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<mods:title id="1EA8630DC98980524EA78F2084497DE7">Kasimovian (late Pennsylvanian) cornute rugose corals from Egypt: taxonomy, facies and palaeogeography of a cool-water fauna from northern Gondwana</mods:title>
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<subSubSection id="898B8B8B7F16FFF7FC91FC92FB32FC0F" box="[813,1232,876,901]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="C12ED8007F16FFF7FC91FC92FB32FC0F" blockId="15.[813,1437,876,1477]" box="[813,1232,876,901]" pageId="15" pageNumber="16">
<heading id="9A666F6C7F16FFF7FC91FC92FB32FC0F" box="[813,1232,876,901]" fontSize="36" level="2" pageId="15" pageNumber="16" reason="3">
Genus
<taxonomicName id="0691A3837F16FFF7FCC5FC93FB28FC0F" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" box="[889,1226,876,901]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FCC5FC93FBF8FC0F" bold="true" box="[889,1050,876,900]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FB9DFC92FB28FC0F" author="Fomichev, V." box="[1057,1226,876,901]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFF7FC91FC72FB49FC6F" pageId="15" pageNumber="16" type="type_taxon">
<paragraph id="C12ED8007F16FFF7FC91FC72FA94FC2F" blockId="15.[813,1437,876,1477]" box="[813,1398,908,933]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FC91FC72FC24FC2F" bold="true" box="[813,966,908,933]" pageId="15" pageNumber="16">
<typeStatus id="1E2A66A27F16FFF7FC91FC72FC84FC2E" box="[813,870,909,933]" pageId="15" pageNumber="16">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0691A3837F16FFF7FC70FC72FA8DFC2F" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[972,1391,908,933]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F16FFF7FC70FC72FB22FC2E" box="[972,1216,909,933]" italics="true" pageId="15" pageNumber="16">Monophyllum sokolovi</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FB7AFC72FA8DFC2F" author="Fomichev, V." box="[1222,1391,908,933]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C12ED8007F16FFF7FC91FC52FB49FC6F" blockId="15.[813,1437,876,1477]" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FC91FC52FBB5FC4F" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[813,1111,940,965]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F16FFF7FC91FC52FCA7FC4F" box="[813,837,941,964]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="F3E504127F16FFF7FCEDFC52FC45FC4E" box="[849,935,941,965]" italics="true" pageId="15" pageNumber="16">sokolovi</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FC12FC52FBB5FC4F" author="Fomichev, V." box="[942,1111,940,965]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
;?
<bibRefCitation id="A500A5F17F16FFF7FBC9FC53FB4BFC4F" author="Fomichev, V." box="[1141,1193,940,964]" pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">1953</bibRefCitation>
M.
<taxonomicName id="0691A3837F16FFF7FB69FC52FC83FC6F" authority="Fomichev 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="parvum">
<emphasis id="F3E504127F16FFF7FB69FC52FACEFC4F" box="[1237,1324,941,964]" italics="true" pageId="15" pageNumber="16">parvum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FA8EFC52FC83FC6F" author="Fomichev, V." pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev 1953</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0691A3837F16FFF7FCD1FC32FB47FC6E" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" box="[877,1189,972,997]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F16FFF7FCD1FC32FC67FC6F" box="[877,901,973,996]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="F3E504127F16FFF7FC2DFC32FC03FC6F" box="[913,993,973,996]" italics="true" pageId="15" pageNumber="16">cassum</emphasis>
Fedorowski, 1987
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFF7FC91FC13FB26FB4E" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="C12ED8007F16FFF7FC91FC13FB26FB4E" blockId="15.[813,1437,876,1477]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FC91FC13FC44FB8F" bold="true" box="[813,934,1004,1028]" pageId="15" pageNumber="16">Diagnosis:</emphasis>
Non-dissepimented, small solitary corals, with major septa radially arranged, may withdraw a little from corallite axis in maturity; cardinal septum shortened from early maturity; counter septum permanently elongated to corallite axis; alar septa slightly elongated at least in some portion of growth; minor septa underdeveloped (after
<bibRefCitation id="A500A5F17F16FFF7FC16FB52FB97FB4E" author="Fedorowski, J." box="[938,1141,1197,1221]" pageId="15" pageNumber="16" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
, p. 67).
