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<document id="2AFF1C4F2CD3A734537DE6648E8431CC" ID-CLB-Dataset="28026" ID-DOI="10.11646/zootaxa.4512.1.1" ID-GBIF-Dataset="26e12e1b-2175-4007-871b-d0b73122e5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607575" ID-ZooBank="E752FB7B-F34C-4D12-B8A2-EA6C791DD6C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586644813" checkinUser="plazi" docAuthor="Kaiser, Christine M., Kaiser, Hinrich &amp; OShea, Mark" docDate="2018" docId="C80EBE29FFD4FFE3FF75F8C20B4BFE2E" docLanguage="en" docName="zootaxa.4512.1.1.pdf" docOrigin="Zootaxa 4512 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Coronella rosenbergii Bleeker 1860" docType="treatment" docVersion="8" lastPageNumber="21" masterDocId="3437C651FFC7FFF6FFE2FE470F12FFFE" masterDocTitle="The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="20" updateTime="1698713152511" updateUser="ExternalLinkService">
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<mods:title id="1F4CFCDBF7494A464AFB5F1F924BB347">The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)</mods:title>
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<treatment id="C80EBE29FFD4FFE3FF75F8C20B4BFE2E" ID-DOI="http://doi.org/10.5281/zenodo.5997363" ID-GBIF-Taxon="154496305" ID-Zenodo-Dep="5997363" LSID="urn:lsid:plazi:treatment:C80EBE29FFD4FFE3FF75F8C20B4BFE2E" httpUri="http://treatment.plazi.org/id/C80EBE29FFD4FFE3FF75F8C20B4BFE2E" lastPageId="21" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="08BD5CB4FFD4FFE5FF75F8C20DCBF91F" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="40180F3FFFD4FFE5FF75F8C20D97F961" blockId="19.[151,645,1669,1695]" box="[151,645,1669,1695]" pageId="19" pageNumber="20">
<heading id="1B50B853FFD4FFE5FF75F8C20D97F961" bold="true" box="[151,645,1669,1695]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<treatmentCitationGroup id="60B72811FFD4FFE5FF75F8C20D97F961" box="[151,645,1669,1695]" pageId="19" pageNumber="20">
<taxonomicName id="87A774BCFFD4FFE5FF75F8C20D97F961" authority="Bleeker, 1860 b: 37" authorityName="Bleeker" authorityPageNumber="37" authorityYear="1860" box="[151,645,1669,1695]" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="rosenbergii">
<emphasis id="72D3D32DFFD4FFE5FF75F8C20D97F961" bold="true" box="[151,645,1669,1695]" pageId="19" pageNumber="20">
<emphasis id="72D3D32DFFD4FFE5FF75F8C20EB2F961" bold="true" box="[151,416,1669,1695]" italics="true" pageId="19" pageNumber="20">Coronella rosenbergii</emphasis>
<treatmentCitation id="C106292EFFD4FFE5FE45F8C20D97F961" author="Bleeker, P." box="[423,645,1669,1695]" page="37" pageId="19" pageNumber="20" year="1860">
<bibRefCitation id="243672CEFFD4FFE5FE45F8C20D72F961" author="Bleeker, P." box="[423,608,1669,1695]" pageId="19" pageNumber="20" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleeker, 1860b</bibRefCitation>
:37
</treatmentCitation>
</emphasis>
</taxonomicName>
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</heading>
</paragraph>
<paragraph id="40180F3FFFD4FFE5FF75F88E0DCBF91F" blockId="19.[151,1436,1737,2029]" box="[151,729,1737,1762]" pageId="19" pageNumber="20">
<emphasis id="72D3D32DFFD4FFE5FF75F88E0E7CF91C" bold="true" box="[151,366,1737,1762]" pageId="19" pageNumber="20">Taxonomic status.</emphasis>
Junior synonym of
<taxonomicName id="87A774BCFFD4FFE5FDACF88D0DC0F91C" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[590,722,1737,1762]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="modestus">
<emphasis id="72D3D32DFFD4FFE5FDACF88D0DC0F91C" box="[590,722,1737,1762]" italics="true" pageId="19" pageNumber="20">S. modestus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFD4FFE5FF25F8AC0C7EF835" pageId="19" pageNumber="20" type="description">
<paragraph id="40180F3FFFD4FFE5FF25F8AC0E86F8FD" blockId="19.[151,1436,1737,2029]" box="[199,404,1771,1796]" pageId="19" pageNumber="20">
<emphasis id="72D3D32DFFD4FFE5FF25F8AC0E5BF8FA" bold="true" box="[199,329,1771,1796]" pageId="19" pageNumber="20">Synonyms.</emphasis>
None.
