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151 lines
18 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zse.96.47792" ID-GBIF-Dataset="46dee71c-5ee5-4a2b-9ca9-c6ec9c73f07c" ID-GBIF-Taxon="182236570" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="A8EDB50ED5705717A80AD87E91F4AE96" ID-ZooBank="C5560B3FB5A0462380EBFE2C483D4774" ModsDocID="1860-0743-96-1-1" checkinTime="1621423348088" checkinUser="pensoft" docAuthor="Tripathy, Basudev, Sajan, Sheikh & Cowie, Robert H." docDate="2019" docId="0B9876B9B61F5FFFB0FF7D225C8A09FC" docLanguage="en" docName="ZoosystEvol 96(1): 1-23" docOrigin="Zoosystematics and Evolution 96 (1)" docPubDate="2019-01-14" docSource="http://dx.doi.org/10.3897/zse.96.47792" docTitle="Pachylabra nevilliana Annandale & Prashad 1921" docType="treatment" docVersion="3" id="A8EDB50ED5705717A80AD87E91F4AE96" lastPageNumber="1" masterDocId="A8EDB50ED5705717A80AD87E91F4AE96" masterDocTitle="Illustrated catalogue of types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the National Zoological Collection of the Zoological Survey of India, with lectotype designations" masterLastPageNumber="23" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1643454254309" updateUser="ExternalLinkService">
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<mods:title>Illustrated catalogue of types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the National Zoological Collection of the Zoological Survey of India, with lectotype designations</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
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<mods:affiliation>Malacology Division, Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sajan, Sheikh</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
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<mods:affiliation>Malacology Division, Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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<mods:nameIdentifier type="email">sksajan.sajan@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cowie, Robert H.</mods:namePart>
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<mods:affiliation>Pacific Biosciences Research Center, University of Hawaii, 3050 Maile Way, Gilmore 408, Honolulu, Hawaii 96822, USA</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2019-01-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>23</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.96.47792</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.47792</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-1</mods:identifier>
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<mods:identifier type="ZooBank">C5560B3FB5A0462380EBFE2C483D4774</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A8EDB50ED5705717A80AD87E91F4AE96</mods:identifier>
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<treatment ID-GBIF-Taxon="182236570" LSID="urn:lsid:plazi:treatment:0B9876B9B61F5FFFB0FF7D225C8A09FC" httpUri="http://treatment.plazi.org/id/0B9876B9B61F5FFFB0FF7D225C8A09FC" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="0B9876B9-B61F-5FFF-B0FF-7D225C8A09FC" authority="Annandale & Prashad, 1921" authorityName="Annandale & Prashad" authorityYear="1921" class="Gastropoda" family="Ampullariidae" genus="Pachylabra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pachylabra nevilliana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nevilliana">Pachylabra nevilliana Annandale & Prashad, 1921</taxonomicName>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Holotype of Pachylabra nevilliana Annandale & Prashad, 1921; NZSI M. 11864 / 2." figureDoi="10.3897/zse.96.47792.figure9" httpUri="https://binary.pensoft.net/fig/371736" pageId="0" pageNumber="1">Figs 9</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Collection labels. A. Pachylabra nevilliana Annandale & Prashad, 1921; NZSI M. 11864 / 2. B. Pila robsoni Prashad, 1925; NZSI M. 2414 (holotype), NZSI M. 21546 / 4 (paratypes)." figureDoi="10.3897/zse.96.47792.figure15" httpUri="https://binary.pensoft.net/fig/371742" pageId="0" pageNumber="1">, 15A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="" authorityName="Annandale & Prashad" authorityYear="1921" class="Gastropoda" family="Ampullariidae" genus="Pachylabra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pachylabra nevilliana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nevilliana">Pachylabra nevilliana</taxonomicName>
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Annandale & Prashad, 1921.
