ACBBF7F9C2" box="[1185,1228,1566,1592]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="30" pageNumber="31">12E</figureCitation>
,
<figureCitation id="D69F03A68623D834DCB5F9ACBA3AF9C2" box="[1238,1281,1566,1592]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="30" pageNumber="31">13E</figureCitation>
,
<figureCitation id="D69F03A68623D834DD69F9ACBA0FF9C2" box="[1290,1332,1566,1592]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="30" pageNumber="31">14E</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238623D834DB5AF9EABC82F975" blockId="30.[825,1475,1623,1898]" pageId="30" pageNumber="31">
=
<taxonomicName id="89A464A08623D834DB2BF9E5BBE4F995" authority="(Bernhauer, 1935)" baseAuthorityName="Bernhauer" baseAuthorityYear="1935" box="[840,1247,1623,1647]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318623D834DB2BF9E5BB2FF995" box="[840,1044,1623,1647]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsi</emphasis>
(Bernhauer, 1935)
</taxonomicName>
nec
<taxonomicName id="89A464A08623D834DD70F9E5BC8EF975" authority="Sharp, 1886" authorityName="Bernh. Nec Sharp" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318623D834DD70F9E5BAF9F995" box="[1299,1474,1623,1647]" italics="true" pageId="30" pageNumber="31">Quedius edwardsi</emphasis>
Sharp, 1886
</taxonomicName>
.
</paragraph>
<paragraph id="4E1B1F238623D834DB36F924BB13F890" blockId="30.[825,1475,1623,1898]" pageId="30" pageNumber="31">
Bernhauer, 1935: 92 (as
<taxonomicName id="89A464A08623D834DC38F924BA31F954" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[1115,1290,1686,1710]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318623D834DC38F924BA31F954" box="[1115,1290,1686,1710]" italics="true" pageId="30" pageNumber="31">Quedius edwardsi</emphasis>
</taxonomicName>
, original description); Korge, 1963 (as
<taxonomicName id="89A464A08623D834DC28F907BA15F937" authorityName="Korge" authorityYear="1963" box="[1099,1326,1717,1741]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DC28F907BA15F937" box="[1099,1326,1717,1741]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
</taxonomicName>
, replacement name); Coiffait and Sáiz 1966: 413 (
<taxonomicName id="89A464A08623D834DCBFF967BA87F917" authorityName="Korge" authorityYear="1963" box="[1244,1468,1749,1773]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DCBFF967BA87F917" box="[1244,1468,1749,1773]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
</taxonomicName>
, redescription without new material or data); Coiffait and Sáiz, 1968: 365 (listed as
<taxonomicName id="89A464A08623D834DC72F8A1BBD6F8D1" authorityName="Korge" authorityYear="1963" box="[1041,1261,1811,1835]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DC72F8A1BBD6F8D1" box="[1041,1261,1811,1835]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
</taxonomicName>
, without new data); Newton 2017: 28 (as
<taxonomicName id="89A464A08623D834DC53F881BA16F8B1" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1072,1325,1843,1867]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DC53F881BA16F8B1" box="[1072,1325,1843,1867]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsianus</emphasis>
</taxonomicName>
, moved from
<taxonomicName id="89A464A08623D834DB5AF8E0BCB2F890" box="[825,905,1874,1898]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318623D834DB5AF8E0BCB2F890" box="[825,905,1874,1898]" italics="true" pageId="30" pageNumber="31">Quedius</emphasis>
</taxonomicName>
to
<taxonomicName id="89A464A08623D834DBCAF8E0BB2CF890" box="[937,1047,1874,1898]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318623D834DBCAF8E0BB2CF890" box="[937,1047,1874,1898]" italics="true" pageId="30" pageNumber="31">Loncovilius</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="4E1B1F238623D835DB5AF83EBE45FD87" blockId="30.[825,1474,1931,1987]" lastBlockId="31.[113,763,144,638]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834DB5AF83EBB7BF859" box="[825,1088,1931,1955]" italics="true" pageId="30" pageNumber="31">
Type material.
<typeStatus id="911FA1818623D834DBBFF83EBB00F859" box="[988,1083,1932,1955]" pageId="30" pageNumber="31" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
(here designated): male, pinned, with genitalia in a separate microvial, with labels as follows: S.
<collectingCountry id="36B35FB38622D835D812FF22BF95FF52" box="[113,174,144,168]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
: Llanquihue prov. F. and M. Edwards. B.M. 1927–63/ Casa Pangue [-41.04, -71.87]
<date id="3A1A39E38622D835D9A2FF1DBD7DFF3D" box="[449,582,175,199]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</date>
./Bernhauer det. [unreadable]/Chicago NHM M. Bernhauer collection/
<taxonomicName id="89A464A08622D835D812FF5CBE5CFEFC" authority="Brh." authorityName="Bernhauer" authorityYear="2024" box="[113,359,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi" status="sp. nov.">Quedius edwardsi Brh.</taxonomicName>
<taxonomicNameLabel id="67E37E4A8622D835D913FF5CBE93FEFC" box="[368,424,238,262]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
/
<taxonomicName id="89A464A08622D835D9D3FF5CBDB0FEFC" authority="Bernhauer" authorityName="Bernhauer" authorityYear="2024" box="[432,651,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">Edwardsi Bernhauer</taxonomicName>
<taxonomicName id="89A464A08622D835DAF5FF5CBDC9FEFC" box="[662,754,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
/
<typeStatus id="911FA1818622D835D812FEBFBE3EFEDF" box="[113,261,269,293]" pageId="31" pageNumber="32" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="89A464A08622D835D976FEBFBDF8FEDF" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[277,707,269,293]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318622D835D976FEBFBD2DFEDF" box="[277,534,269,293]" italics="true" pageId="31" pageNumber="32">Loncovilius edwardsianus</emphasis>
(Korge, 1963)
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
<emphasis id="7CD0C3318622D835D931FEFEBEE7FE9E" box="[338,476,332,356]" italics="true" pageId="31" pageNumber="32">
<typeStatus id="911FA1818622D835D931FEFEBEECFE9E" box="[338,471,332,356]" pageId="31" pageNumber="32" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
male, pinned, with genitalia aưached to the card, with labels as follows: S.
<collectingCountry id="36B35FB38622D835DA5CFED9BD47FE79" box="[575,636,363,387]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
: Llanquihue prov. F. and M. Edwards. B. M. 1927–63/Casa Pangue [-41.04, -71.87]
<date id="3A1A39E38622D835D8B4FE18BE64FE38" box="[215,351,426,450]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</date>
./
<taxonomicName id="89A464A08622D835D908FE18BEF2FE38" box="[363,457,426,450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
<taxonomicName id="89A464A08622D835D983FE18BDB5FE38" authority="Bernh" authorityName="Bernhauer" authorityYear="2024" box="[480,654,426,450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi" status="sp. nov.">
<emphasis id="7CD0C3318622D835D983FE18BD0DFE38" box="[480,566,426,450]" italics="true" pageId="31" pageNumber="32">edwardsi</emphasis>
Bernh
</taxonomicName>
<taxonomicNameLabel id="67E37E4A8622D835DAC4FE18BDCAFE38" box="[679,753,426,450]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
/
<typeStatus id="911FA1818622D835D812FE7BBE3EFE1B" box="[113,261,457,481]" pageId="31" pageNumber="32" type="lectotype">LECTOTYPE</typeStatus>
C.E.Toưenham of
<taxonomicName id="89A464A08622D835D9A6FE7BBDC9FE1B" authority="Bernh. Nec Sharp" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[453,754,457,481]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">Q.edwardsi Bernh.Nec Sharp</taxonomicName>
/
<taxonomicName id="89A464A08622D835D812FE5BBFF2FDFB" box="[113,201,489,513]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
ultor n. n. Toưenham/edwardsi repl. By
<taxonomicName id="89A464A08622D835DA13FE5BBEAFFDDA" authority="Korge 1963 M. K. Nayer" authorityName="Korge" authorityYear="1963" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">edwardsianus Korge 1963 M. K. Ŋayer</taxonomicName>
det. 2005/NHMUK015024785/ PAÞLECTOTYPE
<taxonomicName id="89A464A08622D835D930FD95BDC1FDC5" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[339,762,551,575]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318622D835D930FD95BD6AFDC5" box="[339,593,551,575]" italics="true" pageId="31" pageNumber="32">Loncovilius edwardsianus</emphasis>
(Korge, 1963)
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023. Deposited at NHM.
</paragraph>
<paragraph id="4E1B1F238622D835D812FD2DBD85FD4D" blockId="31.[113,702,671,695]" box="[113,702,671,695]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835D812FD2DBE57FD4D" box="[113,364,671,695]" italics="true" pageId="31" pageNumber="32">Other material eoamined:</emphasis>
Supporting Information, File S7.
</paragraph>
<paragraph id="4E1B1F238622D835D812FD6BBD1CFC97" blockId="31.[113,763,729,878]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835D812FD6BBFECFD0A" box="[113,215,729,752]" italics="true" pageId="31" pageNumber="32">Diagnosis:</emphasis>
Nuchal ridge absent dorsally. Pronotum outline hexagonal. Elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence. Protibiae without sexual dimorphism, both males and females without laterodorsal row of thick spines. Aedeagus as in
<figureCitation id="D69F03A68622D835D9ECFCE4BD1AFC94" box="[399,545,854,878]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="31" pageNumber="32">Figure 10L–N</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238622D835D812FC21BEC9FBBD" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835D812FC21BFD3FC50" box="[113,232,915,938]" italics="true" pageId="31" pageNumber="32">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8622D835D9FAFC24BE88FC50" box="[409,435,918,938]" count="1" pageId="31" pageNumber="32" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318622D835DAC8FC21BD85FC50" box="[683,702,915,938]" italics="true" pageId="31" pageNumber="32">N</emphasis>
= 6]: FBL = 3.95–4.21 (4.1); TL = 7.92–8 (7.95); HW = 1–1.11 (1.05); HL = 1.13–1.25 (1.19); HW/HL = 0.83–0.92 (0.88); PW = 1.15–1.24 (1.21); PL = 1.15–1.25 (1.21); PW/ PL = 0.95–1.03 (1); EW = 1.76–1.9 (1.81); EL = 1.67–1.71 (1.69); PW/HW = 1.11–1.21 (1.15).
</paragraph>
<paragraph id="4E1B1F238622D835D8EEFBFDBE98FAF9" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">
Measurements
<specimenCount id="58A2D4AA8622D835D94BFBE0BE79FB9C" box="[296,322,1106,1126]" count="1" pageId="31" pageNumber="32" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318622D835DA41FBFDBD0EFB9C" box="[546,565,1103,1126]" italics="true" pageId="31" pageNumber="32">N</emphasis>
= 3]: FBL = 4.47– 4.76 (4.59); TL = 7.32–8.56 (7.76); HW = 1.16–1.21 (1.18); HL = 1.3–1.36 (1.33); HW/HL = 0.87–0.9 (0.89); PW = 1.33–1.5 (1.4); PL = 1.31–1.44 (1.37); PW/PL = 1.01– 1.04 (1.02); EW = 1.98–2.13 (2.06); EL = 1.83–1.96 (1.9); PW/HW = 1.15–1.24 (1.18).
</paragraph>
<paragraph id="4E1B1F238622D835D8EEFAB9BFFBFA25" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">Head and pronotum dark brown to black with metallic blue overtones; antennae, mouthparts, pronotum at margins and legs yellowish-brown; elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence, sometimes darker around mesocutellum and at suture; abdominal tergites and sternites brown-black or dark reddish-brown with pale posterior margin.</paragraph>
<paragraph id="4E1B1F238622D835D8EEFA54BCA5FE1B" blockId="31.[113,765,915,1973]" lastBlockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
Head elongate; dorsally and ventrally with iridescence and with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with moderate setiferous punctures of medium and small-size. Eyes small (EYL/HL = x = 0.44), from 1.18 to 1.46 times as long as temples (in lateral view) in males and
<quantity id="895CB2C68622D835D8C2F910BE00F940" box="[161,315,1698,1722]" metricMagnitude="-2" metricUnit="m" metricValue="2.3876" metricValueMax="2.6416" metricValueMin="2.1336" pageId="31" pageNumber="32" unit="in" value="0.94" valueMax="1.04" valueMin="0.84">0.84 to 1.04 in</quantity>
females; distance between eyes about 1.13 times as long as length of eye in males and 1.44 times in females. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.52 to 1.75 times as long as a
<quantity id="895CB2C68622D835DAAAF8ADBDC0F8CD" box="[713,763,1823,1847]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="31" pageNumber="32" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68622D835D970F88DBEACF8AD" box="[275,407,1855,1879]" metricMagnitude="-2" metricUnit="m" metricValue="3.3401" metricValueMax="3.5559999999999996" metricValueMin="3.1242" pageId="31" pageNumber="32" unit="in" value="1.315" valueMax="1.4" valueMin="1.23">1.23 to 1.4 in</quantity>
females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, ending distinctly far from gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
</paragraph>
<paragraph id="4E1B1F238622D835DB26FE5BBB8AFD46" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">Pronotum about as wide as long, convex, hexagonal, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or even up to six times as long as, diameter of punctures. Mesosternum with six macrosetae arranged in a row medially. Metasternum with several macrosetae.</paragraph>
<paragraph id="4E1B1F238622D835DB26FD76BAA1FC62" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 1–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
<paragraph id="4E1B1F238622D835DB26FC2DBB02FAEA" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina usually evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with U-shaped emargination medially (
<figureCitation id="D69F03A68622D835DC9EFB8EBA6FFBAE" box="[1277,1364,1084,1108]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="31" pageNumber="32">Fig. 14E</figureCitation>
); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X apically subtruncate (
<figureCitation id="D69F03A68622D835DB56FB0BBCB0FB2B" box="[821,907,1209,1233]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="31" pageNumber="32">Fig. 12E</figureCitation>
); female tergite X with apical margin distinctly subacute to acute (
<figureCitation id="D69F03A68622D835DBA5FB6BBB21FB0B" box="[966,1050,1241,1265]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="31" pageNumber="32">Fig. 13E</figureCitation>
); ovipositor, its second gonocoxite with three macrosetae medially.
</paragraph>
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Aedeagus as in
<figureCitation id="D69F03A68622D835DB91FAA5BBB0FAD5" box="[1010,1163,1303,1327]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="31" pageNumber="32">Figure 10L–N</figureCitation>
; its total length 1.25–1.31. Median lobe rod-like, its apex in parameral view slightly V-shaped, emarginate, internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a round apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically.
</paragraph>
<paragraph id="4E1B1F238622D835DB49F9BFBCF3F905" blockId="31.[810,1460,1549,1792]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835DB49F9BFBA21F9DF" box="[810,1306,1549,1573]" italics="true" pageId="31" pageNumber="32">
Distribution and habitat:
<taxonomicName id="89A464A08622D835DC47F9BFBA21F9DF" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1060,1306,1549,1573]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">Loncovilius edwardsianus</taxonomicName>
</emphasis>
is known from
<collectingCountry id="36B35FB38622D835DB49F99EBC5FF9BE" box="[810,868,1580,1604]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
, from the
<collectingRegion id="8C60D1C18622D835DBB0F99EBB07F9BE" box="[979,1084,1580,1604]" country="Chile" name="Araucania" pageId="31" pageNumber="32">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18622D835DC2BF99EBBB3F9BE" box="[1096,1160,1580,1604]" country="Chile" name="Aysen" pageId="31" pageNumber="32">Aysén</collectingRegion>
,
<collectingRegion id="8C60D1C18622D835DCF7F99EBBE1F9BE" box="[1172,1242,1580,1604]" country="Chile" name="Biobio" pageId="31" pageNumber="32">Biobío</collectingRegion>
,
<collectingRegion id="8C60D1C18622D835DC85F99EBA69F9BE" box="[1254,1362,1580,1604]" country="Chile" name="Los Lagos" pageId="31" pageNumber="32">Los Lagos</collectingRegion>
, and
<collectingRegion id="8C60D1C18622D835DDEEF99EBC62F999" country="Chile" name="Los Rios" pageId="31" pageNumber="32">Los Ríos</collectingRegion>
regions. In addition, it occurs in the westernmost portions of the
<collectingRegion id="8C60D1C18622D835DB0DF9D9BC85F979" box="[878,958,1643,1667]" country="Argentina" name="Chubut" pageId="31" pageNumber="32">Chubut</collectingRegion>
and
<collectingRegion id="8C60D1C18622D835DB91F9D9BB65F979" box="[1010,1118,1643,1667]" country="Argentina" name="Rio Negro" pageId="31" pageNumber="32">Río Negro</collectingRegion>
provinces of
<collectingCountry id="36B35FB38622D835DC89F9D9BA69F979" box="[1258,1362,1643,1667]" name="Argentina" pageId="31" pageNumber="32">Argentina</collectingCountry>
. Ŋe species occurs from
<quantity id="895CB2C68622D835DBBCF938BB46F95B" box="[991,1149,1674,1698]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="31" pageNumber="32" unit="m" value="700.0" valueMax="1200.0" valueMin="200.0">200 to 1200 m</quantity>
elevation; it is reported from wet sclerophyll and open
<taxonomicName id="89A464A08622D835DC53F91BBB9BF93B" box="[1072,1184,1705,1729]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
<emphasis id="7CD0C3318622D835DC53F91BBB9BF93B" box="[1072,1184,1705,1729]" italics="true" pageId="31" pageNumber="32">Nothofagus</emphasis>
</taxonomicName>
forests along the Valdivian temperate forest ecoregion. It has been collected by sissing leaf liưer and moss.
</paragraph>
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<emphasis id="7CD0C3318622D835DB49F893BBBAF8C3" box="[810,1153,1825,1849]" italics="true" pageId="31" pageNumber="32">
Remarks:
<taxonomicName id="89A464A08622D835DBEDF893BBBAF8C3" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[910,1153,1825,1849]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">Loncovilius edwardsianus</taxonomicName>
</emphasis>
was described under the preoccupied name
<taxonomicName id="89A464A08622D835DBA5F8F3BB56F8A3" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[966,1133,1857,1881]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318622D835DBA5F8F3BB56F8A3" box="[966,1133,1857,1881]" italics="true" pageId="31" pageNumber="32">Quedius edwardsi</emphasis>
</taxonomicName>
based on an unspecified number of
<typeStatus id="911FA1818622D835DB22F8D2BCA1F882" box="[833,922,1888,1912]" pageId="31" pageNumber="32" type="syntype">syntypes</typeStatus>
from Casa Pangue in Llanquihue Province in Southern
<collectingCountry id="36B35FB38622D835DB4AF8CDBC59F86D" box="[809,866,1919,1943]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
. Korge (1963) replaced the name with
<taxonomicName id="89A464A08622D835DC84F832BAB8F86D" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1255,1411,1919,1943]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318622D835DC84F832BAB8F86D" box="[1255,1411,1919,1943]" italics="true" pageId="31" pageNumber="32">Q. edwardsianus</emphasis>
</taxonomicName>
and, interestingly, this species was treated in the genus
<taxonomicName id="89A464A08622D835DD22F82DBAABF84D" box="[1345,1424,1951,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318622D835DD22F82DBAABF84D" box="[1345,1424,1951,1975]" italics="true" pageId="31" pageNumber="32">Quedius</emphasis>
</taxonomicName>
by Coiffait and Sáiz (1966, 1968) who neither studied the type material nor reported any new material. Presumably, all mentioned authors knew this species only from literature because even Sáiz (1971) did not mention it in the fauna of
<collectingCountry id="36B35FB3861DD80ADA44FF5CBD5BFEFC" box="[551,608,238,262]" name="Chile" pageId="32" pageNumber="33">Chile</collectingCountry>
. Ŋe species was transferred to the genus
<taxonomicName id="89A464A0861DD80AD908FEBFBEEDFEDF" box="[363,470,269,293]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C331861DD80AD908FEBFBEEDFEDF" box="[363,470,269,293]" italics="true" pageId="32" pageNumber="33">Loncovilius</emphasis>
</taxonomicName>
much later by Newton (2017). To fix the identity of the species, a male from the Bernhauer’s collection at the FMNH, which is clearly a
<typeStatus id="911FA181861DD80ADA60FEFEBD6FFE9E" box="[515,596,332,356]" pageId="32" pageNumber="33" type="syntype">syntype</typeStatus>
, is here designated as a
<typeStatus id="911FA181861DD80AD8CBFED9BE3EFE79" box="[168,261,363,387]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
of
<emphasis id="7CD0C331861DD80AD947FEDEBEF8FE79" box="[292,451,363,387]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="89A464A0861DD80AD947FEDEBE85FE79" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[292,446,363,387]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="edwardsianus">L. edwardsianus</taxonomicName>
.
</emphasis>
</paragraph>
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<emphasis id="7CD0C3318624D833D897F91FBD43F93D" bold="true" box="[244,632,1709,1736]" pageId="25" pageNumber="26">
(
<figureCitation id="D69F03A68624D833D89CF91FBE77F932" box="[255,332,1709,1736]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." figureDoi="http://doi.org/10.5281/zenodo.14443961" httpUri="https://zenodo.org/record/14443961/files/figure.png" pageId="25" pageNumber="26">Figs 5B</figureCitation>
,
<figureCitation id="D69F03A68624D833D935F91FBE4CF93D" box="[342,375,1709,1735]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." figureDoi="http://doi.org/10.5281/zenodo.14443965" httpUri="https://zenodo.org/record/14443965/files/figure.png" pageId="25" pageNumber="26">7C</figureCitation>
,
<figureCitation id="D69F03A68624D833D9E2F91FBEF6F932" box="[385,461,1709,1736]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="25" pageNumber="26">10D–F</figureCitation>
,
<figureCitation id="D69F03A68624D833D9B5F91FBD39F93D" box="[470,514,1709,1735]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="25" pageNumber="26">12B</figureCitation>
,
<figureCitation id="D69F03A68624D833DA6FF91FBD03F93D" box="[524,568,1709,1735]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="25" pageNumber="26">13B</figureCitation>
,
<figureCitation id="D69F03A68624D833DA21F91FBD57F93D" box="[578,620,1709,1735]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="25" pageNumber="26">14B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238624D833D812F953BEE4F8E2" blockId="25.[113,762,1761,1816]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833D812F953BE6DF903" box="[113,342,1761,1785]" italics="true" pageId="25" pageNumber="26">Zoobank registration:</emphasis>
<uri id="3A3513218624D833D913F953BEE0F8E2" pageId="25" pageNumber="26">
urn:lsid:zoobank.org:act:
<uuid id="3A0225F68624D833DA0DF953BEE0F8E2" pageId="25" pageNumber="26">4DCCE49B-5875-4A73-9DA6-203BC4455E20</uuid>
</uri>
.
</paragraph>
<paragraph id="4E1B1F238624D833D812F889BA48FDDA" blockId="25.[113,763,1850,1999]" lastBlockId="25.[809,1460,144,544]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833D812F889BE32F8A8" box="[113,265,1850,1874]" italics="true" pageId="25" pageNumber="26">Type material.</emphasis>
<materialsCitation id="FECC157E8624D833D97BF888BDC9F86B" collectingDate="1958-12-04" collectorName="Wellington Is. Pto. Eden" country="Chile" location="Fr Kuschel" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<emphasis id="7CD0C3318624D833D97BF888BE43F8A8" box="[280,376,1850,1874]" italics="true" pageId="25" pageNumber="26">
<typeStatus id="911FA1818624D833D97BF888BE48F8A8" box="[280,371,1850,1874]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
:
</emphasis>
male, mounted, with genitalia in a separate microvial, with labels as follows:
<collectorName id="E3517AF58624D833DA07F8EBBFE5F86B" pageId="25" pageNumber="26">Wellington Is. Pto. Eden</collectorName>
[-49.11, -74.40]
<date id="3A1A39E38624D833D9C6F8CABD34F86A" box="[421,527,1912,1936]" pageId="25" pageNumber="26" value="1958-12-04">
<collectingDate id="2A5EC00B8624D833D9C6F8CABD34F86A" box="[421,527,1912,1936]" pageId="25" pageNumber="26" value="1958-12-04">4.XII1958</collectingDate>
</date>
/
<collectingCountry id="36B35FB38624D833DA74F8CABD52F86A" box="[535,617,1912,1936]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
<location id="4B7B49F88624D833DA14F8CBBDC9F86B" LSID="urn:lsid:plazi:treatment:C60DAE358624D830D979F9C9BE3BF9C5:4B7B49F88624D833DA14F8CBBDC9F86B" box="[631,754,1913,1937]" country="Chile" name="Fr Kuschel" pageId="25" pageNumber="26">Fr Kuschel</location>
</materialsCitation>
/
<typeStatus id="911FA1818624D833D812F82ABFC7F84A" box="[113,252,1944,1968]" pageId="25" pageNumber="26" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="89A464A08624D833D96EF82ABECDF84A" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[269,502,1944,1968]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="7CD0C3318624D833D96EF82ABECDF84A" box="[269,502,1944,1968]" italics="true" pageId="25" pageNumber="26">Loncovilius hammondi</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM.
<materialsCitation id="FECC157E8624D833DB1FFF23BA54FDDA" collectingDate="1958-11-10" collectingDateMax="1982-12-29" collectingDateMin="1958-11-10" collectorName="Wellington Is. Pto. Eden &amp; Chiloe S. Pedro &amp; Fr Kuschel &amp; Qued. &amp; Loncovilius' &amp; Cautin Pr. Volcan Villarrica" country="Chile" county="Flowers of Perneuya" elevation="1120" latitude="-49.0" location="Puerto Eden" municipality="Wellington" pageId="25" pageNumber="26" specimenCount="18" specimenCount-female="10" specimenCount-male="8" typeStatus="paratype">
<emphasis id="7CD0C3318624D833DB1FFF23BCD8FF52" box="[892,995,145,168]" italics="true" pageId="25" pageNumber="26">
<typeStatus id="911FA1818624D833DB1FFF23BCE5FF52" box="[892,990,145,168]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectorName id="E3517AF58624D833DB93FF22BBC6FF52" box="[1008,1277,144,168]" pageId="25" pageNumber="26">Wellington Is. Pto. Eden</collectorName>
[-49.11, -74.40]
<date id="3A1A39E38624D833DB49FF1DBCA2FF3D" box="[810,921,175,199]" pageId="25" pageNumber="26" value="1958-12-04">
<collectingDate id="2A5EC00B8624D833DB49FF1DBCA2FF3D" box="[810,921,175,199]" pageId="25" pageNumber="26" value="1958-12-04">4.XII.1958</collectingDate>
</date>
/
<collectingCountry id="36B35FB38624D833DBC2FF1DBCCFFF3D" box="[929,1012,175,199]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
Fr Kuschel [
<specimenCount id="58A2D4AA8624D833DC1DFF1DBB93FF3C" box="[1150,1192,175,199]" count="4" pageId="25" pageNumber="26" type="male">4♂</specimenCount>
<specimenCount id="58A2D4AA8624D833DCD3FF1DBBECFF3C" box="[1200,1239,175,199]" count="8" pageId="25" pageNumber="26" type="female">8♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA8624D833DD53FF1DBA6CFF3C" box="[1328,1367,175,199]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
NHMD ex. NHM;
<specimenCount id="58A2D4AA8624D833DBFCFF7DBCFDFF1C" box="[927,966,207,231]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
CZUG ex. NHM];
<taxonomicName id="89A464A08624D833DCFDFF7DBA2CFF1D" box="[1182,1303,207,231]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
forest/
<collectingCountry id="36B35FB38624D833DD06FF7DBA88FF1D" box="[1381,1459,207,231]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
HE6.
<location id="4B7B49F88624D833DB3DFF5CBCDEFEFC" LSID="urn:lsid:plazi:treatment:C60DAE358624D830D979F9C9BE3BF9C5:4B7B49F88624D833DB3DFF5CBCDEFEFC" box="[862,997,238,262]" country="Chile" county="Flowers of Perneuya" latitude="-49.0" municipality="Wellington" name="Puerto Eden" pageId="25" pageNumber="26">Puerto Eden</location>
Isla
<collectingMunicipality id="AE7F85598624D833DC7AFF5CBBB0FEFC" box="[1049,1163,238,262]" pageId="25" pageNumber="26">Wellington</collectingMunicipality>
[-49.11, -74.40]
<geoCoordinate id="2B9079E48624D833DD21FF5CBA41FEFC" box="[1346,1402,238,262]" degrees="49" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="55555" value="-49.0">49° S</geoCoordinate>
I200 ss.
<date id="3A1A39E38624D833DB26FEBFBC91FEDF" box="[837,938,269,293]" pageId="25" pageNumber="26" value="1958-12-02">
<collectingDate id="2A5EC00B8624D833DB26FEBFBC91FEDF" box="[837,938,269,293]" pageId="25" pageNumber="26" value="1958-12-02">2.xii.1958</collectingDate>
</date>
[
<specimenCount id="58A2D4AA8624D833DBD8FEBFBCD9FEDE" box="[955,994,269,293]" count="1" pageId="25" pageNumber="26" type="female">1♀</specimenCount>
MNNC ex. NHM];
<collectorName id="E3517AF58624D833DCD9FEBFBA67FEDF" box="[1210,1372,269,293]" pageId="25" pageNumber="26">Chiloé S. Pedro</collectorName>
[-43.32, -73.72]
<date id="3A1A39E38624D833DBE2FE9EBCC8FEBE" box="[897,1011,300,324]" pageId="25" pageNumber="26" value="1958-11-10">
<collectingDate id="2A5EC00B8624D833DBE2FE9EBCC8FEBE" box="[897,1011,300,324]" pageId="25" pageNumber="26" value="1958-11-10">10.XI.1958</collectingDate>
</date>
/
<collectingCountry id="36B35FB38624D833DB98FE9EBB74FEBE" box="[1019,1103,300,324]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
<collectorName id="E3517AF58624D833DC3AFE9FBBF4FEBF" box="[1113,1231,301,325]" pageId="25" pageNumber="26">Fr Kuschel</collectorName>
/
<collectingCounty id="A77A67AF8624D833DCB4FE9FBA8FFEBF" box="[1239,1460,301,325]" pageId="25" pageNumber="26">Flowers of Perneưya</collectingCounty>
mucronata/
<collectorName id="E3517AF58624D833DBC4FEFEBCDDFE9E" box="[935,998,332,356]" pageId="25" pageNumber="26">Qued.</collectorName>
sp. 8/‘
<collectorName id="E3517AF58624D833DC40FEFEBB91FE9E" box="[1059,1194,332,356]" pageId="25" pageNumber="26">Loncovilius’</collectorName>
<taxonomicName id="89A464A08624D833DCCCFEFEBCA7FE79" authority="Scheerp. P. M. Hammond" class="Insecta" family="Staphylinidae" genus="Loncovilius" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">cf germaini Scheerp. P.M. Hammond</taxonomicName>
det. 1980 [
<specimenCount id="58A2D4AA8624D833DC75FED9BB05FE78" box="[1046,1086,363,387]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
NHM];
<collectingCountry id="36B35FB38624D833DCFDFED9BBCAFE79" box="[1182,1265,363,387]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
:
<collectorName id="E3517AF58624D833DC9FFED9BCB3FE59" pageId="25" pageNumber="26">Cautín Pr. Volcán Villarrica</collectorName>
[-39.36, -71.94],
<quantity id="895CB2C68624D833DC5BFE39BBB9FE59" box="[1080,1154,395,419]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="25" pageNumber="26" unit="m" value="1120.0">
<elevation id="C589F8108624D833DC5BFE39BBB9FE59" box="[1080,1154,395,419]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="25" pageNumber="26" unit="m" value="1120.0">1120m</elevation>
</quantity>
, site 654
<date id="3A1A39E38624D833DCBDFE38BA49FE58" box="[1246,1394,394,418]" pageId="25" pageNumber="26" value="1982-12-15" valueMax="1982-12-29" valueMin="1982-12-15">
<collectingDate id="2A5EC00B8624D833DCBDFE38BA49FE58" box="[1246,1394,394,418]" pageId="25" pageNumber="26" value="1982-12-15" valueMax="1982-12-29" valueMin="1982-12-15">15-29.xii.1982</collectingDate>
</date>
Noth. Dombeyi-Saxe-gothea with
<taxonomicName id="89A464A08624D833DC2DFE18BA92FE38" authority="A. Newton and M. Nayer" authorityName="A. Newton and M. Nayer" box="[1102,1449,426,450]" class="Magnoliopsida" family="Winteraceae" genus="Drimys" kingdom="Plantae" order="Canellales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Drimys A.Newton and M.Ŋayer</taxonomicName>
/
<collectingMethod id="97E567348624D833DB49FE7BBC89FE1B" box="[810,946,457,481]" pageId="25" pageNumber="26">window trap</collectingMethod>
[
<specimenCount id="58A2D4AA8624D833DBABFE7BBCD4FE1A" box="[968,1007,457,481]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8624D833DB98FE7BBB19FE1A" box="[1019,1058,457,481]" count="1" pageId="25" pageNumber="26" type="female">1♀</specimenCount>
FMNH]. All
<typeStatus id="911FA1818624D833DCA1FE78BA1EFE18" box="[1218,1317,458,482]" pageId="25" pageNumber="26" type="paratype">paratypes</typeStatus>
are supplied with the label: PAÞTYPE
<taxonomicName id="89A464A08624D833DC30FE5BBA0FFDFB" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[1107,1332,489,513]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="7CD0C3318624D833DC30FE5BBA0FFDFB" box="[1107,1332,489,513]" italics="true" pageId="25" pageNumber="26">Loncovilius hammondi</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
</materialsCitation>
.
</paragraph>
<paragraph id="4E1B1F238624D833DB49FDF0BA4EFCCE" blockId="25.[810,1460,577,821]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833DB49FDF0BCABFDA3" box="[810,912,578,601]" italics="true" pageId="25" pageNumber="26">Diagnosis:</emphasis>
Head dorsally with microsculpture of transverse waves; gula with isodiametric microsculpture in the middle; females with a11 distinctly longer than a10 (a11:a10 ratio&gt;1.5). Elytra reddish-brown, disc with coarse non-setiferous punctures and only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII with medial apical emargination in both sexes. Aedeagus as in
<figureCitation id="D69F03A68624D833DC83FCAFBA54FCCF" box="[1248,1391,797,821]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="25" pageNumber="26">Figure 10D–F</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238624D833DB49FCE5BBA5FBF1" blockId="25.[809,1461,854,2006]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833DB49FCE5BC9AFC94" box="[810,929,855,878]" italics="true" pageId="25" pageNumber="26">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8624D833DC32FCEBBB50FC97" box="[1105,1131,857,877]" count="1" pageId="25" pageNumber="26" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318624D833DD07FCE5BA4CFC94" box="[1380,1399,855,878]" italics="true" pageId="25" pageNumber="26">N</emphasis>
= 6]: FBL = 3.5–3.91 (3.67); TL = 6.7–7.31 (7.01); HW = 1.06– 1.16 (1.11); HL = 0.94–1.02 (0.97); HW/HL = 1.12–1.16 (1.14); PW = 1.36–1.51 (1.43); PL = 1.16–1.26 (1.21); PW/ PL = 1.17–1.2 (1.19); EW = 1.65–1.82 (1.76); EL = 1.38–1.63 (1.49); PW/HW = 1.29–1.3 (1.29).
</paragraph>
<paragraph id="4E1B1F238624D833DB26FBA0BBA5FB3D" blockId="25.[809,1461,854,2006]" pageId="25" pageNumber="26">
Measurements
<specimenCount id="58A2D4AA8624D833DB82FBA7BCC0FBD3" box="[993,1019,1045,1065]" count="1" pageId="25" pageNumber="26" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318624D833DCB8FBA1BBD5FBD0" box="[1243,1262,1043,1066]" italics="true" pageId="25" pageNumber="26">N</emphasis>
= 6]: FBL = 3.88– 4.22 (4.04); TL = 7.06–8.88 (7.81); HW = 1.13–1.21 (1.18); HL = 0.98–1.07 (1.03); HW/HL = 1.13–1.19 (1.15); PW = 1.5–1.67 (1.58); PL = 1.23–1.37 (1.31); PW/ PL = 1.18–1.22 (1.20); EW = 1.9–2.3 (2.05); EL = 1.67–1.78 (1.7); PW/HW = 1.31–1.38 (1.33).
</paragraph>
<paragraph id="4E1B1F238624D833DB26FB7CBCEDFABE" blockId="25.[809,1461,854,2006]" pageId="25" pageNumber="26">Head black to brown-black; most of antennae, pronotum, mouthparts, elytra, legs, and abdomen reddish-brown; pronotum, apical antennomeres, tibiae, and disc of abdominal segments darker.</paragraph>
<paragraph id="4E1B1F238624D830DB26FAF9BFCFFEDF" blockId="25.[809,1461,854,2006]" lastBlockId="26.[127,779,144,1171]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL/HL = x = 0.47), from 1.3 to 1.8 times longer than temples (in lateral view); distance between eyes about 1.7 times as long as length of eye in males and 1.68 times in females. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 usually subequal in length; antennomeres 4 to 6 gradually reduced in length; antennomeres 7 to 10 subequal in length; antennomere 11 is from 1.53 to 1.69 times as long as antennomere
<quantity id="895CB2C68624D833DC3EF916BBABF946" box="[1117,1168,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">10 in</quantity>
males and from 1.63 to 1.71 times in females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setiferous punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical.
</paragraph>
<paragraph id="4E1B1F238627D830D8FFFE9FBC3BFDFB" blockId="26.[127,779,144,1171]" pageId="26" pageNumber="27">Pronotum transverse, convex, evenly curved, with one PPDS and with an APP, without SLSP, with several coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setiferous punctate; disc with only a few rows of setiferous punctures and with several coarse non-setiferous punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F238627D830D8FFFDBABFF4FD01" blockId="26.[127,779,144,1171]" pageId="26" pageNumber="27">Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate.</paragraph>
<paragraph id="4E1B1F238627D830D8FFFCB1BE60FB89" blockId="26.[127,779,144,1171]" pageId="26" pageNumber="27">
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternum III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flaưened only in males. Male sternite VIII with a broad V-shaped emargination medially (
<figureCitation id="D69F03A68627D830DA25FC2DBD98FC4D" box="[582,675,927,951]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="26" pageNumber="27">Fig. 14B</figureCitation>
); female sternite VIII with a U-shaped emargination medially; male sternite IX with an obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (
<figureCitation id="D69F03A68627D830D8D9FBAFBE28FBCF" box="[186,275,1053,1077]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="26" pageNumber="27">Fig. 12B</figureCitation>
); female tergite X truncate to slightly arcuate medially (
<figureCitation id="D69F03A68627D830D888FB8EBE7AFBAE" box="[235,321,1084,1108]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="26" pageNumber="27">Fig. 13B</figureCitation>
); ovipositor, its second gonocoxite without macrosetae medially.
</paragraph>
<paragraph id="4E1B1F238627D830D8FFFBC9BD83FB69" blockId="26.[127,779,144,1171]" box="[156,696,1147,1171]" pageId="26" pageNumber="27">
Aedeagus. As in
<figureCitation id="D69F03A68627D830D921FBC9BEE8FB69" box="[322,467,1147,1171]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="26" pageNumber="27">Figure 10D–F</figureCitation>
; its total length ~1.21.
</paragraph>
<paragraph id="4E1B1F238627D830D8E2FB06BD6DFAD0" blockId="26.[128,778,1204,1322]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830D8E2FB06BFCBFB36" box="[129,240,1204,1228]" italics="true" pageId="26" pageNumber="27">Etymology:</emphasis>
Ŋe species epithet isa patronym in recognition of the late Peter. M. Hammond, a prominent coleopterist who also conducted some work on this genus, which was unpublished but which we were able to trace through his labels.
</paragraph>
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<emphasis id="7CD0C3318627D830D8E2FAF9BDABFA99" box="[129,656,1355,1379]" italics="true" pageId="26" pageNumber="27">
Distribution and habitat:
<taxonomicName id="89A464A08627D830D9C0FAF9BDABFA99" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[419,656,1355,1379]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hammondi">Loncovilius hammondi</taxonomicName>
</emphasis>
is known only from
<collectingCountry id="36B35FB38627D830D895FAD9BE0AFA79" box="[246,305,1387,1411]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
, from the
<collectingRegion id="8C60D1C18627D830D9CBFAD9BD2FFA79" box="[424,532,1387,1411]" country="Chile" name="Araucania" pageId="26" pageNumber="27">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18627D830DA41FAD9BDA8FA79" box="[546,659,1387,1411]" country="Chile" name="Los Lagos" pageId="26" pageNumber="27">Los Lagos</collectingRegion>
, as well as from
<collectingRegion id="8C60D1C18627D830D8A0FA38BE03FA58" box="[195,312,1418,1442]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="26" pageNumber="27">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38627D830D9BCFA38BD70FA58" box="[479,587,1418,1442]" name="Antarctica" pageId="26" pageNumber="27">Antarctica</collectingCountry>
regions. Known distribution consists of three widely separated areas, two of them confined to the Valdivian temperate forest and one to the Magellanic subpolar forest. It has been collected using window traps, and directly from flowers of
<taxonomicName id="89A464A08627D830DA58F9BABC32F9E5" box="[571,777,1544,1567]" class="Magnoliopsida" family="Campanulaceae" genus="Pernetya" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Asterales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="mucronata">
<emphasis id="7CD0C3318627D830DA58F9BABC32F9E5" box="[571,777,1544,1567]" italics="true" pageId="26" pageNumber="27">Pernetya mucronata</emphasis>
</taxonomicName>
(
<taxonomicName id="89A464A08627D830D8EBF995BFCEF9C5" box="[136,245,1575,1599]" class="Magnoliopsida" family="Ericaceae" kingdom="Plantae" order="Ericales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="family">Ericaceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="7CD0C3318625D832D893FC49BEF9FBEC" bold="true" box="[240,450,1019,1046]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
(Scheerpeltz, 1933)
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<emphasis id="7CD0C3318625D832D963FB99BDB1FBBF" bold="true" box="[256,650,1067,1093]" pageId="24" pageNumber="25">
(
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,
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,
<figureCitation id="D69F03A68625D832D9EEFB99BEE0FBBF" box="[397,475,1067,1093]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="24" pageNumber="25">10A–C</figureCitation>
,
<figureCitation id="D69F03A68625D832D987FB99BD29FBBF" box="[484,530,1067,1093]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="24" pageNumber="25">12A</figureCitation>
,
<figureCitation id="D69F03A68625D832DA7FFB99BD71FBBF" box="[540,586,1067,1093]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="24" pageNumber="25">13A</figureCitation>
,
<figureCitation id="D69F03A68625D832DA30FB99BD44FBBF" box="[595,639,1067,1093]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="24" pageNumber="25">14A</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238625D832D8E2FBD3BDF4FA69" blockId="24.[129,779,1121,1428]" pageId="24" pageNumber="25">
Germain 1903: 412 (as
<taxonomicName id="89A464A08625D832D914FBD3BD6DFB83" box="[375,598,1121,1145]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="7CD0C3318625D832D914FBD3BD6DFB83" box="[375,598,1121,1145]" italics="true" pageId="24" pageNumber="25">Philonthus cribripennis</emphasis>
</taxonomicName>
, original description); Bernhauer and Schubert 1914: 332 (as
<taxonomicName id="89A464A08625D832DA24FB33BC38FB63" authorityName="Bernhauer &amp; K. Schubert" authorityYear="1914" box="[583,771,1153,1177]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318625D832DA24FB33BC38FB63" box="[583,771,1153,1177]" italics="true" pageId="24" pageNumber="25">Philonthus chilensis</emphasis>
</taxonomicName>
, replacement name); Scheerpeltz 1933: 1344 (as
<taxonomicName id="89A464A08625D832DAC2FB12BFECFB2D" authorityName="Scheerpeltz" authorityYear="1933" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318625D832DAC2FB12BFECFB2D" italics="true" pageId="24" pageNumber="25">Philonthus germaini</emphasis>
</taxonomicName>
, replacement name); Coiffait and Sáiz 1966: 406 (as
<taxonomicName id="89A464A08625D832D8E2FB6DBEF5FB0D" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[129,462,1247,1271]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
<emphasis id="7CD0C3318625D832D8E2FB6DBFD4FB0D" box="[129,239,1247,1271]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318625D832D962FB52BE5EFB0D" box="[257,357,1248,1271]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
)
<emphasis id="7CD0C3318625D832D91BFB52BEF5FB0D" box="[376,462,1248,1271]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
</taxonomicName>
, characters); Coiffait and Sáiz 1968: 365 [as
<taxonomicName id="89A464A08625D832D974FB4CBD5AFAEC" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[279,609,1278,1302]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
<emphasis id="7CD0C3318625D832D974FB4CBEBEFAEC" box="[279,389,1278,1302]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318625D832D9F6FB4DBEC2FAEC" box="[405,505,1279,1302]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
)
<emphasis id="7CD0C3318625D832DA69FB4DBD5AFAEC" box="[522,609,1279,1302]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
</taxonomicName>
<emphasis id="7CD0C3318625D832DA01FB4DBD53FAEC" box="[610,616,1279,1302]" italics="true" pageId="24" pageNumber="25">;</emphasis>
checklist]; Sáiz 1971: 385 [as
<taxonomicName id="89A464A08625D832D978FAAFBD52FACF" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[283,617,1309,1334]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
<emphasis id="7CD0C3318625D832D978FAAFBEB2FACF" box="[283,393,1309,1333]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318625D832D9FFFAACBD3BFACF" box="[412,512,1310,1333]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
)
<emphasis id="7CD0C3318625D832DA70FAACBD52FACF" box="[531,617,1310,1333]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
</taxonomicName>
; listed without new data]; Herman 2001a: 26 (as
<taxonomicName id="89A464A08625D832D9B2FA8FBDA9FAAF" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[465,658,1341,1365]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318625D832D9B2FA8FBDA9FAAF" box="[465,658,1341,1365]" italics="true" pageId="24" pageNumber="25">Loncovilius chilensis</emphasis>
</taxonomicName>
, nomenclature); Herman 2001b: 3083 (as
<taxonomicName id="89A464A08625D832D9AAFAEEBDB0FA8E" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[457,651,1372,1396]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318625D832D9AAFAEEBDB0FA8E" box="[457,651,1372,1396]" italics="true" pageId="24" pageNumber="25">Loncovilius chilensis</emphasis>
</taxonomicName>
, catalogue); Newton 2017: 22 (as
<taxonomicName id="89A464A08625D832D93EFAC9BD1DFA69" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[349,550,1403,1427]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318625D832D93EFAC9BD1DFA69" box="[349,550,1403,1427]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
</taxonomicName>
, nomenclature).
</paragraph>
<paragraph id="4E1B1F238625D832D8E2FA02BDA9FA32" blockId="24.[129,658,1456,1480]" box="[129,658,1456,1480]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D8E2FA02BE7BFA32" box="[129,320,1456,1480]" italics="true" pageId="24" pageNumber="25">Material eoamined:</emphasis>
Supporting Information, File S7.
</paragraph>
<paragraph id="4E1B1F238625D832D8E2FA6CBC3FF94A" blockId="24.[128,778,1501,1713]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D8E2FA6CBFDEFA0F" box="[129,229,1502,1525]" italics="true" pageId="24" pageNumber="25">Diagnosis:</emphasis>
Head dorsally without microsculpture; gula without isodiametric microsculpture in the middle; females with a11 distinctly longer than a10 (a11:a10 ratio &lt;1.5). Elytra, including epipleura, evenly setose and with several coarse non-setiferous punctures. Protibiae sexually dimorphic, with laterodorsal row of thick spines only present in females. Tergite VIII with medial apical emargination only in females. Aedeagus as in
<figureCitation id="D69F03A68625D832DA0EF92BBDC5F94B" box="[621,766,1689,1713]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="24" pageNumber="25">Figure 10A–C</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238625D832D8E2F97BBECEF887" blockId="24.[129,779,1736,1980]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D8E2F97BBFCCF91A" box="[129,247,1737,1760]" italics="true" pageId="24" pageNumber="25">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8625D832D9CBF979BEF9F925" box="[424,450,1739,1759]" count="1" pageId="24" pageNumber="25" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318625D832DAD9F97BBDF6F91A" box="[698,717,1737,1760]" italics="true" pageId="24" pageNumber="25">N</emphasis>
= 6]: FBL = 2.9–3.44 (3.19); TL = 5.6–6.25 (6.01); HW = 0.87– 1.01 (0.94); HL = 0.76–0.86 (0.82); HW/HL = 1.12–1.2 (1.14); PW = 1.11–1.29 (1.21); PL = 0.97–1.18 (1.08); PW/ PL = 1.1–1.14 (1.12); EW = 1.4–1.64 (1.52); EL = 1.16–1.4 (1.3); PW/HW = 1.26–1.33 (1.29).
</paragraph>
<paragraph id="4E1B1F238625D832D8FFF837BB65FD57" blockId="24.[129,779,1736,1980]" lastBlockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">
Measurements
<specimenCount id="58A2D4AA8625D832D954F835BE6AF861" box="[311,337,1927,1947]" count="1" pageId="24" pageNumber="25" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318625D832DA52F837BD7FF866" box="[561,580,1925,1948]" italics="true" pageId="24" pageNumber="25">N</emphasis>
= 6]: FBL = 3.36– 3.87 (3.6); TL = 7.31–7.77 (7.54); HW = 0.96–1.12 (1.03); HL = 0.85–0.95 (0.91); HW/HL = 1.11–1.18 (1.13); PW = 1.25–1.48 (1.34); PL = 1.13–1.28 (1.2); PW/PL = 1.1– 1.16 (1.12); EW = 1.5–1.88 (1.68); EL = 1.38–1.64 (1.49); PW/HW = 1.26–1.33 (1.3).
</paragraph>
<paragraph id="4E1B1F238625D832DB36FD06BBADFCD0" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">Head black to brown-black; mouthparts, antennae, pronotum, legs, disc of elytra, and abdominal segments dark reddish-brown; epipleura, abdominal tergites, and sternites with posterior margin light reddish-brown.</paragraph>
<paragraph id="4E1B1F238625D832DB36FC83BA8EF9E0" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">
Head transverse; dorsally and ventrally glossy with few micropunctures, dorsally without microsculpture, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.46), from 1.4 to 1.86 times longer than the length of temples (in lateral view); distance between eyes about 1.68 times as long as length of the eye in males and 1.8 times in females. Antennal insertions situated closer to frontoclypeus than to eye; antennomeres 2 and 3 subequal in length; antennomeres 5 to 10 subequal in length; antennomere 11 from 1.53 to 1.69 times longer than antennomere
<quantity id="895CB2C68625D832DCE2FBD9BB8DFB79" box="[1153,1206,1131,1155]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="25" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68625D832DD02FBD9BC75FB58" metricMagnitude="-2" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="24" pageNumber="25" unit="in" value="1.55" valueMax="1.6" valueMin="1.5">1.5 to 1.6 in</quantity>
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and entire apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical.
</paragraph>
<paragraph id="4E1B1F238625D832DB36F990BAF8F92C" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">Pronotum transverse, convex, evenly curved, usually with one PPDS with additional paired punctures adjacent to AMRSP (APP), without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra evenly setose and with several coarse non-setiferous punctures.Mesosternumwithfourmacrosetaearrangedinarowmedially.</paragraph>
<paragraph id="4E1B1F238625D832DB36F96CBAF8F84B" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate. Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4withpaleadhesivesetae,withoutterminalplate.</paragraph>
<paragraph id="4E1B1F238624D833D8EEFF22BFE8FDDA" blockId="25.[113,763,144,576]" pageId="25" pageNumber="26">
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line (
<figureCitation id="D69F03A68624D833D81FFF7DBFF1FF1D" box="[124,202,207,231]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="25" pageNumber="26">Fig. 9A</figureCitation>
); in females tergite VIII emarginate medio-apically. Sternite III with basal transverse carina converging evenly at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with deep V-shaped emargination medially (
<figureCitation id="D69F03A68624D833D910FEFEBEF4FE9E" box="[371,463,332,356]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="25" pageNumber="26">Fig. 14A</figureCitation>
); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X subtruncate medially (
<figureCitation id="D69F03A68624D833DA5CFE18BDA1FE38" box="[575,666,426,450]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="25" pageNumber="26">Fig. 12A</figureCitation>
); female tergite X with U-shaped emargination medially (
<figureCitation id="D69F03A68624D833DAE8FE7BBDD3FE1B" box="[651,744,457,481]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="25" pageNumber="26">Fig. 13A</figureCitation>
); ovipositor, its second gonocoxite without macrosetae medially (
<figureCitation id="D69F03A68624D833D81FFDBABFF9FDDA" box="[124,194,520,544]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="25" pageNumber="26">Fig. 9F</figureCitation>
).
</paragraph>
<paragraph id="4E1B1F238624D833D8EEFD95BD8BFDC5" blockId="25.[113,763,144,576]" box="[141,688,551,576]" pageId="25" pageNumber="26">
Aedeagus. As in
<figureCitation id="D69F03A68624D833D950FD95BEFEFDC5" box="[307,453,551,575]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="25" pageNumber="26">Figure 10A–C</figureCitation>
; its total length ~ 1.21.
</paragraph>
<paragraph id="4E1B1F238624D833D812FDD3BFD7FCCE" blockId="25.[113,764,609,821]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833D812FDD3BD6DFD82" box="[113,598,609,633]" italics="true" pageId="25" pageNumber="26">
Distribution and habitat:
<taxonomicName id="89A464A08624D833D9E4FDD3BD6DFD82" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[391,598,609,633]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">Loncovilius germaini</taxonomicName>
</emphasis>
is known only from
<collectingCountry id="36B35FB38624D833D8C9FD32BFDFFD62" box="[170,228,640,664]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
, from the
<collectingRegion id="8C60D1C18624D833D933FD32BE81FD62" box="[336,442,640,664]" country="Chile" name="Araucania" pageId="25" pageNumber="26">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18624D833D9A6FD32BD3DFD62" box="[453,518,640,664]" country="Chile" name="Aysen" pageId="25" pageNumber="26">Aysén</collectingRegion>
, and
<collectingRegion id="8C60D1C18624D833DA5EFD32BD9CFD62" box="[573,679,640,664]" country="Chile" name="Los Lagos" pageId="25" pageNumber="26">Los Lagos</collectingRegion>
regions where it occurs in the eastern portion of the Valdivian temperate forest ecoregion, from
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of elevation. It has been collected by sissing leaf liưer and moss, also by using Malaise and window traps, as well as carrion and dung baited pitfall traps.
</paragraph>
<paragraph id="4E1B1F238624D833D812FCE4BFEDF9A4" blockId="25.[112,763,854,1630]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833D812FCE4BE86FC94" box="[113,445,854,878]" italics="true" pageId="25" pageNumber="26">
Remarks:
<taxonomicName id="89A464A08624D833D889FCE4BE86FC94" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[234,445,854,878]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">Loncovilius germaini</taxonomicName>
</emphasis>
was described as
<taxonomicName id="89A464A08624D833DAF2FCE4BFDEFC77" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="7CD0C3318624D833DAF2FCE4BFDEFC77" italics="true" pageId="25" pageNumber="26">Philonthus cribripennis</emphasis>
</taxonomicName>
based on an unspecified number of
<typeStatus id="911FA1818624D833DA2FFCC4BD9CFC74" box="[588,679,886,910]" pageId="25" pageNumber="26" type="syntype">syntypes</typeStatus>
from an uncertain type locality in
<collectingCountry id="36B35FB38624D833D915FC27BE8AFC57" box="[374,433,917,941]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
. Ŋis name was found to be preoccupied and replaced by another preoccupied name
<taxonomicName id="89A464A08624D833DAF3FC07BE96FC11" authority="Bernhauer and Schubert 1914" authorityName="Bernhauer and Schubert" authorityYear="1914" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318624D833DAF3FC07BDA5FC36" box="[656,670,949,972]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="7CD0C3318624D833DAC9FC06BDC1FC36" box="[682,762,948,972]" italics="true" pageId="25" pageNumber="26">chilensis</emphasis>
Bernhauer and Schubert 1914
</taxonomicName>
; the laưer was replaced again by
<taxonomicName id="89A464A08624D833D811FC41BE93FBF1" authority="Scheerpeltz, 1933" authorityName="Scheerpeltz" authorityYear="1933" box="[114,424,1011,1035]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318624D833D811FC41BFBBFBF0" box="[114,128,1011,1034]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="7CD0C3318624D833D8EDFC41BFDFFBF0" box="[142,228,1011,1034]" italics="true" pageId="25" pageNumber="26">germaini</emphasis>
Scheerpeltz, 1933
</taxonomicName>
. Coiffiat and Sáiz (1966, 1968) and Sáiz (1971) treated
<taxonomicName id="89A464A08624D833D91BFBA1BED0FBD0" authorityName="Scheerpeltz" authorityYear="1933" box="[376,491,1042,1066]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318624D833D91BFBA1BEBCFBD0" box="[376,391,1043,1066]" italics="true" pageId="25" pageNumber="26">L</emphasis>
.
<emphasis id="7CD0C3318624D833D9F6FBA1BED0FBD0" box="[405,491,1043,1066]" italics="true" pageId="25" pageNumber="26">germaini</emphasis>
</taxonomicName>
as a member of the subgenus
<taxonomicName id="89A464A08624D833D8D7FB80BE22FBB3" box="[180,281,1074,1097]" pageId="25" pageNumber="26" rank="subGenus" subGenus="Lienturius">
<emphasis id="7CD0C3318624D833D8D7FB80BE22FBB3" box="[180,281,1074,1097]" italics="true" pageId="25" pageNumber="26">Lienturius</emphasis>
</taxonomicName>
of the genus
<taxonomicName id="89A464A08624D833D9C6FB83BD2FFBB3" authorityName="Germain" authorityYear="1903" box="[421,532,1073,1097]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318624D833D9C6FB83BD2FFBB3" box="[421,532,1073,1097]" italics="true" pageId="25" pageNumber="26">Loncovilius</emphasis>
</taxonomicName>
without adding much data about morphology or distribution of this species, for which even the aedeagus hitherto remained not illustrated. Coiffait and Sáiz (1966) mentioned that the species was known from a female
<typeStatus id="911FA1818624D833D8AEFB1DBE11FB3D" box="[205,298,1199,1223]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
kept in the collection of the Natural History Museum of
<collectingCountry id="36B35FB38624D833D88DFB7CBE12FB1C" box="[238,297,1230,1254]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
. At the same time, Camousseight (1980) did not list this species among the type material in Germain’s collection in this museum. We did not find any material that could be interpreted as types of
<emphasis id="7CD0C3318624D833D9EDFA9FBD33FABE" box="[398,520,1325,1348]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="89A464A08624D833D9EDFA9FBD3FFABE" authorityName="Scheerpeltz" authorityYear="1933" box="[398,516,1325,1348]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">L. germaini</taxonomicName>
.
</emphasis>
Given the uncertainty about type material in the original description and the catalogue by Camousseight (1980), the type status of the female listed by Coiffait and Sáiz (1966) as ‘holotype’ must be verified in the future. For now, we have matched the identity of this species based on the description by Coiffait and Sáiz (1966). As explained by Newton (2017), Herman’s (2001a) interpretation of both names as replacements for only
<taxonomicName id="89A464A08624D833DA79F9BABD9DF9E5" box="[538,678,1543,1567]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="7CD0C3318624D833DA79F9BABD9DF9E5" box="[538,678,1543,1567]" italics="true" pageId="25" pageNumber="26">P. cribripennis</emphasis>
</taxonomicName>
and his resurrection of
<taxonomicName id="89A464A08624D833D973F995BE4CF9C5" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[272,375,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318624D833D973F995BE4CF9C5" box="[272,375,1575,1599]" italics="true" pageId="25" pageNumber="26">P. chilensis</emphasis>
</taxonomicName>
as the valid name for this species was incorrect.
</paragraph>
</subSubSection>
</treatment>
</document>

611
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<emphasis id="7CD0C3318626D831DBBFFEB6BB8AFEE5" bold="true" box="[988,1201,260,287]" italics="true" pageId="27" pageNumber="28">Loncovilius variabilis</emphasis>
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(
<figureCitation id="D69F03A68626D831DBC3FE86BCD4FEB4" box="[928,1007,308,334]" captionStart="Figure 1" captionStartId="1.[113,178,1870,1894]" captionTargetBox="[152,1420,144,1842]" captionTargetId="figure-6@1.[152,1420,144,1842]" captionTargetPageId="1" captionText="Figure 1. Habitats of Loncovilius. A, L. edwardsianus (Korge, 1963) live specimen in the Valdivian Temperate Rainforest in Chiloé, Los Lagos, Chile (Photograph: Matías Gargiulo, through iNaturalist). B, PN Puyehue, Chile, L. lividipennis was collected in mosses on the stones at river banks and under stones in accumulations along banks (photograph: Martin Fikáček). C, L. variabilis sp. nov. found in a flower of the Chiloé archipelago (photograph:Matías Gargiulo, through iNaturalist)." figureDoi="http://doi.org/10.5281/zenodo.14443947" httpUri="https://zenodo.org/record/14443947/files/figure.png" pageId="27" pageNumber="28">Figs 1C</figureCitation>
,
<figureCitation id="D69F03A68626D831DB99FE86BB20FEB4" box="[1018,1051,308,334]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." figureDoi="http://doi.org/10.5281/zenodo.14443961" httpUri="https://zenodo.org/record/14443961/files/figure.png" pageId="27" pageNumber="28">5D</figureCitation>
,
<figureCitation id="D69F03A68626D831DC46FE86BB79FEB4" box="[1061,1090,308,334]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="27" pageNumber="28">9B</figureCitation>
,
<figureCitation id="D69F03A68626D831DC2EFE86BBAFFEB4" box="[1101,1172,308,334]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="27" pageNumber="28">10I–K</figureCitation>
,
<figureCitation id="D69F03A68626D831DCFEFE86BBF7FEB4" box="[1181,1228,308,334]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="27" pageNumber="28">12D</figureCitation>
,
<figureCitation id="D69F03A68626D831DCB5FE86BA3FFEB4" box="[1238,1284,308,334]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="27" pageNumber="28">13D</figureCitation>
,
<figureCitation id="D69F03A68626D831DD6DFE86BA06FEB4" box="[1294,1341,308,334]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="27" pageNumber="28">14D</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238626D831DB49FED7BB90FE66" blockId="27.[810,1459,357,412]" pageId="27" pageNumber="28">
<emphasis id="7CD0C3318626D831DB49FED7BB2EFE87" box="[810,1045,357,381]" italics="true" pageId="27" pageNumber="28">Zoobank registration:</emphasis>
<uri id="3A3513218626D831DC57FED7BB93FE66" pageId="27" pageNumber="28">
urn:lsid:zoobank.org:act:
<uuid id="3A0225F68626D831DD53FED7BB93FE66" pageId="27" pageNumber="28">6008BBBE- 2EF2-4459-8ACD-5AA4E242AA4A</uuid>
</uri>
.
</paragraph>
<paragraph id="4E1B1F238626D836DB49FE0DBD7CF8DF" blockId="27.[809,1460,446,1974]" lastBlockId="28.[127,779,144,1829]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="7CD0C3318626D831DB49FE0DBB23FE2C" box="[810,1048,446,470]" italics="true" pageId="27" pageNumber="28">
Type material.
<typeStatus id="911FA1818626D831DBDAFE0CBB28FE2C" box="[953,1043,446,470]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
:
</emphasis>
male, pinned, with genitalia in a separate microvial, with labels as follows:
<collectingCountry id="36B35FB38626D831DC0DFE6FBBFBFE0F" box="[1134,1216,477,501]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DCAEFE6CBA02FE0C" box="[1229,1337,478,502]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Hualaihué] Palena Pr.: Austral Highway, km 62.9 (
<quantity id="895CB2C68626D831DCCBFE4FBBCBFDEF" box="[1192,1264,509,533]" metricMagnitude="3" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="28" unit="km" value="6.7">6.7 km</quantity>
S Contao turnoff),
<quantity id="895CB2C68626D831DB49FDAEBC5CFDCE" box="[810,871,540,564]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">230m</quantity>
,
<geoCoordinate id="2B9079E48626D831DB0CFDAEBCD5FDCE" box="[879,1006,538,564]" degrees="41" direction="south" minutes="51.155" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="-41.852585">
41°51.155
<emphasis id="7CD0C3318626D831DBB5FDA8BCE7FDC9" box="[982,988,538,563]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DB94FDAEBBBBFDC9" box="[1015,1152,538,564]" degrees="72" direction="west" minutes="42.175" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-72.70292">
72°42.175
<emphasis id="7CD0C3318626D831DC3DFDA8BB5FFDC9" box="[1118,1124,538,563]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
W
</geoCoordinate>
[-41.85, -72.70],
<date id="3A1A39E38626D831DD4EFDAEBA94FDCE" box="[1325,1455,540,564]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">23-25.i.1997</date>
, young secondary Valdivian rainforest;/beating flowers
<taxonomicName id="89A464A08626D831DD08FD89BCB9FD89" class="Magnoliopsida" family="Ericaceae" genus="Tepulia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="stipularis">
<emphasis id="7CD0C3318626D831DD08FD89BCB9FD89" italics="true" pageId="27" pageNumber="28">Tepulia stipularis</emphasis>
</taxonomicName>
(myrtaceae) [
<taxonomicName id="89A464A08626D831DC74FDE9BBB2FD89" box="[1047,1161,603,627]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
] A. Newton and M. Ŋayer FIELD MUSEUM NAT. HIST./
<typeStatus id="911FA1818626D831DCC1FDC8BA0AFD68" box="[1186,1329,634,658]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="89A464A08626D831DD25FDC8BCB9FD4B" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318626D831DD25FDC8BCB9FD4B" italics="true" pageId="27" pageNumber="28">Loncovilius variabilis</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
<emphasis id="7CD0C3318626D831DC8CFD0BBA6FFD2A" box="[1263,1364,697,720]" italics="true" pageId="27" pageNumber="28">
<typeStatus id="911FA1818626D831DC8CFD0BBA74FD2A" box="[1263,1359,697,720]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="36B35FB38626D831DD3DFD0BBA8BFD2B" box="[1374,1456,697,721]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DB53FD6ABCA7FD0A" box="[816,924,728,752]" country="Chile" name="Araucania" pageId="27" pageNumber="28">Araucanía</collectingRegion>
, Vilcún] Cautín Temuco, Cherquenco [~ -38.69, -72.00] I-II.1978 L. E. Peña leg. FIELD MUS. NAT. HIST. [
<specimenCount id="58A2D4AA8626D831DB51FCA5BC60FCD4" box="[818,859,791,815]" count="3" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DB07FCA5BCB0FCD4" box="[868,907,791,815]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38626D831DB9FFCA5BB75FCD5" box="[1020,1102,791,815]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: Reg. XI [
<collectingRegion id="8C60D1C18626D831DCDBFCA5BBC6FCD5" box="[1208,1277,791,815]" country="Chile" name="Aysen" pageId="27" pageNumber="28">Aysén</collectingRegion>
] Queulat N.P. [~ -44.38, -72.25] P.M. Hammond/Below base 1-10.xii.03 shrub flowers/NHM/[
<specimenCount id="58A2D4AA8626D831DBB4FCE7BCC7FC96" box="[983,1020,853,877]" count="5" pageId="27" pageNumber="28" type="male">5♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DC65FCE7BB16FC96" box="[1030,1069,853,877]" count="4" pageId="27" pageNumber="28" type="female">4♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA8626D831DCEBFCE7BB95FC96" box="[1160,1198,853,877]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DCDBFCE7BBE4FC96" box="[1208,1247,853,877]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
NHMD ex. NHM;
<specimenCount id="58A2D4AA8626D831DB49FCC7BC6AFC76" box="[810,849,885,909]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
CZUG ex. NHM]; [same data as above except:] Glacier Path 21.xi-12.xii.03/Malaise trap [
<specimenCount id="58A2D4AA8626D831DCF6FC26BB87FC51" box="[1173,1212,916,940]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
NHM]; [same data as above except:] Base area Beating [or] Base Camp Area 21.xi- 12.xii.03 [
<specimenCount id="58A2D4AA8626D831DBF6FC61BC85FC10" box="[917,958,979,1003]" count="8" pageId="27" pageNumber="28" type="male">8♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DBA9FC61BCCAFC10" box="[970,1009,979,1002]" count="7" pageId="27" pageNumber="28" type="female">7♀</specimenCount>
NHM];
<collectingCountry id="36B35FB38626D831DC3BFC61BB91FC11" box="[1112,1194,979,1003]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DCDCFC61BA0AFC11" box="[1215,1329,979,1003]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Hualaihué] Palena Pr.: Austral Highway, km 62.9 (
<quantity id="895CB2C68626D831DCCFFC40BBD4FBF0" box="[1196,1263,1010,1034]" metricMagnitude="3" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="28" unit="km" value="6.7">6.7km</quantity>
S Contao turnoff),
<quantity id="895CB2C68626D831DB48FBA3BC53FBD3" box="[811,872,1041,1065]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">230m</quantity>
,
<geoCoordinate id="2B9079E48626D831DB15FBA3BCDFFBD3" box="[886,996,1041,1065]" degrees="41" direction="south" minutes="51.15" orientation="latitude" pageId="27" pageNumber="28" precision="9" value="-41.8525">41°51.15’S</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DB90FBA3BB57FBD3" box="[1011,1132,1041,1065]" degrees="72" direction="west" minutes="42.17" orientation="longitude" pageId="27" pageNumber="28" precision="9" value="-72.702835">72°42.17’W</geoCoordinate>
[-41.85, -72.70],
<date id="3A1A39E38626D831DD4DFBA3BA8BFBD3" box="[1326,1456,1041,1065]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">23-25.i.1997</date>
, young secondary Valdivian rainforest;/beating flowers
<taxonomicName id="89A464A08626D831DD08FB83BCB9FB92" class="Magnoliopsida" family="Ericaceae" genus="Tepulia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="stipularis">
<emphasis id="7CD0C3318626D831DD08FB83BCB9FB92" italics="true" pageId="27" pageNumber="28">Tepulia stipularis</emphasis>
</taxonomicName>
(myrtaceae) [
<taxonomicName id="89A464A08626D831DC74FBE2BBB2FB92" box="[1047,1161,1104,1128]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
] A. Newton and M. Ŋayer FIELD MUSEUM NAT. HIST [
<specimenCount id="58A2D4AA8626D831DCF4FBDDBBFBFB7C" box="[1175,1216,1135,1159]" count="5" pageId="27" pageNumber="28" type="male">5♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38626D831DD5EFBDDBA4EFB7D" box="[1341,1397,1135,1159]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
[
<collectingRegion id="8C60D1C18626D831DDEAFBC2BC51FB5D" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
] Llanquihue Lago Chapo [-41.40, -72.52]
<date id="3A1A39E38626D831DD4DFB3DBA88FB5C" box="[1326,1459,1166,1191]" pageId="27" pageNumber="28" value="1985-02">febrero 1985</date>
leg. J. Zuñiga [
<specimenCount id="58A2D4AA8626D831DBDDFB1CBCDDFB3F" box="[958,998,1198,1222]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DB8FFB1CBB28FB3F" box="[1004,1043,1198,1222]" count="3" pageId="27" pageNumber="28" type="female">3♀</specimenCount>
FMNH]; HORNOHUINCO SW. Lago Chapo Lianquihue [
<collectingRegion id="8C60D1C18626D831DC64FB7CBB47FB1C" box="[1031,1148,1230,1254]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Llanquihue, -41.40, -72.52] 21,24-
<date id="3A1A39E38626D831DB0FFB5FBCF3FAFF" box="[876,968,1261,1285]" pageId="27" pageNumber="28" value="1972-12">Dic.1972</date>
Coll:L.E.Pena [
<specimenCount id="58A2D4AA8626D831DC1FFB5FBB98FAFE" box="[1148,1187,1261,1285]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DCD1FB5FBBE3FAFE" box="[1202,1240,1261,1285]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38626D831DD38FB5FBA8BFAFF" box="[1371,1456,1261,1285]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18626D831DB49FABEBCA3FADE" box="[810,920,1292,1316]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
] Rio Gol-Gol Osorno Prov. [~ -40.66, -72.18]/II-8- 57 Peña/
<collectingCountry id="36B35FB38626D831DBEFFA99BCE0FAB9" box="[908,987,1323,1347]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
[green label]/CNCI L2004-28/
<specimenCount id="58A2D4AA8626D831DD54FA9CBA6AFAB8" box="[1335,1361,1326,1346]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
/FMNH-INS 0000 024 263-265 [
<specimenCount id="58A2D4AA8626D831DC51FAF9BB60FA98" box="[1074,1115,1355,1379]" count="3" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
CNC];
<collectingCountry id="36B35FB38626D831DCDBFAF9BA31FA99" box="[1208,1290,1355,1379]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: Chiloé Pr.,
<collectingCountry id="36B35FB38626D831DDF3FAF9BA88FA99" box="[1424,1459,1355,1379]" name="Iceland" pageId="27" pageNumber="28">Isla</collectingCountry>
Chiloé, Compu Alto [-42.87, -73.70],
<date id="3A1A39E38626D831DCA2FAD8BA1AFA78" box="[1217,1313,1386,1410]" pageId="27" pageNumber="28" value="1998-01-18">18.i.1998</date>
, T. Cekalovic leg. FIELD MUS. NAT. HIST./FMNH-INS 0000 024 165-167 [
<specimenCount id="58A2D4AA8626D831DB56FA1BBC60FA3A" box="[821,859,1449,1473]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
FMNH;
<specimenCount id="58A2D4AA8626D831DBABFA1BBCD4FA3A" box="[968,1007,1449,1473]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DB98FA1BBB19FA3A" box="[1019,1058,1449,1473]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
MNNC ex. FMNH];
<collectingCountry id="36B35FB38626D831DD7CFA1BBA4AFA3B" box="[1311,1393,1449,1473]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: Prov. Chiloé:
<collectingCountry id="36B35FB38626D831DB1CFA7ABC99FA1A" box="[895,930,1480,1504]" name="Iceland" pageId="27" pageNumber="28">Isla</collectingCountry>
Chiloé, Compu Alto [-42.87, -73.70]
<date id="3A1A39E38626D831DD51FA7ABAACFA1A" box="[1330,1431,1480,1504]" pageId="27" pageNumber="28" value="1998-01-18">18-I-1998</date>
T. Cekalovic CHIL1C98 001/[Barcode] SM0722732 KUNHM-ENT [
<specimenCount id="58A2D4AA8626D831DB12F9B5BCA3F9E4" box="[881,920,1543,1567]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
SEMC]; [
<collectingCountry id="36B35FB38626D831DC64F9B5BB67F9E5" box="[1031,1116,1543,1567]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18626D831DC04F9B5BBEFF9E5" box="[1127,1236,1543,1567]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Puerto Monư] Prov. Llanquihue Correntoso [~ -41.45, -72.66]
<date id="3A1A39E38626D831DCBCF994BA42F9C4" box="[1247,1401,1574,1598]" pageId="27" pageNumber="28" value="1969-01-22">22-Enero-1969</date>
Coll: L.E.Pena [
<specimenCount id="58A2D4AA8626D831DBF7F9F7BC86F9A6" box="[916,957,1605,1629]" count="2" pageId="27" pageNumber="28" type="male">2♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38626D831DC49F9F7BB47F9A7" box="[1066,1148,1605,1629]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DCEFF9F4BBC1F9A4" box="[1164,1274,1606,1630]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Chonchi] Chiloé Pr.: Miraflores, road to (
<quantity id="895CB2C68626D831DC51F9D7BB4DF987" box="[1074,1142,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="km" value="0.6">0.6km</quantity>
W Hwy 5),
<quantity id="895CB2C68626D831DD6BF9D7BA7EF987" box="[1288,1349,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="28" unit="m" value="130.0">130m</quantity>
,
<geoCoordinate id="2B9079E48626D831DD37F9D7BC0CF966" degrees="42" direction="south" minutes="46.73" orientation="latitude" pageId="27" pageNumber="28" precision="9" value="-42.77883">
42°46.73
<emphasis id="7CD0C3318626D831DDCEF9D1BA88F986" box="[1453,1459,1635,1660]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="2B9079E48626D831DB2AF936BCE9F961" box="[841,978,1666,1692]" degrees="73" direction="west" minutes="47.71" orientation="longitude" pageId="27" pageNumber="28" precision="9" value="-73.795166">
73°47.71
<emphasis id="7CD0C3318626D831DBC0F930BC92F961" box="[931,937,1666,1691]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
W
</geoCoordinate>
[
<geoCoordinate id="2B9079E48626D831DB89F936BB70F966" box="[1002,1099,1668,1692]" degrees="42.7788" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="-42.7788">-42.7788</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DC3CF936BB85F966" box="[1119,1214,1668,1692]" degrees="73.7953" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-73.7953">-73.7953</geoCoordinate>
]
<date id="3A1A39E38626D831DCB5F936BA0EF966" box="[1238,1333,1668,1692]" pageId="27" pageNumber="28" value="1997-01-17">17.i.1997</date>
, secondary Valdivian rainforest:/beating flowering Cald-cluvia paniculata [
<taxonomicName id="89A464A08626D831DB56F971BB3CF921" box="[821,1031,1731,1755]" class="Magnoliopsida" family="Cunoniaceae" genus="Caldcluvia" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="paniculata">
<emphasis id="7CD0C3318626D831DB56F971BB3CF921" box="[821,1031,1731,1755]" italics="true" pageId="27" pageNumber="28">Caldcluvia paniculata</emphasis>
</taxonomicName>
] (
<taxonomicName id="89A464A08626D831DC7CF971BB92F921" box="[1055,1193,1731,1755]" class="Magnoliopsida" family="Cunoniaceae" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Cunoniaceae</taxonomicName>
) A.Newton and M.Ŋayer 994 FIELD MUS. NAT. HIST. [
<specimenCount id="58A2D4AA8626D831DCEBF950BB94F903" box="[1160,1199,1762,1786]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DCDAF950BBDBF903" box="[1209,1248,1762,1786]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38626D831DD38F950BA8BF900" box="[1371,1456,1762,1786]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18626D831DB49F8B0BCA2F8E0" box="[810,921,1794,1818]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Hualaihué] 11
<collectingCountry id="36B35FB38626D831DC2FF8B3BBBFF8E3" box="[1100,1156,1793,1817]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
X, peninsula Huequi under logs termas, Porcelana
<geoCoordinate id="2B9079E48626D831DC76F893BB99F8C3" box="[1045,1186,1823,1849]" degrees="42" direction="south" minutes="27" orientation="latitude" pageId="27" pageNumber="28" precision="1" seconds="30.8" value="-42.458557">
42°27
<emphasis id="7CD0C3318626D831DC32F8ADBB6CF8C2" box="[1105,1111,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
30.8
<emphasis id="7CD0C3318626D831DCE1F8ADBBB6F8C2" box="[1154,1165,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="2B9079E48626D831DCC9F893BA7AF8C2" box="[1194,1345,1823,1849]" degrees="72" direction="west" minutes="27" orientation="longitude" pageId="27" pageNumber="28" precision="1" seconds="15.0" value="-72.45416">
72°27
<emphasis id="7CD0C3318626D831DC86F8ADBBD0F8C2" box="[1253,1259,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
15.0
<emphasis id="7CD0C3318626D831DD74F8ADBA19F8C2" box="[1303,1314,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹʹ</emphasis>
W
</geoCoordinate>
[
<geoCoordinate id="2B9079E48626D831DD2CF892BA94F8C2" box="[1359,1455,1824,1848]" degrees="42.4585" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="-42.4585">-42.4585</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DB49F8F2BCB2F8A2" box="[810,905,1856,1880]" degrees="72.4541" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-72.4541">-72.4541</geoCoordinate>
]
<quantity id="895CB2C68626D831DBFBF8F2BCF4F8AD" box="[920,975,1856,1880]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="40.0">40 m</quantity>
R. Vila leg.
<date id="3A1A39E38626D831DC30F8F2BBF9F8A2" box="[1107,1218,1856,1880]" pageId="27" pageNumber="28" value="2014-12-13">13.xii.2014</date>
/
<taxonomicName id="89A464A08626D831DCA9F8F2BA53F8AD" authority="sp" authorityName="Sp" box="[1226,1384,1856,1880]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp</taxonomicName>
det. V. Assing
<date id="3A1A39E38626D831DB16F8EDBCF1F88D" box="[885,970,1887,1911]" bridgedPair="/" pageId="27" pageNumber="28" value="2015-05">2015/V.</date>
Assing Coll. J Jenkins Shaw det. 2016 [yellow label] [
<specimenCount id="58A2D4AA8626D831DB15F8CDBCA5F86C" box="[886,926,1919,1943]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
NHMW];
<collectingCountry id="36B35FB38626D831DC7AF8CDBB50F86D" box="[1049,1131,1919,1943]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DC1EF8CDBBD6F86D" box="[1149,1261,1919,1943]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Puyehue National Park] Osorno Prov. Parque Nac. Puyehue,
<quantity id="895CB2C68626D831DC95F82CBA0DF84C" box="[1270,1334,1950,1974]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="27" pageNumber="28" unit="km" value="4.1">4.1km</quantity>
E Anticura [-40.66, -72.12],
<quantity id="895CB2C68621D836D956FF22BE49FF52" box="[309,370,144,168]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="28" pageNumber="29" unit="m" value="430.0">430m</quantity>
, trap site 662
<date id="3A1A39E38621D836DA6DFF22BD9AFF52" box="[526,673,144,168]" pageId="28" pageNumber="29" value="1982-12-19" valueMax="1982-12-26" valueMin="1982-12-19">19-26.xii.1982</date>
Valdivian rainfor. A.Newton and M.Ŋayer/Screen- sweeping/
<taxonomicName id="89A464A08621D836DAECFF1DBFA6FF1C" authority="sp." authorityName="Sp." class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp.</taxonomicName>
7 det. A.Newton 1984/
<specimenCount id="58A2D4AA8621D836D9F6FF60BE94FF1C" box="[405,431,210,230]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
/FMNH-INS 0000 024 163 [
<specimenCount id="58A2D4AA8621D836DA81FF7DBC32FF1C" box="[738,777,207,231]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38621D836D885FF5CBE03FEFC" box="[230,312,238,262]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18621D836D92BFF5CBE8DFEFC" box="[328,438,238,262]" country="Chile" name="Los Lagos" pageId="28" pageNumber="29">Los Lagos</collectingRegion>
, Puyehue National Park] Osorno Prov., Puyehue Nat. Pk. Aguas Calientes [-40.73, -72.30],
<quantity id="895CB2C68621D836DAA9FEBFBC3CFEDF" box="[714,775,269,293]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="m" value="500.0">500m</quantity>
,
<date id="3A1A39E38621D836D8E2FE9EBFC5FEBE" box="[129,254,300,324]" pageId="28" pageNumber="29" value="1984-12-20">20.XII.1984</date>
-
<date id="3A1A39E38621D836D966FE9EBE66FEBE" box="[261,349,300,324]" pageId="28" pageNumber="29" value="1985-02-08">8.II.1985</date>
/FMHD#85-930 Derumbes for. Trail, S.andJ. Peck, P#85-45, FIT FIELD MUSEUM NAT HIST/
<specimenCount id="58A2D4AA8621D836DA80FEFDBDC6FE99" box="[739,765,335,355]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
/ FMNH-INS 0000 024 169 [
<specimenCount id="58A2D4AA8621D836D9CBFED9BEF4FE78" box="[424,463,363,387]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38621D836DA5DFED9BDABFE79" box="[574,656,363,387]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18621D836DAC2FEDEBC3FFE79" box="[673,772,363,387]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
] Valdivia Prov.,
<quantity id="895CB2C68621D836D943FE39BE5DFE59" box="[288,358,395,419]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="28" pageNumber="29" unit="km" value="26.0">26 km</quantity>
SE Panguipulli [-39.75, -72.15]
<quantity id="895CB2C68621D836DAAAFE39BC3EFE59" box="[713,773,395,419]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="300.0">300m</quantity>
,
<date id="3A1A39E38621D836D8E2FE18BFC6FE38" box="[129,253,426,450]" pageId="28" pageNumber="29" value="1984-12-16">16.XII.1984</date>
-
<date id="3A1A39E38621D836D966FE18BE51FE38" box="[261,362,426,450]" pageId="28" pageNumber="29" value="1985-02-11">11.II.1985</date>
/FMHD#85-921,
<taxonomicName id="89A464A08621D836DA4DFE18BD9EFE38" box="[558,677,426,450]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
remnant S.andJ. Peck, P#85-34, carrion trap FIELD MUSEUM NAT HIST/
<specimenCount id="58A2D4AA8621D836D8ABFE5EBFD9FDFA" box="[200,226,492,512]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
/FMNH-INS 0000 024 168 [
<specimenCount id="58A2D4AA8621D836DA78FE5BBD78FDFA" box="[539,579,489,513]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38621D836DAD7FE5BBC3DFDFB" box="[692,774,489,513]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18621D836D8E4FDBABFDCFDDA" box="[135,231,520,544]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
] Prov. Valdivia: Parque Oncol (Cruce Sendero Tepual) Rio Cruces [-39.72, -73.27]
<date id="3A1A39E38621D836D9C5FD95BEC5FDC5" box="[422,510,551,575]" pageId="28" pageNumber="29" value="2001-01-08">8-I-2001</date>
T Cekalovic CHIL1CO1 005/[Barcode] SM0723095 KUNHM-ENT [
<specimenCount id="58A2D4AA8621D836DA1AFDF5BD9BFDA4" box="[633,672,583,607]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
SEMC]; [
<collectingCountry id="36B35FB38621D836D8E4FDD4BFE7FD84" box="[135,220,614,638]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D88BFDD4BE73FD84" box="[232,328,614,638]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
] 85/Süd-Chile 31.1.90 Puerto Puyehue [~ -40.65, -72.33] leg.G.H. Schwabe [
<specimenCount id="58A2D4AA8621D836D993FD37BD21FD66" box="[496,538,645,669]" count="2" pageId="28" pageNumber="29" type="male">2♂</specimenCount>
<specimenCount id="58A2D4AA8621D836DA46FD37BD77FD66" box="[549,588,645,669]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
SDEI]; [
<collectingCountry id="36B35FB38621D836DAD2FD37BC3DFD67" box="[689,774,645,669]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D8E2FD17BFC8FD47" box="[129,243,677,701]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836D910FD17BEDBFD47" box="[371,480,677,701]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
] Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836D8B5FD76BE74FD26" box="[214,335,708,732]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
/
<collectingCountry id="36B35FB38621D836D93BFD76BE90FD26" box="[344,427,708,732]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/
<taxonomicName id="89A464A08621D836DA55FD76BDB6FD26" box="[566,653,708,732]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Toưenham TYPE [manuscript name]/NHM [
<specimenCount id="58A2D4AA8621D836DAE2FD51BD92FD00" box="[641,681,739,763]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D8E4FCB1BFE7FCE1" box="[135,220,771,795]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D88AFCB1BE60FCE1" box="[233,347,771,795]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836D98EFCB1BD61FCE1" box="[493,602,771,795]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
] Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836D903FC90BEE2FCC0" box="[352,473,802,826]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
/
<collectingCountry id="36B35FB38621D836D982FC90BD0FFCC0" box="[481,564,802,826]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/
<taxonomicName id="89A464A08621D836DAD1FC90BC32FCC0" box="[690,777,802,826]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Toưenham PAÞTYPE [manuscript name]/NHM [
<specimenCount id="58A2D4AA8621D836D8EAFCD3BF89FC82" box="[137,178,865,889]" count="3" pageId="28" pageNumber="29" type="male">3♂</specimenCount>
<specimenCount id="58A2D4AA8621D836D8D5FCD3BFE6FC82" box="[182,221,865,889]" count="2" pageId="28" pageNumber="29" type="female">2♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D95FFCD3BEAAFC83" box="[316,401,865,889]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D9FBFCD3BD31FC83" box="[408,522,865,889]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836DAE8FCD3BDC7FC83" box="[651,764,865,889]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
]/
<typeStatus id="911FA1818621D836D8E2FC32BFE0FC62" box="[129,219,896,920]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
/Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836DA10FC32BDD0FC62" box="[627,747,896,920]" pageId="28" pageNumber="29" value="1958-11-30">30.XI.1958</date>
/C. E. Toưenham collection. B.M. 1974-587/
<collectingCountry id="36B35FB38621D836DA52FC2DBDB9FC4D" box="[561,642,927,951]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/ nigroflavus Tot [manuscript name]/NHM/PAÞTYPE
<taxonomicName id="89A464A08621D836D8E2FC6CBE68FC0C" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[129,339,990,1014]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D8E2FC6CBE68FC0C" box="[129,339,990,1014]" italics="true" pageId="28" pageNumber="29">Loncovilius variabilis</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023/NHMUK015025170 [
<specimenCount id="58A2D4AA8621D836DAEFFC4FBD88FBEE" box="[652,691,1021,1045]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D8E4FBAFBFE7FBCF" box="[135,220,1053,1077]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D88AFBAFBE60FBCF" box="[233,347,1053,1077]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836D98EFBAFBD61FBCF" box="[493,602,1053,1077]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
] Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836D90DFB8EBEDCFBAE" box="[366,487,1084,1108]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
/
<collectingCountry id="36B35FB38621D836D993FB8EBD78FBAE" box="[496,579,1084,1108]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/Qued. Sp. 5/
<taxonomicName id="89A464A08621D836D8DEFBE9BE5AFB89" authority="sp." authorityName="Sp." box="[189,353,1115,1139]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="30" phylum="Arthropoda" rank="genus">
Loncovilius
<taxonomicNameLabel id="67E37E4A8621D836D926FBEEBE5AFB89" box="[325,353,1116,1139]" pageId="28" pageNumber="29" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
Nov. P.M. Hammond det. 1980/NHM [Blue label]/NHMD 917069 [
<specimenCount id="58A2D4AA8621D836D9A4FBC9BED5FB68" box="[455,494,1147,1171]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHM];
<collectingCountry id="36B35FB38621D836DA30FBC9BD9EFB69" box="[595,677,1147,1171]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836DAD1FBC9BC32FB69" box="[690,777,1147,1171]" country="Chile" name="Region Metropolitana" pageId="28" pageNumber="29">Santiago</collectingRegion>
[
<collectingRegion id="8C60D1C18621D836D8E5FB28BFDAFB48" box="[134,225,1178,1202]" country="Chile" name="Region Metropolitana" pageId="28" pageNumber="29">Santiago</collectingRegion>
Metropolitan]: El Alfalfal [~ -33.51, -70.20]
<date id="3A1A39E38621D836DAABFB28BC31FB48" box="[712,778,1178,1202]" pageId="28" pageNumber="29" value="1977-01">I.1977</date>
L.E.Peña, leg. FIELD MUS. NAT. HIST./FMNH-INS 0000 024 157–162 [
<specimenCount id="58A2D4AA8621D836D97FFB6BBE78FB0A" box="[284,323,1241,1265]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8621D836D928FB6BBE49FB0A" box="[331,370,1241,1265]" count="2" pageId="28" pageNumber="29" type="female">2♀</specimenCount>
FMNH;
<specimenCount id="58A2D4AA8621D836D9B4FB6BBEC5FB0A" box="[471,510,1241,1265]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8621D836DA65FB6BBD16FB0A" box="[518,557,1241,1265]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHMD ex. FMNH;
<specimenCount id="58A2D4AA8621D836D8E2FB4ABF9CFAF5" box="[129,167,1272,1296]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
CZUG ex. FMNH;
<superScript id="B9D1B26B8621D836D91EFB44BEB9FAFE" attach="right" box="[381,386,1270,1284]" fontSize="6" pageId="28" pageNumber="29">*</superScript>
this is potentially a labelling error. Ŋe
<taxonomicName id="89A464A08621D836D8E1FAAABFCFFAD5" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[130,244,1303,1327]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D8E1FAAABFB4FAD5" box="[130,143,1304,1327]" italics="true" pageId="28" pageNumber="29">L</emphasis>
.
<emphasis id="7CD0C3318621D836D8FFFAA5BFCFFAD5" box="[156,244,1303,1327]" italics="true" pageId="28" pageNumber="29">variabilis</emphasis>
</taxonomicName>
model shown in
<figureCitation id="D69F03A68621D836D9C8FAA5BD3AFAD5" box="[427,513,1303,1327]" captionStart="Figure 3" captionStartId="11.[113,178,1776,1800]" captionTargetBox="[114,1458,144,1740]" captionTargetId="figure-6@11.[114,1458,144,1749]" captionTargetPageId="11" captionText="Figure 3. Ecological niche modelling of Loncovilius species. A–D, potential suitable areas predicted with free extrapolation in South America. Binarized projection with 5% threshold for suitability values and mobility-oriented parity analysis (MOP) showing areas for which current model predictions are strictly based on extrapolation. E–H, predictions under shared socio-economic pathways (SSPs) 126, and agreement between different global climate models (GCMs), and MOP agreement among GCMs with strict extrapolative conditions under the low emissions.I–L, predictions under SSPs 585, agreement between different GCMs, and MOP agreement among GCMs with strict extrapolative conditions under the high emissions." figureDoi="http://doi.org/10.5281/zenodo.14443953" httpUri="https://zenodo.org/record/14443953/files/figure.png" pageId="28" pageNumber="29">Figure 3</figureCitation>
does not include this occurrence. Ŋe sets of variables and the calibration results that do include this occurrence can be consulted in the Supporting Information,
<tableCitation id="03262A988621D836D96AFAC7BE59FA77" box="[265,354,1397,1421]" captionStart="Table 1" captionStartId="3.[115,170,893,917]" captionText="Table 1.Taxon sample for the phylogenetic analysis (less column) and resulting taxonomic changes (right column)." pageId="28" pageNumber="29">Table S1</tableCitation>
, and in the Supporting Information, File S6]; [
<collectingCountry id="36B35FB38621D836D8D8FA27BE2BFA57" box="[187,272,1429,1453]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: unknown locality and date]
<collectingCountry id="36B35FB38621D836DA22FA27BD4CFA57" box="[577,631,1429,1453]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
./
<collectingCountry id="36B35FB38621D836DAE1FA27BD83FA57" box="[642,696,1429,1453]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
[Green round label]/Sharp Coll 1905-313./
<taxonomicName id="89A464A08621D836DA46FA06BDBAFA36" box="[549,641,1460,1484]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Toưenham PAÞTYPE [manuscript name]/NHM [
<specimenCount id="58A2D4AA8621D836DA81FA61BC32FA10" box="[738,777,1491,1515]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D8BEFA41BE09F9F1" box="[221,306,1523,1547]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: unknown locality and date]
<collectingCountry id="36B35FB38621D836DA30FA41BDBDF9F1" box="[595,646,1523,1547]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
Sharp Coll./
<taxonomicName id="89A464A08621D836D8E2F9A0BFE3F9D0" box="[129,216,1554,1578]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Brnh (Typ.) [manuscript name]/BRIT. MUSEUM DON. ARROW/B. Typ. ohne Flg.d streifen/Chicago NHMus M.Bernhauer Collection/Bernhauer MS name? Current genus:
<taxonomicName id="89A464A08621D836D97EF9C2BEAEF972" box="[285,405,1648,1672]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Loncovilius</taxonomicName>
teste A.Newton 2000 [
<specimenCount id="58A2D4AA8621D836DA1DF9C2BD9EF97D" box="[638,677,1648,1672]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38621D836D8E5F93DBFE0F95D" box="[134,219,1679,1703]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: unknown locality and date] 53793/Reed/
<collectingCountry id="36B35FB38621D836DAC6F93DBDE0F95D" box="[677,731,1679,1703]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
/Fry Coll. 1905.100/Chicago NHMus M.Bernhauer Collection/
<specimenCount id="58A2D4AA8621D836DA80F900BDC6F93C" box="[739,765,1714,1734]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
/ FMNH-INS 0000 024 171 [
<specimenCount id="58A2D4AA8621D836D9D2F97CBEE3F91F" box="[433,472,1742,1766]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
FMNH]. All
<typeStatus id="911FA1818621D836DA13F97CBDEAF91C" box="[624,721,1742,1766]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
with label: PAÞTYPE
<taxonomicName id="89A464A08621D836D924F95FBD2EF8FF" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[327,533,1773,1797]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D924F95FBD2EF8FF" box="[327,533,1773,1797]" italics="true" pageId="28" pageNumber="29">Loncovilius variabilis</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
</paragraph>
<paragraph id="4E1B1F238621D836D8E2F8F5BD72F8A4" blockId="28.[129,585,1862,1886]" box="[129,585,1862,1886]" pageId="28" pageNumber="29">
<emphasis id="7CD0C3318621D836D8E2F8F5BFDCF8A4" box="[129,231,1863,1886]" italics="true" pageId="28" pageNumber="29">Diagnosis:</emphasis>
Same as
<taxonomicName id="89A464A08621D836D920F8F4BEA7F8A4" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[323,412,1862,1886]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D920F8F4BEA7F8A4" box="[323,412,1862,1886]" italics="true" pageId="28" pageNumber="29">variabilis</emphasis>
</taxonomicName>
group diagnosis.
</paragraph>
<paragraph id="4E1B1F238621D836D8E2F832BA58FEFC" blockId="28.[129,778,1919,1975]" lastBlockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
<emphasis id="7CD0C3318621D836D8E2F832BFCCF86D" box="[129,247,1920,1943]" italics="true" pageId="28" pageNumber="29">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8621D836D9B5F830BECBF86C" box="[470,496,1922,1942]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318621D836D8E2F82DBFAFF84C" box="[129,148,1951,1974]" italics="true" pageId="28" pageNumber="29">N</emphasis>
= 10]: FBL = 2.79–3.19 (2.98); TL = 5.12–6.25 (5.61); HW = 0.72–0.8 (0.77); HL = 0.74–0.83 (0.79); HW/ HL = 0.95–1 (0.97); PW = 0.95–1.05 (1); PL = 0.88–0.98 (0.93); PW/PL = 1.02–1.09 (1.07); EW = 1.31–1.54 (1.38); EL = 1.17–1.43 (1.25); PW/HW = 1.23–1.33 (1.30).
</paragraph>
<paragraph id="4E1B1F238621D836DB36FEBCBB50FE38" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
Measurements
<specimenCount id="58A2D4AA8621D836DC6CFEA2BB12FEDE" box="[1039,1065,272,292]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318621D836DD05FEBCBA42FEDF" box="[1382,1401,270,293]" italics="true" pageId="28" pageNumber="29">N</emphasis>
= 10]: FBL = 3.05–3.53 (3.33); TL = 5.5–7.44 (6.4); HW = 0.79– 0.86 (0.83); HL = 0.82–0.9 (0.85); HW/HL = 0.94–1 (0.97); PW = 1.05–1.17 (1.10); PL = 1–1.08 (1.04); PW/PL = 1.03– 1.09 (1.06); EW = 1.36–1.74 (1.56); EL = 1.22–1.55 (1.43); PW/HW = 1.29–1.39 (1.34).
</paragraph>
<paragraph id="4E1B1F238621D836DB36FE7BBAB0FDC5" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">Head, pronotum, and elytra black to brown-black, head darker; antennae, mouthparts, and legs yellowish-brown; abdominal tergites and sternites variable in coloration, from dark reddish-brown to brown-black with pale posterior margin.</paragraph>
<paragraph id="4E1B1F238621D836DB36FDF5BA6CFA94" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
Head from about as wide as long to, usually, slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.47), from 1.03 to 1.29 times as long as temples (in lateral view); distance between eyes about 1.25 times as long as length of eye in males and 1.34 times in females. Antennomere 3 slightly longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 1.53-1.72 times as long as a
<quantity id="895CB2C68621D836DC2FFC32BB44FC62" box="[1100,1151,896,920]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="28" pageNumber="29" unit="in" value="10.0">10 in</quantity>
males and
<quantity id="895CB2C68621D836DC8CFC32BA52FC62" box="[1263,1385,896,920]" metricMagnitude="-2" metricUnit="m" metricValue="4.0259" metricValueMax="4.318" metricValueMin="3.7338" pageId="28" pageNumber="29" unit="in" value="1.585" valueMax="1.7" valueMin="1.47">1.47-1.70 in</quantity>
females. Basal puncture usually double; parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setiferous punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced to small trace near gular sutures; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with sub-rectangular sclerotized region and entire to slightly notched apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical to fusiform apical palpomere.
</paragraph>
<paragraph id="4E1B1F238621D836DB36FAC4BA12F9D0" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">Pronotum slightly wider than long, convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to 5 or even 10 times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F238621D836DB36F983BBB1F8FF" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. Mesotarsomeres 1–4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres
<quantity id="895CB2C68621D836DC7DF93DBB5DF95D" box="[1054,1126,1679,1703]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="28" pageNumber="29" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
both sexes with terminal plate. Metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.
</paragraph>
<paragraph id="4E1B1F238621D837DB36F8BFBDF5FEBF" blockId="28.[824,1476,144,1985]" lastBlockId="29.[113,765,144,544]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Abdominal tergites III and IV with several rows of sparse and fine punctures of moderate density, without sizeable impunctate area; tergite VIII only slightly emarginate medio-apically in females. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with broad and shallow U-shaped emargination medially (
<figureCitation id="D69F03A68620D837D90AFF22BEF9FF52" box="[361,450,144,168]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="29" pageNumber="30">Fig. 14D</figureCitation>
); female sternite VIII without emargination; male sternite IX without emargination medially (apex subtruncate), its basal portion moderately longer than distal portion; male tergite X with anterior margin subtruncate (
<figureCitation id="D69F03A68620D837D81FFEBFBFEDFEDF" box="[124,214,269,293]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="29" pageNumber="30">Fig. 12D</figureCitation>
); female tergite X apically truncate (
<figureCitation id="D69F03A68620D837DA3DFEBFBD83FEDF" box="[606,696,269,293]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="29" pageNumber="30">Fig. 13D</figureCitation>
); ovipositor, its second gonocoxite with one macroseta medially.
</paragraph>
<paragraph id="4E1B1F238620D837D8EEFEFEBD4AFDDA" blockId="29.[113,765,144,544]" pageId="29" pageNumber="30">
Aedeagus as in
<figureCitation id="D69F03A68620D837D94FFEFEBE8EFE9E" box="[300,437,332,356]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="29" pageNumber="30">Figure 10I–K</figureCitation>
; its total length ~ 0.95. Median lobe rod-like, apex not emarginate; internal sac with median vertical sclerite and pair of short I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe; wider than median lobe; converging into a subrounded apex, slightly projected in the middle; with dark peg setae; in lateral view paramere almost straight, slightly concave apically.
</paragraph>
<paragraph id="4E1B1F238620D837D812FDF3BE83FCEF" blockId="29.[113,763,577,789]" pageId="29" pageNumber="30">
<emphasis id="7CD0C3318620D837D812FDF3BFDAFDA3" box="[113,225,577,601]" italics="true" pageId="29" pageNumber="30">Etymology:</emphasis>
Ŋe species epithet is the Latin adjective
<taxonomicName id="89A464A08620D837DAF6FDF3BDCFFDA3" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[661,756,577,601]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="variabilis">
‘
<emphasis id="7CD0C3318620D837DAF8FDF3BDCFFDA3" box="[667,756,577,601]" italics="true" pageId="29" pageNumber="30">variabilis</emphasis>
</taxonomicName>
’ (variable) and refers to a notable intraspecific morphological variation of this taxon. Ŋis variation can occur between individuals from the same locality, e.g. it can be completely blackishbrown forms and others with a reddish-brown abdomen. Ŋe head punctation also varies, and loss or duplication of paraocular and basal punctures is common.
</paragraph>
<paragraph id="4E1B1F238620D837D812FC85BE22FB93" blockId="29.[113,764,823,1129]" pageId="29" pageNumber="30">
<emphasis id="7CD0C3318620D837D812FC85BD62FCB5" box="[113,601,823,847]" italics="true" pageId="29" pageNumber="30">
Distribution and habitat:
<taxonomicName id="89A464A08620D837D91CFC85BD62FCB5" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[383,601,823,847]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="variabilis">Loncovilius variabilis</taxonomicName>
</emphasis>
is known only from
<collectingCountry id="36B35FB38620D837D8D3FCE4BFD7FC94" box="[176,236,854,878]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
, from the following regions:
<collectingRegion id="8C60D1C18620D837DA54FCE4BD9EFC94" box="[567,677,854,878]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837DAD7FCE4BDCDFC94" box="[692,758,854,878]" country="Chile" name="Aysen" pageId="29" pageNumber="30">Aysén</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837D812FCC4BFDEFC74" box="[113,229,886,910]" country="Chile" name="Los Lagos" pageId="29" pageNumber="30">Los Lagos</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837D896FCC4BE62FC77" box="[245,345,885,909]" country="Chile" name="Los Rios" pageId="29" pageNumber="30">Los Ríos</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837D90AFCC7BED8FC77" box="[361,483,885,909]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="29" pageNumber="30">Magallanes</collectingRegion>
, and Chilean
<collectingCountry id="36B35FB38620D837DAE6FCC7BDCDFC77" box="[645,758,885,909]" name="Antarctica" pageId="29" pageNumber="30">Antarctica</collectingCountry>
, and from Santiago Metropolitan. The species occurs from lowlands to
<quantity id="895CB2C68620D837D896FC06BE73FC31" box="[245,328,948,972]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">1300 m</quantity>
of elevation in the mountains; confined to various
<typeStatus id="911FA1818620D837D88FFC66BE1EFC16" box="[236,293,980,1004]" pageId="29" pageNumber="30">types</typeStatus>
of forests in the Magellanic subpolar and Valdivian temperate forest ecoregions. It has been collected using Malaise traps, and directly from flowers, e.g. from
<taxonomicName id="89A464A08620D837D812FB83BE03FBB3" box="[113,312,1073,1097]" class="Magnoliopsida" family="Myrtaceae" genus="Tepualia" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="stipularis">
<emphasis id="7CD0C3318620D837D812FB83BE03FBB3" box="[113,312,1073,1097]" italics="true" pageId="29" pageNumber="30">Tepualia stipularis</emphasis>
</taxonomicName>
(
<taxonomicName id="89A464A08620D837D92DFB83BEFEFBB3" box="[334,453,1073,1097]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) and
<taxonomicName id="89A464A08620D837DA71FB83BDC1FBB3" box="[530,762,1073,1097]" class="Magnoliopsida" family="Cunoniaceae" genus="Caldcluvia" kingdom="Plantae" order="Oxalidales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="paniculata">
<emphasis id="7CD0C3318620D837DA71FB83BDC1FBB3" box="[530,762,1073,1097]" italics="true" pageId="29" pageNumber="30">Caldcluvia paniculata</emphasis>
</taxonomicName>
(
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).
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264
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<emphasis id="7CD0C3318627D830D94FF9ECBD36F982" bold="true" box="[300,525,1630,1657]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
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(
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,
<figureCitation id="D69F03A68627D830D914F92FBEFEF94D" box="[375,453,1693,1719]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="26" pageNumber="27">10G, H</figureCitation>
,
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,
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,
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)
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male, mounted, with genitalia in a separate microvial, with labels as follows:
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[~ -38.70, -72.01]
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<collectingDate id="2A5EC00B8627D830D97FF8DABE41F87A" box="[284,378,1896,1920]" pageId="26" pageNumber="27" value="1954-07">VII-1954</collectingDate>
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, leg./
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/FMNH-INS 0000 024 367
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/
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<emphasis id="7CD0C3318627D830D927F835BD27F865" box="[324,540,1927,1951]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
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des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
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:
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:
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[~ -38.70, -72.01]
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, leg./
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/FMNH-INS 0000 024 368-369 [
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FMNH]
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;
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E Manzanar
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[-38.44, -71.51]
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</date>
<collectorName id="E3517AF58627D830DCE7FF5CBA19FEFC" box="[1156,1314,238,262]" pageId="26" pageNumber="27">H. F. Howden</collectorName>
/CNCI L2004- 28/
<specimenCount id="58A2D4AA8627D830DB03FEA2BC41FEDE" box="[864,890,272,292]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
/FMNH-INS 0000 024 465 [
<specimenCount id="58A2D4AA8627D830DCC0FEBFBBF0FEDE" box="[1187,1227,269,293]" count="1" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
CNC]
</materialsCitation>
;
<materialsCitation id="FECC157E8627D830DD7DFEBFBA63FE99" collectorName="Coll. L. Pena" country="Chile" location="Chile" pageId="26" pageNumber="27" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<collectingCountry id="36B35FB38627D830DD7DFEBFBA60FEDF" box="[1310,1371,269,293]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
: Tolhuaco [~ -38.23, -71.71]11.I.62/
<collectorName id="E3517AF58627D830DC25FE9FBBECFEBF" box="[1094,1239,301,325]" pageId="26" pageNumber="27">Coll. L. Peña</collectorName>
/
<specimenCount id="58A2D4AA8627D830DC83FE82BBC1FEBE" box="[1248,1274,304,324]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
/FMNH-INS 0000 024 364–366 [
<specimenCount id="58A2D4AA8627D830DBADFEFEBCC3FE99" box="[974,1016,332,356]" count="2" pageId="26" pageNumber="27" type="male">2♂</specimenCount>
FMNH;
<specimenCount id="58A2D4AA8627D830DC3BFEFEBB44FE99" box="[1112,1151,332,356]" count="1" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
MNNC ex. FMNH]
</materialsCitation>
.
</paragraph>
<paragraph id="4E1B1F238627D830DB36FED9BB6EFE38" blockId="26.[824,1474,144,450]" pageId="26" pageNumber="27">
All
<typeStatus id="911FA1818627D830DBE0FEDEBCDDFE7E" box="[899,998,364,388]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
are supplied with the label: PAÞTYPE
<taxonomicName id="89A464A08627D830DB5AFE39BB2DFE58" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[825,1046,395,419]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus">
<emphasis id="7CD0C3318627D830DB5AFE39BB2DFE58" box="[825,1046,395,419]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
</paragraph>
<paragraph id="4E1B1F238627D830DB5AFE56BA3BFD2C" blockId="26.[824,1475,483,727]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830DB5AFE56BCA4FE01" box="[825,927,484,507]" italics="true" pageId="26" pageNumber="27">Diagnosis:</emphasis>
Head dorsally with meshed microsculpture; gula with isodiametric microsculpture in the middle; females with antennomere 11 (a11) distinctly longer than antennomere 10 (a10; a11:a10 ratio&gt;1.5). Elytra reddish-brown, disc glossy with only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII in both sexes with medial apical emargination. Aedeagus as in
<figureCitation id="D69F03A68627D830DC04FD0DBBC1FD2C" box="[1127,1274,703,727]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="26" pageNumber="27">Figure 10G, H</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238627D830DB5AFD4BBB0CFC57" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
<emphasis id="7CD0C3318627D830DB5AFD4BBC8BFCEA" box="[825,944,761,784]" italics="true" pageId="26" pageNumber="27">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8627D830DC02FD49BB40FCF5" box="[1121,1147,763,783]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318627D830DD10FD4BBABDFCEA" box="[1395,1414,761,784]" italics="true" pageId="26" pageNumber="27">N</emphasis>
= 3]: FBL = 3.35–3.59 (3.48); HW = 1.01–1.1 (1.07); HL = 0.85– 0.93 (0.89); HW/HL = 1.18–1.21 (1.19); PW = 1.4– 1.44 (1.42); PL = 1.2–1.25 (1.23); PW/PL = 1.14–1.17 (1.15); EW = 1.67–1.7 (1.68); EL = 1.3–1.44 (1.36); PW/ HW = 1.28–1.39 (1.33).
</paragraph>
<paragraph id="4E1B1F238627D830DB36FC06BB81FB93" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
Measurements
<specimenCount id="58A2D4AA8627D830DB93FC05BB31FC31" box="[1008,1034,951,971]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318627D830DC89FC07BBC6FC36" box="[1258,1277,949,972]" italics="true" pageId="26" pageNumber="27">N</emphasis>
= 4]: FBL = 3.71– 3.93 (3.81); TL = 7.5–7.53 (7.52); HW = 1.17–1.18 (1.18); HL = 0.93–0.97 (0.95); HW/HL = 1.22–1.26 (1.24); PW = 1.54–1.59 (1.56); PL = 1.29–1.37 (1.32); PW/ PL = 1.16–1.19 (1.18); EW = 1.82–1.9 (1.87); EL = 1.46–1.59 (1.54); PW/HW = 1.32–1.35 (1.33).
</paragraph>
<paragraph id="4E1B1F238627D830DB36FBC2BCBAFB3D" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">Head, mouthparts, and legs black to brown-black; pronotum, antennae, elytra, and abdomen dark reddish-brown, elytra lighter.</paragraph>
<paragraph id="4E1B1F238627D830DB36FB7CBB62F84D" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse meshes, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes of medium size (EYL/HL = x = 0.52), from 1.51 to 1.64 times longer than temples (in lateral view); distance between eyes around 1.8 times the length of eye. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 subequal in length; antennomeres 4 to 10 gradually reduced in length; a11 from 1.54 to 1.75 times longer than a
<quantity id="895CB2C68627D830DD2EFA5ABABAF9FA" box="[1357,1409,1512,1536]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68627D830DBC4F9B5BB7CF9E5" box="[935,1095,1543,1567]" metricMagnitude="-2" metricUnit="m" metricValue="4.0893999999999995" metricValueMax="4.2672" metricValueMin="3.9116" pageId="26" pageNumber="27" unit="in" value="1.6099999999999999" valueMax="1.68" valueMin="1.54">1.54 to 1.68 in</quantity>
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical.
</paragraph>
<paragraph id="4E1B1F238626D831D8EEFF22BD10FEBF" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">Pronotum transverse, convex, evenly curved, with one PPDS and an APP, without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setose; disc with only a few rows of setiferous punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F238626D831D8EEFEFEBFFBFDC5" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrallywith apical margin slightly sinuate.</paragraph>
<paragraph id="4E1B1F238626D831D8EEFDF5BD5EFC62" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flaưened only in males. Male sternite VIII with broad V-shaped emargination medially (
<figureCitation id="D69F03A68626D831D910FD51BEEBFD01" box="[371,464,739,763]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="27" pageNumber="28">Fig. 14C</figureCitation>
); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (
<figureCitation id="D69F03A68626D831DA63FCF3BD6CFCA3" box="[512,599,833,857]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="27" pageNumber="28">Fig. 12C</figureCitation>
); female tergite X truncate to slightly arcuate medially (
<figureCitation id="D69F03A68626D831DA61FCD3BD62FC83" box="[514,601,865,889]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="27" pageNumber="28">Fig. 13C</figureCitation>
); ovipositor, its second gonocoxite without macrosetae medially.
</paragraph>
<paragraph id="4E1B1F238626D831D8EEFC2DBDAAFBAE" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">
Aedeagus as in
<figureCitation id="D69F03A68626D831D951FC2DBEF7FC4D" box="[306,460,927,951]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="27" pageNumber="28">Figure 10G, H</figureCitation>
; its total length 1.2. Median lobe bulky, in parameral view with truncated apex; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into M-shape apex, with dark peg setae, in lateral view almost straight and slightly concave apically.
</paragraph>
<paragraph id="4E1B1F238626D831D812FBC7BD11FB36" blockId="27.[113,762,1141,1228]" pageId="27" pageNumber="28">
<emphasis id="7CD0C3318626D831D812FBC7BFDAFB77" box="[113,225,1141,1165]" italics="true" pageId="27" pageNumber="28">Etymology:</emphasis>
Ŋe species epithet is derived from the Latin prefix ‘
<emphasis id="7CD0C3318626D831D814FB27BFAAFB56" box="[119,145,1173,1196]" italics="true" pageId="27" pageNumber="28">im</emphasis>
’ (not) and adjective ‘
<emphasis id="7CD0C3318626D831D90DFB27BEFAFB56" box="[366,449,1173,1196]" italics="true" pageId="27" pageNumber="28">punctate</emphasis>
’ (doưed), referring to the disc of pronotum and elytra mostly impunctate.
</paragraph>
<paragraph id="4E1B1F238626D831D812FB5FBDF2FABE" blockId="27.[112,762,1261,1348]" pageId="27" pageNumber="28">
<emphasis id="7CD0C3318626D831D812FB5FBD05FAFF" box="[113,574,1261,1285]" italics="true" pageId="27" pageNumber="28">
Distribution and habitat:
<taxonomicName id="89A464A08626D831D904FB5FBD05FAFF" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[359,574,1261,1285]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="impunctus">Loncovilius impunctus</taxonomicName>
</emphasis>
is known from the
<collectingRegion id="8C60D1C18626D831D813FABFBE1DFADF" box="[112,294,1293,1317]" country="Chile" name="Araucania" pageId="27" pageNumber="28">Araucanía Region</collectingRegion>
in
<collectingCountry id="36B35FB38626D831D927FABFBE47FADF" box="[324,380,1293,1317]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
where it is confined to Valdivian temperate forest ecoregion. It is unknown how it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>

409
data/DB/45/87/DB4587EEFFB5412BFCDCFECBFE67FD03.xml Normal file
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@ -0,0 +1,409 @@
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<mods:title id="69EE60F1A8C56B693410778A210F07FD">A tenuis relationship: traditional taxonomy obscures systematics and biogeography of the ‘ Acropora tenuis’ (ºcleractinia: Acroporidae) species complex</mods:title>
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<mods:namePart id="1C8457FE4B1266B0B510DE93448E4234">Bridge, Tom C. L.</mods:namePart>
<mods:affiliation id="444DACD4342BC37F53704DCBC27423B0">Biodiversity and Geosciences Program * Museum of Tropical Queensland * Queensland Museum Network * Townsville * QLD * Australia &amp; Australian Research Council Centre of Excellence for Coral Reef ºtudies * James Cook University * Townsville * QLD * Australia &amp; College of ºcience and Engineering * James Cook University * Townsville * QLD * Australia</mods:affiliation>
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<mods:namePart id="EFDD6051073980807488042C27402B1E">Cowman, Peter F.</mods:namePart>
<mods:affiliation id="8B095B1EE2ED8C5C231351CD8CAAF70D">Biodiversity and Geosciences Program * Museum of Tropical Queensland * Queensland Museum Network * Townsville * QLD * Australia &amp; Australian Research Council Centre of Excellence for Coral Reef ºtudies * James Cook University * Townsville * QLD * Australia &amp; College of ºcience and Engineering * James Cook University * Townsville * QLD * Australia</mods:affiliation>
<mods:nameIdentifier id="5DABD4A66C13C705C38D7AF14F1944E0" type="email">tom.bridge@qm.qld.gov.au</mods:nameIdentifier>
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<mods:namePart id="B361F1D97D129986AACE887B8693F82D">Quaưrini, Andrea M.</mods:namePart>
<mods:affiliation id="AE80A8B1F20763EA23DFBEAB9AFA5878">Department of Invertebrate Zoology * National Museum of Natural History * ºmithsonian Institution * Washington * DC * United ºtates</mods:affiliation>
</mods:name>
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<mods:role id="4D704C908FECC35C06F0D0ECBA6012D8">
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<mods:namePart id="9D5BA4D82AB3BBFDADC9817A1A2FF4B6">Bonito, Victor E.</mods:namePart>
<mods:affiliation id="847132AD1BC736F0D897BE8450D6037C">Coral Coast Conservation Center * Votua Village * Baravi * Nadroga * Fiji</mods:affiliation>
</mods:name>
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<mods:role id="775FC6FC1EAE5C2338F1C2E39E89C4DF">
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<mods:namePart id="D5D22485F5152749BA6BAE8B66725D6C">Frederic</mods:namePart>
<mods:affiliation id="1DAA46F426004E9E39134DCCACB04B24">ºesoko ºtation * Tropical Biosphere Research Center * University of the Ryukyus * 3422 ºesoko * Motobu * Okinawa * Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="4A8E353DABF9607C5CD3A2FB04BB00BB">Harii</mods:namePart>
<mods:affiliation id="1F1FBA749A9A3F9D965893B8AF42DF1A">ºesoko ºtation * Tropical Biosphere Research Center * University of the Ryukyus * 3422 ºesoko * Motobu * Okinawa * Japan</mods:affiliation>
</mods:name>
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<mods:roleTerm id="24DC8797AFE24D42D39B7977AC280AC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B1773077C072411191EB34EEC0C2F91">Head, Catherine E. I.</mods:namePart>
<mods:affiliation id="2222191CDE04FCCE3541B296343F3DA9">Institute of Zoology * Zoological ºociety of London * London * United Kingdom &amp; Department of Zoology * University of Oxford * Oxford * United Kingdom</mods:affiliation>
<mods:nameIdentifier id="C1F9DCF12AAECCC435DB0EDDAB70ADEF" type="email">tom.bridge@qm.qld.gov.au</mods:nameIdentifier>
</mods:name>
<mods:name id="8034296B48D0415E046153FE4C40DAB8" type="personal">
<mods:role id="469CB6CDA6FFF2A6B187503D178D4235">
<mods:roleTerm id="61F70B24FDC41C0EF20EC1E0B7ED756F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="88ED0B90FC5F0979C9BA50DD16BB69AA">Hung, Julia Y.</mods:namePart>
<mods:affiliation id="A74A6BB5C4A4500831D1EC994D541F11">College of ºcience and Engineering * James Cook University * Townsville * QLD * Australia</mods:affiliation>
</mods:name>
<mods:name id="7E0067EA68B9BB2120B2F8D04F7034FB" type="personal">
<mods:role id="04A77DE173A88FA4B855301DA5AC112D">
<mods:roleTerm id="9D61C6D4E420685589C4C5CB91B1B4E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="985B2D8FE6CA77E2CBE21BCEFF9737B0">Halafihi, Tuikolongahau</mods:namePart>
<mods:affiliation id="3861F9CC0C0E6E1B1C14551DBA632503">Ministry of Fisheries * Nuku’alofa * Tongatapu * Tonga</mods:affiliation>
</mods:name>
<mods:name id="662E11BFAB3A9731DF9627CE73C7A78E" type="personal">
<mods:role id="846CE1CBA34F0D1FA49695995183817F">
<mods:roleTerm id="629E11BAAFFAFC90F2649CC1DD064EFF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="215D0DEF17820FDDD0498670643F7515">Rongo, Teina</mods:namePart>
<mods:affiliation id="F69AF2748C950C02EAD6330714A56C4D">Kōrero O Te`Ōrau * Rarotonga * Cook Islands</mods:affiliation>
</mods:name>
<mods:name id="24DE94D6BD518C9CA7BFA8055BF9CC03" type="personal">
<mods:role id="24B841262F009552A8942E6564499835">
<mods:roleTerm id="F3C33DADB43D603E34E583B1D326D916">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8FF501001A16A30C4AEFB224E2570E7">Baird, Andrew H.</mods:namePart>
<mods:affiliation id="72714DF7A3163EE86C0FEF7FA91879CA">Biodiversity and Geosciences Program * Museum of Tropical Queensland * Queensland Museum Network * Townsville * QLD * Australia</mods:affiliation>
<mods:nameIdentifier id="2C293A80DA75F09E39AC91A10C923289" type="email">tom.bridge@qm.qld.gov.au</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="55439104BE157777B148F2D3D5A8A686">text</mods:typeOfResource>
<mods:relatedItem id="236B24FF445C6EB56C6025616A07CA5D" type="host">
<mods:titleInfo id="99F96E1CC798C6D092E97115BDC3BBDF">
<mods:title id="B5C42EB9F38A0CBA4E7AE47ED2823171">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="73232262FC88D7FF926FFF3B09E2E9FB">
<mods:date id="DFD0C9187619CA8993E6CC9DBF471BE9">2024</mods:date>
<mods:detail id="2E9A73661D5D4E2D1498895D3F5D76FE" type="pubDate">
<mods:number id="CF1D6A9271C4E55633EADA4FDE0B49DA">2023-07-27</mods:number>
</mods:detail>
<mods:detail id="DFF16F6F04C77568D1C5AD2A9DA5C194" type="volume">
<mods:number id="EE01D1A8FFC06ABCAF0717EB948206BA">202</mods:number>
</mods:detail>
<mods:detail id="D7F710767CFEF73F5F9EEE600067308B" type="issue">
<mods:number id="D8FC4D23F7BFF626D2A150D9C150A462">1</mods:number>
</mods:detail>
<mods:extent id="3231D6D644B23035ACF82A2AC7907C7D" unit="page">
<mods:start id="A699BE773B13D58CF3676E4EDB5CFEAB">1</mods:start>
<mods:end id="0163B0B374A203038907EE80062C0094">24</mods:end>
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</mods:relatedItem>
<mods:location id="2AD04B6850F3127DD787C283D0895493">
<mods:url id="E414C30FB6D846B58EC79A71793AFF61">https://doi.org/10.1093/zoolinnean/zlad062</mods:url>
</mods:location>
<mods:classification id="1EF5F3D7E4EE1C85C999C75944D90403">journal article</mods:classification>
<mods:identifier id="0A18E99800852C920DBA6C66075608C2" type="DOI">10.1093/zoolinnean/zlad062</mods:identifier>
<mods:identifier id="572180CBF87311E48D5D157912E4F51C" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="F4260229BF823AE0C8D7E356CA4FEDE6" type="Zenodo-Dep">14443904</mods:identifier>
</mods:mods>
<treatment id="DB4587EEFFB5412BFCDCFECBFE67FD03" LSID="urn:lsid:plazi:treatment:DB4587EEFFB5412BFCDCFECBFE67FD03" httpUri="http://treatment.plazi.org/id/DB4587EEFFB5412BFCDCFECBFE67FD03" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="1BF66573FFB54129FCDCFECBFB69FE36" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="535336F8FFB54129FCDCFECBFA56FED9" blockId="13.[821,1449,302,329]" box="[821,1449,302,329]" pageId="13" pageNumber="14">
<heading id="081B8194FFB54129FCDCFECBFA56FED9" bold="true" box="[821,1449,302,329]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="6198EAEAFFB54129FCDCFECBFA56FED9" bold="true" box="[821,1449,302,329]" pageId="13" pageNumber="14">
<taxonomicName id="94EC4D7BFFB54129FCDCFECBFA95FED8" ID-CoL="BZW7L" authority="Bonito, Bridge, Fenner &amp; Baird" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[821,1386,302,329]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="tenuissima" status="sp. nov.">
<emphasis id="6198EAEAFFB54129FCDCFECBFBE3FED8" bold="true" box="[821,1052,302,328]" italics="true" pageId="13" pageNumber="14">Acropora tenuissima</emphasis>
Bonito, Bridge, Fenner &amp; Baird
</taxonomicName>
,
<taxonomicNameLabel id="7AAB5791FFB54129FA9AFECAFA56FED9" box="[1395,1449,303,329]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="535336F8FFB54129FCC3FE8AFB69FE36" blockId="13.[810,1459,367,422]" pageId="13" pageNumber="14">
<emphasis id="6198EAEAFFB54129FCC3FE8AFBFBFE17" box="[810,1028,367,391]" italics="true" pageId="13" pageNumber="14">Zoobank registration:</emphasis>
<uri id="277D3AFAFFB54129FBFBFE8AFB69FE36" pageId="13" pageNumber="14">
urn: lsid: zoobank.org:act:
<uuid id="274A0C2DFFB54129FAD6FE8AFB69FE36" pageId="13" pageNumber="14">43573A79- 8333-4FBE-8C78-20E8837CAA10</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="1BF66573FFB5412BFCC3FE2DFE67FD03" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="535336F8FFB54129FCC3FE2DFA92FE6F" blockId="13.[810,1459,456,511]" pageId="13" pageNumber="14">
<materialsCitation id="E3843CA5FFB54129FCC3FE2DFA92FE6F" country="Australia" elevation="7" location="Myrmidon Reef" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
<emphasis id="6198EAEAFFB54129FCC3FE2DFC12FE70" box="[810,1005,456,480]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="8C57885AFFB54129FCC3FE2DFC7AFE70" box="[810,901,456,480]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
<emphasis id="6198EAEAFFB54129FC7CFE2DFC12FE70" bold="true" box="[917,1005,456,480]" italics="true" pageId="13" pageNumber="14">G78705</emphasis>
</emphasis>
* collected from
<elevation id="D8C1D1CBFFB54129FB77FE2DFAEFFE70" box="[1182,1296,456,480]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="m" value="7.0">
<quantity id="94149B1DFFB54129FB77FE2DFB35FE4F" box="[1182,1226,456,479]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="m" value="7.0">7 m</quantity>
depth
</elevation>
on the eastern fore-reef of
<location id="56336023FFB54129FC49FE02FBBDFE6F" LSID="urn:lsid:plazi:treatment:DB4587EEFFB5412BFCDCFECBFE67FD03:56336023FFB54129FC49FE02FBBDFE6F" box="[928,1090,487,511]" country="Australia" name="Myrmidon Reef" pageId="13" pageNumber="14">Myrmidon Reef</location>
* GBR*
<collectingCountry id="2BFB7668FFB54129FB66FE02FB15FE6F" box="[1167,1258,487,511]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
(
<figureCitation id="CBD72A7DFFB54129FB13FE02FABCFE6F" box="[1274,1347,487,511]" captionStart="Figure 5" captionStartId="14.[129,194,1772,1796]" captionTargetBox="[249,1353,144,1744]" captionTargetId="figure-73@14.[249,1353,144,1744]" captionTargetPageId="14" captionText="Figure 5. Acropora tenuissima. Holotype MTQ G78705 in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype." figureDoi="http://doi.org/10.5281/zenodo.14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
* B).
</materialsCitation>
</paragraph>
<paragraph id="535336F8FFB54129FCC3FDC4FB2BFDE7" blockId="13.[810,1459,544,631]" pageId="13" pageNumber="14">
<materialsCitation id="E3843CA5FFB54129FCC3FDC4FA4CFDA8" box="[810,1459,544,568]" country="Fiji" elevation="7" location="Namaqumaqua" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
<emphasis id="6198EAEAFFB54129FCC3FDC4FC6EFDA8" box="[810,913,545,568]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="8C57885AFFB54129FCC3FDC4FC73FDA8" box="[810,908,545,568]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<emphasis id="6198EAEAFFB54129FC7EFDC5FC0FFDA8" bold="true" box="[919,1008,544,568]" pageId="13" pageNumber="14">G78343</emphasis>
*
<elevation id="D8C1D1CBFFB54129FC12FDC4FBA0FDA8" box="[1019,1119,544,568]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="m" value="7.0">
<quantity id="94149B1DFFB54129FC12FDC4FBDDFDA8" box="[1019,1058,545,568]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="m" value="7.0">7 m</quantity>
depth
</elevation>
* fore-reef of
<location id="56336023FFB54129FB0CFDC5FA80FDA8" LSID="urn:lsid:plazi:treatment:DB4587EEFFB5412BFCDCFECBFE67FD03:56336023FFB54129FB0CFDC5FA80FDA8" box="[1253,1407,544,568]" country="Fiji" name="Namaqumaqua" pageId="13" pageNumber="14">Namaqumaqua</location>
*
<collectingCountry id="2BFB7668FFB54129FA64FDC5FA4CFDA8" box="[1421,1459,544,568]" name="Fiji" pageId="13" pageNumber="14">Viti</collectingCountry>
</materialsCitation>
<materialsCitation id="E3843CA5FFB54129FCC3FDA5FC0BFDC8" box="[810,1012,575,600]" collectorName="Levu" country="Fiji" location="Fiji" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
<collectorName id="FE19532EFFB54129FCC3FDA5FCA4FDC8" box="[810,859,576,600]" pageId="13" pageNumber="14">Levu</collectorName>
*
<collectingCountry id="2BFB7668FFB54129FC83FDA5FC73FDC8" box="[874,908,576,600]" name="Fiji" pageId="13" pageNumber="14">Fiji</collectingCountry>
(
<figureCitation id="CBD72A7DFFB54129FC74FDA5FC17FDC7" box="[925,1000,575,600]" captionStart="Figure 5" captionStartId="14.[129,194,1772,1796]" captionTargetBox="[249,1353,144,1744]" captionTargetId="figure-73@14.[249,1353,144,1744]" captionTargetPageId="14" captionText="Figure 5. Acropora tenuissima. Holotype MTQ G78705 in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype." figureDoi="http://doi.org/10.5281/zenodo.14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="13" pageNumber="14">Fig. 5C</figureCitation>
)
</materialsCitation>
; and
<emphasis id="6198EAEAFFB54129FBC7FDDAFB78FDC7" bold="true" box="[1070,1159,575,599]" pageId="13" pageNumber="14">G81179</emphasis>
* collected from
<materialsCitation id="E3843CA5FFB54129FAC6FDA5FB2FFDE7" collectorName="Blue Pools" country="Australia" elevation="1" location="Heron Island" pageId="13" pageNumber="14" specimenCount="1" typeStatus="paratype">
<elevation id="D8C1D1CBFFB54129FAC6FDA5FA65FDC8" box="[1327,1434,576,600]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1.0">
<quantity id="94149B1DFFB54129FAC6FDA5FAA8FDC7" box="[1327,1367,576,600]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1.0">1 m</quantity>
depth
</elevation>
at
<collectorName id="FE19532EFFB54129FCC3FDBAFC6CFDE7" box="[810,915,607,631]" pageId="13" pageNumber="14">Blue Pools</collectorName>
*
<location id="56336023FFB54129FC48FDBAFBDBFDE7" LSID="urn:lsid:plazi:treatment:DB4587EEFFB5412BFCDCFECBFE67FD03:56336023FFB54129FC48FDBAFBDBFDE7" box="[929,1060,607,631]" country="Australia" name="Heron Island" pageId="13" pageNumber="14">Heron Island</location>
* GBR*
<collectingCountry id="2BFB7668FFB54129FB9BFDBAFB2FFDE7" box="[1138,1232,607,631]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="535336F8FFB54129FCC3FD44FC73FD68" blockId="13.[810,1458,673,760]" pageId="13" pageNumber="14">
<emphasis id="6198EAEAFFB54129FCC3FD44FBCEFD29" box="[810,1073,673,697]" italics="true" pageId="13" pageNumber="14">Other material examined:</emphasis>
MTQ: GBR G30675* Davies Reef*
<collectingRegion id="9128F81AFFB54129FCC3FD25FC5DFD48" box="[810,930,704,728]" country="Australia" name="Queensland" pageId="13" pageNumber="14">Queensland</collectingRegion>
*
<collectingCountry id="2BFB7668FFB54129FC52FD25FBE5FD48" box="[955,1050,704,728]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
; G30677* Fitzroy Reef*
<collectingRegion id="9128F81AFFB54129FAD8FD25FA56FD48" box="[1329,1449,704,728]" country="Australia" name="Queensland" pageId="13" pageNumber="14">Queensland</collectingRegion>
*
<collectingCountry id="2BFB7668FFB54129FCC3FD05FC77FD68" box="[810,904,736,760]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
.
</paragraph>
<paragraph id="535336F8FFB54129FCC3FCF7FC25FB32" blockId="13.[810,1459,786,1186]" pageId="13" pageNumber="14">
<emphasis id="6198EAEAFFB54129FCC3FCF7FC72FCBA" box="[810,909,786,810]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="8C57885AFFB54129FCC3FCF7FC78FCBA" box="[810,903,786,810]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
</emphasis>
Colony shape corymbose with determinate growth form. Primary branches long (&gt;
<quantity id="94149B1DFFB54129FB48FCD4FB17FCD9" box="[1185,1256,817,841]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="cm" value="10.0">10 cm</quantity>
)* thin and terete*
<quantity id="94149B1DFFB54129FCC3FCB5FC79FCF8" box="[810,902,848,872]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
in diameter and vertically orientated (
<figureCitation id="CBD72A7DFFB54129FADAFCB5FA7BFCF8" box="[1331,1412,848,872]" captionStart="Figure 5" captionStartId="14.[129,194,1772,1796]" captionTargetBox="[249,1353,144,1744]" captionTargetId="figure-73@14.[249,1353,144,1744]" captionTargetPageId="14" captionText="Figure 5. Acropora tenuissima. Holotype MTQ G78705 in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype." figureDoi="http://doi.org/10.5281/zenodo.14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
* E). Axial corallites tubular* 1.2–2.0 mm outer diameter and 0.8– 1.0 mm inner diameter (
<figureCitation id="CBD72A7DFFB54129FBC1FC6AFB8FFC37" box="[1064,1136,911,935]" captionStart="Figure 5" captionStartId="14.[129,194,1772,1796]" captionTargetBox="[249,1353,144,1744]" captionTargetId="figure-73@14.[249,1353,144,1744]" captionTargetPageId="14" captionText="Figure 5. Acropora tenuissima. Holotype MTQ G78705 in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype." figureDoi="http://doi.org/10.5281/zenodo.14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="13" pageNumber="14">Fig. 5F</figureCitation>
). Radial corallites closely positioned but not touching* predominantly appressed tubular with extended lower lip and large oblique or nariform openings* but become labellate towards branch tips. ºepta on both axial and radial corallites are weakly developed; primary septa up to one-quarter of the radius of the calyx* secondary septa absent or some present as points (
<figureCitation id="CBD72A7DFFB54129FB65FBAEFB2DFBF3" box="[1164,1234,1099,1123]" captionStart="Figure 5" captionStartId="14.[129,194,1772,1796]" captionTargetBox="[249,1353,144,1744]" captionTargetId="figure-73@14.[249,1353,144,1744]" captionTargetPageId="14" captionText="Figure 5. Acropora tenuissima. Holotype MTQ G78705 in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype." figureDoi="http://doi.org/10.5281/zenodo.14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="13" pageNumber="14">Fig. 5F</figureCitation>
* G). Ŋe coenosteum is costate* with some spinule development in the
<typeStatus id="8C57885AFFB54129FACEFB8EFA7DFB13" box="[1319,1410,1131,1155]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
specimen (
<figureCitation id="CBD72A7DFFB54129FC92FB6CFC36FB31" box="[891,969,1161,1185]" captionStart="Figure 5" captionStartId="14.[129,194,1772,1796]" captionTargetBox="[249,1353,144,1744]" captionTargetId="figure-73@14.[249,1353,144,1744]" captionTargetPageId="14" captionText="Figure 5. Acropora tenuissima. Holotype MTQ G78705 in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype." figureDoi="http://doi.org/10.5281/zenodo.14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="13" pageNumber="14">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="535336F8FFB54129FCC3FB24FAD1FAE6" blockId="13.[810,1460,1217,1398]" pageId="13" pageNumber="14">
<emphasis id="6198EAEAFFB54129FCC3FB24FC05FB49" box="[810,1018,1217,1241]" italics="true" pageId="13" pageNumber="14">Molecular phylogeny:</emphasis>
Ŋe
<specimenCount id="45EAFD71FFB54129FBC6FB24FB37FB49" box="[1071,1224,1217,1241]" count="2" pageId="13" pageNumber="14" type="generic">two specimens</specimenCount>
sequenced of this species* from the GBR and
<collectingCountry id="2BFB7668FFB54129FBFFFB05FBCAFB68" box="[1046,1077,1248,1272]" name="Fiji" pageId="13" pageNumber="14">Fiji</collectingCountry>
* are recovered as a reciprocally monophyletic group in all phylogenetic trees (
<figureCitation id="CBD72A7DFFB54129FB3FFB1AFAEAFA88" box="[1238,1301,1279,1304]" captionStart="Figure 2" captionStartId="7.[113,178,1602,1626]" captionTargetBox="[118,1455,148,1529]" captionTargetId="graphics-75@7.[118,1276,159,1526]" captionTargetPageId="7" captionText="Figure 2. Maximum likelihood (ML) phylogeny based on the 50% complete edge-trimmed alignment matrix (2032 loci). A* inset phylogeny depicts the relationships among all six major Acropora clades (sensu Cowman et al. 2020) and the three subclades within clade I. Clade I-C is indicated by the dashed box. Ŋe position of Acropora selago (Clade VI) and Acropora donei (Clade III) are indicated. B* ML phylogeny of clade I-C. Ultrafast bootstrap support and gene/site concordance factors are indicated at the nodes (see key). Primary species hypotheses (PºHs) are indicated by clade labels. Field codes for each specimen (tip labels) also reflect the collection location: Great Barrier Reef (73-* 74-* 76-* GBR); Coral ºea (75-); Lord Howe Island* south-east Australia (81-)* Fiji (101-* FJ)* Tonga (TG); Rarotonga* Cook Islands (CI); Kimbe Bay* Bismarck ºea* Papua New Guinea (69-* KM); Palau (PL); Pohnpei (PN); Ningaloo Reef* Western Australia (88-); Ryukyu Islands* Japan (54-); Chagos (BFC); and Mayoưe (MY)." figureDoi="http://doi.org/10.5281/zenodo.14443913" httpUri="https://zenodo.org/record/14443913/files/figure.png" pageId="13" pageNumber="14">Figs 2</figureCitation>
* 3; ºupporting Information* Figs º1* º2). Ŋe species is sister to
<taxonomicName id="94EC4D7BFFB54129FAF0FAFAFA4CFAA6" authorityName="Hoffmeister" authorityYear="1925" box="[1305,1459,1311,1335]" class="Anthozoa" family="Acroporidae" genus="Acropora" isUncertain="true" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="pagoensis">
<emphasis id="6198EAEAFFB54129FAF0FAFAFACFFAA6" box="[1305,1328,1311,1334]" italics="true" pageId="13" pageNumber="14">A.</emphasis>
aff.
<emphasis id="6198EAEAFFB54129FAB1FAFAFA4CFAA6" box="[1368,1459,1311,1334]" italics="true" pageId="13" pageNumber="14">pagoensis</emphasis>
</taxonomicName>
in the
<collectionCode id="35FDAE3DFFB54129FC80FADAFC70FAC6" box="[873,911,1343,1366]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="14">ML</collectionCode>
reconstructions* but the ºNAPP species tree places
<taxonomicName id="94EC4D7BFFB54129FA75FADBFC56FAE5" authorityName="Hoffmeister" authorityYear="1925" class="Anthozoa" family="Acroporidae" genus="Acropora" isUncertain="true" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="pagoensis">
<emphasis id="6198EAEAFFB54129FA75FADBFA4BFAC5" box="[1436,1460,1342,1365]" italics="true" pageId="13" pageNumber="14">A.</emphasis>
aff.
<emphasis id="6198EAEAFFB54129FCA7FABBFC56FAE5" box="[846,937,1374,1397]" italics="true" pageId="13" pageNumber="14">pagoensis</emphasis>
</taxonomicName>
as sister species to
<taxonomicName id="94EC4D7BFFB54129FB82FABBFB3CFAE5" authorityName="Bridge &amp; Cowman &amp; Quaưrini &amp; Bonito &amp; Frederic &amp; Harii &amp; Head &amp; Hung &amp; Halafihi &amp; Rongo &amp; Baird" authorityYear="2024" box="[1131,1219,1374,1397]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="rongoi">
<emphasis id="6198EAEAFFB54129FB82FABBFB3CFAE5" box="[1131,1219,1374,1397]" italics="true" pageId="13" pageNumber="14">A. rongoi</emphasis>
</taxonomicName>
(
<figureCitation id="CBD72A7DFFB54129FB3AFAB8FAE1FAE6" box="[1235,1310,1373,1398]" captionStart="Figure 3" captionStartId="8.[129,194,876,900]" captionTargetBox="[161,1442,147,848]" captionTargetPageId="8" captionText="Figure 3. Results from multiple species delimitation methods of Clade I-C. A* ºTRUCTURE result for cluster 1 (K = 3) cluster 1i (K = 2) and cluster 1iii (K = 2) in lower panel and cluster 2 (K = 3) in upper panel. Additional ºTRUCTURE plots are available in the ºupporting Information (Fig.º4). B* principal components analysis (PCA) plots for DAPC analysis for cluster 1 in the lower panel and cluster 2 in the upper panel. DAPC clusters are indicated by shape for cluster 1 (K = 3; diamond* triangle and filled circle) and cluster 2 (K = 2; triangles and filled circles). Colours of shapes match the PºH designation from Figure 2. C* unsupervised machine learning method t-ºNE clustering of samples using Gap algorithm clustering for cluster 1 and cluster 2. Genetic clusters are indicated by shape* while colour matches the PºH designation.Cluster 1 is shown in the lower panel; alternative clustering methods and detailed labelled plots are shown in the ºupporting Information (Fig.º5). D* species tree cloudogram of Clade I-C. Cloudogram of the posterior distribution of ºNAPP trees from 2046 ºNPs mined from targeted capture data.Tip labels indicate species designated from phylogenetic and morphological assessment. (see ºupporting Information Fig. º4)." figureDoi="http://doi.org/10.5281/zenodo.14443915" httpUri="https://zenodo.org/record/14443915/files/figure.png" pageId="13" pageNumber="14">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="535336F8FFB5412BFCC3FA6AFE7CFE13" blockId="13.[810,1460,1423,1949]" lastBlockId="15.[113,763,144,387]" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<emphasis id="6198EAEAFFB54129FCC3FA6AFC77FA37" box="[810,904,1423,1447]" italics="true" pageId="13" pageNumber="14">Remarks:</emphasis>
Forms small corymbose clumps* generally
<quantity id="94149B1DFFB54129FADFFA6AFA64FA37" box="[1334,1435,1423,1447]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="13" pageNumber="14" unit="cm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 cm</quantity>
in diameter. Colonies are compact cushions with short branches in high-energy environments. In more sheltered habitats* colonies can form taller* more upright corymbose colonies. Generally* an iridescent yellow–orange to pinkish/orange colour* especially pronounced at the branch tips and axials* but can also be greyish/brown in colour. Polyps lower on the branches are generally extended during the day and brownish in colour. Ŋe species is clearly differentiated morphologically from all other species in clade I-C by its thin* neat branches. Within cluster 1i*
<taxonomicName id="94EC4D7BFFB54129FCC3F92CFC34F970" authorityName="Hoffmeister" authorityYear="1925" box="[810,971,1737,1761]" class="Anthozoa" family="Acroporidae" genus="Acropora" isUncertain="true" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="pagoensis">
<emphasis id="6198EAEAFFB54129FCC3F92CFCBEF970" box="[810,833,1737,1760]" italics="true" pageId="13" pageNumber="14">A.</emphasis>
aff.
<emphasis id="6198EAEAFFB54129FC99F92CFC34F970" box="[880,971,1737,1760]" italics="true" pageId="13" pageNumber="14">pagoensis</emphasis>
</taxonomicName>
has much thicker branches* whereas
<taxonomicName id="94EC4D7BFFB54129FABEF92CFA4CF970" authorityName="Bridge &amp; Cowman &amp; Quaưrini &amp; Bonito &amp; Frederic &amp; Harii &amp; Head &amp; Hung &amp; Halafihi &amp; Rongo &amp; Baird" authorityYear="2024" box="[1367,1459,1737,1760]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="rongoi">
<emphasis id="6198EAEAFFB54129FABEF92CFA4CF970" box="[1367,1459,1737,1760]" italics="true" pageId="13" pageNumber="14">A. rongoi</emphasis>
</taxonomicName>
has thick branches and an indeterminate growth form.
<taxonomicName id="94EC4D7BFFB54129FAB0F90DFC9DF88F" authorityName="Dana" authorityYear="1846" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="tenuis">
<emphasis id="6198EAEAFFB54129FAB0F90DFC9DF88F" italics="true" pageId="13" pageNumber="14">Acropora tenuis</emphasis>
</taxonomicName>
and
<taxonomicName id="94EC4D7BFFB54129FC7CF8EDFB19F88F" authority="Veron &amp; Wallace" authorityName="Veron &amp; Wallace" authorityYear="1984" box="[917,1254,1799,1823]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="azurea">
<emphasis id="6198EAEAFFB54129FC7CF8EDFBC8F88F" box="[917,1079,1800,1823]" italics="true" pageId="13" pageNumber="14">Acropora azurea</emphasis>
Veron &amp; Wallace
</taxonomicName>
* 1984 can co-occur with
<taxonomicName id="94EC4D7BFFB54129FC8BF8C2FC16F8AE" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[866,1001,1831,1854]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB54129FC8BF8C2FC16F8AE" box="[866,1001,1831,1854]" italics="true" pageId="13" pageNumber="14">A. tenuissima</emphasis>
</taxonomicName>
and can have similar colony shape* but
<taxonomicName id="94EC4D7BFFB54129FA75F8C2FC6FF8CE" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB54129FA75F8C2FC6FF8CE" italics="true" pageId="13" pageNumber="14">A. tenuissima</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="94EC4D7BFFB54129FB98F8A3FB3AF8CE" authorityName="Dana" authorityYear="1846" box="[1137,1221,1862,1886]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="tenuis">
<emphasis id="6198EAEAFFB54129FB98F8A3FB3AF8CE" box="[1137,1221,1862,1886]" italics="true" pageId="13" pageNumber="14">A. tenuis</emphasis>
</taxonomicName>
by its thinner branches and smaller corallites* whereas
<taxonomicName id="94EC4D7BFFB54129FB9DF883FB29F8ED" authorityName="Veron &amp; Wallace" authorityYear="1984" box="[1140,1238,1894,1917]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="azurea">
<emphasis id="6198EAEAFFB54129FB9DF883FB29F8ED" box="[1140,1238,1894,1917]" italics="true" pageId="13" pageNumber="14">A. azurea</emphasis>
</taxonomicName>
has appressed radial corallites that are nariform rather than tubular. ºpecimens of
<taxonomicName id="94EC4D7BFFB6412AFF68F8BCFEFAF8E1" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[129,261,1881,1905]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB6412AFF68F8BCFEFAF8E1" box="[129,261,1881,1905]" italics="true" pageId="14" pageNumber="15">A. tenuissima</emphasis>
</taxonomicName>
in the MTQ collection are identified as
<taxonomicName id="94EC4D7BFFB6412AFD46F8BCFF4DF800" baseAuthorityName="MNB" baseAuthorityYear="1941" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nana">
<emphasis id="6198EAEAFFB6412AFD46F8BCFF4DF800" italics="true" pageId="14" pageNumber="15">Acropora nana</emphasis>
</taxonomicName>
(ºtuder* 1878); however* the
<typeStatus id="8C57885AFFB6412AFE07F89DFDB5F800" box="[494,586,1912,1936]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
of
<taxonomicName id="94EC4D7BFFB6412AFD87F89CFF3FF820" authority="(MNB 1941)" baseAuthorityName="MNB" baseAuthorityYear="1941" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nana">
<emphasis id="6198EAEAFFB6412AFD87F89CFD42F800" box="[622,701,1913,1936]" italics="true" pageId="14" pageNumber="15">A. nana</emphasis>
(
<emphasis id="6198EAEAFFB6412AFD39F89CFF4AF820" italics="true" pageId="14" pageNumber="15">MNB 1941</emphasis>
)
</taxonomicName>
has tubular to nariform radial corallites* which extend from the branch at an obtuse angle* rather than the acute to appressed angle of
<taxonomicName id="94EC4D7BFFB6412AFC79F89CFBECF800" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[912,1043,1913,1936]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB6412AFC79F89CFBECF800" box="[912,1043,1913,1936]" italics="true" pageId="14" pageNumber="15">A. tenuissima</emphasis>
</taxonomicName>
. Ŋe coenosteum of the
<taxonomicName id="94EC4D7BFFB6412AFAF8F89CFAA0F800" baseAuthorityName="MNB" baseAuthorityYear="1941" box="[1297,1375,1913,1936]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nana">
<emphasis id="6198EAEAFFB6412AFAF8F89CFAA0F800" box="[1297,1375,1913,1936]" italics="true" pageId="14" pageNumber="15">A. nana</emphasis>
</taxonomicName>
<typeStatus id="8C57885AFFB6412AFA8FF89DFA3DF800" box="[1382,1474,1912,1936]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
also exhibits a heavily ornamented coenosteum* whereas the coenosteum of
<taxonomicName id="94EC4D7BFFB7412BFEE7FF75FE6FFF38" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[270,400,144,168]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB7412BFEE7FF75FE6FFF38" box="[270,400,144,168]" italics="true" pageId="15" pageNumber="16">A. tenuissima</emphasis>
</taxonomicName>
is costate and lacks spinules. A specimen of
<taxonomicName id="94EC4D7BFFB7412BFF3DFF55FEA8FF57" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[212,343,176,199]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB7412BFF3DFF55FEA8FF57" box="[212,343,176,199]" italics="true" pageId="15" pageNumber="16">A. tenuissima</emphasis>
</taxonomicName>
from
<collectingCountry id="2BFB7668FFB7412BFE71FF4AFE45FF57" box="[408,442,175,199]" name="Fiji" pageId="15" pageNumber="16">Fiji</collectingCountry>
is included in the series of images illustrating
<taxonomicName id="94EC4D7BFFB7412BFEFEFF2AFE78FF77" baseAuthorityName="Dana" baseAuthorityYear="1846" box="[279,391,207,231]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="subulata">
<emphasis id="6198EAEAFFB7412BFEFEFF2AFE78FF77" box="[279,391,207,231]" italics="true" pageId="15" pageNumber="16">A. subulata</emphasis>
</taxonomicName>
by
<bibRefCitation id="377D4B09FFB7412BFE44FF2AFD8FFF77" author="Veron JEN" box="[429,624,207,231]" pageId="15" pageNumber="16" refId="ref24743" refString="Veron JEN * ºtafford-ºmith MG * Turak E * DeVantier LM. Corals of the World. 2016. hup: // www. coralsossheworld. org / species _ factsheets / species _ factsheet _ summary / acropora-subulata / (28 June 2022 date last accessed)." type="book" year="2016">
Veron
<emphasis id="6198EAEAFFB7412BFE18FF2AFDDEFF77" box="[497,545,207,231]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2016)
</bibRefCitation>
hưp://www. coralsossheworld.org/media/images/0073_C04_01.jpg; however* the
<typeStatus id="8C57885AFFB7412BFF24FEE8FED6FEB5" box="[205,297,269,293]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
of
<taxonomicName id="94EC4D7BFFB7412BFEA0FEEBFE47FEB5" baseAuthorityName="Dana" baseAuthorityYear="1846" box="[329,440,269,293]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="subulata">
<emphasis id="6198EAEAFFB7412BFEA0FEEBFE47FEB5" box="[329,440,269,293]" italics="true" pageId="15" pageNumber="16">A. subulata</emphasis>
</taxonomicName>
is a fragement of a large tabular or corymbose colony with tightly packed* labellate radial corallites reminiscent of
<taxonomicName id="94EC4D7BFFB7412BFED7FEA9FE00FEF4" authorityName="Brook" authorityYear="1892" box="[318,511,332,356]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="squamosa">
<emphasis id="6198EAEAFFB7412BFED7FEA9FE00FEF4" box="[318,511,332,356]" italics="true" pageId="15" pageNumber="16">Acropora squamosa</emphasis>
</taxonomicName>
(Brook* 1892) but very distinct from
<taxonomicName id="94EC4D7BFFB7412BFF12FE89FE82FE13" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[251,381,364,387]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB7412BFF12FE89FE82FE13" box="[251,381,364,387]" italics="true" pageId="15" pageNumber="16">A. tenuissima</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="07936670FFB6412AFF68F909FD97F8AC" ID-DOI="http://doi.org/10.5281/zenodo.14443921" ID-Zenodo-Dep="14443921" httpUri="https://zenodo.org/record/14443921/files/figure.png" pageId="14" pageNumber="15" startId="14.[129,194,1772,1796]" targetBox="[249,1353,144,1744]" targetPageId="14" targetType="figure">
<paragraph id="535336F8FFB6412AFF68F909FD97F8AC" blockId="14.[129,1467,1772,1852]" pageId="14" pageNumber="15">
<emphasis id="6198EAEAFFB6412AFF68F909FE5BF894" bold="true" box="[129,420,1772,1796]" pageId="14" pageNumber="15">
Figure 5.
<taxonomicName id="94EC4D7BFFB6412AFF34F909FE5BF894" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[221,420,1772,1796]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB6412AFF34F909FE5BF894" bold="true" box="[221,420,1772,1796]" italics="true" pageId="14" pageNumber="15">Acropora tenuissima</emphasis>
</taxonomicName>
</emphasis>
. Holotype MTQ
<emphasis id="6198EAEAFFB6412AFDA2F909FD67F894" box="[587,664,1772,1796]" italics="true" pageId="14" pageNumber="15">G78705</emphasis>
in the field at Myrmidon Reef* Great Barrier Reef (A* B); paratype MTQ G78343 from Namaqumaqua* Viti Levu* Fiji (C); holotype skeletal details (D* E); and scanning electron microscope images of axial corallite (F)* radial corallites (G) and coenosteum (H) of the holotype.
</paragraph>
</caption>
<paragraph id="535336F8FFB7412BFF98FE40FDE5FE6B" blockId="15.[113,763,421,507]" pageId="15" pageNumber="16">
<emphasis id="6198EAEAFFB7412BFF98FE40FF10FE2D" box="[113,239,421,445]" italics="true" pageId="15" pageNumber="16">Distribution:</emphasis>
Currently recorded only from
<collectingCountry id="2BFB7668FFB7412BFDD9FE40FDADFE2D" box="[560,594,421,445]" name="Fiji" pageId="15" pageNumber="16">Fiji</collectingCountry>
and the centralsouthern GBR* but likely to occur elsewhere in the south-west Pacific (ºupporting Information* Fig. º6).
</paragraph>
<paragraph id="535336F8FFB7412BFF98FDF8FE67FD03" blockId="15.[113,762,541,659]" pageId="15" pageNumber="16">
<emphasis id="6198EAEAFFB7412BFF98FDF8FF1EFDA5" box="[113,225,541,565]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
Meaning ‘very thin’ in Latin
<taxonomicName id="94EC4D7BFFB7412BFDEAFDF8FD87FDA4" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[515,632,541,565]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="tenuissima">
*
<emphasis id="6198EAEAFFB7412BFDFBFDF8FD87FDA4" box="[530,632,541,564]" italics="true" pageId="15" pageNumber="16">tenuissima</emphasis>
</taxonomicName>
refers to the thin branches characteristic of this species and which distinguish it from
<taxonomicName id="94EC4D7BFFB7412BFF13FDB9FEB1FDE3" authorityName="Dana" authorityYear="1846" box="[250,334,604,627]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="tenuis">
<emphasis id="6198EAEAFFB7412BFF13FDB9FEB1FDE3" box="[250,334,604,627]" italics="true" pageId="15" pageNumber="16">A. tenuis</emphasis>
</taxonomicName>
and
<taxonomicName id="94EC4D7BFFB7412BFE69FDB9FE32FDE3" box="[384,461,603,627]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="kenti">
<emphasis id="6198EAEAFFB7412BFE69FDB9FE32FDE3" box="[384,461,603,627]" italics="true" pageId="15" pageNumber="16">A. kenti</emphasis>
</taxonomicName>
* both of which co-occur with
<taxonomicName id="94EC4D7BFFB7412BFF98FD9EFF0CFD02" authorityName="Bonito, Bridge, Fenner &amp; Baird" authorityYear="2024" box="[113,243,635,658]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="tenuissima">
<emphasis id="6198EAEAFFB7412BFF98FD9EFF0CFD02" box="[113,243,635,658]" italics="true" pageId="15" pageNumber="16">A. tenuissima</emphasis>
</taxonomicName>
across its range.
</paragraph>
</subSubSection>
</treatment>
</document>