</paragraph>
<paragraph id="40180F3FFFD4FFE5FF25F94B0C7EF835" blockId="19.[151,1436,1737,2029]" pageId="19" pageNumber="20">
<emphasis id="72D3D32DFFD4FFE5FF25F94B0E6FF8DB" bold="true" box="[199,381,1804,1829]" pageId="19" pageNumber="20">Original name.</emphasis>
<taxonomicName id="87A774BCFFD4FFE5FE66F94B0C45F8DB" authority="Rosenbergii Bleeker 1860 b: 37" authorityName="Rosenbergii Bleeker" authorityPageNumber="37" authorityYear="1860" box="[388,855,1804,1829]" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFD4FFE5FE66F94B0D91F8DB" box="[388,643,1804,1829]" italics="true" pageId="19" pageNumber="20">Coronella Rosenbergii</emphasis>
<bibRefCitation id="243672CEFFD4FFE5FD6EF94B0C25F8DB" author="Bleeker, P." box="[652,823,1804,1829]" pageId="19" pageNumber="20" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleeker 1860b</bibRefCitation>
:37
</taxonomicName>
. This species was named for the German naturalist Hermann von Rosenberg (
<date id="341929FFFFD4FFE5FE5FF9690D52F8B8" box="[445,576,1838,1862]" pageId="19" pageNumber="20" value="1817" valueMax="1888">18171888</date>
;
<figureCitation id="D89C13BAFFD4FFE5FDA9F96A0DB1F8B8" box="[587,675,1837,1862]" captionStart="FIGURE 3" captionStartId="9.[151,250,1487,1509]" captionTargetBox="[151,1436,193,1463]" captionTargetId="figure@9.[151,1436,193,1463]" captionTargetPageId="9" captionText="FIGURE 3. Selected 19 th Century authors and collectors of Stegonotus species. (A) Hermann Schlegel (18041884), author of the species name modestus. (B) André Marie Constant Duméril (17741860), co-author with (C) and (D) of the genus name Stegonotus and the species names cucullatus, lividus, and muelleri. (C) Gabriel Bibron (18061848). (D) Auguste Henri André Duméril (18121870). (E) René-Primevère Lesson (17941849), co-collector (with Prosper Garnot; no image available) of the L. cucullatum holotype. (F) Pieter van Oort (18041834), co-collector (with Salomon Müller and Heinrich Macklot; no images available) of the L. lividum syntypes. (G) Pieter Bleeker (18191878), author of the species names greineri (and perhaps collector of its holotype) and rosenbergii. (H) Hermann von Rosenberg (18171888), collector of the Coronella rosenbergii holotype and the person for whom the species was named. (I) Wilhelm Karl Hartwig Peters (18151883), author of the species name samarensis. (J) Fedor Jagor (18161900), collector of the Spilotes samarensis holotype. (K) Albert Karl Ludwig Gotthilf Günther (18301914), author of the genus name Zamenophis as well as the species names holochrous, batjanensis, and australis. The species Stegonotus guentheri was named in Günthers honor. (L) Adolf Bernhard Meyer (18401911), author of the species names magnus and parvus and collector of the type material." httpUri="https://zenodo.org/record/2607577/files/figure.png" pageId="19" pageNumber="20">Fig. 3H</figureCitation>
), who made an herpetological collection along the southern coast of Seram Island. Rosenberg was originally a cartographer and the Acting Civil Administrator of southern Seram (
<bibRefCitation id="243672CEFFD4FFE5FF7CF9370E5AF877" author="Bleeker, P." box="[158,328,1904,1929]" pageId="19" pageNumber="20" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleeker 1860b</bibRefCitation>
;
<bibRefCitation id="243672CEFFD4FFE5FEB1F9370D4DF877" author="Steenis-Kruseman, M. J. van" box="[339,607,1904,1929]" pageId="19" pageNumber="20" pagination="5 - 598" refId="ref46938" refString="Steenis-Kruseman, M. J. van (1950) Alphabetical list of collectors. In: Steenis, C. G. G. J. van (Ed.), Flora Malesiana. Series 1 - Spermatophyta. Folume 1, pp. 5 - 598." type="journal article" year="1950">Steenis-Kruseman 1950</bibRefCitation>
