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<document id="5B55E39BC7605091CB84A19790A1E1B8" ID-DOI="10.1093/zoolinnean/zlad143" ID-ISSN="0024-4082" ID-Zenodo-Dep="14443945" ID-ZooBank="CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734089919291" checkinUser="plazi" docAuthor="Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins &amp; Solodovnikov, Alexey" docDate="2024" docId="C60DAE358623D80ADBD3FA56BEF8FE79" docLanguage="en" docName="zlad143.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlad143" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Loncovilius edwardsianus" docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="3A34D64D863DD82AD863FFB2BF3BFFFA" masterDocTitle="Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="31" updateTime="1734090470362" updateUser="ExternalLinkService">
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<mods:title id="F23898388D448D9147EFE19C78B04020">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="6FF0CA478791D5EFAFBA6E3584F55C53">Reyes-Hernández, José L.</mods:namePart>
<mods:affiliation id="DFC998B35C7F02754B4023630AA4BC49">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
<mods:nameIdentifier id="DA791B111E1BCC75E4F60F976E66FC9A" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C1C520165C17C496E4BCE6213CDF41B7">Hansen, Aslak Kappel</mods:namePart>
<mods:affiliation id="563BBE9210C7086A99ABC66D8516172D">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark &amp; Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:namePart id="01BF3076F4FBD3139DD5DEE7DAB2A617">Shaw, Josh Jenkins</mods:namePart>
<mods:affiliation id="637DBDFEC8FC1C9995974F64C7BA7ABE">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="E0DD92B9B1309D4198048C1AB8441AFB">Solodovnikov, Alexey</mods:namePart>
<mods:affiliation id="AE9B487C15E6E03CD9AD340463B39CAD">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:title id="EAEE82C2B0BE380D73AA4ECFCC03E537">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="B42C3E2C8E672F2D67897DB9C5134DD6">2024</mods:date>
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<mods:title id="E6BBD4CCF5EBE8DC81171DA8CBEEE58D">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="84843A913EA7F67D8D17E3E83B82AF3E">2023-10-28</mods:number>
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<mods:identifier id="B4652B2D7C3AD5EFED725E211D554DC0" type="ISSN">0024-4082</mods:identifier>
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<treatment id="C60DAE358623D80ADBD3FA56BEF8FE79" LSID="urn:lsid:plazi:treatment:C60DAE358623D80ADBD3FA56BEF8FE79" httpUri="http://treatment.plazi.org/id/C60DAE358623D80ADBD3FA56BEF8FE79" lastPageId="32" lastPageNumber="32" pageId="30" pageNumber="31">
<subSubSection id="06BE4CA88623D834DBD3FA56BA77FA04" box="[944,1356,1508,1535]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="4E1B1F238623D834DBD3FA56BA77FA04" blockId="30.[944,1356,1508,1535]" box="[944,1356,1508,1535]" pageId="30" pageNumber="31">
<heading id="1553A84F8623D834DBD3FA56BA77FA04" bold="true" box="[944,1356,1508,1535]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="89A464A08623D834DBD3FA56BA77FA04" ID-CoL="3VYHP" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[944,1356,1508,1535]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DBD3FA56BA77FA04" bold="true" box="[944,1356,1508,1535]" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834DBD3FA56BB94FA05" bold="true" box="[944,1199,1508,1535]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsianus</emphasis>
