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<document id="A0E205D1C4A067188DC52571393AABD4" ID-DOI="10.1093/zoolinnean/zlad143" ID-ISSN="0024-4082" ID-Zenodo-Dep="14443945" ID-ZooBank="CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734089919291" checkinUser="plazi" docAuthor="Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins &amp; Solodovnikov, Alexey" docDate="2024" docId="C60DAE358625D833D893FC49BFEDF9A4" docLanguage="en" docName="zlad143.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlad143" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Loncovilius germaini" docType="treatment" docVersion="2" lastPageNumber="26" masterDocId="3A34D64D863DD82AD863FFB2BF3BFFFA" masterDocTitle="Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="25" updateTime="1734090470362" updateUser="ExternalLinkService">
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<mods:title id="3D9CB471CDE907BE8ABD6603F769E9FB">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="E810DE035F75D78CAEAC06C008E5BA0B">Reyes-Hernández, José L.</mods:namePart>
<mods:affiliation id="17CB9547DA9B8978EBAA50D6E47CFCEF">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
<mods:nameIdentifier id="D1D6D5A0B78CE33750AD76B0B11837BE" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BD3064C702EC4496CF5F0FEAAC9B8604">Hansen, Aslak Kappel</mods:namePart>
<mods:affiliation id="B5E7DD52572C645D7CAFC267B0589ED9">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark &amp; Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:namePart id="2D085A6C5F99FFF64A832CBE55A8FD64">Shaw, Josh Jenkins</mods:namePart>
<mods:affiliation id="2C5531906F1DFB9DF728E899033F8A89">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="F67EA44EE71DC79C7ABAF7BBBE0254AD">Solodovnikov, Alexey</mods:namePart>
<mods:affiliation id="34809C0872D7E10B595FF15590958CC7">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:title id="B1C02B037BBAB8019467F86322D8537B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="7332102BB02DA8B765DF57E467E11D38">2024</mods:date>
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<mods:number id="8EFB80FC45B6C9504F9C89FFC4295199">2023-10-28</mods:number>
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<mods:number id="9334CCBE3B266527DE86E41C50B5BED8">202</mods:number>
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<treatment id="C60DAE358625D833D893FC49BFEDF9A4" LSID="urn:lsid:plazi:treatment:C60DAE358625D833D893FC49BFEDF9A4" httpUri="http://treatment.plazi.org/id/C60DAE358625D833D893FC49BFEDF9A4" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="06BE4CA88625D832D893FC49BDA1FBEF" box="[240,666,1019,1046]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="4E1B1F238625D832D893FC49BDA1FBEF" blockId="24.[240,666,1019,1046]" box="[240,666,1019,1046]" pageId="24" pageNumber="25">
<heading id="1553A84F8625D832D893FC49BDA1FBEF" bold="true" box="[240,666,1019,1046]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="89A464A08625D832D893FC49BDA1FBEF" ID-CoL="3VYHQ" authority="(Scheerpeltz, 1933)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[240,666,1019,1046]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318625D832D893FC49BDA1FBEF" bold="true" box="[240,666,1019,1046]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D893FC49BEF9FBEC" bold="true" box="[240,450,1019,1046]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
(Scheerpeltz, 1933)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="06BE4CA88625D833D963FB99BFEDF9A4" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="4E1B1F238625D832D963FB99BDB1FBBF" blockId="24.[256,650,1067,1093]" box="[256,650,1067,1093]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D963FB99BDB1FBBF" bold="true" box="[256,650,1067,1093]" pageId="24" pageNumber="25">
(
<figureCitation id="D69F03A68625D832D969FB99BE61FBBF" box="[266,346,1067,1093]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." figureDoi="http://doi.org/10.5281/zenodo.14443961" httpUri="https://zenodo.org/record/14443961/files/figure.png" pageId="24" pageNumber="25">Figs 5A</figureCitation>
,
<figureCitation id="D69F03A68625D832D907FB99BEB8FBBF" box="[356,387,1067,1093]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="24" pageNumber="25">9A</figureCitation>
,
<figureCitation id="D69F03A68625D832D9EEFB99BEE0FBBF" box="[397,475,1067,1093]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="24" pageNumber="25">10A–C</figureCitation>
,
<figureCitation id="D69F03A68625D832D987FB99BD29FBBF" box="[484,530,1067,1093]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="24" pageNumber="25">12A</figureCitation>