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<emphasis italics="true" pageId="0" pageNumber="1">Rec. Ind. Mus.</emphasis>
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22 (1): 11-12.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="current taxonomic status">
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<paragraph pageId="0" pageNumber="1">Current taxonomic status.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Annandale & Prashad" baseAuthorityYear="1921" class="Gastropoda" family="Ampullariidae" genus="Pila" kingdom="Animalia" lsidName="Pila nevilliana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nevilliana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pila nevilliana</emphasis>
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</taxonomicName>
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(Annandale & Prashad, 1921), valid species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">"Tranquebar, which is on the east coast of Southern India" [= Tharangambadi, Nagapattinam district, Tamil Nadu, India]; Captain Lewis, leg. (Asiatic Society of Bengal collection).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype (original designation): NZSI M.11864/2 (registered 8 December 1920). Paratypes: NZSI M.25078/5 (registered 13 September 2001, 5 spms), 2 or 3 additional specimens noted by (
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<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill 1877</bibRefCitation>
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: 5, 1885: 4), location unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="shell dimensions">
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<paragraph pageId="0" pageNumber="1">Shell dimensions.</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype: SH 36.1 mm, SW 34.3 mm, AH 27.9 mm, AW 19.6 mm, OH 25.6 mm, OW 16.0 mm. Paratypes: SH 28.5-37.1 mm, SW 26.3-33.5 mm, AH 22.3-28.5 mm, AW 17.3-22.6 mm, OH 20.5-25.5 mm, OW 11.9-15.0 mm</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Annandale, N" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="1" pagination="7 - 12" publicationUrl="https://biodiversitylibrary.org/page/11201530" refId="B2" refString="Annandale, N, Prashad, B, 1921. Materials for a generic revision of the freshwater gastropod molluscs of the Indian Empire. No. 4. The Indian Ampullariidae. Records of the Indian Museum 22 (1): 7 - 12, https://biodiversitylibrary.org/page/11201530" title="Materials for a generic revision of the freshwater gastropod molluscs of the Indian Empire. No. 4. The Indian Ampullariidae." url="https://biodiversitylibrary.org/page/11201530" volume="22" year="1921">Annandale and Prashad (1921</bibRefCitation>
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: 12) explicitly based their description on only the specimens noted by
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<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill (1877</bibRefCitation>
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: 5) as Var. B of
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<taxonomicName authorityName="Reeve" authorityYear="1856" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" kingdom="Animalia" lsidName="Ampullaria nux" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nux">
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<emphasis italics="true" pageId="0" pageNumber="1">Ampullaria nux</emphasis>
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</taxonomicName>
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Reeve, 1856. However, although
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<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill (1877</bibRefCitation>
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: 5) noted eight specimens and
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.23978" author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" refId="B22" refString="Nevill, G, 1885. Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [=Kolkata], x + 306 pp. https://doi.org/10.5962/bhl.title.23978" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [= Kolkata], x + 306 pp." url="https://doi.org/10.5962/bhl.title.23978" year="1885">Nevill (1885</bibRefCitation>
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: 4) noted nine,
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<bibRefCitation author="Annandale, N" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="1" pagination="7 - 12" publicationUrl="https://biodiversitylibrary.org/page/11201530" refId="B2" refString="Annandale, N, Prashad, B, 1921. Materials for a generic revision of the freshwater gastropod molluscs of the Indian Empire. No. 4. The Indian Ampullariidae. Records of the Indian Museum 22 (1): 7 - 12, https://biodiversitylibrary.org/page/11201530" title="Materials for a generic revision of the freshwater gastropod molluscs of the Indian Empire. No. 4. The Indian Ampullariidae." url="https://biodiversitylibrary.org/page/11201530" volume="22" year="1921">Annandale and Prashad (1921</bibRefCitation>
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: 12) provided measurements for only three. Nonetheless, they probably saw all
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<normalizedToken originalValue="Nevill’s">Nevill's</normalizedToken>
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shells, as they noted that they had "extracted the radula from one of the shells examined by Nevill, which have been in Calcutta for at least 60 years, but still in several instances contain the dried animal"; the expression "several instances" suggests that they saw more than the three for which they provided measurements. The specimen figured by