</paragraph>
</subSubSection>
<subSubSection id="898B8B8B7F16FFE9FC91FB32FE7AFB6E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="discussion">
<paragraph id="C12ED8007F16FFF7FC91FB32FAADFA4E" blockId="15.[813,1437,876,1477]" pageId="15" pageNumber="16">
<emphasis id="F3E504127F16FFF7FC91FB32FC78FB6E" bold="true" box="[813,922,1229,1253]" pageId="15" pageNumber="16">Remarks:</emphasis>
In contrast to
<bibRefCitation id="A500A5F17F16FFF7FBE5FB32FB2FFB6F" author="Hill, D." box="[1113,1229,1228,1253]" pageId="15" pageNumber="16" refId="ref26422" refString="Hill, D. (1956). Rugosa. In R. C. Moore (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata (pp. F 233 - F 324). New York." type="journal volume" year="1956">Hill (1956</bibRefCitation>
,
<bibRefCitation id="A500A5F17F16FFF7FB5DFB32FAF7FB6E" author="Hill, D." box="[1249,1301,1229,1253]" pageId="15" pageNumber="16" refId="ref26465" refString="Hill, D. (1981). Supplement 1, Rugosa and Tabulata. In C. Teichert (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata, F 1 - F 762. Boulder / Colorado and Lawrence / Kansas. (Geological Society of America and University of Kansas Press)." type="journal volume" year="1981">1981</bibRefCitation>
), de Groot (1963) and
<bibRefCitation id="A500A5F17F16FFF7FC17FB12FBA2FA8E" author="Weyer, D." box="[939,1088,1260,1285]" pageId="15" pageNumber="16" pagination="755 - 775" refId="ref30369" refString="Weyer, D. (1975). Zur Taxonomie der Antiphyllinae Ilina, 1970. Zeitschrift Der Geologischen Wissenschaften, 3 (6), 755 - 775." type="journal article" year="1975">Weyer (1975)</bibRefCitation>
who listed
<taxonomicName id="0691A3837F16FFF7FB7EFB12FC4AFAAF" authority="Fomichev, 1953" authorityName="Fomichev, Shortened" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB7EFB12FAB8FA8E" box="[1218,1370,1261,1285]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FADEFB12FC4AFAAF" author="Fomichev, V." pageId="15" pageNumber="16" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
as a synonym of
<taxonomicName id="0691A3837F16FFF7FBD3FAF2FA74FAAE" authority="Hudson, 1942" authorityName="Hudson" authorityYear="1942" box="[1135,1430,1293,1317]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FBD3FAF2FB16FAAE" box="[1135,1268,1293,1317]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
<bibRefCitation id="A500A5F17F16FFF7FB41FAF2FA74FAAE" author="Hudson, R. G. S." box="[1277,1430,1293,1317]" pageId="15" pageNumber="16" pagination="257 - 263" refId="ref26658" refString="Hudson, R. G. S. (1942). Fasciculophyllum Thomson and other genera of the &quot; Zaphrentis &quot; omaliusi group of Carboniferous corals. Geological Magazine, 79 (5), 257 - 263." type="journal article" year="1942">Hudson, 1942</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="A500A5F17F16FFF7FC91FAD2FBEAFACE" author="Fedorowski, J." box="[813,1032,1325,1349]" pageId="15" pageNumber="16" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
considered it as a separate genus. This view was followed by
<bibRefCitation id="A500A5F17F16FFF7FBEAFAB2FAEEFAEF" author="Kossovaya, O. L." box="[1110,1292,1356,1381]" pageId="15" pageNumber="16" pagination="71 - 82" refId="ref27287" refString="Kossovaya, O. L. (1996). Correlation of uppermost Carboniferous and Lower Permian Rugose Coral Zones from the Urals to Western North America. Palaios, 11 (1), 71 - 82." type="journal article" year="1996">Kossovaya (1996</bibRefCitation>
, 2001),
<bibRefCitation id="A500A5F17F16FFF7FAD9FAB2FC91FA0E" author="Ogar, V. V." pageId="15" pageNumber="16" pagination="340 - 349" refId="ref28259" refString="Ogar, V. V. (2012). Carboniferous buildups in the Donets Basin (Ukraine). Geologica Belgica, 15 (4), 340 - 349." type="journal article" year="2012">Ogar (2012)</bibRefCitation>
and
<bibRefCitation id="A500A5F17F16FFF7FC17FA92FB95FA0E" author="Wang, X. D. &amp; Yao, L. &amp; Lin, W." box="[939,1143,1389,1413]" pageId="15" pageNumber="16" pagination="165 - 184" refId="ref30038" refString="Wang, X. D., Yao, L., &amp; Lin, W. (2017). Permian rugose corals of the world. In S. G. Lucas, &amp; S. Z. Shen (Eds.), The Permian timescale. Geological Society, London, Special Publications. 450 (2018), 165 - 184." type="journal article" year="2017">Wang et al. (2017)</bibRefCitation>
.