,
<figureCitation id="D69F03A68625D832DA7FFB99BD71FBBF" box="[540,586,1067,1093]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="24" pageNumber="25">13A</figureCitation>
,
<figureCitation id="D69F03A68625D832DA30FB99BD44FBBF" box="[595,639,1067,1093]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="24" pageNumber="25">14A</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238625D832D8E2FBD3BDF4FA69" blockId="24.[129,779,1121,1428]" pageId="24" pageNumber="25">
Germain 1903: 412 (as
<taxonomicName id="89A464A08625D832D914FBD3BD6DFB83" box="[375,598,1121,1145]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="7CD0C3318625D832D914FBD3BD6DFB83" box="[375,598,1121,1145]" italics="true" pageId="24" pageNumber="25">Philonthus cribripennis</emphasis>
</taxonomicName>
, original description); Bernhauer and Schubert 1914: 332 (as
<taxonomicName id="89A464A08625D832DA24FB33BC38FB63" authorityName="Bernhauer &amp; K. Schubert" authorityYear="1914" box="[583,771,1153,1177]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318625D832DA24FB33BC38FB63" box="[583,771,1153,1177]" italics="true" pageId="24" pageNumber="25">Philonthus chilensis</emphasis>
</taxonomicName>
, replacement name); Scheerpeltz 1933: 1344 (as
<taxonomicName id="89A464A08625D832DAC2FB12BFECFB2D" authorityName="Scheerpeltz" authorityYear="1933" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318625D832DAC2FB12BFECFB2D" italics="true" pageId="24" pageNumber="25">Philonthus germaini</emphasis>
</taxonomicName>
, replacement name); Coiffait and Sáiz 1966: 406 (as
<taxonomicName id="89A464A08625D832D8E2FB6DBEF5FB0D" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[129,462,1247,1271]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
<emphasis id="7CD0C3318625D832D8E2FB6DBFD4FB0D" box="[129,239,1247,1271]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318625D832D962FB52BE5EFB0D" box="[257,357,1248,1271]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
)
<emphasis id="7CD0C3318625D832D91BFB52BEF5FB0D" box="[376,462,1248,1271]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
</taxonomicName>
, characters); Coiffait and Sáiz 1968: 365 [as
<taxonomicName id="89A464A08625D832D974FB4CBD5AFAEC" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[279,609,1278,1302]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
<emphasis id="7CD0C3318625D832D974FB4CBEBEFAEC" box="[279,389,1278,1302]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318625D832D9F6FB4DBEC2FAEC" box="[405,505,1279,1302]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
)
<emphasis id="7CD0C3318625D832DA69FB4DBD5AFAEC" box="[522,609,1279,1302]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
</taxonomicName>
<emphasis id="7CD0C3318625D832DA01FB4DBD53FAEC" box="[610,616,1279,1302]" italics="true" pageId="24" pageNumber="25">;</emphasis>
checklist]; Sáiz 1971: 385 [as
<taxonomicName id="89A464A08625D832D978FAAFBD52FACF" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[283,617,1309,1334]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
<emphasis id="7CD0C3318625D832D978FAAFBEB2FACF" box="[283,393,1309,1333]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
(
<emphasis id="7CD0C3318625D832D9FFFAACBD3BFACF" box="[412,512,1310,1333]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
)
<emphasis id="7CD0C3318625D832DA70FAACBD52FACF" box="[531,617,1310,1333]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
</taxonomicName>
; listed without new data]; Herman 2001a: 26 (as
<taxonomicName id="89A464A08625D832D9B2FA8FBDA9FAAF" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[465,658,1341,1365]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318625D832D9B2FA8FBDA9FAAF" box="[465,658,1341,1365]" italics="true" pageId="24" pageNumber="25">Loncovilius chilensis</emphasis>
</taxonomicName>
, nomenclature); Herman 2001b: 3083 (as
<taxonomicName id="89A464A08625D832D9AAFAEEBDB0FA8E" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[457,651,1372,1396]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318625D832D9AAFAEEBDB0FA8E" box="[457,651,1372,1396]" italics="true" pageId="24" pageNumber="25">Loncovilius chilensis</emphasis>
</taxonomicName>
, catalogue); Newton 2017: 22 (as
<taxonomicName id="89A464A08625D832D93EFAC9BD1DFA69" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[349,550,1403,1427]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318625D832D93EFAC9BD1DFA69" box="[349,550,1403,1427]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
</taxonomicName>
, nomenclature).