) but his interest in ornithology connected him to Hermann Schlegel at the RMNH in Leiden. The species description was presented in Dutch. Rosenbergs photograph (
<figureCitation id="D89C13BAFFD4FFE5FB05F9D60A54F854" box="[1255,1350,1937,1962]" captionStart="FIGURE 3" captionStartId="9.[151,250,1487,1509]" captionTargetBox="[151,1436,193,1463]" captionTargetId="figure@9.[151,1436,193,1463]" captionTargetPageId="9" captionText="FIGURE 3. Selected 19 th Century authors and collectors of Stegonotus species. (A) Hermann Schlegel (18041884), author of the species name modestus. (B) André Marie Constant Duméril (17741860), co-author with (C) and (D) of the genus name Stegonotus and the species names cucullatus, lividus, and muelleri. (C) Gabriel Bibron (18061848). (D) Auguste Henri André Duméril (18121870). (E) René-Primevère Lesson (17941849), co-collector (with Prosper Garnot; no image available) of the L. cucullatum holotype. (F) Pieter van Oort (18041834), co-collector (with Salomon Müller and Heinrich Macklot; no images available) of the L. lividum syntypes. (G) Pieter Bleeker (18191878), author of the species names greineri (and perhaps collector of its holotype) and rosenbergii. (H) Hermann von Rosenberg (18171888), collector of the Coronella rosenbergii holotype and the person for whom the species was named. (I) Wilhelm Karl Hartwig Peters (18151883), author of the species name samarensis. (J) Fedor Jagor (18161900), collector of the Spilotes samarensis holotype. (K) Albert Karl Ludwig Gotthilf Günther (18301914), author of the genus name Zamenophis as well as the species names holochrous, batjanensis, and australis. The species Stegonotus guentheri was named in Günthers honor. (L) Adolf Bernhard Meyer (18401911), author of the species names magnus and parvus and collector of the type material." httpUri="https://zenodo.org/record/2607577/files/figure.png" pageId="19" pageNumber="20">Fig. 3H</figureCitation>
; public domain) was taken by the photographer Conrad Spamer in 1882.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFD4FFE2FF25F9930DF4F9AA" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="40180F3FFFD4FFE5FF25F9930C8DF813" blockId="19.[151,1436,1737,2029]" box="[199,927,2004,2029]" pageId="19" pageNumber="20">
<materialsCitation id="F0CF0562FFD4FFE5FF25F9930C8DF813" ID-GBIF-Occurrence="2028140022" box="[199,927,2004,2029]" collectionCode="RMNH" pageId="19" pageNumber="20" specimenCode="RENA 4066" specimenCount="1" typeStatus="holotype">
<emphasis id="72D3D32DFFD4FFE5FF25F9930E29F813" bold="true" box="[199,315,2004,2029]" pageId="19" pageNumber="20">
<typeStatus id="9F1CB19DFFD4FFE5FF25F9930E24F813" box="[199,310,2004,2029]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="26B697FAFFD4FFE5FEA0F9920E8AF813" box="[322,408,2005,2029]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20" type="Museum">RMNH</collectionCode>
.
<specimenCode id="1001A744FFD4FFE5FE79F9920D3AF813" box="[411,552,2004,2029]" collectionCode="RENA" pageId="19" pageNumber="20">RENA 4066</specimenCode>
(
<figureCitation id="D89C13BAFFD4FFE5FDD5F9930D99F813" box="[567,651,2004,2029]" captionStart="FIGURE 10" captionStartId="20.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1050]" captionTargetId="figure@20.[151,1436,193,1050]" captionTargetPageId="20" captionText="FIGURE 10. Holotype of Coronella rosenbergii Bleeker, 1860b, RMNH.RENA 4066, an adult male from Seram Island, Maluku Province, Indonesia. Shown are the entire specimen (TTL = 760 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views." httpUri="https://zenodo.org/record/2607594/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
;
<tableCitation id="0D253A84FFD4FFE5FD74F9930DFAF813" box="[662,744,2004,2029]" captionStart="TABLE 1" captionStartId="60.[147,229,603,624]" captionText="TABLE 1. Name-bearing types of species in the genus Stegonotus. In this list we present basic data, including common measurements and selected scale counts, for all namebearing types (i.e., holotypes—HT, lectotypes—LT, neotypes—NT, syntypes—ST) to provide an easy overview of species. Additional details and the information for paratypes and paralectotypes can be found in the text. Character abbreviations are explained in the main text. For specimens where sex is indicated in red, we used the SCR value and not direct observation to estimate sex. Missing values are indicated by O. Specimen Type SCR SubC Species Synonym Sex SVL TL TTL V SC Dorsals SCE SL IL ILG Number Status (%) Condition LSUMZ admiraltiensis HT M 658 185 843 208 98 32 paired 17-19-15 4+5 8 10 10 93958 MSNG aruensis HT M 582 233 815 190 103 35 paired 17-17-15 3+4 7 8 4 30186 BMNH australis HT M 503 135 638 205 77 27 paired 17-17-15 4+5 8 9 5 1946.1.14.93 AMS australis darnleyensis HT F 720 O O 198 53+ O paired 17-17-15 4+5 9 10 5 R31894 AMS australis plumbeus HT M 1025 250 1275 220 74 25 paired 17-17-15 3+4+5 9 10 5 R31895 BMNH batjanensis HT M 1239 321 1560 235 86 27 paired 17-17-15 3+4+5 8 10 5 1946.1.11.36 FMNH borneensis HT F 783 221 1004 195 79 29 paired 17-17-15 4+5 10 11 5 164746 MNHN cucullatus HT M 769 254 1023 209 91 30 paired 17-17-15 4+5 8 9 5 3412 MTKD a cucullatus magnus ST F O O O 207 74 26 paired O-19-O 4+5 8 O O 573 ZMB b cucullatus magnus ST M 975 265 1240 211 87 29 paired 19-19-17 3+4 8 10 5 8794 MZB derooijae HT M 455 193 648 181 94 34 paired 17-17-15 3+4 7 8 8 Ophi.3288 MWNH diehli HT juv 196 56 252 179 82 31 paired 15-15-15 3+4 7 8 4 1244 NMW dorsalis HT M 805 245 1050 208 88 30 paired 15-15-15 4+5 8 10 5 14861 ZMA 9R florensis HT M 447 109 556 217 65 23 single 21-21-19 3+4+5 10 5 11080 8L BMNH guentheri LT F 683 165 848 192 64 25 paired 15-15-13 4+5 8 9 4 1946.1.11.38 ……continued on the next page" pageId="19" pageNumber="20">Table 1</tableCitation>
), an adult male.
</materialsCitation>
</paragraph>
<caption id="14D85FB7FFD3FFE2FF75FA680DBDFB7C" httpUri="https://zenodo.org/record/2607594/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1071,1093]" targetBox="[151,1436,193,1050]" targetPageId="20">
<paragraph id="40180F3FFFD3FFE2FF75FA680DBDFB7C" blockId="20.[151,1436,1071,1154]" pageId="20" pageNumber="21">
<emphasis id="72D3D32DFFD3FFE2FF75FA680E31FBBA" bold="true" box="[151,291,1071,1093]" pageId="20" pageNumber="21">FIGURE 10.</emphasis>
Holotype of
<taxonomicName id="87A774BCFFD3FFE2FE54FA680C51FBBB" authority="Bleeker, 1860 b" authorityName="Bleeker" authorityYear="1860" box="[438,835,1071,1093]" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rosenbergii">
<emphasis id="72D3D32DFFD3FFE2FE54FA680D8BFBBB" box="[438,665,1071,1093]" italics="true" pageId="20" pageNumber="21">Coronella rosenbergii</emphasis>
Bleeker, 1860b
</taxonomicName>
, RMNH.RENA 4066, an adult male from Seram Island, Maluku Province, Indonesia. Shown are the entire specimen (TTL = 760 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views.
</paragraph>
</caption>
<paragraph id="40180F3FFFD3FFE2FF25FA940B02FA5E" blockId="20.[151,1437,1235,2021]" pageId="20" pageNumber="21">
<emphasis id="72D3D32DFFD3FFE2FF25FA940E74FB12" bold="true" box="[199,358,1235,1260]" pageId="20" pageNumber="21">Type locality.</emphasis>
“Paulohi, Ceram, Molukschen Archipel” [Seram Island,
<collectingRegion id="8263C1DDFFD3FFE2FC0CFA940BA6FB12" box="[1006,1204,1235,1260]" country="Indonesia" name="Maluku Utara" pageId="20" pageNumber="21">Maluku Province</collectingRegion>
,
<collectingCountry id="38B04FAFFFD3FFE2FB22FA940A21FB12" box="[1216,1331,1235,1260]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
].
<bibRefCitation id="243672CEFFD3FFE2FAA7FA940FFDFAEE" author="Bleeker, P." pageId="20" pageNumber="21" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleeker (1860b)</bibRefCitation>
stated specifically that he considered this to be the first collection of reptiles from the southern coast of Seram. The locality name refers to a region in the south-central area of Seram inhabited by speakers of Paulohi, a rare Malayo-Polynesian language (
<bibRefCitation id="243672CEFFD3FFE2FDFDFB780DF4FAA9" author="Stresemann, E." box="[543,742,1343,1367]" pageId="20" pageNumber="21" refId="ref46988" refString="Stresemann, E. (1927) Die Lauterscheinungen in den ambonischen Sprachen. Zeitschrift fur Eingeborenen-Sprachen, Supplement, 10. Reimer, Berlin, Germany." type="book" year="1927">Stresemann 1927</bibRefCitation>
;
<bibRefCitation id="243672CEFFD3FFE2FD10FB070C6FFAA9" author="Wurm, S. A." box="[754,893,1343,1367]" pageId="20" pageNumber="21" pagination="425 - 578" refId="ref47623" refString="Wurm, S. A. (2007) Australasia and the Pacific. In: Moseley, C. (Ed.), Encyclopedia of the World's Endangered Languages. Routledge Publishers, London, United Kingdom, pp. 425 - 578." type="book chapter" year="2007">Wurm 2007</bibRefCitation>
). The most likely location for a collection in the mid-1800s is near the settlement of Amahai (ca.
<geoCoordinate id="259369F8FFD3FFE2FD54FB240C32FA82" box="[694,800,1379,1404]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="5" value="-3.3416">3.3416°S</geoCoordinate>
,
<geoCoordinate id="259369F8FFD3FFE2FCC9FB240CA7FA82" box="[811,949,1379,1404]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="5" value="128.9235">128.9235°E</geoCoordinate>
), a port facing
<collectingCountry id="38B04FAFFFD3FFE2FBB9FB240BA0FA82" box="[1115,1202,1379,1404]" name="Indonesia" pageId="20" pageNumber="21">Ambon</collectingCountry>
, the main population center of the region, which Bleeker visited on
<date id="341929FFFFD3FFE2FD7CFBC00C4BFA5E" box="[670,857,1415,1440]" pageId="20" pageNumber="21" value="1855-10-11">11 October 1855</date>
(
<bibRefCitation id="243672CEFFD3FFE2FC85FBC00B11FA5E" author="Bleeker, P." box="[871,1027,1415,1440]" pageId="20" pageNumber="21" refId="ref42155" refString="Bleeker, P. (1856) Reis door de Minahassa en de Molukschen Archipel, gedaan in de Maanden September en Oktober 1855 in het Gevolg van den Gouverneur Generaal Mr. A. J. Duymaer van Twist. Lange &amp; Co., Batavia [Jakarta, Indonesia], 350 pp." type="book" year="1856">Bleeker 1856</bibRefCitation>
).
</paragraph>
<paragraph id="40180F3FFFD3FFE2FF25FBEC0DF4F9AA" blockId="20.[151,1437,1235,2021]" pageId="20" pageNumber="21">
<emphasis id="72D3D32DFFD3FFE2FF25FBEC0E55FA3A" bold="true" box="[199,327,1451,1476]" pageId="20" pageNumber="21">Collection.</emphasis>
The specimen was collected in southern Seram by Rosenberg, for whom the species was named. Rosenberg was stationed on Seram from
<date id="341929FFFFD3FFE2FD8CFB880C02FA16" box="[622,784,1487,1512]" pageId="20" pageNumber="21" value="1859-04-26">26 April 1859</date>
<date id="341929FFFFD3FFE2FCFDFB880CE6FA16" box="[799,1012,1487,1512]" pageId="20" pageNumber="21" value="1861-12-17">17 December 1861</date>
(
<bibRefCitation id="243672CEFFD3FFE2FBE6FB880BD3FA16" author="Rosenberg, C. B. H. von" box="[1028,1217,1487,1512]" pageId="20" pageNumber="21" pagination="97 - 182" refId="ref46383" refString="Rosenberg, C. B. H. von (1867) Beschrijving van eenige gedeelten van Ceram. Tijdschrift voor Indische Taal-, Land- en Folkenkunde, 16, 97 - 182." type="journal article" year="1867">Rosenberg 1867</bibRefCitation>
;
<bibRefCitation id="243672CEFFD3FFE2FB2DFB880FC2F9F2" author="Steenis-Kruseman, M. J. van" pageId="20" pageNumber="21" pagination="5 - 598" refId="ref46938" refString="Steenis-Kruseman, M. J. van (1950) Alphabetical list of collectors. In: Steenis, C. G. G. J. van (Ed.), Flora Malesiana. Series 1 - Spermatophyta. Folume 1, pp. 5 - 598." type="journal article" year="1950">Steenis-Kruseman 1950</bibRefCitation>
) and, during that time, must have collected for Bleeker, whom he most likely met in Batavia before his deployment. Bleekers species description was dated
<date id="341929FFFFD3FFE2FD11F8500C63F9CE" box="[755,881,1559,1584]" pageId="20" pageNumber="21" value="1860-04">April 1860</date>
, which leads us to believe that the specimen was most likely collected during the second half of 1859.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFD3FFE2FF25F8180B1BF8F3" pageId="20" pageNumber="21" type="key">
<paragraph id="40180F3FFFD3FFE2FF25F8180D16F917" blockId="20.[151,1437,1235,2021]" pageId="20" pageNumber="21">
<emphasis id="72D3D32DFFD3FFE2FF25F8180D78F986" bold="true" box="[199,618,1631,1656]" pageId="20" pageNumber="21">
Key characteristics of the
<materialsCitation id="F0CF0562FFD3FFE2FE1EF8180D78F986" ID-GBIF-Occurrence="2028140012" box="[508,618,1631,1656]" collectionCode="SVL" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
<typeStatus id="9F1CB19DFFD3FFE2FE1EF8180D77F986" box="[508,613,1631,1656]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</emphasis>
O (582) mm SVL + O (178) mm TL = O (760) mm TTL.
<materialsCitation id="F0CF0562FFD3FFE2FB1FF8270E67F960" ID-GBIF-Occurrence="2028140021" collectionCode="SCR" pageId="20" pageNumber="21" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
V
<subScript id="DC230D7AFFD3FFE2FAF3F82A0A0FF985" attach="left" box="[1297,1309,1645,1659]" fontSize="6" pageId="20" pageNumber="21">
<specimenCount id="56A1C4B6FFD3FFE2FAF3F82A0A0FF985" box="[1297,1309,1645,1659]" pageId="20" pageNumber="21" type="male"></specimenCount>
</subScript>
= O (203), SC
<subScript id="DC230D7AFFD3FFE2FF5BF8D30FD7F95C" attach="left" box="[185,197,1684,1698]" fontSize="6" pageId="20" pageNumber="21">
<specimenCount id="56A1C4B6FFD3FFE2FF5BF8D30FD7F95C" box="[185,197,1684,1698]" pageId="20" pageNumber="21" type="male"></specimenCount>
</subScript>
= O (80),
<collectionCode id="26B697FAFFD3FFE2FEA2F8C10E67F960" box="[320,373,1670,1694]" country="USA" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography, University of California" pageId="20" pageNumber="21" type="Herbarium">SCR</collectionCode>
</materialsCitation>
<materialsCitation id="F0CF0562FFD3FFE2FE94F8D30B51F960" ID-GBIF-Occurrence="2028140023" box="[374,1091,1669,1698]" collectionCode="IL" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<subScript id="DC230D7AFFD3FFE2FE94F8D30E90F95C" attach="left" box="[374,386,1684,1698]" fontSize="6" pageId="20" pageNumber="21">
<specimenCount id="56A1C4B6FFD3FFE2FE94F8D30E90F95C" box="[374,386,1684,1698]" pageId="20" pageNumber="21" type="male"></specimenCount>
</subScript>
= O (0.28), D = O (17-17-15), SL
<subScript id="DC230D7AFFD3FFE2FCECF8D30C05F95C" attach="left" box="[782,791,1684,1698]" fontSize="6" pageId="20" pageNumber="21">E</subScript>
= O (3+4), SL = 7 (7),
<collectionCode id="26B697FAFFD3FFE2FBCAF8C00B51F960" box="[1064,1091,1671,1694]" pageId="20" pageNumber="21">IL</collectionCode>
</materialsCitation>
= O (9), IL
<subScript id="DC230D7AFFD3FFE2FB2EF8D30BC5F95C" attach="left" box="[1228,1239,1684,1698]" fontSize="6" pageId="20" pageNumber="21">G</subScript>
= O (4).
<bibRefCitation id="243672CEFFD3FFE2FAA7F8C20FFDF93B" author="Bleeker, P." pageId="20" pageNumber="21" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleeker (1860b)</bibRefCitation>
remarked that the tail was roughly one quarter the length of the entire body, which fits well with our measurements (TL/TTL = 0.23).
</paragraph>
<paragraph id="40180F3FFFD3FFE2FF25F8B30B1BF8F3" blockId="20.[151,1437,1235,2021]" box="[199,1033,1780,1805]" pageId="20" pageNumber="21">
<emphasis id="72D3D32DFFD3FFE2FF25F8B30D45F8F3" bold="true" box="[199,599,1780,1805]" pageId="20" pageNumber="21">Key characteristics of the species.</emphasis>
See the account of
<taxonomicName id="87A774BCFFD3FFE2FCD1F8B20CA5F8F3" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[819,951,1780,1805]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="modestus">
<emphasis id="72D3D32DFFD3FFE2FCD1F8B20CA5F8F3" box="[819,951,1780,1805]" italics="true" pageId="20" pageNumber="21">S. modestus</emphasis>
</taxonomicName>
above.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFD3FFE3FF25F95F0B4BFE2E" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="40180F3FFFD3FFE3FF25F95F0B4BFE2E" blockId="20.[151,1437,1235,2021]" lastBlockId="21.[151,1436,151,464]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="72D3D32DFFD3FFE2FF25F95F0E5DF8CF" bold="true" box="[199,335,1816,1841]" pageId="20" pageNumber="21">Comments.</emphasis>
<bibRefCitation id="243672CEFFD3FFE2FEBEF95F0D08F8CF" author="Bleeker, P." box="[348,538,1816,1841]" pageId="20" pageNumber="21" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleeker (1860b)</bibRefCitation>
specifically mentioned that the specimen had “een flaauwe gele dwarsche nekband” [a faint yellow cross neckband], which is a key characteristic of
<taxonomicName id="87A774BCFFD3FFE2FC0AF97A0B7CF8AB" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1000,1134,1852,1877]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="modestus">
<emphasis id="72D3D32DFFD3FFE2FC0AF97A0B7CF8AB" box="[1000,1134,1852,1877]" italics="true" pageId="20" pageNumber="21">S. modestus</emphasis>
</taxonomicName>
. In his treatment of
<collectingRegion id="8263C1DDFFD3FFE2FAB5F97A0A8EF8AA" box="[1367,1436,1853,1876]" country="Indonesia" name="Papua Barat" pageId="20" pageNumber="21">Papua</collectingRegion>
New Guinean
<taxonomicName id="87A774BCFFD3FFE2FEDEF9260D83F887" authority=", McDowell (1972)" authorityName=", McDowell" authorityYear="1972" box="[316,657,1888,1913]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFD3FFE2FEDEF9260EA4F886" box="[316,438,1889,1912]" italics="true" pageId="20" pageNumber="21">Stegonotus</emphasis>
,
<bibRefCitation id="243672CEFFD3FFE2FE24F9270D83F887" author="McDowell, S. B." box="[454,657,1888,1913]" pageId="20" pageNumber="21" pagination="6 - 26" refId="ref45102" refString="McDowell, S. B. (1972) The species of Stegonotus (Serpentes, Colubridae) in Papua New Guinea. Zoologische Mededelingen, 47 (2), 6 - 26." type="journal article" year="1972">McDowell (1972)</bibRefCitation>
</taxonomicName>
included a footnote below his account for
<taxonomicName id="87A774BCFFD3FFE2FB9DF9260A17F887" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1151,1285,1888,1913]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="modestus">
<emphasis id="72D3D32DFFD3FFE2FB9DF9260A17F887" box="[1151,1285,1888,1913]" italics="true" pageId="20" pageNumber="21">S. modestus</emphasis>
</taxonomicName>
, in which he stated that “Bleeker based his description [of
<taxonomicName id="87A774BCFFD3FFE2FD78F9C20C29F863" authorityName="Bleeker" authorityYear="1860" box="[666,827,1924,1949]" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rosenbergii">
<emphasis id="72D3D32DFFD3FFE2FD78F9C20C29F863" box="[666,827,1924,1949]" italics="true" pageId="20" pageNumber="21">C. rosenbergii</emphasis>
</taxonomicName>
] on two specimens,” one from the museum in Leiden (RMNH 4066) and one from London (BMNH 1946.1.13.88). McDowell further opined that
<bibRefCitation id="243672CEFFD3FFE2FAF6F9EF0E01F81B" author="Boulenger, G. A." pageId="20" pageNumber="21" refId="ref42396" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Nat. Hist.) Folume I. Containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae, part. Taylor &amp; Francis, London, United Kingdom, 448 pp." type="book" year="1893">Boulengers (1893:367)</bibRefCitation>
listing of the latter as “Type of
<taxonomicName id="87A774BCFFD3FFE2FD66F98A0C37F81B" authorityName="Bleeker" authorityYear="1860" box="[644,805,1996,2021]" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rosenbergii">
<emphasis id="72D3D32DFFD3FFE2FD66F98A0C37F81B" box="[644,805,1996,2021]" italics="true" pageId="20" pageNumber="21">C. rosenbergii</emphasis>
</taxonomicName>
” constituted a
<typeStatus id="9F1CB19DFFD3FFE2FC31F98B0B2BF81B" box="[979,1081,1996,2021]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
designation. We carefully read
<bibRefCitation id="243672CEFFD2FFE3FF75FED00E76FF4E" author="Bleeker, P." box="[151,356,151,176]" pageId="21" pageNumber="22" pagination="35 - 38" refId="ref42237" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article" year="1860">Bleekers (1860b)</bibRefCitation>
species description and found no indication that Bleeker had two specimens in hand when he described the species. Furthermore, whereas Bleekers type locality is very specific, the one provided by
<bibRefCitation id="243672CEFFD2FFE3FACAFEFB0FF3FF06" author="Boulenger, G. A." pageId="21" pageNumber="22" refId="ref42396" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Nat. Hist.) Folume I. Containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae, part. Taylor &amp; Francis, London, United Kingdom, 448 pp." type="book" year="1893">Boulenger (1893)</bibRefCitation>
for BMNH 1946.1.13.88 is vague, listing merely “Ceram.” Lastly, the original collection number of the BMNH specimen is 63.12.4.9, indicating that the specimen reached London with a shipment from Bleeker in 1863, three years after the original description was published. Our interpretation of the
<emphasis id="72D3D32DFFD2FFE3FBDEFF600B65FEBE" box="[1084,1143,295,320]" italics="true" pageId="21" pageNumber="22">Code</emphasis>
leads us to conclude that
<bibRefCitation id="243672CEFFD2FFE3FF75FF0B0E60FE9A" author="Boulenger, G. A." box="[151,370,332,357]" pageId="21" pageNumber="22" refId="ref42396" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Nat. Hist.) Folume I. Containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae, part. Taylor &amp; Francis, London, United Kingdom, 448 pp." type="book" year="1893">Boulengers (1893)</bibRefCitation>
notation that BMNH 1946.1.13.88 (formerly 63.12.4.9) should be considered a
<typeStatus id="9F1CB19DFFD2FFE3FAE1FF0A0A48FE9B" box="[1283,1370,333,357]" pageId="21" pageNumber="22" type="syntype">syntype</typeStatus>
of
<taxonomicName id="87A774BCFFD2FFE3FA60FF0A0E04FE76" authorityName="Bleeker" authorityYear="1860" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="rosenbergii">
<emphasis id="72D3D32DFFD2FFE3FA60FF0A0E04FE76" italics="true" pageId="21" pageNumber="22">C. rosenbergii</emphasis>
</taxonomicName>
is not supported by evidence in the original description or from the specimen itself. We therefore disagree with
<bibRefCitation id="243672CEFFD2FFE3FED8FFD30D14FE52" author="McDowell, S. B." box="[314,518,404,429]" pageId="21" pageNumber="22" pagination="6 - 26" refId="ref45102" refString="McDowell, S. B. (1972) The species of Stegonotus (Serpentes, Colubridae) in Papua New Guinea. Zoologische Mededelingen, 47 (2), 6 - 26." type="journal article" year="1972">McDowell (1972)</bibRefCitation>
and hereby invalidate the purported
<typeStatus id="9F1CB19DFFD2FFE3FC51FFD30B0BFE53" box="[947,1049,404,429]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
designation (
<emphasis id="72D3D32DFFD2FFE3FB55FFD30BE0FE53" box="[1207,1266,404,429]" italics="true" pageId="21" pageNumber="22">Code</emphasis>
, Article 74.2). There is but a single type specimen of
<taxonomicName id="87A774BCFFD2FFE3FDA5FFFF0DF5FE2E" authorityName="Bleeker" authorityYear="1860" box="[583,743,439,464]" class="Reptilia" family="Colubridae" genus="Coronella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="rosenbergii">
<emphasis id="72D3D32DFFD2FFE3FDA5FFFF0DF5FE2E" box="[583,743,439,464]" italics="true" pageId="21" pageNumber="22">C. rosenbergii</emphasis>
</taxonomicName>
, and it is the
<typeStatus id="9F1CB19DFFD2FFE3FC9CFFF00CF2FE2E" box="[894,992,439,464]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
in Leiden.
</paragraph>
</subSubSection>
</treatment>
</document>