(Korge, 1963)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="06BE4CA88623D80ADBDEF9ACBEF8FE79" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="4E1B1F238623D834DBDEF9ACBA04F9C2" blockId="30.[957,1343,1566,1592]" box="[957,1343,1566,1592]" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834DBDEF9ACBA04F9C2" bold="true" box="[957,1343,1566,1592]" pageId="30" pageNumber="31">
(
<figureCitation id="D69F03A68623D834DBA4F9ACBB2DF9C2" box="[967,1046,1566,1592]" captionStart="Figure 1" captionStartId="1.[113,178,1870,1894]" captionTargetBox="[152,1420,144,1842]" captionTargetId="figure-6@1.[152,1420,144,1842]" captionTargetPageId="1" captionText="Figure 1. Habitats of Loncovilius. A, L. edwardsianus (Korge, 1963) live specimen in the Valdivian Temperate Rainforest in Chiloé, Los Lagos, Chile (Photograph: Matías Gargiulo, through iNaturalist). B, PN Puyehue, Chile, L. lividipennis was collected in mosses on the stones at river banks and under stones in accumulations along banks (photograph: Martin Fikáček). C, L. variabilis sp. nov. found in a flower of the Chiloé archipelago (photograph:Matías Gargiulo, through iNaturalist)." figureDoi="http://doi.org/10.5281/zenodo.14443947" httpUri="https://zenodo.org/record/14443947/files/figure.png" pageId="30" pageNumber="31">Figs 1A</figureCitation>
,
<figureCitation id="D69F03A68623D834DC43F9ACBB7BF9C2" box="[1056,1088,1566,1592]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." figureDoi="http://doi.org/10.5281/zenodo.14443963" httpUri="https://zenodo.org/record/14443963/files/figure.png" pageId="30" pageNumber="31">6A</figureCitation>
,
<figureCitation id="D69F03A68623D834DC29F9ACBBACF9C2" box="[1098,1175,1566,1592]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="30" pageNumber="31">10L–N</figureCitation>
,
<figureCitation id="D69F03A68623D834DCC2F9ACBBF7F9C2" box="[1185,1228,1566,1592]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="30" pageNumber="31">12E</figureCitation>
,
<figureCitation id="D69F03A68623D834DCB5F9ACBA3AF9C2" box="[1238,1281,1566,1592]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="30" pageNumber="31">13E</figureCitation>
,
<figureCitation id="D69F03A68623D834DD69F9ACBA0FF9C2" box="[1290,1332,1566,1592]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="30" pageNumber="31">14E</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4E1B1F238623D834DB5AF9EABC82F975" blockId="30.[825,1475,1623,1898]" pageId="30" pageNumber="31">
=
<taxonomicName id="89A464A08623D834DB2BF9E5BBE4F995" authority="(Bernhauer, 1935)" baseAuthorityName="Bernhauer" baseAuthorityYear="1935" box="[840,1247,1623,1647]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318623D834DB2BF9E5BB2FF995" box="[840,1044,1623,1647]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsi</emphasis>
(Bernhauer, 1935)
</taxonomicName>
nec
<taxonomicName id="89A464A08623D834DD70F9E5BC8EF975" authority="Sharp, 1886" authorityName="Bernh. Nec Sharp" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318623D834DD70F9E5BAF9F995" box="[1299,1474,1623,1647]" italics="true" pageId="30" pageNumber="31">Quedius edwardsi</emphasis>
Sharp, 1886
</taxonomicName>
.