</paragraph>
<paragraph id="4E1B1F238625D832D8E2FA02BDA9FA32" blockId="24.[129,658,1456,1480]" box="[129,658,1456,1480]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D8E2FA02BE7BFA32" box="[129,320,1456,1480]" italics="true" pageId="24" pageNumber="25">Material eoamined:</emphasis>
Supporting Information, File S7.
</paragraph>
<paragraph id="4E1B1F238625D832D8E2FA6CBC3FF94A" blockId="24.[128,778,1501,1713]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D8E2FA6CBFDEFA0F" box="[129,229,1502,1525]" italics="true" pageId="24" pageNumber="25">Diagnosis:</emphasis>
Head dorsally without microsculpture; gula without isodiametric microsculpture in the middle; females with a11 distinctly longer than a10 (a11:a10 ratio &lt;1.5). Elytra, including epipleura, evenly setose and with several coarse non-setiferous punctures. Protibiae sexually dimorphic, with laterodorsal row of thick spines only present in females. Tergite VIII with medial apical emargination only in females. Aedeagus as in
<figureCitation id="D69F03A68625D832DA0EF92BBDC5F94B" box="[621,766,1689,1713]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="24" pageNumber="25">Figure 10A–C</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238625D832D8E2F97BBECEF887" blockId="24.[129,779,1736,1980]" pageId="24" pageNumber="25">
<emphasis id="7CD0C3318625D832D8E2F97BBFCCF91A" box="[129,247,1737,1760]" italics="true" pageId="24" pageNumber="25">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8625D832D9CBF979BEF9F925" box="[424,450,1739,1759]" count="1" pageId="24" pageNumber="25" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318625D832DAD9F97BBDF6F91A" box="[698,717,1737,1760]" italics="true" pageId="24" pageNumber="25">N</emphasis>
= 6]: FBL = 2.9–3.44 (3.19); TL = 5.6–6.25 (6.01); HW = 0.87– 1.01 (0.94); HL = 0.76–0.86 (0.82); HW/HL = 1.12–1.2 (1.14); PW = 1.11–1.29 (1.21); PL = 0.97–1.18 (1.08); PW/ PL = 1.1–1.14 (1.12); EW = 1.4–1.64 (1.52); EL = 1.16–1.4 (1.3); PW/HW = 1.26–1.33 (1.29).
</paragraph>
<paragraph id="4E1B1F238625D832D8FFF837BB65FD57" blockId="24.[129,779,1736,1980]" lastBlockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">
Measurements
<specimenCount id="58A2D4AA8625D832D954F835BE6AF861" box="[311,337,1927,1947]" count="1" pageId="24" pageNumber="25" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318625D832DA52F837BD7FF866" box="[561,580,1925,1948]" italics="true" pageId="24" pageNumber="25">N</emphasis>
= 6]: FBL = 3.36– 3.87 (3.6); TL = 7.31–7.77 (7.54); HW = 0.96–1.12 (1.03); HL = 0.85–0.95 (0.91); HW/HL = 1.11–1.18 (1.13); PW = 1.25–1.48 (1.34); PL = 1.13–1.28 (1.2); PW/PL = 1.1– 1.16 (1.12); EW = 1.5–1.88 (1.68); EL = 1.38–1.64 (1.49); PW/HW = 1.26–1.33 (1.3).
</paragraph>
<paragraph id="4E1B1F238625D832DB36FD06BBADFCD0" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">Head black to brown-black; mouthparts, antennae, pronotum, legs, disc of elytra, and abdominal segments dark reddish-brown; epipleura, abdominal tergites, and sternites with posterior margin light reddish-brown.</paragraph>
<paragraph id="4E1B1F238625D832DB36FC83BA8EF9E0" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">
Head transverse; dorsally and ventrally glossy with few micropunctures, dorsally without microsculpture, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.46), from 1.4 to 1.86 times longer than the length of temples (in lateral view); distance between eyes about 1.68 times as long as length of the eye in males and 1.8 times in females. Antennal insertions situated closer to frontoclypeus than to eye; antennomeres 2 and 3 subequal in length; antennomeres 5 to 10 subequal in length; antennomere 11 from 1.53 to 1.69 times longer than antennomere
<quantity id="895CB2C68625D832DCE2FBD9BB8DFB79" box="[1153,1206,1131,1155]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="25" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68625D832DD02FBD9BC75FB58" metricMagnitude="-2" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="24" pageNumber="25" unit="in" value="1.55" valueMax="1.6" valueMin="1.5">1.5 to 1.6 in</quantity>
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and entire apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical.
</paragraph>
<paragraph id="4E1B1F238625D832DB36F990BAF8F92C" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">Pronotum transverse, convex, evenly curved, usually with one PPDS with additional paired punctures adjacent to AMRSP (APP), without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra evenly setose and with several coarse non-setiferous punctures.Mesosternumwithfourmacrosetaearrangedinarowmedially.</paragraph>
<paragraph id="4E1B1F238625D832DB36F96CBAF8F84B" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate. Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4withpaleadhesivesetae,withoutterminalplate.</paragraph>
<paragraph id="4E1B1F238624D833D8EEFF22BFE8FDDA" blockId="25.[113,763,144,576]" pageId="25" pageNumber="26">
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line (
<figureCitation id="D69F03A68624D833D81FFF7DBFF1FF1D" box="[124,202,207,231]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="25" pageNumber="26">Fig. 9A</figureCitation>
); in females tergite VIII emarginate medio-apically. Sternite III with basal transverse carina converging evenly at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with deep V-shaped emargination medially (
<figureCitation id="D69F03A68624D833D910FEFEBEF4FE9E" box="[371,463,332,356]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="25" pageNumber="26">Fig. 14A</figureCitation>
); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X subtruncate medially (
<figureCitation id="D69F03A68624D833DA5CFE18BDA1FE38" box="[575,666,426,450]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="25" pageNumber="26">Fig. 12A</figureCitation>
); female tergite X with U-shaped emargination medially (
<figureCitation id="D69F03A68624D833DAE8FE7BBDD3FE1B" box="[651,744,457,481]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="25" pageNumber="26">Fig. 13A</figureCitation>
); ovipositor, its second gonocoxite without macrosetae medially (
<figureCitation id="D69F03A68624D833D81FFDBABFF9FDDA" box="[124,194,520,544]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." figureDoi="http://doi.org/10.5281/zenodo.14443975" httpUri="https://zenodo.org/record/14443975/files/figure.png" pageId="25" pageNumber="26">Fig. 9F</figureCitation>
).
</paragraph>
<paragraph id="4E1B1F238624D833D8EEFD95BD8BFDC5" blockId="25.[113,763,144,576]" box="[141,688,551,576]" pageId="25" pageNumber="26">
Aedeagus. As in
<figureCitation id="D69F03A68624D833D950FD95BEFEFDC5" box="[307,453,551,575]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="25" pageNumber="26">Figure 10A–C</figureCitation>
; its total length ~ 1.21.
</paragraph>
<paragraph id="4E1B1F238624D833D812FDD3BFD7FCCE" blockId="25.[113,764,609,821]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833D812FDD3BD6DFD82" box="[113,598,609,633]" italics="true" pageId="25" pageNumber="26">
Distribution and habitat:
<taxonomicName id="89A464A08624D833D9E4FDD3BD6DFD82" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[391,598,609,633]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">Loncovilius germaini</taxonomicName>
</emphasis>
is known only from
<collectingCountry id="36B35FB38624D833D8C9FD32BFDFFD62" box="[170,228,640,664]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
, from the
<collectingRegion id="8C60D1C18624D833D933FD32BE81FD62" box="[336,442,640,664]" country="Chile" name="Araucania" pageId="25" pageNumber="26">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18624D833D9A6FD32BD3DFD62" box="[453,518,640,664]" country="Chile" name="Aysen" pageId="25" pageNumber="26">Aysén</collectingRegion>
, and
<collectingRegion id="8C60D1C18624D833DA5EFD32BD9CFD62" box="[573,679,640,664]" country="Chile" name="Los Lagos" pageId="25" pageNumber="26">Los Lagos</collectingRegion>
regions where it occurs in the eastern portion of the Valdivian temperate forest ecoregion, from
<quantity id="895CB2C68624D833D9D2FD0CBD6EFD2C" box="[433,597,702,726]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.45" metricValueMin="0.75" pageId="25" pageNumber="26" unit="m" value="1100.0" valueMax="1450.0" valueMin="750.0">750 to 1450 m</quantity>
of elevation. It has been collected by sissing leaf liưer and moss, also by using Malaise and window traps, as well as carrion and dung baited pitfall traps.
</paragraph>
<paragraph id="4E1B1F238624D833D812FCE4BFEDF9A4" blockId="25.[112,763,854,1630]" pageId="25" pageNumber="26">
<emphasis id="7CD0C3318624D833D812FCE4BE86FC94" box="[113,445,854,878]" italics="true" pageId="25" pageNumber="26">
Remarks:
<taxonomicName id="89A464A08624D833D889FCE4BE86FC94" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[234,445,854,878]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">Loncovilius germaini</taxonomicName>
</emphasis>
was described as
<taxonomicName id="89A464A08624D833DAF2FCE4BFDEFC77" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="7CD0C3318624D833DAF2FCE4BFDEFC77" italics="true" pageId="25" pageNumber="26">Philonthus cribripennis</emphasis>
</taxonomicName>
based on an unspecified number of
<typeStatus id="911FA1818624D833DA2FFCC4BD9CFC74" box="[588,679,886,910]" pageId="25" pageNumber="26" type="syntype">syntypes</typeStatus>
from an uncertain type locality in
<collectingCountry id="36B35FB38624D833D915FC27BE8AFC57" box="[374,433,917,941]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
. Ŋis name was found to be preoccupied and replaced by another preoccupied name
<taxonomicName id="89A464A08624D833DAF3FC07BE96FC11" authority="Bernhauer and Schubert 1914" authorityName="Bernhauer and Schubert" authorityYear="1914" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318624D833DAF3FC07BDA5FC36" box="[656,670,949,972]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="7CD0C3318624D833DAC9FC06BDC1FC36" box="[682,762,948,972]" italics="true" pageId="25" pageNumber="26">chilensis</emphasis>
Bernhauer and Schubert 1914
</taxonomicName>
; the laưer was replaced again by
<taxonomicName id="89A464A08624D833D811FC41BE93FBF1" authority="Scheerpeltz, 1933" authorityName="Scheerpeltz" authorityYear="1933" box="[114,424,1011,1035]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318624D833D811FC41BFBBFBF0" box="[114,128,1011,1034]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="7CD0C3318624D833D8EDFC41BFDFFBF0" box="[142,228,1011,1034]" italics="true" pageId="25" pageNumber="26">germaini</emphasis>
Scheerpeltz, 1933
</taxonomicName>
. Coiffiat and Sáiz (1966, 1968) and Sáiz (1971) treated
<taxonomicName id="89A464A08624D833D91BFBA1BED0FBD0" authorityName="Scheerpeltz" authorityYear="1933" box="[376,491,1042,1066]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="7CD0C3318624D833D91BFBA1BEBCFBD0" box="[376,391,1043,1066]" italics="true" pageId="25" pageNumber="26">L</emphasis>
.