<bibRefCitation id="A500A5F17F16FFF7FB3FFA92FAB9FA0E" author="Fedorowski, J." box="[1155,1371,1389,1413]" pageId="15" pageNumber="16" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski (1987a)</bibRefCitation>
mentioned the following differences to
<taxonomicName id="0691A3837F16FFF7FB08FA72FADBFA2E" authorityName="Hudson" authorityYear="1942" box="[1204,1337,1421,1445]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB08FA72FADBFA2E" box="[1204,1337,1421,1445]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
</taxonomicName>
and further similar antiphyllinid genera (see also
<tableCitation id="8C13EDBB7F16FFF7FB4EFA52FAA2FA4E" box="[1266,1344,1453,1477]" captionStart="Table 1" captionStartId="7.[151,209,684,707]" captionTargetPageId="7" captionText="Table 1 Morphological comparison of genera in Antiphyllinae Ilina, 1970 present in the Aheimer Formation, western side of Gulf of Suez, Egypt including Bradyphyllum Grabau, 1928 for comparison with morphologically close Lytvolasma Soshkina, (based on Fedorowski, 1987a and Chwieduk,2013)" pageId="15" pageNumber="16">Table 1</tableCitation>
):
</paragraph>
<paragraph id="C12ED8007F16FFF7FCFEFA11FB68F96D" blockId="15.[834,1437,1518,1894]" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FCDEFA11FC1AF98D" authorityName="Fomichev, Shortened" authorityYear="1953" box="[866,1016,1518,1542]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FCDEFA11FC1AF98D" box="[866,1016,1518,1542]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="0691A3837F16FFF7FB28FA11FAF5F98D" authorityName="Hudson" authorityYear="1942" box="[1172,1303,1518,1542]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB28FA11FAF5F98D" box="[1172,1303,1518,1542]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
</taxonomicName>
by: (1) the strongly and permanently elongated counter septum reaching the corallite axis, even on the calice floor, where it may form an incipient columella, (2) shortening of the cardinal septum not only on the calice wall as in
<taxonomicName id="0691A3837F16FFF7FC77F971FBACF92D" authorityName="Hudson" authorityYear="1942" box="[971,1102,1678,1702]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FC77F971FBACF92D" box="[971,1102,1678,1702]" italics="true" pageId="15" pageNumber="16">Rotiphyllum</emphasis>
</taxonomicName>
, but also on the calice floor, (3) the triangular shape of at least peripheral parts of the cardinal fossula at maturity.
</paragraph>
<paragraph id="C12ED8007F16FFE9FCFEF911FE58FEE8" blockId="15.[834,1437,1518,1894]" lastBlockId="17.[173,775,235,547]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<taxonomicName id="0691A3837F16FFF7FCDEF911FC1AF88D" authorityName="Fomichev, Shortened" authorityYear="1953" box="[866,1016,1774,1798]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FCDEF911FC1AF88D" box="[866,1016,1774,1798]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="0691A3837F16FFF7FB3AF911FAFCF88D" authorityName="Grabau" authorityYear="1928" box="[1158,1310,1774,1798]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FB3AF911FAFCF88D" box="[1158,1310,1774,1798]" italics="true" pageId="15" pageNumber="16">Bradyphyllum</emphasis>
</taxonomicName>
by the permanently elongated counter septum that reaches the corallite axis and tends to form an incipient columella. Although
<taxonomicName id="0691A3837F16FFF7FBA0F8B1FB50F8ED" authorityName="Fomichev, Shortened" authorityYear="1953" box="[1052,1202,1870,1894]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F16FFF7FBA0F8B1FB50F8ED" box="[1052,1202,1870,1894]" italics="true" pageId="15" pageNumber="16">Monophyllum</emphasis>
</taxonomicName>
develops a tendency to form an axial area free of septa, like most of the genera included in
<taxonomicName id="0691A3837F08FFE9FE12FEF4FDDDFEA8" box="[430,575,267,291]" class="Anthozoa" family="Antiphyllidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="35" phylum="Cnidaria" rank="subFamily" subFamily="Antiphyllinae">Antiphyllinae</taxonomicName>