</paragraph>
<paragraph id="4E1B1F238623D834DB36F924BB13F890" blockId="30.[825,1475,1623,1898]" pageId="30" pageNumber="31">
Bernhauer, 1935: 92 (as
<taxonomicName id="89A464A08623D834DC38F924BA31F954" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[1115,1290,1686,1710]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318623D834DC38F924BA31F954" box="[1115,1290,1686,1710]" italics="true" pageId="30" pageNumber="31">Quedius edwardsi</emphasis>
</taxonomicName>
, original description); Korge, 1963 (as
<taxonomicName id="89A464A08623D834DC28F907BA15F937" authorityName="Korge" authorityYear="1963" box="[1099,1326,1717,1741]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DC28F907BA15F937" box="[1099,1326,1717,1741]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
</taxonomicName>
, replacement name); Coiffait and Sáiz 1966: 413 (
<taxonomicName id="89A464A08623D834DCBFF967BA87F917" authorityName="Korge" authorityYear="1963" box="[1244,1468,1749,1773]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DCBFF967BA87F917" box="[1244,1468,1749,1773]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
</taxonomicName>
, redescription without new material or data); Coiffait and Sáiz, 1968: 365 (listed as
<taxonomicName id="89A464A08623D834DC72F8A1BBD6F8D1" authorityName="Korge" authorityYear="1963" box="[1041,1261,1811,1835]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DC72F8A1BBD6F8D1" box="[1041,1261,1811,1835]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
</taxonomicName>
, without new data); Newton 2017: 28 (as
<taxonomicName id="89A464A08623D834DC53F881BA16F8B1" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1072,1325,1843,1867]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318623D834DC53F881BA16F8B1" box="[1072,1325,1843,1867]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsianus</emphasis>
</taxonomicName>
, moved from
<taxonomicName id="89A464A08623D834DB5AF8E0BCB2F890" box="[825,905,1874,1898]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318623D834DB5AF8E0BCB2F890" box="[825,905,1874,1898]" italics="true" pageId="30" pageNumber="31">Quedius</emphasis>
</taxonomicName>
to
<taxonomicName id="89A464A08623D834DBCAF8E0BB2CF890" box="[937,1047,1874,1898]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318623D834DBCAF8E0BB2CF890" box="[937,1047,1874,1898]" italics="true" pageId="30" pageNumber="31">Loncovilius</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="4E1B1F238623D835DB5AF83EBE45FD87" blockId="30.[825,1474,1931,1987]" lastBlockId="31.[113,763,144,638]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="7CD0C3318623D834DB5AF83EBB7BF859" box="[825,1088,1931,1955]" italics="true" pageId="30" pageNumber="31">
Type material.
<typeStatus id="911FA1818623D834DBBFF83EBB00F859" box="[988,1083,1932,1955]" pageId="30" pageNumber="31" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
(here designated): male, pinned, with genitalia in a separate microvial, with labels as follows: S.
<collectingCountry id="36B35FB38622D835D812FF22BF95FF52" box="[113,174,144,168]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
: Llanquihue prov. F. and M. Edwards. B.M. 1927–63/ Casa Pangue [-41.04, -71.87]
<date id="3A1A39E38622D835D9A2FF1DBD7DFF3D" box="[449,582,175,199]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</date>
./Bernhauer det. [unreadable]/Chicago NHM M. Bernhauer collection/
<taxonomicName id="89A464A08622D835D812FF5CBE5CFEFC" authority="Brh." authorityName="Bernhauer" authorityYear="2024" box="[113,359,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi" status="sp. nov.">Quedius edwardsi Brh.</taxonomicName>
<taxonomicNameLabel id="67E37E4A8622D835D913FF5CBE93FEFC" box="[368,424,238,262]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
/
<taxonomicName id="89A464A08622D835D9D3FF5CBDB0FEFC" authority="Bernhauer" authorityName="Bernhauer" authorityYear="2024" box="[432,651,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">Edwardsi Bernhauer</taxonomicName>
<taxonomicName id="89A464A08622D835DAF5FF5CBDC9FEFC" box="[662,754,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
/
<typeStatus id="911FA1818622D835D812FEBFBE3EFEDF" box="[113,261,269,293]" pageId="31" pageNumber="32" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="89A464A08622D835D976FEBFBDF8FEDF" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[277,707,269,293]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318622D835D976FEBFBD2DFEDF" box="[277,534,269,293]" italics="true" pageId="31" pageNumber="32">Loncovilius edwardsianus</emphasis>
(Korge, 1963)
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
<emphasis id="7CD0C3318622D835D931FEFEBEE7FE9E" box="[338,476,332,356]" italics="true" pageId="31" pageNumber="32">
<typeStatus id="911FA1818622D835D931FEFEBEECFE9E" box="[338,471,332,356]" pageId="31" pageNumber="32" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
male, pinned, with genitalia aưached to the card, with labels as follows: S.