<emphasis id="7CD0C3318624D833D9F6FBA1BED0FBD0" box="[405,491,1043,1066]" italics="true" pageId="25" pageNumber="26">germaini</emphasis>
</taxonomicName>
as a member of the subgenus
<taxonomicName id="89A464A08624D833D8D7FB80BE22FBB3" box="[180,281,1074,1097]" pageId="25" pageNumber="26" rank="subGenus" subGenus="Lienturius">
<emphasis id="7CD0C3318624D833D8D7FB80BE22FBB3" box="[180,281,1074,1097]" italics="true" pageId="25" pageNumber="26">Lienturius</emphasis>
</taxonomicName>
of the genus
<taxonomicName id="89A464A08624D833D9C6FB83BD2FFBB3" authorityName="Germain" authorityYear="1903" box="[421,532,1073,1097]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318624D833D9C6FB83BD2FFBB3" box="[421,532,1073,1097]" italics="true" pageId="25" pageNumber="26">Loncovilius</emphasis>
</taxonomicName>
without adding much data about morphology or distribution of this species, for which even the aedeagus hitherto remained not illustrated. Coiffait and Sáiz (1966) mentioned that the species was known from a female
<typeStatus id="911FA1818624D833D8AEFB1DBE11FB3D" box="[205,298,1199,1223]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
kept in the collection of the Natural History Museum of
<collectingCountry id="36B35FB38624D833D88DFB7CBE12FB1C" box="[238,297,1230,1254]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
. At the same time, Camousseight (1980) did not list this species among the type material in Germain’s collection in this museum. We did not find any material that could be interpreted as types of
<emphasis id="7CD0C3318624D833D9EDFA9FBD33FABE" box="[398,520,1325,1348]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="89A464A08624D833D9EDFA9FBD3FFABE" authorityName="Scheerpeltz" authorityYear="1933" box="[398,516,1325,1348]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">L. germaini</taxonomicName>
.
</emphasis>
Given the uncertainty about type material in the original description and the catalogue by Camousseight (1980), the type status of the female listed by Coiffait and Sáiz (1966) as ‘holotype’ must be verified in the future. For now, we have matched the identity of this species based on the description by Coiffait and Sáiz (1966). As explained by Newton (2017), Herman’s (2001a) interpretation of both names as replacements for only
<taxonomicName id="89A464A08624D833DA79F9BABD9DF9E5" box="[538,678,1543,1567]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cribripennis">
<emphasis id="7CD0C3318624D833DA79F9BABD9DF9E5" box="[538,678,1543,1567]" italics="true" pageId="25" pageNumber="26">P. cribripennis</emphasis>
</taxonomicName>
and his resurrection of
<taxonomicName id="89A464A08624D833D973F995BE4CF9C5" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[272,375,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="7CD0C3318624D833D973F995BE4CF9C5" box="[272,375,1575,1599]" italics="true" pageId="25" pageNumber="26">P. chilensis</emphasis>
</taxonomicName>
as the valid name for this species was incorrect.
</paragraph>
</subSubSection>
</treatment>
</document>
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