(Illina, 1970), its counter septum remains elongated, which is not the case in
<taxonomicName id="0691A3837F08FFE9FEA0FEB4FE56FEE8" authorityName="Grabau" authorityYear="1928" box="[284,436,331,355]" class="Anthozoa" family="Antiphyllidae" genus="Bradyphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FEA0FEB4FE56FEE8" box="[284,436,331,355]" italics="true" pageId="17" pageNumber="18">Bradyphyllum</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="95EE88887F09FFE8FF1BF9F8FD24F880" ID-DOI="http://doi.org/10.5281/zenodo.12003288" ID-Zenodo-Dep="12003288" httpUri="https://zenodo.org/record/12003288/files/figure.png" pageId="16" pageNumber="17" startId="16.[167,198,1543,1563]" targetBox="[291,1292,238,1527]" targetPageId="16" targetType="figure">
<paragraph id="C12ED8007F09FFE8FF1BF9F8FD24F880" blockId="16.[167,1422,1543,1803]" pageId="16" pageNumber="17">
<emphasis id="F3E504127F09FFE8FF1BF9F8FF10F990" bold="true" box="[167,242,1543,1563]" pageId="16" pageNumber="17">Fig. 9 A</emphasis>
<taxonomicName id="0691A3837F09FFE8FF4BF9F8FE28F990" authority="El-Desouky &amp; Herbig &amp; Kora, 2023" authorityName="El-Desouky &amp; Herbig &amp; Kora" authorityYear="2023" box="[247,458,1543,1563]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Cnidaria" rank="species" species="galalaensis" status="sp. nov.">
<emphasis id="F3E504127F09FFE8FF4BF9F8FE28F990" box="[247,458,1543,1563]" italics="true" pageId="16" pageNumber="17">Monophyllum galalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8D6B9697F09FFE8FE72F9F8FE19F990" box="[462,507,1543,1563]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
(Holotype specimen, Rah 9).
<emphasis id="F3E504127F09FFE8FD4CF9F8FCEBF990" bold="true" box="[752,777,1543,1563]" pageId="16" pageNumber="17">A1</emphasis>
Lateral external view of the corallite, positions of thin-sections indicated in A1.
<emphasis id="F3E504127F09FFE8FF67F9DDFF16F9BD" bold="true" box="[219,244,1570,1590]" pageId="16" pageNumber="17">A2</emphasis>
A calice view of the paratype specimen (RAh 41), with eroded rim, shows an elongated counter septum that form a columella (arrow).
<emphasis id="F3E504127F09FFE8FF1BF9C2FF06F9DA" bold="true" box="[167,228,1597,1617]" pageId="16" pageNumber="17">A3A5</emphasis>
Successive transverse thin-sections of the neanic stage,showing the pseudo-radial to pinnate arrangement of major septa, a long thin ended cardinal septum, and a longer counter septum with a swollen end in a dense stereocolumn.
<emphasis id="F3E504127F09FFE8FC4DF9A8FBC3F9E0" bold="true" box="[1009,1057,1623,1643]" pageId="16" pageNumber="17">A67</emphasis>
Two successive mature sections, showing a short cardinal septum.
<emphasis id="F3E504127F09FFE8FEC4F98DFE4AF9F7" bold="true" box="[376,424,1646,1670]" pageId="16" pageNumber="17">
A8
<superScript id="36E475487F09FFE8FE2DF991FE4AF9F7" attach="left" box="[401,424,1646,1660]" fontSize="6" pageId="16" pageNumber="17">1, 2</superScript>
</emphasis>
Transverse thin-section in the late mature stage below the calice,showing radially arranged major septa that retreat a little leaving a narrow free axial area occupied by the elongated part of the counter septum. Note,
<emphasis id="F3E504127F09FFE8FC49F972FBF4F91C" bold="true" box="[1013,1046,1673,1697]" pageId="16" pageNumber="17">
A8
<superScript id="36E475487F09FFE8FBB2F976FBF4F91C" attach="left" box="[1038,1046,1673,1687]" fontSize="6" pageId="16" pageNumber="17">2</superScript>
</emphasis>
is a redrawing of section
<emphasis id="F3E504127F09FFE8FB52F972FAEDF91C" bold="true" box="[1262,1295,1673,1697]" pageId="16" pageNumber="17">
A8
<superScript id="36E475487F09FFE8FABBF976FAEDF91C" attach="none" box="[1287,1295,1673,1687]" fontSize="6" pageId="16" pageNumber="17">1</superScript>
</emphasis>
. Paratype specimen (RAh 13). External alar view of the corallite,positions of thin-sections are indicated in white lines.
<emphasis id="F3E504127F09FFE8FB93F958FBA5F930" bold="true" box="[1071,1095,1703,1723]" pageId="16" pageNumber="17">B2</emphasis>
Nepionic stage.
<emphasis id="F3E504127F09FFE8FB6FF958FB09F930" bold="true" box="[1235,1259,1703,1723]" pageId="16" pageNumber="17">B3</emphasis>
Early neanic stage.
<emphasis id="F3E504127F09FFE8FF1BF93DFF34F95D" bold="true" box="[167,214,1730,1750]" pageId="16" pageNumber="17">B45</emphasis>
Two partly compressed successive mature sections (ephebic stage), show a distinctly long counter septum.
<emphasis id="F3E504127F09FFE8FBD5F93DFB63F95D" bold="true" box="[1129,1153,1730,1750]" pageId="16" pageNumber="17">B6</emphasis>
Last mature section—more or less in the calice—showing a detached columella that is separated from the long counter septum. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa
</paragraph>
</caption>
<paragraph id="C12ED8007F08FFE9FF11FE93FD36FDA8" blockId="17.[173,775,235,547]" pageId="17" pageNumber="18">
<taxonomicName id="0691A3837F08FFE9FF71FE94FE81FE08" authorityName="Fomichev, Shortened" authorityYear="1953" box="[205,355,363,387]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FF71FE94FE81FE08" box="[205,355,363,387]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="0691A3837F08FFE9FE49FE94FEE2FE28" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE49FE94FD73FE08" box="[501,657,363,387]" italics="true" pageId="17" pageNumber="18">Actinophrentis</emphasis>
<bibRefCitation id="A500A5F17F08FFE9FD26FE94FEE2FE28" author="Fomichev, V." pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
by (1) the radial arrangement of septa in the mature portions of
<taxonomicName id="0691A3837F08FFE9FE0AFE54FDAEFE48" authorityName="Fomichev, Shortened" authorityYear="1953" box="[438,588,427,451]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE0AFE54FDAEFE48" box="[438,588,427,451]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
vs. the pinnate arrangement of the major septa lasting up to the calice floor in
<taxonomicName id="0691A3837F08FFE9FEE8FE14FE12FD88" authorityName="Fomichev" authorityYear="1953" box="[340,496,491,515]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FEE8FE14FE12FD88" box="[340,496,491,515]" italics="true" pageId="17" pageNumber="18">Actinophrentis</emphasis>
</taxonomicName>
and (2) the lacking tendency of
<taxonomicName id="0691A3837F08FFE9FE92FDF4FE28FDA8" authorityName="Fomichev" authorityYear="1953" box="[302,458,523,547]" class="Anthozoa" family="Antiphyllidae" genus="Actinophrentis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE92FDF4FE28FDA8" box="[302,458,523,547]" italics="true" pageId="17" pageNumber="18">Actinophrentis</emphasis>
</taxonomicName>
to form a free axial area.
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BFDB2FD6DFD4F" blockId="17.[151,776,588,1253]" pageId="17" pageNumber="18">
Species described by
<bibRefCitation id="A500A5F17F08FFE9FE24FDB2FDB4FDEE" author="Fomichev, V." box="[408,598,588,613]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev (1953)</bibRefCitation>
to be “
<taxonomicName id="0691A3837F08FFE9FD07FDB2FF38FD0E" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Metriophyllidae" genus="Stereolasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FD07FDB2FF38FD0E" italics="true" pageId="17" pageNumber="18">Stereolasma</emphasis>
</taxonomicName>
” might belong to
<taxonomicName id="0691A3837F08FFE9FE0EFD92FDD5FD0E" authorityName="Hudson" authorityYear="1942" box="[434,567,621,645]" class="Anthozoa" family="Metriophyllidae" genus="Rotiphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE0EFD92FDD5FD0E" box="[434,567,621,645]" italics="true" pageId="17" pageNumber="18">Rotiphyllum</emphasis>
</taxonomicName>
(in toto, see
<bibRefCitation id="A500A5F17F08FFE9FD64FD92FF29FD2F" author="Hill, D." pageId="17" pageNumber="18" refId="ref26422" refString="Hill, D. (1956). Rugosa. In R. C. Moore (Ed.), Treatise on Invertebrate Paleontology, Part F, Coelenterata (pp. F 233 - F 324). New York." type="journal volume" year="1956">Hill, 1956</bibRefCitation>
; de Groot, 1963), to
<taxonomicName id="0691A3837F08FFE9FE6CFD72FD8AFD2E" authorityName="Fomichev, Shortened" authorityYear="1953" box="[464,616,653,677]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FE6CFD72FD8AFD2E" box="[464,616,653,677]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
and/or other Antiphyllinidae (
<bibRefCitation id="A500A5F17F08FFE9FEF1FD52FDFBFD4E" author="Fedorowski, J." box="[333,537,685,709]" pageId="17" pageNumber="18" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
, p. 36, 68).