<collectingCountry id="36B35FB38622D835DA5CFED9BD47FE79" box="[575,636,363,387]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
: Llanquihue prov. F. and M. Edwards. B. M. 1927–63/Casa Pangue [-41.04, -71.87]
<date id="3A1A39E38622D835D8B4FE18BE64FE38" box="[215,351,426,450]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</date>
./
<taxonomicName id="89A464A08622D835D908FE18BEF2FE38" box="[363,457,426,450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
<taxonomicName id="89A464A08622D835D983FE18BDB5FE38" authority="Bernh" authorityName="Bernhauer" authorityYear="2024" box="[480,654,426,450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi" status="sp. nov.">
<emphasis id="7CD0C3318622D835D983FE18BD0DFE38" box="[480,566,426,450]" italics="true" pageId="31" pageNumber="32">edwardsi</emphasis>
Bernh
</taxonomicName>
<taxonomicNameLabel id="67E37E4A8622D835DAC4FE18BDCAFE38" box="[679,753,426,450]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
/
<typeStatus id="911FA1818622D835D812FE7BBE3EFE1B" box="[113,261,457,481]" pageId="31" pageNumber="32" type="lectotype">LECTOTYPE</typeStatus>
C.E.Toưenham of
<taxonomicName id="89A464A08622D835D9A6FE7BBDC9FE1B" authority="Bernh. Nec Sharp" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[453,754,457,481]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">Q.edwardsi Bernh.Nec Sharp</taxonomicName>
/
<taxonomicName id="89A464A08622D835D812FE5BBFF2FDFB" box="[113,201,489,513]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
ultor n. n. Toưenham/edwardsi repl. By
<taxonomicName id="89A464A08622D835DA13FE5BBEAFFDDA" authority="Korge 1963 M. K. Nayer" authorityName="Korge" authorityYear="1963" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">edwardsianus Korge 1963 M. K. Ŋayer</taxonomicName>
det. 2005/NHMUK015024785/ PAÞLECTOTYPE
<taxonomicName id="89A464A08622D835D930FD95BDC1FDC5" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[339,762,551,575]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318622D835D930FD95BD6AFDC5" box="[339,593,551,575]" italics="true" pageId="31" pageNumber="32">Loncovilius edwardsianus</emphasis>
(Korge, 1963)
</taxonomicName>
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023. Deposited at NHM.
</paragraph>
<paragraph id="4E1B1F238622D835D812FD2DBD85FD4D" blockId="31.[113,702,671,695]" box="[113,702,671,695]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835D812FD2DBE57FD4D" box="[113,364,671,695]" italics="true" pageId="31" pageNumber="32">Other material eoamined:</emphasis>
Supporting Information, File S7.
</paragraph>
<paragraph id="4E1B1F238622D835D812FD6BBD1CFC97" blockId="31.[113,763,729,878]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835D812FD6BBFECFD0A" box="[113,215,729,752]" italics="true" pageId="31" pageNumber="32">Diagnosis:</emphasis>
Nuchal ridge absent dorsally. Pronotum outline hexagonal. Elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence. Protibiae without sexual dimorphism, both males and females without laterodorsal row of thick spines. Aedeagus as in
<figureCitation id="D69F03A68622D835D9ECFCE4BD1AFC94" box="[399,545,854,878]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="31" pageNumber="32">Figure 10L–N</figureCitation>
.
</paragraph>
<paragraph id="4E1B1F238622D835D812FC21BEC9FBBD" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835D812FC21BFD3FC50" box="[113,232,915,938]" italics="true" pageId="31" pageNumber="32">Description:</emphasis>
Measurements
<specimenCount id="58A2D4AA8622D835D9FAFC24BE88FC50" box="[409,435,918,938]" count="1" pageId="31" pageNumber="32" type="male">♂</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318622D835DAC8FC21BD85FC50" box="[683,702,915,938]" italics="true" pageId="31" pageNumber="32">N</emphasis>
= 6]: FBL = 3.95–4.21 (4.1); TL = 7.92–8 (7.95); HW = 1–1.11 (1.05); HL = 1.13–1.25 (1.19); HW/HL = 0.83–0.92 (0.88); PW = 1.15–1.24 (1.21); PL = 1.15–1.25 (1.21); PW/ PL = 0.95–1.03 (1); EW = 1.76–1.9 (1.81); EL = 1.67–1.71 (1.69); PW/HW = 1.11–1.21 (1.15).
