Illustrated catalogue of types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the National Zoological Collection of the Zoological Survey of India, with lectotype designations
Author
Tripathy, Basudev
Malacology Division, Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India
Author
Sajan, Sheikh
https://orcid.org/0000-0002-2785-6824
Malacology Division, Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India
sksajan.sajan@gmail.com
Author
Cowie, Robert H.
Pacific Biosciences Research Center, University of Hawaii, 3050 Maile Way, Gilmore 408, Honolulu, Hawaii 96822, USA
text
Zoosystematics and Evolution
2019
2019-01-14
96
1
1
23
http://dx.doi.org/10.3897/zse.96.47792
journal article
http://dx.doi.org/10.3897/zse.96.47792
1860-0743-1-1
C5560B3FB5A0462380EBFE2C483D4774
A8EDB50ED5705717A80AD87E91F4AE96
Pachylabra nevilliana Annandale & Prashad, 1921
Figs 9
, 15A
Pachylabra nevilliana
Annandale & Prashad, 1921.
Rec. Ind. Mus.
22 (1): 11-12.
Current taxonomic status.
Pila nevilliana
(Annandale & Prashad, 1921), valid species.
Type locality.
"Tranquebar, which is on the east coast of Southern India" [= Tharangambadi, Nagapattinam district, Tamil Nadu, India]; Captain Lewis, leg. (Asiatic Society of Bengal collection).
Type material.
Holotype (original designation): NZSI M.11864/2 (registered 8 December 1920). Paratypes: NZSI M.25078/5 (registered 13 September 2001, 5 spms), 2 or 3 additional specimens noted by (
Nevill 1877
: 5, 1885: 4), location unknown.
Shell dimensions.
Holotype: SH 36.1 mm, SW 34.3 mm, AH 27.9 mm, AW 19.6 mm, OH 25.6 mm, OW 16.0 mm. Paratypes: SH 28.5-37.1 mm, SW 26.3-33.5 mm, AH 22.3-28.5 mm, AW 17.3-22.6 mm, OH 20.5-25.5 mm, OW 11.9-15.0 mm
Remarks.
Annandale and Prashad (1921
: 12) explicitly based their description on only the specimens noted by
Nevill (1877
: 5) as Var. B of
Ampullaria nux
Reeve, 1856. However, although
Nevill (1877
: 5) noted eight specimens and
Nevill (1885
: 4) noted nine,
Annandale and Prashad (1921
: 12) provided measurements for only three. Nonetheless, they probably saw all
Nevill's
shells, as they noted that they had "extracted the radula from one of the shells examined by Nevill, which have been in Calcutta for at least 60 years, but still in several instances contain the dried animal"; the expression "several instances" suggests that they saw more than the three for which they provided measurements. The specimen figured by
Annandale and Prashad (1921
: fig. 2) as the
"type-shell"
(= holotype) is the specimen in NZSI M.11864/2; this specimen matches the illustrations in all regards, including, the erosion/corrosion of the apex, wear of the periostracum and appearance of the operculum. Its measurements are within the likely margin of error of those given by Annandale and Prashad (1920: 12, middle column). Annandale and Prashad (1920: 11) based their description only on
Nevill's
shells, either eight (
Nevill 1877
: 5) or nine (
Nevill 1885
: 4). They gave only a single catalogue number (11864/2) for the
"Type-specimens"
(plural). This lot now contains only the holotype. An additional lot, NZSI M.25078/5, labelled as paratypes contains five shells.
Cowie (2015
: 43) considered these as only "possible paratypes". However, further study indicates that they can indeed be considered part of the type series because 1) the dimensions of at least two of them approximate the additional dimensions given by Annandale and Prashad; 2)
"Tranquebar"
is written on the shells; and 3)
"type"
is written in the original register alongside the entry for this lot.
Figure 9.
Holotype of
Pachylabra nevilliana
Annandale & Prashad, 1921; NZSI M.11864/2.