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<bibRefCitation author="Annandale, N" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="1" pagination="7 - 12" publicationUrl="https://biodiversitylibrary.org/page/11201530" refId="B2" refString="Annandale, N, Prashad, B, 1921. Materials for a generic revision of the freshwater gastropod molluscs of the Indian Empire. No. 4. The Indian Ampullariidae. Records of the Indian Museum 22 (1): 7 - 12, https://biodiversitylibrary.org/page/11201530" title="Materials for a generic revision of the freshwater gastropod molluscs of the Indian Empire. No. 4. The Indian Ampullariidae." url="https://biodiversitylibrary.org/page/11201530" volume="22" year="1921">Annandale and Prashad (1921</bibRefCitation>
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: fig. 2) as the
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<normalizedToken originalValue="“type-shell”">"type-shell"</normalizedToken>
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(= holotype) is the specimen in NZSI M.11864/2; this specimen matches the illustrations in all regards, including, the erosion/corrosion of the apex, wear of the periostracum and appearance of the operculum. Its measurements are within the likely margin of error of those given by Annandale and Prashad (1920: 12, middle column). Annandale and Prashad (1920: 11) based their description only on
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<normalizedToken originalValue="Nevill’s">Nevill's</normalizedToken>
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shells, either eight (
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<bibRefCitation author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" publicationUrl="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" refId="B21" refString="Nevill, G, 1877. Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp. http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" title="Catalogue of Mollusca in the Indian Museum, Calcutta. Fasciculus E. Indian Museum, Calcutta [= Kolkata], [4] + 42 pp." url="http://media.api.aucklandmuseum.com/id/media/p/1bf9a093da0f3bd874f966ce7d0483bf7904668f?rendering=original.pdf" year="1877">Nevill 1877</bibRefCitation>
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: 5) or nine (
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.23978" author="Nevill, G" journalOrPublisher="Archives of Natural History" pageId="0" pageNumber="1" refId="B22" refString="Nevill, G, 1885. Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [=Kolkata], x + 306 pp. https://doi.org/10.5962/bhl.title.23978" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). Indian Museum, Calcutta [= Kolkata], x + 306 pp." url="https://doi.org/10.5962/bhl.title.23978" year="1885">Nevill 1885</bibRefCitation>
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: 4). They gave only a single catalogue number (11864/2) for the
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<normalizedToken originalValue="“Type-specimens”">"Type-specimens"</normalizedToken>
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(plural). This lot now contains only the holotype. An additional lot, NZSI M.25078/5, labelled as paratypes contains five shells.
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3940.1.1" author="Cowie, RH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 92" refId="B8" refString="Cowie, RH, 2015. The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda. Zootaxa 3940 (1): 1 - 92, DOI: https://doi.org/10.11646/zootaxa.3940.1.1" title="The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda." url="https://doi.org/10.11646/zootaxa.3940.1.1" volume="3940" year="2015">Cowie (2015</bibRefCitation>
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: 43) considered these as only "possible paratypes". However, further study indicates that they can indeed be considered part of the type series because 1) the dimensions of at least two of them approximate the additional dimensions given by Annandale and Prashad; 2)
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<normalizedToken originalValue="“Tranquebar”">"Tranquebar"</normalizedToken>
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is written on the shells; and 3)
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<normalizedToken originalValue="“type”">"type"</normalizedToken>
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is written in the original register alongside the entry for this lot.
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</paragraph>
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<caption doi="10.3897/zse.96.47792.figure9" httpUri="https://binary.pensoft.net/fig/371736" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
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Holotype of
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<taxonomicName authority="" authorityName="Annandale & Prashad" authorityYear="1921" class="Gastropoda" family="Ampullariidae" genus="Pachylabra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pachylabra nevilliana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nevilliana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pachylabra nevilliana</emphasis>
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</taxonomicName>
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Annandale & Prashad, 1921; NZSI M.11864/2.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |