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<mods:title id="F36AA86955DBBB168732EF67CF37D853">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<taxonomicName id="89A464A08626D831DBBFFEB6BB8AFEE5" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[988,1201,260,287]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="7CD0C3318626D831DBBFFEB6BB8AFEE5" bold="true" box="[988,1201,260,287]" italics="true" pageId="27" pageNumber="28">Loncovilius variabilis</emphasis>
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(
<figureCitation id="D69F03A68626D831DBC3FE86BCD4FEB4" box="[928,1007,308,334]" captionStart="Figure 1" captionStartId="1.[113,178,1870,1894]" captionTargetBox="[152,1420,144,1842]" captionTargetId="figure-6@1.[152,1420,144,1842]" captionTargetPageId="1" captionText="Figure 1. Habitats of Loncovilius. A, L. edwardsianus (Korge, 1963) live specimen in the Valdivian Temperate Rainforest in Chiloé, Los Lagos, Chile (Photograph: Matías Gargiulo, through iNaturalist). B, PN Puyehue, Chile, L. lividipennis was collected in mosses on the stones at river banks and under stones in accumulations along banks (photograph: Martin Fikáček). C, L. variabilis sp. nov. found in a flower of the Chiloé archipelago (photograph:Matías Gargiulo, through iNaturalist)." figureDoi="http://doi.org/10.5281/zenodo.14443947" httpUri="https://zenodo.org/record/14443947/files/figure.png" pageId="27" pageNumber="28">Figs 1C</figureCitation>
,
<figureCitation id="D69F03A68626D831DB99FE86BB20FEB4" box="[1018,1051,308,334]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." figureDoi="http://doi.org/10.5281/zenodo.14443961" httpUri="https://zenodo.org/record/14443961/files/figure.png" pageId="27" pageNumber="28">5D</figureCitation>
,
<figureCitation id="D69F03A68626D831DC46FE86BB79FEB4" box="[1061,1090,308,334]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (AE); ovipositor (FH). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="27" pageNumber="28">9B</figureCitation>
,
<figureCitation id="D69F03A68626D831DC2EFE86BBAFFEB4" box="[1101,1172,308,334]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. AC, Loncovilius germaini. DF, L. hammondi. G, H, L. impunctus. IK, L. variabilis. LN, L. edwardsianus. OQ, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="27" pageNumber="28">10IK</figureCitation>
,
<figureCitation id="D69F03A68626D831DCFEFE86BBF7FEB4" box="[1181,1228,308,334]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="27" pageNumber="28">12D</figureCitation>
,
<figureCitation id="D69F03A68626D831DCB5FE86BA3FFEB4" box="[1238,1284,308,334]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="27" pageNumber="28">13D</figureCitation>
,
<figureCitation id="D69F03A68626D831DD6DFE86BA06FEB4" box="[1294,1341,308,334]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="27" pageNumber="28">14D</figureCitation>
)
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Type material.
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male, pinned, with genitalia in a separate microvial, with labels as follows:
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, Hualaihué] Palena Pr.: Austral Highway, km 62.9 (
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S Contao turnoff),
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41°51.155
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S
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72°42.175
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[-41.85, -72.70],
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, young secondary Valdivian rainforest;/beating flowers
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<emphasis id="7CD0C3318626D831DD08FD89BCB9FD89" italics="true" pageId="27" pageNumber="28">Tepulia stipularis</emphasis>
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(myrtaceae) [
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] A. Newton and M. Ŋayer FIELD MUSEUM NAT. HIST./
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<emphasis id="7CD0C3318626D831DD25FDC8BCB9FD4B" italics="true" pageId="27" pageNumber="28">Loncovilius variabilis</emphasis>
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des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
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:
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, Vilcún] Cautín Temuco, Cherquenco [~ -38.69, -72.00] I-II.1978 L. E. Peña leg. FIELD MUS. NAT. HIST. [
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FMNH];
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: Reg. XI [
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] Queulat N.P. [~ -44.38, -72.25] P.M. Hammond/Below base 1-10.xii.03 shrub flowers/NHM/[
<specimenCount id="58A2D4AA8626D831DBB4FCE7BCC7FC96" box="[983,1020,853,877]" count="5" pageId="27" pageNumber="28" type="male">5♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DC65FCE7BB16FC96" box="[1030,1069,853,877]" count="4" pageId="27" pageNumber="28" type="female">4♀</specimenCount>
NHM;
<specimenCount id="58A2D4AA8626D831DCEBFCE7BB95FC96" box="[1160,1198,853,877]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DCDBFCE7BBE4FC96" box="[1208,1247,853,877]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
NHMD ex. NHM;
<specimenCount id="58A2D4AA8626D831DB49FCC7BC6AFC76" box="[810,849,885,909]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
CZUG ex. NHM]; [same data as above except:] Glacier Path 21.xi-12.xii.03/Malaise trap [
<specimenCount id="58A2D4AA8626D831DCF6FC26BB87FC51" box="[1173,1212,916,940]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
NHM]; [same data as above except:] Base area Beating [or] Base Camp Area 21.xi- 12.xii.03 [
<specimenCount id="58A2D4AA8626D831DBF6FC61BC85FC10" box="[917,958,979,1003]" count="8" pageId="27" pageNumber="28" type="male">8♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DBA9FC61BCCAFC10" box="[970,1009,979,1002]" count="7" pageId="27" pageNumber="28" type="female">7♀</specimenCount>
NHM];
<collectingCountry id="36B35FB38626D831DC3BFC61BB91FC11" box="[1112,1194,979,1003]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DCDCFC61BA0AFC11" box="[1215,1329,979,1003]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Hualaihué] Palena Pr.: Austral Highway, km 62.9 (
<quantity id="895CB2C68626D831DCCFFC40BBD4FBF0" box="[1196,1263,1010,1034]" metricMagnitude="3" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="28" unit="km" value="6.7">6.7km</quantity>
S Contao turnoff),
<quantity id="895CB2C68626D831DB48FBA3BC53FBD3" box="[811,872,1041,1065]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">230m</quantity>
,
<geoCoordinate id="2B9079E48626D831DB15FBA3BCDFFBD3" box="[886,996,1041,1065]" degrees="41" direction="south" minutes="51.15" orientation="latitude" pageId="27" pageNumber="28" precision="9" value="-41.8525">41°51.15S</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DB90FBA3BB57FBD3" box="[1011,1132,1041,1065]" degrees="72" direction="west" minutes="42.17" orientation="longitude" pageId="27" pageNumber="28" precision="9" value="-72.702835">72°42.17W</geoCoordinate>
[-41.85, -72.70],
<date id="3A1A39E38626D831DD4DFBA3BA8BFBD3" box="[1326,1456,1041,1065]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">23-25.i.1997</date>
, young secondary Valdivian rainforest;/beating flowers
<taxonomicName id="89A464A08626D831DD08FB83BCB9FB92" class="Magnoliopsida" family="Ericaceae" genus="Tepulia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="stipularis">
<emphasis id="7CD0C3318626D831DD08FB83BCB9FB92" italics="true" pageId="27" pageNumber="28">Tepulia stipularis</emphasis>
</taxonomicName>
(myrtaceae) [
<taxonomicName id="89A464A08626D831DC74FBE2BBB2FB92" box="[1047,1161,1104,1128]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
] A. Newton and M. Ŋayer FIELD MUSEUM NAT. HIST [
<specimenCount id="58A2D4AA8626D831DCF4FBDDBBFBFB7C" box="[1175,1216,1135,1159]" count="5" pageId="27" pageNumber="28" type="male">5♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38626D831DD5EFBDDBA4EFB7D" box="[1341,1397,1135,1159]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
[
<collectingRegion id="8C60D1C18626D831DDEAFBC2BC51FB5D" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
] Llanquihue Lago Chapo [-41.40, -72.52]
<date id="3A1A39E38626D831DD4DFB3DBA88FB5C" box="[1326,1459,1166,1191]" pageId="27" pageNumber="28" value="1985-02">febrero 1985</date>
leg. J. Zuñiga [
<specimenCount id="58A2D4AA8626D831DBDDFB1CBCDDFB3F" box="[958,998,1198,1222]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DB8FFB1CBB28FB3F" box="[1004,1043,1198,1222]" count="3" pageId="27" pageNumber="28" type="female">3♀</specimenCount>
FMNH]; HORNOHUINCO SW. Lago Chapo Lianquihue [
<collectingRegion id="8C60D1C18626D831DC64FB7CBB47FB1C" box="[1031,1148,1230,1254]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Llanquihue, -41.40, -72.52] 21,24-
<date id="3A1A39E38626D831DB0FFB5FBCF3FAFF" box="[876,968,1261,1285]" pageId="27" pageNumber="28" value="1972-12">Dic.1972</date>
Coll:L.E.Pena [
<specimenCount id="58A2D4AA8626D831DC1FFB5FBB98FAFE" box="[1148,1187,1261,1285]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DCD1FB5FBBE3FAFE" box="[1202,1240,1261,1285]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38626D831DD38FB5FBA8BFAFF" box="[1371,1456,1261,1285]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18626D831DB49FABEBCA3FADE" box="[810,920,1292,1316]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
] Rio Gol-Gol Osorno Prov. [~ -40.66, -72.18]/II-8- 57 Peña/
<collectingCountry id="36B35FB38626D831DBEFFA99BCE0FAB9" box="[908,987,1323,1347]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
[green label]/CNCI L2004-28/
<specimenCount id="58A2D4AA8626D831DD54FA9CBA6AFAB8" box="[1335,1361,1326,1346]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
/FMNH-INS 0000 024 263-265 [
<specimenCount id="58A2D4AA8626D831DC51FAF9BB60FA98" box="[1074,1115,1355,1379]" count="3" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
CNC];
<collectingCountry id="36B35FB38626D831DCDBFAF9BA31FA99" box="[1208,1290,1355,1379]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: Chiloé Pr.,
<collectingCountry id="36B35FB38626D831DDF3FAF9BA88FA99" box="[1424,1459,1355,1379]" name="Iceland" pageId="27" pageNumber="28">Isla</collectingCountry>
Chiloé, Compu Alto [-42.87, -73.70],
<date id="3A1A39E38626D831DCA2FAD8BA1AFA78" box="[1217,1313,1386,1410]" pageId="27" pageNumber="28" value="1998-01-18">18.i.1998</date>
, T. Cekalovic leg. FIELD MUS. NAT. HIST./FMNH-INS 0000 024 165-167 [
<specimenCount id="58A2D4AA8626D831DB56FA1BBC60FA3A" box="[821,859,1449,1473]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
FMNH;
<specimenCount id="58A2D4AA8626D831DBABFA1BBCD4FA3A" box="[968,1007,1449,1473]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DB98FA1BBB19FA3A" box="[1019,1058,1449,1473]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
MNNC ex. FMNH];
<collectingCountry id="36B35FB38626D831DD7CFA1BBA4AFA3B" box="[1311,1393,1449,1473]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: Prov. Chiloé:
<collectingCountry id="36B35FB38626D831DB1CFA7ABC99FA1A" box="[895,930,1480,1504]" name="Iceland" pageId="27" pageNumber="28">Isla</collectingCountry>
Chiloé, Compu Alto [-42.87, -73.70]
<date id="3A1A39E38626D831DD51FA7ABAACFA1A" box="[1330,1431,1480,1504]" pageId="27" pageNumber="28" value="1998-01-18">18-I-1998</date>
T. Cekalovic CHIL1C98 001/[Barcode] SM0722732 KUNHM-ENT [
<specimenCount id="58A2D4AA8626D831DB12F9B5BCA3F9E4" box="[881,920,1543,1567]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
SEMC]; [
<collectingCountry id="36B35FB38626D831DC64F9B5BB67F9E5" box="[1031,1116,1543,1567]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18626D831DC04F9B5BBEFF9E5" box="[1127,1236,1543,1567]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Puerto Monư] Prov. Llanquihue Correntoso [~ -41.45, -72.66]
<date id="3A1A39E38626D831DCBCF994BA42F9C4" box="[1247,1401,1574,1598]" pageId="27" pageNumber="28" value="1969-01-22">22-Enero-1969</date>
Coll: L.E.Pena [
<specimenCount id="58A2D4AA8626D831DBF7F9F7BC86F9A6" box="[916,957,1605,1629]" count="2" pageId="27" pageNumber="28" type="male">2♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38626D831DC49F9F7BB47F9A7" box="[1066,1148,1605,1629]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DCEFF9F4BBC1F9A4" box="[1164,1274,1606,1630]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Chonchi] Chiloé Pr.: Miraflores, road to (
<quantity id="895CB2C68626D831DC51F9D7BB4DF987" box="[1074,1142,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="km" value="0.6">0.6km</quantity>
W Hwy 5),
<quantity id="895CB2C68626D831DD6BF9D7BA7EF987" box="[1288,1349,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="28" unit="m" value="130.0">130m</quantity>
,
<geoCoordinate id="2B9079E48626D831DD37F9D7BC0CF966" degrees="42" direction="south" minutes="46.73" orientation="latitude" pageId="27" pageNumber="28" precision="9" value="-42.77883">
42°46.73
<emphasis id="7CD0C3318626D831DDCEF9D1BA88F986" box="[1453,1459,1635,1660]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="2B9079E48626D831DB2AF936BCE9F961" box="[841,978,1666,1692]" degrees="73" direction="west" minutes="47.71" orientation="longitude" pageId="27" pageNumber="28" precision="9" value="-73.795166">
73°47.71
<emphasis id="7CD0C3318626D831DBC0F930BC92F961" box="[931,937,1666,1691]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
W
</geoCoordinate>
[
<geoCoordinate id="2B9079E48626D831DB89F936BB70F966" box="[1002,1099,1668,1692]" degrees="42.7788" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="-42.7788">-42.7788</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DC3CF936BB85F966" box="[1119,1214,1668,1692]" degrees="73.7953" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-73.7953">-73.7953</geoCoordinate>
]
<date id="3A1A39E38626D831DCB5F936BA0EF966" box="[1238,1333,1668,1692]" pageId="27" pageNumber="28" value="1997-01-17">17.i.1997</date>
, secondary Valdivian rainforest:/beating flowering Cald-cluvia paniculata [
<taxonomicName id="89A464A08626D831DB56F971BB3CF921" box="[821,1031,1731,1755]" class="Magnoliopsida" family="Cunoniaceae" genus="Caldcluvia" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="paniculata">
<emphasis id="7CD0C3318626D831DB56F971BB3CF921" box="[821,1031,1731,1755]" italics="true" pageId="27" pageNumber="28">Caldcluvia paniculata</emphasis>
</taxonomicName>
] (
<taxonomicName id="89A464A08626D831DC7CF971BB92F921" box="[1055,1193,1731,1755]" class="Magnoliopsida" family="Cunoniaceae" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Cunoniaceae</taxonomicName>
) A.Newton and M.Ŋayer 994 FIELD MUS. NAT. HIST. [
<specimenCount id="58A2D4AA8626D831DCEBF950BB94F903" box="[1160,1199,1762,1786]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8626D831DCDAF950BBDBF903" box="[1209,1248,1762,1786]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38626D831DD38F950BA8BF900" box="[1371,1456,1762,1786]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18626D831DB49F8B0BCA2F8E0" box="[810,921,1794,1818]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Hualaihué] 11
<collectingCountry id="36B35FB38626D831DC2FF8B3BBBFF8E3" box="[1100,1156,1793,1817]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
X, peninsula Huequi under logs termas, Porcelana
<geoCoordinate id="2B9079E48626D831DC76F893BB99F8C3" box="[1045,1186,1823,1849]" degrees="42" direction="south" minutes="27" orientation="latitude" pageId="27" pageNumber="28" precision="1" seconds="30.8" value="-42.458557">
42°27
<emphasis id="7CD0C3318626D831DC32F8ADBB6CF8C2" box="[1105,1111,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
30.8
<emphasis id="7CD0C3318626D831DCE1F8ADBBB6F8C2" box="[1154,1165,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="2B9079E48626D831DCC9F893BA7AF8C2" box="[1194,1345,1823,1849]" degrees="72" direction="west" minutes="27" orientation="longitude" pageId="27" pageNumber="28" precision="1" seconds="15.0" value="-72.45416">
72°27
<emphasis id="7CD0C3318626D831DC86F8ADBBD0F8C2" box="[1253,1259,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
15.0
<emphasis id="7CD0C3318626D831DD74F8ADBA19F8C2" box="[1303,1314,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹʹ</emphasis>
W
</geoCoordinate>
[
<geoCoordinate id="2B9079E48626D831DD2CF892BA94F8C2" box="[1359,1455,1824,1848]" degrees="42.4585" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="-42.4585">-42.4585</geoCoordinate>
,
<geoCoordinate id="2B9079E48626D831DB49F8F2BCB2F8A2" box="[810,905,1856,1880]" degrees="72.4541" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-72.4541">-72.4541</geoCoordinate>
]
<quantity id="895CB2C68626D831DBFBF8F2BCF4F8AD" box="[920,975,1856,1880]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="40.0">40 m</quantity>
R. Vila leg.
<date id="3A1A39E38626D831DC30F8F2BBF9F8A2" box="[1107,1218,1856,1880]" pageId="27" pageNumber="28" value="2014-12-13">13.xii.2014</date>
/
<taxonomicName id="89A464A08626D831DCA9F8F2BA53F8AD" authority="sp" authorityName="Sp" box="[1226,1384,1856,1880]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp</taxonomicName>
det. V. Assing
<date id="3A1A39E38626D831DB16F8EDBCF1F88D" box="[885,970,1887,1911]" bridgedPair="/" pageId="27" pageNumber="28" value="2015-05">2015/V.</date>
Assing Coll. J Jenkins Shaw det. 2016 [yellow label] [
<specimenCount id="58A2D4AA8626D831DB15F8CDBCA5F86C" box="[886,926,1919,1943]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
NHMW];
<collectingCountry id="36B35FB38626D831DC7AF8CDBB50F86D" box="[1049,1131,1919,1943]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18626D831DC1EF8CDBBD6F86D" box="[1149,1261,1919,1943]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
, Puyehue National Park] Osorno Prov. Parque Nac. Puyehue,
<quantity id="895CB2C68626D831DC95F82CBA0DF84C" box="[1270,1334,1950,1974]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="27" pageNumber="28" unit="km" value="4.1">4.1km</quantity>
E Anticura [-40.66, -72.12],
<quantity id="895CB2C68621D836D956FF22BE49FF52" box="[309,370,144,168]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="28" pageNumber="29" unit="m" value="430.0">430m</quantity>
, trap site 662
<date id="3A1A39E38621D836DA6DFF22BD9AFF52" box="[526,673,144,168]" pageId="28" pageNumber="29" value="1982-12-19" valueMax="1982-12-26" valueMin="1982-12-19">19-26.xii.1982</date>
Valdivian rainfor. A.Newton and M.Ŋayer/Screen- sweeping/
<taxonomicName id="89A464A08621D836DAECFF1DBFA6FF1C" authority="sp." authorityName="Sp." class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp.</taxonomicName>
7 det. A.Newton 1984/
<specimenCount id="58A2D4AA8621D836D9F6FF60BE94FF1C" box="[405,431,210,230]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
/FMNH-INS 0000 024 163 [
<specimenCount id="58A2D4AA8621D836DA81FF7DBC32FF1C" box="[738,777,207,231]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38621D836D885FF5CBE03FEFC" box="[230,312,238,262]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18621D836D92BFF5CBE8DFEFC" box="[328,438,238,262]" country="Chile" name="Los Lagos" pageId="28" pageNumber="29">Los Lagos</collectingRegion>
, Puyehue National Park] Osorno Prov., Puyehue Nat. Pk. Aguas Calientes [-40.73, -72.30],
<quantity id="895CB2C68621D836DAA9FEBFBC3CFEDF" box="[714,775,269,293]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="m" value="500.0">500m</quantity>
,
<date id="3A1A39E38621D836D8E2FE9EBFC5FEBE" box="[129,254,300,324]" pageId="28" pageNumber="29" value="1984-12-20">20.XII.1984</date>
-
<date id="3A1A39E38621D836D966FE9EBE66FEBE" box="[261,349,300,324]" pageId="28" pageNumber="29" value="1985-02-08">8.II.1985</date>
/FMHD#85-930 Derumbes for. Trail, S.andJ. Peck, P#85-45, FIT FIELD MUSEUM NAT HIST/
<specimenCount id="58A2D4AA8621D836DA80FEFDBDC6FE99" box="[739,765,335,355]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
/ FMNH-INS 0000 024 169 [
<specimenCount id="58A2D4AA8621D836D9CBFED9BEF4FE78" box="[424,463,363,387]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
FMNH];
<collectingCountry id="36B35FB38621D836DA5DFED9BDABFE79" box="[574,656,363,387]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18621D836DAC2FEDEBC3FFE79" box="[673,772,363,387]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
] Valdivia Prov.,
<quantity id="895CB2C68621D836D943FE39BE5DFE59" box="[288,358,395,419]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="28" pageNumber="29" unit="km" value="26.0">26 km</quantity>
SE Panguipulli [-39.75, -72.15]
<quantity id="895CB2C68621D836DAAAFE39BC3EFE59" box="[713,773,395,419]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="300.0">300m</quantity>
,
<date id="3A1A39E38621D836D8E2FE18BFC6FE38" box="[129,253,426,450]" pageId="28" pageNumber="29" value="1984-12-16">16.XII.1984</date>
-
<date id="3A1A39E38621D836D966FE18BE51FE38" box="[261,362,426,450]" pageId="28" pageNumber="29" value="1985-02-11">11.II.1985</date>
/FMHD#85-921,
<taxonomicName id="89A464A08621D836DA4DFE18BD9EFE38" box="[558,677,426,450]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
remnant S.andJ. Peck, P#85-34, carrion trap FIELD MUSEUM NAT HIST/
<specimenCount id="58A2D4AA8621D836D8ABFE5EBFD9FDFA" box="[200,226,492,512]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
/FMNH-INS 0000 024 168 [
<specimenCount id="58A2D4AA8621D836DA78FE5BBD78FDFA" box="[539,579,489,513]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
FMNH];
<collectingCountry id="36B35FB38621D836DAD7FE5BBC3DFDFB" box="[692,774,489,513]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: [
<collectingRegion id="8C60D1C18621D836D8E4FDBABFDCFDDA" box="[135,231,520,544]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
] Prov. Valdivia: Parque Oncol (Cruce Sendero Tepual) Rio Cruces [-39.72, -73.27]
<date id="3A1A39E38621D836D9C5FD95BEC5FDC5" box="[422,510,551,575]" pageId="28" pageNumber="29" value="2001-01-08">8-I-2001</date>
T Cekalovic CHIL1CO1 005/[Barcode] SM0723095 KUNHM-ENT [
<specimenCount id="58A2D4AA8621D836DA1AFDF5BD9BFDA4" box="[633,672,583,607]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
SEMC]; [
<collectingCountry id="36B35FB38621D836D8E4FDD4BFE7FD84" box="[135,220,614,638]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D88BFDD4BE73FD84" box="[232,328,614,638]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
] 85/Süd-Chile 31.1.90 Puerto Puyehue [~ -40.65, -72.33] leg.G.H. Schwabe [
<specimenCount id="58A2D4AA8621D836D993FD37BD21FD66" box="[496,538,645,669]" count="2" pageId="28" pageNumber="29" type="male">2♂</specimenCount>
<specimenCount id="58A2D4AA8621D836DA46FD37BD77FD66" box="[549,588,645,669]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
SDEI]; [
<collectingCountry id="36B35FB38621D836DAD2FD37BC3DFD67" box="[689,774,645,669]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D8E2FD17BFC8FD47" box="[129,243,677,701]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836D910FD17BEDBFD47" box="[371,480,677,701]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
] Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836D8B5FD76BE74FD26" box="[214,335,708,732]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
/
<collectingCountry id="36B35FB38621D836D93BFD76BE90FD26" box="[344,427,708,732]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/
<taxonomicName id="89A464A08621D836DA55FD76BDB6FD26" box="[566,653,708,732]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Toưenham TYPE [manuscript name]/NHM [
<specimenCount id="58A2D4AA8621D836DAE2FD51BD92FD00" box="[641,681,739,763]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D8E4FCB1BFE7FCE1" box="[135,220,771,795]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D88AFCB1BE60FCE1" box="[233,347,771,795]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836D98EFCB1BD61FCE1" box="[493,602,771,795]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
] Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836D903FC90BEE2FCC0" box="[352,473,802,826]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
/
<collectingCountry id="36B35FB38621D836D982FC90BD0FFCC0" box="[481,564,802,826]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/
<taxonomicName id="89A464A08621D836DAD1FC90BC32FCC0" box="[690,777,802,826]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Toưenham PAÞTYPE [manuscript name]/NHM [
<specimenCount id="58A2D4AA8621D836D8EAFCD3BF89FC82" box="[137,178,865,889]" count="3" pageId="28" pageNumber="29" type="male">3♂</specimenCount>
<specimenCount id="58A2D4AA8621D836D8D5FCD3BFE6FC82" box="[182,221,865,889]" count="2" pageId="28" pageNumber="29" type="female">2♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D95FFCD3BEAAFC83" box="[316,401,865,889]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D9FBFCD3BD31FC83" box="[408,522,865,889]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836DAE8FCD3BDC7FC83" box="[651,764,865,889]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
]/
<typeStatus id="911FA1818621D836D8E2FC32BFE0FC62" box="[129,219,896,920]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
/Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836DA10FC32BDD0FC62" box="[627,747,896,920]" pageId="28" pageNumber="29" value="1958-11-30">30.XI.1958</date>
/C. E. Toưenham collection. B.M. 1974-587/
<collectingCountry id="36B35FB38621D836DA52FC2DBDB9FC4D" box="[561,642,927,951]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/ nigroflavus Tot [manuscript name]/NHM/PAÞTYPE
<taxonomicName id="89A464A08621D836D8E2FC6CBE68FC0C" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[129,339,990,1014]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D8E2FC6CBE68FC0C" box="[129,339,990,1014]" italics="true" pageId="28" pageNumber="29">Loncovilius variabilis</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023/NHMUK015025170 [
<specimenCount id="58A2D4AA8621D836DAEFFC4FBD88FBEE" box="[652,691,1021,1045]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D8E4FBAFBFE7FBCF" box="[135,220,1053,1077]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836D88AFBAFBE60FBCF" box="[233,347,1053,1077]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
and Chilean
<collectingCountry id="36B35FB38621D836D98EFBAFBD61FBCF" box="[493,602,1053,1077]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
] Welling Is. Pto. Eden [-49.11, -74.40]
<date id="3A1A39E38621D836D90DFB8EBEDCFBAE" box="[366,487,1084,1108]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
/
<collectingCountry id="36B35FB38621D836D993FB8EBD78FBAE" box="[496,579,1084,1108]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
Fr Kuschel/Qued. Sp. 5/
<taxonomicName id="89A464A08621D836D8DEFBE9BE5AFB89" authority="sp." authorityName="Sp." box="[189,353,1115,1139]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="30" phylum="Arthropoda" rank="genus">
Loncovilius
<taxonomicNameLabel id="67E37E4A8621D836D926FBEEBE5AFB89" box="[325,353,1116,1139]" pageId="28" pageNumber="29" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
Nov. P.M. Hammond det. 1980/NHM [Blue label]/NHMD 917069 [
<specimenCount id="58A2D4AA8621D836D9A4FBC9BED5FB68" box="[455,494,1147,1171]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHM];
<collectingCountry id="36B35FB38621D836DA30FBC9BD9EFB69" box="[595,677,1147,1171]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
:
<collectingRegion id="8C60D1C18621D836DAD1FBC9BC32FB69" box="[690,777,1147,1171]" country="Chile" name="Region Metropolitana" pageId="28" pageNumber="29">Santiago</collectingRegion>
[
<collectingRegion id="8C60D1C18621D836D8E5FB28BFDAFB48" box="[134,225,1178,1202]" country="Chile" name="Region Metropolitana" pageId="28" pageNumber="29">Santiago</collectingRegion>
Metropolitan]: El Alfalfal [~ -33.51, -70.20]
<date id="3A1A39E38621D836DAABFB28BC31FB48" box="[712,778,1178,1202]" pageId="28" pageNumber="29" value="1977-01">I.1977</date>
L.E.Peña, leg. FIELD MUS. NAT. HIST./FMNH-INS 0000 024 157162 [
<specimenCount id="58A2D4AA8621D836D97FFB6BBE78FB0A" box="[284,323,1241,1265]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8621D836D928FB6BBE49FB0A" box="[331,370,1241,1265]" count="2" pageId="28" pageNumber="29" type="female">2♀</specimenCount>
FMNH;
<specimenCount id="58A2D4AA8621D836D9B4FB6BBEC5FB0A" box="[471,510,1241,1265]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
<specimenCount id="58A2D4AA8621D836DA65FB6BBD16FB0A" box="[518,557,1241,1265]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHMD ex. FMNH;
<specimenCount id="58A2D4AA8621D836D8E2FB4ABF9CFAF5" box="[129,167,1272,1296]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
CZUG ex. FMNH;
<superScript id="B9D1B26B8621D836D91EFB44BEB9FAFE" attach="right" box="[381,386,1270,1284]" fontSize="6" pageId="28" pageNumber="29">*</superScript>
this is potentially a labelling error. Ŋe
<taxonomicName id="89A464A08621D836D8E1FAAABFCFFAD5" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[130,244,1303,1327]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D8E1FAAABFB4FAD5" box="[130,143,1304,1327]" italics="true" pageId="28" pageNumber="29">L</emphasis>
.
<emphasis id="7CD0C3318621D836D8FFFAA5BFCFFAD5" box="[156,244,1303,1327]" italics="true" pageId="28" pageNumber="29">variabilis</emphasis>
</taxonomicName>
model shown in
<figureCitation id="D69F03A68621D836D9C8FAA5BD3AFAD5" box="[427,513,1303,1327]" captionStart="Figure 3" captionStartId="11.[113,178,1776,1800]" captionTargetBox="[114,1458,144,1740]" captionTargetId="figure-6@11.[114,1458,144,1749]" captionTargetPageId="11" captionText="Figure 3. Ecological niche modelling of Loncovilius species. AD, potential suitable areas predicted with free extrapolation in South America. Binarized projection with 5% threshold for suitability values and mobility-oriented parity analysis (MOP) showing areas for which current model predictions are strictly based on extrapolation. EH, predictions under shared socio-economic pathways (SSPs) 126, and agreement between different global climate models (GCMs), and MOP agreement among GCMs with strict extrapolative conditions under the low emissions.IL, predictions under SSPs 585, agreement between different GCMs, and MOP agreement among GCMs with strict extrapolative conditions under the high emissions." figureDoi="http://doi.org/10.5281/zenodo.14443953" httpUri="https://zenodo.org/record/14443953/files/figure.png" pageId="28" pageNumber="29">Figure 3</figureCitation>
does not include this occurrence. Ŋe sets of variables and the calibration results that do include this occurrence can be consulted in the Supporting Information,
<tableCitation id="03262A988621D836D96AFAC7BE59FA77" box="[265,354,1397,1421]" captionStart="Table 1" captionStartId="3.[115,170,893,917]" captionText="Table 1.Taxon sample for the phylogenetic analysis (less column) and resulting taxonomic changes (right column)." pageId="28" pageNumber="29">Table S1</tableCitation>
, and in the Supporting Information, File S6]; [
<collectingCountry id="36B35FB38621D836D8D8FA27BE2BFA57" box="[187,272,1429,1453]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: unknown locality and date]
<collectingCountry id="36B35FB38621D836DA22FA27BD4CFA57" box="[577,631,1429,1453]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
./
<collectingCountry id="36B35FB38621D836DAE1FA27BD83FA57" box="[642,696,1429,1453]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
[Green round label]/Sharp Coll 1905-313./
<taxonomicName id="89A464A08621D836DA46FA06BDBAFA36" box="[549,641,1460,1484]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Toưenham PAÞTYPE [manuscript name]/NHM [
<specimenCount id="58A2D4AA8621D836DA81FA61BC32FA10" box="[738,777,1491,1515]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
NHM]; [
<collectingCountry id="36B35FB38621D836D8BEFA41BE09F9F1" box="[221,306,1523,1547]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: unknown locality and date]
<collectingCountry id="36B35FB38621D836DA30FA41BDBDF9F1" box="[595,646,1523,1547]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
Sharp Coll./
<taxonomicName id="89A464A08621D836D8E2F9A0BFE3F9D0" box="[129,216,1554,1578]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
nigroflavus Brnh (Typ.) [manuscript name]/BRIT. MUSEUM DON. ARROW/B. Typ. ohne Flg.d streifen/Chicago NHMus M.Bernhauer Collection/Bernhauer MS name? Current genus:
<taxonomicName id="89A464A08621D836D97EF9C2BEAEF972" box="[285,405,1648,1672]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Loncovilius</taxonomicName>
teste A.Newton 2000 [
<specimenCount id="58A2D4AA8621D836DA1DF9C2BD9EF97D" box="[638,677,1648,1672]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
FMNH]; [
<collectingCountry id="36B35FB38621D836D8E5F93DBFE0F95D" box="[134,219,1679,1703]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
: unknown locality and date] 53793/Reed/
<collectingCountry id="36B35FB38621D836DAC6F93DBDE0F95D" box="[677,731,1679,1703]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
/Fry Coll. 1905.100/Chicago NHMus M.Bernhauer Collection/
<specimenCount id="58A2D4AA8621D836DA80F900BDC6F93C" box="[739,765,1714,1734]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
/ FMNH-INS 0000 024 171 [
<specimenCount id="58A2D4AA8621D836D9D2F97CBEE3F91F" box="[433,472,1742,1766]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
FMNH]. All
<typeStatus id="911FA1818621D836DA13F97CBDEAF91C" box="[624,721,1742,1766]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
with label: PAÞTYPE
<taxonomicName id="89A464A08621D836D924F95FBD2EF8FF" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[327,533,1773,1797]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D924F95FBD2EF8FF" box="[327,533,1773,1797]" italics="true" pageId="28" pageNumber="29">Loncovilius variabilis</emphasis>
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
</paragraph>
<paragraph id="4E1B1F238621D836D8E2F8F5BD72F8A4" blockId="28.[129,585,1862,1886]" box="[129,585,1862,1886]" pageId="28" pageNumber="29">
<emphasis id="7CD0C3318621D836D8E2F8F5BFDCF8A4" box="[129,231,1863,1886]" italics="true" pageId="28" pageNumber="29">Diagnosis:</emphasis>
Same as
<taxonomicName id="89A464A08621D836D920F8F4BEA7F8A4" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[323,412,1862,1886]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318621D836D920F8F4BEA7F8A4" box="[323,412,1862,1886]" italics="true" pageId="28" pageNumber="29">variabilis</emphasis>
</taxonomicName>
group diagnosis.
</paragraph>
<paragraph id="4E1B1F238621D836D8E2F832BA58FEFC" blockId="28.[129,778,1919,1975]" lastBlockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
<emphasis id="7CD0C3318621D836D8E2F832BFCCF86D" box="[129,247,1920,1943]" italics="true" pageId="28" pageNumber="29">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8621D836D9B5F830BECBF86C" box="[470,496,1922,1942]" count="1" pageId="28" pageNumber="29" type="male"></specimenCount>
[minmax (average);
<emphasis id="7CD0C3318621D836D8E2F82DBFAFF84C" box="[129,148,1951,1974]" italics="true" pageId="28" pageNumber="29">N</emphasis>
= 10]: FBL = 2.793.19 (2.98); TL = 5.126.25 (5.61); HW = 0.720.8 (0.77); HL = 0.740.83 (0.79); HW/ HL = 0.951 (0.97); PW = 0.951.05 (1); PL = 0.880.98 (0.93); PW/PL = 1.021.09 (1.07); EW = 1.311.54 (1.38); EL = 1.171.43 (1.25); PW/HW = 1.231.33 (1.30).
</paragraph>
<paragraph id="4E1B1F238621D836DB36FEBCBB50FE38" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
Measurements
<specimenCount id="58A2D4AA8621D836DC6CFEA2BB12FEDE" box="[1039,1065,272,292]" count="1" pageId="28" pageNumber="29" type="female"></specimenCount>
[minmax (average);
<emphasis id="7CD0C3318621D836DD05FEBCBA42FEDF" box="[1382,1401,270,293]" italics="true" pageId="28" pageNumber="29">N</emphasis>
= 10]: FBL = 3.053.53 (3.33); TL = 5.57.44 (6.4); HW = 0.79 0.86 (0.83); HL = 0.820.9 (0.85); HW/HL = 0.941 (0.97); PW = 1.051.17 (1.10); PL = 11.08 (1.04); PW/PL = 1.03 1.09 (1.06); EW = 1.361.74 (1.56); EL = 1.221.55 (1.43); PW/HW = 1.291.39 (1.34).
</paragraph>
<paragraph id="4E1B1F238621D836DB36FE7BBAB0FDC5" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">Head, pronotum, and elytra black to brown-black, head darker; antennae, mouthparts, and legs yellowish-brown; abdominal tergites and sternites variable in coloration, from dark reddish-brown to brown-black with pale posterior margin.</paragraph>
<paragraph id="4E1B1F238621D836DB36FDF5BA6CFA94" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
Head from about as wide as long to, usually, slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.47), from 1.03 to 1.29 times as long as temples (in lateral view); distance between eyes about 1.25 times as long as length of eye in males and 1.34 times in females. Antennomere 3 slightly longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 1.53-1.72 times as long as a
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males and
<quantity id="895CB2C68621D836DC8CFC32BA52FC62" box="[1263,1385,896,920]" metricMagnitude="-2" metricUnit="m" metricValue="4.0259" metricValueMax="4.318" metricValueMin="3.7338" pageId="28" pageNumber="29" unit="in" value="1.585" valueMax="1.7" valueMin="1.47">1.47-1.70 in</quantity>
females. Basal puncture usually double; parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setiferous punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced to small trace near gular sutures; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with sub-rectangular sclerotized region and entire to slightly notched apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical to fusiform apical palpomere.
</paragraph>
<paragraph id="4E1B1F238621D836DB36FAC4BA12F9D0" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">Pronotum slightly wider than long, convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to 5 or even 10 times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
<paragraph id="4E1B1F238621D836DB36F983BBB1F8FF" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. Mesotarsomeres 14 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres
<quantity id="895CB2C68621D836DC7DF93DBB5DF95D" box="[1054,1126,1679,1703]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="28" pageNumber="29" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
both sexes with terminal plate. Metatarsomeres 24 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.
</paragraph>
<paragraph id="4E1B1F238621D837DB36F8BFBDF5FEBF" blockId="28.[824,1476,144,1985]" lastBlockId="29.[113,765,144,544]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Abdominal tergites III and IV with several rows of sparse and fine punctures of moderate density, without sizeable impunctate area; tergite VIII only slightly emarginate medio-apically in females. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with broad and shallow U-shaped emargination medially (
<figureCitation id="D69F03A68620D837D90AFF22BEF9FF52" box="[361,450,144,168]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="29" pageNumber="30">Fig. 14D</figureCitation>
); female sternite VIII without emargination; male sternite IX without emargination medially (apex subtruncate), its basal portion moderately longer than distal portion; male tergite X with anterior margin subtruncate (
<figureCitation id="D69F03A68620D837D81FFEBFBFEDFEDF" box="[124,214,269,293]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="29" pageNumber="30">Fig. 12D</figureCitation>
); female tergite X apically truncate (
<figureCitation id="D69F03A68620D837DA3DFEBFBD83FEDF" box="[606,696,269,293]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="29" pageNumber="30">Fig. 13D</figureCitation>
); ovipositor, its second gonocoxite with one macroseta medially.
</paragraph>
<paragraph id="4E1B1F238620D837D8EEFEFEBD4AFDDA" blockId="29.[113,765,144,544]" pageId="29" pageNumber="30">
Aedeagus as in
<figureCitation id="D69F03A68620D837D94FFEFEBE8EFE9E" box="[300,437,332,356]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. AC, Loncovilius germaini. DF, L. hammondi. G, H, L. impunctus. IK, L. variabilis. LN, L. edwardsianus. OQ, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="29" pageNumber="30">Figure 10IK</figureCitation>
; its total length ~ 0.95. Median lobe rod-like, apex not emarginate; internal sac with median vertical sclerite and pair of short I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe; wider than median lobe; converging into a subrounded apex, slightly projected in the middle; with dark peg setae; in lateral view paramere almost straight, slightly concave apically.
</paragraph>
<paragraph id="4E1B1F238620D837D812FDF3BE83FCEF" blockId="29.[113,763,577,789]" pageId="29" pageNumber="30">
<emphasis id="7CD0C3318620D837D812FDF3BFDAFDA3" box="[113,225,577,601]" italics="true" pageId="29" pageNumber="30">Etymology:</emphasis>
Ŋe species epithet is the Latin adjective
<taxonomicName id="89A464A08620D837DAF6FDF3BDCFFDA3" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[661,756,577,601]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="7CD0C3318620D837DAF8FDF3BDCFFDA3" box="[667,756,577,601]" italics="true" pageId="29" pageNumber="30">variabilis</emphasis>
</taxonomicName>
(variable) and refers to a notable intraspecific morphological variation of this taxon. Ŋis variation can occur between individuals from the same locality, e.g. it can be completely blackishbrown forms and others with a reddish-brown abdomen. Ŋe head punctation also varies, and loss or duplication of paraocular and basal punctures is common.
</paragraph>
<paragraph id="4E1B1F238620D837D812FC85BE22FB93" blockId="29.[113,764,823,1129]" pageId="29" pageNumber="30">
<emphasis id="7CD0C3318620D837D812FC85BD62FCB5" box="[113,601,823,847]" italics="true" pageId="29" pageNumber="30">
Distribution and habitat:
<taxonomicName id="89A464A08620D837D91CFC85BD62FCB5" authorityName="Reyes-Hernández &amp; Hansen &amp; Shaw &amp; Solodovnikov" authorityYear="2024" box="[383,601,823,847]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="variabilis">Loncovilius variabilis</taxonomicName>
</emphasis>
is known only from
<collectingCountry id="36B35FB38620D837D8D3FCE4BFD7FC94" box="[176,236,854,878]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
, from the following regions:
<collectingRegion id="8C60D1C18620D837DA54FCE4BD9EFC94" box="[567,677,854,878]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837DAD7FCE4BDCDFC94" box="[692,758,854,878]" country="Chile" name="Aysen" pageId="29" pageNumber="30">Aysén</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837D812FCC4BFDEFC74" box="[113,229,886,910]" country="Chile" name="Los Lagos" pageId="29" pageNumber="30">Los Lagos</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837D896FCC4BE62FC77" box="[245,345,885,909]" country="Chile" name="Los Rios" pageId="29" pageNumber="30">Los Ríos</collectingRegion>
,
<collectingRegion id="8C60D1C18620D837D90AFCC7BED8FC77" box="[361,483,885,909]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="29" pageNumber="30">Magallanes</collectingRegion>
, and Chilean
<collectingCountry id="36B35FB38620D837DAE6FCC7BDCDFC77" box="[645,758,885,909]" name="Antarctica" pageId="29" pageNumber="30">Antarctica</collectingCountry>
, and from Santiago Metropolitan. The species occurs from lowlands to
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of elevation in the mountains; confined to various
<typeStatus id="911FA1818620D837D88FFC66BE1EFC16" box="[236,293,980,1004]" pageId="29" pageNumber="30">types</typeStatus>
of forests in the Magellanic subpolar and Valdivian temperate forest ecoregions. It has been collected using Malaise traps, and directly from flowers, e.g. from
<taxonomicName id="89A464A08620D837D812FB83BE03FBB3" box="[113,312,1073,1097]" class="Magnoliopsida" family="Myrtaceae" genus="Tepualia" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="stipularis">
<emphasis id="7CD0C3318620D837D812FB83BE03FBB3" box="[113,312,1073,1097]" italics="true" pageId="29" pageNumber="30">Tepualia stipularis</emphasis>
</taxonomicName>
(
<taxonomicName id="89A464A08620D837D92DFB83BEFEFBB3" box="[334,453,1073,1097]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) and
<taxonomicName id="89A464A08620D837DA71FB83BDC1FBB3" box="[530,762,1073,1097]" class="Magnoliopsida" family="Cunoniaceae" genus="Caldcluvia" kingdom="Plantae" order="Oxalidales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="paniculata">
<emphasis id="7CD0C3318620D837DA71FB83BDC1FBB3" box="[530,762,1073,1097]" italics="true" pageId="29" pageNumber="30">Caldcluvia paniculata</emphasis>
</taxonomicName>
(
<taxonomicName id="89A464A08620D837D81AFBE3BE36FB93" box="[121,269,1105,1129]" class="Magnoliopsida" family="Cunoniaceae" kingdom="Plantae" order="Oxalidales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Cunoniaceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>