</paragraph>
<paragraph id="C12ED8007F08FFE9FF2BFD33FE7AFB6E" blockId="17.[151,776,588,1253]" pageId="17" pageNumber="18">
<emphasis id="F3E504127F08FFE9FF2BFD33FDDEFD6E" bold="true" box="[151,572,716,741]" pageId="17" pageNumber="18">Geographic and stratigraphic range:</emphasis>
<taxonomicName id="0691A3837F08FFE9FDFFFD32FD39FD6E" authorityName="Fomichev, Shortened" authorityYear="1953" box="[579,731,717,741]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="genus">
<emphasis id="F3E504127F08FFE9FDFFFD32FD39FD6E" box="[579,731,717,741]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
</taxonomicName>
is a long ranging but relatively rare genus. It is known from the Bashkirian of the Donets Basin (
<bibRefCitation id="A500A5F17F08FFE9FDF2FCF2FCE2FCAF" author="Fomichev, V." box="[590,768,780,805]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
;
<bibRefCitation id="A500A5F17F08FFE9FF2BFCD2FEB3FCCF" author="Kossovaya, O. L." box="[151,337,812,837]" pageId="17" pageNumber="18" pagination="187 - 199" refId="ref27258" refString="Kossovaya, O. L. (1986). The mid-Carboniferous rugose coral recovery. Geological Society London, Special Publication, 102, 187 - 199." type="journal article" year="1986">Kossovaya, 1986</bibRefCitation>
,
<figureCitation id="59AAC4857F08FFE9FED9FCD3FE4AFCCF" box="[357,424,812,836]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
;
<bibRefCitation id="A500A5F17F08FFE9FE06FCD3FD9DFCCE" author="Fedorowski, J." box="[442,639,812,837]" pageId="17" pageNumber="18" pagination="247 - 316" refId="ref25858" refString="Fedorowski, J. (2022). Bashkirian Rugosa (Anthozoa) from the Donets Basin (Ukraine). Part 12. Concluding considerations. Acta Geologica Polonica, 72 (3), 247 - 316." type="journal article" year="2022">Fedorowski, 2022</bibRefCitation>
,
<figureCitation id="59AAC4857F08FFE9FD2FFCD3FD35FCCE" box="[659,727,812,837]" captionStart="Fig" captionStartId="3.[167,198,1519,1539]" captionTargetBox="[286,1300,239,1503]" captionTargetId="figure-143@3.[283,1303,236,1506]" captionTargetPageId="3" captionText="Fig. 2 A Lithostratigraphic succession of the Aheimer Formation studied along the eastern cliffs of the Northern Galala Plateau after Kora and Mansour (1992). B Enlargement of the mudstone dominated basal shale part of the lower member of the Aheimer Formation studied herein (see also Fig. 3A)" figureDoi="http://doi.org/10.5281/zenodo.12003190" httpUri="https://zenodo.org/record/12003190/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
:
<taxonomicName id="0691A3837F08FFE9FD55FCD2FE4BFCEF" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F08FFE9FD55FCD2FCE3FCCF" box="[745,769,813,836]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="F3E504127F08FFE9FF2BFCB3FF0FFCEF" box="[151,237,844,868]" italics="true" pageId="17" pageNumber="18">sokolovi</emphasis>
<bibRefCitation id="A500A5F17F08FFE9FF45FCB3FE4BFCEF" author="Fomichev, V." box="[249,425,844,868]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0691A3837F08FFE9FE07FCB2FD1AFCEF" authority="Fomichev, 1953" authorityName="Fomichev" authorityYear="1953" box="[443,760,844,868]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="parvum">
<emphasis id="F3E504127F08FFE9FE07FCB2FE31FCEF" box="[443,467,845,868]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="F3E504127F08FFE9FE5AFCB2FDDFFCEF" box="[486,573,845,868]" italics="true" pageId="17" pageNumber="18">parvum</emphasis>
<bibRefCitation id="A500A5F17F08FFE9FDF5FCB3FD1AFCEF" author="Fomichev, V." box="[585,760,844,868]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
</taxonomicName>
), the Moscovian of the same region (
<bibRefCitation id="A500A5F17F08FFE9FDA8FC93FD5CFC0F" author="Fomichev, V." box="[532,702,876,900]" pageId="17" pageNumber="18" refId="ref26079" refString="Fomichev, V. (1953). Rugose corals and stratigraphy of Middle and Upper Carboniferous and Middle Permian deposits of the Donets Basin. GeoI. Inst. (VESEGEI), 622 pp." type="book" year="1953">Fomichev, 1953</bibRefCitation>
: original designation of the
<typeStatus id="1E2A66A27F08FFE9FE26FC72FE2BFC2E" box="[410,457,909,933]" pageId="17" pageNumber="18">type</typeStatus>
species
<taxonomicName id="0691A3837F08FFE9FD91FC72FD49FC2E" authorityName="Fomichev" authorityYear="1953" box="[557,683,909,933]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="sokolovi">
<emphasis id="F3E504127F08FFE9FD91FC72FD49FC2E" box="[557,683,909,933]" italics="true" pageId="17" pageNumber="18">M. sokolovi</emphasis>
</taxonomicName>
;
<bibRefCitation id="A500A5F17F08FFE9FD00FC72FEEDFC4E" author="Kossovaya, O. L." pageId="17" pageNumber="18" pagination="187 - 199" refId="ref27258" refString="Kossovaya, O. L. (1986). The mid-Carboniferous rugose coral recovery. Geological Society London, Special Publication, 102, 187 - 199." type="journal article" year="1986">Kossovaya, 1986</bibRefCitation>
,
<figureCitation id="59AAC4857F08FFE9FE9EFC52FE86FC4F" box="[290,356,941,965]" captionStart="Fig" captionStartId="12.[167,198,1342,1362]" captionTargetBox="[285,1299,238,1327]" captionTargetId="figure-189@12.[283,1303,236,1329]" captionTargetPageId="12" captionText="Fig. 7 Lytvolasma aheimerensis n. sp. A External view of the specimen RA (2), showing a straight corallite with partly eroded calice and non-preserved apical part, positions of thin-sections indicated.BD Successive transverse thin-sections in the early preserved growth stages, shows very thick major septa joining together near the centre.The cardinal septum is deformed in D. EG Successive thin-sections in the early mature stages show the formation of a stereozone with a free axial area,shortening of the cardinal septum with maturity and the longest counter septum with a rhopaloid end. HI Two transverse thin-sections below the calice, deformed due to rejuvenation in H and lateral compaction in I. Scale bars: 2.5 mm, except A: 5 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003254" httpUri="https://zenodo.org/record/12003254/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
;
<bibRefCitation id="A500A5F17F08FFE9FECAFC52FE10FC4E" author="Ogar, V. V." box="[374,498,941,965]" pageId="17" pageNumber="18" pagination="340 - 349" refId="ref28259" refString="Ogar, V. V. (2012). Carboniferous buildups in the Donets Basin (Ukraine). Geologica Belgica, 15 (4), 340 - 349." type="journal article" year="2012">Ogar, 2012</bibRefCitation>
) and the Moscovian of Novaya Zemlya (fide
<bibRefCitation id="A500A5F17F08FFE9FE3BFC32FDA5FC6E" author="Fedorowski, J." box="[391,583,973,997]" pageId="17" pageNumber="18" pagination="87 - 160" refId="ref25455" refString="Fedorowski, J. (1981). Carboniferous corals: Distribution and sequence. Acta Palaeontologica Polonica, 26 (2), 87 - 160." type="journal article" year="1981">Fedorowski, 1981</bibRefCitation>
and
<bibRefCitation id="A500A5F17F08FFE9FD3EFC32FF29FB8E" author="Wang, X. D. &amp; Yang, S. R. &amp; Yao, L. &amp; Sugiyama, T. &amp; Hu, K. Y." pageId="17" pageNumber="18" pagination="603 - 63" refId="ref29952" refString="Wang, X. D., Yang, S. R., Yao, L., Sugiyama, T., &amp; Hu, K. Y. (2022). Carboniferous biostratigraphy of rugose corals. In S. G. Lucas, J. W. Schneider, X. Wang, S. Nikolaeva (Eds), The Carboniferous timescale. Geological Society, London, Special Publications. 512 (2022). 603 - 63." type="journal article" year="2022">Wang et al., 2022</bibRefCitation>
). It is also known from the PennsylvanianPermian transition and the Wolfcampian (Asselianmiddle Artinskian) of southwest Texas (
<bibRefCitation id="A500A5F17F08FFE9FE07FBD2FD64FBCE" author="Fedorowski, J." box="[443,646,1069,1093]" pageId="17" pageNumber="18" pagination="1 - 271" refId="ref25513" refString="Fedorowski, J. (1987 a). Upper Palaeozoic rugose corals from southwestern Texas and adjacent areas: Gaptank Formation and Wolfcampian corals, Part 1. Palaeontologica Polonica, 48, 1 - 271." type="journal article" year="1987">Fedorowski, 1987a</bibRefCitation>
:
<taxonomicName id="0691A3837F08FFE9FD2FFBD2FEB7FBEE" authority="Fedorowski, 1987" authorityName="Fedorowski" authorityYear="1987" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="cassum">
<emphasis id="F3E504127F08FFE9FD2FFBD2FD49FBCF" box="[659,683,1069,1092]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="F3E504127F08FFE9FD0BFBD2FCE5FBCF" box="[695,775,1069,1092]" italics="true" pageId="17" pageNumber="18">cassum</emphasis>
Fedorowski, 1987
</taxonomicName>
). Kossovaya et al., (2001,
<figureCitation id="59AAC4857F08FFE9FDD2FBB2FD49FBEE" box="[622,683,1101,1125]" captionStart="Fig" captionStartId="9.[167,198,1520,1540]" captionTargetBox="[287,1299,246,1507]" captionTargetId="figure-67@9.[283,1303,236,1508]" captionTargetPageId="9" captionText="Fig. 6 A Actinophrentis crassithecata n. sp.: (Holotype specimen RAh (57). A1 External view showing a thick walled, slightly curved corallite with septal furrows ornamenting the external wall. Position of thin-sections indicated. A2 Transverse thin-section near the non-preserved apical part, shows thick major septa meeting in the centre and adaxially thinning cardinal septum. A3 Transverse thin-section in the middle part of the corallite, shows major septa meeting in the centre and noticeable fossular breaks. A4 Last adult transverse thin section just below the calice shows septa withdrawn a little from the centre.A5 Details of the external wall of A3. B1, B2 External lateral views of the specimen RAh (11) showing a thick walled,slightly curved corallite with septal furrows and concentric growth lines on the external wall. C Lytvolasma cf. canadense Fedorowski and Bamber,2001, specimen RAh (66). C1, C2 Transverse thin-section in the middle mature part of the corallite, shows a short cardinal septum and a long counter septum with a rhopaloid axial end. C3 Transverse thin-section in the calice, shows a long counter septum. Scale bars: 2.5 mm, except A2A5: 2 mm. Black dots indicate the positions of cardinal septum (below), counter septum (above) and the two alar septa" figureDoi="http://doi.org/10.5281/zenodo.12003244" httpUri="https://zenodo.org/record/12003244/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
) figured?
<taxonomicName id="0691A3837F08FFE9FF1DFB92FE83FB0F" box="[161,353,1133,1157]" class="Anthozoa" family="Hapsiphyllidae" genus="Monophyllum" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F3E504127F08FFE9FF1DFB92FEDBFB0E" box="[161,313,1133,1157]" italics="true" pageId="17" pageNumber="18">Monophyllum</emphasis>
sp.
</taxonomicName>
from the late Artinskian of the Urals and/or northern Timan Ridge (not further specified). Herein, we report the genus from the Kasimovian west of the Gulf of
<collectingRegion id="035516E27F08FFE9FEAEFB32FEAAFB6E" box="[274,328,1229,1253]" country="Egypt" name="As Suways" pageId="17" pageNumber="18">Suez</collectingRegion>
,
<collectingCountry id="B98698907F08FFE9FEEEFB32FE76FB6E" box="[338,404,1229,1253]" name="Egypt" pageId="17" pageNumber="18">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>