</paragraph>
<paragraph id="4E1B1F238622D835D8EEFBFDBE98FAF9" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">
Measurements
<specimenCount id="58A2D4AA8622D835D94BFBE0BE79FB9C" box="[296,322,1106,1126]" count="1" pageId="31" pageNumber="32" type="female">♀</specimenCount>
[min–max (average);
<emphasis id="7CD0C3318622D835DA41FBFDBD0EFB9C" box="[546,565,1103,1126]" italics="true" pageId="31" pageNumber="32">N</emphasis>
= 3]: FBL = 4.47– 4.76 (4.59); TL = 7.32–8.56 (7.76); HW = 1.16–1.21 (1.18); HL = 1.3–1.36 (1.33); HW/HL = 0.87–0.9 (0.89); PW = 1.33–1.5 (1.4); PL = 1.31–1.44 (1.37); PW/PL = 1.01– 1.04 (1.02); EW = 1.98–2.13 (2.06); EL = 1.83–1.96 (1.9); PW/HW = 1.15–1.24 (1.18).
</paragraph>
<paragraph id="4E1B1F238622D835D8EEFAB9BFFBFA25" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">Head and pronotum dark brown to black with metallic blue overtones; antennae, mouthparts, pronotum at margins and legs yellowish-brown; elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence, sometimes darker around mesocutellum and at suture; abdominal tergites and sternites brown-black or dark reddish-brown with pale posterior margin.</paragraph>
<paragraph id="4E1B1F238622D835D8EEFA54BCA5FE1B" blockId="31.[113,765,915,1973]" lastBlockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
Head elongate; dorsally and ventrally with iridescence and with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with moderate setiferous punctures of medium and small-size. Eyes small (EYL/HL = x = 0.44), from 1.18 to 1.46 times as long as temples (in lateral view) in males and
<quantity id="895CB2C68622D835D8C2F910BE00F940" box="[161,315,1698,1722]" metricMagnitude="-2" metricUnit="m" metricValue="2.3876" metricValueMax="2.6416" metricValueMin="2.1336" pageId="31" pageNumber="32" unit="in" value="0.94" valueMax="1.04" valueMin="0.84">0.84 to 1.04 in</quantity>
females; distance between eyes about 1.13 times as long as length of eye in males and 1.44 times in females. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.52 to 1.75 times as long as a
<quantity id="895CB2C68622D835DAAAF8ADBDC0F8CD" box="[713,763,1823,1847]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="31" pageNumber="32" unit="in" value="10.0">10 in</quantity>
males and from
<quantity id="895CB2C68622D835D970F88DBEACF8AD" box="[275,407,1855,1879]" metricMagnitude="-2" metricUnit="m" metricValue="3.3401" metricValueMax="3.5559999999999996" metricValueMin="3.1242" pageId="31" pageNumber="32" unit="in" value="1.315" valueMax="1.4" valueMin="1.23">1.23 to 1.4 in</quantity>
females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, ending distinctly far from gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
</paragraph>
<paragraph id="4E1B1F238622D835DB26FE5BBB8AFD46" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">Pronotum about as wide as long, convex, hexagonal, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or even up to six times as long as, diameter of punctures. Mesosternum with six macrosetae arranged in a row medially. Metasternum with several macrosetae.</paragraph>
<paragraph id="4E1B1F238622D835DB26FD76BAA1FC62" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 1–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
<paragraph id="4E1B1F238622D835DB26FC2DBB02FAEA" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina usually evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with U-shaped emargination medially (
<figureCitation id="D69F03A68622D835DC9EFB8EBA6FFBAE" box="[1277,1364,1084,1108]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443985" httpUri="https://zenodo.org/record/14443985/files/figure.png" pageId="31" pageNumber="32">Fig. 14E</figureCitation>
); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X apically subtruncate (
<figureCitation id="D69F03A68622D835DB56FB0BBCB0FB2B" box="[821,907,1209,1233]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443979" httpUri="https://zenodo.org/record/14443979/files/figure.png" pageId="31" pageNumber="32">Fig. 12E</figureCitation>
); female tergite X with apical margin distinctly subacute to acute (
<figureCitation id="D69F03A68622D835DBA5FB6BBB21FB0B" box="[966,1050,1241,1265]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." figureDoi="http://doi.org/10.5281/zenodo.14443981" httpUri="https://zenodo.org/record/14443981/files/figure.png" pageId="31" pageNumber="32">Fig. 13E</figureCitation>
); ovipositor, its second gonocoxite with three macrosetae medially.
</paragraph>
<paragraph id="4E1B1F238622D835DB26FAA5BA66FA11" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
Aedeagus as in
<figureCitation id="D69F03A68622D835DB91FAA5BBB0FAD5" box="[1010,1163,1303,1327]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." figureDoi="http://doi.org/10.5281/zenodo.14443977" httpUri="https://zenodo.org/record/14443977/files/figure.png" pageId="31" pageNumber="32">Figure 10L–N</figureCitation>
; its total length 1.25–1.31. Median lobe rod-like, its apex in parameral view slightly V-shaped, emarginate, internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a round apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically.
</paragraph>
<paragraph id="4E1B1F238622D835DB49F9BFBCF3F905" blockId="31.[810,1460,1549,1792]" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835DB49F9BFBA21F9DF" box="[810,1306,1549,1573]" italics="true" pageId="31" pageNumber="32">
Distribution and habitat:
<taxonomicName id="89A464A08622D835DC47F9BFBA21F9DF" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1060,1306,1549,1573]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">Loncovilius edwardsianus</taxonomicName>
</emphasis>
is known from
<collectingCountry id="36B35FB38622D835DB49F99EBC5FF9BE" box="[810,868,1580,1604]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
, from the
<collectingRegion id="8C60D1C18622D835DBB0F99EBB07F9BE" box="[979,1084,1580,1604]" country="Chile" name="Araucania" pageId="31" pageNumber="32">Araucanía</collectingRegion>
,
<collectingRegion id="8C60D1C18622D835DC2BF99EBBB3F9BE" box="[1096,1160,1580,1604]" country="Chile" name="Aysen" pageId="31" pageNumber="32">Aysén</collectingRegion>
,
<collectingRegion id="8C60D1C18622D835DCF7F99EBBE1F9BE" box="[1172,1242,1580,1604]" country="Chile" name="Biobio" pageId="31" pageNumber="32">Biobío</collectingRegion>
,
<collectingRegion id="8C60D1C18622D835DC85F99EBA69F9BE" box="[1254,1362,1580,1604]" country="Chile" name="Los Lagos" pageId="31" pageNumber="32">Los Lagos</collectingRegion>
, and
<collectingRegion id="8C60D1C18622D835DDEEF99EBC62F999" country="Chile" name="Los Rios" pageId="31" pageNumber="32">Los Ríos</collectingRegion>
regions. In addition, it occurs in the westernmost portions of the
<collectingRegion id="8C60D1C18622D835DB0DF9D9BC85F979" box="[878,958,1643,1667]" country="Argentina" name="Chubut" pageId="31" pageNumber="32">Chubut</collectingRegion>
and
<collectingRegion id="8C60D1C18622D835DB91F9D9BB65F979" box="[1010,1118,1643,1667]" country="Argentina" name="Rio Negro" pageId="31" pageNumber="32">Río Negro</collectingRegion>
provinces of
<collectingCountry id="36B35FB38622D835DC89F9D9BA69F979" box="[1258,1362,1643,1667]" name="Argentina" pageId="31" pageNumber="32">Argentina</collectingCountry>
. Ŋe species occurs from
<quantity id="895CB2C68622D835DBBCF938BB46F95B" box="[991,1149,1674,1698]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="31" pageNumber="32" unit="m" value="700.0" valueMax="1200.0" valueMin="200.0">200 to 1200 m</quantity>
elevation; it is reported from wet sclerophyll and open
<taxonomicName id="89A464A08622D835DC53F91BBB9BF93B" box="[1072,1184,1705,1729]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
<emphasis id="7CD0C3318622D835DC53F91BBB9BF93B" box="[1072,1184,1705,1729]" italics="true" pageId="31" pageNumber="32">Nothofagus</emphasis>
</taxonomicName>
forests along the Valdivian temperate forest ecoregion. It has been collected by sissing leaf liưer and moss.
</paragraph>
<paragraph id="4E1B1F238622D80ADB49F893BEF8FE79" blockId="31.[809,1459,1825,1975]" lastBlockId="32.[127,780,144,387]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="7CD0C3318622D835DB49F893BBBAF8C3" box="[810,1153,1825,1849]" italics="true" pageId="31" pageNumber="32">
Remarks:
<taxonomicName id="89A464A08622D835DBEDF893BBBAF8C3" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[910,1153,1825,1849]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">Loncovilius edwardsianus</taxonomicName>
</emphasis>
was described under the preoccupied name
<taxonomicName id="89A464A08622D835DBA5F8F3BB56F8A3" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[966,1133,1857,1881]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="7CD0C3318622D835DBA5F8F3BB56F8A3" box="[966,1133,1857,1881]" italics="true" pageId="31" pageNumber="32">Quedius edwardsi</emphasis>
</taxonomicName>
based on an unspecified number of
<typeStatus id="911FA1818622D835DB22F8D2BCA1F882" box="[833,922,1888,1912]" pageId="31" pageNumber="32" type="syntype">syntypes</typeStatus>
from Casa Pangue in Llanquihue Province in Southern
<collectingCountry id="36B35FB38622D835DB4AF8CDBC59F86D" box="[809,866,1919,1943]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
. Korge (1963) replaced the name with
<taxonomicName id="89A464A08622D835DC84F832BAB8F86D" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1255,1411,1919,1943]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
<emphasis id="7CD0C3318622D835DC84F832BAB8F86D" box="[1255,1411,1919,1943]" italics="true" pageId="31" pageNumber="32">Q. edwardsianus</emphasis>
</taxonomicName>
and, interestingly, this species was treated in the genus
<taxonomicName id="89A464A08622D835DD22F82DBAABF84D" box="[1345,1424,1951,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C3318622D835DD22F82DBAABF84D" box="[1345,1424,1951,1975]" italics="true" pageId="31" pageNumber="32">Quedius</emphasis>
</taxonomicName>
by Coiffait and Sáiz (1966, 1968) who neither studied the type material nor reported any new material. Presumably, all mentioned authors knew this species only from literature because even Sáiz (1971) did not mention it in the fauna of
<collectingCountry id="36B35FB3861DD80ADA44FF5CBD5BFEFC" box="[551,608,238,262]" name="Chile" pageId="32" pageNumber="33">Chile</collectingCountry>
. Ŋe species was transferred to the genus
<taxonomicName id="89A464A0861DD80AD908FEBFBEEDFEDF" box="[363,470,269,293]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7CD0C331861DD80AD908FEBFBEEDFEDF" box="[363,470,269,293]" italics="true" pageId="32" pageNumber="33">Loncovilius</emphasis>
</taxonomicName>
much later by Newton (2017). To fix the identity of the species, a male from the Bernhauer’s collection at the FMNH, which is clearly a
<typeStatus id="911FA181861DD80ADA60FEFEBD6FFE9E" box="[515,596,332,356]" pageId="32" pageNumber="33" type="syntype">syntype</typeStatus>
, is here designated as a
<typeStatus id="911FA181861DD80AD8CBFED9BE3EFE79" box="[168,261,363,387]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
of
<emphasis id="7CD0C331861DD80AD947FEDEBEF8FE79" box="[292,451,363,387]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="89A464A0861DD80AD947FEDEBE85FE79" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[292,446,363,387]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="edwardsianus">L. edwardsianus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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